taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
B3264F4BFF82704EADB9FCD7FA5FFBFC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673340/files/figure.png	https://doi.org/10.5281/zenodo.6673340	FIGURE 2. Tityus (Atreus) forcipula (Gervais, 1843), habitus. A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. Scale bar= 10 mm.	FIGURE 2. Tityus (Atreus) forcipula (Gervais, 1843), habitus. A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. Scale bar= 10 mm.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF82704EADB9FCD7FA5FFBFC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673342/files/figure.png	https://doi.org/10.5281/zenodo.6673342	FIGURE 3. Tityus (Atreus) moralensis sp. nov., habitus. A, B. Male holotype (MUSENUV-Ar 2104). A. Dorsal view. B. Ventral view. C, D. Female paratype (ICN-1402). C. Dorsal view. D. Ventral view. Scale bar= 15 mm.	FIGURE 3. Tityus (Atreus) moralensis sp. nov., habitus. A, B. Male holotype (MUSENUV-Ar 2104). A. Dorsal view. B. Ventral view. C, D. Female paratype (ICN-1402). C. Dorsal view. D. Ventral view. Scale bar= 15 mm.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF82704EADB9FCD7FA5FFBFC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673360/files/figure.png	https://doi.org/10.5281/zenodo.6673360	FIGURE 9. Tityus (Atreus) Gervais, 1843, pectinal area. A, B. Tityus (Atreus) forcipula (Gervais, 1843).A. Male (MUSENUV- Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402).	FIGURE 9. Tityus (Atreus) Gervais, 1843, pectinal area. A, B. Tityus (Atreus) forcipula (Gervais, 1843).A. Male (MUSENUV- Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402).	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF82704EADB9FCD7FA5FFBFC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673368/files/figure.png	https://doi.org/10.5281/zenodo.6673368	FIGURE 12. Tityus (Atreus) Gervais, 1843, metasoma, dorsal view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm.	FIGURE 12. Tityus (Atreus) Gervais, 1843, metasoma, dorsal view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF82704EADB9FCD7FA5FFBFC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673372/files/figure.png	https://doi.org/10.5281/zenodo.6673372	FIGURE 13. Tityus (Atreus) Gervais, 1843, metasoma, ventral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm.	FIGURE 13. Tityus (Atreus) Gervais, 1843, metasoma, ventral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF82704EADB9FCD7FA5FFBFC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673376/files/figure.png	https://doi.org/10.5281/zenodo.6673376	FIGURE 15. Tityus (Atreus) Gervais, 1843, telson, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 1.5 mm.	FIGURE 15. Tityus (Atreus) Gervais, 1843, telson, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 1.5 mm.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF8D7051ADB9FB73FC47F816.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673334/files/figure.png	https://doi.org/10.5281/zenodo.6673334	FIGURE 1. Phylogenetic relationships of Tityus species inferred from a combined dataset of 28S and COI sequences. The topology is recovered from the IQ-TREE analysis. Squares at nodes indicate the Shimodaira–Hasegawa approximate likelihood-ratio test (left), the ultrafast bootstrap (middle) and the parsimony bootstrap (right). All sequences obtained in this study are in bold.	FIGURE 1. Phylogenetic relationships of Tityus species inferred from a combined dataset of 28S and COI sequences. The topology is recovered from the IQ-TREE analysis. Squares at nodes indicate the Shimodaira–Hasegawa approximate likelihood-ratio test (left), the ultrafast bootstrap (middle) and the parsimony bootstrap (right). All sequences obtained in this study are in bold.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF8D7051ADB9FB73FC47F816.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673340/files/figure.png	https://doi.org/10.5281/zenodo.6673340	FIGURE 2. Tityus (Atreus) forcipula (Gervais, 1843), habitus. A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. Scale bar= 10 mm.	FIGURE 2. Tityus (Atreus) forcipula (Gervais, 1843), habitus. A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. Scale bar= 10 mm.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF8D7051ADB9FB73FC47F816.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673348/files/figure.png	https://doi.org/10.5281/zenodo.6673348	FIGURE 4. Tityus (Atreus) Gervais, 1843, carapace sculpturing, dorsal aspect. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 2.5 mm.	FIGURE 4. Tityus (Atreus) Gervais, 1843, carapace sculpturing, dorsal aspect. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 2.5 mm.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF8D7051ADB9FB73FC47F816.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673350/files/figure.png	https://doi.org/10.5281/zenodo.6673350	FIGURE 5. Tityus (Atreus) Gervais, 1843, femur sculpturing. A–D. Tityus (Atreus) forcipula (Gervais, 1843). A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. E–H. Tityus (Atreus) moralensis sp. nov. E, F. Male holotype (MUSENUV-Ar 2104). E. Dorsal view. F. Ventral view. G, H. Female paratype (ICN-1402). G. Dorsal view. H. Ventral view. Scale bars= 2 mm.	FIGURE 5. Tityus (Atreus) Gervais, 1843, femur sculpturing. A–D. Tityus (Atreus) forcipula (Gervais, 1843). A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. E–H. Tityus (Atreus) moralensis sp. nov. E, F. Male holotype (MUSENUV-Ar 2104). E. Dorsal view. F. Ventral view. G, H. Female paratype (ICN-1402). G. Dorsal view. H. Ventral view. Scale bars= 2 mm.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF8D7051ADB9FB73FC47F816.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673354/files/figure.png	https://doi.org/10.5281/zenodo.6673354	FIGURE 6. Tityus (Atreus) Gervais, 1843, patella sculpturing. A–D. Tityus (Atreus) forcipula (Gervais, 1843). A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. E–H. Tityus (Atreus) moralensis sp. nov. E, F. Male holotype (MUSENUV-Ar 2104). E. Dorsal view. F. Ventral view. G, H. Female paratype (ICN-1402). G. Dorsal view. H. Ventral view. Scale bars= 2 mm.	FIGURE 6. Tityus (Atreus) Gervais, 1843, patella sculpturing. A–D. Tityus (Atreus) forcipula (Gervais, 1843). A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. E–H. Tityus (Atreus) moralensis sp. nov. E, F. Male holotype (MUSENUV-Ar 2104). E. Dorsal view. F. Ventral view. G, H. Female paratype (ICN-1402). G. Dorsal view. H. Ventral view. Scale bars= 2 mm.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF8D7051ADB9FB73FC47F816.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673356/files/figure.png	https://doi.org/10.5281/zenodo.6673356	FIGURE 7. Tityus (Atreus) Gervais, 1843,female chela sculpturing.A–C. Tityus(Atreus) forcipula (Gervais,1843) (MUSENUV- Ar 436). A. External view. B. Dorsal view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (ICN-1402). D. External view. E. Dorsal view. F. Ventral view. Scale bars= 3 mm.	FIGURE 7. Tityus (Atreus) Gervais, 1843,female chela sculpturing.A–C. Tityus(Atreus) forcipula (Gervais,1843) (MUSENUV- Ar 436). A. External view. B. Dorsal view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (ICN-1402). D. External view. E. Dorsal view. F. Ventral view. Scale bars= 3 mm.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF8D7051ADB9FB73FC47F816.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673358/files/figure.png	https://doi.org/10.5281/zenodo.6673358	FIGURE 8. Tityus (Atreus) Gervais, 1843, male chela sculpturing. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. External view. B. Dorsal view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. External view. E. Dorsal view. F. Ventral view. Scale bars= 3 mm.	FIGURE 8. Tityus (Atreus) Gervais, 1843, male chela sculpturing. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. External view. B. Dorsal view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. External view. E. Dorsal view. F. Ventral view. Scale bars= 3 mm.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF8D7051ADB9FB73FC47F816.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673360/files/figure.png	https://doi.org/10.5281/zenodo.6673360	FIGURE 9. Tityus (Atreus) Gervais, 1843, pectinal area. A, B. Tityus (Atreus) forcipula (Gervais, 1843).A. Male (MUSENUV- Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402).	FIGURE 9. Tityus (Atreus) Gervais, 1843, pectinal area. A, B. Tityus (Atreus) forcipula (Gervais, 1843).A. Male (MUSENUV- Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402).	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF8D7051ADB9FB73FC47F816.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673364/files/figure.png	https://doi.org/10.5281/zenodo.6673364	FIGURE 10. Tityus (Atreus) Gervais, 1843, sternites IV–VII. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402).	FIGURE 10. Tityus (Atreus) Gervais, 1843, sternites IV–VII. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402).	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF8D7051ADB9FB73FC47F816.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673366/files/figure.png	https://doi.org/10.5281/zenodo.6673366	FIGURE 11. Tityus (Atreus) Gervais, 1843, hemispermatophore. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. Posterior view. B. Lateral view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. Posterior view. E. Lateral view. F. Ventral view. Abbreviations= B, body; Bl, basal lobe; El, external lobe; F, flagellum; Fo, foot; iL, internal lobe; Pref, pars reflexa; Prec, pars recta. Scale bars= 2 mm. Scale bars= 2 mm.	FIGURE 11. Tityus (Atreus) Gervais, 1843, hemispermatophore. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. Posterior view. B. Lateral view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. Posterior view. E. Lateral view. F. Ventral view. Abbreviations= B, body; Bl, basal lobe; El, external lobe; F, flagellum; Fo, foot; iL, internal lobe; Pref, pars reflexa; Prec, pars recta. Scale bars= 2 mm. Scale bars= 2 mm.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF8D7051ADB9FB73FC47F816.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673368/files/figure.png	https://doi.org/10.5281/zenodo.6673368	FIGURE 12. Tityus (Atreus) Gervais, 1843, metasoma, dorsal view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm.	FIGURE 12. Tityus (Atreus) Gervais, 1843, metasoma, dorsal view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF8D7051ADB9FB73FC47F816.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673372/files/figure.png	https://doi.org/10.5281/zenodo.6673372	FIGURE 13. Tityus (Atreus) Gervais, 1843, metasoma, ventral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm.	FIGURE 13. Tityus (Atreus) Gervais, 1843, metasoma, ventral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF8D7051ADB9FB73FC47F816.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673374/files/figure.png	https://doi.org/10.5281/zenodo.6673374	FIGURE 14. Tityus (Atreus) Gervais, 1843, metasoma, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm.	FIGURE 14. Tityus (Atreus) Gervais, 1843, metasoma, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF8D7051ADB9FB73FC47F816.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673376/files/figure.png	https://doi.org/10.5281/zenodo.6673376	FIGURE 15. Tityus (Atreus) Gervais, 1843, telson, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 1.5 mm.	FIGURE 15. Tityus (Atreus) Gervais, 1843, telson, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 1.5 mm.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF8D7051ADB9FB73FC47F816.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673378/files/figure.png	https://doi.org/10.5281/zenodo.6673378	FIGURE 16. Distribution map of the Tityus forcipula species group. A. South America. B. Records area. C. Species records.	FIGURE 16. Distribution map of the Tityus forcipula species group. A. South America. B. Records area. C. Species records.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF93705CADB9FA30FCF2FEE4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673334/files/figure.png	https://doi.org/10.5281/zenodo.6673334	FIGURE 1. Phylogenetic relationships of Tityus species inferred from a combined dataset of 28S and COI sequences. The topology is recovered from the IQ-TREE analysis. Squares at nodes indicate the Shimodaira–Hasegawa approximate likelihood-ratio test (left), the ultrafast bootstrap (middle) and the parsimony bootstrap (right). All sequences obtained in this study are in bold.	FIGURE 1. Phylogenetic relationships of Tityus species inferred from a combined dataset of 28S and COI sequences. The topology is recovered from the IQ-TREE analysis. Squares at nodes indicate the Shimodaira–Hasegawa approximate likelihood-ratio test (left), the ultrafast bootstrap (middle) and the parsimony bootstrap (right). All sequences obtained in this study are in bold.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF93705CADB9FA30FCF2FEE4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673342/files/figure.png	https://doi.org/10.5281/zenodo.6673342	FIGURE 3. Tityus (Atreus) moralensis sp. nov., habitus. A, B. Male holotype (MUSENUV-Ar 2104). A. Dorsal view. B. Ventral view. C, D. Female paratype (ICN-1402). C. Dorsal view. D. Ventral view. Scale bar= 15 mm.	FIGURE 3. Tityus (Atreus) moralensis sp. nov., habitus. A, B. Male holotype (MUSENUV-Ar 2104). A. Dorsal view. B. Ventral view. C, D. Female paratype (ICN-1402). C. Dorsal view. D. Ventral view. Scale bar= 15 mm.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF93705CADB9FA30FCF2FEE4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673348/files/figure.png	https://doi.org/10.5281/zenodo.6673348	FIGURE 4. Tityus (Atreus) Gervais, 1843, carapace sculpturing, dorsal aspect. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 2.5 mm.	FIGURE 4. Tityus (Atreus) Gervais, 1843, carapace sculpturing, dorsal aspect. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 2.5 mm.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF93705CADB9FA30FCF2FEE4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673350/files/figure.png	https://doi.org/10.5281/zenodo.6673350	FIGURE 5. Tityus (Atreus) Gervais, 1843, femur sculpturing. A–D. Tityus (Atreus) forcipula (Gervais, 1843). A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. E–H. Tityus (Atreus) moralensis sp. nov. E, F. Male holotype (MUSENUV-Ar 2104). E. Dorsal view. F. Ventral view. G, H. Female paratype (ICN-1402). G. Dorsal view. H. Ventral view. Scale bars= 2 mm.	FIGURE 5. Tityus (Atreus) Gervais, 1843, femur sculpturing. A–D. Tityus (Atreus) forcipula (Gervais, 1843). A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. E–H. Tityus (Atreus) moralensis sp. nov. E, F. Male holotype (MUSENUV-Ar 2104). E. Dorsal view. F. Ventral view. G, H. Female paratype (ICN-1402). G. Dorsal view. H. Ventral view. Scale bars= 2 mm.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF93705CADB9FA30FCF2FEE4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673354/files/figure.png	https://doi.org/10.5281/zenodo.6673354	FIGURE 6. Tityus (Atreus) Gervais, 1843, patella sculpturing. A–D. Tityus (Atreus) forcipula (Gervais, 1843). A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. E–H. Tityus (Atreus) moralensis sp. nov. E, F. Male holotype (MUSENUV-Ar 2104). E. Dorsal view. F. Ventral view. G, H. Female paratype (ICN-1402). G. Dorsal view. H. Ventral view. Scale bars= 2 mm.	FIGURE 6. Tityus (Atreus) Gervais, 1843, patella sculpturing. A–D. Tityus (Atreus) forcipula (Gervais, 1843). A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. E–H. Tityus (Atreus) moralensis sp. nov. E, F. Male holotype (MUSENUV-Ar 2104). E. Dorsal view. F. Ventral view. G, H. Female paratype (ICN-1402). G. Dorsal view. H. Ventral view. Scale bars= 2 mm.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF93705CADB9FA30FCF2FEE4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673356/files/figure.png	https://doi.org/10.5281/zenodo.6673356	FIGURE 7. Tityus (Atreus) Gervais, 1843,female chela sculpturing.A–C. Tityus(Atreus) forcipula (Gervais,1843) (MUSENUV- Ar 436). A. External view. B. Dorsal view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (ICN-1402). D. External view. E. Dorsal view. F. Ventral view. Scale bars= 3 mm.	FIGURE 7. Tityus (Atreus) Gervais, 1843,female chela sculpturing.A–C. Tityus(Atreus) forcipula (Gervais,1843) (MUSENUV- Ar 436). A. External view. B. Dorsal view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (ICN-1402). D. External view. E. Dorsal view. F. Ventral view. Scale bars= 3 mm.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF93705CADB9FA30FCF2FEE4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673358/files/figure.png	https://doi.org/10.5281/zenodo.6673358	FIGURE 8. Tityus (Atreus) Gervais, 1843, male chela sculpturing. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. External view. B. Dorsal view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. External view. E. Dorsal view. F. Ventral view. Scale bars= 3 mm.	FIGURE 8. Tityus (Atreus) Gervais, 1843, male chela sculpturing. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. External view. B. Dorsal view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. External view. E. Dorsal view. F. Ventral view. Scale bars= 3 mm.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF93705CADB9FA30FCF2FEE4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673360/files/figure.png	https://doi.org/10.5281/zenodo.6673360	FIGURE 9. Tityus (Atreus) Gervais, 1843, pectinal area. A, B. Tityus (Atreus) forcipula (Gervais, 1843).A. Male (MUSENUV- Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402).	FIGURE 9. Tityus (Atreus) Gervais, 1843, pectinal area. A, B. Tityus (Atreus) forcipula (Gervais, 1843).A. Male (MUSENUV- Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402).	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF93705CADB9FA30FCF2FEE4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673364/files/figure.png	https://doi.org/10.5281/zenodo.6673364	FIGURE 10. Tityus (Atreus) Gervais, 1843, sternites IV–VII. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402).	FIGURE 10. Tityus (Atreus) Gervais, 1843, sternites IV–VII. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402).	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF93705CADB9FA30FCF2FEE4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673366/files/figure.png	https://doi.org/10.5281/zenodo.6673366	FIGURE 11. Tityus (Atreus) Gervais, 1843, hemispermatophore. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. Posterior view. B. Lateral view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. Posterior view. E. Lateral view. F. Ventral view. Abbreviations= B, body; Bl, basal lobe; El, external lobe; F, flagellum; Fo, foot; iL, internal lobe; Pref, pars reflexa; Prec, pars recta. Scale bars= 2 mm. Scale bars= 2 mm.	FIGURE 11. Tityus (Atreus) Gervais, 1843, hemispermatophore. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. Posterior view. B. Lateral view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. Posterior view. E. Lateral view. F. Ventral view. Abbreviations= B, body; Bl, basal lobe; El, external lobe; F, flagellum; Fo, foot; iL, internal lobe; Pref, pars reflexa; Prec, pars recta. Scale bars= 2 mm. Scale bars= 2 mm.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF93705CADB9FA30FCF2FEE4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673368/files/figure.png	https://doi.org/10.5281/zenodo.6673368	FIGURE 12. Tityus (Atreus) Gervais, 1843, metasoma, dorsal view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm.	FIGURE 12. Tityus (Atreus) Gervais, 1843, metasoma, dorsal view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF93705CADB9FA30FCF2FEE4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673372/files/figure.png	https://doi.org/10.5281/zenodo.6673372	FIGURE 13. Tityus (Atreus) Gervais, 1843, metasoma, ventral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm.	FIGURE 13. Tityus (Atreus) Gervais, 1843, metasoma, ventral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF93705CADB9FA30FCF2FEE4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673374/files/figure.png	https://doi.org/10.5281/zenodo.6673374	FIGURE 14. Tityus (Atreus) Gervais, 1843, metasoma, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm.	FIGURE 14. Tityus (Atreus) Gervais, 1843, metasoma, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF93705CADB9FA30FCF2FEE4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673376/files/figure.png	https://doi.org/10.5281/zenodo.6673376	FIGURE 15. Tityus (Atreus) Gervais, 1843, telson, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 1.5 mm.	FIGURE 15. Tityus (Atreus) Gervais, 1843, telson, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 1.5 mm.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF93705CADB9FA30FCF2FEE4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673378/files/figure.png	https://doi.org/10.5281/zenodo.6673378	FIGURE 16. Distribution map of the Tityus forcipula species group. A. South America. B. Records area. C. Species records.	FIGURE 16. Distribution map of the Tityus forcipula species group. A. South America. B. Records area. C. Species records.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
B3264F4BFF93705CADB9FA30FCF2FEE4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6673380/files/figure.png	https://doi.org/10.5281/zenodo.6673380	FIGURE 17. Tityus (Atreus) moralensis sp. nov. A. Male holotype, live habitus. B. Microhabitat in El Cerrito, El Moral pathway in the Cordillera Central, Colombia.	FIGURE 17. Tityus (Atreus) moralensis sp. nov. A. Male holotype, live habitus. B. Microhabitat in El Cerrito, El Moral pathway in the Cordillera Central, Colombia.	2022-06-21	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy		Zenodo	biologists	Pinto-Da-Rocha, Ricardo;Cabra-García, Jimmy			
