taxonID	namePublishedIn	acceptedNameUsageID	parentNameUsageID	originalNameUsageID	kingdom	phylum	class	order	family	genus	taxonRank	scientificName	scientificNameAuthorship	canonicalName	verbatimScientificName	basionymAuthors	basionymYear	combinationAuthors	combinationYear	taxonomicStatus	nomenclaturalStatus	references	taxonRemarks
B276DC1EFFA0FFE5FF0DF922FAF306E0.taxon					Animalia	Arthropoda	Insecta	Coleoptera	Scarabaeidae	Macroderes	species	Macroderes amplior Frolov & Scholtz 2005	Frolov & Scholtz 2005	Macroderes amplior	Macroderes amplior Frolov & Scholtz, 2005			Frolov & Scholtz	2005			https://treatment.plazi.org/id/B276DC1EFFA0FFE5FF0DF922FAF306E0	
B276DC1EFFA1FFE5FF0DFC07FEBD03BB.taxon					Animalia	Arthropoda	Insecta	Coleoptera	Scarabaeidae	Macroderes	species	Macroderes arrowi Janssens 1939	Janssens 1939	Macroderes arrowi	Macroderes arrowi Janssens, 1939			Janssens	1939			https://treatment.plazi.org/id/B276DC1EFFA1FFE5FF0DFC07FEBD03BB	
B276DC1EFFA6FFE1FF0DFA45FD8A0405.taxon					Animalia	Arthropoda	Insecta	Coleoptera	Scarabaeidae	Macroderes	species	Macroderes bias Olivier 1789	Olivier 1789	Macroderes bias	Macroderes bias Olivier, 1789			Olivier	1789			https://treatment.plazi.org/id/B276DC1EFFA6FFE1FF0DFA45FD8A0405	
B276DC1EFFA6FFE1FF0DFA45FD8A0405.taxon.syn1		B276DC1EFFA6FFE1FF0DFA45FD8A0405.taxon			Animalia	Arthropoda	Insecta	Coleoptera	Scarabaeidae	Scarabaeus	species	Scarabaeus bias Olivier 1789	Olivier 1789	Scarabaeus bias	Scarabaeus bias Olivier, 1789: 187			Olivier	1789	synonym		https://treatment.plazi.org/id/B276DC1EFFA6FFE1FF0DFA45FD8A0405	
B276DC1EFFA6FFE1FF0DFA45FD8A0405.taxon.syn2		B276DC1EFFA6FFE1FF0DFA45FD8A0405.taxon			Animalia	Arthropoda	Insecta	Coleoptera	Scarabaeidae	Macroderes	species	Macroderes pilula Sharp 1880	Sharp 1880	Macroderes pilula	Macroderes pilula Sharp, 1880: 38			Sharp	1880	synonym		https://treatment.plazi.org/id/B276DC1EFFA6FFE1FF0DFA45FD8A0405	
B276DC1EFFA5FFEEFF0DF8D5FBC7003D.taxon	Abdalla, Ishtiag H., Deschodt, Christian M., Scholtz, Clarke H., Sole, Catherine L. (2018): An update to the taxonomy of the genus Macroderes Westwood 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) with descriptions of new species from South Africa. Zootaxa 4504 (1): 41-75, DOI: 10.11646/zootaxa.4504.1.3				Animalia	Arthropoda	Insecta	Coleoptera	Scarabaeidae	Macroderes	species	Macroderes cederbergensis Abdalla, Deschodt, Scholtz, and Sole 2018	Abdalla, Deschodt, Scholtz, and Sole 2018	Macroderes cederbergensis	Macroderes cederbergensis Abdalla & Deschodt			Abdalla & Deschodt & Scholtz & Sole	2018		spec. nov.	https://treatment.plazi.org/id/B276DC1EFFA5FFEEFF0DF8D5FBC7003D	
B276DC1EFFA5FFEEFF0DF8D5FBC7003D.taxon.syn1		B276DC1EFFA5FFEEFF0DF8D5FBC7003D.taxon			Animalia	Arthropoda	Insecta	Coleoptera	Scarabaeidae	Macroderes	species	Macroderes minutus Frolov & Scholtz 2005	Frolov & Scholtz 2005	Macroderes minutus	Macroderes minutus Frolov & Scholtz, 2005: 385			Frolov & Scholtz	2005	synonym		https://treatment.plazi.org/id/B276DC1EFFA5FFEEFF0DF8D5FBC7003D	
B276DC1EFFAAFFEFFF0DFA36FBD507C1.taxon					Animalia	Arthropoda	Insecta	Coleoptera	Scarabaeidae	Macroderes	species	Macroderes cornutus Frolov & Scholtz 2005	Frolov & Scholtz 2005	Macroderes cornutus	Macroderes cornutus Frolov & Scholtz, 2005			Frolov & Scholtz	2005			https://treatment.plazi.org/id/B276DC1EFFAAFFEFFF0DFA36FBD507C1	
B276DC1EFFA8FFEDFF0DF9B0FBAE003D.taxon					Animalia	Arthropoda	Insecta	Coleoptera	Scarabaeidae	Macroderes	species	Macroderes endroedyi Frolov & Scholtz 2005	Frolov & Scholtz 2005	Macroderes endroedyi	endroedyi Frolov & Scholtz, 2005			Frolov & Scholtz	2005			https://treatment.plazi.org/id/B276DC1EFFA8FFEDFF0DF9B0FBAE003D	
B276DC1EFFA9FFEAFF0DFA36FBA7075D.taxon					Animalia	Arthropoda	Insecta	Coleoptera	Scarabaeidae	Macroderes	species	Macroderes fornicatus Sharp 1880	Sharp 1880	Macroderes fornicatus	Macroderes fornicatus Sharp, 1880			Sharp	1880			https://treatment.plazi.org/id/B276DC1EFFA9FFEAFF0DFA36FBA7075D	
B276DC1EFFAEFFE8FF0DF89DFB3905BD.taxon					Animalia	Arthropoda	Insecta	Coleoptera	Scarabaeidae	Macroderes	species	Macroderes foveatus Frolov & Scholtz 2005	Frolov & Scholtz 2005	Macroderes foveatus	Macroderes foveatus Frolov & Scholtz, 2005			Frolov & Scholtz	2005			https://treatment.plazi.org/id/B276DC1EFFAEFFE8FF0DF89DFB3905BD	
B276DC1EFFACFFE9FF0DF894FF23003D.taxon	Abdalla, Ishtiag H., Deschodt, Christian M., Scholtz, Clarke H., Sole, Catherine L. (2018): An update to the taxonomy of the genus Macroderes Westwood 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) with descriptions of new species from South Africa. Zootaxa 4504 (1): 41-75, DOI: 10.11646/zootaxa.4504.1.3				Animalia	Arthropoda	Insecta	Coleoptera	Scarabaeidae	Macroderes	species	Macroderes gifboomi Abdalla, Deschodt, Scholtz, and Sole 2018	Abdalla, Deschodt, Scholtz, and Sole 2018	Macroderes gifboomi	Macroderes gifboomi Abdalla & Scholtz			Abdalla & Deschodt & Scholtz & Sole	2018		spec. nov.	https://treatment.plazi.org/id/B276DC1EFFACFFE9FF0DF894FF23003D	
B276DC1EFFADFFF7FF0DFA36FA9404E8.taxon					Animalia	Arthropoda	Insecta	Coleoptera	Scarabaeidae	Macroderes	species	Macroderes greeni Kirby 1818	Kirby 1818	Macroderes greeni	Macroderes greeni Kirby, 1818			Kirby	1818			https://treatment.plazi.org/id/B276DC1EFFADFFF7FF0DFA36FA9404E8	
B276DC1EFFADFFF7FF0DFA36FA9404E8.taxon.syn1		B276DC1EFFADFFF7FF0DFA36FA9404E8.taxon			Animalia	Arthropoda	Insecta	Coleoptera	Scarabaeidae	Onthophagus	species	Onthophagus greeni Kirby 1818	Kirby 1818	Onthophagus greeni	Onthophagus greeni Kirby, 1818: 397			Kirby	1818	synonym		https://treatment.plazi.org/id/B276DC1EFFADFFF7FF0DFA36FA9404E8	
B276DC1EFFB0FFF5FF0DF90DFEF9010E.taxon	Abdalla, Ishtiag H., Deschodt, Christian M., Scholtz, Clarke H., Sole, Catherine L. (2018): An update to the taxonomy of the genus Macroderes Westwood 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) with descriptions of new species from South Africa. Zootaxa 4504 (1): 41-75, DOI: 10.11646/zootaxa.4504.1.3				Animalia	Arthropoda	Insecta	Coleoptera	Scarabaeidae	Macroderes	species	Macroderes leipoldti Abdalla, Deschodt, Scholtz, and Sole 2018	Abdalla, Deschodt, Scholtz, and Sole 2018	Macroderes leipoldti	Macroderes leipoldti Abdalla & Deschodt			Abdalla & Deschodt & Scholtz & Sole	2018		spec. nov.	https://treatment.plazi.org/id/B276DC1EFFB0FFF5FF0DF90DFEF9010E	
B276DC1EFFB1FFF3FF0DFB60FD9E0420.taxon					Animalia	Arthropoda	Insecta	Coleoptera	Scarabaeidae	Macroderes	species	Macroderes minutus Frolov & Scholtz 2005	Frolov & Scholtz 2005	Macroderes minutus	Macroderes minutus Frolov & Scholtz, 2005			Frolov & Scholtz	2005			https://treatment.plazi.org/id/B276DC1EFFB1FFF3FF0DFB60FD9E0420	
B276DC1EFFB7FFF0FF0DF960FF3D0155.taxon					Animalia	Arthropoda	Insecta	Coleoptera	Scarabaeidae	Macroderes	species	Macroderes mutilans Kolbe 1908	Kolbe 1908	Macroderes mutilans	Macroderes mutilans Kolbe, 1908			Kolbe	1908			https://treatment.plazi.org/id/B276DC1EFFB7FFF0FF0DF960FF3D0155	
B276DC1EFFB4FFF0FF0DFBA9FC0B02AA.taxon					Animalia	Arthropoda	Insecta	Coleoptera	Scarabaeidae	Macroderes	species	Macroderes namakwanus Frolov & Scholtz 2005	Frolov & Scholtz 2005	Macroderes namakwanus	Macroderes namakwanus Frolov & Scholtz, 2005			Frolov & Scholtz	2005			https://treatment.plazi.org/id/B276DC1EFFB4FFF0FF0DFBA9FC0B02AA	
B276DC1EFFB5FFFFFF0DFA3AFF3D0725.taxon					Animalia	Arthropoda	Insecta	Coleoptera	Scarabaeidae	Macroderes	species	Macroderes nitidus Harold / 1877	Harold / 1877	Macroderes nitidus	Macroderes nitidus Harold, 1877			Harold /	1877			https://treatment.plazi.org/id/B276DC1EFFB5FFFFFF0DFA3AFF3D0725	
B276DC1EFFBBFFFCFF0DFD44FC810458.taxon	Abdalla, Ishtiag H., Deschodt, Christian M., Scholtz, Clarke H., Sole, Catherine L. (2018): An update to the taxonomy of the genus Macroderes Westwood 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) with descriptions of new species from South Africa. Zootaxa 4504 (1): 41-75, DOI: 10.11646/zootaxa.4504.1.3				Animalia	Arthropoda	Insecta	Coleoptera	Scarabaeidae	Macroderes	species	Macroderes oreatus Abdalla, Deschodt, Scholtz, and Sole 2018	Abdalla, Deschodt, Scholtz, and Sole 2018	Macroderes oreatus	Macroderes oreatus Abdalla & Deschodt			Abdalla & Deschodt & Scholtz & Sole	2018		spec. nov.	https://treatment.plazi.org/id/B276DC1EFFBBFFFCFF0DFD44FC810458	
B276DC1EFFB8FFFCFF0DFE9FFBC80135.taxon					Animalia	Arthropoda	Insecta	Coleoptera	Scarabaeidae	Macroderes	species	Macroderes politulus Preudhomme de Borre 1880	Preudhomme de Borre 1880	Macroderes politulus	Macroderes politulus Preudhomme de Borre, 1880			Preudhomme de Borre	1880			https://treatment.plazi.org/id/B276DC1EFFB8FFFCFF0DFE9FFBC80135	
B276DC1EFFB8FFFDFF0DFB34FE360778.taxon	Abdalla, Ishtiag H., Deschodt, Christian M., Scholtz, Clarke H., Sole, Catherine L. (2018): An update to the taxonomy of the genus Macroderes Westwood 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) with descriptions of new species from South Africa. Zootaxa 4504 (1): 41-75, DOI: 10.11646/zootaxa.4504.1.3				Animalia	Arthropoda	Insecta	Coleoptera	Scarabaeidae	Macroderes	species	Macroderes porselinus Abdalla, Deschodt, Scholtz, and Sole 2018	Abdalla, Deschodt, Scholtz, and Sole 2018	Macroderes porselinus	Macroderes porselinus Abdalla			Abdalla & Deschodt & Scholtz & Sole	2018		spec. nov.	https://treatment.plazi.org/id/B276DC1EFFB8FFFDFF0DFB34FE360778	
B276DC1EFFB9FFFAFF0DFD75FC2304E8.taxon	Abdalla, Ishtiag H., Deschodt, Christian M., Scholtz, Clarke H., Sole, Catherine L. (2018): An update to the taxonomy of the genus Macroderes Westwood 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) with descriptions of new species from South Africa. Zootaxa 4504 (1): 41-75, DOI: 10.11646/zootaxa.4504.1.3				Animalia	Arthropoda	Insecta	Coleoptera	Scarabaeidae	Macroderes	species	Macroderes soleiana Abdalla, Deschodt, Scholtz, and Sole 2018	Abdalla, Deschodt, Scholtz, and Sole 2018	Macroderes soleiana	Macroderes soleiana Abdalla & Deschodt			Abdalla & Deschodt & Scholtz & Sole	2018		spec. nov.	https://treatment.plazi.org/id/B276DC1EFFB9FFFAFF0DFD75FC2304E8	
B276DC1EFFB9FFFAFF0DFD75FC2304E8.taxon.syn1		B276DC1EFFB9FFFAFF0DFD75FC2304E8.taxon			Animalia	Arthropoda	Insecta	Coleoptera	Scarabaeidae	Macroderes	species	Macroderes nitidus Harold / 1877	Harold / 1877	Macroderes nitidus	Macroderes nitidus Harold, 1877: 384			Harold /	1877	synonym		https://treatment.plazi.org/id/B276DC1EFFB9FFFAFF0DFD75FC2304E8	
B276DC1EFFBEFFFAFF0DFE0AFF2302CD.taxon	Abdalla, Ishtiag H., Deschodt, Christian M., Scholtz, Clarke H., Sole, Catherine L. (2018): An update to the taxonomy of the genus Macroderes Westwood 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) with descriptions of new species from South Africa. Zootaxa 4504 (1): 41-75, DOI: 10.11646/zootaxa.4504.1.3				Animalia	Arthropoda	Insecta	Coleoptera	Scarabaeidae	Macroderes	species	Macroderes tortuosus Abdalla, Deschodt, Scholtz, and Sole 2018	Abdalla, Deschodt, Scholtz, and Sole 2018	Macroderes tortuosus	Macroderes tortuosus Abdalla and Scholtz			Abdalla & Deschodt & Scholtz & Sole	2018		spec. nov.	https://treatment.plazi.org/id/B276DC1EFFBEFFFAFF0DFE0AFF2302CD	
B276DC1EFFBFFFFBFF0DFF67FE2600CE.taxon					Animalia	Arthropoda	Insecta	Coleoptera	Scarabaeidae	Macroderes	species	Macroderes undulatus Preudhomme de Borre 1880	Preudhomme de Borre 1880	Macroderes undulatus	Macroderes undulatus Preudhomme de Borre, 1880			Preudhomme de Borre	1880			https://treatment.plazi.org/id/B276DC1EFFBFFFFBFF0DFF67FE2600CE	
B276DC1EFFBFFFF9FF0DFA2EFE290418.taxon					Animalia	Arthropoda	Insecta	Coleoptera	Scarabaeidae	Macroderes	species	Macroderes pristinus Sharp 1880	Sharp 1880	Macroderes pristinus	Macroderes pristinus Sharp, 1880			Sharp	1880			https://treatment.plazi.org/id/B276DC1EFFBFFFF9FF0DFA2EFE290418	
B276DC1EFFBDFFF9FF0DFE52FEE506CD.taxon					Animalia	Arthropoda	Insecta	Coleoptera	Scarabaeidae	Macroderes	species	Macroderes dubius Peringuey 1901	Peringuey 1901	Macroderes dubius	Macroderes dubius Péringuey, 1901			Peringuey	1901			https://treatment.plazi.org/id/B276DC1EFFBDFFF9FF0DFE52FEE506CD	
B276DC1EFFBDFFF9FF0DFC2CFE0B034F.taxon					Animalia	Arthropoda	Insecta	Coleoptera	Scarabaeidae	Macroderes	species	Macroderes spectabilis Peringuey 1901	Peringuey 1901	Macroderes spectabilis	Macroderes spectabilis Péringuey, 1901			Peringuey	1901			https://treatment.plazi.org/id/B276DC1EFFBDFFF9FF0DFC2CFE0B034F	
