taxonID	type	description	language	source
B17287F5E077FF9D4EB8888CFBE514B9.taxon	description	General distribution. The species is only known from the Ethiopian highlands.	en	Kuhlmann, Michael (2020): New records of the bee genus Colletes Latreille 1802 (Hymenoptera: Apoidea Colletidae) in Ethiopia with description of a new species. Zootaxa 4722 (6): 583-590, DOI: 10.11646/zootaxa.4722.6.5
B17287F5E077FF9F4EB88DBBFDA917B6.taxon	description	(Figs 1 – 2)	en	Kuhlmann, Michael (2020): New records of the bee genus Colletes Latreille 1802 (Hymenoptera: Apoidea Colletidae) in Ethiopia with description of a new species. Zootaxa 4722 (6): 583-590, DOI: 10.11646/zootaxa.4722.6.5
B17287F5E077FF9F4EB88DBBFDA917B6.taxon	diagnosis	Diagnosis. Colletes michaelis belongs to the taxonomically challenging C. rufitarsis species-group. The group comprises nine described species of which including C. michaelis four occur in Ethiopia (see Kuhlmann & Pauly 2013). The female of C. michaelis differs from the other described species of the group by a combination of the following characters: fine and dense punctation of T 1 (Fig. 1 d) (only C. senkelensis Kuhlmann 2013 has finer punctation, see Kuhlmann & Pauly 2013) and lack of oily bluish shine of metasomal terga. The male can be recognized by the typical shape of S 7 (Fig. 2 c) in combination with the lack of an oily bluish shine of the finely and densely punctured metasomal terga (Fig. 2 b).	en	Kuhlmann, Michael (2020): New records of the bee genus Colletes Latreille 1802 (Hymenoptera: Apoidea Colletidae) in Ethiopia with description of a new species. Zootaxa 4722 (6): 583-590, DOI: 10.11646/zootaxa.4722.6.5
B17287F5E077FF9F4EB88DBBFDA917B6.taxon	description	Description. Female. Bl = 10.0 – 11.0 mm. Head. Head 1.25 x wider than long. Integument black except mandible partly dark reddish-brown. Face except clypeus densely covered with long, yellowish-grey, erect hairs. Clypeus convex, with shallow longitudinal median depression, supraclypeal area elevated, rectangular. Clypeus coarsely and densely punctate; surface between punctures in the upper part slightly shagreened otherwise smooth and shiny; Supraclypeal area with coarse but shallow puncatation, matt to slightly shiny (Fig. 1 b). Malar area medially about 1 / 3 as long as width of mandible base, finely striate. Antenna black, ventrally yellowish to dark brown. Mesosoma. Integument black. Mesoscutal disc coarsely and mostly densely punctate (i <0.5 d), shagreened and matt, sometimes medially shiny (Fig. 1 c). Scutellum as mesoscutum but punctation mostly finer (Fig. 1 c). Mesoscutum, scutellum, metanotum, mesepisternum and propodeum covered with long orange-brown erect hairs (Fig. 1 a). Wings. Yellow- ish; wing venation yellowish to dark brown. Legs. Integument dark to light reddish-brown. Vestiture whitish to yellowish, scopa yellowish-white. Metasoma. Integument black except depressed apical margins of terga reddish to yellowish translucent (Fig. 1 d). T 1 anteriorly densely covered with long, yellowish-white erect hairs; apical tergal hair band of T 1 medially narrower, on T 2 – T 5 broad and yellowish-white. Terga densely (i <d) and finely punctate, between punctures smooth and shiny (Fig. 1 d). Male. Bl = 9.5 – 11.0 mm. Head. Head 1.27 x wider than long. Integument black except mandible partly dark reddish-brown. Face densely covered with long, yellowish-brown, erect hairs. Malar area medially about 2 / 3 long as width of mandible base, finely striate. Antenna black, ventrally dark brown. Mesosoma. Integument black. Mesoscutal disc irregularly densely punctate (i <0.5 d), shagreened and matt. Mesoscutum, scutellum, metanotum, mesepisternum and propodeum covered with long yellowish-brown, erect hairs (Fig. 2 a). Wings. Slightly yellow- ish-brown; wing venation yellowish to dark brown. Legs. Integument dark reddish-brown to yellowish-brown (tarsi). Vestiture whitish to yellowish-brown. Metasoma. Integument black except depressed apical margins of terga yellowish translucent. T 1 completely and T 2 medially loosely covered with long but apically successively shorter, erect yellowish-brown hairs (Fig. 2 b); apical tergal hair bands broad, yellowish-brown. Terga densely and finely punctate (i <d), between punctures smooth and shiny (Fig. 2 b). Terminalia. Genitalia and S 7 as illustrated (Figs. 2 c – d).	en	Kuhlmann, Michael (2020): New records of the bee genus Colletes Latreille 1802 (Hymenoptera: Apoidea Colletidae) in Ethiopia with description of a new species. Zootaxa 4722 (6): 583-590, DOI: 10.11646/zootaxa.4722.6.5
B17287F5E077FF9F4EB88DBBFDA917B6.taxon	materials_examined	Material examined (87 specimens). Type material: female Lectotype, DEMOCRATIC REPUBLIC OF THE CONGO: Lubumbashi, Katanga, 11 ° 45 ′ S, 27 ° 40 ′ E, 12.1.1921, Mich. Bequaert Coll. (BMNH). Additional material: ETHIOPIA: 1 ♀, SNNPR, Chebera Churchura NP, E 36 ° 40 ′ N 07 ° 01 ′, 28. x. 2018, M. Kas- parek (RCMK); 4 ♂, 30 km NW Negele, E 39 ° 26 ′ N 05 ° 32 ′, 1480 m, 25. v. 2015, J. Halada (OÖLM, RCMK). DEMO- CRATIC REPUBLIC OF THE CONGO: 1 ♂, P. N. A. Mutsora, [E 29 ° 44 ′ N 00 ° 19 ′], 1939, Hackars (MRAC); 1 ♀, Rutshuru, [E 29 ° 26 ′ S 01 ° 10 ′], x. 1937, J. Ghesquière (RCMK) (labelled as PT C. tropicus); 2 ♂, Bassin Lu- kuga, [E 28 ° S 05 °], iv. – vii. 1934, De Saeger (MRAC) (labelled as HT, PT C. tropicus); 1 ♂, Lubumbashi, Katanga, E 27 ° 40 ′ S 11 ° 45 ′, 31. i. 1921, Mich. Bequaert (BMNH); 1 ♂, idem, 27. i. 1921 (RCMK). UGANDA: 1 ♀, 1 ♂, En- tebbe [E 32 ° 27 ′ N 00 ° 03 ′], 21. i. 1913, C. C. Gowdey (BMNH, RCMK) (labelled as PT C. tropicus); 1 ♀, 1 ♂, idem, 1. i. 1913 (BMNH, RCMK) (labelled as PT C. tropicus); 1 ♀, Kampala, [E 32 ° 34 ′ N 00 ° 19 ′], 23. ix. 1913, C. C. Gowdey (BMNH) (labelled as PT C. tropicus); 1 ♀, 2 ♂, near Kampala, [E 32 ° 33 ′ N 00 ° 19 ′], 2007, T. Munyuli (SANC, RCMK); 2 ♀, Ankole, Kichwamba, [E 30 ° 11 ′ N 00 ° 43 ′], 23. – 29. iv. 1968, P. J. Spangler (USNM, RCMK); 1 ♂, 20. – 30. ix. 1918, C. C. Gowdey (BMNH); 1 ♂, 1918, C. C. Gowdey (BMNH). KENYA: 2 ♂, Katutu-Kithioko, [E 37 ° 50 ′ S 01 ° 10 ′], 27. xi. 1999, M. Snizek (RCMK); 6 ♂, Nairobi, Salama env., [E 36 ° 48 ′ S 01 ° 17 ′], 25. xi. 1999 (OÖLM, RCMK); 1 ♂, Karen, Nairobi, [E 36 ° 42 ′ S 01 ° 19 ′], 2000 m, 17. vi. 1967, Michener (RCMK); 1 ♂, Ikutha, [E 38 ° 10 ′ S 02 ° 03 ′], 1898 (NHMW). BURUNDI: 1 ♂, 9 mi. N of Nyanza Lac, [E 29 ° 35 ′ S 04 ° 20 ′], 800 m, 1. vii. 1958, E. S. Ross & R. E. Leech (RCMK). TANZANIA: 1 ♂, prov. Kagera, Kimisi Game Reserve, 30 km S Karagwe, E 31 ° 04 ′ 46 ′′ S 01 ° 52 ′ 59 ′′, 1600 m, 25. – 26. xi. 2017, J. Halada (RCMK); 1 ♀, Kigoma pr., 25 km SSE Kasulu, [E 30 ° 11 ′ S 04 ° 44 ′], 1200 m, 28. xii. 2006, J. Halada (RCMK); 1 ♂, W of Kiberashi, Kitwei plain, [E 37 ° 25 ′ S 05 ° 23 ′], 16. iii. 2002, M. Snizek (RCMK); 1 ♀, Sagara mts., [E 36 ° 30 ′ S 06 ° 52 ′], 12. xii. 1997, M. Snizek (RCMK); 1 ♂, Mbeya pr., 70 km NW Tunduma, [E 32 ° 25 ′ S 08 ° 58 ′], 1500 m, 2. i. 2007, M. Kadlecova (OÖLM) (labelled as PT C. shonae). ZAMBIA: 1 ♂, Mkushi env. E, [E 29 ° 23 ′ S 13 ° 36 ′], 16. – 18. xii. 2004, M. Snizek (RCMK); 1 ♂, 40 km N Kabwe, Sangala school env., [E 28 ° 18 ′ S 14 ° 09 ′], 19. xii. 2004, M. Snizek (OÖLM); 2 ♂, 120 km N Lusaka, [E 28 ° 09 ′ S 14 ° 39 ′]; 12. – 14. xii. 2002, J. Halada (RCMK) (labelled as PT C. shonae); 1 ♂, 10 km S Mazambuka, [E 27 ° 45 ′ S 15 ° 54 ′]; 28. xii. 2002, J. Halada (RCMK) (labelled as PT C. shonae). MALAWI: 1 ♀, Rumpi, [E 33 ° 51 ′ S 11 ° 01 ′], 12. iv. 1967, C. D. Michener (SEMC) (labelled as PT C. shonae); 1 ♂, 6 mi. N of Ekwendeni, [E 33 ° 52 ′ S 11 ° 22 ′], 1200 m, 21. ii. 1958, E. S. Ross & R. E. Leech (RCMK) (labelled as PT C. shonae); 2 ♂, Dedza, [E 34 ° 19 ′ S 14 ° 22 ′], 20. iv. 1967, C. D. Michener (RCMK); 1 ♀, 85 km SE Lilongwe, Dedza, [E 34 ° 19 ′ S 14 ° 22 ′], 5. – 12. i. 2002, J. Halada (RCMK); 1 ♂, Mulanje Mts. env., [E 35 ° 30 ′ S 16 ° 00 ′], 23. – 26. xii. 2001, J. Halada (RCMK); 1 ♀, 2 ♂, Mlanje, [E 35 ° 30 ′ S 16 ° 02 ′], 6. iv. 1967, C. D. Michener (SEMC, RCMK) (labelled PT C. shonae). MOZAMBIQUE: 1 ♀, 65 km S Vlongné, [E 34 ° 19 ′ S 15 ° 13 ′], 1250 m, 8. xii. 2005, J. Halada (OÖLM) (labelled as PT C. shonae); 1 ♂, 100 km NE Tete, [E 34 ° 20 ′ S 15 ° 26 ′]; 910 m, 7. xii. 2005, J. Halada (OÖLM); 1 ♀, 6 ♂, 30 km NW Catandica, [E 33 ° 07 ′ S 17 ° 47 ′], 630 m, 12. xii. 2005, J. Halada (OÖLM, RCMK) (labelled as PT C. shonae); 4 ♂, idem, 17. – 18. xii. 2003 (OÖLM, RCMK) (labelled as PT C. shonae); 1 ♂, Manica pr., 65 km W Chimolo, [E 32 ° 55 ′ S 18 ° 57 ′], 10. – 11. xii. 2003, J. Halada (OÖLM) (labelled as PT C. shonae); 1 ♂, 40 km SW Chimolo, [E 33 ° 11 ′ S 19 ° 23 ′], 650 m, 16. xii. 2005, J. Halada (RCMK). ZIMBABWE: 3 ♂, 30 km W. Harare, [E 30 ° 45 ′ S 17 ° 48 ′], 22. xii. 1998, J. Halada (OÖLM, RCMK) (labelled as HT, PT C. shonae); 3 ♀, 4 ♂, Salisbury, Mashonaland, [E 31 ° 03 ′ S 17 ° 49 ′], January 1900, G. A. K. Marshal (BMNH, RCMK) (labelled as PT C. shonae); 3 ♂, 70 km SW Gweru, Nalatale Ruins env., E 29 ° 22 ′ S 19 ° 46 ′, December 2011, J. Halada (OÖLM, RCMK); 1 ♂, Bulawayo, [E 28 ° 34 ′ S 20 ° 10 ′], 2. xii. 1922, Rhodesia Museum (RCMK) (labelled as PT C. shonae); 1 ♂, idem, 26. xii. 1922 (RCMK) (labelled as PT C. shonae); 3 ♂, idem, 23. ii. 1913, G. Arnold (SDEI, RCMK) (la- belled as PT C. shonae). General distribution. This species is widely distributed in tropical and subtropical parts of eastern Africa from Ethiopia to Zimbabwe. Floral hosts. Scopal pollen loads of three females collected at three different sites (2 x Malawi, 1 x Tanzania) were analyzed. All samples only contained Asteraceae pollen suggesting that C. michaelis might be a specialist (oligolectic) bee species. Seasonal activity (first – last observations). Throughout its range C. michaelis was recorded in every month of the year except for August and depending on climatic conditions seems to be active all year round.	en	Kuhlmann, Michael (2020): New records of the bee genus Colletes Latreille 1802 (Hymenoptera: Apoidea Colletidae) in Ethiopia with description of a new species. Zootaxa 4722 (6): 583-590, DOI: 10.11646/zootaxa.4722.6.5
B17287F5E077FF9F4EB88DBBFDA917B6.taxon	discussion	Remarks. Colletes michaelis shows some slight morphological variation throughout its range. When I started studying Afrotropical Colletes and only few isolated specimens were available this variation suggested the existence of two additional closely related taxa. They were named and labelled as type specimens of C. shonae and C. tropicus, respectively, but not formally described and can still be found as such in the collections mentioned under “ Ma- terial examined ” (HT — Holotype, PT — Paratype (s )). Later intermediate forms turned up supporting the assumption of only a single variable species.	en	Kuhlmann, Michael (2020): New records of the bee genus Colletes Latreille 1802 (Hymenoptera: Apoidea Colletidae) in Ethiopia with description of a new species. Zootaxa 4722 (6): 583-590, DOI: 10.11646/zootaxa.4722.6.5
B17287F5E072FF994EB88C33FD63128A.taxon	description	(Fig. 3)	en	Kuhlmann, Michael (2020): New records of the bee genus Colletes Latreille 1802 (Hymenoptera: Apoidea Colletidae) in Ethiopia with description of a new species. Zootaxa 4722 (6): 583-590, DOI: 10.11646/zootaxa.4722.6.5
B17287F5E072FF994EB88C33FD63128A.taxon	diagnosis	Diagnosis. Colletes meneliki belongs to the C. malmus species-group that comprises seven described species (C. malmus (Cameron 1905), C. mackiae Cockerell 1932, C. malleatus Cockerell 1933, C. phenax Cockerell 1946, C. plebeius Cockerell 1946, C. sordescens Cockerell 1933, C. sororcula Cockerell 1936) and at least 16 undescribed ones that are mostly from southern Africa. The group is characterised by a more or less rounded S 7 in males (cf. Fig. 3 c) in combination with an oily bluish shine of variable intensity on metasomal terga in most species. The new species is among the smallest in this group and unique because of the very dispersed and fine punctation of mesoscutum and metasomal terga (Fig. 3 b). The combination of shiny, hardly punctured terga and the characteristic shape of S 7 is distinctive and there are no species that C. meneliki can be confused with.	en	Kuhlmann, Michael (2020): New records of the bee genus Colletes Latreille 1802 (Hymenoptera: Apoidea Colletidae) in Ethiopia with description of a new species. Zootaxa 4722 (6): 583-590, DOI: 10.11646/zootaxa.4722.6.5
B17287F5E072FF994EB88C33FD63128A.taxon	description	Description. Female. Unknown. Male. Bl = 7.0 – 7.5 mm. Head. Head 1.30 x wider than long. Integument black except tip of mandible partly dark reddish-brown. Face densely covered with long, white to yellowish-white, erect hairs, on vertex partly dark brown hairs. Malar area medially about half as long as width of mandible base, finely striate. Antenna black. Mesosoma. Integument black. Mesoscutal disc almost impunctate, smooth and shiny. Scutellum sparsely punctate, smooth and shiny. Mesoscutum, scutellum, metanotum, mesepisternum and propodeum covered with long, greyish-white erect hairs, on disc of mesoscutum and apical part of scutellum blackish hairs intermixed (Fig. 3 a). Wings. Transparent; wing venation brown to blackish. Legs. Integument black; apical tarsi yellowish-brown. Vestiture greyish-white. Metasoma. Integument black with slight oily bluish shine; depressed apical tergal margins yellowish to reddish translucent (Fig. 3 b). Disc of T 1 sparsely covered with long, erect greyish-white hairs (Fig. 3 a, b); apical tergal hair bands broad (Fig. 3 a, b). Terga sparsely and very finely punctate (i = 2 – 4 d), between punctures smooth and shiny (Fig. 3 b). Terminalia. Genitalia and S 7 as illustrated (Figs. 3 c – d).	en	Kuhlmann, Michael (2020): New records of the bee genus Colletes Latreille 1802 (Hymenoptera: Apoidea Colletidae) in Ethiopia with description of a new species. Zootaxa 4722 (6): 583-590, DOI: 10.11646/zootaxa.4722.6.5
B17287F5E072FF994EB88C33FD63128A.taxon	materials_examined	Type material (3 specimens). Holotype, ♂, ETHIOPIA, Sidam prov., 50 km NE Mega, [E 38 ° 27 ′ N 04 ° 24 ′], 1420 m, 24. iv. 2007, J. Halada (OÖLM). Paratypes: 1 ♂, ETHIOPIA, Oromia State, 25 km SE Mega, E 38 ° 26 ′ 53 ′′ N 03 ° 55 ′ 37 ′′, 1400 m, 15. iv. 2016, J. Halada (RCMK); 1 ♂, ETHIOPIA, Sidam prov., near Bitata [E 39 ° 28 ′ N 05 ° 28 ′], 1480 m, 27. iv. 2007, J. Halada (RCMK).	en	Kuhlmann, Michael (2020): New records of the bee genus Colletes Latreille 1802 (Hymenoptera: Apoidea Colletidae) in Ethiopia with description of a new species. Zootaxa 4722 (6): 583-590, DOI: 10.11646/zootaxa.4722.6.5
B17287F5E072FF994EB88C33FD63128A.taxon	etymology	Etymology. This species is named after Menelik I., the legendary first emperor of the Ethiopian Empire, illegitimate son of King Salomo and the Queen of Sheba and founder of the Salomonic dynasty. General distribution. Only known from south eastern Ethiopia. Floral hosts. unknown. Seasonal activity (first – last observations). IV.	en	Kuhlmann, Michael (2020): New records of the bee genus Colletes Latreille 1802 (Hymenoptera: Apoidea Colletidae) in Ethiopia with description of a new species. Zootaxa 4722 (6): 583-590, DOI: 10.11646/zootaxa.4722.6.5
B17287F5E073FF9A4EB88E3DFB4510DA.taxon	description	General distribution. The species is known from desert regions of East and Northeast Africa.	en	Kuhlmann, Michael (2020): New records of the bee genus Colletes Latreille 1802 (Hymenoptera: Apoidea Colletidae) in Ethiopia with description of a new species. Zootaxa 4722 (6): 583-590, DOI: 10.11646/zootaxa.4722.6.5
B17287F5E070FF9A4EB8881CFB3C132D.taxon	discussion	Remarks. The male collected in 1911 by Kristensen in Harrar is labeled exactly like the type material housed in the museums in Berlin and Frankfurt (Kuhlmann 1998) and presumably belongs to the original type series (see Kuhlmann 2018). General distribution. The species is apparently endemic to the highlands of Ethiopia.	en	Kuhlmann, Michael (2020): New records of the bee genus Colletes Latreille 1802 (Hymenoptera: Apoidea Colletidae) in Ethiopia with description of a new species. Zootaxa 4722 (6): 583-590, DOI: 10.11646/zootaxa.4722.6.5
