taxonID	type	description	language	source
B0587B3CA103FFBEBEDAFBEDFE42F975.taxon	materials_examined	Material examined. Holotype. ♀, [TANZANIA], Tanganyika, 14 mi S. of Sumbawanga, elev. 1750 m, 16. ii. [19] 58, (E S. Ross & R E. Leech) (CASC).	en	Azevedo, Celso O., Mugrabi, Daniele F. (2014): Three new species of Tuberepyris Lanes et Azevedo (Hymenoptera, Bethylidae), with amended diagnosis of the genus. Zootaxa 3794 (4): 556-564, DOI: 10.11646/zootaxa.3794.4.6
B0587B3CA103FFBEBEDAFBEDFE42F975.taxon	distribution	Distribution. Tanzania.	en	Azevedo, Celso O., Mugrabi, Daniele F. (2014): Three new species of Tuberepyris Lanes et Azevedo (Hymenoptera, Bethylidae), with amended diagnosis of the genus. Zootaxa 3794 (4): 556-564, DOI: 10.11646/zootaxa.3794.4.6
B0587B3CA103FFBABEDAF8E1FA91F865.taxon	description	Description. Female (macropterous form). Body length 2.5 mm. LFW 1.1 mm. Color. Head, mesosoma, scape dark castaneous almost black; antenna, mandible, legs (except tarsi), metasoma dark castaneous; wing venation castaneous; palpi, tarsi light castaneous; wings subhyaline. Head (Figs 1 – 3). Mandible with three apical teeth; lower and median teeth broad and rounded, upper teeth narrow and sharpened. Clypeus short; slightly projected, median lobe trapezoidal, apical margin angularly concave; median elevation and carina absent. Suture fronto-clypeal absent. Eye flat. First four antennomeres in ratio of about 9: 5: 3: 4; flagellomere I 1.0 x as long as wide; flagellomere IX 1.3 x as long as wide. Frons slightly coriaceous with small and sparse punctures. WH 0.80 x LH; WF 1.08 x HE; VOL 1.0 x HE; OOL 2.06 x WOT. Frontal angle of ocellar triangle right; posterior ocellus distant from vertex crest 1.0 x DAO. Vertex straight; corner rounded. First two maxillar palpomeres distinctly shorter and wider than others, III-IV distal ones progressively wider apicad. Labial palpomeres shorter than maxillar ones. Mesosoma (Fig. 1). Pronotal disc 0.62 x as long as wide; anterior and posterior carina absent. Anterior margin of pronotal collar without groove. Mesoscutum with parapsidal furrow 0.43 x as long as mesoscutum. Mesoscutummesoscutellar sulcus absent. Forewing fully developed; 3.5 x as long as wide; subcostal vein 0.26 x as long as LFW; anal vein 0.4 x subcostal vein; pterostigma small and triangular (Fig. 11). Hind wing with tree distal hamuli (Fig. 12). Propodeal disc areolate, anterior portion with elongated cells, 1.18 x as long as wide; lateral margin convergent in dorsal view; transverse posterior and lateral carinae absent; median carina absent; lateral of propodeum coriaceous with spiracles circular; declivity of propodeum coriaceous without median or lateral carinae. Propleuron reaching anterior margin of pronotal disc in dorsal view. Mesopleuron (Fig. 10) bulging; subtegular groove defined only by the upper margin; lower and anterior foveae opened, lower margin of both foveae fused and not surpassing half of mesopleuron length; mesopleural pit inconspicuous and shallow. Profemur 2.25 x longer than wide. Metasoma. Polished, shining, first sternite short, not visible beyond petiole. Tergite II slightly shorter tergite I, posterior margin slightly excavated medially. Female (brachypterous form). Similar to macropterous form, except by: body length 1.67 mm. LFW 0.73 mm. First four antennal segments in ratio of 7: 4: 2: 3; flagellomere I 0.6 x as long as wide. WH 0.85 x LH; WF 1.16 x HE; OOL l. 54 x WOT. Forewing (Fig. 18) short; subcostal vein 0.23 x as long as LFW; anal vein 0.2 x subcostal vein; pterostigma very small and triangular. Hind wing with two hamuli (Fig. 19). Propodeal disc 1.15 x as long as wide, without median discal carina. Profemur 2.12 x longer than wide. Metasoma 1.13 x mesosoma; tergite II shorter tergite I, with posterior margin evenly convex. Male. Body length 1.98 mm. LFW 1.18 mm. Color. Head, mesosoma, scape dark castaneous almost black; antenna, mandible, legs (except tarsi), metasoma dark castaneous; wing venation castaneous; palpi, tarsi light castaneous; wings hyaline. Head (Fig. 13). Mandible with three apical teeth; median tooth narrow and sharp; lower and upper teeth broad and rounded. Clypeus short; slightly projected, median lobe trapezoidal, apical margin angularly concave; median elevation and carina absent. Carina fronto-clypeal present. Eye flat. First four antennomeres in ratio of about 8: 8: 5: 7; flagellomere I 1.25 x as long as wide; flagellomere IX 2.0 x as long as wide. Frons slightly coriaceous with small and sparse punctures. WH 0.92 x LH; WF 1.25 x HE; VOL 0.92 x HE; OOL 1.08 x WOT. Frontal angle of ocellar triangle obtuse; posterior ocellus distant from vertex crest 0.40 x DAO. Vertex straight. First two maxillar palpomeres distinctly shorter and wider than others, III-IV distal ones progressively wider apicad. Labial palpomeres shorter than maxillar ones. Mesosoma (Fig. 13). Pronotal disc 0.48 x as long as wide; anterior and posterior carina absent. Anterior margin of pronotal collar without groove. Mesoscutum with parapsidal furrow 0.42 x as long as mesoscutum. Scutellum without groove neither pits. Forewing fully developed (Fig. 14); subcostal vein 0.25 x as long as LFW; anal vein 0.25 x subcostal vein; pterostigma small and triangular. Hind wing with three distal hamuli (Fig. 15). Propodeal disc areolate, 1.15 x as long as wide; lateral margin convergent in dorsal view; transverse posterior and lateral carinae present; median carina absent; lateral of propodeum imbricate with spiracles circular; declivity of propodeum imbricate without median or lateral carinae. Propleuron reaching anterior margin of pronotal disc in dorsal view. Mesopleuron bulging; subtegular groove defined only by the upper margin; lower and anterior foveae opened, lower margin of both foveae fused and not surpassing half of mesopleuron length; mesopleural pit inconspicuous and shallow. Profemur 2.27 x longer than wide. Metasoma. Polished and shining. Tergite II shorter tergite I, with posterior margin evenly convex. Genitalia (Figs 16 – 17) with paramere slightly shorter than basiparamere, very wide, overlapping each other, dorsal half membranous and translucent, apical margin somewhat truncate, dorsal and ventral margins convex in lateral view; digitus wide, progressively narrowing apicad, apex rounded; cuspis slightly longer than digitus; aedeagus diamond-shaped, with pair of rounded apical lobes, apex surpassing cuspis apex and not reaching paramere apex; genital ring with dorsal half projected basad medially.	en	Azevedo, Celso O., Mugrabi, Daniele F. (2014): Three new species of Tuberepyris Lanes et Azevedo (Hymenoptera, Bethylidae), with amended diagnosis of the genus. Zootaxa 3794 (4): 556-564, DOI: 10.11646/zootaxa.3794.4.6
B0587B3CA103FFBABEDAF8E1FA91F865.taxon	materials_examined	Material examined. Holotype. ♀. MADAGASCAR. Toliara Prov [ince], California Acad [emy] of Siences, Parc National d’Andohahela, Forêt de Manantalinjo, 150 m, 33.6 km 63 º ENE Amboasary, 7.6 km 99 º E Hazofotsy, spiny forest thicket, 24 º 49 ’ 1 ” S 46 º 36 ’ 36 ” E, 12 – 16. I. 2002, sifted litter, Fisher, Griswold et al. coll., California Acad [emy] of Siences, BLF 4810 [code] (CASENT 2111681). Allotype: ♂, MADAGASCAR: Tulear Province, Andohahela Nat’l Park, Ihazofotsey Parcelle III, 24 º 49.85 ’ S 46 º 32.17 ’ E, 20 May- 3 June 2003, California Acad [emy] of Science, colls: M. Irwin, F. Parker, R. Harin’Hala, elev [ation] 80 m, Malaise trap in dry spiny forest, MA – 02 – 21 – 20 (CASENT 2112271). Paratype: ♀, Fiherenana, 100 m, gallery forest, 23 º 10 ’ 37 ” S 43 º 57 ’ 39 ” E, 3 – 7. XI. 2002, sifted litter (leaf mold, rotten wood), Frontier Wildemess Project colls., MGF 043 code (CASENT 2111003).	en	Azevedo, Celso O., Mugrabi, Daniele F. (2014): Three new species of Tuberepyris Lanes et Azevedo (Hymenoptera, Bethylidae), with amended diagnosis of the genus. Zootaxa 3794 (4): 556-564, DOI: 10.11646/zootaxa.3794.4.6
B0587B3CA103FFBABEDAF8E1FA91F865.taxon	discussion	Remarks. Both macropterous and brachypterous forms are very similar to each other. They have VOL 1.0 x HE, the forewing are 3.5 x as long as wide.	en	Azevedo, Celso O., Mugrabi, Daniele F. (2014): Three new species of Tuberepyris Lanes et Azevedo (Hymenoptera, Bethylidae), with amended diagnosis of the genus. Zootaxa 3794 (4): 556-564, DOI: 10.11646/zootaxa.3794.4.6
B0587B3CA103FFBABEDAF8E1FA91F865.taxon	distribution	Distribution. Madagascar.	en	Azevedo, Celso O., Mugrabi, Daniele F. (2014): Three new species of Tuberepyris Lanes et Azevedo (Hymenoptera, Bethylidae), with amended diagnosis of the genus. Zootaxa 3794 (4): 556-564, DOI: 10.11646/zootaxa.3794.4.6
B0587B3CA103FFBABEDAF8E1FA91F865.taxon	etymology	Etymology. The specific epithet nihilus refers to the reduction integument texture and sculpture.	en	Azevedo, Celso O., Mugrabi, Daniele F. (2014): Three new species of Tuberepyris Lanes et Azevedo (Hymenoptera, Bethylidae), with amended diagnosis of the genus. Zootaxa 3794 (4): 556-564, DOI: 10.11646/zootaxa.3794.4.6
B0587B3CA104FFB9BEDAFF51FEBCFAD9.taxon	description	Description. Female. Body length 2.45 mm. LFW 1.16 mm. Body entirely flattened, head 0.3 x as high as long in lateral view. Color. Head, dorsum and lateral of mesosoma, antenna dark castaneous; venter of mesosoma, mandible, legs (except tarsi), wing venation, posterior half of metasoma castaneous; palpi, tarsi, anterior half of metasoma light castaneous; wings hyaline. Head (Figs 4 – 5). Mandible slender, with four visible sharpened apical teeth; lowermost one large. Clypeus short; slightly projected, median lobe trapezoidal with apical margin roundly concave; medially elevated, but not carinate. Carina fronto-clypeal absent. Eye flat, with minute and sparse setae. First four antennomeres in ratio of about 11: 6: 3: 4; flagellomere I as long as wide; flagellomere IX distinctly longer than wide. Frons coriaceous with just few small punctures. LH 1.4 x WH; WF 0.94 x HE; VOL 1.03 x HE; OOL 1.43 x WOT. Frontal angle of ocellar triangle obtuse; ocellar triangle almost touching vertex. Vertex straight; corner rounded. Side of head almost parallel behind eye. Occipital carina absent. First two maxillar palpomeres distinctly shorter and wider than others, III-IV distal ones progressively wider apicad. Labial palpomeres shorter than maxillar ones. Mesosoma (Fig. 4 – 6). Pronotal disc with top surface 2.2 x as long as mesoscutum; ecarinate. Pronotal collar without groove. Mesoscutum with parapsidal furrow narrow, straight, absent on anterior half of mesoscutum, almost reaching tegula. Mesoscutum-mesoscutellar sulcus absent. Forewing fully developed; 3.1 x as long as wide; subcostal vein 0.27 x as long as LFW; median vein present in proximal region, 0.1 x as long as subcostal vein; pterostigma small and triangular (Fig. 21). Hind wing with three distal hamuli (Fig. 22). Propodeal disc coriaceous, 1.15 x as long as median width, progressively narrowing posteriorly; transverse posterior and lateral carinae absent; median carina absent; lateral of propodeum coriaceous with spiracles circular; declivity of propodeum coriaceous without median or lateral carinae. Anterior margin of propleuron almost reaching anterior margin of pronotal disc level in dorsal view. Mesopleuron (Fig. 20) with subtegular groove poorly outlined; lower and anterior foveae absent, mesopleural pit inconspicuous and shallow. Profemur 2.27 x longer than wide. Metasoma. Not petiolate; polished; 1.55 x longer than mesosoma. Tergite II slightly longer tergite I, posterior margin slightly excavated medially. Male unknown.	en	Azevedo, Celso O., Mugrabi, Daniele F. (2014): Three new species of Tuberepyris Lanes et Azevedo (Hymenoptera, Bethylidae), with amended diagnosis of the genus. Zootaxa 3794 (4): 556-564, DOI: 10.11646/zootaxa.3794.4.6
B0587B3CA104FFB9BEDAFF51FEBCFAD9.taxon	materials_examined	Material examined. Holotype. ♀. SOUTH AFRICA, E Cape, Blauwe Krans Farm, (12.8 km 216 º SW Kirkwood), 33 º 30.747 ’ S 25 º 24.644 ’ E, 13 - 15. Feb 2001, S van Noort, Malaise trap, VB 01 - A 3 T-M 43, Valley Bushweld (goat trashed) (SAM-HYM P 043936).	en	Azevedo, Celso O., Mugrabi, Daniele F. (2014): Three new species of Tuberepyris Lanes et Azevedo (Hymenoptera, Bethylidae), with amended diagnosis of the genus. Zootaxa 3794 (4): 556-564, DOI: 10.11646/zootaxa.3794.4.6
B0587B3CA104FFB9BEDAFF51FEBCFAD9.taxon	distribution	Distribution. South Africa.	en	Azevedo, Celso O., Mugrabi, Daniele F. (2014): Three new species of Tuberepyris Lanes et Azevedo (Hymenoptera, Bethylidae), with amended diagnosis of the genus. Zootaxa 3794 (4): 556-564, DOI: 10.11646/zootaxa.3794.4.6
B0587B3CA104FFB9BEDAFF51FEBCFAD9.taxon	etymology	Etymology. The specific epithet codex is a noun in apposition and refers to the short size of the median vein of the forewing.	en	Azevedo, Celso O., Mugrabi, Daniele F. (2014): Three new species of Tuberepyris Lanes et Azevedo (Hymenoptera, Bethylidae), with amended diagnosis of the genus. Zootaxa 3794 (4): 556-564, DOI: 10.11646/zootaxa.3794.4.6
B0587B3CA104FFB8BEDAFA02FAF2FC96.taxon	description	Description. Female. Body length 2.5 mm. LFW 1.5 mm. Body entirely flattened, head 0.36 x as high as long in lateral view. Color. Head and mesosoma dark castaneous almost black; legs (except tarsi), metasoma dark castaneous; antenna, mandible, wing venation castaneous; palpi, tarsi light castaneous; wings subhyaline. Head (Figs 7 – 9). Mandible wide apically, with four visible sharpened apical teeth; lowermost one distinctly larger. Clypeus short; slightly projected, median lobe trapezoidal with rounded corner, medially elevated, but not carinate. Suture fronto-clypeal present. Eye slightly bulging laterad, with minute and sparse setae. First four antennomeres in ratio of about 37: 16: 8: 10; flagellomere I about as long as wide; flagellomere IX distinctly longer than wide. Frons coriaceous with just few small and shallow punctures. LH 1.07 x WH; WF 0.91 x HE; VOL 0.6 x HE; OOL 1.29 x WOT. Frontal angle of ocellar triangle obtuse; ocellar triangle almost touching vertex. Vertex badly convex; corner rounded. Side of head behind eye slightly diverging anterad. Occipital carina absent. First two maxillar palpomeres distinctly shorter and wider than others, III-IV distal ones progressively wider apicad. Labial palpomeres shorter than maxillar ones. Mesosoma (Fig. 7 – 9). Pronotal disc with top surface 1.87 x as long as mesoscutum; ecarinate. Pronotal collar without groove. Mesoscutum with parapsidal furrow narrow, straight, absent on anterior third of mesoscutum. Mesoscutum-mesoscutellar sulcus narrow, angled laterally. Forewing fully developed; 3.05 x as long as wide; subcostal vein 0.28 x as long as LFW; median vein present in apical region, 0.3 x as long as subcostal vein; pterostigma small and subtriangular (Fig. 23). Hind wing with three distal hamuli (Fig. 25). Propodeal disc areolate-coriaceous, 1.12 x as long as median width, progressively narrowing posteriorly; transverse posterior and lateral carinae present; median carina complete and well-defined; lateral of propodeum areolate-coriaceous with circular spiracles; declivity of propodeum areolate-coriaceous without median or lateral carinae. Anterior margin of propleuron almost reaching anterior margin of pronotal disc level in dorsal view. Mesopleuron (Fig. 23) with subtegular groove poorly outlined; lower and anterior foveae absent, mesopleural pit slightly deep. Profemur 2.25 x longer than wide. Metasoma. Not petiolate; polished; 1.28 x longer than mesosoma. Tergite II short, only 0.76 x as long as tergite I, posterior margin slightly excavated medially. Male unknown.	en	Azevedo, Celso O., Mugrabi, Daniele F. (2014): Three new species of Tuberepyris Lanes et Azevedo (Hymenoptera, Bethylidae), with amended diagnosis of the genus. Zootaxa 3794 (4): 556-564, DOI: 10.11646/zootaxa.3794.4.6
B0587B3CA104FFB8BEDAFA02FAF2FC96.taxon	materials_examined	Material examined. Holotype. ♀, CENTRAL AFRICAN REPUBLIC, Prefecture Sangha-Mbaéré, Parc National de Dzanga-Ndoki, Mabéa Bai, 21.4 km 53 º NE Bayanga, 3 º 02.01 ’ N 16 º 24.57 ’ E, 510 m, 2 - 3. v. 2001, S van Noort, Malaise trap, CAR 01 - M 16, Lowland Rainforest, marsh clearing (ISAM).	en	Azevedo, Celso O., Mugrabi, Daniele F. (2014): Three new species of Tuberepyris Lanes et Azevedo (Hymenoptera, Bethylidae), with amended diagnosis of the genus. Zootaxa 3794 (4): 556-564, DOI: 10.11646/zootaxa.3794.4.6
B0587B3CA104FFB8BEDAFA02FAF2FC96.taxon	distribution	Distribution. Central African Republic.	en	Azevedo, Celso O., Mugrabi, Daniele F. (2014): Three new species of Tuberepyris Lanes et Azevedo (Hymenoptera, Bethylidae), with amended diagnosis of the genus. Zootaxa 3794 (4): 556-564, DOI: 10.11646/zootaxa.3794.4.6
B0587B3CA104FFB8BEDAFA02FAF2FC96.taxon	etymology	Etymology. The specific epithet hamus is a noun in apposition and refers to the shaped of wing venation.	en	Azevedo, Celso O., Mugrabi, Daniele F. (2014): Three new species of Tuberepyris Lanes et Azevedo (Hymenoptera, Bethylidae), with amended diagnosis of the genus. Zootaxa 3794 (4): 556-564, DOI: 10.11646/zootaxa.3794.4.6
