taxonID	type	description	language	source
B0685C00FFAA0908C98AFBFDFC3610D9.taxon	diagnosis	Diagnosis. The adults of Berotha are characterized by the slightly falcate forewing, the long antennal scape with a length-width ratio generally of 3: 1 to 5: 1, and the complex of male fused gonocoxites, gonapophyses and gonostyli 10 (= paramere-mediuncus complex) with a basal sclerite and a bow-shaped structure formed by bundled bristles (hereafter often referred to as a bristle-bow).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFAA0908C98AFBFDFC3610D9.taxon	distribution	Distribution. China; India; Indonesia; Malaysia; Philippines; Sri Lanka.	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA9090AC98AFE94FC8510F1.taxon	description	(Figs. 1 – 2)	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA9090AC98AFE94FC8510F1.taxon	diagnosis	Diagnosis. This species is characterized by the male tergum 9 + ectoprocts rounded distally, the sternum 9 in lateral view distinctly tapering posteriad, the complex of fused gonocoxites, gonapophyses and gonostyli 10 with distinct bristle-bow, reaching to segment 8 anteriorly.	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA9090AC98AFE94FC8510F1.taxon	description	Description. Male. Body length 7.0 mm; forewing length 11.5 mm, hindwing length 10.0 mm. Head generally yellowish brown with dark brown spots and yellow setae. Compound eyes blackish brown. Antennae yellow with yellowish setae; scape slightly darker, with brown spots on distal margin, and with brownish setae ventrally, length-width ratio approximately 4: 1. Prothorax generally yellowish brown with dark brown spots, setae yellow dorsally but dark brown ventrally; pronotum with dense blackish brown spots laterally, and with one black longitudinal marking medially. Mesothorax yellowish brown dorsally, metathorax brown. Legs yellow with dark brown spots and brown setae. Forewing falcate on apex, slightly smoky brown, but darker along distal margin, with brown spots on branching points of longitudinal veins; pterostigma narrowly elongate, reddish brown, with hyaline parts at middle. Longitudinal veins pale yellow, interrupted by many brownish spots; crossveins brown. Hindwing slightly paler than forewing, immaculate. Abdomen brown with pale yellow setae. Male genitalia. Tergum 9 + ectoprocts broad, rounded distally. Sternum 9 in lateral view tapering posteriad, subtriangular. Complex of fused gonocoxites, gonapophyses and gonostyli 10 (= paramere-mediuncus complex) forming a short bow formed by bundled bristles, reaching to segment 8 anteriorly. Hypandrium internum small, subtriangular. Female. Body length 6.0 mm; forewing length 11.0 mm, hindwing length 10.2 mm. Female genitalia. Hypocaudae almost twice as long as basal part of gonocoxites 9. Bursa-copulatrixspermatheca complex comprising a small but strongly sclerotized ball-shaped element and a basal coil with varing lumens in width.	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA9090AC98AFE94FC8510F1.taxon	materials_examined	Materials examined. Holotype ♂, China: Fujian, Dehua, Shuikou [25 ° 70.060 ' N, 118 ° 45.242 ' E], 6. XI. 1974, Chikun Yang (CAU). Paratype ♀, same data as holotype (CAU).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA9090AC98AFE94FC8510F1.taxon	distribution	Distribution. China (Fujian).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA9090AC98AFE94FC8510F1.taxon	discussion	Remarks. The genitalia of both male and female type specimens are lost. No additional specimens of this species were available for our study. The above description was made according to the remaining parts of the type specimens and the original description in Yang & Liu (1999).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA8090DC98AFA14FE7B12AE.taxon	description	(Figs. 3 – 4, 12 – 18)	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA8090DC98AFA14FE7B12AE.taxon	diagnosis	Diagnosis. This species is characterized by the male tergum 9 + ectoprocts rounded distally, the sternum 9 short, posteriorly truncate, the complex of fused gonocoxites, gonapophyses and gonostyli 10 with distinct bristle-bow, reaching to segment 8 anteriorly.	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA8090DC98AFA14FE7B12AE.taxon	description	Description. Male. Body length 8.3 mm; forewing length 13.0 mm, hindwing length 11.6 mm. Head generally pale yellow with brownish setae, blackish brown dotted on vertex. Compound eyes blackish brown. Antennae pale yellow with yellowish setae, but setae on scape and pedicel pale brown and longer than flagellum; scape and pedicel with blackish brown spots; scape long and thick, length-width ratio approximately 5: 1. Prothorax generally pale yellow with blackish brown spots, setae yellow dorsally but dark brown ventrally; pronotum with dense blackish brown spots. Meso- and metathorax pale yellow with blackish brown spots dorsally. Legs pale yellow with dark brown spots and generally brown setae. Forewing distinctly convex in pterostigmal area, and falcate on apex, slightly smoky brown, slightly darker along distal margin, with brown spots on branching points of longitudinal veins; pterostigma narrowly elongate, brown, with hyaline parts at middle, paler on distal margin. Longitudinal veins pale yellow but paler on distal margin, interrupted by many brownish spots; crossveins brown. Hindwing slightly paler than forewing, immaculate. Abdomen yellow with blackish brown spots and pale yellow setae. Male genitalia. Tergum 9 + ectoprocts rounded distally in lateral view. Trichobothria reduced. Sternum 9 short, shorter than sternum 8, posteriorly truncate. Gonocoxites 9 paired, rod-shaped proximally, with hook-like apex. Complex of fused gonocoxites, gonapophyses and gonostyli 10 (= paramere-mediuncus complex) with a basal sclerite and a short bow formed by bundled bristles, reaching to segment 8 anteriorly. Fused gonocoxites 11 (= gonarcus) forming an arched sclerite. Hypandrium internum subtriangular. Female. Body length 6.8 mm; forewing length 9.5 mm, hindwing length 7.5 mm. Female genitalia. Sternum 7 mostly reduced, gonocoxites 7 present as paired domed sclerites, bearing long setae. Gonapophyses 7 as a pair of smooth sclerite discs. Gonocoxites 8 in ventral view narrow, ribbon-shaped, strongly sclerotized, medially with paired membranous processus directed anteroventrad, nearly subtriangular. Gonapophyses 8 (= pudiculum) forming a tiny arcuate sclerite ventrally. Tergum 9 + ectoprocts slightly shorter than tergum 8, ventrally tapering and bearing long setae. Gonocoxites 9 paired, club-shaped, bearing long setae; hypocaudae almost 1.5 times as long as basal part of gonocoxites 9. Bursa-copulatrix-spermatheca complex comprising a small but strongly sclerotized ball-shaped element and a basal coil with lumens of different width.	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA8090DC98AFA14FE7B12AE.taxon	materials_examined	Materials examined. 1 ♂, China: Yunnan, Mengla [21 ° 46.528 ' N, 101 ° 57.153 ' E], 800 m, 11. IV. 1981, Fasheng Li (CAU); 1 ♀, China: Yunnan, Menghai [21 ° 96.343 ' N, 100 ° 45.944 ' E], 18. IV. 1981, Chikun Yang (CAU).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA8090DC98AFA14FE7B12AE.taxon	distribution	Distribution. China (Yunnan).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA8090DC98AFA14FE7B12AE.taxon	discussion	Remarks. The primary type of this species is a female specimen, which was collected in Menghai, Yunnan on April 17, 1981. Unfortunately, the primary type cannot be currently located. We examined one male and one female deposited in CAU. They were collected from Menghai and Mengla of Yunnan in 1981, and probably belong to the specimens of B. bannana which were originally described by Yang (1986). Although the presently examined specimens are not the types, the identification of them as B. bannana could be verified based on the characters as described in Yang (1986).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFAE090EC98AFC28FBC21782.taxon	description	(Figs. 5 – 6, 19 – 23, 35 – 37)	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFAE090EC98AFC28FBC21782.taxon	diagnosis	Diagnosis. The species is characterized by the male sternum 8 posteriorly with a median digitiform processus directed posteroventrad, the largely paired male sternum 9 with one to four little teeth on each lateral lobe, the ventral portion of gonocoxites 9 in lateral view nearly right-angled and the presence of large bristle-bow of complex of fused gonocoxites, gonapophyses and gonostyli 10 (= paramere-mediuncus complex).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFAE090EC98AFC28FBC21782.taxon	description	Description. Male. Body length 6.0 – 7.0 mm; forewing length 9.2 – 10.5 mm, hindwing length 8.0 – 10.0 mm. Head generally yellow with blackish brown spots; setae yellow dorsally but dark brown on frons and clypeus. Compound eyes blackish brown. Antennae pale yellow with yellowish setae; setae on scape and pedicel longer than those of flagellum, yellow dorsally, dark brown ventrally; scape long and thick, length-width ratio approximately 5: 1. Prothorax generally pale yellow with blackish brown setae ventrally. Meso- and metathorax slightly paler than prothorax dorsally; pro- and mesonotum with blackish brown spots. Legs pale yellow with brown setae, tibia and tarsus with blackish brown spots. Forewing falcate on apex, slightly smoky brown, slightly darker along distal margin and crossveins; pterostigma narrowly elongate, reddish brown, with hyaline parts at middle. Longitudinal veins pale yellow, interrupted by many brownish spots, darker on branching points of longitudinal veins; crossveins brown. Hindwing slightly paler than forewing, immaculate; longitudinal veins pale brown, crossveins dark brown. Abdomen yellow with blackish brown spots and black brown setae; each pregenital segment darker on posterior margin. Male genitalia. Sternum 8 posteriorly with a median digitiform processus directed posteroventrad. Tergum 9 + ectoprocts in lateral view nearly shell-shaped. Trichobothria reduced. Sternum 9 largely paired, with subtriangular lateral lobes, which are connected by a sclerotized bar, apex of lateral lobes with one to four little teeth. Gonocoxites 9 paired, with hook-like apex. Complex of fused gonocoxites, gonapophyses and gonostyli 10 (= paramere-mediuncus complex) forming a basal sclerite and a long bow formed by 7 – 8 long bundled bristles, reaching to segment 7 anteriorly and ending as a bundle of free bristles terminally. Fused gonocoxites 11 (= gonarcus) arched, reaching to segment 8. Hypandrium internum large, subtriangular. Female. Body length 7.8 mm; forewing length 11.8 mm, hindwing length 10.5 mm. Female genitalia. Sternum 7 mostly reduced. Gonocoxites 7 present as paired domed sclerites, bearing long setae. Gonapophyses 7 as a pair of smooth sclerite discs. Gonocoxites 8 in ventral view narrow, ribbon-shaped, strongly sclerotized, with paired membranous processus directed anteroventrad, nearly semicircular. Gonapophyses 8 (= pudiculum) forming a tiny arc-shaped sclerite in ventral view. Tergum 9 + ectoprocts slightly shorter than tergum 8, ventrally slightly tapering and bearing long setae. Gonocoxites 9 paired, club-shaped, bearing long setae; hypocaudae almost 5 times as long as basal part of gonocoxites 9 in ventral view. Bursacopulatrix-spermatheca complex comprising a small but strongly sclerotized globular element, and a basal coil with lumens of different width, ending as broad bursa.	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFAE090EC98AFC28FBC21782.taxon	materials_examined	Materials examined. 1 ♂, China: Hainan, Mt. Limushan [19 ° 26.772 ' N, 109 ° 78.684 ' E], 3. IV. 2013, Wang & Jiang (CAU); 1 ♂, 1 ♀, China: Hainan, Mt. Diaoluoshan [18 ° 79.376 ' N, 109 ° 87.094 ' E], 700 m, 09. V. 2011, Wenxuan Bi (CAU); 1 ♂, China: Hainan, Mt. Diaoluoshan [18 ° 79.376 ' N, 109 ° 87.094 ' E], 900 m, 30. V. 2007, Jingxian Liu (CAU); 1 ♀, China: Hainan, Jianfengling [18 ° 71.495 ' N, 108 ° 87.511 ' E], 26. II. 1982, Chikun Zhang (CAU).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFAE090EC98AFC28FBC21782.taxon	distribution	Distribution. China (Hainan).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFAE090EC98AFC28FBC21782.taxon	discussion	Remarks. This species appears to be closely related to B. spetana from Taiwan in having similar male sternum 8 posteriorly with a median digitiform processus, the largely paired male sternum 9 with little teeth on each lateral lobe, the complex of fused gonocoxites, gonapophyses and gonostyli 10 forming a large bristle-bow, reaching to segment 7 anteriorly and the female fused gonocoxites 8 in ventral view narrow and ribbon-shaped, with paired membranous semicircular processus directed anteroventrad, but it can be distinguished from the latter species by the male gonocoxites 9 ventral portion in lateral view nearly right-angled, the processus of sternum 8 in ventral view extremely short and the female tergum 9 + ectoprocts in lateral view tapering ventrally. In B. spetana the male gonocoxites 9 ventral portion in lateral view nearly arcuately convex, the processus of sternum 8 distinct and the female tergum 9 + ectoprocts in lateral view nearly square. It is notable that the number of little teeth on lateral lobes of male sternum 9 is variable among conspecific individuals. Of the three specimens we examined, the left lateral lobe of male sternum 9 bears one, two or four teeth, while the right lateral lobe bears three, three or four teeth (Figs. 35 – 37).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFAC090EC98AFC85FEE411CC.taxon	diagnosis	Diagnosis. The species is characterized by the male sternum 8 posteriorly with a median digitiform processus directed posteroventrad; the largely paired male sternum 9 with two or three little teeth on each lateral lobe; the ventral portion of gonocoxites 9 in lateral view nearly arcuately convex and the presence of large bristle-bow complex of fused gonocoxites, gonapophyses and gonostyli 10 (= paramere-mediuncus complex).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFAC090EC98AFC85FEE411CC.taxon	distribution	Distribution. China (Taiwan).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFAC090EC98AFC85FEE411CC.taxon	discussion	Remarks. This species was sufficiently described in U. Aspöck et al. (2013) and a re-description here is unnecessary.	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFAC0901C98AFAD0FF2B158A.taxon	diagnosis	Diagnosis. This species is characterized by the male tergum 9 + ectoprocts in lateral view with posterodorsal portion nearly right-angled, and with posteroventral portion arcuately convex; the complex of fused gonocoxites, gonapophyses and gonostyli 10 with short bristle-bow, reaching to segment 9 anteriorly.	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFAC0901C98AFAD0FF2B158A.taxon	description	Description. Male. Body length 7.0 – 9.0 mm; forewing length 11.0 mm, hindwing length 10.0 mm. Head generally yellowish brown with brown spots and brown setae; setae longer ventrally. Compound eyes blackish brown. Antennae long, scape with long setae, length-width ratio approximately 3: 1. Prothorax generally yellowish, with dense brown spots and setae dorsally. Pronotum with dense blackish setae ventrally. Legs yellowish brown. Forewing falcate on apex, slightly smoky yellow; pterostigma reddish brown. Longitudinal veins pale yellow, interrupted by many brownish spots; crossveins brown. Hindwing slightly paler than forewing, with brown spots only on pterostigma and some branching points of longitudinal veins. Abdomen pale brown dorsally and yellowish brown ventrally. Male genitalia. Tergum 9 + ectoprocts in lateral view with posterodorsal portion nearly right-angled and posteroventral portion arcuately convex. Complex of fused gonocoxites, gonapophyses and gonostyli 10 (= paramere-mediuncus complex) with a basal sclerite and a short bow formed by bundled bristles, reaching to segment 9 anteriorly. Hypandrium internum extremely small. Female. Unknown.	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFAC0901C98AFAD0FF2B158A.taxon	distribution	Distribution. China (Zhejiang).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFAC0901C98AFAD0FF2B158A.taxon	discussion	Remarks. The primary types of this species cannot be found currently. No additional specimens of this species were available for our study. The above description was made according to the original description in Yang & Liu (1995).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA30901C98AFE8DFEC8170F.taxon	diagnosis	Diagnosis. This species is characterized by the male’s extremely tiny bristle-bow complex of fused gonocoxites, gonapophyses and gonostyli 10 (= paramere-mediuncus complex).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA30901C98AFE8DFEC8170F.taxon	distribution	Distribution. China (Taiwan).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA30901C98AFE8DFEC8170F.taxon	discussion	Remarks. This species was sufficiently described in U. Aspöck et al. (2013), a re-description is unnecessary in this paper.	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA30900C98AFD10FC4C12BA.taxon	description	(Figs. 7, 24 – 26)	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA30900C98AFD10FC4C12BA.taxon	diagnosis	Diagnosis. The new species is characterized by the male tergum 9 + ectoprocts rounded distally, the sternum 9 short, distinctly concaved posteriad; the complex of fused gonocoxites, gonapophyses and gonostyli 10 with distinct bristle-bow, reaching to segment 8 anteriorly.	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA30900C98AFD10FC4C12BA.taxon	description	Description. Male. Body length 8.2 mm; forewing length 12.0 mm, hindwing length 10.5 mm. Head generally pale yellow with blackish brown spots. Compound eyes blackish brown. Antennae pale yellow with yellowish setae, but setae on scape and pedicel pale brown and longer than flagellum; scape and pedicel with blackish brown spots on distal margin; scape long and thick, length-width ratio approximately 5: 1. Prothorax generally pale yellow with blackish brown spots; setae yellow dorsally but dark brown ventrally; pronotum with dense blackish brown spots. Meso- and metathorax generally yellow, mesonotum with blackish brown spots medially, metanotum immaculate. Legs pale yellow with dark brown spots and brown setae. Forewing distinctly convex in pterostigmal area, and falcate on apex, slightly smoky brown, slightly darker along distal margin, with brown spots on branching points of longitudinal veins; pterostigma narrowly elongate, reddish brown, with hyaline parts at middle, paler on distal margin. Longitudinal veins pale yellow but paler on distal margin, interrupted by many brownish spots; crossveins brown. Hindwing slightly paler than forewing. Abdomen yellow with blackish brown spots and pale yellow setae. Male genitalia. Tergum 9 + ectoprocts subquadrate in lateral view. Trichobothria reduced. Sternum 9 short, slightly shorter than sternum 8, distinctly concaved posteriad. Gonocoxites 9 paired, rod-shaped proximally, with hook-like apex. Complex of fused gonocoxites, gonapophyses and gonostyli 10 (= paramere-mediuncus complex) with a basal sclerite and a short bow formed by bundled bristles, reaching to segment 8 anteriorly. Fused gonocoxites 11 (= gonarcus) forming an arched sclerite. Hypandrium internum subtriangular. Female. Unknown.	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA30900C98AFD10FC4C12BA.taxon	materials_examined	Material examined. Holotype ♂, China: Guangdong, Mt. Dinghushan Nature Reserve [23 ° 16.592 ' N, 112 ° 54.091 ' E], 36 m, 27. VI. 2017, light trap, Yisheng Zhao (CAU).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA30900C98AFD10FC4C12BA.taxon	distribution	Distribution. China (Guangdong).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA30900C98AFD10FC4C12BA.taxon	etymology	Etymology. The new species is named after the Guangdong Province where it was discovered.	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA30900C98AFD10FC4C12BA.taxon	discussion	Remarks. This species appears to be closely related to B. bannana from Yunnan in having similar gonocoxites 9 of hook-like apex, the complex of fused gonocoxites, gonapophyses and gonostyli 10 (= paramere-mediuncus complex) of a short bristles-bow, the fused gonocoxites 11 (= gonarcus) of arched sclerite and the hypandrium internum subtriangular, but it can be distinguished from the latter species by the male sternum 9 posteriorly concaved. In B. bannana the male sternum 9 is posteriorly truncate.	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA10903C98AF9A3FD5E1284.taxon	diagnosis	Diagnosis. The adults of Isoscelipteron are characterized by the strongly falcate forewings with inconspicuous pterostigma and the complex of fused gonocoxites, gonapophyses and gonostyli 10 (= paramere-mediuncus complex) with large spiral part composed of numerous loops.	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA10903C98AF9A3FD5E1284.taxon	distribution	Distribution. Albania; Australia (Queensland); Bulgaria; China; Cyprus; France; Greece; India; Indonesia; Iran; Israel; Italy; Japan; Lebanon; Macedonia; Malaysia (Borneo); Morocco; Philippines; Portugal; Romania; Solomon Islands; Spain; Turkey and Vietnam.	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA00904C98AF8B5FAF11244.taxon	description	(Figs. 8, 27 – 28)	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA00904C98AF8B5FAF11244.taxon	diagnosis	Diagnosis. This species is characterized by the male tergum 9 + ectoprocts in lateral view posteriorly arcuately convex, the fused gonocoxites 11 (= gonarcus) medially with a ring-shaped lobe, the torulus slightly arched.	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA00904C98AF8B5FAF11244.taxon	description	Description. Male. Body length 7.0 – 7.5 mm; forewing length 9.0 – 10.0 mm, hindwing length 7.0 – 10.0 mm. Head generally brownish yellow with yellowish setae, but setae dark brown on frons and clypeus. Compound eyes blackish brown. Antennae yellow with yellowish setae; setae on scape longer and darker than those on pedicel and flagellum; scape thick, length-width ratio approximately 3: 1. Prothorax generally pale brown with setae yellowish dorsally, but darker ventrally; pronotum pale brown, laterally with blackish brown spots. Meso- and metanotum pale brown; mesonotum with some blackish brown spots medially; metanotum immaculate. Legs pale yellow with pale brown setae; tibia and tarsus with dark brown spots, profemur with several pale brown spots; metafemur proximally with one dark brown spot. Forewing falcate on apex, slightly smoky brown, slightly darker along posterior margin, with brown spots on branching points of several longitudinal veins; pterostigma indistinct, brown, but some individuals with some reddish brown spots. Longitudinal veins pale yellow, interrupted by many brownish spots; crossveins brown. Hindwing slightly paler than forewing, immaculate. Abdomen generally yellow, tergum pale brown with some brown spots. Male genitalia. Tergum 9 + ectoprocts slightly longer than tergum 8, in lateral view posteriorly arcuately convex. Trichobothria reduced. Sternum 9 slightly shorter than sternum 8, slightly concaved posteriad. Hypomeres membranous, sack-shaped, directed posteriorly and tapering distally. Gonocoxites 9 paired, with hook-like apex. Complex of fused gonocoxites, gonapophyses and gonostyli 10 (= paramere-mediuncus complex) with large spiral part composed of nearly 14 loops, anteriorly reaching to segment 7, posteriorly ending at segment 9. Fused gonocoxites 11 (= gonarcus) in lateral view nearly rod-shaped, in ventral view with a ring-shaped sclerite medially. Torulus arched. Hypandrium internum large. Female genitalia (description after Yang & Liu 1995). Tergum 9 + ectoprocts ventrally slightly tapering and bearing long setae. Gonocoxites 9 paired, club-shaped; hypocaudae almost twice as long as basal part of gonocoxites 9 in lateral view. Bursa-copulatrix-spermatheca complex comprising a small but strongly sclerotized globular element and a basal coil with varying lumen in width.	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA00904C98AF8B5FAF11244.taxon	materials_examined	Materials examined. 1 ♂, China: Zhejiang, Mt. Tianmushan [30 ° 31.635 ' N, 119 ° 44.619 ' E], 14. IX. 2012 (CAU); 1 ♂, China: Anhui, Mt. Huangshan, Yungu Temple [30 ° 11.966 ' N, 118 ° 19.285 ' E], 890 m, 22. VII. 1977, Chikun Yang (CAU); 3 ♂, China: Zhejiang, 21. VII. 1995, Malaise trap, Chikun Yang (CAU); 11 ♂, China: Zhejiang, 2. IX. 1995, Malaise trap, Chikun Yang (CAU).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA00904C98AF8B5FAF11244.taxon	distribution	Distribution. China (Zhejiang, Anhui).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA00904C98AF8B5FAF11244.taxon	discussion	Remarks. The primary types of this species cannot currently be found. The types were collected from Mt. Gutianshan, Zhejiang, on August 2, 1992. The specimens herein examined were collected from adjacent areas near the type locality of I. dictyophilum and show identical morphological characters to that in type specimens (see Yang & Liu 1995), which thereby confirmed our identification of this species as I. dictyophilum. Yang & Liu (1995) noted that this species is usually attracted by the Malaise trap but the females are rarely collected. All specimens we examined were males and no female specimens were available for our study.	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA60907C98AF848FEB6133A.taxon	diagnosis	Diagnosis. This species is characterized by the male tergum 9 + ectoprocts posteriorly truncate and the gonocoxites 11 narrow medially (Yang & Liu 1999).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA60907C98AF848FEB6133A.taxon	description	Description. Male. Body length 8.0 mm; forewing length 11.0 mm, hindwing length 9.0 mm. Head generally yellowish brown. Compound eyes blackish brown. Labrum with a pair of brown large spots. Antennae brown; scape thick, length-width ratio approximately 2: 1. Prothorax yellowish brown with some spots; pronotum pale brown. Legs yellow with long setae, tibia with many brown spots. Forewing falcate on apex, slightly smoky yellow. Longitudinal veins yellowish brown, interrupted by many brownish spots; crossveins brown. Hindwing slightly paler than forewing, veins yellowish brown. Abdomen brown. Male genitalia. Tergum 9 + ectoprocts posteriorly truncate. Complex of fused gonocoxites, gonapophyses and gonostyli 10 (= paramere-mediuncus complex) with large spiral part composed of nearly 15 loops, anteriorly reaching to segment 7, posteriorly ending to segment 9. Gonocoxites 11 (= gonarcus) narrow at middle.	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA60907C98AF848FEB6133A.taxon	distribution	Distribution. China (Fujian). Remarks. The primary types of this species cannot currently be found. No specimens of this species were available for our study. The above description was made according to the original description and illustration in Yang & Liu (1999).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA50907C98AF95AFC451283.taxon	diagnosis	Diagnosis. This species is characterized by the male tergum 9 + ectoprocts in lateral view slightly elongated and tapering posteroventrad, the sternum 9 distinctly concaved posteriad, the hypomeres subquadrate, the gonocoxites 11 (= gonarcus) arched.	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA50907C98AF95AFC451283.taxon	distribution	Distribution. China (Taiwan).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA50907C98AF95AFC451283.taxon	discussion	Remarks. The male of this species was described in U. Aspöck & H. Aspöck (1980) and the female was described in U. Aspöck et al. (2013), re-description is unnecessary.	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA40918C98AFA14FB2114FA.taxon	description	(Figs. 9 – 10, 29 – 32)	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA40918C98AFA14FB2114FA.taxon	diagnosis	Diagnosis. This species is characterized by the male tergum 9 + ectoprocts posteroventrally with short obtuse processus, the fused gonocoxites 11 (= gonarcus) with median incision, and the torulus subquadrate.	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA40918C98AFA14FB2114FA.taxon	description	Description. Male. Body length 9.0 – 10.0 mm; forewing length 12.5 – 13.5 mm, hindwing length 10.5 – 11.5 mm. Head generally pale yellow with yellowish setae, but setae on frons and clypeus pale brown. Compound eyes blackish brown. Antennae yellow with yellowish setae; setae on scape longer than those on pedicel and flagellum; scape thick, length-width ratio approximately 3: 1. Prothorax generally pale yellow; pronotum laterally brown with several reddish brown spots. Meso- and metathorax pale yellowish brown with yellowish setae, meso- and metanotum laterally brown. Legs pale yellow, tibia distally with some dark brown spots; setae generally pale yellow, but pale brown on tibia. Forewing distinctly convex in pterostigmal area, and falcate on apex, slightly smoky brown and darker along distal margin, with grayish brown spots on branching points of many longitudinal veins and along outer gradate series; pterostigma narrowly elongate, slightly reddish brown, with some brownish spots; setae grayish on inner margin, yellowish on outer margin. Longitudinal veins pale yellow, interrupted by many brownish spots; crossveins dark brown. Hindwing slightly paler than forewing, immaculate. Abdomen yellowish brown, pregenital segments dorsally brown. Male genitalia. Tergum 9 + ectoprocts as long as tergum 8, in lateral view nearly semicircular, posteroventrally produced into a short obtuse processus. Sternum 9 slightly shorter than sternum 8, slightly concaved posteriad. Gonocoxites 9 paired, with hook-like apex. Complex of fused gonocoxites, gonapophyses and gonostyli 10 (= paramere-mediuncus complex) with large spiral part composed of numerous loops, anteriorly reaching into segment 7. Fused gonocoxites 11 (= gonarcus) arched, with median incision. Torulus subquadrate. Hypandrium internum large subtriangular. Female. Body length 6.6 – 9.0 mm; forewing length 11.0 – 13.0 mm, hindwing length 10.0 – 11.5 mm. Female genitalia. Sternum 7 mostly reduced, gonocoxites 7 present as a pair of domed sclerites; gonapophyses 7 as hairless plates. Gonocoxites 8 ventrally with a pair of membranous digitiform processus directed anteroventrad, connected by a tiny transverse sclerite. Gonapophyses 8 forming a tiny sclerite (pudiculum). Tergum 9 + ectoprocts nearly as long as tergum 8, ventrally tapering and bearing long setae. Gonocoxites 9 paired, club-shaped; hypocaudae almost three times as long as basal part of gonocoxites 9 in ventral view. Bursacopulatrix-spermatheca complex comprising a small but strongly sclerotized globular element and a basal coil with varying lumen in widths.	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA40918C98AFA14FB2114FA.taxon	materials_examined	Materials examined. 1 ♂, China: Shandong, Mt. Kunyushan, Taiboding [37 ° 30.117 ' N, 121 ° 74.140 ' E], 18. VIII. 2014, Tingting Zhang (CAU); 2 ♂, China: Shandong, Mt. Kunyushan, Taiboding [37 ° 30.117 ' N, 121 ° 74.140 ' E], IX. 2014, Malaise trap, Tingting Zhang (CAU); 2 ♂, China: Guizhou, Mt. Yulongshan [27 ° 73.054 ' N, 107 ° 64.168 ' E], VIII. 2017, Malaise trap, Mengqing Wang (CAU); 2 ♂, China: Zhejiang, 21. VII. 1995, Malaise trap, Chikun Yang (CAU); 1 ♂, China: Zhejiang, 18. VII. 1995, Malaise trap, Chikun Yang (CAU); 1 ♂, China: Zhejiang, 10. IX. 1995, Malaise trap, Chikun Yang (CAU); 5 ♀, China: Shandong, Mt. Kunyushan [37 ° 30.117 ' N, 121 ° 74.140 ' E], 09. V. 2011, Malaise trap, Tingting Zhang (CAU); 1 ♀, China: Guizhou, Mt. Fanjingshan [27 ° 69.965 ' N, 108 ° 83.955 ' E], 02. VIII. 2001, Xinli Wang (CAU); 1 ♀, China: Sichuan, Mt. Emeishan, Qingyinge [29 ° 57.572 ' N, 103 ° 40.711 ' E], 710 m, 27. VIII. 1961, Chikun Yang (CAU); 1 ♀, China: Zhejiang, 2. IX. 1995, Malaise trap, Chikun Yang (CAU).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA40918C98AFA14FB2114FA.taxon	distribution	Distribution. China (Guizhou, Shandong, Shanghai, Sichuan, Zhejiang).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFA40918C98AFA14FB2114FA.taxon	discussion	Remarks. The holotype of this species is a female from Shanghai and is deposited in the Muséum National d'Histoire Naturelle, Paris. It has been studied, redescribed, and illustrated by U. Aspöck & H. Aspöck (1991). Moreover, in that paper also a male from Shanghai has been described and illustrated in detail. Based on these descriptions and illustrations, the present material in this paper can be clearly assigned to I. pectinatum. Aside from Shanghai, Shandong and Zhejiang (both near the type locality), I. pectinatum is now known also from Guizhou and Sichuan in southwestern China, thus it can be considered a widespread species.	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFBA0918C98AFA53FCD5130E.taxon	distribution	Distribution. China (Taiwan).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFBA0918C98AFA53FCD5130E.taxon	discussion	Remarks. Primary type material has probably been lost. This species is possibly identical with I. formosense (Krüger, 1922) (see discussion in U. Aspöck et al. 2013).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFBA091DC98AF90EFC791411.taxon	description	(Figs. 11, 33 – 34)	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFBA091DC98AF90EFC791411.taxon	diagnosis	Diagnosis. The new species is characterized by the male tergum 9 + ectoprocts with posterodorsal portion nearly right-angled and with posteroventral portion slightly elongated and tapering, the fused gonocoxites 11 (= gonarcus) medially with a ring-shaped lobe, and the torulus broadly arched.	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFBA091DC98AF90EFC791411.taxon	description	Description. Male. Body length 7.0 – 7.5 mm; forewing length 9.0 – 10.0 mm, hindwing length 7.0 – 10.0 mm. Head generally brownish yellow with yellowish setae, but setae on frons and clypeus dark brown. Compound eyes blackish brown. Antennae yellow with yellowish setae; setae on scape longer and darker than those of pedicel and flagellum; scape thick, length-width ratio approximately 3: 1. Prothorax generally pale brown with yellowish setae but darker ventrally, pronotum pale brown, laterally with blackish brown spots. Meso- and metathorax pale brown, mesonotum with some blackish brown spots medially; metanotum immaculate. Legs pale yellow with pale brown setae; tibia and tarsus with dark brown spots, profemur with several pale brown spots; metafemur proximally with a dark brown spot. Forewing falcate on apex, slightly smoky brown but slightly darker along posterior margin, with brown spots on branching points of several longitudinal veins; pterostigma indistinct, brown. Longitudinal veins pale yellow, interrupted by many brownish spots; crossveins brown. Hindwing slightly paler than forewing, immaculate. Abdomen generally yellow, tergum pale brown with some brown spots. Male genitalia. Tergum 9 + ectoprocts with posterodorsal portion nearly right-angled, slightly elongated and tapering posteroventrad. Trichobothria reduced. Sternum 9 slightly shorter than sternum 8, slightly concaved posteriad. Hypomeres membranous, directed posteriorly and tapering distally. Gonocoxites 9 paired, with hooklike apex. Complex of fused gonocoxites, gonapophyses and gonostyli 10 (= paramere-mediuncus complex) with large spiral part composed of nearly 14 loops, anteriorly reaching to segment 7, posteriorly ending at segment 9. Fused gonocoxites 11 (= gonarcus) in lateral view nearly rod-shaped, in ventral view medially with a ring-shaped lobe. Torulus broadly arched. Hypandrium internum large. Female. Unknown.	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFBA091DC98AF90EFC791411.taxon	materials_examined	Materials examined. Holotype ♂, China: Guizhou, Mt. Yulongshan [27 ° 73.054 ' N, 107 ° 64.168 ' E], VIII. 2017, Malaise trap, Mengqing Wang (CAU). Paratypes: 1 ♂, China: Guizhou, Mt. Yulongshan [27 ° 73.054 ' N, 107 ° 64.168 ' E], IX. 2017, Malaise trap, Mengqing Wang (CAU); 1 ♂, China: Guizhou, Tiancheng [27 ° 78.845 ' N, 107 ° 59.255 ' E], IX. 2017, Malaise trap, Mengqing Wang (CAU).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFBA091DC98AF90EFC791411.taxon	distribution	Distribution. China (Guizhou).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFBA091DC98AF90EFC791411.taxon	etymology	Etymology. The species name refers to the shape of tergum 9 + ectoprocts (lat. acutus = acute, caudatus = tailed).	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
B0685C00FFBA091DC98AF90EFC791411.taxon	discussion	Remarks. This species appears to be closely related to I. dictyophilum from Zhejiang and Anhui in having generally similar male genitalia, but it can be distinguished from the latter species by the male tergum 9 + ectoprocts with posterodorsal portion nearly right-angled and with posteroventral portion elongated and tapering. In I. dictyophilum the male tergum 9 + ectoprocts is posteriorly convex.	en	Li, Di, Aspöck, Horst, Aspöck, Ulrike, Liu, Xingyue (2018): A review of the beaded lacewings (Neuroptera: Berothidae) from China. Zootaxa 4500 (2): 235-257, DOI: 10.11646/zootaxa.4500.2.5
