taxonID	type	description	language	source
B7401262FF9A7914FF4DFE73FC60FD46.taxon	description	Figs 1 – 5 Type material. Holotype: TAIWAN • ♂; Hsinchu Co., Jianshi, Beidelaman; 24 ° 44 ' 15.7 " N, 121 ° 16 ' 58.8 " E; 1 May 2019; Y. C. Liao leg.; Helicia cochinchinensis; NCHU, dry mounted. Paratypes: TAIWAN • 42 ♂, 25 ♀, 12 immatures; same data as for holotype • 3 ♂, 3 ♀; same data as for holotype, but NMNS • 3 ♂, 3 ♀; same data as for holotype, but NHMB • 2 ♂, 1 ♀; Hsinchu Co., Jianshi, Beidelaman; 24 ° 44 ' 15.7 " N, 121 ° 16 ' 58.8 " E; 30 Apr. 2011; C. T. Tang leg.; Helicia cochinchinensis • 3 ♂, 2 ♀; Taichung City, Daxueshan forest road: 16 K; 24 ° 14 ' 37.3 " N, 120 ° 54 ' 47.7 " E; 8 Mar. 2012; Y. C. Liao leg.; Helicia rengetiensis • 1 ♂, 1 ♀; Taichung City, Daxueshan forest road: 16 K; 24 ° 14 ' 37.3 " N, 120 ° 54 ' 47.7 " E; 21 Feb. 2013; Y. C. Liao leg.; Helicia rengetiensis • 5 immatures; Taichung City, Daxueshan forest road: 20.5 K; 24 ° 14 ' 41.1 " N, 120 ° 55 ' 20.0 " E; 22 Jan. 2014; B. C. Lai leg.; Helicia rengetiensis • 15 ♂, 18 ♀, 12 immatures; Taichung City, Daxueshan forest road: 20.5 K; 24 ° 14 ' 41.1 " N, 120 ° 55 ' 20.0 " E; 28 Jul. 2017; Y. C. Liao leg.; Helicia rengetiensis • 1 ♀, 26 immatures; Hsinchu Co., Jianshi, Beidelaman; 24 ° 44 ' 15.7 " N, 121 ° 16 ' 58.8 " E; 23 Apr. 2018; Y. C. Liao leg.; Helicia cochinchinensis • 2 ♂, 4 ♀, 25 immatures; Taoyuan City, Fuxing, Dongyanshan Forest Recreation Area; 24 ° 49 ' 10.9 " N 121 ° 24 ' 40.2 " E; 29 Apr. 2019; Y. C. Liao leg.; Helicia cochinchinensis • 7 immatures; Hsinchu Co., Jianshi, Lidongshan; 24 ° 41 ' 23.0 " N 121 ° 18 ' 31.0 " E; 29 Apr. 2019; Y. C. Liao leg.; Helicia cochinchinensis • 1 immature; Xinbei City, Sanxia, The Great Roots Forestry Resort; 24 ° 52 ' 24.3 " N 121 ° 24 ' 22.5 " E; 10 Dec. 2020; Y. C. Liao leg.; Helicia formosana Description. Adults (Figs 1, 5 A). Coloration. Body color yellowish brown (Fig. 1). Newly emerged individuals light green. A white stripe on dorsum of abdomen. Antennae yellow with apices of segments 4, 6 and 8 dark brown, and entire segments 9 and 10 black. Compound eyes reddish brown. Ocelli orange. Legs brown. Forewing and hindwing transparent. Structure. Body median-sized, length from anterior head margin to tip of folded forewing 3.7 – 4.9 mm, female larger; covered in long fine setae. Head (Fig. 1 B, D) slightly narrower than thorax, inclined in a 45 ° angle from longitudinal body axis. Vertex (Fig. 2 A) 1.7 – 1.9 times as wide as long, moderately concave at posterior margin. Genal processes prominent (Fig. 2 A), 0.6 – 0.8 times as long as vertex along mid-line, divergent, conical, blunt at apex, pubescent. Antenna (Fig. 2 C, E) slender, 10 - segmented, 1.7 – 1.9 times as long as head width, relative length of flagellar segments as 1.0: 0.4: 0.3: 0.3: 0.3: 0.3: 0.2: 0.2, with a single rhinarium on each of segments 4, 6, 8 and 9; longer, pointed terminal seta 1.1 times and shorter, truncate terminal seta 0.2 times as long as segment 10. Thorax weakly arched dorsally. Pronotum deflexed from mesothorax in a 45 ° angle. Legs slender. Meracanthus well developed, horn-shaped, acute at apex (Fig. 2 F); metatibia 1.2 – 1.3 times as long as head width, slightly inflated basally with small spines, with 1 + 2 apical spurs. Forewing (Fig. 2 B) 5.3 – 6.0 times as long as head width, 2.7 – 2.9 times as long as wide, widest slightly distal to the middle; wing apex subacute, lying in cell m 1 near apex of vein M 1 + 2; vein R + M + Cu strictly trifurcating into veins R, M and Cu; vein Rs moderately long, irregularly, concavely curved to fore margin of wing; vein M weakly curved with moderately long diverging branches; cell m 1 triangular; vein Cu 1 a slightly curved; cell cu 1 same size as cell m 1; line connecting apices of veins Rs and Cu 1 a proximal of bifurcation of vein M; surface spinules absent; radular spinules present along wing margin in the middle of cells m 1, m 2 and cu 1. Hindwing (Fig. 2 D) 0.6 times as long and 0.5 times as wide as forewing. Male terminalia (Fig. 3 A – C). Proctiger tubular, in profile broadly convex posteriorly, covered in long setae except for basal third laterally (Fig. 3 A). Subgenital plate subglobular, with long setae laterally and ventrally. Paramere (Fig. 3 B) about as long as proctiger; in profile lamellar, irregularly narrowing to apex which is acute and weakly directed inwardly; inner face beset with long setae. Distal segment of aedeagus (Fig. 3 C) about as long as paramere, apical third inflated, spoon-shaped; sclerotized end tube of ductus ejaculatorius short, sinuous. Female terminalia (Fig. 3 D) cuneate, short. Proctiger with sinuate dorsal margin and pointed apex, longer than subgenital plate; with long setae in the middle and apically; circumanal ring one fourth as long as proctiger, consisting of one row of pores (Fig. 3 D). Subgenital plate, in profile, irregularly triangular, acute at apex; beset in long hairs laterally and ventrally. Dorsal valvulae cuneate, ventral valvulae straight lacking teeth. Measurements (range, mean ± SD) in mm (5 males, 5 females). Body length (including forewing) ♂ 3.72 – 4.14, 3.98 ± 0.17; ♀ 4.56 – 4.90, 4.68 ± 0.13. Head width ♂ 0.63 – 0.66, 0.65 ± 0.01; ♀ 0.68 – 0.71, 0.69 ± 0.01. Vertex width ♂ 0.38 – 0.39, 0.38 ± 0.01; ♀ 0.40 – 0.41, 0.41 ± 0.01 Vertex length ♂ 0.20 – 0.23, 0.22 ± 0.01; ♀ 0.24 – 0.25, 0.24 ± 0.01. Genal cone length ♂ 0.16 – 0.19, 0.17 ± 0.01; ♀ 0.17 – 0.19, 0.18 ± 0.01. Antenna length ♂ 1.08 – 1.24, 1.18 ± 0.06; ♀ 1.20 – 1.26, 1.23 ± 0.03. Metatibia length ♂ 0.74 – 0.80, 0.77 ± 0.02; ♀ 0.85 – 0.90, 0.88 ± 0.02. Forewing length ♂ 3.28 – 3.58, 3.47 ± 0.12; ♀ 3.92 – 4.21, 4.05 ± 0.11. Forewing width ♂ 1.18 – 1.27, 1.24 ± 0.04; ♀ 1.47 – 1.59, 1.51 ± 0.05. Fifth instar immatures (Figs 4, 5 B). Coloration. General color light green. Body (Fig. 4 A) form oval, 1.5 – 1.6 times as long as wide; sclerotized dorsally, membranous ventrally. Dorsal body surface covered in short normal setae or acute sectasetae; margin of head, forewing (Fig. 4 C) and hindwing pads, as well as caudal plate covered with long, very slender, acute sectasetae which are relatively densely spaced (distance between setae about 0.8 – 1.0 times their length). Antenna (Fig. 4 F) weakly curved; 8 - segmented; scape and pedicel much thicker than flagellum; relative length of flagellar segments as 1.0: 0.6: 0.3: 0.5: 0.4: 1.8; with a single subapical rhinarium on each of segments 4 and 6, and two on segment 8. Legs moderately long, femur slightly shorter than tibiotarsus; tarsus with two well-developed claws, tarsal arolium (Fig. 4 B) longer than claws, triangular with a cleft in the anterior margin, lacking pedicel. Forewing pad 3.5 times long as broad, 2.2 – 2.5 times as long as antenna; humeral lobe relatively short, reaching hind margin of eye. Caudal plate broadly rounded caudally, 0.9 – 1.0 times as long as wide. Circumanal ring (Fig. 4 D, E) relatively wide, two lobes, each robust with round apex, 0.4 times as wide as caudal plate; in ventral position close to hind of caudal plate; outer ring composed of 1 row of pores. Measurements (range, mean ± SD) in mm (5 immatures). Body length 2.05 – 3.00, 2.51 ± 0.35. Head width 0.78 – 0.80, 0.79 ± 0.01. Antenna length 0.48 – 0.53, 0.50 ± 0.02. Metatibiotarsus length 0.63 – 0.70, 0.66 ± 0.03. Forewing pad length 1.11 – 1.23, 1.18 ± 0.05. Caudal plate length 0.95 – 1.45, 1.11 ± 0.20. Caudal plate width 1.10 – 1.48, 1.24 ± 0.14. Circumanal ring width 0.44 (n = 1).	en	Liao, Yi-Chang, Yang, Man-Miao (2025): Trioza burckhardti sp. nov. (Hemiptera: Triozidae) - the first record of jumping plant-lice associated with Proteaceae in North Hemisphere. Zootaxa 5727 (1): 140-147, DOI: 10.11646/zootaxa.5727.1.9, URL: https://doi.org/10.11646/zootaxa.5727.1.9
B7401262FF9A7914FF4DFE73FC60FD46.taxon	etymology	Etymology. The species name derives from Dr. Daniel Burckhardt, in honor of his significant contributions to the systematics of Psylloidea.	en	Liao, Yi-Chang, Yang, Man-Miao (2025): Trioza burckhardti sp. nov. (Hemiptera: Triozidae) - the first record of jumping plant-lice associated with Proteaceae in North Hemisphere. Zootaxa 5727 (1): 140-147, DOI: 10.11646/zootaxa.5727.1.9, URL: https://doi.org/10.11646/zootaxa.5727.1.9
B7401262FF9A7914FF4DFE73FC60FD46.taxon	distribution	Distribution. Taiwan.	en	Liao, Yi-Chang, Yang, Man-Miao (2025): Trioza burckhardti sp. nov. (Hemiptera: Triozidae) - the first record of jumping plant-lice associated with Proteaceae in North Hemisphere. Zootaxa 5727 (1): 140-147, DOI: 10.11646/zootaxa.5727.1.9, URL: https://doi.org/10.11646/zootaxa.5727.1.9
B7401262FF9A7914FF4DFE73FC60FD46.taxon	biology_ecology	Host plant. Helicia rengetiensis Masam., Helicia formosana Hemsl., Helicia cochinchinensis Lour. Biology. This species is univoltine, with adults emerging from February to April. It usually induces leaf curling galls (Fig. 5 C, D) but can occasionally induce flower galls (Fig. 5 E, F), even on the same individual plant. The galls are semi-open, each containing multiple immatures inside. This species inhabits lowland mountainous areas, occurring in the forest understory at elevations of 600 to 1,500 m elevation.	en	Liao, Yi-Chang, Yang, Man-Miao (2025): Trioza burckhardti sp. nov. (Hemiptera: Triozidae) - the first record of jumping plant-lice associated with Proteaceae in North Hemisphere. Zootaxa 5727 (1): 140-147, DOI: 10.11646/zootaxa.5727.1.9, URL: https://doi.org/10.11646/zootaxa.5727.1.9
