taxonID	type	description	language	source
B44A87E57C16FFA0FF069D5CFC99FF56.taxon	description	(Fig. 2 a, 3 a, 4, 7 a, 8 a)	en	Pelingen, Arthien Lovell (2025): Dr. Wolfram Mey’s Stonefly (Insecta: Plecoptera) Collection from Luzon, Philippines: Part II. Leuctridae. Zootaxa 5604 (1): 52-60, DOI: 10.11646/zootaxa.5604.1.5, URL: https://doi.org/10.11646/zootaxa.5604.1.5
B44A87E57C16FFA0FF069D5CFC99FF56.taxon	materials_examined	Holotype: ♂, Philippines, Luzon \ Benguet, 25. XI. 1997 \ Adunot-river Untenauf \ leg. Mey, Ebert, Nuss (PNM). Paratypes: 2 ♂, Luzon, Quezon \ Infanta, 9. IV. 1997 \ leg. Mey; 1 ♂, Mt. Banahaw \ Kinabuhayan \ 17 - 19. III. 2000 \ leg. Mey & Richter (AdMU)	en	Pelingen, Arthien Lovell (2025): Dr. Wolfram Mey’s Stonefly (Insecta: Plecoptera) Collection from Luzon, Philippines: Part II. Leuctridae. Zootaxa 5604 (1): 52-60, DOI: 10.11646/zootaxa.5604.1.5, URL: https://doi.org/10.11646/zootaxa.5604.1.5
B44A87E57C16FFA0FF069D5CFC99FF56.taxon	diagnosis	Diagnosis. The lateral projections of the tergum 10 of males bifurcate, with the top spine appearing longer than the lower one in lateral view. The cercus bears long setations, and in dorsal aspect, the epiproct looks like an elongated oval form.	en	Pelingen, Arthien Lovell (2025): Dr. Wolfram Mey’s Stonefly (Insecta: Plecoptera) Collection from Luzon, Philippines: Part II. Leuctridae. Zootaxa 5604 (1): 52-60, DOI: 10.11646/zootaxa.5604.1.5, URL: https://doi.org/10.11646/zootaxa.5604.1.5
B44A87E57C16FFA0FF069D5CFC99FF56.taxon	description	Description. Male adult habitus. Forewing length: 4.5 – 4.8 mm (n = 3). Head light brown to brown, wider than pronotum; compound eyes black; antennae light brown; mouthparts brown. Pronotum light brown with dark rugosities; wings subhyaline with darker veins; legs brown. Abdomen light brown. Male terminalia. Tergum 9 strongly sclerotized at anterior margin, weakly sclerotized posteriorly, ornamented throughout. Sternum 9 longer than wide, distal portion with narrow trapezoidal subgenital plate, the plate wider than longer; vesicle in ventral view dark brown, ovoid, covered with long, dense hairs. Tergum 10 from lateral view the upper spine appears longer than the lower spine, with well-sclerotized, bifurcate lateral projections that terminate in two sharp, parallel-sided points in the dorsal aspect. Central plate sclerotized and trilobed, lateral lobes an elongated subtrapezoid, medial lobe darkly pigmented. Cercus twice longer than wide, curved dorsally, with setation, without terminal spine. Epiproct strongly sclerotized and darkly sclerotized laterally, oval in dorsal view, longer than wide. Subanal lobe with prominent basal sclerotization, ambiguous ventral furrows, membranous lateral edges, and apex.	en	Pelingen, Arthien Lovell (2025): Dr. Wolfram Mey’s Stonefly (Insecta: Plecoptera) Collection from Luzon, Philippines: Part II. Leuctridae. Zootaxa 5604 (1): 52-60, DOI: 10.11646/zootaxa.5604.1.5, URL: https://doi.org/10.11646/zootaxa.5604.1.5
B44A87E57C16FFA0FF069D5CFC99FF56.taxon	etymology	Etymology. The species is named after its current known distribution in the Luzon Island, Philippines.	en	Pelingen, Arthien Lovell (2025): Dr. Wolfram Mey’s Stonefly (Insecta: Plecoptera) Collection from Luzon, Philippines: Part II. Leuctridae. Zootaxa 5604 (1): 52-60, DOI: 10.11646/zootaxa.5604.1.5, URL: https://doi.org/10.11646/zootaxa.5604.1.5
B44A87E57C16FFA0FF069D5CFC99FF56.taxon	distribution	Distribution. This species is known so far only from Benguet and Quezon provinces in the Philippines.	en	Pelingen, Arthien Lovell (2025): Dr. Wolfram Mey’s Stonefly (Insecta: Plecoptera) Collection from Luzon, Philippines: Part II. Leuctridae. Zootaxa 5604 (1): 52-60, DOI: 10.11646/zootaxa.5604.1.5, URL: https://doi.org/10.11646/zootaxa.5604.1.5
B44A87E57C16FFA0FF069D5CFC99FF56.taxon	discussion	Remarks. The novel species is a typical of R. malayana group, sensu Sivec et al. (2008). The male appears to be closely connected to R. malayana and more similar with R. meyi having alike bifurcated tergum 10 lateral projections and strongly hooked epiprocts. However, R. luzonensis sp. nov. can be differentiated from R. malayana with the upper spine of bifurcate lateral projections on tergum 10 being longer than the lower one in lateral view; while R. luzonensis sp. nov. differentiated from R. meyi based on a bigger and more ovoid epiproct on its dorsal view. In addition, the posterior margin of sternum 9 is curved smoothly in R. luzonensis sp. nov. while R. meyi projects curving posteriorly. Its subanal lobes are also more elongated than that of R. meyi. Lastly, R. luzonensis sp. nov. lacks a terminal spine on its cerci in contrast to R. malayana.	en	Pelingen, Arthien Lovell (2025): Dr. Wolfram Mey’s Stonefly (Insecta: Plecoptera) Collection from Luzon, Philippines: Part II. Leuctridae. Zootaxa 5604 (1): 52-60, DOI: 10.11646/zootaxa.5604.1.5, URL: https://doi.org/10.11646/zootaxa.5604.1.5
B44A87E57C14FFA7FF069FABFD54FE52.taxon	description	(Fig. 2 b, 3 b, 5, 7 b, 8 b)	en	Pelingen, Arthien Lovell (2025): Dr. Wolfram Mey’s Stonefly (Insecta: Plecoptera) Collection from Luzon, Philippines: Part II. Leuctridae. Zootaxa 5604 (1): 52-60, DOI: 10.11646/zootaxa.5604.1.5, URL: https://doi.org/10.11646/zootaxa.5604.1.5
B44A87E57C14FFA7FF069FABFD54FE52.taxon	materials_examined	Holotype: ♂, Philippines, Luzon \ Zambales Mts, Coto \ 9 - 10. XI. 1998, 150 m \ leg. Mey & Speidel (PNM). Paratype: 1 ♂, same data as holotype (AdMU)	en	Pelingen, Arthien Lovell (2025): Dr. Wolfram Mey’s Stonefly (Insecta: Plecoptera) Collection from Luzon, Philippines: Part II. Leuctridae. Zootaxa 5604 (1): 52-60, DOI: 10.11646/zootaxa.5604.1.5, URL: https://doi.org/10.11646/zootaxa.5604.1.5
B44A87E57C14FFA7FF069FABFD54FE52.taxon	diagnosis	Diagnosis. Male adult of this species is characterized by the bifurcated lateral projections of the tergum and the epiproct, which is curved rather shallowly. So far, this is the only Philippine species without a vesicle.	en	Pelingen, Arthien Lovell (2025): Dr. Wolfram Mey’s Stonefly (Insecta: Plecoptera) Collection from Luzon, Philippines: Part II. Leuctridae. Zootaxa 5604 (1): 52-60, DOI: 10.11646/zootaxa.5604.1.5, URL: https://doi.org/10.11646/zootaxa.5604.1.5
B44A87E57C14FFA7FF069FABFD54FE52.taxon	description	Description. Male adult habitus. Forewing length is 4.5 – 5.0 mm. Head brown to dark brown, wider than pronotum; compound eyes black; antennae brown to light brown; mouthparts brown. Pronotum light brown with slightly dark rugosities; wings subhyaline with darker veins; legs brown. Abdomen brown. Male terminalia. Tergum 9 well sclerotized at anterior margin, less so posteriorly which terminates in a wide, produced sclerite with tiny granules. Sternum 9 longer than wide, distal portion with narrow trapezoidal subgenital plate, the plate wider than longer; vesicle. Tergum 10 with well sclerotized, bifurcating equilateral projections. Central plate sclerotized and trilobed, lateral lobes subtriangular, together with a circular; medial lobe darkly pigmented, with a small knob-like projection. Cercus twice as long as wide, curved dorsally, with setations, without terminal spine. Epiproct sclerotized, and rather shallowly curved in lateral view. Subanal lobe with sclerotized lateral margins and apex membranous, ventral furrows vague.	en	Pelingen, Arthien Lovell (2025): Dr. Wolfram Mey’s Stonefly (Insecta: Plecoptera) Collection from Luzon, Philippines: Part II. Leuctridae. Zootaxa 5604 (1): 52-60, DOI: 10.11646/zootaxa.5604.1.5, URL: https://doi.org/10.11646/zootaxa.5604.1.5
B44A87E57C14FFA7FF069FABFD54FE52.taxon	etymology	Etymology. The specific epithet refers to the absence of a vesicle on the 9 th sternum and a terminal spine on the cerci which distinguishes it from all other Philippine Rhopalopsole.	en	Pelingen, Arthien Lovell (2025): Dr. Wolfram Mey’s Stonefly (Insecta: Plecoptera) Collection from Luzon, Philippines: Part II. Leuctridae. Zootaxa 5604 (1): 52-60, DOI: 10.11646/zootaxa.5604.1.5, URL: https://doi.org/10.11646/zootaxa.5604.1.5
B44A87E57C14FFA7FF069FABFD54FE52.taxon	distribution	Distribution. This species is known so far only from Zambales Province, Central Luzon, Philippines.	en	Pelingen, Arthien Lovell (2025): Dr. Wolfram Mey’s Stonefly (Insecta: Plecoptera) Collection from Luzon, Philippines: Part II. Leuctridae. Zootaxa 5604 (1): 52-60, DOI: 10.11646/zootaxa.5604.1.5, URL: https://doi.org/10.11646/zootaxa.5604.1.5
B44A87E57C14FFA7FF069FABFD54FE52.taxon	discussion	Remarks. The novel species is a typical of R. malayana group, sensu Sivec et al. (2008). The male appears to be closely related to R. malayana and R. meyi having the same bifurcating tergum 10 lateral projections. However, R. absentia sp. nov. can be differentiated from both R. malayana and R. meyi with its epiproct that is only slightly curved as compared to regularly rounded hooks.	en	Pelingen, Arthien Lovell (2025): Dr. Wolfram Mey’s Stonefly (Insecta: Plecoptera) Collection from Luzon, Philippines: Part II. Leuctridae. Zootaxa 5604 (1): 52-60, DOI: 10.11646/zootaxa.5604.1.5, URL: https://doi.org/10.11646/zootaxa.5604.1.5
B44A87E57C12FFA5FF069CC7FE2EFCDF.taxon	description	(Fig. 2 c, 3 c, 6, 7 c, 8 c)	en	Pelingen, Arthien Lovell (2025): Dr. Wolfram Mey’s Stonefly (Insecta: Plecoptera) Collection from Luzon, Philippines: Part II. Leuctridae. Zootaxa 5604 (1): 52-60, DOI: 10.11646/zootaxa.5604.1.5, URL: https://doi.org/10.11646/zootaxa.5604.1.5
B44A87E57C12FFA5FF069CC7FE2EFCDF.taxon	materials_examined	Holotype: ♂, Philippines, Luzon, Quezon \ Infanta, 9. IV. 1997 \ leg. Mey (PNM). Paratypes: 1 ♂, same data as holotype; 3 ♂, Mt. Banahaw \ Kinabuhayan \ 17 - 19. III. 2000 \ leg. Mey & Richter (AdMU)	en	Pelingen, Arthien Lovell (2025): Dr. Wolfram Mey’s Stonefly (Insecta: Plecoptera) Collection from Luzon, Philippines: Part II. Leuctridae. Zootaxa 5604 (1): 52-60, DOI: 10.11646/zootaxa.5604.1.5, URL: https://doi.org/10.11646/zootaxa.5604.1.5
B44A87E57C12FFA5FF069CC7FE2EFCDF.taxon	diagnosis	Diagnosis. Male adults of this species are characterized by the lateral projections of the tergum 10 composed of a giant dorsal spine and a tiny ventral tubercle. The epiproct is small and elliptical in the dorsal view.	en	Pelingen, Arthien Lovell (2025): Dr. Wolfram Mey’s Stonefly (Insecta: Plecoptera) Collection from Luzon, Philippines: Part II. Leuctridae. Zootaxa 5604 (1): 52-60, DOI: 10.11646/zootaxa.5604.1.5, URL: https://doi.org/10.11646/zootaxa.5604.1.5
B44A87E57C12FFA5FF069CC7FE2EFCDF.taxon	description	Description. Male adult habitus. Forewing length is 4.0 – 4.2 mm (n = 5). Head dark brown, wider than pronotum; compound eyes black; antennae light brown; mouthparts brownish to brown. Pronotum light brown without dark rugosities; wings subhyaline with darker veins; legs light brown. Abdomen brown to light brown. Male terminalia. Tergum 9 well sclerotized anteriorly, less so posteriorly, this margin terminating in a wide, produced sclerite with tiny granules. Sternum 9 longer than wide, distal portion with narrow trapezoidal subgenital plate, the plate wider than long; vesicle in ventral view dark brown, ovoid, covered with long, dense hairs. Tergum 10 with well sclerotized lateral projections terminating into a giant dorsal spine and a tiny ventral tubercle. Central plate sclerotized and trilobed, lateral lobes subtriangular, together with a circular; medial lobe darkly pigmented, with a small knob-like projection. Cercus twice as long as wide, curved dorsally, with setation, without terminal spine. Epiproct darkly sclerotized, and strongly curved cephalic in lateral view. Subanal lobe with sclerotized lateral margins and apex membranous, ventral furrows vague.	en	Pelingen, Arthien Lovell (2025): Dr. Wolfram Mey’s Stonefly (Insecta: Plecoptera) Collection from Luzon, Philippines: Part II. Leuctridae. Zootaxa 5604 (1): 52-60, DOI: 10.11646/zootaxa.5604.1.5, URL: https://doi.org/10.11646/zootaxa.5604.1.5
B44A87E57C12FFA5FF069CC7FE2EFCDF.taxon	etymology	Etymology. The specific epithet refers to the cephalically curved epiproct in lateral view as the distinguishing characteristic from R. andreae and R. cayasan.	en	Pelingen, Arthien Lovell (2025): Dr. Wolfram Mey’s Stonefly (Insecta: Plecoptera) Collection from Luzon, Philippines: Part II. Leuctridae. Zootaxa 5604 (1): 52-60, DOI: 10.11646/zootaxa.5604.1.5, URL: https://doi.org/10.11646/zootaxa.5604.1.5
B44A87E57C12FFA5FF069CC7FE2EFCDF.taxon	distribution	Distribution. This species is known so far only from Quezon Province, Philippines.	en	Pelingen, Arthien Lovell (2025): Dr. Wolfram Mey’s Stonefly (Insecta: Plecoptera) Collection from Luzon, Philippines: Part II. Leuctridae. Zootaxa 5604 (1): 52-60, DOI: 10.11646/zootaxa.5604.1.5, URL: https://doi.org/10.11646/zootaxa.5604.1.5
B44A87E57C12FFA5FF069CC7FE2EFCDF.taxon	discussion	Remarks. The novel species is a typical of R. malayana group, sensu Sivec et al. (2008). It can be possibly confused with R. andreae and R. cayasan having tergum 10 lateral projections with a giant dorsal spine. However, R. rotundus sp. nov. can be differentiated from R. andreae with its epiproct being more cephalically curved as compared to the R. andreae epiproct which is more straightly projected caudally in lateral view (Sivec et al. 2008: 31). Interestingly, R. cayasan can also be differentiated from R. rotundus due to its shorter upright epiproct, in lateral view (Sivec et al. 2008: 33)	en	Pelingen, Arthien Lovell (2025): Dr. Wolfram Mey’s Stonefly (Insecta: Plecoptera) Collection from Luzon, Philippines: Part II. Leuctridae. Zootaxa 5604 (1): 52-60, DOI: 10.11646/zootaxa.5604.1.5, URL: https://doi.org/10.11646/zootaxa.5604.1.5
