identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
B400FD2EFFB20900FF102D9EFD3B8ACF.text	B400FD2EFFB20900FF102D9EFD3B8ACF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudocleobis Pocock 1900	<div><p>Genus Pseudocleobis Pocock, 1900</p><p>Pseudocleobis Pocock, 1900: 304; Kraepelin, 1901: 108; Roewer, 1934: 602; Mello-Leitão, 1938a: 23; Muma, 1971a: 14–16; Muma, 1976: 28.</p><p>Tetracleobis Roewer, 1934: 605; Muma, 1976: 28 (synonymized by Maury, 1976: 93).</p><p>Type species: Cleobis andinus Pocock, 1899, by original designation.</p><p>Distribution: South American transition zone (except Paramo biogeographic province), the Central Chilean and Patagonian sub-regions (Andean region) and the Western Chaco district (Neotropical region) (sensu Morrone, 2014, 2015).</p><p>Revised diagnosis: Pseudocleobis can be distinguished from all other ammotrechid genera, except Dasycleobis and Chinchippus (see note below), by the following combination of characters: Chelicerae fixed finger with FD, FSD, FM, FSM and FP teeth on median series (Fig. 6A); with RFM, RFSM, RFP, RFSP teeth on retrofondal series (Fig. 7B) and with PFM, PFSM, PFP, PFSP teeth on profondal series (Fig. 6A). Movable finger with MM, MSM, MP teeth, without MPL tooth (Figs. 6A; 7B). Females and juveniles without dorsal hump on fixed finger (Fig. 6A). Fixed finger of males highly modified and usually with a conspicuous flagellar groove. Pedipalps coated with bifurcated tip setae and with long lateroventral spiniform setae (as long or longer than the width of pedipalp article) on femur, tibia and basitarsus (Fig. 6C; usually four pairs on basitarsus and five pairs on tibia); telotarsus slightly pearshaped, fixed, and without spiniform setae (Fig. 6C). Tibia II and III without dorsoapical spiniform seta. Basitarsus II and III with two retrolateral (i.e., RL-b, RL-sd) and one retrodorsal distal (i.e., RD-d) spiniform setae; the distal retrolateral spiniform setae is oriented ventrally (Fig. 6D). Telotarsus II and III divided in two tarsomeres with the following spiniform setal formula 1.2.2/2.2 (Fig. 6E). Telotarsus IV divided in four tarsomeres with the following spiniform setal formula 2.2-2-2/2.2 (Fig. 6F). Males of some species have long filiform ctenidia on sternites III and IV (see discussion).</p><p>Note: A recent contribution (Iuri et al 2021) suggest a close relationship between Pseudocleobis, Dasycleobis Mello-Leito, 1940 and Chinchippus Chamberlin, 1920 . The three genera share most of the diagnostic morphologic characters mentioned here. The genus Dasycleobis, known from the high Andean central west Argentina, have more plumose setae on the cheliceral fixed finger (two or more complete rows) and a lateral folding on the flagellum absent on the known Pseudocleobis species. Chinchippus is currently known from the coastal Peruvian desert and could be distinguished from Pseudocleobis by its tracheal system (see Iuri et al 2021). Further analysis is necessary to establish clear limits between these genera.</p></div>	https://treatment.plazi.org/id/B400FD2EFFB20900FF102D9EFD3B8ACF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Iuri, Hernán A.;Iglesias, Mónica S.	Iuri, Hernán A., Iglesias, Mónica S. (2022): The genus Pseudocleobis Pocock, 1900 (Solifugae: Ammotrechidae) in transitional Patagonia-Monte deserts, with descriptions of two new species. Zootaxa 5094 (3): 435-460, DOI: 10.11646/zootaxa.5094.3.4
B400FD2EFFB1090FFF102AC2FB5C889F.text	B400FD2EFFB1090FFF102AC2FB5C889F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudocleobis profanus Iuri & Iglesias 2022	<div><p>Pseudocleobis profanus Iuri, sp. nov.</p><p>(Figs. 3A, B; 4; 7C, D; 9C; 10C; 11B; 12D–F; 14. Table 1)</p><p>Pseudocleobis huinca: Maury 1976 misidentification (in part)</p><p>Type material. Holotype: ARGENTINA: Río Negro: ♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-68.05639&amp;materialsCitation.latitude=-38.815594" title="Search Plazi for locations around (long -68.05639/lat -38.815594)">Cinco Saltos</a>, behind the cemetery [38º48’56,14’’S 68º03’23,00’’W], 11–12.I.2014. H.A. Iuri coll., (MACN-Ar 38425) ; Paratypes: ARGENTINA: Río Negro: 4♂, Same data as holotype (MACN-Ar 38430, 38474, 38480, 38491); 1♂, Same locality and collector as holotype, 7.I.2012 (MACN-Ar 38493).</p><p>Other examined material. ARGENTINA: Neuquén: 1♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-70.36188&amp;materialsCitation.latitude=-38.52351" title="Search Plazi for locations around (long -70.36188/lat -38.52351)">Las Lajas</a> [38º31’24,63’’S 70º21’42,76’’W], 16.I.1967, P. San Martín coll., (MACN-Ar 6873; recorded by Maury 1976 as P. huinca) ; Río Negro: 1♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-67.58417&amp;materialsCitation.latitude=-39.0283" title="Search Plazi for locations around (long -67.58417/lat -39.0283)">General Roca</a> [39º01’41,89’’S 67º35’03,00’’W], I.1962, A. Bachmann coll., (MACN-Ar 6871; paratype of P. huinca, misidentified by Maury (1976)) ; 25♂, 10♀, same locality and collector as holotype, 6–9.I.2016, (MACN-Ar) .</p><p>Etymology: The name comes from the Latin adjective ‘ profanus ’, non-sacred, given that the type specimens were found near a cemetery. It’s an adjective in the nominative singular.</p><p>Diagnosis: Pseudocleobis profanus sp. nov. can be distinguished from all other Pseudocleobis except P. huinca and P. bardensis by the male movable finger mucron with a narrow dorsal crest and shovel-like apex (Figs. 7D; 11B; 12D), and the flagellum with a prolateral subcircular subapical row of filaments (Fig. 9C). It can be distinguished from P. bardensis by the more prominent dorsal crest and more prominent shovel-like apex of movable finger mucron (Figs. 7C–D) and the apex of fixed finger mucron ending in a smooth curve till the tip (Figs. 7C–D). It can be distinguished from P. huinca by the shape of fixed finger mucron, with a shorter and not bi-convex proventral flange (Figs. 7C, D).</p><p>Description: Male: measurements in Table 1. Color in vivo. As in Fig. 3. Color in 96 % ethanol. Prosoma: propeltidium brown with central portion, two areas on each side of the eyes and two areas in the posterior margin paler (as Fig. 6B). Lateral lobes brown. Ocular tubercle black. Insertion of setae yellowish. Para-, meso- and metapeltidium brown, insertion of setae yellowish. Meso- and metapeltidium with two pale oval areas. Chelicerae: brown with three longitudinal pale oval areas. Insertion of setae yellowish. Pedipalps: brown with ventral surface paler. Coxae pale yellowish. Legs: leg I, II and III; Brown, with ventral surface paler. Telotarsus, coxa and trochanter pale yellowish. Leg IV; with a similar color pattern, but the telotarsus is more pigmented. Malleoli: pale yellowish.</p><p>Opisthosoma: tergites brown with insertion of setae yellowish. Pleurites and sternites pale yellowish.</p><p>Morphology and Chaetotaxy: Prosoma: propeltidium slightly wider than long with bifurcated tip setae of different sizes (most of them pointing backward). Lateral lobes, partially separated by dorsal grooves. Median plagula, and anterior and posterior arci, with a transverse row of different-sized bifurcated tip setae. Meso- and metapeltidium wider than long, with bifurcated tip setae.</p><p>Chelicerae: Dentition and processes: fixed finger: median series with FD, FSD, FM, FSM and FP teeth (Fig. 7D), the FSD tooth is very tiny and sometimes absent; the FP is the biggest tooth on fixed finger; the FD is very tiny but present in all specimens examined; retrofondal series with RFM, RFSM, RFP, RFSP teeth and profondal series with PFM, PFSM, PFP, PFSP teeth. The fixed finger mucron is long, robust, and with a wide prolateral flagellar groove (with conspicuous prodorsal and proventral flanges; Figs. 11B; 12D), and it ends in a smooth curve till the tip (Fig. 7C, D). Movable finger: with MM, MSM and MP teeth (Fig. 7D); the MM similar or bigger than FM; the MP is the biggest tooth of the chelicera. The mucron is highly modified with a high, narrow dorsal crest and a shovel-like apex (Figs. 7C, D; 11B). Chaetotaxy: prolateral surface: The pvd series consists of two rows of plumose setae. The distal row is incomplete, usually restricted to the area of the profondal teeth (Fig. 12E). The second row extends from the pic to FM tooth. The pvsd series consists of one row of blunt setae that usually extends from the PFSP to the FSM. The pm series consists of small plumose hairs. The pdp series consists of four blunt setae. The pmpc and the pv consists of small hairs. Retrolateral surface: The rlm series consists of different-sized bifurcated tip setae, with the proximal group pointing backward. The rlf series consists of simple-tip, anteriorly directed, setae.</p><p>Flagellum: the flagellum is pear-shaped with a wide apex and concave ventral edge (Fig. 9C). Attachment ring of flagellum large, located at the level between the FSM and RFM teeth. The edge is thin with some irregular prolongations. On the retrolateral surface there is a subcircular subapical row of long fine filaments (Fig. 9A–B). The prolateral surface is smooth, not fringed (Fig. 12D).</p><p>Pedipalps (as Figs. 6C): all segments with several bifurcated tip setae and some tapered setae (Fig. 14B). Femur with a prolateral row of four long ventral spiniform setae. Tibia with 2.2.2.2.2 long ventral spiniform setae, the retrolateral row smaller and more similar in size, the prolateral row longer and arranged in increasing size I=V, II=IV, III; there is one pair of apical setae that seems like a weak pair of spiniform setae, but the insertion socket is different being completely circular and not strongly elevated. Basitarsus with 2.2.2.2 long ventral spiniform setae, the retrolateral smaller than the prolateral, but more equally sized in each row than those of the tibia; the proximal pair is weaker but can be distinguished; with some distal clubbed setae. Telotarsus with some clubbed setae (Fig. 14C), some slit sensilla (Fig. 14A), and retrodorsal pores.</p><p>Legs: all legs coated with bifurcated tip setae of different size; the trochanters and femora possess some robust bifurcated tip setae. The arolium of walking legs is very small. Leg I: without claws nor spiniform setae. Basitarsus and telotarsus with some clubbed setae (Fig. 14F) and bifurcated tip setae. With dorsal pores on the retrodorsal surface (Fig. 14D) and a short, blunt, apical seta (Fig. 14E). Leg II and III: tibia with 1.2 lateroventral spiniform setae; basitarsus with 1 retrodorsal distal spiniform seta (i.e. RD-d), 1.1 retrolateral spiniform setae (i.e. RL-b, RL-sd) (as Fig. 6D), and 1.2 lateroventral spiniform setae (i.e. RV-d, PV-d, PV-sd); telotarsus divided into two tarsomeres with the following spiniform formula: 1.2.2/2.2 (as Fig. 6E); the distal subdivision is relatively weak but complete, corresponding to the small terminal tarsomere that is placed on a sub-ventral position. Leg IV: tibia with 1.1.2 ventral spiniform setae; basitarsus with 1.1.2 ventral spiniform setae; telotarsus divided in four tarsomeres with the following spiniform setae formula: 2.2-2-2/2.2; the distal subdivision same as leg II and III, the basal subdivisions are weak folds but perceptible due the articulation (as Figs. 6F; 13B; 15G).</p><p>Maleolli (Fig.14 G-I): the surface is similar to the surface of the setae (Fig. 14H). With some short filaments, particularly on the lateral margins (Fig. 14I).</p><p>Opisthosoma: tergites and sternites with bifurcated tip setae. With some large, thick, tapered tip setae (ctenidia) on sternite III and IV. The number of ctenidia is 2-2 on spiracular sternite I and 3-3 on spiracular sternite II. The microsetae (Fig. 14J–L) are present and placed in the same pattern as described by Iuri et al. (2014), with two pairs on sternites II, III and IV, and a single pair on sternite V.</p><p>Female: morphology and chaetotaxy similar to males, except by the wider propeltidium, wider chelicerae, and more robust body in general. Living specimens with body coloration pattern similar to that of female Pseudocleobis huinca as in Fig. 2A. The posterior pair of spiniform setae of basitarsus and tibia is more spiniform than males. Opisthosomal sternite III and IV without ctenidia. Genital plate (Fig. 10C): similar to the plate of P. huinca and P. bardensis, being flat, with a typical posterior opening resembling an omega symbol (Ω) (as Fig. 10A–E) and with a median posterior sclerite (upward arrow in Fig. 10C). There are two anterior small plates on the opening (transversal arrows in Fig. 10C). The lateral margins are slightly convex (never concave).</p><p>Note: some studied females have the genital plate shrivelled (see Fig. 10D–E) and, in some of these plates, the median sclerite and the small plaques are missing. It is probable that those were females that had recently copulated. Genital plate damage after copulation was reported in some solifuge species (Peretti &amp; Wilemart 2007, HruškováMartišová et al. 2010).</p><p>Distribution: the only three known localities of P. profanus sp. nov. belong to the Austral district of Monte biogeographic province and the Austral Payunia district of Patagonian biogeographic province (Figs. 1A, C; 4). Pseudocleobis profanus is apparently absent southern to the Negro river, and the distribution of this species may be limited southerly by the Negro and Limay rivers. On the other hand, its northern limit is unknown as the area between the Neuquén and Colorado rivers and the Septentrional Payunia district are poorly sampled.</p></div>	https://treatment.plazi.org/id/B400FD2EFFB1090FFF102AC2FB5C889F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Iuri, Hernán A.;Iglesias, Mónica S.	Iuri, Hernán A., Iglesias, Mónica S. (2022): The genus Pseudocleobis Pocock, 1900 (Solifugae: Ammotrechidae) in transitional Patagonia-Monte deserts, with descriptions of two new species. Zootaxa 5094 (3): 435-460, DOI: 10.11646/zootaxa.5094.3.4
B400FD2EFFBE090CFF102972FD7E89EE.text	B400FD2EFFBE090CFF102972FD7E89EE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudocleobis huinca : Maury 1976	<div><p>Pseudocleobis huinca Maury, 1976</p><p>(Figs. 2A–C; 4; 6A–C, F; 7A, B; 9A, B; 10A, D, E; 11A; 12A–C; 13A–E. Table 1, table 2)</p><p>Pseudocleobis huinca: Maury 1976: 94, 96–99; Figs. 18–20. Galiano &amp; Maury 1979: 329. Maury 1984: 75; Figs. 2–3.</p><p>Not Pseudocleobis huinca: Maury 1976: 98–99 (misidentification in part, see notes below).</p><p>Type material (examined). Holotype: ARGENTINA: Río Negro: ♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-66.1625&amp;materialsCitation.latitude=-40.67972" title="Search Plazi for locations around (long -66.1625/lat -40.67972)">Valcheta</a> [40º40’47,00’’S 66º09’45,00’’W], 23.I.1975, E.A. Maury, A. Toth, P. Domínguez and C. Césari coll., (MACN-Ar 6865) . Paratypes: ARGENTINA: Río Negro: 1♀, same data as Holotype (MACN-Ar 6866, Allotype); 1♂, 2♀, same data as Holotype (MACN-Ar 6867); 1♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-67.26052&amp;materialsCitation.latitude=-40.507023" title="Search Plazi for locations around (long -67.26052/lat -40.507023)">Ramos Mexía</a> [40º30’25,28’’S 67º15’37,87’’W], Río Negro, 22.I.1975, E.A. Maury, A. Toth, P. Domínguez and C. Césari coll., (MACN-Ar 6868) ; 1♂, Arroyo de la Ventana [41º40’04,91’’S 66º05’55,85’’W], Río Negro, same date and collectors of Holotype, (MACN-Ar 6869); 1♂, 1♀, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-67.58417&amp;materialsCitation.latitude=-39.0283" title="Search Plazi for locations around (long -67.58417/lat -39.0283)">Paso Córdoba</a> [39º06’47,46’’S 67º37’34,08’’W], Río Negro, 28.I.1974, E.A. Maury coll., (MACN-Ar 6870); 2♂, General Roca [39º01’41,89’’S 67º35’03,00’’W], Río Negro, I.1962, A. Bachmann coll., (MACN-Ar 6871) .</p><p>Other examined material. ARGENINA: Chubut: 1♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-64.49952&amp;materialsCitation.latitude=-42.46455" title="Search Plazi for locations around (long -64.49952/lat -42.46455)">Península Valdéz</a> [42º27’52,38’’S 64º29’58,29’’W], XII.1967, P. Williner coll., (MACN-Ar 6874) ; Río Negro: 2♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-66.144684&amp;materialsCitation.latitude=-40.688633" title="Search Plazi for locations around (long -66.144684/lat -40.688633)">Valcheta</a>, near the railroad station [40º41’19,07’’S 66º08’40,88’’W], 29–31.I.2013, H.A. Iuri and D.L. Iuri coll., (MACN-Ar) ; 10♂, 2♀, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-66.17051&amp;materialsCitation.latitude=-40.666355" title="Search Plazi for locations around (long -66.17051/lat -40.666355)">Valcheta</a>, near the abandoned fluorite factory [40°39’58,87”S 66°10’13,84”W], 10–12.I.2015, H.A. Iuri coll., (MACN-Ar) ; 2♂, 5♀, same locality and collector, 17.I.2016 (MACN-Ar); 19♂, and some ♀ and juv, Paso Córdoba, near the <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-67.620125&amp;materialsCitation.latitude=-39.11282" title="Search Plazi for locations around (long -67.620125/lat -39.11282)">Nautical Club</a> “ Julio A. Trulls ” [39º06’46,16’’S 67º37’12,45’’W], 13–15.I.2014, H.A. Iuri coll., (MACN-Ar) ; 71♂, 21♀, 3 immature, same locality and collector, 16–21.I.2015, (MACN-Ar); 40♂, 34♀, 8 immature, same locality and collector, 11–12.I.2016, (MACN-Ar); 1♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-68.05639&amp;materialsCitation.latitude=-38.815594" title="Search Plazi for locations around (long -68.05639/lat -38.815594)">Cinco Saltos</a>, behind the cemetery [38º48’56,14’’S 68º03’23,00’’W], 11–12.I.2014, H.A. Iuri coll., (MACN-Ar) ; 1♂, 3♀, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-65.024345&amp;materialsCitation.latitude=-41.646267" title="Search Plazi for locations around (long -65.024345/lat -41.646267)">Playas Doradas</a>, “sendero del cuis” [41°38’46.55”S 65° 1’27.64”W], 8–10.I.2015, H.A. Iuri coll., (MACN-Ar) ; 5♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-65.38688&amp;materialsCitation.latitude=-41.604614" title="Search Plazi for locations around (long -65.38688/lat -41.604614)">Sierra Grande</a>, near the cemetery [41º36’16.62’’S 65º23’12.77’’W], 19–20.I.2016, H.A. Iuri coll., (MACN-Ar) ; 1♂, 5♀, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-68.08547&amp;materialsCitation.latitude=-40.824455" title="Search Plazi for locations around (long -68.08547/lat -40.824455)">Los Menucos</a>, near the cemetery [40º49’28.04’’S 68º5’7.70’’W], 13–16.I.2016, H.A. Iuri coll., (MACN-Ar) .</p><p>Notes: One male from Bajada “El Marucho”, Neuquén (MACN-Ar 6872) recorded by Maury (1976) as P. huinca actually corresponds to P. mauryi sp. nov., while one of the males paratypes from General Roca, Río Negro (MACN-Ar 6871), and the male from Las Lajas, Neuquén (MACN-Ar 6873), also recorded by Maury (1976) as P. huinca, correspond to P. profanus sp. nov. The specimens from Sierra Grande (MACN-Ar 6875) and from Cerro Áspero (MACN-Ar 6876), recorded by Maury (1976) as P. huinca, are juveniles Pseudocleobis but could not be assigned to this species.</p><p>Diagnosis: Pseudocleobis huinca can be distinguished from all other Pseudocleobis except P. bardensis and P. profanus sp. nov. by the male movable finger mucron with a narrow dorsal crest and shovel-like apex (Fig. 7B; 12A), and the flagellum with a prolateral subcircular subapical row of filaments (Fig. 7A, B). It can be distinguished from P. bardensis by the movable finger mucron with a more prominent median crest and more prominent shovellike apex (Figs. 7A–B; 11A; 12A), and from both, P. bardensis and P. profanus sp. nov., by the shape of the fixed finger mucron, with a more long and bi-convex proventral flange (Figs. 7A–B; 11A; 12A) ending in a small hooklike tip.</p><p>Note: Maury (1976) proposed the difference in the width of the lateral lobe to distinguish between the female genital plate of P. huinca and P. bardensis . However, studying many female specimens of P. huinca we have observed that the width of the posterior lobes is not useful to separate these species.Also, the females captured with the males of P. profanus sp. nov. are morphologically similar to the females of P. huinca . Thus, the genital plate allows distinguishing P. huinca females from females of other species such as P. andinus, P. alticola, P. solitarius, but not from P. bardensis and P. profanus sp. nov.</p><p>Description: see Maury (1976); male and female similar to P. profanus sp. nov. and P. bardensis except for the diagnostic characters. Measurements in Table 1.</p><p>Variability: males have a highly reduced FSD tooth, usually vestigial or absent. Variations on females usually consist of additional, very small, denticles on the secondary teeth series. For the variability of the dentition on both sexes see Table 2.</p><p>Distribution: the records of P. huinca correspond to the Austral district of the Monte biogeographic province (Figs. 1A,B,D; 4), and seem to be restricted to this biogeographic area. This species is found southern to the Negro river, with only two records northern to this river, and no records northern to the Colorado river (the Colorado river approximately matches the northern limit of the Monte Austral district). Pseudocleobis huinca seems to be absent at the west, between the Limay and Neuquén rivers.</p></div>	https://treatment.plazi.org/id/B400FD2EFFBE090CFF102972FD7E89EE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Iuri, Hernán A.;Iglesias, Mónica S.	Iuri, Hernán A., Iglesias, Mónica S. (2022): The genus Pseudocleobis Pocock, 1900 (Solifugae: Ammotrechidae) in transitional Patagonia-Monte deserts, with descriptions of two new species. Zootaxa 5094 (3): 435-460, DOI: 10.11646/zootaxa.5094.3.4
B400FD2EFFBC090BFF102EF8FBF589EF.text	B400FD2EFFBC090BFF102EF8FBF589EF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudocleobis bardensis : Maury 1976	<div><p>Pseudocleobis bardensis Maury, 1976</p><p>(Figs. 4; 7E, F; 9D; 10B. Table 1)</p><p>Pseudocleobis bardensis: Maury 1976: 94, 99–101; Figs. 21–23.</p><p>Type material (examined). Holotype: ARGENTINA: Mendoza: ♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-69.80445&amp;materialsCitation.latitude=-35.868866" title="Search Plazi for locations around (long -69.80445/lat -35.868866)">Bardas Blancas</a> [35º52’07,93’’S 69º48’16,02’’W], Malargüe, 6–7.I.1975, E. Maury, A. Toth, P. Domínguez and C. Césari coll., (MACN-Ar 6877) ; Paratypes: ARGENTINA: Mendoza: 1♀, same data as Holotype (MACN-Ar 6878, Allotype) .</p><p>Other examined material: ARGENTINA: Mendoza: 1 ♀, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-69.819084&amp;materialsCitation.latitude=-35.802063" title="Search Plazi for locations around (long -69.819084/lat -35.802063)">Near</a> “Las Brujas” cavern [35º48’07,43’’S 69º49’08,72’’W], Bardas Blancas, Malargüe, 26.III.1975, E. Maury coll., (MACN-Ar 6879) ; 1 ♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-69.58291&amp;materialsCitation.latitude=-35.47502" title="Search Plazi for locations around (long -69.58291/lat -35.47502)">Malargüe</a> [35º28’30,08’’S 69º34’58,47’’W], 23.I.1979, E. Maury coll., (MACN-Ar) ; 1 ♂, Estancia Yaucha [34º12’19,22’’S 69º11’46,43’’W], A. Roig leg., (MACN-Ar).</p><p>Diagnosis: Pseudocleobis bardensis can be distinguished from all other Pseudocleobis except P. huinca and P. profanus sp. nov. by the male movable finger mucron with a narrow dorsal crest and shovel-like apex, and the flagellum with a prolateral subcircular subapical row of filaments. It can be distinguished from P. huinca and P. profanus sp. nov. by the less prominent dorsal crest and shovel-like apex of the movable finger mucron (Fig. 7E–F), and by the fixed finger mucron more robust, with the apex turning almost straight downward, forming a dorsal angle (Fig. 7E–F).</p><p>Description: see Maury (1976), otherwise, male and female similar to P. huinca and P. profanus sp. nov. except for the diagnostic characters. On males the MM tooth is of the same size or smaller than FM tooth, and the tiny FSD tooth may appear fused at the FD tooth base. Measurements in Table 1.</p><p>Distribution: Pseudocleobis bardensis was recorded from the Huarpe and Cuyan districts of the Cuyan High Andean biogeographic province (Fig. 4), at an altitudinal range of 1400–1600 m.a.s.l.</p></div>	https://treatment.plazi.org/id/B400FD2EFFBC090BFF102EF8FBF589EF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Iuri, Hernán A.;Iglesias, Mónica S.	Iuri, Hernán A., Iglesias, Mónica S. (2022): The genus Pseudocleobis Pocock, 1900 (Solifugae: Ammotrechidae) in transitional Patagonia-Monte deserts, with descriptions of two new species. Zootaxa 5094 (3): 435-460, DOI: 10.11646/zootaxa.5094.3.4
B400FD2EFFB90917FF1028AFFD668A7F.text	B400FD2EFFB90917FF1028AFFD668A7F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudocleobis mauryi Iuri & Iglesias 2022	<div><p>Pseudocleobis mauryi Iuri, sp. nov.</p><p>(Figs. 4; 8A, B; 9E; 11C; 12G–I. Table 1)</p><p>Pseudocleobis huinca: Maury 1976 misidentification (in part)</p><p>Type material: Holotype: ARGENTINA: Neuquén: ♂, “Area Natural Protegida Cuchillo Curá”, near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-70.38732&amp;materialsCitation.latitude=-38.612022" title="Search Plazi for locations around (long -70.38732/lat -38.612022)">Las Lajas</a>, Neuquén, Argentina [38º36’43,28’’S 70º23’14,36’’W], 16–22.I.2012, H.A. Iuri coll., (MACN-Ar 38419) ; Paratypes: ARGENTINA: Neuquén: 2 ♂, same data as holotype (MACN-Ar 38470, 38472) .</p><p>Other examined material: ARGENTINA: Neuquén: 1♂, Bajada “El Marucho” [39º26’16,06’’S 70º09’58,60’’W], 8.XII.1965, S. Schajovskoy coll., (MACN-Ar 6872, recorded by Maury 1976 as P. huinca) .</p><p>Etymology: The species name is a patronym in honor to Dr. Emilio A. Maury, former director of the Aracnological Division of the Museo Argentino de Ciencias Naturales “Bernardino Rivadavia”, who was the most important taxonomist of the order in South America, and who has made the most significant contributions on this genus.</p><p>Diagnosis: Pseudocleobis mauryi sp. nov. resembles P. huinca, P. bardensis and P. profanus sp. nov. in the body coloration pattern (darker in P. mauryi) and the general shape of the flagellum, but can be distinguished from them by the following characters: the ventral margin of fixed finger mucron is straight directed downwards and presents a tiny distal notch (Fig. 8B, DN) that makes the FD, FSD and FM be placed in a more vertical line and more anteriorly directed (Fig. 8A, B). The apex of the flagellum has a fringed retrolateral surface, with long filaments (Fig. 9E). The margins at the apex are folded so that the ventral and dorsal margins almost touch each other. The attachment ring is slightly smaller and placed, at level between the FP and the PFM tooth. The movable finger mucron is strongly curved upwards and the apex is acuminate. The movable finger teeth are close to each other (Figs. 8A, B). The difference of size between the MM and MP teeth is bigger than in any other species of Pseudocleobis .</p><p>Description: Male: measurements in Table 1. Color: Similar pattern to P. huinca, P. bardensis and P. profanus sp. nov., but darker in general.</p><p>Morphology and Chaetotaxy: as described for P. profanus sp. nov., except by the following details: Fixed finger: the fixed finger mucron is robust but much shorter than the movable finger mucron (Fig. 8A), and has a wide prolateral flagellar grove (Fig. 12G). The proventral flange is straight directed downwards and curved at the base forming a tiny notch (DN in Fig. 8B). The FD, FSD and FM teeth are strongly anteriorly directed, being placed almost in vertical line (Fig. 8A, B). The FSD tooth is very reduced (Fig. 8B). Movable finger: the MP is the biggest tooth of the chelicera; the MM is smaller than the FM tooth. The movable finger mucron is long and strongly curved upwards. Flagellum: the flagellum has several long filaments on the retrolateral surface of the apex (Figs. 9E). The apex is folded so that the ventral and dorsal margins almost touch each other. The attachment ring is at level between the FP and the PFM tooth (Fig. 8B). Pedipalps: the pedipalps have a similar spiniform setae armature as P. huinca, P. bardensis and P. profanus sp. nov. but with the basal pair of the tibia and basitarsus more setiform. Additionally, the other spiniform setae of tibia and basitarsus are more robust (but not longer) and more equally-sized.</p><p>Opisthosoma: with several bifurcated tip setae of different sizes and some filiform ctenidia on sternite III and IV. The ctenidia are weaker than in P. huinca, P. bardensis and P. profanus sp. nov. and more difficult to distinguish at optic microscope. We observed 2-2 on sternite III and 4-4 on sternite IV.</p><p>Female: Unknown.</p><p>Variability: one studied male (MACN-Ar 6872) has an anomalous right chelicera with an additional row of two fondal teeth.</p><p>Distribution: records of P. mauryi sp. nov. are from the Austral Payunia district of the Patagonian biogeographic province (Figs. 1C; 4), at about 900 masl.</p></div>	https://treatment.plazi.org/id/B400FD2EFFB90917FF1028AFFD668A7F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Iuri, Hernán A.;Iglesias, Mónica S.	Iuri, Hernán A., Iglesias, Mónica S. (2022): The genus Pseudocleobis Pocock, 1900 (Solifugae: Ammotrechidae) in transitional Patagonia-Monte deserts, with descriptions of two new species. Zootaxa 5094 (3): 435-460, DOI: 10.11646/zootaxa.5094.3.4
B400FD2EFFA60910FF102A52FB348B3A.text	B400FD2EFFA60910FF102A52FB348B3A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudocleobis solitarius : Maury 1976	<div><p>Pseudocleobis solitarius Maury, 1976</p><p>(Figs. 2D–F; 5; 6D, E; 8C–E; 9F; 10F; 11D; 12J–L; 15. Table 1, table 2)</p><p>Pseudocleobis solitarius: Maury 1976: 87–94 –101, figs.: 24–25; Maury 1983: 172.</p><p>Type material (examined). Holotype: ARGENTINA: Río Negro: ♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-67.62614&amp;materialsCitation.latitude=-39.113182" title="Search Plazi for locations around (long -67.62614/lat -39.113182)">Paso Córdoba</a> [39º06’47,46’’S 67º37’34,08’’W], General Roca, I.1961, A. Bachmann coll., (MACN-Ar 6886) ; Paratypes: ARGENTINA: Río Negro: 1♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-64.458496&amp;materialsCitation.latitude=-40.104084" title="Search Plazi for locations around (long -64.458496/lat -40.104084)">General Conesa</a> [40º06’14,71’’S 64º27’30,59’’W], 16.II.1941, M. Birabén coll., (MACN-Ar 6887) .</p><p>Other examined material: ARGENTINA: Mendoza: 3♂, 1♀, El Nihuil [35º01’40,53’’S 68º40’20,51’’W], 8.XI.1952, R. Cotta coll., (MACN-Ar 7589); Neuquén: 1♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-68.56503&amp;materialsCitation.latitude=-39.069313" title="Search Plazi for locations around (long -68.56503/lat -39.069313)">Arroyito</a> [39º04’09,53’’S 68º33’54,11’’W], 7.I.1986. M. &amp; P. Gentilé coll., (MACN-Ar) ; Río Negro: 1♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-64.93888&amp;materialsCitation.latitude=-40.73023" title="Search Plazi for locations around (long -64.93888/lat -40.73023)">San Antonio Oeste</a> [40º43’48,82’’S 64º56’19,98’’W], 19.I.1994, A. Roig coll., (MACN-Ar) ; 7♂, 11♀, 1 immature, Paso Córdoba, near the <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-67.620125&amp;materialsCitation.latitude=-39.11282" title="Search Plazi for locations around (long -67.620125/lat -39.11282)">Nautical Club</a> “Julio A. Trulls” [39º06’46,16’’S 67º37’12,45’’W], 16–21.I.2015, H.A. Iuri coll., (MACN-Ar) ; 5♂, 2♀, same locality and collector, I-2016, (MACN-Ar); 5♂, 1♀, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-66.17051&amp;materialsCitation.latitude=-40.666355" title="Search Plazi for locations around (long -66.17051/lat -40.666355)">Valcheta</a>, near the abandoned fluorite factory [40°39’58,87”S 66°10’13,84”W], 10– 12.I.2015, H.A. Iuri coll., (MACN-Ar) ; 3♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-65.024345&amp;materialsCitation.latitude=-41.646267" title="Search Plazi for locations around (long -65.024345/lat -41.646267)">Playas Doradas</a>, “sendero del cuis” [41°38’46.55”S 65° 1’27.64”W], 8–10.I.2015, H.A. Iuri coll., (MACN-Ar) .</p><p>Material tentatively assigned to P. solitarius: Argentina: San Juan: 1♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-67.8425&amp;materialsCitation.latitude=-30.163805" title="Search Plazi for locations around (long -67.8425/lat -30.163805)">Ichigualasto</a> [30º09’49,70’’S 67º50’32,98’’W], 17.XII.1979, E.A. Maury &amp; A. Roig coll., (MACN-Ar 7590) ; 1♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-67.65907&amp;materialsCitation.latitude=-31.589355" title="Search Plazi for locations around (long -67.65907/lat -31.589355)">Bermejo</a> [31º35’21,68’’S 67º39’32,67’’W], 15.XII.1979, E.A. Maury &amp; A. Roig coll., (MACN-Ar 7591) .</p><p>Diagnosis: The flagellum attached at level between FSD and FM teeth (Fig. 8D), and the stylet-like fixed finger mucron on males (Figs. 8C; 12J) are characteristic for this species. Pseudocleobis solitarius resembles P. mustersi, in the shape on the flagellum-mucron complex. However, P. mustersi has other unique modifications on the chelicerae such as a bifurcated movable finger mucron. All the others Pseudocleobis examined have a wider shaped flagellum with an attachment ring at level between the FSM and FP teeth or posterior.</p><p>Note: The female of P. solitarius has a wide, bell-shaped genital plate with a large oval opening in the middle (Fig. 10F). This shape is different from all other described females of Pseudocleobis . However, as the female genital plate has been proved to be uninformative at the species level in some cases (see P. huinca, P. bardensis and P. profanus), and the females of some species from southern Patagonia are unknown (e.g., P. mustersi, P. levii) we prefer not mention it as diagnostic for the species.</p><p>Redescription: Male: measurements in Table 1. Color in vivo (Fig. 2E–F). Specimens of the population from the type locality possess a whitish body coloration pattern with beige legs.</p><p>Color in 96 % ethanol. The pattern in alcohol is quite similar to the live specimens, except that the body turns more whitish and legs turn darker.</p><p>Morphology and Chaetotaxy: General body chaetotaxy as described for P. profanus . Fixed finger (Figs. 8C–D; 12J): the fixed finger mucron is much narrower than the movable finger mucron (Fig. 8C, D). The FSD smaller than FD and FM, but generally well developed, not vestigial (Fig. 8D). The FSM and FM teeth are separated by a short diastema. Movable finger (Fig. 8C–D): the movable finger is robust and possesses a wide dorsal crest on the mucron. The three teeth (MM, MSM, MP) are widely separated from each other. Flagellum (Figs. 8D; 9F; 12J): the flagellum in very narrow (Fig. 9F) with small attachment ring placed at level between FSD and FM teeth (Fig. 8D). The flagellum reaches almost the tip of mucron (Figs. 8D; 12J). The central portion of the prolateral surface possesses some prominent filaments (Fig. 12J). The membrane, in general, is thicker than the flagellum of the other species treated here, and with some flat filaments in the retrolateral surface. The apex has some surface roughness and a few, very small, projections. Pedipalps (Fig. 15A–C): chaetotaxy as described for P. profanus sp. nov. except that the spiniform setae are more setiform (thin and undulate), more equally sized (especially those of the tibia), and the clubbed setae were not seen. Leg I (Fig. 15D–F): similar to P. profanus, but the clubbed setae (Fig. 15F) are less abundant. Opisthosoma: ctenidia: 5-5 on sternite III and 4-4 on sternite IV (Fig. 15J).</p><p>Female: Measurements in Table 1. Color in vivo: As in Fig. 2D. Morphology and chaetotaxy similar to male, except by wider propeltidium, wider chelicerae, and more robust body in general (Fig. 2D). Pigment pattern similar to males. The spiniform setae of the pedipalps are more spiniform than in males (especially those of the tibia). Genital plate (Fig. 10F): the genital plate is relative wide, bell-shaped. There is a large oval opening in the middle, covered distally by two whitish rigid plates. The lateral margins are straight or concave. Short, thick, bifurcated tip setae are placed on the lateral margins.</p><p>Variability: Dentition variability on both sexes is detailed in Table 2. A careful examination shows that the dark brown areas on propeltidium and chelicerae of P. huinca, are whitish in the specimens of P. solitarius from Paso Córdoba. That coloration allows to easily distinguishing it from P. huinca (found in sympatry in the type locality). However, specimens of P. solitarius, from other populations (i.e., Playas Doradas and Valcheta), possess some brown pigment similar to P. huinca (also found in sympatry on these localities), so they cannot be easily distinguished by the coloration in these areas.</p><p>Distribution: The records of P. solitarius are from the Austral and Septentrional districts of the Monte biogeographic province (Figs. 1A–B, D; 5). However, we found females of Pseudocleobis with the P. solitarius genital plate shape at the MACN collection, from the Chubut and Santa Cruz districts of the Patagonian biogeographic province. As we could not study males from these localities, these females are included in the distribution map as solitarius shape. The males recorded by Maury (1983) for San Juan province have some slight differences in the shape of cheliceral fingers mucra, and they could belong to a different species. However, due to the scarcity of material, we maintain these here as P. solitarius .</p><p>Examined material of Pseudocleobis sp. (females solitarius shape): ARGENTINA: Chubut: 1♀, 15 km S of Los Tamariscos, 19.I.1977, E.A. Maury and Petriccha coll., (MACN-Ar) ; 1♀, Puerto Lobos, 1.II.2016, E.A. Maury coll., (MACN-Ar) ; 1♀, Puerto Lobos, Arroyo Verde, 25–26.I.1975, E.A. Maury, C. Césari and P. Domínguez coll., (MACN-Ar) ; 1♀, Puerto Madryn, XII.1981, P. Goloboff coll., (MACN-Ar) ; 1♀, Telsen, XII.1981, P. Goloboff coll., (MACN-Ar) ; 1♀, Rada Tilly, near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-67.55205&amp;materialsCitation.latitude=-45.952316" title="Search Plazi for locations around (long -67.55205/lat -45.952316)">Punta del Marqués</a> [45°57’8.33’’S 67°33’7.35’’W], 28–30.I.2015, H.A. Iuri coll., (MACN-Ar) ; Río Negro: 1♀, 5 km of Moligue, 1500 masl, 20.I.1973, (MACN-Ar) ; 1♀, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-65.38688&amp;materialsCitation.latitude=-41.604614" title="Search Plazi for locations around (long -65.38688/lat -41.604614)">Sierra Grande</a>, near the cemetary [41º36’16.62’’S 65º23’12.77’’W], 19–20.I.2016, H.A. Iuri coll., (MACN-Ar) .</p></div>	https://treatment.plazi.org/id/B400FD2EFFA60910FF102A52FB348B3A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Iuri, Hernán A.;Iglesias, Mónica S.	Iuri, Hernán A., Iglesias, Mónica S. (2022): The genus Pseudocleobis Pocock, 1900 (Solifugae: Ammotrechidae) in transitional Patagonia-Monte deserts, with descriptions of two new species. Zootaxa 5094 (3): 435-460, DOI: 10.11646/zootaxa.5094.3.4
