taxonID	type	description	language	source
BA492440124EFFF0FCD9FC7FFCEFA8D6.taxon	description	(Figs. 1 A, 2, 3 and 4 A-C) urn: lsid: zoobank. org: act: 549 E 3797 - 8024 - 4 C 89 - A 13 C- 547166490 EAA Type series: Holotype: male, Sungkang, Jing Ying Village, Ren-ai Township, 2 - VII- 2014, T. Yoshida leg. (NMNS, male genitalia illustrated). Paratypes: TAIWAN: [New Taipei City] 1 female & 1 ex., Wulai District, 4 - VI- 1970, Y. Hori leg. (EUMJ). [Yilan County] 2 exs, Fushan Botanical Garden, Yuanshan Township, 25 - 27 - IX- 2000, L. Papp, L. Peregovits & L. Ronkay leg., at light (HNHM; studied by MK); 2 exs, Cueifong Lake, Nan-ao Township, alt. 1950 m, 1 - IV- 2004, T. Kurihara leg. (ELKU). [Nantou County] 1 ex., Gaofeng, 7 - 8 - V- 2009, J. Aoki leg. (ELKU); 3 males, 1 female & 2 exs, Tun-yuan-trail, Jing Ying Village, Ren-ai Township, 1 - VII- 2014, T. Yoshida leg. (ELKU); 3 exs, same locality, 6 - VII- 2014, T. Yoshida leg. (ELKU); 10 exs, same data as holotype. (ELKU); 1 male, Tsiufeng, Ren-ai Township, 10 - VII- 1966, H. Kamiya leg. (KUM). [Chiayi County] 1 ex., Alishan, Alishan Township, 1 - VIII- 1968, K. Y. leg. (ELKU); 1 ex., Fenchihu, Zhuqi Township, 22 - VII- 1966, H. Kamiya leg. (KUM); 2 exs, same locality, 23 - VII- 1966, H. Kamiya leg. (KUM); 1 ex., same locality, 24 - VII- 1966, H. Kamiya leg. (KUM); 1 ex., same locality, 22 - VI- 1968, M. Tomokuni leg. (EUMJ). [Kaohsiung City] 1 ex., Shanping, Liouguei District, 20 - 21 - III- 1980, T. Niisato leg. (EUMJ); 1 ex., Mt. Su-shan (Shi Shan) near Liouguei, Taoyuan District, 25 - IV- 1986, K. Baba leg. (KUM). Etymology: The specific name is derived from Taiwan, the type locality of the new species. Diagnosis: This species resembles P. piceus Grouvelle, 1882; it is distinguished by the distal portions of parameres bearing dense short pubescence, in comparison with distal portions of parameres bearing only two apical setae in P. piceus. This species also resembles P. harmandi but is easily distinguished by the presence of a row of long erect setae along each lateral elytral margin, the expanded anterior angles of pronotum and the stick-shaped and elongate parameres (cone-shaped and stout in P. harmandi) (Figs. 3 and 4). Description: Body length: 2.83 - 3.43 mm (n = 20). Coloration: (Figs. 1 A and 2). Head and pronotum dark brown. Elytra somewhat lighter colored, with a quadrate black macula on each elytron at middle, connected by narrow horizontal band at posterior 1 / 3, round maculae at posterior 1 / 4, connected to lateral margins, darkened around humeri and end of elytra. Antennae dark brown, 7 th and / or 8 th to 10 th antennomere darker, 11 th (apex) yellowish-brown. Head: (Fig. 2 A and B). Wide, HW / HL 1.28 - 1.43; IE / HL 0.86 - 0.98. Temples somewhat enlarged behind eyes, narrowed gradually toward basis. Eyes of moderate size, slightly prominent. Dorsal surface with coarse punctation. Antennae long; antennomeres with pubescence of moderate to large length; distal portion of 7 th to 10 th and entire 11 th antennomeres with short pubescence, very dense on 8 th to 11 th antennomeres; approximate ratios of antennomere lengths of holotype as follows: 2.6: 1.0: 1.2: 1.3: 1.3: 1.5: 1.4: 1.2: 1.1: 1.0: 1.7. Pronotum: (Figs. 2 A, B and 4 B). Subquadrate, widest near anterior margin, straightly narrowed toward base, lateral margins slightly flattened; lateral areas of basis somewhat flattened, PW / PL 1.18 - 1.27. Punctation on pronotal disk as on vertex; pubescence composed of numerous setae of moderate length, very long setae on teeth of lateral margins and anterior and posterior angles (Fig. 4 B). Anterior angle with two small teeth and few small tubercles; lateral margin with some very small teeth, anterior tooth largest; posterior angle with small tooth (Fig. 4 B). Elytra: (Figs. 2 A and 4 C). Elongate-oval, EW / BL 0.40 - 0.45. Rows of punctures wider than interstices. Pubescence composed of numerous semi-erect setae of medium length, some very long, erect setae along each lateral margin, longer toward humeral swelling, with some long setae on lateral margin (Fig. 4 C). Male genitalia: (Fig. 3). Spiculum gastrale (Fig. 3 A) with long and very thin strut, diverging widely around apex; branches slightly widened, connected by membrane; lateral sclerites elongate, curved inwards around apices, with short and thin strut. Parameres (Fig. 3 B) stick-shaped, elongate, depressed at base, protruding dorsally at inner basal angles, narrowed at basal 1 / 3 to 2 / 3, dorsally densely punctate, punctures of various size on basal 1 / 3, with a short seta on each inner margin of basal 1 / 3, dorsally densely with short setae on distal 1 / 2 or 1 / 3 to apices, with a long seta and a very long seta on each apex. Phallobase (Fig. 3 B) long, upper layer deeply and widely incised at distal 1 / 3, with incision covered by a membrane, lower layer thin, extending toward base. Penis (Fig. 3 B) long, sub-parallel, gradually narrowed around apex, with rounded protrusion at apex, finely punctate near apex. Internal sac recurved around apex, with a thin ringed structure and a thin apical strut, densely armed with numerous spines on apical half near space of the ringed structure, with two spinous parallel armed lines around middle, with dense, minute spines near middle. Distribution: Taiwan. Remarks: This new species is distributed in high altitudes of Taiwan and occurs sympatrically with P. harmandi. On Tun-yuan-trail, the species was collected from dead leaves of broad-leaved trees.	en	Yoshida 1, Takahiro, Karner, Michael, Hirowatari, Toshiya (2018): Chtenopteryx canariensis Salcedo-Vargas & Guerrero-Kommritz 2000. Zoological Studies 57 (18): 1-18, DOI: 10.6620/ZS.2018.57-18, URL: http://dx.doi.org/10.5281/zenodo.13316922
BA492440124AFFFCFCF9FE1FFF74AC96.taxon	description	(Figs. 1 B, 5 and 6)	en	Yoshida 1, Takahiro, Karner, Michael, Hirowatari, Toshiya (2018): Chtenopteryx canariensis Salcedo-Vargas & Guerrero-Kommritz 2000. Zoological Studies 57 (18): 1-18, DOI: 10.6620/ZS.2018.57-18, URL: http://dx.doi.org/10.5281/zenodo.13316922
BA492440124AFFFCFCF9FE1FFF74AC96.taxon	materials_examined	Type series: Lectotype: male, “ Ceylan | Horn. ”; “ Type ” [red label]; “ Museum Paris | Collection | Grouvelle ”; “ Psammoecus | delicatus | ty. | Grouv ” [Grouvelle’s hand] (MNHN; studied by MK in 2015, here designated). Paralectotypes: 4 females, “ Sumatra | Palembang ”; “ delicatus ” [printed label] (MNHN; studied by MK in 2015, deposited in the collection of Grouvelle and Horn). Lectotype of P. nitidior syn. nov.: male, “ Java ”; “ Type ” [red label]; “ nitidior | ty. Grouv ”. (MNHN; studied by TY in 2017, here designated). Paralectotypes of P. nitidior syn. nov.: 6 exs, same label data as the lectotype. (MNHN; studied by TY in 2017, here designated). Lectotype of P. raffrayi syn. nov.: female, “ Singapore ”; “ Type ” [red label]; “ Raffrayi | ty. A. Grouv ”. (MNHN; studied by TY in 2017, here designated). Lectotype of P. x-notatus syn. nov.: male, “ H. | Sauter. | Akau | Formosa | 1 - 10.12.07 [round label] ”; “ Psammoechus [sic] | x-nigrum | type. Grouv. ” [not Grouvelle’s hand]; “ Museum Leiden. | Psammoe | cus | x notatus | Det: Grouv ”; “ type ” [blue label]; “ Cat. No | 5 ” (RMNH; studied by MK in 2014, here designated). Paralectotypes of P. x-notatus syn. nov.: 2 exs, same label data as the lectotype. (MNHN; studied by TY in 2017, here designated); 1 female, “ H. | Sauter. | Kosempo | Formosa | Jan. 08 ” [round label]; “ Psammoecus | x nigrum | G. Grouv ” [Grouvelle’s hand]; “ type ”; “ Museum Leiden. | Psammoecus | x notatus | Det: Grouv ”; “ type ” [blue label]; “ Type ” [red label] (RMNH; studied by MK in 2014, here designated); 1 male and 1 ex., same label data. (MNHN; studied by TY in 2017, here designated). Specimens examined: TAIWAN: [Miaoli County] 1 male (Catalog No. NMNS ENT 3235 - 1443), Sanyi, 14 - 16 - VI- 1999, C. S. Lin & W. T. Yang leg., UV Light. (NMNS). [Taichung City] 2 exs, Wufeng, Wufeng Disctrict, 24 - IV- 1992, Smetana leg. (MHNG; studied by MK); 1 ex., Fengyuan, 22 - IV- 1990, Smetana leg. (MHNG; studied by MK). [Nantou County] 1 male (Catalog No. NMNS ENT 1602 - 131), Chushan (= Zhushan Township), 8 - 9 - XI- 1993, C. S. Lin & W. T. Yang leg., UV Light. (NMNS); 1 ex. (Catalog No. NMNS ENT 3513 - 284), Lienhuachih, Yuchi Township, 6 - VII- 12 - VIII- 1998, C. S. Lin & W. T. Yang leg., Malaise Trap (NMNS); 1 ex., NW Yuchi, Yuchi Township, 8 - X- 2009, Dányi & Lazányi leg. (HNHM: studied by MK); 1 ex. (Catalog No. NMNS ENT 1582 - 328), Fenghuangku, Lugu Township, 20 - 22 - IX- 1993, C. S. Lin & M. L. Chan leg., UV Light (NMNS); 1 male (Catalog No. 1337 - 5), Shih-tzu-tou, 6 - IX- 1987, I. C. Hsu leg., Sweeping Net. (NMNS). [Chiayi County] 1 ex., Chuchi, Zhuqi Township, 20 - VII- 1966, H. Kamiya leg. (KUM). [Kaohsiung City] 7 exs, Kosempo (= Jiaxian District), I- 1908, H. Sauter leg. (ZMHB, HNHM; studied by MK); 1 male and 4 exs, same label data. (SDEI); 1 ex., Mt. Ta Yuen Shan near Liouguei, 5 - VI- 1989, K. Baba leg. (KUM); 1 ex., Liu Kui (= Liouguei), 2 - VIII- 1989, K. Baba leg. (KUM); 1 ex., same locality, 30 - III- 1986, K. Baba leg. (EUMJ); 1 ex., Shanping Forest Recreation Area, near Liukuei, 19 - 21 - XI- 2009, Ronkay & Merkl leg. (HNHM; studied by MK). [Taitung County] 8 exs, Pilam (= Taitung), II- 1908, H. Sauter leg. (HNHM; studied by MK). [Pingtung County] 11 exs, Akau (= Pingtung City), 1 - 10 - XII- 1907, H. Sauter leg. (HNHM; studied by MK); 7 exs, same locality, 1 - 10 - XII- 1907, H. Sauter leg. (ZMHB; studied by MK); 2 males and 8 exs, same locality, XII- 1907, H. Sauter leg. (SDEI); 1 ex. (Catalog No. NMNS ENT 6920 - 303), Nanjenshan, Manzhou Township, 1 - 19 - IV- 2009, M. L. Jeng & T. R. Chen leg., SLAM Trap. (NMNS); 1 ex. (Catalog No. NMNS ENT 6920 - 82), same locality & same period, M. L. Jeng & T. R. Chen leg., Flight Interception Trap. (NMNS); 2 exs (Catalog No. NMNS ENT 6469 - 40 & 48), same locality, 21 - VI- 4 - VII- 2009, M. L. Jeng & T. R. Chen leg., Flight Interception Trap. (NMNS); 1 male & 1 ex. (Catalog No. NMNS ENT 6469 - 486 & 503), same locality, 4 - 14 - VII- 2009, M. L. Jeng & T. R. Chen leg., Flight Interception Trap. (NMNS); 1 ex. (Catalog No. NMNS ENT 6674 - 322), same locality, 27 - IX- 12 - X- 2009, M. L. Jeng & T. R. Chen leg., Flight Interception Trap. (NMNS); 1 male (Catalog No. NMNS ENT 6632 - 57), Manchou (= Manzhou) Lite Chulaoshushan, 30 - VIII- 14 - IX- 2009, M. L. Jeng & T. R. Chen leg., Malaise Trap. (NMNS); 1 ex. (Catalog No. NMNS ENT 6579 - 378), same locality, 12 - 26 - IX- 2009, M. L. Jeng & T. R. Chen leg., Malaise Trap. (NMNS); 1 male (Catalog No. NMNS ENT 6525 - 85), Manchou (= Manzhou) Lite No: 169 Hsien Rd. 3.7 km, 19 - IV- 1 - V- 2009, M. L. Jeng & T. R. Chen leg., Flight Interception Trap. (NMNS); 1 ex., Sichongxi, Checheng Township, 1 - I- 1986, K. Baba leg. (EUMJ); 1 male. (Catalog No. NMNS ENT 6632 - 57), Manchou (= Manzhou) Lite Chulaoshushan, 30 - X- 14 - IX- 2009, M. L. Jeng & T. R. Chen leg., Malaise Trap. (NMNS). Diagnosis: This species differs from the other Taiwanese species by the very long and slender teeth on lateral margins of its pronotum, the pattern of these teeth, the pubescence composed of moderate and long setae on head, pronotum and elytra, and its large, strongly protruding eyes. Psammoecus delicatus is related to P. complexus Pal, 1985, but can easily be distinguished by the different shape of the lateral pronotal teeth (see Pal 1985). Description: Body length: 2.38 - 2.98 mm (n = 16). Coloration: (Figs. 1 B and 5). Head and pronotum yellowish-brown. Elytra yellowish-brown with variable x-shaped blackish-brown maculae covering posterior 2 / 3 (Fig. 5 A and B); posterior half of maculae in lighter color specimen obscure or absent; sometimes posterior 1 / 3 of elytra darkened (Fig. 5 C). Antennae yellowish-brown basally, 7 th to 10 th antennomere sometimes darkened, 11 th antennomere lighter colored, rarely also 10 th antennomere lighter colored (Fig. 5 C). Head: (Fig. 5). Wide, HW / HL 1.34 - 1.47; IE / HL 0.79 - 0.89. Temples gradually narrowed toward bases. Eyes large, well prominent. Dorsal surface punctation sparse and weak, pubescence composed of moderate and long setae. Antennae long; antennomeres with moderate and long, erect setae; distal portion of 7 th to 10 th and entire 11 th antennomere covered with short pubescence, dense on 8 th to 11 th; approximate ratios of antennomere lengths of lectotype as follows: 3.0: 1.0: 1.3: 1.4: 1.4: 1.5: 1.2: 1.2: 1.2: 1.2: 2.4. Pronotum: (Fig. 5). Wide, point of greatest width variable, between anterior 1 / 3 and middle, distinctly narrowed toward base; lateral areas slightly flattened, PW / PL 1.32 - 1.42. Punctation on dorsal surface denser and slightly stronger than on vertex. Pubescence composed of numerous moderate and fewer long setae, long seta on each tooth and tubercle on anterior, lateral and posterior margins and anterior and posterior angles. Anterior angle with a few large tubercles; lateral margin with some dorso-laterally bent short to long teeth; posterior angle with a very small tooth and a small tubercle. Elytra: (Fig. 5). Oval, EW / BL 0.39 - 0.44. Rows of punctures slightly wider than interstices. Pubescence composed of many semi-erect moderate setae, most of interstices with rows of long erect setae, very long erect setae in a row along lateral margins, setae longer toward humeral swelling, anterior setae on small tubercles, with some long setae on lateral margin. Male genitalia: (Fig. 6). Spiculum gastrale (Fig. 6 A) with short and thick strut, enlarged around base, diverging widely near middle; branches obscure and thin, connected by membrane; lateral sclerites elongate triangular, enlarged inwards near anterior 1 / 3. Parameres (Fig. 6 B) small and stout, fused basally, rounded at apices, apex sometimes somewhat angular, dorsally with some small punctures around middle of inner margins, densely covered with setae of various length near apices, dorsally with some short, sparse setae near inner margins of apical 1 / 3, with a long apicolateral seta. Phallobase (Fig. 6 B) long, subparallel, basal half of upper layer diverging, the branches widening distally, with membrane between bases of the branches, lower layer narrow, exposed around base. Penis (Fig. 6 C) long, gradually narrowed toward apex, somewhat rounded at apex, with fine dense punctation around apex. Internal sac with very long and thin structure partly ringed around apex, with a pair of small spines near middle (sometimes absent), with a pair of large spines around base. Distribution: Taiwan; India; Sri Lanka; Singapore; Indonesia. Remarks: The external morphological characters as well as the male genital structures of P. x-notatus and P. nitidior type specimens show the typical morphology of P. delicatus. Hence, we conclude that these species are junior synonyms of P. delicatus. Although Grouvelle’s description of P. raffrayi suggests that at least two syntypes should be present, only one female specimen could be found and studied. Since P. delicatus is well characterized by its external morphological characters, we have no doubt that P. raffrayi is a junior synonym. The unusual morphology of the spiculum gastrale is remarkable compared to that of other Psammoecus. This may indicate that this species belongs to a distinct species group. Studies of extensive material from the Oriental region revealed numerous closely related yet undescribed species.	en	Yoshida 1, Takahiro, Karner, Michael, Hirowatari, Toshiya (2018): Chtenopteryx canariensis Salcedo-Vargas & Guerrero-Kommritz 2000. Zoological Studies 57 (18): 1-18, DOI: 10.6620/ZS.2018.57-18, URL: http://dx.doi.org/10.5281/zenodo.13316922
BA4924401246FFFEFF1FF9DFFE6FA836.taxon	description	(Figs. 1 C, 4 D-F and 7)	en	Yoshida 1, Takahiro, Karner, Michael, Hirowatari, Toshiya (2018): Chtenopteryx canariensis Salcedo-Vargas & Guerrero-Kommritz 2000. Zoological Studies 57 (18): 1-18, DOI: 10.6620/ZS.2018.57-18, URL: http://dx.doi.org/10.5281/zenodo.13316922
BA4924401246FFFEFF1FF9DFFE6FA836.taxon	materials_examined	S p e c i m e n s e x a m i n e d: J A PA N: 1 m a l e, Yoshin, Tanzawa, Kanagawa Prefecture, 26 - V- 1989, Y. Hirano leg. (EUMJ; formerly holotype of Psammoecus boreas). TAIWAN: [Yilan County] 1 male & 1 ex., Taipingshan, Datong Township, alt. 1950 m, 1 - IV- 2004, T. Kurihara leg. (EUMJ). [Hualien County] 1 ex., Piluhsi For. Rec. Area, 22 - V- 1999, 2150 m, S. E. Halbert & C. W. L. O’Brien leg., Night Beating (FSCA). [Nantou County] 1 ex., Pi Lu Chieh, alt. 2400 m, 6 - IX- 1986, K. Baba leg. (EUMJ); 1 ex., Gaofeng, 7 - 8 - V- 2009, J. Aoki leg. (ELKU); 4 exs, Wushe, Ren-ai Township, 16 - III- 1983, H. & M. Townes leg. (FSCA); 2 exs, Meifeng, Ren-ai Township, 3 - V- 1983, Henry K. Townes leg., Insect Flight Trap. (FSCA); 2 males, 1 female & 16 exs, Tun-yuan-trail, Jing Ying Village, Ren-ai Township, 5 - VII- 2014, T. Yoshida leg. (ELKU); 1 male & 4 exs, same locality, 6 - VII- 2014, T. Yoshida leg. (ELKU); 1 ex., N 24 ° 02 ' 53.0 " E 121 ° 12 ' 55.5 ", Ren-ai Township, 6 - VIII- 2008, Barclay, Mendel & Ewers leg. (BMNH; studied by MK). [Chiayi County] 1 ex., Fenchihu, Zhuqi Township, 10 - VII- 1966, H. Kamiya leg. (KUM); 5 exs, same locality, 16 - VII- 1966, H. Kamiya leg. (KUM); 2 exs, same locality, 23 - VII- 1966, H. Kamiya leg. (KUM); 7 exs, same locality, 24 - VII- 1966, H. Kamiya leg. (KUM); 1 ex., same locality, 27 - VII- 1966, H. Kamiya leg. (KUM); 1 ex., same locality, 28 - VII- 1966, H. Kamiya leg. (KUM). [Taitung County] 5 exs, Taimali-shan (Mt.), Jinfeng Township, alt. ca. 1300 m, 22 ° 38 ' N 120 ° 57 ' E, 23 - VI- 2015, J. Yamasako leg. (ELKU). [Kaohsiung City] 1 male, vic. Fujieda (= Tengzhi) nr. Taoyuan, Taoyuan District, at light, 6 - IV- 1976, K. Ushijima leg. (EUMJ).	en	Yoshida 1, Takahiro, Karner, Michael, Hirowatari, Toshiya (2018): Chtenopteryx canariensis Salcedo-Vargas & Guerrero-Kommritz 2000. Zoological Studies 57 (18): 1-18, DOI: 10.6620/ZS.2018.57-18, URL: http://dx.doi.org/10.5281/zenodo.13316922
BA4924401246FFFEFF1FF9DFFE6FA836.taxon	diagnosis	Diagnosis: This species resembles P. taiwanensis sp. nov. but is easily distinguished by the absence of a row of long erect setae along each lateral elytral margin, the weak anterior angles of pronotum and the cone-shaped and stout parameres (see Yoshida and Hirowatari 2014). This species also resembles P. trimaculatus and other closely related species, but it differs from them by the shorter lateral teeth of its pronotum.	en	Yoshida 1, Takahiro, Karner, Michael, Hirowatari, Toshiya (2018): Chtenopteryx canariensis Salcedo-Vargas & Guerrero-Kommritz 2000. Zoological Studies 57 (18): 1-18, DOI: 10.6620/ZS.2018.57-18, URL: http://dx.doi.org/10.5281/zenodo.13316922
BA4924401246FFFEFF1FF9DFFE6FA836.taxon	description	Coloration: (Figs. 1 C and 7). Head and pronotum yellowish-brown. Elytra sometimes a little lighter colored, with variable black maculae near posterior 1 / 3; round maculae around middle of each elytron, another round one near posterior 1 / 3 close to elytral suture (Fig. 7 C): maculae sometimes connected to each other (Fig. 7 A) or forming a wide v-shaped covering posterior half of elytra (Fig. 7 D). Antennae yellowish-brown basally, 9 th and 10 th antennomere black, 11 th yellowish-brown, sometimes darkened.	en	Yoshida 1, Takahiro, Karner, Michael, Hirowatari, Toshiya (2018): Chtenopteryx canariensis Salcedo-Vargas & Guerrero-Kommritz 2000. Zoological Studies 57 (18): 1-18, DOI: 10.6620/ZS.2018.57-18, URL: http://dx.doi.org/10.5281/zenodo.13316922
BA4924401246FFFEFF1FF9DFFE6FA836.taxon	distribution	Distribution: Taiwan; Japan; Thailand (new record); Nepal; India.	en	Yoshida 1, Takahiro, Karner, Michael, Hirowatari, Toshiya (2018): Chtenopteryx canariensis Salcedo-Vargas & Guerrero-Kommritz 2000. Zoological Studies 57 (18): 1-18, DOI: 10.6620/ZS.2018.57-18, URL: http://dx.doi.org/10.5281/zenodo.13316922
BA4924401246FFFEFF1FF9DFFE6FA836.taxon	discussion	Remarks: This species occurs sympatrically with P. taiwanensis sp. nov. Psammoecus boreas, now synonymized with P. harmandi, was described from Japan based on diagnostic differences from P. harmandi according to a redescription of P. harmandi by Pal (1985). Yoshida and Hirowatari (2014) pointed out that P. harmandi differed from P. boreas by its shorter antennae and slightly transverse 10 th antennomere. Although we could not examine the type of P. harmandi, we had an opportunity to examine a male specimen of P. harmandi from northern India and found these two species to be indistinguishable. This species is widely distributed; specimens from Japan, Taiwan, Thailand, Nepal, and India have been studied. Except for Hokkaido (Japan) where the northern limit of its distribution might be reached, they are exclusively distributed in high altitudes. Compared to the coloration described by Pal (1985), the Taiwanese specimens show considerable intraspecific variation. We have not seen specimens outside of Taiwan with strongly enlarged black maculae that cover the posterior half of the elytra (Fig. 7 D). Examples for the variability are given in figure 7.	en	Yoshida 1, Takahiro, Karner, Michael, Hirowatari, Toshiya (2018): Chtenopteryx canariensis Salcedo-Vargas & Guerrero-Kommritz 2000. Zoological Studies 57 (18): 1-18, DOI: 10.6620/ZS.2018.57-18, URL: http://dx.doi.org/10.5281/zenodo.13316922
BA4924401244FFFEFF1BFEBFFAA8ACF6.taxon	description	(Fig. 1 D)	en	Yoshida 1, Takahiro, Karner, Michael, Hirowatari, Toshiya (2018): Chtenopteryx canariensis Salcedo-Vargas & Guerrero-Kommritz 2000. Zoological Studies 57 (18): 1-18, DOI: 10.6620/ZS.2018.57-18, URL: http://dx.doi.org/10.5281/zenodo.13316922
BA4924401244FFFEFF1BFEBFFAA8ACF6.taxon	materials_examined	Type series: Lectotype: male, “ N. Guinea | Hatam Vll | Beccari 1875 ”; “ SYNTYPUS | Psammoecus | dentatus | Grouvelle, 1882 ” (MSNG; studied by MK in 2015 and TY in 2017, here designated). Paralectotypes: 1 male and 1 female, same label data as lectotype. (MSNG; studied by MK in 2015, here designated). Specimens examined: JAPAN: 1 male, Kûra, Ishigaki Island, Okinawa Prefecture, 22 - IX- 2012, T. Yoshida leg. (EUMJ; formerly holotype of P. scitus). TAIWAN: [Hualien County] 1 ex., Liyuchih, 14 - VIII- 1966, H. Kamiya leg. (KUM); 2 exs, same locality, 15 - VIII- 1966, H. Kamiya leg. (KUM); 2 exs, Juisui (Naka spa) (= Ruisui Township), 27 - XI- 1962, K. Baba leg. (EUMJ); 2 exs, Fu Yuan For. Rec. Area, 28 - V- 1997, C. W. & L. B. O’Brien leg. (FSCA). [Nantou County] 1 male (Catalog No. NMNS ENT 1582 - 60), Fenghuangku, Lugu Township, 20 - 22 - IX- 1993, C. S. Lin & M. L. Chan leg., UV Light (NMNS). [Chiayi County] 1 ex. (Catalog No. NMNS ENT 425 - 406), Yunghsing, Dapu Township, 15 - XII- 1988, K. W. Huang leg., Sweeping Net (NMNS). [Taitung County] 2 males & 3 exs, Doulan, Donghe Township, 9 - IV- 2012, J. Yamasako leg. (ELKU); 1 ex., Linchan, Daren Township, 6 - VI- 2013, S. Tsuyuki leg. (ELKU); 5 exs, Jhiben Hot Spring, Beinan Township, 24 - V- 1969, S. Hisamatsu leg. (EUMJ); 1 ex., Lutao (= Green Island), Green Island Township, 26 - 28 - III- 1998, M. Sato leg (EUMJ); 3 exs, Island Lanyu (= Orchid Island), Lanyu Township, 8 - IV- 1978, Kôzô Murakami leg. (EUMJ); 15 exs, Yung Hsin Farm, Lanyu Island (= Orchid Island), Lanyu Township, 29 - 30 - VII- 2008, Mendel & Barclay leg. (BMNH; studied by MK). [Kaohsiung City] 1 ex., Kosempo (= Jiaxian District), I- 1908, H. Sauter leg. (ZMHB; studied by MK); 1 ex., Paiyunshan (= Baiyunshan), near Chiashien (= Jiaxian), Jiaxian District, 21 - III- 1977, Yutaka Notsu leg. (EUMJ); 2 exs, Fa Kuo Shan, near Liu Kui (= Liouguei), alt. 800 m, 6 - X- 1986, K. Baba leg. (EUMJ); 1 ex., Liu Kui (= Liouguei), 30 - III- 1986, K. Baba leg. (EUMJ); 1 ex., same locality, 2 - IV- 1986, K. Baba leg. (EUMJ); 1 ex., same locality, 4 - VIII- 1978, T. Niisato leg. (EUMJ). [Tainan City] 1 ex., Kantoushan, Dongshan District, 17 - II- 2013, W. - C. Liao leg. (TARI). [Pingtung County] 2 exs, Hengchun, 4 - VIII- 1966, H. Kamiya leg. (KUM); 1 ex. (Catalog No. NMNS ENT 6215 - 77), Chungchienlu, Mudan Township, 27 - 28 - VII- 2009, W. T. Yang & K. W. Huang leg., UV Light Trap (NMNS); 1 male & 1 female, Dong Yuan Forest Recreation Area, Dong Yuan Village, Mudan Township, Pingtung City, 10 - VII- 2014, T. Yoshida leg. (ELKU); 2 exs (Catalog No. NMNS ENT 6259 - 276 & 580), Chiupeng, Manzhou Township, 19 - IV- 1 - V- 2009, M. L. Jeng & T. J. Chen leg., Flighintercept Net. (NMNS); 2 exs, Kenting, 4 - IX- 1983, J. B. Happner leg. (FSCA); 1 ex., Kenting Park, 17 - VIII- 1968, M. Nishikawa leg. (EUMJ); 1 ex., same locality, 17 - III- 1977, Yutaka Notsu leg. (EUMJ). Diagnosis: This species can be distinguished from other Taiwanese Psammoecus species by the small, even-sized lateral teeth on its pronotum and the irregular distances between the adjacent teeth. Distribution: Taiwan; Japan; New Guinea. Remarks: This species was described as P. scitus by Yoshida and Hirowatari (2014). This study examines many specimens in this species, including P. dentatus and P. scitus type specimens. We found no significant differences between these two species in their morphology, including their male genitalia; thus, P. scitus is synonymized with P. dentatus. Specimens from an extremely large area have been studied in the course of related revisionary work. Psammoecus dentatus may be recorded from many more South East Asian localities.	en	Yoshida 1, Takahiro, Karner, Michael, Hirowatari, Toshiya (2018): Chtenopteryx canariensis Salcedo-Vargas & Guerrero-Kommritz 2000. Zoological Studies 57 (18): 1-18, DOI: 10.6620/ZS.2018.57-18, URL: http://dx.doi.org/10.5281/zenodo.13316922
BA4924401244FFF9FC9EFA7FFDAEAF56.taxon	description	(Fig. 1 E) Psammaechus [sic] trimaculatus Motschulsky, 1858: Etud. Ent. 7: 45. Type locality: Sri Lanka. (referring to Hetschko (1930 ))	en	Yoshida 1, Takahiro, Karner, Michael, Hirowatari, Toshiya (2018): Chtenopteryx canariensis Salcedo-Vargas & Guerrero-Kommritz 2000. Zoological Studies 57 (18): 1-18, DOI: 10.6620/ZS.2018.57-18, URL: http://dx.doi.org/10.5281/zenodo.13316922
BA4924401244FFF9FC9EFA7FFDAEAF56.taxon	diagnosis	Diagnosis: This species is closely similar to P. triguttatus and P. labyrinthicus, and it is difficult to distinguish them by their external characters. This species can be distinguished from the former by the larger basis of parameres and the wider distance between the posterior margin of the phallobase and the deepest point of incision of the anterior margin of the phallobase, and from the latter by the larger bases of parameres and the wider protuberances on the inner margins of the branches of the anterior phallobase (see Yoshida and Hirawatari 2014). Distribution: Taiwan; Japan (Yoshida and Hirowatari 2014); Russia (Kovalev 2016); Madagascar (Karner 2012); Mauritius, France (Réunion), South Africa, Tanzania, Uganda (Karner 2014); Nepal, India and Bhutan (Pal 1985). Remarks: Pal (1985) provided a redescription of this species, Yoshida and Hirowatari (2014) provided a detailed description of male genital structures of this species. Only male genitalia provide reliable diagnostic characters when separating P. trimaculatus from closely related species. Accordingly, Yoshida and Hirowatari (2014) only considered data from specimens that were determined by study of genital structures to be reliable. The same practice was applied in the present study.	en	Yoshida 1, Takahiro, Karner, Michael, Hirowatari, Toshiya (2018): Chtenopteryx canariensis Salcedo-Vargas & Guerrero-Kommritz 2000. Zoological Studies 57 (18): 1-18, DOI: 10.6620/ZS.2018.57-18, URL: http://dx.doi.org/10.5281/zenodo.13316922
BA4924401243FFF9FC26FB5FFB87AF87.taxon	diagnosis	Diagnosis: This species is closely similar to P. trimaculatus and P. triguttatus (see the diagnosis of P. trimaculatus and Yoshida and Hirawatari 2014).	en	Yoshida 1, Takahiro, Karner, Michael, Hirowatari, Toshiya (2018): Chtenopteryx canariensis Salcedo-Vargas & Guerrero-Kommritz 2000. Zoological Studies 57 (18): 1-18, DOI: 10.6620/ZS.2018.57-18, URL: http://dx.doi.org/10.5281/zenodo.13316922
BA4924401243FFF9FC26FB5FFB87AF87.taxon	distribution	Distribution: Taiwan; Japan.	en	Yoshida 1, Takahiro, Karner, Michael, Hirowatari, Toshiya (2018): Chtenopteryx canariensis Salcedo-Vargas & Guerrero-Kommritz 2000. Zoological Studies 57 (18): 1-18, DOI: 10.6620/ZS.2018.57-18, URL: http://dx.doi.org/10.5281/zenodo.13316922
BA4924401243FFF9FC26FB5FFB87AF87.taxon	discussion	Remarks: Yoshida and Hirowatari (2014) redescribed this species in detail with the male genital structure.	en	Yoshida 1, Takahiro, Karner, Michael, Hirowatari, Toshiya (2018): Chtenopteryx canariensis Salcedo-Vargas & Guerrero-Kommritz 2000. Zoological Studies 57 (18): 1-18, DOI: 10.6620/ZS.2018.57-18, URL: http://dx.doi.org/10.5281/zenodo.13316922
BA4924401243FFF9FF4AF99FFB87AD16.taxon	description	(Fig. 1 F)	en	Yoshida 1, Takahiro, Karner, Michael, Hirowatari, Toshiya (2018): Chtenopteryx canariensis Salcedo-Vargas & Guerrero-Kommritz 2000. Zoological Studies 57 (18): 1-18, DOI: 10.6620/ZS.2018.57-18, URL: http://dx.doi.org/10.5281/zenodo.13316922
BA4924401243FFF9FF4AF99FFB87AD16.taxon	diagnosis	Diagnosis: This species is closely similar to P. trimaculatus and P. labyrinthicus (see the diagnosis of P. trimaculatus and Yoshida and Hirawatari 2014). Distribution: Taiwan; Japan. Remarks: Yoshida and Hirowatari (2014) redescribed this species in detail using the male genital structure.	en	Yoshida 1, Takahiro, Karner, Michael, Hirowatari, Toshiya (2018): Chtenopteryx canariensis Salcedo-Vargas & Guerrero-Kommritz 2000. Zoological Studies 57 (18): 1-18, DOI: 10.6620/ZS.2018.57-18, URL: http://dx.doi.org/10.5281/zenodo.13316922
BA4924401243FFFBFC2BF8CFFE36AAF6.taxon	description	(Fig. 1 H)	en	Yoshida 1, Takahiro, Karner, Michael, Hirowatari, Toshiya (2018): Chtenopteryx canariensis Salcedo-Vargas & Guerrero-Kommritz 2000. Zoological Studies 57 (18): 1-18, DOI: 10.6620/ZS.2018.57-18, URL: http://dx.doi.org/10.5281/zenodo.13316922
BA4924401243FFFBFC2BF8CFFE36AAF6.taxon	materials_examined	Type series: Lectotype of P. stultus syn. nov.: male, “ H. | Sauter. | Akau | Formosa | 1 - 10.12.07 ” [round label]; “ Psammoechus [sic] | stultus, Grouv. | type. ” [not Grouvelle’s hand]; “ Museum Leiden. | Psammoecus | stultus | Det: Grouv. ” [not Grouvelle’s hand]; “ type ” [blue label]; “ Type ” [red label] (RMNH; studied by MK in 2017, here designated). Paralectotypes of P. stultus syn. nov.: 1 male and 1 ex., same label data as the lectotype. (MNHN; studied by TY in 2017, here designated) Specimens examined: TAIWAN: [Taoyuan City] 1 ex., Upper Plain, Fuxing District, 600 m, N 24 ° 41 ' 12.1 " E 121 ° 23 ' 39.3 ", 11 - IV- 1988, Miller, Stange & Wang leg. (FSCA). [Taichung City] 6 exs, Wufeng, Wufeng Disctrict, 14 - IV- 1990, Smetana leg. (MHNG; studied by MK); 1 ex., same locality, 24 - IV- 1992, Smetana leg. (MHNG; studied by MK). [Penghu County] 1 ex. (Catalog No. NMNS ENT 3422 - 828), Makung (= Magong), 19 - 21 - VI- 2000, C. S. Lin & W. T. Yang leg., Sweeping Net. (NMNS); 1 ex. (Catalog No. NMNS ENT 3422 - 523), same date & locality, C. S. Lin & W. T. Yang leg., UV Light. (NMNS). [Taitung County] 2 exs, Pilam (= Taitung), II- 1908, H. Sauter leg. (HNHM; studied by MK); 1 ex., Doulan, Donghe Township, 9 - IV- 2012, J. Yamasako leg. (ELKU); 4 exs, Nanlion, Green Island Township, alt. ca. 200 m, 4 - IV- 2004, T. Kurihara leg. (ELKU); 1 ex., 4 km SE Yeh-Ying Village, Lanyu Island (= Orchid Island), Lanyu Township, 22 - V- 2007, Martin & Quicke leg. (BMNH; studied by MK); 8 exs, Yung Hsin Farm, Lanyu Island (= Orchid Island), Lanyu Township, 29 - 30 - VII- 2008, Mendel & Barclay leg. (BMNH; studied by MK). [Kaohsiung City] 2 ex., Kosempo (= Jiaxian District), I- 1908, H. Sauter leg. (HNHM, ZMHB; studied by MK); 1 ex., Liu Kui (= Liouguei), 4 - VI- 1989, K. Baba leg. (KUM); 1 ex., same locality, 30 - III- 1986, K. Baba leg. (EUMJ); 1 ex., same locality, at light, 4 - VIII- 1978, T. Niisato leg. (EUMJ); 1 ex., Minsheng Road, Liougui District, at light, 9 - VI- 1977, K. Ushijima leg. (EUMJ); 1 ex., Tsai Tie Ku, near Liu Kui (= Liouguei), 6 - X- 1986, K. Baba leg. (EUMJ); 1 ex., Kosempo (= Jiaxian District), I- 1908, H. Sauter leg. (SDEI); 4 exs, same locality, VIII- 1909, H. Sauter leg. (MNHN; studied by MK in 2017); 1 male, 2 females and 6 exs, same label data. (SDEI). [Pingtung County] 2 exs, Akau (= Pingtung City), 1 - 10 - XII- 1907, H. Sauter leg. (HNHM, ZMHB; studied by MK); 2 males and 5 exs, same locality, XII- 1907, H. Sauter leg. (SDEI); 1 ex., Haikou, N 22 ° 06 ' 10.4 " E 120 ° 42 ' 15.4 ", 24 - XI- 2001, L, Stange & H. Wang leg. (FSCA); 1 ex. (Catalog No. NMNS ENT 7063 - 354), Nanjenshan, Manzhou Township, 11 - 25 - X- 2009, M. L. Jeng & T. R. Chen leg., Flight Interception Trap. (NMNS); 1 ex. (Catalog No. NMNS ENT 6824 - 375), Manchou (= Manzhou) Lite Chulaoshushan, 11 - 25 - X- 2009, M. L. Jeng & T. R. Chen leg., Flight Interception Trap. (NMNS); 2 exs (Catalog No. 1566 - 192 & 262), Yingdar, 19 - 20 - V- 1993, C. S. Lin & W. T. Yang leg., UV Light. (NMNS); 3 exs (Catalog No. 1565 - 202, 218 & 238), Kenting, 17 - 18 - V- 1993, C. S. Lin & W. T. Yang leg., UV Light. (NMNS); 7 exs, same locality, 1 - 5 - IX- 1983, J. B. Happner leg. (FSCA); 1 ex. (Catalog No. 2695 - 5), Kenting Park, 25 - 26 - VIII- 1997, C. S. Lin & W. T. Yang leg., UV Light. (NMNS); 1 ex. (Catalog No. NMNS ENT 4388 - 539), Hengchun, Kenting Park, 7 - 8 - IV- 2004, C. S. Lin & W. T. Yang leg., UV Light Trap. (NMNS); 1 ex., Kenting Natural Reserve, 4 - 6 - X- 2000, Papp, Peregovits & Ronka leg. (HNHM; studied by MK); 1 ex. (Catalog No. NMNS ENT 6849 - 29), Sheting, Hengchun, 1 - 19 - IV- 2009, M. L. Jeng & T. R. Cheng leg., Flight Interception Trap. (NMNS); 1 ex. (Catalog No. NMNS ENT 4810 - 445), same locality, 16 - 17 - II- 2005, C. S. Lin & W. T. Yang leg., UV Light Trap. (NMNS). Diagnosis: This species can be distinguished from other Taiwanese Psammoecus species by the short oval habitus and the black elytra with yellow maculae. Distribution: Taiwan; Japan; Philippines; India; Sri Lanka; Malaysia; Indonesia; Madagascar; France (Réunion); Seychelles. Remarks: Pal (1985) and Karner (2012) provided redescriptions of this species, and Yoshida and Hirowatari (2014) provided a detailed description of male genital structure of this species. In this study, we examined morphology-including male genitalia of P. stultus type specimens - and found P. stultus should be synonymized with P. simonis. Grouvelle (1892) initially described this species as ‘ simonis’, but later Grouvelle (1908) referred to it as ‘ simoni’, and all subsequent authors have used the latter spelling (Hetschko 1930; Pal 1985; Sen Gupta and Pal 1996; Hirano 2009 2010; Karner 2012 2014; Yoshida and Hirowatari 2014 2015). According to Article 33 of the International Code of Zoological Nomenclature, if the original spelling was changed without demonstrably intentional emendation, the subsequent spelling should be regarded as incorrect and not an available name except in the case that the subsequent spelling is in prevailing usage. Otherwise, Article 23.9.1 of the code states that prevailing usage of the junior synonym must be maintained when the senior synonym has not been used as a valid name after 1899, and the junior synonym has been used for a particular taxon as its presumed valid name in at least 25 works, published by at least 10 authors in the immediately preceding 50 years and encompassing a span of not less than 10 years. In the case of this species, the subsequent spelling ‘ simoni’ changed without demonstrably intentional emendation should be regarded as an incorrect spelling, and it has not been in prevailing usage like the requirement in the Article 23.9.1 of the code. Therefore, ‘ simonis’ is deemed to be valid name of this species.	en	Yoshida 1, Takahiro, Karner, Michael, Hirowatari, Toshiya (2018): Chtenopteryx canariensis Salcedo-Vargas & Guerrero-Kommritz 2000. Zoological Studies 57 (18): 1-18, DOI: 10.6620/ZS.2018.57-18, URL: http://dx.doi.org/10.5281/zenodo.13316922
BA4924401241FFFBFF1CFC7FFB28AC16.taxon	description	(Fig. 1 I)	en	Yoshida 1, Takahiro, Karner, Michael, Hirowatari, Toshiya (2018): Chtenopteryx canariensis Salcedo-Vargas & Guerrero-Kommritz 2000. Zoological Studies 57 (18): 1-18, DOI: 10.6620/ZS.2018.57-18, URL: http://dx.doi.org/10.5281/zenodo.13316922
BA4924401241FFFBFF1CFC7FFB28AC16.taxon	materials_examined	Specimens examined: TAIWAN: [New Taipei City] 1 male, Kueishan (= Guishan) ~ Wulai, Wulai District, 5 - VI- 1970, Y. Hori leg. (EUMJ); 2 exs, Wulai, Wulai District, 4 - VI- 1970, Y. Hori leg. (EUMJ); 2 exs, Fushan, Wulai District, 10 - IX- 2001, J. & L. Stange leg. (FSCA); 3 exs, Fushan, Wulai District, alt. 450 m, 1 - 10 - XI- 2001, H. Wang leg., Malaise Trap in hardwood forest. (FSCA); 1 ex., Fushan, Wulai District, alt. 600 m, 28 - III- 2004, T. Kurihara leg. (ELKU); 1 ex., same locality, 29 - III- 2004, T. Kurihara leg. (EUMJ). [Taipei City] 2 exs, Yangmingshan, Beitou District, 24 - VIII- 1966, H. Kamiya leg. (KUM). [Yilan County] 1 ex., Fushan Botanical Garden, Yuanshan Township, 23 - 24 - VII- 2005, Galsworthy leg. (BMNH; studied by MK); 1 ex., Fushan Botanical Garden, Yuanshan Township, 25 - 27 - IX- 2000, L. Papp, L. Peregovits & L. Ronkay leg. (HNHM; studied by MK); 1 ex., Yuanshan, Yuanshan Township, 26 - III- 2009, H. - J. Chen leg. (TARI). [Nantou County] 1 ex., NW Lushan, Ren-ai Township, 8 - VIII- 2008, Barclay & Mendel leg. (BMNH; studied by MK); 1 ex. (Catalog No. NMNS ENT 5237 - 5688), Chunyang, Ren-ai Township, 12 - III- 9 - IV- 2002, C. S. Lin & W. T. Yang leg., Malaise Trap (KCN) (NMNS); 1 male (Catalog No. NMNS ENT 5237 - 3268), same locality, 9 - IV- 7 - V- 2002, C. S. Lin & W. T. Yang leg. (NMNS); 1 female & 1 ex., Penpuchi, Ren-ai Township, 13 - VII- 1966, H. Kamiya leg. (KUM); 1 ex., same locality, 14 - VII- 1966, H. Kamiya leg. (KUM). [Chiayi County] 1 male, Fenchihu, Zhuqi Township, 21 - III- 1968, T. Okadome leg. (EUMJ). [Kaohsiung City] 2 exs (Catalog No. NMNS ENT 6574 - 447 & 457), Shanping, Liougui District, 5 - 6 - XI- 2010, W. T. Yang leg., UV Light Trap (NMNS). [Pingtung County] 3 males & 2 exs, Dong Yuan Forest Recreation Area, Dong Yuan Village, Mudan Township, Pingtung City, 12 - VII- 2014, T. Yoshida leg. (ELKU).	en	Yoshida 1, Takahiro, Karner, Michael, Hirowatari, Toshiya (2018): Chtenopteryx canariensis Salcedo-Vargas & Guerrero-Kommritz 2000. Zoological Studies 57 (18): 1-18, DOI: 10.6620/ZS.2018.57-18, URL: http://dx.doi.org/10.5281/zenodo.13316922
BA4924401241FFFBFF1CFC7FFB28AC16.taxon	diagnosis	Diagnosis: This species is somewhat similar to P. taiwanensis sp. nov., and they share the following character states: the short lateral teeth on pronotum of identical size and the row of long erect setae along each lateral margin of elytra. This species can be easily distinguished from P. taiwanensis sp. nov. by the weakly curved anterior margin of pronotum and the not impressed dorsolateral pronotum.	en	Yoshida 1, Takahiro, Karner, Michael, Hirowatari, Toshiya (2018): Chtenopteryx canariensis Salcedo-Vargas & Guerrero-Kommritz 2000. Zoological Studies 57 (18): 1-18, DOI: 10.6620/ZS.2018.57-18, URL: http://dx.doi.org/10.5281/zenodo.13316922
BA4924401241FFFBFF1CFC7FFB28AC16.taxon	distribution	Distribution: Taiwan; Japan.	en	Yoshida 1, Takahiro, Karner, Michael, Hirowatari, Toshiya (2018): Chtenopteryx canariensis Salcedo-Vargas & Guerrero-Kommritz 2000. Zoological Studies 57 (18): 1-18, DOI: 10.6620/ZS.2018.57-18, URL: http://dx.doi.org/10.5281/zenodo.13316922
BA4924401241FFFBFF1CFC7FFB28AC16.taxon	discussion	Remarks: Yoshida and Hirowatari (2013) described this species in detail with the male genital structure. This species was originally described from the Nansei Islands, southeastern Japan. Yoshida and Hirowatari (2013) suggested that this species originated from the southern part of continental China, based on its distribution. The data presented here show that P. hiranoi is distributed throughout Taiwan.	en	Yoshida 1, Takahiro, Karner, Michael, Hirowatari, Toshiya (2018): Chtenopteryx canariensis Salcedo-Vargas & Guerrero-Kommritz 2000. Zoological Studies 57 (18): 1-18, DOI: 10.6620/ZS.2018.57-18, URL: http://dx.doi.org/10.5281/zenodo.13316922
