taxonID	type	description	language	source
BA2A87DCFF8FFFD1F2E4FDFEFC4EFA4F.taxon	description	(Figs. 1 A, 6 A, 16 D, 23 A, 29 B, D, 32 D, 36 E) Zoobank. org / urn: lsid: zoobank. org: act: C 3 A 41 F 65 - E 096 - 48 F 6 - BADE- 0 FFB 4 E 24 A 78 A	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF8FFFD1F2E4FDFEFC4EFA4F.taxon	materials_examined	Type Locality. Mt. Baker, Washington, USA.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF8FFFD1F2E4FDFEFC4EFA4F.taxon	description	Description. Habitus: Length 8.2 – 10.1 mm. Dark brown to reddish brown (Fig. 1 A); antennae and palpi reddish brown; elytral epiplurae sometimes lighter. Head: Broad, ratio of width across eyes to length of head about 5: 4. Interantennal groove broadly and deeply impressed. Anteocellar foveae large, deeply impressed. Eyes glabrous (Fig. 36 E). Antennomeres 5 – 10 at least 2 times longer than wide; antennomeres 4 – 11 each with many sensory pits with papilliform projections. Ocelli present. Nuchal constriction vague. Maxillary palpi as in Fig. 32 D. Mandibular molar area with L-shaped row of setae (Fig. 29 B). Labrum with sensory pores along entire surface. Labial palpi with 3 rd palpomere 1.6 – 1.8 times longer than 2 nd. Thorax: Pronotum wide (Fig. 16 D), length to width ratio = 0.61 – 0.65; ratio of pronotal width to head width = 1.72 – 1.79; maximum width subequal to elytral width at humeral angle; punctures on dorsal surface separated by average distance slightly greater than 1 puncture diameter; lateral margins broadly explanate anterad and posterad lateral foveae; lateral foveae deeply impressed. Elytra with humeral angles projecting (Fig. 29 D); epipleural carina projecting; 2.0 – 2.3 times longer than pronotum; apical margins broadly convex. Wings fully developed in most individuals, brachypterous in very few. Mesosternum with projecting tooth; longitudinal carina along midline of mesosternum complete. Legs: All tibiae with dense pubescence to apex or with small subglabrous region at apex, subglabrous apex of mesotibia less than or equal to length of basal mesotarsomere. Metatrochanter without tooth on apical margin. Abdomen: Wing-folding spicules on tergites IV and V broad, transverse, narrowly separated; tergite VI lacking wing-folding spicules. Aedeagus: Length 1.60 – 1.73 mm. Median lobe abruptly narrowed just before apex (Fig. 23 A). Parameres subparallel near base, nearly evenly convex for remainder; ending past apex of median lobe. Internal sac oblong; lightly sclerotized, microspinules denser on basal half.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF8FFFD1F2E4FDFEFC4EFA4F.taxon	materials_examined	Type Specimens. Holotype male (UAMObs: Ento: 235822) and allotype female (UAMObs: Ento: 235823) labeled as follows: WASH., Mt. Baker, Heather Meadows, 23. VII. 1979, 4000 ’, J. M. & B. A. Campbell / HOLOTYPE ³ (or ALLOTYPE ♀) Phlaeopterus bakerensis desig. L. J. Mullen and J. M. Campbell CNC No. 18468 [red label]. Both specimens are in CNC. Paratypes (184) deposited in AMNH, CAS, CNC, CSCA, FMNH, MCZ, RBCM, ROM, UCRC, and USNM.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF8FFFD1F2E4FDFEFC4EFA4F.taxon	distribution	Distribution. Phlaeopterus bakerensis is known only from Mt. Baker and Mount Baker-Snoqualmie National Forest, Washington (Fig. 6 A) at elevations of 1,220 – 1,500 m. We found one additional specimen of this species in the J. Jarrige collection, Schmitt box 359, in the MNHN, with nearly illegible label data that we tentatively interpret as follows: Austin Pass N. W. Cass. OR. [illegible, possibly niv co.] 1600 m 27. VII. 62 G. D. / Phlaeopterus m. h. [possibly a reference to Melville Hatch] / Phlaeopterus bakerensis n. sp. det. L. Mullen 2015. This specimen suggests that P. bakerensis may also occur in Oregon, but this should be regarded with suspicion unless confirmed by additional collection records. This specimen was not included in the range map of P. bakerensis due to our uncertain interpretation of the locality data. Bionomics. Adults were collected at night on the surface of snowfields during August, where they were observed feeding on arthropods (mostly Diptera) or during the day under rocks at edges of snowfields.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF8FFFD1F2E4FDFEFC4EFA4F.taxon	discussion	Remarks. Phlaeopterus bakerensis is, to our knowledge, the largest species in the subfamily Omaliinae. It can be easily distinguished from other Phlaeopterus species by its large size, broad pronotum, and projecting humeral angles of the elytra. This species has not been collected since 1979 (arctos. database. museum / saved / Phlaeopterus % 5 Fbakerensis) and so would be an ideal candidate for recollection efforts.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF8FFFD1F2E4FDFEFC4EFA4F.taxon	etymology	Etymology. This species was named for the type locality of Mt. Baker, Washington.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF8FFFCBF0F7F9C7FC18FEAE.taxon	description	(Figs. 1 B, 6 B, 16 B, 21 F, 22 A)	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF8FFFCBF0F7F9C7FC18FEAE.taxon	description	P. cavicollis, male (UAMObs: Ento: 235795), D) P. czerskyi, female. Scale (UAMObs: Ento: 235496). Scale UAMObs: Ento: 232406), C) P. frosti, male (UAMObs: Ento: 235798), D) P. fusconiger, female (bars = 1 mm. Ento: 234115), C) P. kavanaughi Scale bars = 1 mm.: Ento: 234056)., female (UAMObs: Ento: 232423), D) P. lagrandeuri, female (UAMObs Fig. 4. (UAMObs: Ento: 233445). Scale Phlaeopterus species, habitus. A) P. loganensis, female (UAMObs: Ento: 233072), B) P. 235802), C) P. obsoletus, male (UAMObs: Ento: 233776), D) P. occidentalis, female (bars = 1 mm. longipennis, male UAMObs: Ento: (UAMObs: Ento: 233301). Scale bars = 1 mm.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF8FFFCBF0F7F9C7FC18FEAE.taxon	materials_examined	Type Locality. Colorado, USA.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF8FFFCBF0F7F9C7FC18FEAE.taxon	description	Redescription. Habitus: Length 5.6 – 7.7 mm. Dark brown to reddish brown; lateral margins of pronotum often lighter (Fig. 1 B); head, elytra, palpi, and antennae sometimes lighter. Head: Moderately broad, ratio of width across eyes to length of head = 1.0 – 1.3. Interantennal groove broadly and deeply impressed. Anteocellar foveae large, deeply impressed. Eyes glabrous or with less than 10 scattered setae near ventral margin. Antennomeres 5 – 10 at least 2 times longer than wide; antennomeres 4 – 11 each with many sensory pits with papilliform projections. Ocelli present. Nuchal constriction vague. Mandibular molar area with L-shaped row of setae. Labrum with sensory pores along entire surface. Labial palpi with 3 rd palpomere 1.6 – 1.8 times longer than palpomere 2. Thorax: Pronotum broad (Fig. 16 B), length to width ratio = 0.63 – 0.73; ratio of pronotal width to head width 1.5 – 1.61; maximum width subequal to elytral width at humeral angles; punctures on dorsal surface separated by average distance equal to twice diameter of a puncture; lateral margins explanate posterad lateral foveae, narrowly explanate (P. castaneus castaneus) to not explanate (P. castaneus cascadiensis) anterad lateral foveae; lateral foveae deeply impressed. Elytra with humeral angles convex; epipleural carina not projecting; 2.2 – 2.5 times longer than pronotum; apical margins broadly convex, uniformly dark reddish brown (P. castaneus castaneus) or with sutural area lighter reddish brown than remainder of elytral surface (P. castaneus cascadiensis). Wings fully developed in most individuals, brachypterous in very few. Mesosternum with projecting tooth; longitudinal carina along midline of mesosternum complete but not strongly carinate. Legs: All tibiae with dense pubescence from base to apex. Metatrochanter without tooth on apical margin. Abdomen: Wing-folding spicules on tergites IV and V broad, transverse, narrowly separated; tergite VI lacking wing-folding patches. Aedeagus: Length 1.20 – 1.36 mm for P. castaneus cascadiensis (Fig. 21 F), 1.32 – 1.48 mm for P. castaneus castaneus (Fig. 22 A). Median lobe narrowed abruptly just before apex, with longitudinal carina at apex. Parameres narrow. Internal sac variable; 2 / 3 to half length of median lobe; covered in microspinules; with or without subapical transverse fold.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF8FFFCBF0F7F9C7FC18FEAE.taxon	materials_examined	Type Specimens. Tilea castanea: Lectotype male (here designated, UAMObs: Ento: 235783) labeled as follows: Col. / ³ / CASEY bequest 1925 / TYPE USNM 48100 / [Tilea] Castanea [sic] / LECTOTYPE ³ Phlaeopterus castanea (Casey) des. 1984, J. M. Campbell. Paralectotype male (here designated) labeled as follows: CASEY bequest 1925 / castanea PARATYPE USNM 48110. We designated these specimens as lectotype and paralectotype to clarify the application of this name to this taxon and because a holotype was not designated in the series examined by Casey (1893), which comprised two specimens. Phlaeopterus cascadiensis: Holotype male (UAMObs: Ento: 235219) and allotype female (UAMObs: Ento: 235777) labeled as follows: Mt. Rainier, WASH., Tipsoo Lake, Aug. 2, 1938, M. H. Hatch / TYPE ³ (or ALLOTYPE ♀) Phlaeopterus cascadiensis 1951 – M. H. Hatch. All four specimens are in the USNM.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF8FFFCBF0F7F9C7FC18FEAE.taxon	distribution	Distribution. Phlaeopterus castaneus castaneus is known from the Rocky Mountains of Alberta south to Colorado and west to eastern Oregon, eastern Washington, and Garibaldi Provincial Park, British Columbia (Fig. 6 B). Phlaeopterus castaneus cascadiensis is known from the Aleutian Islands and southeastern Alaska, and the Pacific Coast and Cascade Ranges of British Columbia south to Oregon (Fig. 6 B). We confirm the surprising occurrence of this species on Unalaska Island from a single specimen (UAMObs: Ento: 233345). Bionomics. This species has been collected at elevations of 500 – 2,500 m at the edges of cold, fast streams and sometimes at the edges of snowfields and lakes fed by melting snow.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF8FFFCBF0F7F9C7FC18FEAE.taxon	discussion	Remarks. Phlaeopterus castaneus can be distinguished from all other Phlaeopterus species by the shape of the pronotum, tibiae evenly pubescent to apex, mesosternum weakly but completely carinate, and the form of the aedeagus. We chose to classify P. cascadiensis as a subspecies of P. castaneus primarily because of the abundance of intermediate forms at the zone of overlap of the two taxa. The two subspecies can be distinguished throughout most of their range by the length and microspinules of the internal sac of the aedeagus, the explanate lateral margins of the pronotum, and, to some extent, the coloration of the elytra. However, in Garibaldi and Manning Provincial Parks, intermediate states of microspinules of the internal sac and the explanate lateral margins of the pronotum can be found. The two subspecies are diagnosable throughout their range except in the zone of overlap in Garibaldi and Manning Provincial Parks, British Columbia.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF95FFCFF0F5FE97FC2DF960.taxon	description	(Figs. 1 C, 7 A, 16 C, 22 B – D)	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF95FFCFF0F5FE97FC2DF960.taxon	materials_examined	Type Locality. British Columbia, Canada.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF95FFCFF0F5FE97FC2DF960.taxon	description	Redescription. Habitus: Length 6.3 – 9.2 mm. Brown to reddish brown; elytra and lateral and basal margins of pronotum often lighter; palpi and epipleura sometimes lighter (Fig. 1 C). Head: Broad, ratio of width across eyes to length of head about 5: 4. Interantennal groove broadly and deeply impressed. Anteocellar foveae large, deeply impressed. Eyes glabrous or with less than 10 scattered setae near ventral margin. Antennomeres 5 – 10 at least 2 times longer than wide; antennomeres 4 – 11 each with many sensory pits with papilliform projections. Ocelli present. Nuchal constriction vague. Mandibular molar area with L-shaped row of setae. Labrum with sensory pores along entire surface. Labial palpi with 3 rd palpomere 1.6 – 1.8 times longer than palpomere 2. Thorax: Pronotum wide (Fig. 16 C), length to width ratio = 0.61 – 0.68; ratio of pronotal width to head width = 1.49 – 1.72; maximum width subequal to elytral width at humeral angle; punctures on dorsal surface separated by average distance equal to approximately twice as great as diameter of a puncture; lateral margins broadly explanate anterad and posterad lateral foveae; lateral foveae deeply impressed. Elytra with humeral angles convex; epipleural carina not projecting; 2.2 – 2.7 times longer than pronotum; apical margins broadly convex to nearly truncate. Wings fully developed in most individuals, brachypterous in very few. Mesosternum with projecting tooth, tooth more strongly notched on posterior margin than on anterior margin; longitudinal carina along midline of mesosternum complete. Legs: All tibiae with dense pubescence to apex or with small subglabrous region at apex. Metatrochanter without tooth on apical margin. Abdomen: Wing-folding spicules on tergites IV and V broad, transverse, narrowly separated; tergite VI lacking wing-folding spicules. Aedeagus: Somewhat variable (Fig. 22 B – D), 1.28 – 1.72 mm long. Median lobe convexly converging from base to midline, gradually converging from midline to apex. Parameres diverging from base to near apex, then converging towards apex of median lobe. Internal sac elongate, rectangular to narrowest near midline, subapical transverse fold present or absent.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF95FFCFF0F5FE97FC2DF960.taxon	materials_examined	Type Specimens. Tilea cavicollis: Lectotype male (here designated, UAMObs: Ento: 235220) and paralectotype female (here designated, UAMObs: Ento: 232466) labeled as follows: Columbie britannique / Syntype / LECTOTYPE ³ (or PARALECTOTYPE ♀) Tilea cavicollis Fvl., desig. 1984 J. M. Campbell / Phlaeopterus cavicollis (Fvl.), det. 1984, J. M. Campbell. The paralectotype female is missing Fauvel’ s handwritten determination label but otherwise has identical label data. We designated these specimens as lectotype and paralectotype to clarify the application of this name to this taxon and because a holotype was not designated in the series examined by Fauvel (1878), which comprised an unknown number of specimens. Both specimens are in the RBINS. Phloeopterus [sic] longipalpus: Lectotype male (here designated to clarify the application of this name to this taxon, UAMObs: Ento: 235764) labeled as follows: Oct. 3 / Placer Co. CAL. / ³ / T. longipalpus Cas. / CASEY bequest 1925 / TYPE USNM 48106 / Lectotype ³ Phlaeopterus longipalpus Casey des. 1984, J. M. Campbell. We designated this specimen as a lectotype because a holotype was not designated in the series examined by Casey (1885). The Casey Collection contains seven other specimens collected from “ Placer Co., Cal. ” with a label stating “ # 2 and # 7 may be paratypes ”. However, Casey (1885) examined at least one female, and these two specimens are males, so we could not determine which two of the seven specimens should be paralectotypes. The lectotype is in the USNM. Tilea brevipennis: Holotype male (UAMObs: Ento: 235786) labeled as follows: Wy / ³ / CASEY bequest 1925 / TYPE USNM 48109 / brevipennis / HOLOTYPE ³ Phlaeopterus brevipennis Casey. This specimen is in the USNM.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF95FFCFF0F5FE97FC2DF960.taxon	distribution	Distribution. Phlaeopterus cavicollis is widely distributed in western North America from southern California north through the Pacific Coast and Cascade Ranges and the Rocky Mountains of British Columbia to southeastern Alaska, and east to Wyoming and Colorado (Fig. 7 A). Adults have been collected during June – October, and in November in California, at 600 – 3,500 m elevation. Bionomics. Adults have been observed feeding on wind-blown insects (mostly Diptera) on the surface of snowfields, breeding on the surface of snowfields, and rarely flying. Adults have also been collected under rocks at the edges of snowfields and in moss or under rocks at the edges of cold, fast streams or springs at lower elevations.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF95FFCFF0F5FE97FC2DF960.taxon	discussion	Remarks. Phlaeopterus cavicollis can be distinguished from all other Phlaeopterus species except for P. bakerensis by having lateral margins of the pronotum broadly and subequally explanate anterad and posterad the lateral foveae. Phlaeopterus cavicollis can be distinguished from P. bakerensis by its smaller size and lacking projecting humeral angles of the elytra. We chose to synonymize P. brevipennis and P. longipalpus under P. cavicollis due to their shared characters (see Redescription) and the presence of intermediate forms. The morphological variation that has previously been used to differentiate these three species is as follows. Specimens of P. longipalpus from the Sierra Nevada Mountains, the Trinity Alps, and Mt. Lassen, California have the tibiae evenly pubescent to the apex and the internal sac of the aedeagus roughly rectangular with a subapical transverse fold (Fig. 22 D). Specimens of P. cavicollis from Alaska, British Columbia, and the Pacific Coast and Cascade Ranges south to northern California usually also have the tibiae evenly pubescent to the apex and the internal sac of the aedeagus roughly triangular but lack a subapical transverse fold (Fig. 22 B). Most specimens (P. brevipennis) from the Rocky Mountains, Washington, Montana, and Colorado have the apex of the tibiae narrowly glabrous (but a few have pubescence evenly distributed to the apex) and the internal sac of the aedeagus without a subapical transverse fold (Fig. 22 C) and with margins narrowed near the midline. Some specimens intermediate between P. cavicollis and P. brevipennis from eastern Oregon, eastern Washington, and Idaho cannot be assigned to either name with certainty. It is difficult to be certain whether these populations should be regarded as a single, broadly distributed, variable species or three distinct species. Given the available morphological evidence, we chose to regard them as a single species. Genetic analysis would be useful in testing these synonymizations, but at present no sequence data are available for this species.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF90FFCEF2F5FF1CFCA1FCCD.taxon	materials_examined	Type Locality. Eastern Siberia, Russia.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF90FFCEF2F5FF1CFCA1FCCD.taxon	description	Redescription. Habitus: Length 4.0 – 4.3 mm. Dark brown to reddish brown; abdomen darkest (Fig. 1 D); antennae, mouthparts, and legs often yellowish brown. Head: Moderately wide, width across eyes greater than head length. Interantennal groove vaguely impressed. Anteocellar foveae moderately impressed. Eyes moderately pubescent on ventral half, with more than 10 setae. Antennomeres 5 – 10 1.2 – 1.8 times longer than wide. Ocelli present. Nuchal constriction vague. Mandibles with row of setae (Fig. 30 B). Labrum with sensory pores along entire surface (Fig. 30 A). Mentum as in Fig. 30 D. Maxilla as in Fig. 30 E. Labial palpi with 3 rd palpomere greater than 2 times longer than 2 nd palpomere (Fig. 30 C). Thorax: Pronotum moderately narrow, length to width ratio = 0.80; ratio of pronotal width to head width = 1.5; maximum width slightly less than width of elytra at bases; dorsal surface very coarsely, sparsely punctate, punctures separated by average distance 1 – 2 times diameter of a puncture; lateral margins narrowly explanate posterad lateral foveae, not explanate anterad lateral foveae; lateral foveae vaguely impressed, contiguous with lateral margins. Elytra with humeral angles convex; epipleural carina not projecting; about 1.7 times longer than pronotum; apical margins convex. Wings brachypterous, reduced to short lobes. Mesosternum with projecting tooth; longitudinal carina along midline of mesosternum distinct, strongly and roughly carinate anteriorly, becoming vague posteriorly. Legs: All tibiae evenly pubescent to apex. Metatrochanter without tooth on apical margin. Abdomen: Wingfolding spicules absent from all tergites. Apical palisade fringe on tergite VII absent. Aedeagus: Median lobe somewhat triangular, apex acute (Fig. 26). Parameres diverging from base to apical third, then slightly converging to apex. Internal sac heavily sclerotized, covered in microspinules.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF90FFCEF2F5FF1CFCA1FCCD.taxon	materials_examined	Type Specimens. Holotype male labeled as follows: 14. VI. [19] 96 / XaMap-flaðau [Khamar- Daban Mts.] / KoMapnucĸnň [Komarinskiy] / xp. [Mt.] h = 1900 / fflasspnu A. [Shavrin A. leg.] Holotype / Lesteva / czerskyi / Shavrin A. 2000. Paratype female labeled as follows: XaMap-flaðau [Khamar- Daban Mts.] / cp. T. Baðxn [middle flow of Babkha River] [51 ° 46 ʹ N, 103 ° 95 ʹ E] / 8 – 14. V. 1999 / fflasspnu A. / [Shavrin A. leg.] / Paratype / Lesteva / czerskyi / Shavrin A. 2000. The holotype is deposited in the ZIN. The paratype is deposited in the CSH.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF90FFCEF2F5FF1CFCA1FCCD.taxon	distribution	Distribution. Phlaeopterus czerskyi is known only from the Khamar-Daban Mountains of eastern Siberia (Fig. 7 B). Bionomics. Adults have been collected at 900 – 1,900 m elevation during May – August in wet moss at the edges of small streams.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF90FFCEF2F5FF1CFCA1FCCD.taxon	discussion	Remarks. Phlaeopterus czerskyi is the only species of the genus known outside of northwestern North America. The occurrence of P. czerskyi in eastern Siberia confirms Casey’ s (1893) speculation: “ [Phlaeopterus] is the most conspicuous of the endemic North American genera of Omaliini, although probably occurring also in Siberia … ” It is easily distinguished from all other Phlaeopterus species by the crenulate lateral margins of the pronotum, the especially coarse and sparse punctation of the pronotum and elytra, the lack of wing-folding spicules, and the brachypterous wings. It most closely resembles P. lagrandeuri and can be distinguished from P. lagrandeuri by characters discussed in the remarks section of that species.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF90FFC2F0E2FC6FFC7EFB88.taxon	description	(Figs. 2 A, 8 A, 17 E, H, 20 F, 25 D, 29 C, 37 J – K, 39 E – F, 41 C, 42 F) Zoobank. org / urn: lsid: zoobank. org: act: 6 D 8522 FE-F 60 F- 4086 - 98 BA-CEE 3411 A 5 F 18	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF90FFC2F0E2FC6FFC7EFB88.taxon	materials_examined	Type Locality. Yoho National Park, British Columbia, Canada.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF90FFC2F0E2FC6FFC7EFB88.taxon	description	Description. Habitus: Length 4.8 – 6.4 mm. Dark brown to black; tarsi, apex of tibiae, and antennae often lighter, reddish brown to dark brown (Fig. 2 A). Head: Moderately narrow, ratio of width across eyes subequal to length of head (Fig. 59). Interantennal groove shallowly impressed. Anteocellar foveae large, deeply impressed. Eyes glabrous or with less than 10 scattered setae near ventral margin. Antennomeres 5 – 10 at least 2 times longer than wide; antennomeres 4 – 11 each with many sensory pits with papilliform projections (Fig. 37 J – K). Ocelli present. Nuchal constriction vague. Mandibular molar area with L-shaped row of setae (Fig. 29 C). Maxilla as in Fig. 39 E – F. Hypopharynx as in Fig. 41 C. Labrum with sensory pores along entire surface. Labial palpi with 3 rd palpomere 1.6 – 1.8 times longer than palpomere 2. Gula as in Fig. 42 F. Thorax: Pronotum narrow (Fig. 17 E), length to width ratio = 0.88 – 0.95; ratio of pronotal width to head width = 1.27 – 1.41; maximum width less than elytral width at base; bakerensis, E) P. smetanai, F) P. occidentalis, G) P. olympicus, H) P. loganensis. punctures on dorsal surface separated by average distance slightly greater than diameter of a puncture; without vague impression near midline on ventral surface (as in P. filicornis); lateral margins deflexed posterad and anterad lateral foveae; lateral foveae vague, contiguous with and somewhat obscured by deflexed lateral margin. Elytra with humeral angles convex; epipleural carina not projecting; 2.4 – 2.7 times longer than pronotum; apical margins sexually dimorphic, broadly convex in males but somewhat prolonged at suture in females. Wings fully developed in most individuals, brachypterous in very few. Mesosternum with projecting tooth; longitudinal carina along midline of mesosternum complete but vague (Fig. 20 F). Legs: Apices of all tibiae glabrous, length of subglabrous apex of metatibia as ratio to metatibial length = 6.1 – 11.5. Metatrochanter without tooth on apical margin. Abdomen: Wing-folding spicules on tergites IV and V broad, transverse, nearly contiguous; tergite VI with small, nearly circular wing-folding patches (Fig. 17 H). Aedeagus: Length 0.90 – 1.05 mm. Median lobe triangular, apex somewhat acute (Fig. 25 D). Parameres nearly straight. Internal sac expanded at base; moderately sclerotized; evenly covered in microspinules.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF90FFC2F0E2FC6FFC7EFB88.taxon	materials_examined	Type Specimens. Holotype male (UAMObs: Ento: 235834) and allotype female (UAMObs: Ento: 235835) labeled as follows: Alta., 20 mi. SW Kananaskis F. E. S. [Forest Experiment Station], 7000 ’, Snow Ridge, VII- 31 - 1971, J. M. & B. A. Campbell / Holotype ³ (or Allotype ♀) Phlaeopterus elongatus, desig. L. J. Mullen and J. M. Campbell, CNC No. 18462 (red label). Both specimens are deposited in the CNC. Paratypes (n = 122) deposited in the AMNH, CAS, CNC, CSCA, FMNH, MCZ, and USNM.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF90FFC2F0E2FC6FFC7EFB88.taxon	distribution	Distribution. Phlaeopterus elongatus is known from the Rocky Mountains from Wyoming north to Summit Lake, British Columbia, in the Cascade Range and Selkirk Mountains of British Columbia, and the Alaska Range and Talkeetna Mountains of Alaska (Fig. 8 A). Bionomics. Adults have been collected at 1,100 – 3,200 m elevation during June – September from particularly cold habitats, including the edges of snowfields, lakes fed by melting snow, and sometimes from wet moss in the splash zones of waterfalls or the edges of streams. Unlike other new species described herein that have not been re-collected since the late 1970 s or early 1980 s, we collected P. elongatus extensively in the Alaska Range and Talkeetna Mountains during the summers of 2013 – 2014, suggesting these populations are persisting in Alaska.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF90FFC2F0E2FC6FFC7EFB88.taxon	discussion	Remarks. Phlaeopterus elongatus can be distinguished from all other Phlaeopterus species by its elongate form, especially narrow pronotum, and the presence of wing-folding spicules on tergite VI. This species has been misidentified as P. filicornis in most collections but can be differentiated from P. filicornis by the aforementioned characters as well as its smaller size, darker color, shallower interantennal groove, longer glabrous region of the mesotibia, and aedeagus with a narrower median lobe.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF90FFC2F0E2FC6FFC7EFB88.taxon	etymology	Etymology. The name of this species is derived from the Latin elongate, referring to the particularly elongate shape of the body.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF9CFFC4F0D6FB8BFCAEFA1A.taxon	description	(Figs. 2 B, 8 B, 17 C, 25 C, 29 G, 31 H)	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF9CFFC4F0D6FB8BFCAEFA1A.taxon	materials_examined	Type Locality. Placer County, California, USA.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF9CFFC4F0D6FB8BFCAEFA1A.taxon	description	Redescription. Habitus: Length 5.9 – 7.7 mm. Dark brown, almost black, palpi, elytra, and glabrous apices of tibiae lighter (Fig. 2 B). Head: Moderately narrow, ratio of width across eyes subequal to head length. Interantennal groove deeply impressed. Anteocellar foveae large, deeply impressed. Eyes glabrous or with less than 10 scattered setae near ventral margin. Antennomeres 5 – 10 at least 2 times longer than wide; antennomeres 4 – 11 each with many sensory pits with papilliform projections. Ocelli present. Nuchal constriction vague. Mandibular molar area with L-shaped row of setae. Labrum with sensory pores along entire surface (Fig. 31 H). Labial palpi with 3 rd palpomere 1.6 – 1.8 times longer than palpomere 2. Thorax: Pronotum moderately narrow (Fig. 17 C), length to width ratio = 0.76 – 0.81; ratio of pronotal width to head width = 1.35 – 1.43; maximum width less than width of elytra at bases; punctures on dorsal surface separated by average distance equal to greater than diameter of a puncture; with vague impression near midline on ventral surface; lateral margins deflexed posterad and anterad lateral foveae; lateral foveae moderately impressed. Elytra with humeral angles convex; epipleural carina not projecting; 2.4 – 3.2 times longer than pronotum; apical margins sexually dimorphic, elongated at suture and slightly diverging in females (Fig. 29 G) and broadly convex and not diverging in males. Wings fully developed in most individuals, brachypterous in very few. Mesosternum with projecting tooth; longitudinal carina along midline of mesosternum complete. Legs: All tibiae glabrous, length of subglabrous apex of metatibia as ratio to metatibial length = 4.9 – 8.8. Metatrochanter without tooth on apical margin. Abdomen: Wing-folding spicules on tergites IV and V broad, transverse, nearly contiguous; tergite VI lacking wing-folding patches. Aedeagus: Length 1.25 – 1.38 mm. Median lobe somewhat triangular, apex acutely triangular (Fig. 25 C). Parameres diverging from lagrandeuri, B) P. houkae, C cascadiensis.) P. longipennis, D) P. obsoletus, E) P. kavanaughi, F) P. castaneus base until just before apex. Internal sac elongate, lightly sclerotized, evenly covered in small microspinules.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF9CFFC4F0D6FB8BFCAEFA1A.taxon	materials_examined	Type Specimens. Phlaeopterus filicornis. Lectotype male (here designated to clarify the application of this name to this taxon, UAMObs: Ento: 235784) labeled as follows: Sept. / Placer Co. Cal. / ³ / CASEY bequest 1925 / TYPE USNM 48108 / filicornis Casey / LECTOTYPE ³ Tilea filicornis Csy. des. 1982, J. M. Campbell. Paralectotype male (UAMObs: Ento: 235785) labeled as follows: Sept. / Placer Co. Cal. / CASEY bequest 1925 / CASEY det. 2 filicornis / PARALECTOTYPE ³ Tilea filicornis Csy., des. 1982, J. M. Campbell. The Casey Collection in the USNM contains these two males of the species. Tilea rufitarsis. Lectotype male (here designated to clarify the application of this name to this taxon) labeled as follows: Siskiyou Co. Cal. / ³ / CASEY bequest 1925 / TYPE USNM 48107 / rufitarsis / LECTOTYPE ³ Tilea rufitarsis Csy. des. 1982, J. M. Campbell. Paralectotype female labeled as follows: Siskiyou Co. Cal. / ♀ / CASEY bequest 1925 / rufitarsis 2 PARATYPE USNM 48107 / PARALECTOTYPE ♀ Tilea rufitarsis Csy. des. 1982, J. M. Campbell. Paralectotype female labeled as follows: Siskiyou Co. Cal. / ♀ / CASEY bequest 1925 / rufitarsis 2 / PARATYPE USNM 48107 / PARALECTOTYPE ♀ Tilea rufitarsis Csy. des. 1982, J. M. Campbell. The Casey Collection contains these three specimens.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF9CFFC4F0D6FB8BFCAEFA1A.taxon	distribution	Distribution. Phlaeopterus filicornis is known only from California in the Sierra Nevada and Cascade Range (Fig. 8 B). Bionomics. Adults have been collected at 1,820 – 3,440 m elevation from July – September under rocks near cold, fast-flowing streams or at the edges of permanent or long-lasting snowfields, and in moss at the edges of streams or in the splash zones of waterfalls.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF9CFFC4F0D6FB8BFCAEFA1A.taxon	discussion	Remarks. Phlaeopterus filicornis can be distinguished from all other Phlaeopterus species by the shape of the pronotum (length: width ratio, lateral margins, and lateral foveae), sexually dimorphic shape of the elytral apices, glabrous apices of the tibiae, and form of the aedeagus. It is highly similar to P. hatchi, from which it differs by the sexually dimorphic shape of the elytral apices, pronotum with a vague impression near the midline on the ventral surface, shape and size of the median lobe of the aedeagus, microspinules of the internal sac of the aedeagus, and a slight difference in body length. We examined the lectotypes of P. filicornis and P. rufitarsis and were not able to find any significant differences between the two specimens. Therefore, we chose to synonymize P. rufitarsis under P. filicornis.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF9AFFFBF128FA2BFC4EFA70.taxon	description	(Figs. 2 C, 9 A, 17 B, 18 A, 20 D, 25 A, 36 D)	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF9AFFFBF128FA2BFC4EFA70.taxon	materials_examined	Type Locality. Mt. Rainier, Washington, USA.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF9AFFFBF128FA2BFC4EFA70.taxon	description	Redescription. Habitus: Length 5.8 – 7.2 mm. Reddish brown to black; legs, elytral epipleura, tarsi, and apices of tibiae lighter reddish brown to yellowish brown (Fig. 2 C). Head: Slightly broader than long, ratio of width across eyes to head length = 4: 3. Interantennal groove broadly and deeply impressed. Anteocellar foveae large, deeply impressed. Eyes glabrous or with less than 10 scattered hairs near ventral margin (Fig. 36 D). Antennomeres 5 – 10 at least 2 times longer than wide; antennomeres 4 – 11 each with many sensory pits with papilliform projections. Ocelli present. Nuchal constriction vague. Mandibular molar area with L-shaped row of setae. Labrum with sensory pores along entire surface. Labial palpi with 3 rd palpomere 1.6 – 1.8 times longer than palpomere 2. Thorax: Pronotum moderately wide (Fig. 17 B), length to width ratio = 0.68 – 0.72; ratio of pronotal width to head width = 1.54 – 1.67; maximum width subequal to elytral width at humeral angles; punctures separated by distance equal to slightly greater than diameter of a puncture; lateral margins explanate posterad but not anterad lateral foveae; lateral foveae deeply impressed. Elytra with humeral angles convex; epipleural carina not projecting; 2.1 – 2.2 times longer than pronotum; apical margins convex or subtruncate. Wings fully developed in most indivuals, brachypterous in very few. Mesosternum with projecting tooth; longitudinal carina along midline of mesosternum distinct on anterior half, distinct or reduced on posterior half (Fig. 20 D). Legs: Apices of all tibiae glabrous, subglabrous apex of mesotibia subequal to length of basal 4 mesotarsomeres (Fig. 18 A), length of subglabrous apical region of metatibia as ratio to metatibial length in males = 4.9 – 8.4, in females = 4.2 – 5.5. Metatrochanter without tooth on apical margin. Abdomen: Wing-folding spicules on tergites IV and V; shape of wing-folding patches on tergite V broadly oval and narrowly separate. Aedeagus: Length 1.23 – 1.40 mm. Median lobe somewhat triangular with sides narrowing from base to apex; apex not carinate (Fig. 25 A). Parameres extending slightly beyond apex of median lobe. Internal sac rectangular and elongate; heavily sclerotized, uniformly covered with microspinules; lacking subapical transverse fold.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF9AFFFBF128FA2BFC4EFA70.taxon	materials_examined	Type Specimens. Holotype male (UAMObs: Ento: 235778) and allotype female (UAMObs: Ento: 235779) labeled as follows: Mt. Rainier, WASH., below Sluskin Falls, Aug. 23, 1930 M. H. Hatch TYPE ³ (or ALLOTYPE ♀), Phlaeopterus frosti, 1951 – M. H. Hatch. Both specimens are in the USNM.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF9AFFFBF128FA2BFC4EFA70.taxon	distribution	Distribution. Phlaeopterus frosti is found in the Cascade Range from Oregon north to Manning Provincial Park, British Columbia, the Olympic Mountains of Washington, the Pacific Coast Range, Selkirk Mountains, and southern Rocky Mountains of British Columbia, and the Kenai Mountains of Alaska (Fig. 9 A). Bionomics. Adults have been collected at 50 – 2,050 m elevation during June – August. This species has been collected on the surface of permanent or long-lasting snowfields at night in Washington and Oregon, where they forage immediately after dusk, and during the day in British Columbia. They have also been collected under rocks at the edges of snowfields and cold, fast-flowing streams, in moss growing in the splash-zones of waterfalls, and occasionally in the gravelly, marshy edges of high elevation lakes. On the surface of snowfields, they have been observed feeding on small insects (mostly Diptera) stranded on the surface of the snow.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFF9AFFFBF128FA2BFC4EFA70.taxon	discussion	Remarks. Phlaeopterus frosti can be distinguished from all other Phlaeopterus species by the shape of the pronotum (length: width ratio, lateral margins, and lateral foveae), the glabrous region at the apex of the tibiae, and the form of the aedeagus. This species is very similar to P. fusconiger but can be distinguished by the characters discussed in the Remarks section of the latter species. These two species are sympatric in the Pacific Coast Range from the Olympic Mountains of Washington north to the Kenai Mountains of Alaska.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA5FFFAF099F9C7FC8AFA15.taxon	description	(Figs. 2 D, 9 B, 17 A, 18 B, 24 D)	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA5FFFAF099F9C7FC8AFA15.taxon	materials_examined	Type Locality. Unalaska Island, Alaska, USA.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA5FFFAF099F9C7FC8AFA15.taxon	description	Redescription. Habitus: Length 5.8 – 7.2 mm. Dark brown to reddish brown; legs, elytral epipleura, and tarsi often lighter; apices of tibiae, tarsi, and palpi even lighter reddish brown; elytra usually slightly lighter than pronotum (Fig. 2 D). Head: Slightly broader than long, ratio of width across eyes to head length = 4: 3. Interantennal groove broadly and deeply impressed. Anteocellar foveae large, deeply impressed. Eyes glabrous or with less than 10 scattered setae near ventral margin. Antennomeres 5 – 10 at least 2 times longer than wide; antennomeres 4 – 11 each with many sensory pits with papilliform projections. Ocelli present. Nuchal constriction vague. Mandibular molar area with L-shaped row of setae. Labrum with sensory pores along entire surface. Labial palpi with 3 rd palpomere 1.6 – 1.8 times longer than palpomere 2. Thorax: Pronotum moderately wide (Fig. 17 A), length to width ratio = 0.68 – 0.72; ratio of pronotal width to head width = 1.54 – 1.67; maximum width subequal to elytral width at humeral angles; punctures separated by distance equal to slightly greater than diameter of a puncture; lateral margins explanate posterad but not anterad lateral foveae; lateral foveae deeply impressed. Elytra with humeral angles convex; epipleural carina not projecting; 2.1 – 2.2 times longer than pronotum; apical margins convex or subtruncate. Wings fully developed in most individuals, brachypterous in very few. Mesosternum with projecting tooth, tooth more strongly notched on posterior margin than on anterior margin; longitudinal carina along midline of mesosternum distinct on anterior half, reduced on posterior half. Legs: Apices of all tibiae glabrous, subglabrous apex of mesotibia subequal to length of basal 3 mesotarsomeres (Fig. 18 B), length of glabrous apical region of metatibia as ratio to metatibial length in males = 7.4 – 12.2, in females = 6.6 – 8.6. Metatrochanter without tooth on apical margin. Abdomen: Wing-folding spicules on tergites IV and V; shape of wing-folding patches on tergite V broadly oval and narrowly separate. Aedeagus: Length 1.23 – 1.43 mm long. Median lobe with margins concavely constricted near midline (Fig. 24 D). Parameres convex, converging past apex of median lobe. Internal sac rectangular; lightly sclerotized, uniformly covered with microspinules; lacking subapical transverse fold.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA5FFFAF099F9C7FC8AFA15.taxon	materials_examined	Type Specimen. Holotype female (UAMObs: Ento: 235794) labeled as follows: Unalaschka, Phlaeopterus fusconiger Motsch., Unalaschka [Unalaska Island] / Phlaeopterus fusconiger Mots., HOLOTYPE, exam. 1984 J. M. Campbell. As of 1984, the holotype was confirmed by JMC to be located in the collection of the ZMM. The specimen is glued to a transparent plate and badly damaged with both elytra and apical five antennomeres of the right antenna, right middle and hind legs, and apical four metatarsomeres missing. In 2015, Alexey V. Shavrin (A. Shavrin, personal communication) visited the collection and could not locate the holotype. Although the holotype is possibly missing, this species is currently in no danger of being confused with similar species, thus a neotype is not needed.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA5FFFAF099F9C7FC8AFA15.taxon	distribution	Distribution. Phlaeopterus fusconiger is widely distributed along the northwest coast of North America, from the Olympic Mountains of Washington north through the Pacific Coast Range, on Vancouver Island and the Haida Gwaii, to the Aleutian Islands and Nome, Alaska (Fig. 9 B). Bionomics. Adults have been collected at near sea level in Alaska up to 2,000 m in mainland Washington and British Columbia during June – August. The Nome record (UAMObs: Ento: 234463) is the northernmost record for any Phlaeopterus species.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA5FFFAF099F9C7FC8AFA15.taxon	discussion	Remarks. Phlaeopterus fusconiger can be distinguished from all other Phlaeopterus species by the shape of the pronotum, the glabrous region at the apex of the tibiae, and the aedeagus. This species is very similar to P. frosti but can be distinguished by the shorter glabrous region of the mesotibia, slightly lighter color, more strongly developed anterior margins of the mesosternal notch, and the aedeagus with the internal sac shorter, broader, less heavily sclerotized, and with the apex less acutely narrowed. These two species are sympatric in the coastal range from the Olympic Mountains of Washington north to the Kenai Mountains of Alaska. We have confirmed the surprising occurrence of P. fusconiger in Nome, Alaska by examination of a single specimen collected in Nome by George Ball in 1958 (UAMObs: Ento: 234463). However, we are aware of no additional specimens of Phlaeopterus from Nome, or anywhere on the Seward Peninsula. This remote population would be interesting to confirm and compare genetically to other populations of this species.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA4FFF9F0EFFA17FCD4FD22.taxon	description	(Figs. 3 A, 10 A, 17 D, 25 B, 29 F, 35 C – D) Zoobank. org / urn: lsid: zoobank. org: act: FF 1 FC 2 B 7 - 035 B- 4479 - 88 D 4 - 6 E 3 B 9 C 93 B 407 Hatch 1957: 59 [misidentification as P. rufitarsis].	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA4FFF9F0EFFA17FCD4FD22.taxon	materials_examined	Type Locality. Mt. Begbie, British Columbia, Canada.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA4FFF9F0EFFA17FCD4FD22.taxon	description	Description. Habitus: Length 5.5 – 6.9 mm. Dark brown, almost black; palpi, elytra, and glabrous apices of tibiae lighter (Fig. 3 A). Head: Moderately narrow, ratio of width across eyes subequal to head length. Interantennal groove shallowly impressed (Fig. 35 C – D). Anteocellar foveae large, deeply impressed. Eyes glabrous or with less than 10 scattered setae near ventral margin. Antennomeres 5 – 10 at least 2 times longer than wide; antennomeres 4 – 11 each with many sensory pits with papilliform projections. Ocelli present. Nuchal constriction vague. Mandibular molar area with Lshaped row of setae. Labrum with sensory pores along entire surface. Labial palpi with 3 rd palpomere 1.6 – 1.8 times longer than palpomere 2. Thorax: Pronotum moderately narrow (Fig. 17 D), length to width ratio = 0.78 – 0.84; ratio of pronotal width to head width = 1.30 – 1.39; maximum width less than width of elytra at bases; punctures on dorsal surface separated by average distance equal to greater than diameter of single puncture; without vague impression near midline on ventral surface (as in P. filicornis); lateral margins deflexed posterad and anterad lateral foveae; lateral foveae moderately impressed. Elytra with humeral angles convex; epipleural carina not projecting; 2.4 – 3.0 times longer than pronotum; apical margins not sexually dimorphic, broadly convex and not diverging in both sexes (Fig. 29 F). Wings fully developed in most individuals, brachypterous in very few. Mesosternum with projecting tooth; longitudinal carina along midline of mesosternum complete. Legs: Apices of all tibiae glabrous, length of subglabrous apex of metatibia as ratio to metatibial length = 4.4 – 7.8. Metatrochanter without tooth on apical margin. Abdomen: Wing-folding spicules on tergites IV and V broad, transverse, nearly contiguous; tergite VI lacking wing-folding patches. Aedeagus: Length 1.28 – 1.40 mm. Median lobe short, with lateral margins subparallel then acutely triangular nearing apex (Fig. 25 B). Internal sac elongate, nearly as long as median lobe, moderately heavily sclerotized, covered with microspinules but with distinctly larger microspinules in patches near apex.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA4FFF9F0EFFA17FCD4FD22.taxon	materials_examined	Type Specimens. Holotype male (UAMObs: Ento: 235832) and allotype female (UAMObs: Ento: 235833) labeled as follows: B. C., Mt. Begbie, 6300 ’, VIII – 30 – 1971, J. M. Campbell / HOLOTYPE ³ (or ALLOTYPE ♀) Phlaeopterus hatchi, desig. L. J. Mullen and J. M. Campbell, CNC No. 18463 (red label). Both specimens are deposited in the CNC. Paratypes (n = 73) are deposited in the AMNH, CAS, CNC, CSCA, FMNH, MCZ, and USNM.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA4FFF9F0EFFA17FCD4FD22.taxon	distribution	Distribution. Phlaeopterus hatchi is known from northern California north through the Cascade Range, the Mission Mountains of Montana, the Rocky Mountains of British Columbia, and in Alaska by a single specimen (UAMObs: Ento: 232742) collected by A. Ord in 2012 between Haines and Juneau (Fig. 10 A). Bionomics. Adults have been collected at 1,100 – 2,130 m elevation during July – August. Adults were mostly collected from under stones at the edges of small, cold streams, but sometimes were taken under rocks near snowfields or in splash zones of waterfalls and moss along the edges of streams.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA4FFF9F0EFFA17FCD4FD22.taxon	discussion	Remarks. Phlaeopterus hatchi can be distinguished from all other Phlaeopterus species by the shape of the pronotum, the glabrous apices of the tibiae, and the aedeagus. It is similar to P. filicornis, from which it differs by characters discussed in the Remarks section of that species.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA4FFF9F0EFFA17FCD4FD22.taxon	etymology	Etymology. This species is named in honor of Dr. Melville H. Hatch.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA7FFF8F0C2FD74FEA0FAEF.taxon	description	(Figs. 3 B, 10 B, 15 F, 17 F, 19 F, 21 B, 28 F, 31 F, 33 F, 34 C – D, 36 B, 37 F – G, 39 A – B, 40 F, 42 D)	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA7FFF8F0C2FD74FEA0FAEF.taxon	materials_examined	Type Locality. Mt. Baker, Washington, USA.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA7FFF8F0C2FD74FEA0FAEF.taxon	description	Redescription. Habitus: Length 3.1 – 4.0 mm. Dark brown; elytra usually lighter; legs, antennae, and mouthparts yellowish brown; abdomen usually reddish brown (Fig. 3 B). Head: Width moderate, width across eyes to head length slightly wider than long (Fig. 34 C – D). Interantennal groove vague or absent at least across midline of head. Anteocellar foveae vague, shallowly impressed. Eyes moderately pubescent on ventral half, with more than 10 setae (Fig. 36 B). Antennomeres 3 – 10 each 1.6 – 2.0 times longer than wide; antennomeres 4 – 11 each with many sensory pits with papilliform projections (Fig. 37 F – G). Ocelli present. Nuchal constriction absent. Epipharynx as in Fig. 33 F. Hypopharynx as in Fig. 40 F. Mandibular molar area with L-shaped row of setae (Fig. 28 F). Labrum with sensory pores along anterior margin only (Fig. 31 F). Maxilla with lacinia and galea as in Fig. 39 A – B. Labial palpi with 3 rd palpomere 1.2 – 1.5 times longer than 2 nd palpomere. Gula as in Fig. 42 D. Thorax: Pronotum narrow (Fig. 15 F), length to width ratio = 0.7 – 0.8; ratio of pronotal width to head width = 1.3 – 1.4; maximum width subequal to width of elytra at bases; punctures separated by average distance equal to slightly less than diameter of a puncture; lateral margins narrowly explanate posterad lateral fovea, not explanate anterad lateral foveae; lateral foveae moderately impressed, contiguous with lateral margins. Elytra with humeral angles convex; epipleural carina not projecting; 1.9 – 2.1 times longer than pronotum; apical margins convex. Wings brachypterous. Mesosternum with projecting tooth; longitudinal carina along midline of mesosternum present but vague, especially near midline and posterior margin (Fig. 19 F). Legs: All tibiae evenly pubescent to apex. Metatrochanter without tooth on apical margin. Abdomen: Tergites IVand Veach with wing-folding spicules very broad, nearly contiguous or contiguous in shape of single transverse band (Fig. 17 F). Apical palisade fringe on tergite VII absent. Aedeagus: Length 0.65 – 0.75 mm. Median lobe with lateral margins subparallel from base to past midline, then abruptly narrowed, apex acute (Fig. 21 B). Parameres broad, parallel and nearly straight. Internal sac with 2 dense patches of spicules lateroapically.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA7FFF8F0C2FD74FEA0FAEF.taxon	materials_examined	Type Specimen. Holotype female (UAMObs: Ento: 235216) labeled as follows: Mt. Baker Lodge, Aug. 26, 1935, WASH., M. H. Hatch / Det. 1954, H. Houk / not Vellica longipennis Csy., compared with type, M. H. Hatch 1958 / TYPE ♀ Phlaeopterus houkae 1951 – M. H. Hatch / Vellica longipennis Csy., compared with Fall coll., M. H. Hatch 1952. The specimen is deposited in the USNM.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA7FFF8F0C2FD74FEA0FAEF.taxon	distribution	Distribution. Phlaeopterus houkae is known broadly from the Pacific Coast and Cascade Ranges, from Siskiyou County, California and Oregon north to the Alaska Range in Alaska (Fig. 10 B). It is also known coastally from Baranof Island, Prince of Wales Island, Haida Gwaii, Vancouver Island, and the Olympic Mountains. Bionomics. Adults have been collected at 90 – 2,400 m elevation during June – August in wet moss at the edge of cold water, usually small streams and pools.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA7FFF8F0C2FD74FEA0FAEF.taxon	discussion	Remarks. Phlaeopterus houkae can be distinguished from all other Phlaeopterus species by its small size, presence of ocelli, vague anteocellar foveae, slight nuchal constriction, shape and punctation of the pronotum, shape of the wing-folding spicules on tergites IV and V, lack of wing-folding spicules on tergite VI, and the unique structure of the internal sac of the aedeagus.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA6FFF8F2EEFAA6FB19F960.taxon	description	(Figs. 3 C, 11 A, 16 A, 21 E, 35 A – B, 41 B) Zoobank. org / urn: lsid: zoobank. org: act: 6 CE 135 BC- 1612 - 4382 - 92 FD- 6 F 6 FB 45 A 6 DBD	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA6FFF8F2EEFAA6FB19F960.taxon	materials_examined	Type Locality. Lassen National Park, California, USA.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA6FFF8F2EEFAA6FB19F960.taxon	description	Description. Habitus: Length 6.2 – 7.6 mm. Dark brown to reddish brown to yellowish brown; head and abdomen nearly black, elytra usually distinctly lighter reddish to yellowish brown; pronotal margins often also distinctly lighter reddish brown to yellowish brown; palpi, antennae, and legs reddish brown to yellowish brown (Fig. 3 C). Head: Moderately broad (Fig. 35 A – B), ratio of width across eyes to head length = 1.0 – 1.3. Interantennal groove broadly and deeply impressed. Anteocellar foveae large, deeply impressed. Eyes glabrous or with less than 10 scattered setae near ventral margin. Antennomeres 5 – 10 at least 2 times longer than wide; antennomeres 4 – 11 each with many sensory pits with papilliform projections. Ocelli present. Nuchal constriction vague. Mandibular molar area with L-shaped row of setae. Hypopharynx as in Fig. 41 B. Labrum with sensory pores along entire surface. Labial palpi with 3 rd palpomere 1.6 – 1.8 times longer than 2 nd palpomere. Thorax: Pronotum broad (Fig. 16 A), length to width ratio = 0.71 – 0.79; ratio of pronotal width to head width = 1.49 – 1.67; maximum width subequal to elytral width across humeral angles; punctures on dorsal surface separated by average distance slightly greater than diameter of a puncture; lateral margins explanate posterad lateral foveae, not explanate anterad lateral foveae; lateral foveae deeply impressed. Elytra with humeral angles convex; epipleural carina not projecting; 2.1 – 2.6 times longer than pronotum. Wings fully developed in most individuals, brachypterous in very few. Mesosternum with projecting tooth; longitudinal carina along midline of mesosternum complete but not strongly carinate. Legs: All tibiae with dense pubescence from base to apex. Metatrochanter without tooth on apical margin. Abdomen: Wing-folding spicules on tergites IV and V broad, transverse, narrowly separated; tergite VI lacking wing-folding patches. Aedeagus: Length 1.16 – 1.34 mm. Median lobe approximately evenly, somewhat convexly narrowed from base to apex; with vague ventral longitudinal carina at apex (Fig. 21 E). Parameres narrow. Internal sac elongate, tube-like, moderately sclerotized; with distinct median transverse fold.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA6FFF8F2EEFAA6FB19F960.taxon	materials_examined	Type Specimens. Holotype male (UAMObs: Ento: 235218) and allotype female (UAMObs: Ento: 235831) labeled as follows: CAL. Lassen N. P., base of Summit Tr., 8500 ’, 16. VII. 1979, JM & BA Campbell / HOLOTYPE ³ (or ALLOTYPE ♀) Phlaeopterus kavanaughi, desig. L. J. Mullen and J. M. Campbell, CNC No. 18464 (red label). Both specimens are deposited in the CNC. Paratypes (n = 183) are deposited in AMNH, NHMUK, CAS, CSCA, FMNH, MCZ, OSAC, USNM, and UCRC.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA6FFF8F2EEFAA6FB19F960.taxon	distribution	Distribution. Phlaeopterus kavanaughi is known from the Sierra Nevada and the Cascade Range of California north to Oregon (Fig. 11 A). Bionomics. Adults have been collected during July – August at 1,830 – 3,400 m elevation, with a single specimen (UAMObs: Ento: 234158) collected at 90 m in Multnomah County, Oregon. This species has been collected at night on snowfields, where	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA1FFFEF0ECFF1CFCF7F960.taxon	description	(Figs. 3 D, 11 B, 15 E, 19 E, 21 A, 28 E, 31 E, 33 E, 34 A – B, 36 A, 38 E – F, 40 E, 41 D, 42 C)	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA1FFFEF0ECFF1CFCF7F960.taxon	discussion	Remarks. Phlaeopterus kavanaughi can be distinguished from all other Phlaeopterus species by the tibiae pubescent to the apices, the shape of the pronotal lateral margins, the ventral longitudinal carina of the median lobe of the aedeagus, the shape and median transverse fold of the internal sac of the aedeagus, and the usually yellowish brown elytra and pronotal lateral margins that are paler than remainder of the body.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA1FFFEF0ECFF1CFCF7F960.taxon	etymology	Etymology. This species is named in honor of Dr. David H. Kavanaugh, California Academy of Sciences, San Francisco, California, USA.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA1FFFEF0ECFF1CFCF7F960.taxon	materials_examined	Type Locality. Carkeek Park, Seattle, Washington, USA.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA1FFFEF0ECFF1CFCF7F960.taxon	description	Redescription. Habitus: Length 3.5 – 4.7 mm. Dark brown to reddish brown; legs, mouthparts, antennae, and elytra usually lighter (Fig. 3 D); elytra with distinctive yellow to reddish yellow area ranging from small spot on humeri to nearly covering elytral disc. Head: Moderately narrow, width across eyes to head length slightly wider than long (Fig. 34 A – B). Interantennal groove large, deeply impressed. Anteocellar foveae small, shallowly impressed. Eyes moderately pubescent on ventral half, with more than 10 setae (Fig. 36 A). Antennomeres 5 – 10 1.2 – 1.8 times longer than wide; antennomeres 4 – 11 each with many sensory pits with papilliform projections. Ocelli present. Nuchal constriction vague. Mandibular molar area with L-shaped row of setae and additional oblique row of setae (Fig. 28 E). Labrum with sensory pores along anterior margin only (Fig. 31 E). Maxilla with lacinia and galea as in Fig. 38 E – F. Epipharynx as in Fig. 33 E. Hypopharynx as in Fig. 40 E. Labial palpi with 3 rd palpomere 1.1 – 1.3 times longer than 2 nd palpomere. Gula as in Fig. 42 C. Thorax: Pronotum moderately narrow (Fig. 15 E), length to width ratio = 0.68 – 0.72; ratio of pronotal width to head width = 1.54 – 1.67; maximum width less than elytral width at bases; dorsal surface coarsely, sparsely punctate, punctures separated by average distance slightly less than diameter of a puncture; lateral margins narrowly explanate posterad lateral fovea, not explanate anterad lateral foveae; lateral foveae moderately impressed, contiguous with lateral margins. Elytra with humeral angles convex; epipleural carina not projecting; 2.2 – 2.5 times longer than pronotum; apical margins convex. Wings fully developed. Mesosternum with projecting tooth; longitudinal carina along midline of mesosternum absent or vague. Legs: All tibiae evenly pubescent to apex. Metatrochanter with tooth on apical margin (Fig. 41 D). Protibia slightly sinuate on medioventral margin in male. Abdomen: Wing-folding spicules on tergites IV and V broad, transverse, nearly contiguous; tergite VI without wing-folding spicules. Apical palisade fringe on tergite VII absent. Aedeagus: Length 0.63 – 0.70 mm. Median lobe triangular. Parameres broad, diverging until just past midline then converging towards apex of median lobe (Fig. 21 A). Internal sac irregular in shape, with 2 apical patches of thick setae.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA1FFFEF0ECFF1CFCF7F960.taxon	materials_examined	Type Specimens. Holotype male (UAMObs: Ento: 235215) labeled as follows: Seattle, WASH., Carkeek Park, Bel., V. 3.1949 / Type ³, Phlaeopterus lagrandeuri, 1951, M. H. Hatch. Allotype female (UAMObs: Ento: 235775) labeled as follows: Corvallis, ORE., VI. 1.1946, KM & DM Fender / ALLOTYPE ♀ Phlaeopterus lagrandeuri 1951, M. H. Hatch / Marys Peak. Both specimens are deposited in the USNM.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA1FFFEF0ECFF1CFCF7F960.taxon	distribution	Distribution. Phlaeopterus lagrandeuri is widely distributed from the Kenai Peninsula of Alaska, British Columbia, including Haida Gwaii and Vancouver Island, south through Washington and Oregon, and one specimen from Calahan, California (UAMObs: Ento: 232633) (Fig. 11 B). Bionomics. Adults have been collected from near sea level to 2,200 m elevation during May – August.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA1FFFEF0ECFF1CFCF7F960.taxon	discussion	Remarks. Phlaeopterus lagrandeuri is easily distinguished from all other Phlaeopterus species by the especially coarse, sparse punctation of the pronotum and elytra, yellow spot on each elytron (although occasionally vague or absent), tooth on the apical margin of the metatrochanter, form of the mandibles, and the unique structure of the internal sac of the aedeagus. This species most closely resembles Siberian P. czerskyi but can be distinguished by many characters including the more shallowly impressed anteocellar foveae, less coarse punctation, lateral margins of the pronotum smooth, not crenulate, fully developed wings, and longer elytra.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA3FFF3F29CFC73FD4DFD48.taxon	materials_examined	Type Locality. Logan Pass, Glacier National Park, Montana, USA.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA3FFF3F29CFC73FD4DFD48.taxon	description	Redescription. Habitus: Length 5.8 – 7.4 mm. Dark brown to black; head often black, pronotum and elytra lighter; antennae, palpi, and legs even lighter, legs becoming progressively lighter from base to apex (Fig. 4 A). Head: Width across eyes to head length subequal to slightly wider than long. Interantennal groove broadly and shallowly impressed. Anteocellar foveae large, deeply impressed. Eyes glabrous or with less than 10 scattered setae near ventral margin. Antennomeres 5 – 10 at least 2 times longer than wide; antennomeres 4 – 11 each with many sensory pits with papilliform projections. Ocelli present. Nuchal constriction vague. Mandibular molar area with L-shaped row of setae. Labrum with sensory pores along entire surface. Labial palpi with 3 rd palpomere 1.6 – 1.8 times longer than palpomere 2. Thorax: Pronotum moderately wide (Fig. 16 H), length to width ratio = 0.68 – 0.75; ratio of pronotal width to head width = 1.52 – 1.67; maximum width subequal to elytral width at humeral angle; punctures separated by distance slightly less than diameter of a puncture on center of disc, almost contiguous towards margin; lateral margins explanate posterad but not anterad lateral foveae; lateral foveae deeply impressed. Elytra with humeral angles convex; epipleural carina not projecting; 2.2 – 2.7 times longer than pronotum; apical margins broadly triangular, elongated at suture (Fig. 29 E). Wings fully developed in most individuals, brachypterous in very few. Mesosternum with projecting tooth; longitudinal carina along midline of mesosternum indistinct. Legs: Apices of all tibiae without pubescence, length of subglabrous apex of metatibia as ratio to metatibial length in males = 10.2 – 17.3, in females = 8.1 – 13.0. Metatrochanter without tooth on apical margin. Abdomen: Wing-folding spicules on tergites IV and V broad, narrowly separated; wing-folding patches on tergite VI small, circular to slightly transverse. Apical palisade fringe on tergite VII present. Aedeagus: Length 1.40 – 1.55 mm long. Median lobe narrow, elongate, lateral margins constricted and subparallel near midline, expanding anterad and posterad midline (Fig. 24 C). Parameres narrow, convexly curved. Internal sac elongate, lightly sclerotized, dorsal half evenly, lightly covered in microspinules.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA3FFF3F29CFC73FD4DFD48.taxon	materials_examined	Type Specimens. Holotype male (UAMObs: Ento: 235773) and allotype female (UAMObs: Ento: 235774) labeled as follows: G. N. P. Mont., Logan Pass, Aug. 22, 1939, M. H. Hatch / TYPE ³ Bionomics. Adults have been collected during June – September at 600 – 2,400 m elevation in wet moss and piles of rocks at the edges of fast, cold streams, in the splash-zone of waterfalls and cascades, and under rocks at the edges of snowfields or snowmelt ponds. Specimens collected at low elevations were associated with springs or fast streams that remain cold to lower elevations.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA3FFF3F29CFC73FD4DFD48.taxon	discussion	Remarks. Phlaeopterus loganensis can be distinguished from all other Phlaeopterus species by having the apices of the elytra triangular in both the male and the female (Fig. 29 E). Although P. elongatus and P. filicornis have the elytral apices triangular in the female, no other species of the genus has the elytral apices triangular in both sexes. Phlaeopterus loganensis is most similar to P. olympicus, and these two species can be distinguished by characters discussed in the Remarks section of the latter species.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFADFFF1F0E5FCC9FDCEF960.taxon	description	(Figs. 4 B, 12 B, 15 G, 20 A, 21 C, 32 C, 36 C, 42 E)	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFADFFF1F0E5FCC9FDCEF960.taxon	distribution	Distribution. Phlaeopterus loganensis is known only from the Rocky Mountains of Idaho, Montana, and British Columbia, the Selkirk Mountains of British Columbia, and Banff and Jasper National Parks, Alberta (Fig. 12 A).	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFADFFF1F0E5FCC9FDCEF960.taxon	materials_examined	Type Locality. Placer County, California, USA.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFADFFF1F0E5FCC9FDCEF960.taxon	description	Redescription. Habitus: Length 2.7 – 3.6 mm. Light brown to dark brown, legs and antennae sometimes lighter (Fig. 4 B). Head: Width across eyes to head length slightly wider than long. Interantennal groove absent. Anteocellar foveae moderately large, deeply impressed. Eyes moderately pubescent on ventral half, with more than 10 setae (Fig. 36 C). Antennomeres 3 – 10 each 1.3 – 1.9 times longer than wide, antennomeres 4 – 11 each with many sensory pits with groups of pore-like openings. Ocelli absent. Nuchal constriction vague. Mandibular molar area with L-shaped row of setae. Maxillary palpus as in Fig. 32 C. Labrum with sensory pores along anterior margin only. Labial palpi with 3 rd palpomere 1.4 – 1.8 times longer than palpomere 2. Gula as in Fig. 42 E. Thorax: Pronotum narrow (Fig. 15 G), length to width ratio = 0.79 – 0.87; ratio of pronotal width to head width = 1.11 – 1.33; maximum width slightly narrower than elytra at base; punctures separated by average distance subequal to or slightly greater than diameter of a puncture; lateral margins very narrowly explanate just posterad lateral fovea, otherwise not at all explanate anterad or posterad lateral foveae. Foveae broad, moderately deeply impressed, narrowly separated from undescribed, D) Unamis sp., Phlaeopterus smetanai. obsoletus, H) E) Phlaeopterus lagrandeuri, F) Phlaeopterus houkae, G) Phlaeopterus sac small, narrowly elongate; with patch of large microspinules.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFADFFF1F0E5FCC9FDCEF960.taxon	materials_examined	Type Specimens. Lectotype male (here designated to clarify the application of this name to this taxon) labeled as follows: Oct. / Placer Co., CAL. / TYPE USNM 48113 / Vellica longipennis Cas. / LECTOTYPE ³ Vellica longipennis Csy., desig. 1984, J. M. Campbell. The Casey collection also contains six paralectotypes, all labeled “ Siskiyou Co., Cal. ” All specimens are deposited in the USNM.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFADFFF1F0E5FCC9FDCEF960.taxon	distribution	Distribution. Phlaeopterus longipennis is known from northern California in the Sierra Nevada, Cascade, and Trinity Mountains, through Oregon to Mt. Rainier National Park, Washington (Fig. 12 B). Bionomics. Adults have been collected at 500 – 3,000 m elevation during May – August in wet moss at the edges of cold streams.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFADFFF1F0E5FCC9FDCEF960.taxon	discussion	Remarks. Phlaeopterus longipennis can be differentiated from all other Phlaeopterus species except P. obsoletus by its absence of ocelli. Phlaeopterus longipennis is externally similar to P. obsoletus, but it differs significantly in aedeagal characters and can be externally differentiated by the punctation of the head and pronotum, its slightly shorter elytra, and its smaller overall size. Casey (1885) stated that Vellica belongs between Lesteva and Phlaeopterus, and compared Vellica to Lesteva, noting that the two genera differ in the structure of the maxillary and labial palpi and the length of the elytra. However, Casey (1885) did not make any morphological comparisons between Phlaeopterus and Vellica. Phlaeopterus longipennis matches the diagnosis of the genus Phlaeopterus given herein and so is moved from the monotypic genus Vellica, thus making Vellica a synonym under Phlaeopterus. This transfer was also suggested by Newton et al. (2000). lateral margins. Elytra with humeral angles convex; epipleural carina not projecting; 2.2 – 2.4 times longer than pronotum; apical margins convex. Wings developed or brachypterous. Mesosternum with projecting tooth; longitudinal carina along midline of mesosternum present, becoming vague near posterior margin (Fig. 20 A). Abdomen: Tergites IV and V each with wing-folding spicules very broad, nearly contiguous or contiguous in shape of single transverse band. Apical palisade fringe on tergite VII absent. Legs: All tibiae evenly pubescent to apex. Metatrochanter without tooth on apical margin. Aedeagus: Length 0.68 – 0.78 mm. Median lobe narrowly triangular. Parameres narrow; diverging near base then parallel (Fig. 21 C). Internal	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFAFFFF6F0FDFB6EFE25F960.taxon	description	(Figs. 4 C, 13 A, 15 H, 17 G, 20 B, 21 D, 29 A, 31 G, 33 G, 34 E – F, 37 H – I, 39 C – D, 41 A) Zoobank. org / urn: lsid: zoobank. org: act: D 2 D 70 CE 0 - DDEC- 4834 - 857 E-DC 6 D 80 FFD 18 D Hatch 1957: 461 [misidentification as V. longipennis].	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFAFFFF6F0FDFB6EFE25F960.taxon	materials_examined	Type Locality. Berkeley Park, Mt. Rainer National Park, Washington, USA.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFAFFFF6F0FDFB6EFE25F960.taxon	description	Description. Habitus: Length 2.9 – 3.9 mm. Light brown to dark brown, legs lighter, antennae sometimes lighter (Fig. 4 C). Head: Width across eyes to head length slightly wider than long (Fig. 34 E – F). Interantennal groove absent. Anteocellar foveae moderately large, deeply impressed. Eyes moderately pubescent on ventral half, with more than 10 setae. Antennomeres 3 – 10 each 1.3 – 1.9 times longer than wide, antennomeres 4 – 11 each with many sensory pits with groups of pore-like openings (Fig. 37 H – I). Ocelli absent. Nuchal constriction vague. Mandibular molar area with Lshaped row of setae. Epipharynx as in Fig. 33 G. Maxillary palpus as in Fig. 39 C – D. Hypopharynx as in Fig. 41 A. Labrum with sensory pores along anterior margin only (Fig. 31 G). Labial palpi with 3 rd palpomere 1.4 – 1.8 times longer than palpomere 2. Thorax: Pronotum narrow (Fig. 15 H), length to width ratio = 0.79 – 0.88; ratio of pronotal width to head width = 1.25 – 1.33; maximum width slightly narrower than elytra at base; punctures separated by average distance subequal to or slightly greater than diameter of a puncture; lateral margins not explanate anterad or posterad lateral fovea; lateral foveae moderately deeply impressed, narrowly separated from lateral margins. Elytra with humeral angles convex; epipleural carina not projecting; 2.0 – 2.3 times longer than pronotum; apical margins convex. Wings developed or brachypterous. Mesosternum with projecting tooth; longitudinal carina along midline of mesosternum vaguely present, becoming obscured on posterior half (Fig. 20 B). Legs: All tibiae evenly pubescent to apex. Metatrochanter without tooth on apical margin. Abdomen: Tergites IV and V with wing-folding spicules very broad, nearly contiguous or contiguous in shape of single transverse band. Apical palisade fringe on tergite VII absent. Aedeagus: Length 0.52 – 0.63 mm. Median lobe short, oblong, sides subparallel, apex broadly convex. Parameres diverging in basal half (Fig. 21 D). Internal sac large; with two mediolateral patches of microspinules.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFAFFFF6F0FDFB6EFE25F960.taxon	materials_examined	Type Specimens. Holotype male (UAMObs: Ento: 235217) and allotype female (UAMObs: Ento: 235837) labeled as follows: WASH, Mt. Rainer N. P., Berkeley Park, 3.5 mi. W. Sunrise, 6700 ’, 8. VIII. 1975, J. M. & B. A. Campbell / HOLOTYPE ³ (or ALLOTYPE ♀) Phlaeopterus obsoletus desig. L. J. Mullen and J. M. Campbell, CNC No. 18470 (red label). Both specimens are deposited in the CNC. Paratypes (n = 878) are deposited in the AMNH, BMNH, CAS, CNC, CSCA, FMNH, MCZ, OSAC, and USNM.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFAFFFF6F0FDFB6EFE25F960.taxon	distribution	Distribution. Phlaeopterus obsoletus is known from the Cascade Range in British Columbia, Oregon, Washington, and California, and from Jasper National Park, Alberta (Fig. 13 A). Bionomics. Adults have been collected at 1,000 – 2,130 m elevation during May – August in wet moss at the edges of streams or in the splash zone of waterfalls.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFAFFFF6F0FDFB6EFE25F960.taxon	discussion	Remarks. Phlaeopterus obsoletus can be distinguished from all other Phlaeopterus species except P. longipennis by the absence of ocelli. Phlaeopterus obsoletus and P. longipennis are very similar but can be differentiated by characters discussed in the Remarks section of the latter species. This species has been misidentified as P. longipennis in most collections as well as by Hatch (1957).	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFAFFFF6F0FDFB6EFE25F960.taxon	etymology	Etymology. This species name is derived from the Latin adjective for effaced or indistinct, referring to the absence of ocelli.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA8FFF4F0C4FA93FBDCF960.taxon	description	(Figs. 4 D, 13 B, 16 F, 20 E, 23 D, 24 A) Zoobank. org / urn: lsid: zoobank. org: act: 7 D 45457 E-C 5 D 0 - 4 CF 2 - 83 FB-BAC 51055 CDCF Casey 1893: 402 [misidentification as T. cavicollis].	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA8FFF4F0C4FA93FBDCF960.taxon	materials_examined	Type Locality. Mt. Rainier National Park, Washington, USA.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA8FFF4F0C4FA93FBDCF960.taxon	description	Description. Habitus: Length 6.1 – 8.2 mm. Reddish brown to black; elytra sometimes lighter than pronotum; palpi, legs, and antennae reddishbrown to black (Fig. 4 D). Head: Width across eyes to head length subequal to slightly wider than long. Interantennal groove broadly and moderately deeply impressed. Anteocellar foveae large, deeply impressed. Eyes glabrous or with less than 10 scattered setae near ventral margin. Antennomeres 5 – 10 at least 2 times longer than wide; antennomeres 4 – 11 each with many sensory pits with papilliform projections. Ocelli present. Nuchal constriction vague. Mandibular molar area with Lshaped row of setae. Labrum with sensory pores along entire surface. Labial palpi with 3 rd palpomere 1.6 – 1.8 times longer than palpomere 2. Thorax: Pronotum moderately wide (Fig. 16 F), length to width ratio = 0.63 – 0.71; ratio of pronotal width to head width = 1.45 – 1.70; maximum width subequal to elytral width at humeral angles; punctures on dorsal surface separated by distance slightly less than to slightly more than diameter of a puncture; lateral margins explanate posterad lateral foveae, narrowly explanate anterad lateral foveae, narrowing to not at all explanate at apex; lateral foveae deeply impressed. Elytra with humeral angles convex; epipleural carina not projecting; 2.2 – 2.6 times longer than pronotum; apical margins broadly convex. Wings fully developed in most individuals, brachypterous in very few. Mesosternum with projecting tooth; longitudinal carina along entire midline of mesosternum (Fig. 20 E). Legs: Apices of all tibia without pubescence, length of subglabrous apex of metatibia as ratio to metatibial length in males = 7.6 – 28.0, in females = 8.6 – 19.4. Metatrochanter without tooth on apical margin. Abdomen: Wing-folding spicules on tergites IV and V broad, transverse, narrowly separated; tergite VI with wing-folding spicules. Aedeagus: Length 1.24 – 1.50 mm. Median lobe irregularly narrowing from base to abruptly truncate apex. Parameres diverging from base to apical fourth that strongly converges towards apex of median lobe (Figs. 23 D, 24 A). Internal sack lightly sclerotized; with subapical transverse fold; apical 2 / 3 covered in microspinules except for smooth area of transverse fold.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA8FFF4F0C4FA93FBDCF960.taxon	materials_examined	Type Specimens. Holotype male (UAMObs: Ento: 235826) and allotype female (UAMObs: Ento: 235827) labeled as follows: WASH., Mt. Rainier N. P., end of West Side Rd., 3. VIII. 1979, 3700 ’, J. M. & B. A. Campbell / HOLOTYPE ³ (or AL- LOTYPE ♀) Phlaeopterus occidentalis desig. L. J. Mullen and J. M. Campbell CNC No. 18466 (red label). Both specimens are in the CNC, Ottawa. Paratypes (n = 232) are deposited in the AMNH, CAS, CNC, CSCA, FMNH, MCZ, ROM, USNM, and UCRC.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA8FFF4F0C4FA93FBDCF960.taxon	distribution	Distribution. Phlaeopterus occidentalis is known from southern California north through Oregon, Washington, and British Columbia to southeastern Alaska (Fig. 13 B). A single specimen (UAM: Ento: 332755) was collected from Great Basin National Park, Nevada. We attempted to re-collect this species from the 1968 locality (UAMObs: Ento: 233463) near Haines, Alaska, in the summer of 2014 but were unsuccessful. Bionomics. Adults of this species have been collected at 300 – 3,200 m, a broader elevation range than any other species in the genus. In British Columbia and Alaska, the species has only been collected at 300 – 1,800 m. Adults have been collected during May – October in California and Oregon, but only during June – August in Washington and Alaska. This species has been collected under rocks and in moss in the splash-zone at the edges of small streams or waterfalls.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA8FFF4F0C4FA93FBDCF960.taxon	discussion	Remarks. Specimens from Vancouver Island vary from those of the mainland by having the base of the internal sac of the aedeagus broader and the apex of the internal sac concave. The aedeagus of the Vancouver Island (Fig. 24 A) and mainland (Fig. 23 D) populations are illustrated. There is also a slight difference in the length of the posterior tibiae vs. posterior tarsus and in the length of the antennomeres approaching the apex. Furthermore, the mainland population shows geographic variation in the aedeagus, with the aedeagus becoming slightly longer and the apex of the median lobe progressively more narrowly triangular moving from south to north. We deem these differences insufficient to warrant splitting these populations into multiple species or subspecies. Casey (1893) erroneously referred to a specimen of P. occidentalis as T. cavicollis in his treatment of the latter species, likely because the type series of the latter species was mixed. This error was fixed by a lectotype designation for P. cavicollis. The specimen examined by Casey (1893) is labeled as follows: Vanc. / ♀ / CASEY bequest 1925 / cavicollis Fvl. / May be from original type series. – see Casey, Col. notices V. 1893, p. 402.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFA8FFF4F0C4FA93FBDCF960.taxon	etymology	Etymology. This species is named for its wide distribution in northwestern North America.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFB5FFEBF2E3FF1CFC63FA20.taxon	description	(Figs. 5 A, 14 A, 16 G, 20 C, 24 B, 36 F) Zoobank. org / urn: lsid: zoobank. org: act: 7 B 1 AF 64 C- 7 E 3 E- 4214 - 834 B- 02849 F 1 FA 285	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFB5FFEBF2E3FF1CFC63FA20.taxon	materials_examined	Type Locality. Olympic National Park, Washington, USA.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFB5FFEBF2E3FF1CFC63FA20.taxon	description	Description. Habitus: Length 6.3 – 8.2 mm. Dark brown to reddish brown; legs, elytral epipleurae, and palpi brown to reddish brown; elytra often lighter than pronotum (Fig. 5 A). Head: Width across eyes to head length subequal to slightly wider than long. Interantennal groove broadly and shallowly impressed. Anteocellar foveae large, deeply impressed. Eyes glabrous or with less than 10 setae near ventral margin (Fig. 36 F). Antennomeres 5 – 10 at least 2 times longer than wide; antennomeres 4 – 11 each with many sensory pits with papilliform projections. Ocelli present. Nuchal constriction vague. Mandibular molar area with L-shaped row of setae. Labrum with sensory pores along entire surface. Labial palpi with 3 rd palpomere 1.6 – 1.8 times longer than palpomere 2. Thorax: Pronotum moderately wide (Fig. 16 G), length to width ratio = 0.68 – 0.72; ratio of pronotal width to head width = 1.52 – 1.59; maximum width subequal to elytral width at humeral angle; punctures on dorsal surface separated by distance equal to diameter of a puncture; lateral margins explanate posterad but not anterad lateral foveae; lateral foveae deeply impressed. Elytra with humeral angles convex; epipleural carina not projecting; 2.2 – 2.5 times longer than pronotum; apical margins broadly convex. Wings fully developed in most individuals, brachypterous in very few. Mesosternum with projecting tooth; longitudinal carina along midline of mesosternum indistinct (Fig. 20 C). Legs: Apices of all tibia without pubescence, length of subglabrous apex of metatibia as ratio to metatibial length = 8.3 – 20.0. Metatrochanter without tooth on apical margin. Abdomen: Wing-folding spicules on tergites IV and V broad, transverse, narrowly separated; tergite VI lacking wing-folding spicules. Aedeagus: Length 1.50 – 1.60 mm. Median lobe broadly triangular, margins narrowing more sharply nearing apex. Parameres not or barely curved towards apex of median lobe, originating near middle of median lobe (Fig. 24 B). Internal sac broadly triangular, lightly sclerotized, covered with microspinules.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFB5FFEBF2E3FF1CFC63FA20.taxon	materials_examined	Type Specimens. Holotype male (UAMObs: Ento: 235824) and allotype female (UAMObs: Ento: 235825) labeled as follows: WASH. Olympic N. P., 0.5 mi S Lake No. 8 (7 Lakes Basin) 5000 ’, 16. VIII. 1979, J. M. and B. A. Campbell / HOLOTYPE ³ (or ALLOTYPE ♀) Phlaeopterus olympicus, desig. L. J. Mullen and J. M. Campbell, CNC No. 18464 (red label). Both specimens are in the CNC. Paratypes (n = 305) are deposited in the AMNH, CAS, CNC, CSCA, FMNH, MCZ, and USNM.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFB5FFEBF2E3FF1CFC63FA20.taxon	distribution	Distribution. Phlaeopterus olympicus is known only from the Olympic Mountains of Washington State, inside Olympic National Park, and from a single specimen (UAMObs: Ento: 234159) from Harts Pass, Okanagan County, Washington (Fig. 14 A). Bionomics. Adults of this species have been collected during July – August at 650 – 1,970 m elevation, with most specimens collected at 1,500 – 1,800 m. Adults have been collected as they were feeding on arthropods on the surface of snowfields at night and during the day under wet rocks and gravel at the edges of snowfields or in moss or wet rocks at the edges of cold streams flowing from snowfields. Some specimens have been collected at lower elevations under stones or in moss along the edges of the Soleduck River or in moss and under stones at the edges of streams draining high-elevation lakes.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFB5FFEBF2E3FF1CFC63FA20.taxon	discussion	Remarks. Phlaeopterus olympicus can be distinguished from all other Phlaeopterus species by the lack of a mesosternal carina but presence of a mesosternal tooth, its pronotal shape, and the glabrous apices of the tibiae. Phlaeopterus olympicus is most similar to P. loganensis, from which it may be distinguished by having the apex of the elytra broadly convex, and by the shape of the aedeagus. Phlaeopterus olympicus and P. loganensis are the only two species of the genus completely lacking a carina along the midline of the mesosternum, although the carina is vague, especially on the posterior half, in other species. Phlaeopterus olympicus has not been collected since 1984, despite three other species having been collected recently from the Olympic Mountains: P. fusconiger as recently as 2014 (UAM: Ento: 299528), P. cavicollis as recently as 2008 (UAMObs: Ento: 232734), and P. lagrandeuri as recently as 1996 (UAMObs: Ento: 234415).	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFB5FFEBF2E3FF1CFC63FA20.taxon	etymology	Etymology. The specific epithet olympicus is derived from the type locality.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFB5FFEAF08CFA61FCD3FC7B.taxon	description	(Figs. 5 B, 14 B, 16 E, 23 B – C, 33 H) Zoobank. org / urn: lsid: zoobank. org: act: B 4 AB 0 AFD-A 8 C 3 - 4922 - 8957 - C 875 B 57264 E 8	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFB5FFEAF08CFA61FCD3FC7B.taxon	materials_examined	Type Locality. Mt. Hood National Forest, Oregon, USA.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFB5FFEAF08CFA61FCD3FC7B.taxon	description	Description. Habitus: Length 6.0 – 9.0 mm. Black, usually darker than any other Phlaeopterus species; legs, palpi, and antennae sometimes lighter, dark brown (Fig. 5 B). Head: Broad, ratio of width across eyes to head length about 5: 4. Interantennal groove broadly and deeply impressed. Anteocellar foveae large, deeply impressed. Eyes glabrous or with less than 10 scattered setae near ventral margin. Antennomeres 5 – 10 at least 2 times longer than wide; antennomeres 4 – 11 each with many sensory pits with papilliform projections. Ocelli present. Nuchal constriction vague. Epipharynx as in Fig. 33 H. Mandibular molar area with L-shaped row of setae. Labrum with sensory pores along entire surface. Labial palpi with 3 rd palpomere 1.6 – 1.8 times longer than palpomere 2. Thorax: Pronotum somewhat broad (Fig. 16 E), length to width ratio = 0.63 – 0.77; ratio of pronotal width to head width = 1.49 – 1.67; maximum width subequal to elytral width at humeral angles; punctures on dorsal surface separated by average distance slightly greater than diameter of a puncture; lateral margins explanate posterad lateral foveae, progressively more narrowly explanate from lateral foveae to just before apex; lateral foveae deeply impressed. Elytra with humeral angles convex; epipleural carina not projecting; 2.3 – 2.6 times longer than pronotum; apical margins broadly convex. Wings fully developed in most individuals, brachypterous in very few. Mesosternum with projecting tooth, tooth more strongly notched on posterior margin than on anterior margin; longitudinal carina along midline of mesosternum complete. Legs: All tibiae with dense pubescence to apex or with small glabrous region at apex, if with glabrous region, length of glabrous apex of mesotibia shorter than length of basal mesotarsomere. Metatrochanter without tooth on apical margin. Abdomen: Wing-folding spicules on tergites IV and V broad, transverse, narrowly separated; tergite VI lacking wingfolding patches. Aedeagus: Length 1.28 – 1.67 mm. Median lobe roughly triangular, evenly to convexly narrowed from base to apex. Parameres slightly diverging from base to near apex then converging; ending just beyond to distinctly beyond apex of median lobe (Fig. 23 B – C). Internal sac elongate, roughly rectangular, narrowed from base to apex; apex subtruncate; lightly sclerotized; more densely covered in microspinules on apical half.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFB5FFEAF08CFA61FCD3FC7B.taxon	materials_examined	Type Specimens. Holotype male (UAMObs: Ento: 235828) and paratype female (UAMObs: Ento: 235829) labeled as follows: ORE., Mt. Hood N. F., Tilly Jane Cr., 7000 ’ 31. VII. 1979, J. M. & B. A. Campbell / HOLOTYPE ³ (or ALLOTYPE ♀) Phlaeopterus smetanai, desig. L. J. Mullen and J. M. Campbell CNC No. 18465 (red label). Both specimens are in the CNC. Paratypes (n = 504) are deposited in the AMNH, CAS, CNC, CSCA, FMNH, MCZ, NHMUK, OSAC, RBC, ROM, UCRC, and USNM.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFB5FFEAF08CFA61FCD3FC7B.taxon	distribution	Distribution. Phlaeopterus smetanai is broadly distributed in the northwestern USA from Washington east to Idaho and south to Colorado, Arizona, and southern California (Fig. 14 B). Bionomics. Adults have been collected during April – October at elevations of 360 – 3,000 m. Adults have been collected under rocks and in moss from the edges of cold streams or in the splash zones of waterfalls, as well as in gravel at the edges of springs and in one case from under rocks near a spring that had recently stopped flowing.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFB5FFEAF08CFA61FCD3FC7B.taxon	discussion	Remarks. Phlaeopterus smetanai is more uniformly black than any other Phlaeopterus species. It is most similar to P. occidentalis, from which it can be distinguished by its darker color, the subglabrous region at the apex of the mesotibia shorter, and by aedeagal characters, including lacking the subapical transverse fold of the latter species. Members of a population near Payson, Arizona differ from other specimens by their shape and larger size of the aedeagus (1.60 – 1.67 mm vs. 1.28 – 1.52 mm; Payson population aedeagus in Fig. 23 C, main population aedeagus in Fig. 23 B). Genetic analysis could help determine if this population is a distinct species.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
BA2A87DCFFB5FFEAF08CFA61FCD3FC7B.taxon	etymology	Etymology. This species is named in honor of Dr. Aleš Smetana, Biosystematics Research Institute, Ottawa, Canada.	en	Mullen, Logan J., Campbell, J. M., Sikes, Derek S. (2018): Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). The Coleopterists Bulletin 72: 1, DOI: 10.1649/0010-065X-72.mo4.1, URL: http://dx.doi.org/10.1649/0010-065x-72.mo4.1
