taxonID	type	description	language	source
B8748B02FFF6FF85FF63B03CFDACF861.taxon	materials_examined	Type species: Nargus taiwanensis Perreau, 1996 a, by original designation.	en	Wang, Cheng-Bin, Růžička, Jan, Zhou, Hong-Zhang (2015): Nargus (Eunargus) celli sp. nov. (Coleoptera: Leiodidae: Cholevinae: Cholevini), a new species from China. Zootaxa 4012 (3): 570-580, DOI: 10.11646/zootaxa.4012.3.10
B8748B02FFF6FF85FF63B03CFDACF861.taxon	diagnosis	Diagnosis. Large body size compared to other subgenera; pronotal hind corners absent, completely rounded; protibiae without fringe of small spines on apical and dorsal margin, outer spurs visible; aedeagus with median lobe divided into two parts: dorsal and ventral laminae; internal sac with isolate and extremely developed tooth, approximately as long as median lobe of aedeagus (Perreau, 1996 a; 1996 b; 1997).	en	Wang, Cheng-Bin, Růžička, Jan, Zhou, Hong-Zhang (2015): Nargus (Eunargus) celli sp. nov. (Coleoptera: Leiodidae: Cholevinae: Cholevini), a new species from China. Zootaxa 4012 (3): 570-580, DOI: 10.11646/zootaxa.4012.3.10
B8748B02FFF6FF85FF63B03CFDACF861.taxon	distribution	Distribution. China (Fig. 1).	en	Wang, Cheng-Bin, Růžička, Jan, Zhou, Hong-Zhang (2015): Nargus (Eunargus) celli sp. nov. (Coleoptera: Leiodidae: Cholevinae: Cholevini), a new species from China. Zootaxa 4012 (3): 570-580, DOI: 10.11646/zootaxa.4012.3.10
B8748B02FFF5FF82FF63B011FE70FD31.taxon	materials_examined	Material examined. CHINA, Fujian: 1 ♂, Kuatun [ca. 27 ° 40 ′ N 117 ° 40 ′ E], 8. IV. 1946, Tschung sen. leg. // Nargus (Eunargus) / franki Perreau, 1997 / Jan Růžička det. 2005 (CJFR); 1 ♂, same data as previous (CJRZ); 1 ♀, Kuatun, 25. IV. [19] 46, Tschung sen. leg. // Nargus (Eunargus) / franki Perreau, 1997 / Jan Růžička det. 2005 (CJFR); CHINA, Sichuan: 1 ♂, 2 ♀, Wolong Natur. Res. [ca. 31 ° 06 ′ N 103 ° 15 ′ E], 900 m, litter, 23. V. 1994, Kurbatov [leg.] // Nargus (Eunargus) / franki Perreau, 1997 / Jan Růžička det. 2005 (CJFR); 1 ♀, same data as previous (CJRZ).	en	Wang, Cheng-Bin, Růžička, Jan, Zhou, Hong-Zhang (2015): Nargus (Eunargus) celli sp. nov. (Coleoptera: Leiodidae: Cholevinae: Cholevini), a new species from China. Zootaxa 4012 (3): 570-580, DOI: 10.11646/zootaxa.4012.3.10
B8748B02FFF5FF82FF63B011FE70FD31.taxon	description	Redescription. Male. EBL: 3.8 mm. Length of different body parts: HL: AL: PL: ELL = 0.6: 1.6: 0.9: 2.1 mm; width: HW: EW: PW: ELW = 1.0: 0.1: 1.4: 1.6 mm. Proportion of antennomeres from base to tip in Μm (length × width): 175 × 87, 164 × 68, 157 × 73, 125 × 67, 118 × 69, 112 × 75, 139 × 83, 92 × 86, 126 × 100, 130 × 108, 191 × 105. Habitus (Fig. 2 A) oval, rather convex and sublustrous. Moderately pigmented: mostly yellowish-brown; antennomeres 6 to 10 somewhat darker; each elytron with a large blackish-brown spot at middle. Dorsum continually covered with fine, recumbent, sallow pubescence. Head retractile, HW / HL = 1.5. Surface covered with fine punctures, separated about 5 – 6 times of their diameter, interspaces with reticulate microsculpture. Frontoclypeal suture distinct. Clypeus subtrapezoidal, anterior margin substraight. Compound eyes well developed, EW / HW = 0.1. Antennae (Fig. 3 A) slender and long, AL / HW = 1.7; all antennomeres longer than wide; antennomere 3 slightly shorter than 2; antennomere 6 with length / width = 1.5; 11 th longest, rather elongated. Pronotum (Fig. 3 B) transverse, widest at about basal 2 / 5, PW / PL = 1.6. Sides rounded, regularly narrowing forward and backward from the widest point; hind corners absent, completely rounded. Posterior margin shallowly and widely emarginate in middle. Surface covered with fine punctures, interspaces with reticulate microsculpture. Elytra widely oval, widest at about basal 3 / 8, ELL / EW = 1.3. Sides regularly curved, gradually narrowing from the widest point to apices; apices (Fig. 3 G) widely subrounded. Sutural striae present. Surface covered with fine punctures, interspaces with sparse microtrichiae. Metathoracic wings fully developed. Prolegs thin and slender, with basal three protarsomeres (Fig. 3 C) expanded: TW / BTW = 1.3. Protibiae (Fig. 3 E) slightly dilated around middle and weakly narrowed towards apex. Profemora wider than protibiae. Mesotibiae slightly curved, mesotarsi simply linear. Metatibiae straight. Abdominal ventrite VIII (Fig. 3 I) very shallowly and widely emarginate on posterior margin. Genital segment (Fig. 3 J) long and elliptical, with spiculum gastrale weakly sclerotized; genital plate narrow; tergite IX rounded ventro-apically. Aedeagus (Figs. 4 A – C) with median lobe divided into two laminae: in dorsal view (Fig. 4 A), dorsal lamina extended over entire width of median lobe, strongly acuminated into a narrow, obliquely truncated apex, and bearing ca. 4 long and strong setae subapically on right side; while ventral lamina strongly and transversely reduced, shorter than dorsal lamina, situated at right side, with hook-like apex. Large tooth of internal sac extremely developed, slender and sinuate, projecting outward from median lobe. Parameres slender, slightly longer than median lobe, distinctly constricted below apices, each with two apical setae. In lateral view (Fig. 4 C), median lobe regularly bent ventrad, large tooth of internal sac upturned, and parameres almost with equal width throughout whole length. Internal sac with many minute denticles in basal part. Female. Similar to male in general appearance (Fig. 2 B), but can be distinguished from following characters: protarsi (Fig. 3 D) simply linear; protibiae (Fig. 3 E) more slender; elytral apex (Fig. 3 H) slightly wider; ventrite VIII (Fig. 4 D) gently rounded at posterior margin, and with widely rounded spiculum ventrale; valvifer (Figs. 4 E & F) with 1 lateral seta; coxite (Figs. 4 E & F) with 3 subapical setae and 1 fine lateral seta; stylus (Figs. 4 E & F) minute, cylindrical, with 1 long apical seta. Affinities. Nargus (Eunargus) franki Perreau is easily distinguished from other members of this subgenus by the combination of the following characters: body size small (3.8 mm); mostly yellowish-brown, each elytron with a large blackish-brown spot at the middle; metathoracic wings fully developed; aedeagus with dorsal lamina slender, with narrow apex; ventral lamina situated at right side in dorsal view, shorter than dorsal lamina; large tooth of internal sac simple, projected outward from median lobe; parameres slender, slightly longer than median lobe, and in lateral view almost with equal width throughout whole length.	en	Wang, Cheng-Bin, Růžička, Jan, Zhou, Hong-Zhang (2015): Nargus (Eunargus) celli sp. nov. (Coleoptera: Leiodidae: Cholevinae: Cholevini), a new species from China. Zootaxa 4012 (3): 570-580, DOI: 10.11646/zootaxa.4012.3.10
B8748B02FFF5FF82FF63B011FE70FD31.taxon	distribution	Distribution. China (Fujian, Sichuan) (Fig. 1).	en	Wang, Cheng-Bin, Růžička, Jan, Zhou, Hong-Zhang (2015): Nargus (Eunargus) celli sp. nov. (Coleoptera: Leiodidae: Cholevinae: Cholevini), a new species from China. Zootaxa 4012 (3): 570-580, DOI: 10.11646/zootaxa.4012.3.10
B8748B02FFF5FF82FF63B011FE70FD31.taxon	discussion	Remarks. We also examined two female paratypes, labeled “ CHINA, Zhejiang: Lin’an County, 500 m, W. Tianmu Shan N. R. [ca. 30 ° 20 ′ N 119 ° 25 ′ E], 16. V. 1996, J. Cooter // Cryptomeria - Liriodendron dominant high forest leaf litter. Winkler extractor // PARATYPE / NARGUS (EUNARGUS) FRANKI n. sp. / M. Perreau det. 1997 (CJRZ, OUMNH) ”. After dissection, we found its metathoracic wings strongly reduced and ventrite VIII with narrowly subtriangular spiculum ventrale that is obviously different from Nargus (Eunargus) franki. Therefore, we think that they can represent an undescribed species very close to N. (E.) franki. Until discovery of male, we refrain from its formal description.	en	Wang, Cheng-Bin, Růžička, Jan, Zhou, Hong-Zhang (2015): Nargus (Eunargus) celli sp. nov. (Coleoptera: Leiodidae: Cholevinae: Cholevini), a new species from China. Zootaxa 4012 (3): 570-580, DOI: 10.11646/zootaxa.4012.3.10
B8748B02FFF1FF82FF63B4A6FDC4FB54.taxon	materials_examined	Material examined. None. Affinities. Nargus (Eunargus) taiwanensis Perreau, according to the original description, is easily to distinguish from other members of this subgenus by the combination of the following characters: body size small (3.6 mm) and uniformly brown, including elytra; metathoracic wings absent; apical angles of elytra elongated into two very strong teeth that overlap each other; aedeagus with dorsal lamina robust, with straight, slightly assymetrical apex; ventral lamina situated at left side in dorsal view, distinctly longer than dorsal lamina; large tooth of internal sac strongly sinuate, not projected outward from median lobe; parameres relatively slender, distinctly shorter than median lobe, and in lateral view gradually widened towards subapex and then narrowed apically.	en	Wang, Cheng-Bin, Růžička, Jan, Zhou, Hong-Zhang (2015): Nargus (Eunargus) celli sp. nov. (Coleoptera: Leiodidae: Cholevinae: Cholevini), a new species from China. Zootaxa 4012 (3): 570-580, DOI: 10.11646/zootaxa.4012.3.10
B8748B02FFF1FF82FF63B4A6FDC4FB54.taxon	distribution	Distribution. China (Taiwan) (Fig. 1).	en	Wang, Cheng-Bin, Růžička, Jan, Zhou, Hong-Zhang (2015): Nargus (Eunargus) celli sp. nov. (Coleoptera: Leiodidae: Cholevinae: Cholevini), a new species from China. Zootaxa 4012 (3): 570-580, DOI: 10.11646/zootaxa.4012.3.10
B8748B02FFF1FF8DFF63B34AFD3AFCA0.taxon	materials_examined	Type locality. CHINA, Guangxi: Napo, Defu Nature Reserve [ca. 23 ° 17 ′ N 105 ° 47 ′ E], 1400 m. Type material. Holotype: CHINA, Guangxi: ♂, Napo, Defu Nature Reserve, 1400 m, aspirator, 3 – 5. VI. 1998, Min Wu leg. (IZ-CAS).	en	Wang, Cheng-Bin, Růžička, Jan, Zhou, Hong-Zhang (2015): Nargus (Eunargus) celli sp. nov. (Coleoptera: Leiodidae: Cholevinae: Cholevini), a new species from China. Zootaxa 4012 (3): 570-580, DOI: 10.11646/zootaxa.4012.3.10
B8748B02FFF1FF8DFF63B34AFD3AFCA0.taxon	description	Description. Male. EBL: 4.4 mm. Length of different body parts: HL: AL: PL: ELL = 0.7: 1.8: 1.1: 2.4 mm; width: HW: EW: PW: ELW = 1.0: 0.1: 1.6: 2.0 mm. Proportion of antennomeres from base to tip in Μm (length × width): 181 × 86, 167 × 67, 188 × 66, 146 × 62, 145 × 67, 124 × 72, 155 × 90, 92 × 85, 139 × 100, 139 × 103, 216 × 98. Habitus (Fig. 2 C) oval, rather convex and sublustrous. Well pigmented: mostly dark brown; mouthparts, basal two and ultimate antennomeres, protarsi and apex of meso- and metatarsi more or less yellowish. Dorsum continually covered with fine, recumbent, sallow pubescence. Head retractile, HW / HL = 1.4. Surface covered with fine punctures, separated about 3 – 5 times of their diameter, interspaces with reticulate microsculpture. Frontoclypeal suture distinct. Clypeus subtrapezoidal, anterior margin substraight. Compound eyes well developed, EW / HW = 0.1. Antennae (Fig. 5 A) very slender and very long, AL / HW = 1.8; all antennomeres longer than wide; antennomere 3 distinctly longer than 2; antennomere 6 with length / width = 1.7; antennomere 11 longest, rather elongated. Pronotum (Fig. 5 B) transverse, widest around middle, PW / PL = 1.6. Sides rounded, regularly narrowing forward and backward from the widest point; hind corners absent, completely rounded. Posterior margin shallowly and narrowly emarginate in middle. Surface covered with fine punctures, interspaces with reticulate microsculpture. Elytra widely oval, widest at about basal 1 / 3, ELL / EW = 1.2. Sides regularly curved, gradually narrowing from the widest point to apices; apices (Fig. 5 E) widely truncated and with inner angle obtusely rounded. Sutural striae present. Surface covered with fine punctures, interspaces with microtrichiae. Metathoracic wings fully developed. Prolegs thin and slender, with basal three protarsomeres (Fig. 5 C) weakly expanded: TW / BTW = 1.1. Protibiae (Fig. 5 D) slightly dilated below middle and weakly narrowed towards apex. Profemora wider than protibiae. Mesotibiae slightly curved, mesotarsi simply linear. Metatibiae substraight. Abdominal ventrite VIII (Fig. 5 F) deeply emarginate beside large median protrusion on posterior margin. Genital segment (Fig. 5 G) wide and subtriangular, with spiculum gastrale small; genital plate narrow; tergite IX widely rounded ventro-apically. Aedeagus (Figs. 6 A – F) with median lobe divided into two laminae: in dorsal view (Figs. 6 A & D), dorsal lamina extended over entire width of median lobe, abruptly turning to left side at apical part where covered with micro-setae, and bearing ca. 4 long and strong setae on left side above middle; while ventral lamina slender, strongly and transversely reduced, slightly shorter than dorsal lamina, situated at left side. Large tooth of internal sac extremely developed and with two apices, left one short and straight, right one long and turned to right side at apical part in dorsal view. Parameres rather wide and robust, slightly longer than median lobe, each with two apical setae. In lateral view (Figs. 6 C & F), median lobe strongly bent ventrad and parameres very widened around middle. Internal sac with many minute denticles in basal part. Female. Unknown. Affinities. Nargus (Eunargus) celli sp. nov. is easily distinguished from other members in this subgenus by the combination of the following characters: distintly larger body size (4.4 mm) and uniformly dark brown, including elytra; metathoracic wings fully developed; aedeagus with dorsal lamina robust, abruptly turning left in apical part in dorsal view; ventral lamina straight, situated at left side in dorsal view, slightly shorter than dorsal lamina; large tooth of internal sac with two apices, not projected outwards from median lobe; parameres wide and robust, slightly longer than median lobe, and in lateral view very widened around middle.	en	Wang, Cheng-Bin, Růžička, Jan, Zhou, Hong-Zhang (2015): Nargus (Eunargus) celli sp. nov. (Coleoptera: Leiodidae: Cholevinae: Cholevini), a new species from China. Zootaxa 4012 (3): 570-580, DOI: 10.11646/zootaxa.4012.3.10
B8748B02FFF1FF8DFF63B34AFD3AFCA0.taxon	etymology	Etymology. The specific epithet is from the name of “ Cell ”, a fictional character, an insect-like humanoid in the Dragon Ball manga series created by Akira Toriyama.	en	Wang, Cheng-Bin, Růžička, Jan, Zhou, Hong-Zhang (2015): Nargus (Eunargus) celli sp. nov. (Coleoptera: Leiodidae: Cholevinae: Cholevini), a new species from China. Zootaxa 4012 (3): 570-580, DOI: 10.11646/zootaxa.4012.3.10
B8748B02FFF1FF8DFF63B34AFD3AFCA0.taxon	distribution	Distribution. China (Guangxi) (Fig. 1).	en	Wang, Cheng-Bin, Růžička, Jan, Zhou, Hong-Zhang (2015): Nargus (Eunargus) celli sp. nov. (Coleoptera: Leiodidae: Cholevinae: Cholevini), a new species from China. Zootaxa 4012 (3): 570-580, DOI: 10.11646/zootaxa.4012.3.10
