identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
BE00878FFFAB810846BB8B03FAF3651A.text	BE00878FFFAB810846BB8B03FAF3651A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Holotrichia burmeisteri Brenske 1892	<div><p>Holotrichia burmeisteri Brenske, 1892</p><p>Although Holotrichia Hope, 1837 is a well-used name, it is a junior synonym of Ancylonycha Dejean, 1833, and an application to the International Commission on Zoological Nomenclature is needed to conserve Holotrichia as the valid name (Bousquet &amp; Bouchard 2013; Allsopp &amp; Schoolmeesters 2024).</p><p>Brenske (1892) described Holotrichia burmeisteri Brenske, 1892 from specimens labelled Philippines, Palau and Austrl. (which he presumed to be Australia). He highlighted three specimens but probably saw more; no holotype was designated. ZMB holds six specimens (Figs 2 – 7), five males and one of unknown gender, all of which were labelled as syntypes in 2007. Five are labelled as from Brenske’s collection (three as “cotypes” and two as “types”), and one from Schaufuss’ collection (labelled “Type”). All bear identification labels in Brenske’s handwriting (compare Figs 2–7 with Horn &amp; Kahle (1935 –37, tafel III, fig. 6)). To stabilise the nomenclature, I here designate a lectotype and have labelled it and the paralectotypes.</p><p>Type series. Lectotype ♂ (here designated) (Fig. 2): Aedeagus and spiculum gastrale dissected out and mounted on card with one set of tarsomeres; Hist.-Coll. Coleoptera Nr. 100783 Holotrichia burmeisteri Brenske Philippinen. Coll. Schaufuss Zool. Mus. Berlin [printed, yellow label] | 100783 [printed] | Schaufuss [printed] | Type [printed, faded red label] | A 40 Tus Philippen [handwritten] | Burmeisteri Brenske Philippinen [handwritten] | Holotrichia (subg burmeisteri m. [handwritten] | SYNTYPUS Holotrichia burmeisteri Brenske, 1892 labelled by MNHUB 2007 [printed, red label] | MfN URI http://coll.mfn-berlin.de/u/09eb4f [printed with QR code]; in ZMB.</p><p>Paralectotypes (Figs 3–7): 1♂, aedeagus and spiculum gastrale dissected out and mounted on card; Burmeisteri Brsk [handwritten] | Philippin coll. [illegible word] [handwritten] | Coll. Brenske [printed, black border] | SYNTYPUS Holotrichia burmeisteri Brenske, 1892 labelled by MNHUB 2007 [printed, red label] | MfN URI http://coll.mfn-berlin.de/u/09fcc3 [printed with QR code]; 1♂, Philippin [handwritten] | Coll. Brenske [printed, black border] | Burmeisteri cotyp. Brsk [handwritten]; Burmeisteri Brsk [handwritten]; SYNTYPUS Holotrichia burmeisteri Brenske, 1892 labelled by MNHUB 2007 [printed, red label] | MfN URI http://coll.mfn-berlin.de/u/09c3ee [printed with QR code]; 1♂, Burmeisteri typ. Brsk [handwritten] | Philippin coll. [illegible word] [handwritten] | Coll. Brenske [printed, black border] | SYNTYPUS Holotrichia burmeisteri Brenske, 1892 labelled by MNHUB 2007 [printed, red label] | MfN URI http://coll.mfn-berlin.de/u/09ed20 [printed with QR code]; 1♂, Palau Ins. Coll. Felsche [handwritten] | Burmeisteri cotyp. Brsk [handwritten] | Coll. Brenske [printed, black border] | SYNTYPUS Holotrichia burmeisteri Brenske, 1892 labelled by MNHUB 2007 [printed, red label] | MfN URI http://coll.mfn-berlin.de/u/09fe5b [printed with QR code]; 1♂, Aedeagus and spiculum gastrale dissected out and mounted on card; Austrl. [printed] | Burmeisteri cotyp Brsk [handwritten] | Coll. Brenske [printed, black border] | SYNTYPUS Holotrichia burmeisteri Brenske, 1892 labelled by MNHUB 2007 [printed, red label] | MfN URI http://coll.mfn-berlin.de/u/0a0b80 [printed with QR code]; all in ZMB.</p><p>Brenske (1892) and Matsumoto (2008) thought that the presumed Australian record (labelled as “Austrl.”, Fig. 7) was doubtful. No specimens of H. burmeisteri were recorded by Britton (1978) in his comprehensive revision of the Australian Melolonthini, with the only record of Holotrichia spp. being a single, damaged, presumably transported specimen of H. serrata (Fabricius, 1781) by Blackburn (1890) [as Neolepidiota obscura Blackburn, 1890]. No further specimens of H. burmeisteri are known from the Philippines and the location was more recently considered doubtful by Matsumoto (2008). Brenske’s record presumably refers to when Spain administered the Caroline Islands as Nuevas Filipinas. Matsumoto (2008) concluded that the type series came from Palau.</p></div>	https://treatment.plazi.org/id/BE00878FFFAB810846BB8B03FAF3651A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Allsopp, Peter G.	Allsopp, Peter G. (2025): Holotrichia burmeisteri Brenske, 1892 and allies (Coleoptera: Scarabaeidae: Melolonthinae: Rhizotrogini): a tale of four genera and four countries. Zootaxa 5631 (1): 166-178, DOI: 10.11646/zootaxa.5631.1.8, URL: https://doi.org/10.11646/zootaxa.5631.1.8
BE00878FFFA9810846BB8883FB3E6691.text	BE00878FFFA9810846BB8883FB3E6691.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Metatrogus Britton 1978	<div><p>Transfer to Metatrogus Britton, 1978</p><p>Matsumoto (2008) examined the type series of H. burmeisteri, as well as additional specimens from Babelthuap, and transferred the species to the genus Metatrogus Britton, 1978 . Metatrogus comprises four other species (Britton 1978; Allsopp 1999; Weir et al. 2019), all from eastern Queensland, Australia and 3200–4300 km south of Palau.</p><p>Matsumoto (2008) based his transfer to the Melolonthina (now Melolonthini) mainly on the wing venation of H. burmeisteri and, presumably, on superficial similarities in the shape of the aedeagi but made no mention of examining any Metatrogus specimens. He curiously added “Britton’s definition of Metatrogus applies very well to this species [ burmeisteri] except for the following points: (1) ‘hypopleuron’ of pronotum, abdomen, pygidium and propygidium without any adpressed white hairs; (2) anterior margin of pronotum rimmed throughout”.</p><p>Comparison of the type series of H. burmeisteri and specimens of the four Australian species of Metatrogus ( M. castaneus Britton, 1978, M. lukei Allsopp, 1999, M. praeceps Britton, 1978, M. septuosus Britton, 1978; specimens in QM) shows that this transfer was incorrect. Amongst other characters, H. burmeisteri has: unmodified, elongate setae (Fig. 9) on the venter, whereas Metatrogus have flattened, white, adpressed setae (Fig. 11); the labrum deeply indented and forming two lobes (Fig. 8), whereas Metatrogus has the labrum indented to less than half length and the indentation is broadly rounded (Fig. 10); the antennal club is of three lamellae, whereas it is of three, five or six lamellae in Metatrogus; the metepisternum narrow (ratio 5:1 to 6:1) and metepimeron small (Fig. 9), whereas the metepisternum is rhomboid and the metepimeron is larger in Metatrogus (Fig. 11); parameres where the apices do not cross over and without a basal backward projection on each (Fig. 19), whereas three of the Metatrogus spp. have the apices of the parameres crossing over, and all have a basal backward projection on each paramere (Britton 1978, figs. 74–82; Allsopp 1999, fig. 3).</p><p>In essence, H. burmeisteri clearly keys to the Rhizotrogini, whereas Metatrogus falls within the Melolonthini (Lacroix 2010, pp. 19–20). No endemic Rhizotrogini are known from Australia (Weir et al. 2019).</p></div>	https://treatment.plazi.org/id/BE00878FFFA9810846BB8883FB3E6691	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Allsopp, Peter G.	Allsopp, Peter G. (2025): Holotrichia burmeisteri Brenske, 1892 and allies (Coleoptera: Scarabaeidae: Melolonthinae: Rhizotrogini): a tale of four genera and four countries. Zootaxa 5631 (1): 166-178, DOI: 10.11646/zootaxa.5631.1.8, URL: https://doi.org/10.11646/zootaxa.5631.1.8
BE00878FFFA2810246BB8A35FA4165A2.text	BE00878FFFA2810246BB8A35FA4165A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cheberdord Allsopp 2025	<div><p>Cheberdord Allsopp, new genus</p><p>urn:lsid:zoobank.org:act: D2AF10B0-FDBE-4D0E-A3E8-1CC24283AEA7</p><p>Type species. Holotrichia burmeisteri Brenske, 1892, hereby designated.</p><p>Description. Body 17.5–23.0 mm long. Colour yellow-brown with head, pronotum, scutellum and profemora brown-red, pronotum may be dark brown; elytra ranging from light yellow-brown to a mahogany brown. Mentum flat. Labrum strongly concave, appearing bilobed, level with anterior face of clypeus. Antennae of 10 antennomeres, club of 3 long lamellae, antennomere 7 may be flattened and leaf-like, club of male about 0.85-1 times length of scape, that of female about 0.5 times as long as scape. Anterior face of clypeus flat, indistinctly punctured; anterior clypeal margin arcuately rounded, reflexed, sometimes weakly but distinctly emarginate medially. Upper clypeus and frons with large, round punctures separated by less than twice their diameters. Pronotum wider than base of elytra, anterior margin broadly, weakly emarginate, posterior margin sinuate with a barely perceptible marginal line, lateral margin crenate, broadly, strongly rounded, anterolateral angle bluntly angulate, posterolateral angle slightly produced, blunt, surface smooth, shining, fine punctures separated by 1–3 times their diameters. Elytra smooth, shining, evenly punctured, strial punctures scarcely separable from other punctures, intervals 1, 3, 5 and 7 feebly convex, each puncture with a small, scarcely visible, white seta; epipleuron with a single row of intermixed long and short setae or with just short setae (0.5 mm long) forming a fringe visible in dorsal view, long setae about 1.0 mm long basally, decreasing in length towards apex. Pygidium nearly triangular, roughly sculptured with elongate punctures scabrous and wrinkled in apical third. Metasternum densely pubescent, punctured, midline complete. Abdomen with sterna 2–5 connate medially, very fine, setigerous punctures present, becoming sparse medially, denser laterally. Protibia tridentate in apical half, anterior spur long, slender, parallel sided, pointed or blunt, slightly shorter than or as long as protarsomere 1. Meso- and metatibiae each with a row of sharp serrations on outer margin next to body, inner spur shorter than or about as long as outer, both pointed. Tarsal claws with a small, sharp tooth near base, directed toward base of claw. Parameres each with a basal backwardly-directed tooth, apex dilated with ventral and dorsal apices rounded and feebly emarginate.</p><p>Differential diagnosis. No contemporary comparison of the Oriental and far eastern Palaearctic genera of the Rhizotrogini exists. As discussed above, the shape of the parameres distinguishes Cheberdord Allsopp, new genus from other genera.</p><p>Etymology. The name is derived from the Palauan word for beetle (https://www.tekinged.com/?lookup=beetle&amp;direction=ep) and is to be treated as feminine.</p><p>Key to species</p><p>1 Anterior margin of clypeus evenly rounded with no trace of median emargination; parameres as in Figs 16, 19–21 ............................................................. Cheberdord burmeisteri (Brenske, 1892), new combination</p><p>- Anterior margin of clypeus distinctly emarginate medially.................................................... 2</p><p>2 Antennomere 7 flattened and leaf-like; epipleuron with a single row of intermixed long and short setae forming a fringe visible in dorsal view, long setae about 1.0 mm long basally, decreasing in length toward apex; parameres as in Fig. 17 ................................................................... Cheberdord dybasi (Gordon, 1979), new combination</p><p>- Antennomere 7 feebly flattened; epipleuron with a single row of short setae visible in dorsal view, setae less than 0.5 mm long basally, decreasing slightly in length toward apex; parameres as in Fig. 18 ................................................................................................. Cheberdord furtiva (Gordon, 1979), new combination</p></div>	https://treatment.plazi.org/id/BE00878FFFA2810246BB8A35FA4165A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Allsopp, Peter G.	Allsopp, Peter G. (2025): Holotrichia burmeisteri Brenske, 1892 and allies (Coleoptera: Scarabaeidae: Melolonthinae: Rhizotrogini): a tale of four genera and four countries. Zootaxa 5631 (1): 166-178, DOI: 10.11646/zootaxa.5631.1.8, URL: https://doi.org/10.11646/zootaxa.5631.1.8
BE00878FFFA3810246BB8950FCFC67ED.text	BE00878FFFA3810246BB8950FCFC67ED.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cheberdord burmeisteri (Brenske 1892) Allsopp 2025	<div><p>Cheberdord burmeisteri (Brenske, 1892), new combination</p><p>(Figs 2–9, 12, 16, 19–22, 24)</p><p>Holotrichia burmeisteri Brenske, 1892: 187 .</p><p>Metatrogus burmeisteri: Matsumoto 2008: 11.</p><p>Lepidiota carolinensis Arrow, 1939: 86, new synonymy .</p><p>Description. Cartwright &amp; Gordon (1971: 280). Dorsum Figs 2–7, 12, 22; aedeagus Figs 16, 19–21.</p><p>Distribution. Palau. Babelthuap: Badeldaob [Babelthuap]; Melokeiok [Melekeok, 7.52°N, 134.60°E]; Ngeremlengui [7.53°N, 134.56°E]; Ogiwal [Nigiwal, 7.57°N 134.62°E; Airhi [Airai, 7.36°N 134.54°E]; Koror [7.34°N 134.48°E]. Guam (intercepted in transit, Fig. 24).</p></div>	https://treatment.plazi.org/id/BE00878FFFA3810246BB8950FCFC67ED	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Allsopp, Peter G.	Allsopp, Peter G. (2025): Holotrichia burmeisteri Brenske, 1892 and allies (Coleoptera: Scarabaeidae: Melolonthinae: Rhizotrogini): a tale of four genera and four countries. Zootaxa 5631 (1): 166-178, DOI: 10.11646/zootaxa.5631.1.8, URL: https://doi.org/10.11646/zootaxa.5631.1.8
BE00878FFFA3810246BB8A9CFE84668A.text	BE00878FFFA3810246BB8A9CFE84668A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cheberdord dybasi (Gordon 1971) Allsopp 2025	<div><p>Cheberdord dybasi (Gordon, 1971), new combination</p><p>(Figs 13, 17)</p><p>Lepidiota dybasi Gordon in Cartwright &amp; Gordon, 1971: 280.</p><p>Description. Cartwright &amp; Gordon (1971: 280). Dorsum Fig. 13; aedeagus Fig. 17.</p><p>Distribution. Palau. Peleliu, including Amaingal Mt [ca. 7°N, 134.2°E] and east and west coasts (Cartwright &amp; Gordon 1971).</p></div>	https://treatment.plazi.org/id/BE00878FFFA3810246BB8A9CFE84668A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Allsopp, Peter G.	Allsopp, Peter G. (2025): Holotrichia burmeisteri Brenske, 1892 and allies (Coleoptera: Scarabaeidae: Melolonthinae: Rhizotrogini): a tale of four genera and four countries. Zootaxa 5631 (1): 166-178, DOI: 10.11646/zootaxa.5631.1.8, URL: https://doi.org/10.11646/zootaxa.5631.1.8
BE00878FFFA0810146BB8CD0FC4760CF.text	BE00878FFFA0810146BB8CD0FC4760CF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cheberdord furtiva (Gordon 1971) Allsopp 2025	<div><p>Cheberdord furtiva (Gordon, 1971), new combination</p><p>(Figs 14, 18)</p><p>Lepidiota furtiva Gordon in Cartwright &amp; Gordon, 1971: 282.</p><p>Description. Cartwright &amp; Gordon (1971: 282). Dorsum Fig. 14; aedeagus Fig. 18.</p><p>Distribution. Palau. Ngurukdapel [Nguruktabel, 7.26°N, 134.41°E];Auluptagel [Ulebschel, 7.31°N, 134.48°E]; Peleliu, Amaingal Mt [ca. 7°N, 134.2°E]; Koror; North Central Islet.</p></div>	https://treatment.plazi.org/id/BE00878FFFA0810146BB8CD0FC4760CF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Allsopp, Peter G.	Allsopp, Peter G. (2025): Holotrichia burmeisteri Brenske, 1892 and allies (Coleoptera: Scarabaeidae: Melolonthinae: Rhizotrogini): a tale of four genera and four countries. Zootaxa 5631 (1): 166-178, DOI: 10.11646/zootaxa.5631.1.8, URL: https://doi.org/10.11646/zootaxa.5631.1.8
