taxonID	type	description	language	source
05796986007AAFC5A591AD66FA9BC2F1.taxon	diagnosis	Diagnosis. Sibon bevridgelyi is placed in the genus Sibon based on phylogenetic evidence (Fig. 3) and on having the labial beneath primary temporal conspicuosly higher than other labials. The species differs from all described species of Sibon based on the following combination of characters: (1) 15 / 15 / 15 smooth dorsals with enlarged vertebral row (1.3 - 1.7 times as wide as adjacent rows); (2) seven supralabials with 4 th and 5 th contacting orbit or eight supralabials with 5 th and 6 th contacting orbit; (3) one pair of infralabials in contact behind symphysial; (4) postmental absent; (5) 175 - 193 ventrals in males, 193 in the single female; (6) 80 - 94 divided subcaudals in males, 98 in the single female; (7) dorsal and ventral ground color pale yellow with or without irregular black bands, and with a black stippled disruptive pattern of irregular rusty to reddish brown blotches that are separated from each other by light interspaces (Figs 6, 2 b, c); bands incomplete and stippling not prominent or absent on ventral surfaces; head heavily speckled or blotched with black or rusty pigment; eyes light slate blue to pale goldenrod with black speckles and reticulations; (8) 349 - 732 mm SVL in males, 786 mm in the single female; (9) 124 - 268 mm TL in males, 204 mm in the single female.	en	Arteaga, Alejandro, Salazar-Valenzuela, David, Mebert, Konrad, Penafiel, Nicolas, Aguiar, Gabriela, Sa ́ nchez-Nivicela, Juan C., Pyron, R. Alexander, Colston, Timothy J., Cisneros-Heredia, Diego F., Yanez-Munoz, Mario H., Venegas, Pablo J., Guayasamin, Juan M., Torres-Carvajal, Omar (2018): Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru. ZooKeys 766: 79-147, DOI: http://dx.doi.org/10.3897/zookeys.766.24523, URL: http://dx.doi.org/10.3897/zookeys.766.24523
05796986007AAFC5A591AD66FA9BC2F1.taxon	description	Description of holotype. Adult male, SVL 602 mm, tail length 186 mm (31 % SVL); head length 20.9 mm (3 % SVL) from tip of snout to commissure of mouth; head width 12.4 mm (59 % head length) taken at broadest point; snout-orbit distance 21 mm; head distinct from neck; snout short, blunt in dorsal and lateral outline; rostral 3.5 mm wide, broader than high; internasals 1.9 mm wide, broader than long; prefrontals 4.4 mm wide, longer than broad, entering orbit; supraocular 4.4 mm long, longer than broad; frontal 4.1 mm long, pentagonal and rounded, in contact with prefrontals, supraoculars, and parietals; parietals 7.7 mm long, longer than broad; nasal weakly divided, in contact with first three supralabials, loreal, prefrontal, internasal, and rostral; loreal 3.7 mm long, longer than high, entering the orbit; eye diameter 3.9 mm; pupil semi-elliptical; no preocular; two postoculars; temporals 1 + 3 on the right side, 2 + 3 on the left side; eight supralabials with 5 th and 6 th contacting orbit on the right side, seven supralabials with 4 th and 5 th contacting orbit on the left side; symphysial separated from chinshields by the first pair of infralabials; nine infralabials, 1 - 7 contacting chinshields; anterior pair of chinshields broader than long, posterior pair longer than broad; dorsal scales in 15 / 15 / 15 rows, smooth, without apical pits; 184 ventrals; 80 divided subcaudals; cloacal plate single.	en	Arteaga, Alejandro, Salazar-Valenzuela, David, Mebert, Konrad, Penafiel, Nicolas, Aguiar, Gabriela, Sa ́ nchez-Nivicela, Juan C., Pyron, R. Alexander, Colston, Timothy J., Cisneros-Heredia, Diego F., Yanez-Munoz, Mario H., Venegas, Pablo J., Guayasamin, Juan M., Torres-Carvajal, Omar (2018): Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru. ZooKeys 766: 79-147, DOI: http://dx.doi.org/10.3897/zookeys.766.24523, URL: http://dx.doi.org/10.3897/zookeys.766.24523
05796986007AAFC5A591AD66FA9BC2F1.taxon	distribution	Distribution. Northwestern Peru in the department of Piura, and southwestern Ecuador in the provinces of Azuay, Chimborazo, El Oro, Guayas, Los Rios and Manabi at elevations between 5 and 1206 m (Fig. 8).	en	Arteaga, Alejandro, Salazar-Valenzuela, David, Mebert, Konrad, Penafiel, Nicolas, Aguiar, Gabriela, Sa ́ nchez-Nivicela, Juan C., Pyron, R. Alexander, Colston, Timothy J., Cisneros-Heredia, Diego F., Yanez-Munoz, Mario H., Venegas, Pablo J., Guayasamin, Juan M., Torres-Carvajal, Omar (2018): Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru. ZooKeys 766: 79-147, DOI: http://dx.doi.org/10.3897/zookeys.766.24523, URL: http://dx.doi.org/10.3897/zookeys.766.24523
05796986007AAFC5A591AD66FA9BC2F1.taxon	etymology	Etymology. The specific epithet honors the late Prof. Beverly S. Ridgely, life-long birder and conservationist, and father of Robert S. Ridgely, well known in Ecuadorian ornithological circles and co-author of The Birds of Ecuador. Though he never got to visit Buenaventura, from afar Bev continued to delight in the conservation successes of Fundacion Jocotoco, which now owns and manages one of the few protected areas where the Vulnerable Sibon bevridgelyi is known to occur.	en	Arteaga, Alejandro, Salazar-Valenzuela, David, Mebert, Konrad, Penafiel, Nicolas, Aguiar, Gabriela, Sa ́ nchez-Nivicela, Juan C., Pyron, R. Alexander, Colston, Timothy J., Cisneros-Heredia, Diego F., Yanez-Munoz, Mario H., Venegas, Pablo J., Guayasamin, Juan M., Torres-Carvajal, Omar (2018): Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru. ZooKeys 766: 79-147, DOI: http://dx.doi.org/10.3897/zookeys.766.24523, URL: http://dx.doi.org/10.3897/zookeys.766.24523
E171318E6131E49E2B33DE6D11DDCBE9.taxon	diagnosis	Diagnosis. Dipsas bobridgelyi is placed in the genus Dipsas based on phylogenetic evidence (Fig. 3), and the absence of a labial that is noticeably higher than other labials and in contact with the postocular, primary, and secondary temporals. The species differs from all described species of Dipsas based on the following combination of characters: (1) 15 / 15 / 15 smooth dorsals with enlarged vertebral row (2.1 - 2.2 times as wide as adjacent rows); (2) loreal and prefrontal in contact with orbit; (3) 9 supralabials with 4 th and 5 th contacting orbit; (4) one pair of infralabials in contact behind symphysial; (5) 180 - 201 ventrals in males, 178 - 184 in females; (6) 95 - 117 divided subcaudals in males, 96 - 98 in females; (7) dorsal and ventral color made up of 30 - 35 bold black body rings (up to 7 - 12 vertebral scales long) separated from each other by narrow (up to 3 - 4 vertebral scales long) dingy white interspaces; dorsal aspect of interspaces heavily speckled with rusty and black pigment; ventral surfaces lacking speckling; ground color of head dingy white with various degrees of scattered black pigment that coalesce on the top of the head, and various degrees of rusty speckling concentrated on the snout, nape and sides of the head; iris rich dark brown; (8) 372 - 478 mm SVL in males, 286 - 404 mm in females; (9) 158 - 212 mm TL in males, 117 - 158 mm in females.	en	Arteaga, Alejandro, Salazar-Valenzuela, David, Mebert, Konrad, Penafiel, Nicolas, Aguiar, Gabriela, Sa ́ nchez-Nivicela, Juan C., Pyron, R. Alexander, Colston, Timothy J., Cisneros-Heredia, Diego F., Yanez-Munoz, Mario H., Venegas, Pablo J., Guayasamin, Juan M., Torres-Carvajal, Omar (2018): Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru. ZooKeys 766: 79-147, DOI: http://dx.doi.org/10.3897/zookeys.766.24523, URL: http://dx.doi.org/10.3897/zookeys.766.24523
E171318E6131E49E2B33DE6D11DDCBE9.taxon	description	Description of holotype. Adult male, SVL 372 mm, tail length 158 mm (43 % SVL); head length 15.1 mm (4 % SVL) from tip of snout to commissure of mouth; head width 8.1 mm (54 % head length) taken at broadest point; snout-orbit distance 4.3 mm; head distinct from neck; snout short, blunt in dorsal and lateral outline; rostral 2.4 mm wide, broader than high; internasals 2.3 mm wide, broader than long; prefrontals 2.5 mm wide, longer than broad and contacting orbit; supraocular 3.2 mm long, longer than broad; frontal 3.9 mm long, hexagonal, in contact with prefrontals, supraoculars, and parietals; parietals 4.7 mm long, longer than broad; nasal divided, in contact with first three supralabials, loreal, prefrontal, internasal, and rostral; loreal 1.8 mm long, slightly higher than long, entering the orbit; eye diameter 2.7 mm; pupil semi-elliptical; no preocular; two postoculars; temporals 2 + 3; nine supralabials, 4 th and 5 th contacting orbit; symphysial separated from chinshields by the first pair of infralabials; 13 infralabials, 1 - 7 contacting chinshields; anterior pair of chinshields longer than broad, posterior pair broader than long; dorsal scales in 15 / 15 / 15 rows, smooth, without apical pits; 182 ventrals; 101 divided subcaudals; cloacal plate single.	en	Arteaga, Alejandro, Salazar-Valenzuela, David, Mebert, Konrad, Penafiel, Nicolas, Aguiar, Gabriela, Sa ́ nchez-Nivicela, Juan C., Pyron, R. Alexander, Colston, Timothy J., Cisneros-Heredia, Diego F., Yanez-Munoz, Mario H., Venegas, Pablo J., Guayasamin, Juan M., Torres-Carvajal, Omar (2018): Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru. ZooKeys 766: 79-147, DOI: http://dx.doi.org/10.3897/zookeys.766.24523, URL: http://dx.doi.org/10.3897/zookeys.766.24523
E171318E6131E49E2B33DE6D11DDCBE9.taxon	distribution	Distribution. Foothills of the southwestern Ecuadorian Andes in the provinces of Azuay and El Oro, and northwestern Peruvian Andes in the department of Tumbes, at elevations between 39 and 572 m (Fig. 4).	en	Arteaga, Alejandro, Salazar-Valenzuela, David, Mebert, Konrad, Penafiel, Nicolas, Aguiar, Gabriela, Sa ́ nchez-Nivicela, Juan C., Pyron, R. Alexander, Colston, Timothy J., Cisneros-Heredia, Diego F., Yanez-Munoz, Mario H., Venegas, Pablo J., Guayasamin, Juan M., Torres-Carvajal, Omar (2018): Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru. ZooKeys 766: 79-147, DOI: http://dx.doi.org/10.3897/zookeys.766.24523, URL: http://dx.doi.org/10.3897/zookeys.766.24523
E171318E6131E49E2B33DE6D11DDCBE9.taxon	etymology	Etymology. This species is named in honor of Dr. Robert " Bob " S. Ridgely, a leading ornithologist and distinguished conservationist who has dedicated almost 50 years of his life to the study and conservation of birds and biodiversity across Latin America. Bob is the President of Rainforest Trust and for the past twenty years has been a major driver of conservation in Ecuador through Fundacion Jocotoco, which he helped establish twenty years ago. In 1980, Bob visited the type locality of Dipsas bobridgelyi (Buenaventura, meaning " good fortune "), now known to be a key area for the conservation of biodiversity. Bob embarked on conservation and worked diligently to raise funds through Rainforest Trust for the past 18 years to purchase private properties and establish what is now the Reserva Buenaventura of Fundacion Jocotoco.	en	Arteaga, Alejandro, Salazar-Valenzuela, David, Mebert, Konrad, Penafiel, Nicolas, Aguiar, Gabriela, Sa ́ nchez-Nivicela, Juan C., Pyron, R. Alexander, Colston, Timothy J., Cisneros-Heredia, Diego F., Yanez-Munoz, Mario H., Venegas, Pablo J., Guayasamin, Juan M., Torres-Carvajal, Omar (2018): Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru. ZooKeys 766: 79-147, DOI: http://dx.doi.org/10.3897/zookeys.766.24523, URL: http://dx.doi.org/10.3897/zookeys.766.24523
4807FA902BB26EA2A026CB0F1D905251.taxon	diagnosis	Diagnosis. Dipsas georgejetti is placed in the genus Dipsas based on phylogenetic evidence (Fig. 3) and the absence of a labial that is noticeably higher than other labials and in contact with the postocular, primary and secondary temporals. The species differs from all described species of Dipsas based on the following combination of characters: (1) 15 / 15 / 15 smooth dorsals with a slightly enlarged vertebral row (1 - 1.4 times as wide as adjacent rows); (2) loreal and prefrontal in contact with orbit; (3) 7 supralabials with 4 th and 5 th (3 th- 5 th in DHMECN 11646) contacting orbit; (4) no infralabials in contact behind symphysial; (5) 172 - 180 ventrals in males, 177 in one female; (6) 69 - 86 divided subcaudals in males, 58 in one female; (7) dorsal ground color light sandy brown with a pattern of 53 - 61 drab to brown black-edged middorsal blotches that are wider (6 - 7 vertebral scales long) and solid down to the edges of the ventrals on the first one third of the body, but becoming narrower (1 - 3 vertebral scales long) and broken up laterally towards the tail; interspaces finely speckled with brown pigment; ground color of the head light sandy brown with bold dark brown to black irregular blotches scattered on head plates and edging supralabials; ventral surfaces sandy brown with fine black speckling; iris sandy brown with dense dark brown speckling; (8) 270 - 711 mm SVL in males, 856 mm in one female; (9) 87 - 170 mm TL in males, 150 mm in one female.	en	Arteaga, Alejandro, Salazar-Valenzuela, David, Mebert, Konrad, Penafiel, Nicolas, Aguiar, Gabriela, Sa ́ nchez-Nivicela, Juan C., Pyron, R. Alexander, Colston, Timothy J., Cisneros-Heredia, Diego F., Yanez-Munoz, Mario H., Venegas, Pablo J., Guayasamin, Juan M., Torres-Carvajal, Omar (2018): Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru. ZooKeys 766: 79-147, DOI: http://dx.doi.org/10.3897/zookeys.766.24523, URL: http://dx.doi.org/10.3897/zookeys.766.24523
4807FA902BB26EA2A026CB0F1D905251.taxon	description	Description of holotype. Adult male, SVL 315 mm, TL 87 mm (28 % SVL); head length 13.6 mm (4 % SVL) from tip of snout to commissure of mouth; head width 8.4 mm (62 % head length) taken at broadest point; snout-orbit distance 3.5 mm; head distinct from neck; snout short, blunt in dorsal and lateral outline; rostral 2.0 mm wide, broader than high; internasals 1.7 mm wide, broader than long; prefrontals 2.5 mm wide, longer than broad and contacting orbit; supraocular 3.4 mm long, longer than broad; frontal 3.3 mm long, pentagonal, in contact with prefrontals, supraoculars, and parietals; parietals 5.5 mm long, longer than broad; nasal divided, in contact with first two supralabials, loreal, prefrontal, internasal, and rostral; loreal 1.7 mm long, slightly higher than long, entering orbit; eye diameter 2.8 mm; pupil semi-elliptical; no preocular; two postoculars; temporals 2 + 2; seven supralabials, 4 th and 5 th contacting orbit; symphysial in contact with first pair of chinshields; nine infralabials, 1 - 6 contacting chinshields; anterior pair of chinshields longer than broad, posterior pair broader than long; dorsal scales in 15 / 15 / 15 rows, smooth, without apical pits; 178 ventrals; 69 divided subcaudals; cloacal plate single.	en	Arteaga, Alejandro, Salazar-Valenzuela, David, Mebert, Konrad, Penafiel, Nicolas, Aguiar, Gabriela, Sa ́ nchez-Nivicela, Juan C., Pyron, R. Alexander, Colston, Timothy J., Cisneros-Heredia, Diego F., Yanez-Munoz, Mario H., Venegas, Pablo J., Guayasamin, Juan M., Torres-Carvajal, Omar (2018): Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru. ZooKeys 766: 79-147, DOI: http://dx.doi.org/10.3897/zookeys.766.24523, URL: http://dx.doi.org/10.3897/zookeys.766.24523
4807FA902BB26EA2A026CB0F1D905251.taxon	distribution	Distribution. Deciduous and semideciduous forests along the central Pacific coast in Ecuador in the provinces of Manabi and Guayas, at elevations between 5 and 317 m (Fig. 5).	en	Arteaga, Alejandro, Salazar-Valenzuela, David, Mebert, Konrad, Penafiel, Nicolas, Aguiar, Gabriela, Sa ́ nchez-Nivicela, Juan C., Pyron, R. Alexander, Colston, Timothy J., Cisneros-Heredia, Diego F., Yanez-Munoz, Mario H., Venegas, Pablo J., Guayasamin, Juan M., Torres-Carvajal, Omar (2018): Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru. ZooKeys 766: 79-147, DOI: http://dx.doi.org/10.3897/zookeys.766.24523, URL: http://dx.doi.org/10.3897/zookeys.766.24523
4807FA902BB26EA2A026CB0F1D905251.taxon	etymology	Etymology. The specific name georgejetti honors George Jett, who has been a long-time donor to Rainforest Trust and has supported the reserves of Fundacion Jocotoco in Ecuador. He is an international traveler with a passion for reptiles, amphibians, and birds.	en	Arteaga, Alejandro, Salazar-Valenzuela, David, Mebert, Konrad, Penafiel, Nicolas, Aguiar, Gabriela, Sa ́ nchez-Nivicela, Juan C., Pyron, R. Alexander, Colston, Timothy J., Cisneros-Heredia, Diego F., Yanez-Munoz, Mario H., Venegas, Pablo J., Guayasamin, Juan M., Torres-Carvajal, Omar (2018): Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru. ZooKeys 766: 79-147, DOI: http://dx.doi.org/10.3897/zookeys.766.24523, URL: http://dx.doi.org/10.3897/zookeys.766.24523
5E40BCBB394FC41BF1550480EDEEB597.taxon	diagnosis	Diagnosis. Dipsas oswaldobaezi is placed in the genus Dipsas based on phylogenetic evidence (Fig. 3) and the absence of a labial that is noticeably higher than other labials and in contact with the postocular, primary and secondary temporals. The species differs from all described species of Dipsas based on the following combination of characters: (1) 15 / 15 / 15 smooth dorsals with a slightly enlarged vertebral row (1 - 1.2 times as wide as adjacent rows); (2) loreal and prefrontal in contact with orbit; (3) six supralabials with 3 rd and 4 th contacting orbit; (4) no infralabials in contact behind symphysial; (5) 163 - 179 ventrals in males, 177 - 179 in females; (6) 68 - 70 divided subcaudals in males, 65 - 66 in females; (7) dorsal ground color light sandy brown with a pattern of 55 - 63 drab to brown black-edged middorsal blotches that are wider (7 - 9 vertebral scale rows) and solid down to the edges of the ventrals on the first one third of the body, but becoming narrower (1 - 3 vertebral scales long) and broken up laterally towards the tail; interspaces finely speckled with brown pigment; ground color of the head light sandy brown with a thin light cream nuchal collar and bold dark brown to black irregular blotches scattered on head plates and edging supralabials; ventral surfaces sandy brown with fine black speckling (Fig. 13 b); iris sandy brown with dense dark brown speckling; (8) 277 - 348 mm SVL in males, 407 - 428 mm in females; (9) 85 - 114 mm TL in males, 110 - 122 mm in females.	en	Arteaga, Alejandro, Salazar-Valenzuela, David, Mebert, Konrad, Penafiel, Nicolas, Aguiar, Gabriela, Sa ́ nchez-Nivicela, Juan C., Pyron, R. Alexander, Colston, Timothy J., Cisneros-Heredia, Diego F., Yanez-Munoz, Mario H., Venegas, Pablo J., Guayasamin, Juan M., Torres-Carvajal, Omar (2018): Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru. ZooKeys 766: 79-147, DOI: http://dx.doi.org/10.3897/zookeys.766.24523, URL: http://dx.doi.org/10.3897/zookeys.766.24523
5E40BCBB394FC41BF1550480EDEEB597.taxon	description	Description of holotype. Adult female, SVL 277 mm, tail length 85 mm (31 % SVL); head length 9.5 mm (3.4 % SVL) from tip of snout to commissure of mouth; head width 7.3 mm (76 % head length) taken at broadest point; snout-orbit distance 3.3 mm; head distinct from neck; snout short, blunt in dorsal and lateral outline; rostral 2.1 mm wide, broader than high; internasals 1.2 mm wide, broader than long; prefrontals 2.2 mm wide, slightly broader than long and contacting orbit; supraocular 2.6 mm long, longer than broad; frontal 2.9 mm long, pentagonal, in contact with prefrontals, supraoculars, and parietals; parietals 4.2 mm long, longer than broad; nasal not divided, in contact with first supralabial, loreal, prefrontal, internasal, and rostral; loreal 1.3 mm long, longer than high, entering orbit; eye diameter 2.2 mm; pupil semi-elliptical; no preocular; two postoculars; temporals 2 + 2; 6 supralabials, 3 rd and 4 th contacting orbit; symphysial separated from chinshields by the first pair of infralabials; 9 / 8 (right / left) infralabials, 1 - 6 / 1 - 5 contacting chinshields; both pairs of chinshields longer than broad; dorsal scales in 15 / 15 / 15 rows, smooth, without apical pits; 179 ventrals; 70 divided subcaudals; cloacal plate single.	en	Arteaga, Alejandro, Salazar-Valenzuela, David, Mebert, Konrad, Penafiel, Nicolas, Aguiar, Gabriela, Sa ́ nchez-Nivicela, Juan C., Pyron, R. Alexander, Colston, Timothy J., Cisneros-Heredia, Diego F., Yanez-Munoz, Mario H., Venegas, Pablo J., Guayasamin, Juan M., Torres-Carvajal, Omar (2018): Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru. ZooKeys 766: 79-147, DOI: http://dx.doi.org/10.3897/zookeys.766.24523, URL: http://dx.doi.org/10.3897/zookeys.766.24523
5E40BCBB394FC41BF1550480EDEEB597.taxon	distribution	Distribution. Deciduous and semideciduous lowland to lower montane forests and dry lowland shrublands in southwestern Ecuador (provinces of Loja and El Oro) and northwestern Peru (department of Tumbes), at elevation between 39 and 1289 m (Fig. 5).	en	Arteaga, Alejandro, Salazar-Valenzuela, David, Mebert, Konrad, Penafiel, Nicolas, Aguiar, Gabriela, Sa ́ nchez-Nivicela, Juan C., Pyron, R. Alexander, Colston, Timothy J., Cisneros-Heredia, Diego F., Yanez-Munoz, Mario H., Venegas, Pablo J., Guayasamin, Juan M., Torres-Carvajal, Omar (2018): Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru. ZooKeys 766: 79-147, DOI: http://dx.doi.org/10.3897/zookeys.766.24523, URL: http://dx.doi.org/10.3897/zookeys.766.24523
5E40BCBB394FC41BF1550480EDEEB597.taxon	etymology	Etymology. The specific name oswaldobaezi honors Dr. Oswaldo Baez, a renowned Ecuadorian biologist and researcher who has dedicated his life to the teaching of science, scientific thinking, and the conservation of nature. Oswaldo Baez has played a major role in science education in Ecuador through many popular science articles and books.	en	Arteaga, Alejandro, Salazar-Valenzuela, David, Mebert, Konrad, Penafiel, Nicolas, Aguiar, Gabriela, Sa ́ nchez-Nivicela, Juan C., Pyron, R. Alexander, Colston, Timothy J., Cisneros-Heredia, Diego F., Yanez-Munoz, Mario H., Venegas, Pablo J., Guayasamin, Juan M., Torres-Carvajal, Omar (2018): Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru. ZooKeys 766: 79-147, DOI: http://dx.doi.org/10.3897/zookeys.766.24523, URL: http://dx.doi.org/10.3897/zookeys.766.24523
5096F079C40CAF2380FAF36B53016546.taxon	diagnosis	Diagnosis. Dipsas klebbai is placed in the genus Dipsas based on phylogenetic evidence (Fig. 3), and the absence of a labial that is noticeably higher than other labials and in contact with the postocular, primary and secondary temporals. The species differs from all described species of Dipsas based on the following combination of characters: (1) 15 / 15 / 15 smooth dorsals with enlarged vertebral row (1.5 - 1.8 times as wide as adjacent rows); (2) one loreal and one preocular in contact with orbit; (3) 9 - 11 supralabials with (usually) 4 th to 6 th contacting orbit; (4) one pair of infralabials in contact behind symphysial; (5) 181 - 201 ventrals in males, 187 - 194 in females; (6) 99 - 123 divided subcaudals in males, 98 - 106 in females; (7) dorsal and ventral ground color light brown with various degrees of fine black speckling and 27 - 36 dark brown to black, cream-edged oblong blotches that are longer that interspaces and become smaller towards the tail (Fig. 2 m, n); on first half of body, the dark bands meet ventrally to form full body rings; on second half they fail to meet ventrally; head black with different degrees of whitish edging on the labial scales, and a thin (1 - 2 scales long) cream to light brown irregular nuchal collar; dorsal blotches usually incomplete ventrally, extending far onto ventrals and occasionally fusing midventrally; cream edges of neighboring blotches fused in first 6 - 9 blotches; (8) 401 - 749 mm SVL in males, 525 - 630 mm in females; (9) 169 - 330 mm TL in males, 209 - 240 mm in females.	en	Arteaga, Alejandro, Salazar-Valenzuela, David, Mebert, Konrad, Penafiel, Nicolas, Aguiar, Gabriela, Sa ́ nchez-Nivicela, Juan C., Pyron, R. Alexander, Colston, Timothy J., Cisneros-Heredia, Diego F., Yanez-Munoz, Mario H., Venegas, Pablo J., Guayasamin, Juan M., Torres-Carvajal, Omar (2018): Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru. ZooKeys 766: 79-147, DOI: http://dx.doi.org/10.3897/zookeys.766.24523, URL: http://dx.doi.org/10.3897/zookeys.766.24523
5096F079C40CAF2380FAF36B53016546.taxon	description	Description of holotype. Adult male, SVL 608 mm, tail length 262 mm (43 % SVL); head length 20.3 mm (3 % SVL) from tip of snout to commissure of mouth; head width 12.7 mm (62 % head length) taken at broadest point; snout-orbit distance 5.4 mm; head distinct from neck; snout short, blunt in dorsal and lateral outline; rostral 4.0 mm wide, broader than high; internasals 2.6 mm wide, as broad as long; prefrontals 3.9 mm wide, broader than long, excluded from entering orbit by preocular; supraocular 4.3 mm long, broader than long; frontal 4.5 mm long, hexagonal, in contact with prefrontals, supraoculars, and parietals; parietals 6.6 mm long, longer than broad; nasal divided, in contact with first two supralabials, loreal, prefrontal, internasal, and rostral; loreal 2.6 mm long, slightly longer than high, entering orbit; eye diameter 4.5 mm; pupil semi-elliptical; one preocular; two postoculars; temporals 2 + 2; ten supralabials, 5 th and 6 th contacting orbit; symphysial separated from chinshields by the first pair of infralabials; 14 infralabials, 2 - 7 contacting chinshields; anterior pair of chinshields longer than broad, posterior pair broader than long; dorsal scales in 15 / 15 / 15 rows, smooth, without apical pits; 188 ventrals; 116 divided subcaudals; cloacal plate single.	en	Arteaga, Alejandro, Salazar-Valenzuela, David, Mebert, Konrad, Penafiel, Nicolas, Aguiar, Gabriela, Sa ́ nchez-Nivicela, Juan C., Pyron, R. Alexander, Colston, Timothy J., Cisneros-Heredia, Diego F., Yanez-Munoz, Mario H., Venegas, Pablo J., Guayasamin, Juan M., Torres-Carvajal, Omar (2018): Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru. ZooKeys 766: 79-147, DOI: http://dx.doi.org/10.3897/zookeys.766.24523, URL: http://dx.doi.org/10.3897/zookeys.766.24523
5096F079C40CAF2380FAF36B53016546.taxon	distribution	Distribution. Endemic to the eastern slopes of the Ecuadorian Andes in the provinces of Napo and Sucumbios at elevations between 1246 and 2120 m (Fig. 4).	en	Arteaga, Alejandro, Salazar-Valenzuela, David, Mebert, Konrad, Penafiel, Nicolas, Aguiar, Gabriela, Sa ́ nchez-Nivicela, Juan C., Pyron, R. Alexander, Colston, Timothy J., Cisneros-Heredia, Diego F., Yanez-Munoz, Mario H., Venegas, Pablo J., Guayasamin, Juan M., Torres-Carvajal, Omar (2018): Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru. ZooKeys 766: 79-147, DOI: http://dx.doi.org/10.3897/zookeys.766.24523, URL: http://dx.doi.org/10.3897/zookeys.766.24523
5096F079C40CAF2380FAF36B53016546.taxon	etymology	Etymology. Named after Casey Klebba, in recognition of his appreciation of and passion for Andean wildlife, and his invaluable support of AA's field expeditions to remote areas of Ecuador. After a visit to Peru in 2011, Casey became an active supporter of conservation and scientific projects in Ecuador.	en	Arteaga, Alejandro, Salazar-Valenzuela, David, Mebert, Konrad, Penafiel, Nicolas, Aguiar, Gabriela, Sa ́ nchez-Nivicela, Juan C., Pyron, R. Alexander, Colston, Timothy J., Cisneros-Heredia, Diego F., Yanez-Munoz, Mario H., Venegas, Pablo J., Guayasamin, Juan M., Torres-Carvajal, Omar (2018): Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru. ZooKeys 766: 79-147, DOI: http://dx.doi.org/10.3897/zookeys.766.24523, URL: http://dx.doi.org/10.3897/zookeys.766.24523
7DEB677F8A6F967EBB6789ACA937054E.taxon	diagnosis	Diagnosis. Dipsas palmeri differs from all described species of Dipsas based on the following combination of characters: (1) 15 / 15 / 15 smooth dorsals with enlarged vertebral row; (2) one loreal and one preocular in contact with orbit; (3) 8 - 10 supralabials with (usually) 4 th to 6 th contacting orbit; (4) one pair of infralabials in contact behind symphysial; (5) 172 - 202 ventrals in males, 181 - 200 in females; (6) 91 - 118 divided subcaudals in males, 86 - 102 in females; (7) dorsal and ventral ground color light brown with various degrees of fine black speckling and with 32 - 41 brown to blackish, white-edged circular blotches that are longer than interspaces in the first half of the body, but shorter in the second half (Figs 1 r, s); adult head gray with different degrees of whitish edging on the labial scales, and a thin (1 - 2 scales long) white to light grayish brown irregular parietal collar; dorsal blotches incomplete ventrally, extending marginally onto ventrals but not fusing midventrally; (8) 215 - 907 mm SVL in males, 642 - 1187 mm in females; (9) 78 - 390 mm TL in males, 246 - 298 mm in females.	en	Arteaga, Alejandro, Salazar-Valenzuela, David, Mebert, Konrad, Penafiel, Nicolas, Aguiar, Gabriela, Sa ́ nchez-Nivicela, Juan C., Pyron, R. Alexander, Colston, Timothy J., Cisneros-Heredia, Diego F., Yanez-Munoz, Mario H., Venegas, Pablo J., Guayasamin, Juan M., Torres-Carvajal, Omar (2018): Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru. ZooKeys 766: 79-147, DOI: http://dx.doi.org/10.3897/zookeys.766.24523, URL: http://dx.doi.org/10.3897/zookeys.766.24523
7DEB677F8A6F967EBB6789ACA937054E.taxon	distribution	Distribution. Eastern slopes of the Ecuadorian and Peruvian Andes south of the Jatunyacu-Napo river valley in Ecuador and north of the Huancabamba depression at elevations between 1211 and 2282 m (Fig. 4).	en	Arteaga, Alejandro, Salazar-Valenzuela, David, Mebert, Konrad, Penafiel, Nicolas, Aguiar, Gabriela, Sa ́ nchez-Nivicela, Juan C., Pyron, R. Alexander, Colston, Timothy J., Cisneros-Heredia, Diego F., Yanez-Munoz, Mario H., Venegas, Pablo J., Guayasamin, Juan M., Torres-Carvajal, Omar (2018): Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru. ZooKeys 766: 79-147, DOI: http://dx.doi.org/10.3897/zookeys.766.24523, URL: http://dx.doi.org/10.3897/zookeys.766.24523
88FB43CB53DB280639129EFCDF2D17D3.taxon	diagnosis	Diagnosis. Dipsas peruana differs from all described species of Dipsas based on the following combination of characters: (1) 15 / 15 / 15 smooth dorsals with moderately enlarged vertebral row; (2) one loreal and one preocular in contact with orbit; (3) 8 - 9 supralabials with 4 - 6 or 3 - 5 contacting orbit; (4) one pair of infralabials in contact behind symphysial; (5) 177 - 200 ventrals in males, 180 - 203 in females; (6) 75 - 127 divided subcaudals in males, 79 - 105 in females; (7) dorsal and ventral ground color brown to dark brown (light brown in juveniles) with 33 - 43 blackish brown to complete black, white to cream edged circular to vertically elliptical blotches that are longer than interspaces; head dark brown with dingy cream reticulations and different degrees of whitish edging on the labial scales, and a thin (1 - 3 scales long) white to light grayish brown irregular nuchal collar; dorsal blotches extending marginally onto ventrals and rarely fusing midventrally; (8) 199 mm SVL in males, 610 - 725 mm in females; (9) 85 mm TL in males, 155 - 241 mm in females.	en	Arteaga, Alejandro, Salazar-Valenzuela, David, Mebert, Konrad, Penafiel, Nicolas, Aguiar, Gabriela, Sa ́ nchez-Nivicela, Juan C., Pyron, R. Alexander, Colston, Timothy J., Cisneros-Heredia, Diego F., Yanez-Munoz, Mario H., Venegas, Pablo J., Guayasamin, Juan M., Torres-Carvajal, Omar (2018): Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru. ZooKeys 766: 79-147, DOI: http://dx.doi.org/10.3897/zookeys.766.24523, URL: http://dx.doi.org/10.3897/zookeys.766.24523
88FB43CB53DB280639129EFCDF2D17D3.taxon	distribution	Distribution. Eastern slopes of the Peruvian and Bolivian Andes south of the Huancabamba depression at elevations between 1279 and 2671 m (Fig. 4).	en	Arteaga, Alejandro, Salazar-Valenzuela, David, Mebert, Konrad, Penafiel, Nicolas, Aguiar, Gabriela, Sa ́ nchez-Nivicela, Juan C., Pyron, R. Alexander, Colston, Timothy J., Cisneros-Heredia, Diego F., Yanez-Munoz, Mario H., Venegas, Pablo J., Guayasamin, Juan M., Torres-Carvajal, Omar (2018): Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru. ZooKeys 766: 79-147, DOI: http://dx.doi.org/10.3897/zookeys.766.24523, URL: http://dx.doi.org/10.3897/zookeys.766.24523
82CA92F6A92ABB2757CEAA1E2EFC2E47.taxon	diagnosis	Diagnosis. Dipsas latifrontalis differs from all described species of Dipsas based on the following combination of characters: (1) 15 / 15 / 15 smooth dorsals with moderately enlarged vertebral row; (2) one loreal and one preocular in contact with orbit; (3) 8 - 10 supralabials with 3 rd to 6 th contacting orbit; (4) one pair of infralabials in contact behind symphysial; (5) 192 ventrals in one male (CVULA 7883), 194 in the female holotype; (6) 109 divided subcaudals in the single male, 95 in the female holotype; (7) dorsal and ventral ground color bronze (light brown in juveniles) with 32 - 36 dark reddish brown to black, circular to vertically elliptical blotches that are longer than interspaces and white to cream edged on first half of body; head grayish brown to black with different degrees of whitish edging on the labial scales, and with or without a thin (1 - 2 scales long) dingy white irregular nuchal collar; dorsal blotches extending marginally onto ventrals and occasionally fusing on the anterior part of the body; (8) 800 mm SVL in the holotype female; (9) 220 mm TL in the holotype female.	en	Arteaga, Alejandro, Salazar-Valenzuela, David, Mebert, Konrad, Penafiel, Nicolas, Aguiar, Gabriela, Sa ́ nchez-Nivicela, Juan C., Pyron, R. Alexander, Colston, Timothy J., Cisneros-Heredia, Diego F., Yanez-Munoz, Mario H., Venegas, Pablo J., Guayasamin, Juan M., Torres-Carvajal, Omar (2018): Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru. ZooKeys 766: 79-147, DOI: http://dx.doi.org/10.3897/zookeys.766.24523, URL: http://dx.doi.org/10.3897/zookeys.766.24523
82CA92F6A92ABB2757CEAA1E2EFC2E47.taxon	distribution	Distribution. Known only from two localities in the Venezuelan Andes and one in the Northern Colombian Andes at elevations between 1000 and 1400 m (Fig. 4).	en	Arteaga, Alejandro, Salazar-Valenzuela, David, Mebert, Konrad, Penafiel, Nicolas, Aguiar, Gabriela, Sa ́ nchez-Nivicela, Juan C., Pyron, R. Alexander, Colston, Timothy J., Cisneros-Heredia, Diego F., Yanez-Munoz, Mario H., Venegas, Pablo J., Guayasamin, Juan M., Torres-Carvajal, Omar (2018): Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru. ZooKeys 766: 79-147, DOI: http://dx.doi.org/10.3897/zookeys.766.24523, URL: http://dx.doi.org/10.3897/zookeys.766.24523
