taxonID	type	description	language	source
BD2187DB72140A64E199B154FF0AC60E.taxon	description	(Figs. 1 – 6) Diagnostic Combination. Lateral lobes (¼ adnasalia) clearly delineated (Fig. 1); lamellae clypeales conical (Fig. 1); antenna shorter than HW; lateral elongation of antennomere 3 short, A 39 / A 4, 0.20; mandible not toothed; primary pore ANg inserted proximally; setae AN 2 and AN 3 not articulated at same level on antennomere 3; prementum with 2 additional setae; head appendages with several spine-like secondary setae; metathoracic legs about 1.20 times as long as prothoracic legs, with additional spiniform setae on each article; tibiae not pseudo-chelate; spinulae on ventral margin of tibiae (lacking on metatibia) and tarsi spine-like (Fig. 5); primary setae CO 6 and CO 7 inserted distally on procoxa; posterodorsal margin of tibiae with a row of additional natatory setae (1 st instar) (Fig. 6); seta TI 6 lacking on protibia, short and spine-like on mesotibia and hair-like on metatibia; trochanter with. 15 secondary setae (Table 2); last abdominal segment subtrapezoidal, truncate along posterior margin, with spine-like additional setae present (1 st instar); siphon faintly delineated (Fig. 2); urogomphus longer than last abdominal segment, with additional setae (1 st instar) (Fig. 2) and secondary setae (2 nd and 3 rd instar). Description: 1 st Instar (Fig. 1). Color. Head capsule predominantly yellow with some small brownish maculae laterally; head appendages predominantly pale yellow except articulation of A 1 and A 2, dark brown; thoracic and abdominal terga predominantly yellow with a few small dark brown maculae; legs and urogomphi predominantly yellow. Head (Fig. 1). HL ¼ 0.83 – 0.88 mm (mean ¼ 0.85 mm, n ¼ 6); HW ¼ 0.80 – 0.85 mm (mean ¼ 0.83 mm, n ¼ 6); FCL ¼ 0.40 – 0.46 mm (mean ¼ 0.44 mm, n ¼ 6). Cephalic capsule (Fig. 1) obovate, about as long as broad (HL / HW ¼ 1.01 – 1.07), lateral lobes (¼ adnasalia) distinct; apical margin of frontoclypeus with 19 – 20 spatulate lamellae clypeales. Antenna slightly shorter than HW (length of antenna / HW ¼ 0.89 – 0.95); A 2. A 3 » A 4. A 1, A 2 / A 3 ¼ 1.06 – 1.20; A 39 / A 4 ¼ 0.13 – 0.16. Mandible. Inner margin not toothed. Maxilla. Galea 0.71 – 0.73 times length of palpomere 1; length of MX 3 / length of MX 2 ¼ 1.27 – 1.28. Chaetotaxy and porotaxy. Parietals lacking additional setae; prementum with 2 additional setae; antennomere 4 lacking an additional pore; primary pore ANg inserted proximally; primary seta AN 2 articulated distally, not at level of seta AN 3. Legs (Figs. 3 – 6). Tibiae of normal shape (i. e., not pseudo-chelate); metathoracic legs about 1.20 times length of prothoracic legs, and about 3.50 times HW; femur. coxa. tibia » tarsus. trochanter (metathoracic leg); spinulae on ventral margin of tibiae and tarsi spine-like and strongly developed, those on protarsus articulated on a black carina (visible under a dissecting microscope); longest of metathoracic claws 0.36 – 0.37 times as long as metatarsus. Chaetotaxy and porotaxy. Setae CO 6 and CO 7 inserted distally on all legs; seta FE 6 hair-like on each femur; seta TI 6 lacking on protibia, short and spine-like on mesotibia, and elongate and hair-like on metatibia. Additional setae present on each article (Table 1). Abdomen (Fig. 2). LLAS ¼ 0.43 – 0.50 mm (mean ¼ 0.46 mm, n ¼ 6). Segment 8 subtrapezoidal dorsally, shorter than HW, LLAS / HW ¼ 0.53 – 0.58, truncate posteriorly. Siphon faintly delineated. Urogomphus (Fig. 2). Length of urogomphus ¼ 1.00 – 1.02 mm (mean ¼ 1.01 mm, n ¼ 6), 2.02 – 2.39 times as long as LLAS, 1.18 – 1.27 times as long as HW. Chaetotaxy and porotaxy. Urogomphus with several additional spine-like setae. 2 nd Instar (Figs. cf. Alarie et al. 2001). Color. Head capsule predominantly yellow with a blotchy pattern comprising several large brownish maculae; head appendages predominantly pale yellow except A 1 dark brown; thoracic and abdominal terga predominantly yellow with a blotchy pattern; legs creamy white to pale yellow; urogomphi predominantly yellow. Head. HL ¼ 1.32 – 1.35 mm (mean ¼ 1.34, n ¼ 3); HW ¼ 1.18 – 1.23 mm (mean ¼ 1.21 mm, n ¼ 3); FCL ¼ 0.66 mm (mean ¼ 0.66 mm, n ¼ 3). Cephalic capsule obovate, slightly longer than broad (HL / HW ¼ 1.08 – 1.14), lateral lobes (¼ adnasalia) delineated; apical margin of frontoclypeus with 35 – 39 spatulate lamellae clypeales; parietals with 8 – 10 lateral spines. Antenna shorter than HW (length of antenna / HW ¼ 0.77 – 0.80); A 2. A 1 subequal to A 3. A 4, A 2 / A 3 ¼ 1.11 – 1.23; A 39 / A 4 ¼ 0.11 – 0.13. Mandible. Inner margin not toothed. Maxilla. Galea 0.63 – 0.71 times length of MX 1; length of MX 3 / length of MX 2 ¼ 0.92 – 1.09. Chaetotaxy and porotaxy. Prementum with 2 additional setae; primary pore ANg inserted proximally; primary seta AN 2 articulated distally, not at level of seta AN 3; head appendages with several secondary spine-like setae: antennomere 1 (7 – 10), antennomere 2 (9), maxillary palpomere 1 (7), maxillary palpomere 2 (6 – 7), labial palpomere 1 (10). Legs. Tibiae of normal shape (i. e., not pseudo-chelate); metathoracic legs about 1.20 times length of prothoracic legs, and about 3.50 times HW; femur. coxa. tibia » tarsus. trochanter (metathoracic leg); length of metafemur / length metacoxa about 1.20; longest of metathoracic claws 0.31 times as long as metatarsus; spinulae on ventral margin of tibiae and tarsi spine-like and strongly developed (lacking on metatibia). Chaetotaxy and porotaxy. Seta CO 6 always distad; seta CO 7 inserted proximally on procoxa and distally on meso- and metacoxa; position and number of secondary setae (Table 1). Abdomen. LLAS ¼ 0.83 – 0.96 mm (mean ¼ 0.88 mm, n ¼ 3). Segment 8 subtrapezoidal dorsally, shorter than HW, LLAS / HW ¼ 0.68 – 0.82, truncate posteriorly. Siphon lacking. Urogomphus. Length of urogomphus ¼ 1.47 – 1.54 mm (mean ¼ 1.49 mm, n ¼ 3), 1.54 – 1.80 times as long as LLAS, 1.19 – 1.25 times as long as HW. Chaetotaxy and porotaxy. Urogomphus with several secondary setae. 3 rd Instar. As 2 nd instar except as follows. Color. Mottled pattern denser; urogomphus predominantly dark brown. Head. HL ¼ 1.93 – 2.02 mm (mean ¼ 1.98 mm, n ¼ 2); HW ¼ 1.69 – 1.72 mm (mean ¼ 1.71 mm, n ¼ 2); FCL ¼ 0.81 – 0.83 mm (mean ¼ 0.82 mm, n ¼ 2). Cephalic capsule slightly longer than broad (HL / HW ¼ 1.13 – 1.17); apical margin of frontoclypeus with 40 – 45 spatulate lamellae clypeales; parietals with 11 – 12 lateral spines. Antenna. Length of antenna / HW ¼ 0.66 – 0.70; A 2. A 1 » A 3. A 4, A 2 / A 3 ¼ 1.23 – 1.31; A 39 / A 4 ¼ 0.13 – 0.14. Maxilla. Galea 0.61 – 0.70 times length of MX 1; length of MX 3 / length of MX 2 ¼ 0.85 – 0.86. Legs. Metathoracic legs about 1.20 times length of prothoracic legs, and about 3.30 times HW; femur. coxa. tibia » tarsus. trochanter (metathoracic leg); longest of metathoracic claws 0.30 – 0.31 times as long as metatarsus. Chaetotaxy and porotaxy. Position and number of secondary setae (Table 2). Abdomen. LLAS ¼ 1.20 – 1.39 mm (mean ¼ 1.29 mm; n ¼ 2); LLAS / HW ¼ 0.70 – 0.82. Urogomphus. Length of urogomphus ¼ 1.72 – 1.93 mm (mean ¼ 1.83 mm, n ¼ 2), 1.24 – 1.61 times as long as LLAS, 1.02 – 1.12 times as long as HW. Source of Associated Material. The larvae of Allomatus nannup were collected in association with adults at the following locality: WEST AUSTRALIA, Mokerdillup Brook, 10 km. W. Bridgetown, 3480940 E, 116839170 S, 5. x. 1999, S. Halse coll; Blackwood river nr Bridgetown, 3385980 E, 11680980 S, 21. ix. 2000, CHS Watts coll.	en	Alarie, Yves, Watts, Chris H. S. (2003): Larval Morphology of Allomatus nannup Watts (Coleoptera: Adephaga: Dytiscidae). The Coleopterists Bulletin 57 (3): 255-264, DOI: 10.1649/551, URL: http://dx.doi.org/10.1649/551
