taxonID	type	description	language	source
C35EF101FFDDC425FF50FD34FE9FFBDF.taxon	description	Figs 1, 2	en	Ross, Andrew J., Hancock, David L., Mcalister, Erica, Zhou, Jiale (2024): A fly in the ointment: the re-discovered holotype of ‘ Trypeta’ cylindrica Walker, 1853 belongs to the genus Coelopacidia Enderlein (Diptera: Tephritidae), not Chyliza Fallén (Psilidae). Zootaxa 5529 (3): 593-600, DOI: 10.11646/zootaxa.5529.3.10, URL: http://dx.doi.org/10.11646/zootaxa.5529.3.10
C35EF101FFDDC425FF50FD34FE9FFBDF.taxon	materials_examined	Holotype. NHMUK 013448450, male, William Saunders Collection. Type locality. India, precise location not known.	en	Ross, Andrew J., Hancock, David L., Mcalister, Erica, Zhou, Jiale (2024): A fly in the ointment: the re-discovered holotype of ‘ Trypeta’ cylindrica Walker, 1853 belongs to the genus Coelopacidia Enderlein (Diptera: Tephritidae), not Chyliza Fallén (Psilidae). Zootaxa 5529 (3): 593-600, DOI: 10.11646/zootaxa.5529.3.10, URL: http://dx.doi.org/10.11646/zootaxa.5529.3.10
C35EF101FFDDC425FF50FD34FE9FFBDF.taxon	description	Redescription Male. Length from tip of head (excluding antennae) to tip of abdomen 10.2 mm, wing length 9.4 mm; head length 1.6 mm (excluding antennae). Head with 2 pairs of frontal and 1 pair of orbital setae, all weak; ocellar setae very weak and hair-like; antennae about as long as face, first flagellomere fulvous, third flagellomere pale yellow, elongate and apically rounded; arista long, black and bare. Thorax largely reddish brown with postpronotal lobes and narrow postsutural lateral vittae yellow; anepisternal stripe broadly yellow and extending as a narrow dorsal stripe to postpronotal lobe; chaetotaxy with well developed postpronotal, anterior and posterior notopleural, supra-alar and postalar setae and moderately developed dorsocentral setae placed behind line of supra-alar setae; I anepisternal seta; scutellum red-brown with 2 pairs of setae, basal and apical. Wing largely hyaline with a very faint yellowish tinge; pterostigma yellow; r-m crossvein at apical third of cell dm; dm-cu crossvein with a narrow infuscation, broadest posteriorly; apical brown spot across vein R 4 + 5 as broad as long in apical portion of cell r 2 + 3. Legs yellow; fore femora with 2 short, black anterior spines in distal half; mid tibiae with 2 distinct apical spines. Abdomen elongate, reddish brown with darker staining.	en	Ross, Andrew J., Hancock, David L., Mcalister, Erica, Zhou, Jiale (2024): A fly in the ointment: the re-discovered holotype of ‘ Trypeta’ cylindrica Walker, 1853 belongs to the genus Coelopacidia Enderlein (Diptera: Tephritidae), not Chyliza Fallén (Psilidae). Zootaxa 5529 (3): 593-600, DOI: 10.11646/zootaxa.5529.3.10, URL: http://dx.doi.org/10.11646/zootaxa.5529.3.10
C35EF101FFDDC425FF50FD34FE9FFBDF.taxon	discussion	Remarks The Coelopacidia specimen, NHMUK 013448450, is very similar to Walker’s (1853) original description and Westwood’s figures of ‘ Trypeta ’ cylindrica in general habitus and particularly in that it possesses brown wing-tips. The specimen also has distinct pterostigmata on the wings, long, black setae on the thorax and head, and a long, curved arista on the antenna, which are clearly seen in Westwood’s figures. From this it is clear that this specimen is the correct holotype. Although the number of specimens was not expressly stated, the precise nature of Walker’s original description and subsequent illustrations implied only a single specimen was available, here accepted as the ‘ Holotype by monotypy’ under Article 73.1.2 of the International Code of Zoological Nomenclature (ICZN 1999). This specimen, a male, also bears a handwritten [original?] label that is largely indecipherable but appears to indicate ‘ Cape Camp’. Bezzi (1913) suggested a relationship with Adrama Walker and his statement of ‘ Type in London’ therefore indicates a reference to this specimen. Hardy’s (1959) redescription does not match the original description and illustrations, so he had misidentified the “ type ” (see below). Later, Hardy (1977) listed Coelopacidia cylindrica (Walker) as a new combination in the Tephritidae, so he had subsequently correctly identified the holotype but did not refer to it as such. Kapoor (1993) also listed Coelopacidia cylindrica in the Tephritidae, however he referred to both Hardy (1959) and Hardy (1977), effectively referring to both specimens. Hancock & Drew (2005), David et al. (2013, with figures) and Hancock (2019) referred to this specimen but they did not consider it was the holotype of a previously named species. This species is very similar to C. vivax (Munro, 1932) from South Africa, differing only in the broader apical spot on the wing. The locality ‘ Cape Camp’ is ambiguous and the possibility remains that the type locality of ‘ East Indies [India] ’ is erroneous and the specimen is actually from the Cape Province of South Africa, with C. vivax a potential synonym.	en	Ross, Andrew J., Hancock, David L., Mcalister, Erica, Zhou, Jiale (2024): A fly in the ointment: the re-discovered holotype of ‘ Trypeta’ cylindrica Walker, 1853 belongs to the genus Coelopacidia Enderlein (Diptera: Tephritidae), not Chyliza Fallén (Psilidae). Zootaxa 5529 (3): 593-600, DOI: 10.11646/zootaxa.5529.3.10, URL: http://dx.doi.org/10.11646/zootaxa.5529.3.10
C35EF101FFDBC427FF50FAC4FE03FE17.taxon	description	Figs 3, 4	en	Ross, Andrew J., Hancock, David L., Mcalister, Erica, Zhou, Jiale (2024): A fly in the ointment: the re-discovered holotype of ‘ Trypeta’ cylindrica Walker, 1853 belongs to the genus Coelopacidia Enderlein (Diptera: Tephritidae), not Chyliza Fallén (Psilidae). Zootaxa 5529 (3): 593-600, DOI: 10.11646/zootaxa.5529.3.10, URL: http://dx.doi.org/10.11646/zootaxa.5529.3.10
C35EF101FFDBC427FF50FAC4FE03FE17.taxon	materials_examined	Holotype. NHMUK 013444154, female. Type locality. Peradeniya, Sri Lanka. Other material. Specimens listed by Shatalkin (1998) under ‘ Chyliza cylindrica ’, including the misidentified ‘ type’ (NHMUK 013444153).	en	Ross, Andrew J., Hancock, David L., Mcalister, Erica, Zhou, Jiale (2024): A fly in the ointment: the re-discovered holotype of ‘ Trypeta’ cylindrica Walker, 1853 belongs to the genus Coelopacidia Enderlein (Diptera: Tephritidae), not Chyliza Fallén (Psilidae). Zootaxa 5529 (3): 593-600, DOI: 10.11646/zootaxa.5529.3.10, URL: http://dx.doi.org/10.11646/zootaxa.5529.3.10
C35EF101FFDBC427FF50FAC4FE03FE17.taxon	description	Description See Shatalkin (1998) under ‘ Chyliza cylindrica’. Holotype, length from tip of head (excluding antennae) to tip of abdomen 5.2 mm, wing-length 3.8 mm.	en	Ross, Andrew J., Hancock, David L., Mcalister, Erica, Zhou, Jiale (2024): A fly in the ointment: the re-discovered holotype of ‘ Trypeta’ cylindrica Walker, 1853 belongs to the genus Coelopacidia Enderlein (Diptera: Tephritidae), not Chyliza Fallén (Psilidae). Zootaxa 5529 (3): 593-600, DOI: 10.11646/zootaxa.5529.3.10, URL: http://dx.doi.org/10.11646/zootaxa.5529.3.10
C35EF101FFDBC427FF50FAC4FE03FE17.taxon	discussion	Remarks Specimen 013444153 matches Hardy’s (1959) description of the “ type ” of Trypeta cylindrica Walker, particularly in that it is in ‘ very poor condition’ with ‘ completely hyaline wings’ and the ‘ thorax is densely covered with recumbent white pile, ’ but does not match the original description by Walker (1853) and figures by Westwood, so was misidentified. When Steyskal (1977) listed ‘ cylindrica ’ under ‘ unplaced species of Platystomatidae’ he referred to Hardy (1959), so was therefore referring to this specimen. This specimen was subsequently referred to as the ‘ type’ of Chyliza cylindrica (Walker) and the species was re-described by Shatalkin (1998), who included Chyliza pallidipes Lamb, 1917 as a junior synonym. Shatalkin’s description was based on a total of 12 specimens from Sri Lanka, India and Thailand, including the holotype of C. pallidipes. Given that the ‘ type’ had been misidentified and the name combination ‘ Chyliza cylindrica (Walker) ’ was not formally described as a valid species then the ‘ Type’ and ‘ Holotype’ labels were therefore incorrectly attributed. The green-circled ‘ Type’ label was added in 1940 when the collection was moved to safety at the beginning of World War II and the red-bordered ‘ Holotype’ label very recently (A. Pont, pers. comm.). However, the green-circled ‘ Type’ labels were added arbitrarily and were often placed on specimens that were not types. The holotype of C. pallidipes (NHMUK 013444154, Fig. 4 A, B), and the misidentified ‘ type’ are clearly conspecific and are very similar in length, from tip of head (excluding antennae) to tip of abdomen, being 5.2 mm for 013444154 and 5.4 mm for 013444153. Chyliza pallidipes Lamb stat. rev. is thus taken out of synonymy and is the valid name for this species.	en	Ross, Andrew J., Hancock, David L., Mcalister, Erica, Zhou, Jiale (2024): A fly in the ointment: the re-discovered holotype of ‘ Trypeta’ cylindrica Walker, 1853 belongs to the genus Coelopacidia Enderlein (Diptera: Tephritidae), not Chyliza Fallén (Psilidae). Zootaxa 5529 (3): 593-600, DOI: 10.11646/zootaxa.5529.3.10, URL: http://dx.doi.org/10.11646/zootaxa.5529.3.10
