identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
34B24800D54859F1BB4D94F7A303214D.text	34B24800D54859F1BB4D94F7A303214D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyphocallipus africanus Stoev & Akkari 2025	<div><p>Cyphocallipus africanus sp. nov.</p><p>Figs 1, 2</p><p>Diagnosis.</p><p>A species of the genus Cyphocallipus, differing from C. excavatus in having a semicircular rounded protective lamella at the distal part of telopodite (vs subrectangular in C. excavatus), bearing smaller fold at the margin than in C. excavatus; coxosternal process m with apically rounded and not distally expanded process k, and a much shorter process j (1 / 3 of process k vs 1 / 2 of process k in C. excavatus); coxosternal process i with a straight distal part; coxosternal process i ’ broad, incised distally, posterior part of distal incision oval, posterior proximal margin triangularly oval; sternal lamella similar to C. excavatus in position but lower. Table 1 provides a comparison of the main taxonomic characters between Cyphocallipus africanus sp. nov. and C. excavatus .</p><p>Etymology.</p><p>The specific epithet emphasizes the discovery of the first native member of the order Callipodida on the African continent. Adjective.</p><p>Material examined.</p><p>• Holotype: adult male, Morocco, Tanger-Tetouan-Al Hoceima region, Chefchaouen Province, Bab Taza, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-5.1054444&amp;materialsCitation.latitude=35.068695" title="Search Plazi for locations around (long -5.1054444/lat 35.068695)">cave Maala Abladir</a>, 35°04'07.3"N, 5°06'19.6"W, 14.VIII.2005, leg. G. E. S. [Grupo de Exploraciones Subterráneas of Priego], NHMW MY 10657.</p><p>Description.</p><p>Body cylindrical, length circa 56 mm, maximal body diameter circa 2.85 mm. 58 PTs + Telson. Colour in life unknown; colour of preserved specimen: head frons and vertex yellow alveolate. Male head concave with the characteristic for the genus pair of knobs on epicranium (Fig. 1 A, B); Eyes: 53 ommatidia in 10 rows (Fig. 1 A). Organ of Tömösváry twice as large as largest ommatidium. All pleurotergites yellow, with light-brown transverse band on posterior margin (Fig. 1 C). Antennae broken, brownish, ratio: 2&gt; 3&gt; 5&gt; 4&gt; 6&gt; 7 (vs 5&gt; 2&gt; 4&gt; 6&gt; 3&gt; 1&gt; 7). Legs yellow; claws darker. Pleurotergal crests touching each other at anterior part of pleurotergite (prozonum) and becoming separate and thinning towards pleurotergite end (Fig. 1 C); 5 + 5 pleurotergal crests between ozopores on midbody pleurotergites. Ozopores situated between crests 5 and 6, very small and difficult to spot.</p><p>Legs: Leg-pairs 1 and 2 reduced and more setose than the rest, with postfemoral and tibial brushes and tarsal combs. Leg-pairs 3–5 with postfemoral and tibial brushes; legs 1–5 (also postgonopodal legs) with prefemur, femur, postfemur and tibia micropapillate; leg-pair 2 with posterior gonopore, prefemora of legs 3–7 slightly enlarged and covered with dense setae. Coxal sacs large and conspicuous at least until leg-pair 12.</p><p>Telson (Fig. 1 D): anal valves: with smaller dorsal and larger ventral plates; dorsal plates with one microseta each; hypoproct tripartite, median sclerite largest, subtrapezoidal, bearing a pair of macrosetae situated in the middle. Lateral sclerites smaller, subtriangular, with one seta each. Epiproct: with two moderately long spinnerets.</p><p>Gonopods (Figs 1 E, F, 2 A – F): gonopod comprising a long and arcuate telopodite (T), attached at the inner posterior end of gonocoxa and pointing posterioventrad, accompanied by a flagelliform accessory process (pseudoflagellum - F), following the curvature of telopodite. Distal part of telopodite enlarged, with a rounded buckle (bu) at the base of the protective lamella (Pl). Protective lamella covering partially the solenomere (s) and parasolenomere (ps); lobe more rounded at margin, semicircular, with a very small fold at the margin. Sternal lamella large, broad, posterior to the telopodite and the coxal processes, less developed and lower compared to C. excavatus; Coxosternum with three large processes i ’, i, and m. Posterior coxosternal process i ’ broad, incised distally, posterior part of distal incision oval, posterior proximal margin triangularly oval; coxosternal process m forked into processes j and k; process j is 1 / 3 of the length of process k; k apically rounded and not distally expanded as in C. excavatus . Coxosternal process i straight.</p><p>Female. Unknown.</p><p>Distribution.</p><p>Known only from the type locality in Morocco. Bab Taza is a small town in Talassemtane National Park in the Rif Mountains (Moutaouakil pers. comm.) (Fig. 3). The cave Maala Abladir has relatively large entrance (Fig. 4) and two floors. Clay covers much of the area, and the cave ends with a pool (Fig. 5). In a paper summarizing the results of studies on hypogean beetles of the family Cholevidae in Morocco, Fresneda and Fadrique (2006) reported the Speonemadus maroccanus (Jeannel, 1936) from the Maala Abladir. To our knowledge, this is the only other animal hitherto recorded from the cave.</p><p>Habitat.</p><p>Although collected from a cave, the new species does not exhibit any traits of troglomorphism, leading us to presume that it was only seeking shelter there.</p><p>Comments.</p><p>The specimen is heavily infested by fungi (Fig. 1 A, B). Most likely they are representatives of the ectoparasitic fungi of the order Laboulbeniales Engler, 1898, as described in other members of Callipodida from the Iberian Peninsula (Reboleira and Enghoff 2015).</p></div>	https://treatment.plazi.org/id/34B24800D54859F1BB4D94F7A303214D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Stoev, Pavel;Akkari, Nesrine	Stoev, Pavel, Akkari, Nesrine (2025): Cyphocallipus africanus sp. nov.: first native callipodid millipede from continental Africa (Diplopoda, Callipodida). ZooKeys 1253: 321-341, DOI: 10.3897/zookeys.1253.161612
581D550E3BFE5315B3A1C85A221AA2BF.text	581D550E3BFE5315B3A1C85A221AA2BF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyphocallipus excavatus Verhoeff 1909	<div><p>Cyphocallipus excavatus Verhoeff, 1909</p><p>Figs 6, 7, 8, 9</p><p>Material examined.</p><p>• 1 male, Spain, Almería, Sierra de Gádor, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-2.7581444&amp;materialsCitation.latitude=36.880424" title="Search Plazi for locations around (long -2.7581444/lat 36.880424)">Canchal Antenas II, MSS, Félix</a>, 36°52'49.53"N, 2°45'29.32"W, 1300 m, 11.XII.2018, Centro de Investigación de Colecciones Científicas de la Universidad de Almería, Cecoual-Dpto. Biol. Geol. col. Pablo Barranco leg. (Figs 3 A – D, 4 A – E) ; • 1 male, 3 females, Spain, Málaga Province, Cómpeta, Casa de la Mina, Pine Forest, 15.I.2006, leg. D. Kime, det. J-P. Mauriès (National Museum of Natural Hostory, Sofia – Myriapoda collection) (Fig. 3 E, F) ; • 1 male, Spain, Órgiva, 7.X.1977, Briganti, Parodi, Zoia leg., ex. Mus. Verona, G. Osella ded. 1983 (Scanning electron micrographs) .</p><p>Description.</p><p>This species has already been redescribed by Mauriès (1978), Hoffman (2009), and Spelda (2015). However, here, we present habitus images of the species and document the species gonopod morphology, using both light micrography and scanning electron microscope imagery. Thus, we provide more accurate and detailed information on key taxonomic features (Figs 6 – 9). Table 1 summarizes the main distinguishing characteristics of the two species.</p></div>	https://treatment.plazi.org/id/581D550E3BFE5315B3A1C85A221AA2BF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Stoev, Pavel;Akkari, Nesrine	Stoev, Pavel, Akkari, Nesrine (2025): Cyphocallipus africanus sp. nov.: first native callipodid millipede from continental Africa (Diplopoda, Callipodida). ZooKeys 1253: 321-341, DOI: 10.3897/zookeys.1253.161612
6F65A34840DC5D2387D2B44C5D06C9FD.text	6F65A34840DC5D2387D2B44C5D06C9FD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyphocallipus Verhoeff 1909	<div><p>Genus Cyphocallipus Verhoeff, 1909</p><p>Diagnosis.</p><p>Body with 53–59 PTs. Forehead of male concave; a pair of small knobs present on epicranium of both known species of Cyphocallipus; pleurotergites: with longitudinal crests, which are spaced apart in their posterior part; metazona not much elevated compared to prozona (in Lusitanipus and Dorycallipus posterior part are elevated); Collum: with longitudinal furrows and indistinct setae in the posterior half and at the side flaps; Coxal sacs: from 3 rd to 22 nd pair of legs. Gonopods: with a large sternal lamella and three coxosternal processes, two of them much longer than the 3 rd between them, the foremost with an excavation for the telopodite; telopodite long and thin pseudoflagellum following its curvature. Female (unknown in C. africanus): posterior lateral edge of second metazonum produced caudal as a large subtriangular lobe over lower side of third. Second leg-pair greatly reduced (Verhoeff 1909; Hoffman 2009; Spelda 2015).</p><p>Comment.</p><p>In his review of the members of the order Callipodida from the Iberian Peninsula, Spelda (2015) provided a comprehensive bibliographic catalogue of the records of the genus Cyphocallipus and C. excavatus, along with photographs of key taxonomic characteristics of the species. Therefore, we do not include a bibliographic review here.</p></div>	https://treatment.plazi.org/id/6F65A34840DC5D2387D2B44C5D06C9FD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Stoev, Pavel;Akkari, Nesrine	Stoev, Pavel, Akkari, Nesrine (2025): Cyphocallipus africanus sp. nov.: first native callipodid millipede from continental Africa (Diplopoda, Callipodida). ZooKeys 1253: 321-341, DOI: 10.3897/zookeys.1253.161612
