identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
C213FF65FFF4F640519FF8C0FAB7FEEC.text	C213FF65FFF4F640519FF8C0FAB7FEEC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lepidostoma coreanum	<div><p>The L. coreanum Species Complex of the genus Lepidostoma Rambur</p><p>The males of the L. coreanum Species Complex are morphologically characterized by unique genitalia, scapes of antennae, and wings of males as follows: brown. Abdominal segment IX short. Segment X composed of slender mesal, lateral and ventral arms but lateral and ventral arms often short or reduced completely. Inferior appendages setose, composed of broad, semicircular or subquadrate main articles, bar-like or comb-like dorsal hooks and short bar-shaped inner hooks. Scapes of antennae very setose, thick and long, 1.5–2.0 times as long as head, each with short mesal process. Wings apically round, covered with fine setae, scales, and spindle-like setae, often reduced in venation particularly in coastal area. The species complex consists of Lepidostoma coreanum (Kumanski &amp; Weaver 1992), L. niigataense sp. nov., L. hattorii sp. nov., L. yuwanense sp. nov., and L. ishigakiense sp. nov.</p></div>	https://treatment.plazi.org/id/C213FF65FFF4F640519FF8C0FAB7FEEC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ito, Tomiko	Ito, Tomiko (2016): The Lepidostoma coreanum Species Complex (Trichoptera, Lepidostomatidae) in the Asian Far East. Zootaxa 4061 (4): 397-417, DOI: 10.11646/zootaxa.4061.4.5
C213FF65FFF5F647519FFE07FD60FADC.text	C213FF65FFF5F647519FFE07FD60FADC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lepidostoma coreanum (Kumanski & Weaver 1992) Kumanski & Weaver 1992	<div><p>Lepidostoma coreanum (Kumanski &amp; Weaver 1992)</p><p>(Figs. 1–6, 21)</p><p>Indocrunoecia coreana Kumanski &amp; Weaver 1992, 163–165, male, female, North Korea (Province Pyongan Pukudo). Dinarthrum coreanum: Ito 1998, male, female, larva, case, Russian Far East (Primorye, Khasanski), Japan (Honshu: Shizuoka (Amagi-yugashima-cho), Wakayama); Ito et al. 2002, 23, Japan (Shikoku).</p><p>Lepidostoma coreanum: Weaver 2002, 180; Ito 2005, 504, 507, 512, male, larva.</p><p>Male (Figs. 1–5). Specimens from Primorye (Fig. 1). Antennae about 5.5 mm long, scape setose, thick and long, 1.5 times as long as head, short mesal process near middle of each scape. Maxillary palps extremely setose, each 2- segmented, 0.8 mm long in total, basal segment curved anteriorly at half its length with tuft of very long setae apically, long apical segment emerging near middle of basal segment. Labial palps each 3-segmented, slender, 1.0 mm long in total.</p><p>Wings covered with fine setae, short scales, and spindle-like setae; spindle-like setae lined and circled in coastal area; forewings each with discoidal cell, thyridial cell, and apical forks 1 and 2, Cu2 indistinct at basal half and fused to Cu1 and A1+2+3 apically, arculus large; hind wings each with fork 1, Cu2 fused to Cu1 apically, A1+2+3 also fused to Cu1 apically; forewings and hind wings each 5.0– 5.2 mm long and 4.7–4.8 mm long, respectively (n=3).</p><p>Genitalia. Abdominal segment IX (IX) short, dorsum longer than venter. Segment X (X) composed of slender mesal (ma), lateral (la), and ventral arms (va); mesal arms longest and directed caudad, lateral arms directed caudad and distinctly shorter than mesal arms, ventral arms curved slightly dorsocaudad with length subequal to lateral arms; 3 arms subacute apically. Inferior appendages (ia) setose, each composed of main article (maia), dorsal hook (dh), and inner hook (ih); basal half of main article broad in lateral view, semicircular with serrate posterior margin in ventral view; apical half of main article bar-like, curved mesad, mesocaudal apex rather acute; dorsal hook directed laterocaudad, length subequal to main article; inner hook much shorter than dorsal hook, bar-like and directed laterocaudad. Phallic apparatus thick, apical half membranous with inner U-shaped sclerite, paramere absent.</p><p>Variation (Figs. 2–5). Specimens of all 4 localities in Japan exhibit some variations. In a male from Amagiyugashima-cho, Shizuoka (Fig. 2), the genitalia are almost identical to those of males from the type locality in North Korea (Kumanski &amp; Weaver 1992) and from Primorye (Fig. 1), but the wing venation and maxillary palps are a little different from those of the later; in the Amagi-yugashima-cho specimens, Ma and Mp are stalked, crossvein m-cu and the thyridial cell are absent in each forewing; crossvein m-cu and the apical parts of Cu2 and A1+A2+A3 are lacking in each hind wing; the basal segment of each maxillary palp is thickened apically and the long apical segment emerges from the basal 2/3 of the basal segment.</p><p>In a male from Wakayama (Fig. 3), the lateral arms of segment X are reduced, hump-like; the basal 2/3 of Cu2 is lacking in each forewing.</p><p>In the males from Ehime (Fig. 4) and Kochi (Fig. 5), the lateral arms of segment X are reduced to a low hump, the main article of each inferior appendage has a round convex posteromesal margin in ventral view. The maxillary palps are setose, short, and unsegmented. Cubital and anal veins of forewings and hind wings are not fused apically and an arculus is absent in cubital area and some crossveins are lacking. Cu1 has a short apical fork 5 in each hind wing of a specimen from Kochi.</p><p>Female (Fig. 6 A–F). Specimens from Primorye. Antennae about 5.5 mm long, scape setose, thick, long and 2 times as long as head, mesal processes absent. Maxillary palps each 5-segmented and about 1 mm long, labial palps each 3-segmented and 0.6 mm long; all palps slender.</p><p>Wings covered with short fine setae, typical venation in females of the family; forewings with apical forks 1, 2, 3, 5, discoidal and thyridial cells, and arculus; hind wings with forks 1, 2 and 5. Lengths of forewings and hind wings on each side 5.1–6.0 mm and 4.0–5.0 mm, respectively (n=5).</p><p>Lateral ends of tergite VIII broad. Lateral plates weakly sclerotized, trianglular. Subgenital plate sclerotized, subquadrate with slightly concave posterior margin and middle depression on anterior margin. Vaginal apparatus oval and slightly sclerotized.</p><p>Larva (Figs. 6 G, H). Specimens from Primorye. Head width and body length of final instar larva up to 0.7 mm and 7 mm, respectively. Head dark brown, with numerous short secondary setae on dorsum in early instars. Pronotum dark brown. Mesonotal plate brown with some light colored dots on anterior half and light brown on posterior half. Chaetotaxy of thorax: 35–40 setae on pronotum; 7, about 12 and about 12 setae on setal areas (sa) 1, sa 2 and sa 3, respectively, of mesonotum; and 1, about 5, and about 12 on sa 1, sa 2, and sa 3, respectively, of metonotum. Single tracheal gills present in subdorsal rows near posterior margins of segments II–VI, in subventral rows near posterior margins of segments II–VII, and in subdorsal rows near anterior margins of segments II–III (or II–IV or II–V). Single subventral gill present near anterior margin of segment II in some specimens. Other characters as in L. robustum (Ito 1984) .</p><p>Case of larva (Fig. 6 I). Specimens from Primorye. Cylindrical, slightly curved and slightly tapered posteriorly, composed of sand grains with round posterior opening. Length up to 10 mm.</p><p>Remarks. The males of this species have some variations locally but they are clearly distinguished from those of other related species by their elongated ventral arms of segment X. Records of this species from Mie, Honshu, Japan, by Kawase and Morita (2001, 2010) and Morita (2008, 2011) are uncertain because the specimens could not be examined with our present taxonomic knowledge.</p><p>Specimens examined. Russian Far East, Primorye, Khasanski: 4 males, small water flow near Vitya Village, 30.vi.1988, T.S. Vshivkova; 5 larvae, same locality, 11.v.1996, T.S. Vshivkova &amp; T. Ito; 1 male, 6 females, 2 pupae (1 male, 1 female), same locality, 11.v.1996 (larvae), reared and preserved in alcohol in vi.1996, T.S. Vshivkova &amp; T. Ito. Japan, Honshu, Shizuoka: 1 male, 1 female, 2 pupae (females), Izu-shi, Amagi-yugashimacho, Kami-funabara, Funabara-gawa, 550 m, 21.iv.1996 (larvae), reared and preserved in alcohol on 24.vi.1996, T. Hattori &amp; T. Ito. Wakayama: 1 male, Kozagawa-cho, Hirai, 20.v.1993, N. Kuhara, light trap. Shikoku, Ehime: 3 males, 1 female, Uchiko-machi, Odamiyama, Miyamaso-mukai, small water flow, 24.vii–8.viii.2000, E. Yamamoto, Malaise trap; 3 females, Kuma-kogen-cho, Yurano-no-mori, 30.vi–31.vii.2007, E. Yamamoto, Malaise trap. Kochi: 1 male, Okawa-mura, Funato, Yoshino-gawa, 378 m, 27.iii.2004, K. Nio.</p><p>Distribution (Fig. 21). North Korea (Province Pyongan Pukudo), Russian Far East (Primorye, Khasanski), Japan (central Honshu, Shikoku).</p><p>Habitat. Adults were collected from small water flows in forests.</p><p>Japanese name. Korea-sunatsutsu-tobikera.</p></div>	https://treatment.plazi.org/id/C213FF65FFF5F647519FFE07FD60FADC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ito, Tomiko	Ito, Tomiko (2016): The Lepidostoma coreanum Species Complex (Trichoptera, Lepidostomatidae) in the Asian Far East. Zootaxa 4061 (4): 397-417, DOI: 10.11646/zootaxa.4061.4.5
C213FF65FFF2F64B519FFA17FD1EFA3B.text	C213FF65FFF2F64B519FFA17FD1EFA3B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lepidostoma niigataense	<div><p>Lepidostoma niigataense sp. nov.</p><p>(Figs. 7–11, 21)</p><p>Dinarthrum coreanum: Ito 1998, male, Japan (Honshu: Miyagi, Yamagata, Kanagawa), in part, misidentification. Lepidostomatidae gen. sp.: Nozaki 1997, 37, Japan (Honshu: Kanagawa).</p><p>Diagnosis. Males of this species are very similar to those of Lepidostoma coreanum (Kumanski &amp; Weaver 1992), but are clearly discriminated from the later by the triangular lateral arms and greatly reduced ventral arms of tergum X. Females of this species are difficult to discriminate from those of L. coreanum .</p><p>Male (Figs. 7–11). Type specimens (Fig. 7). Antennae about 6 mm long, each with scape setose, thick and long, 2 times as long as head, short mesal process near middle. Maxillary palps extremely setose, each 2-segmented, 0.6 mm long in total, basal segment curved anterad at basal 2/3 with tuft of very long setae apically, long apical segment emerging near middle of basal segment. Labial palps each 3-segmented, slender, 0.8 mm long in total.</p><p>Wings covered with fine setae, short scales, and spindle-like setae; spindle-like setae lining and encircling especially the cubital area; forewings each with apical forks 1 and 2 and discoidal cells, Cu1 and Cu2 fused with each other for apical 1/4; hind wings each with apical fork 1, Cu unbranched, and vein A lacking; forewings and hind wings each 5.2–6.2 mm long and 4.7–5.0 mm long, respectively (n=3).</p><p>Genitalia. Abdominal segment IX short, dorsum slightly longer than venter. Segment X composed of mesal arms, lateral arms and reduced ventral arms; mesal arms long, more or less compressed laterally, directed caudad with subacute apices; lateral arms triangular or subquadrate, about 1/3 as long as mesal arms; ventral arms reduced to short slender bars. Inferior appendages setose, each composed of main article, dorsal hook, and inner hook; basal half of main article broad in lateral view, broad with serrate posteromesal hump in ventral view; apical half of main article thick bar, curved mesad with comb-like apex; dorsal hook bar-like, directed caudad, slightlly shorter than main article; inner hook much shorter and more slender than dorsal hook, bar-like and directed laterocaudad. Phallic apparatus thick, membranous in apical half with inner U-shaped sclerite, paramere absent.</p><p>Variation (Figs. 8–11). Specimens collected from all 4 localities of Japan show some variations. In a male from Yamagata (Fig. 8), the genitalia and wings are almost identical with those of males from the type locality in Niigata (Fig. 7), but maxillary palps different from those of the later, with the apical segment short, about 1/2 as long as the basal segment.</p><p>In the males from Miyagi (Fig. 9), the mesal arms of segment X are somewhat short relative to those of males from the type locality, the ventral arms of segment X are reduced to low triangular lobes, the main article of each inferior appendage is round and without posteromesal humps in ventral view. Thyridial cell present in each forewing and second segment of each maxillary palp very short.</p><p>In the males from Kanagawa (Fig. 10), the mesal arms of segment X are rather short and the ventral arms are reduced to low triangular lobes; the main article of each inferior appendage has a cleft apically. The wings are almost the same as those of the type specimens, but crossvein m (Ma-Mp) and vein Cu2 are present in each hind wing. The apical segment of each maxillary palp is short, about 1/3 as long as the basal segment.</p><p>In the males from Aichi (Fig. 11), the ventral arms of segment X are reduced to low triangular lobes. The wings are almost the same as those of the type specimens, but cross vein m (Ma-Mp) and apical fork 3 are present in each hind wing. Apical segment of each maxillary palp very short.</p><p>Female (Fig. 7 I–K). Type specimens. Antennal length indeterminate due to specimen condition, scapes setose, thick, long, and 2 times as long as head, mesal processes absent. Maxillary palps each 5-segmented and about 1 mm long, labial palps each 3-segmented and 0.6 mm long, all palps slender.</p><p>Wings covered with short fine setae, venation typical of other species of Lepidostomatidae; forewings and hind wings each 5.2–6.4 mm long and 4.4–5.4 mm long, respectively (n=2).</p><p>Lateral ends of tergite VIII broad. Lateral plates sclerotized, narrow, long rectangular. Subgenital plate weakly sclerotized, subquadrate, with slightly concave posterior margin and deeply concave anterior margin. Vaginal apparatus oval and slightly sclerotized.</p><p>Immature stages. Unknown.</p><p>Holotype: Japan, Honshu, Niigata: 1 male, Minami-uonuma-shi, Yamato-machi, Mizunashi-gawa, 37˚08’26”N, 139˚01’19”E, 300 m a.s.l., 4.vii.1996, T. Hattori, light trap (CBM-ZI 151250).</p><p>Paratypes. 2 males, 2 females, same data as holotype (CBM-ZI 151251 – 151254).</p><p>Other specimens examined. Japan, Honshu, Niigata: 1 male, 5 females, same data as holotype. Yamagata: 1 male, Nagai-shi, Nogawa, 21.viii.1996, T. Hosoe. Miyagi: 1 male, 1 female, Sendai-shi, near Futakuchi-onsen, hygropetric zone, 7.vi.1974, T. Hattori; 1 male, 1 female, same locality, 1.vi.1994, T. Ito. Kanagawa: 1 male, Yamakita-machi, Shiroishi-zawa, 15.vi.1984, T. Nozaki; 2 males, 2 females, Kiyokawa-mura, Shiomizu-gawa, 690 m, 5.vi.1994, T. Hattori. Aichi: 3 males, Shinshiro-shi, Shimoyoshida, Adera, hygropetric zone on walking trail to Adera-no-taki Fall, 27.vii.2012, H. Nishimoto.</p><p>Etymology. The specific name refers to the name of the type locality.</p><p>Distribution (Fig. 21). Japan (northen and central Honshu).</p><p>Habitat. Adults were collected from small streams in forests.</p><p>Japanese name. Niigata-sunatsutsu-tobikera.</p></div>	https://treatment.plazi.org/id/C213FF65FFF2F64B519FFA17FD1EFA3B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ito, Tomiko	Ito, Tomiko (2016): The Lepidostoma coreanum Species Complex (Trichoptera, Lepidostomatidae) in the Asian Far East. Zootaxa 4061 (4): 397-417, DOI: 10.11646/zootaxa.4061.4.5
C213FF65FFFEF64E519FFA4AFD1DFC09.text	C213FF65FFFEF64E519FFA4AFD1DFC09.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lepidostoma hattorii	<div><p>Lepidostoma hattorii sp. nov.</p><p>(Figs. 12–17, 21)</p><p>Dinarthrum coreanum: Ito 1998, male, Japan (Honshu: Yamanashi, Shizuoka), in part, misidentification. Morita 2002, 37, Japan (Honshu: Mie). Morita 2003, 26, Japan (Honshu: Mie), misidentification.</p><p>Lepidostoma coreanum: Morita 2009, 140, Japan (Honshu: Mie), misidentification.</p><p>Lepidostoma sp.: Kawase &amp; Hayashi 2010, 86, Japan (Honshu: Shimane).</p><p>Diagnosis. Males of this species are similar to those of Lepidostoma niigataense sp. nov., but clearly discriminated from the later by triangular mesal arms and reduced lateral and ventral arms of tergum X. Females of this species are also similar to those of L. niigataense and difficult to discriminate.</p><p>Male (Figs. 12–17). Type specimens (Fig. 12). Antennal length indeterminate due to specimen condition, each scape setose, thick and long, 1.5 times as long as head, with short mesal process near middle of scape. Maxillary palps very setose, unsegmented, comb-like, 0.6 mm long. Labial palps each 3-segmented, slender, 1.0 mm long in total.</p><p>Wings covered with fine setae, short scales, and spindle-like setae; spindle-like setae lining cubital area; forewings each with apical forks 1 and 2 and discoidal cell, Cu1 fused to Mp at apical 1/4; hind wings each with apical fork 1, Cu1 fused to Mp at apical 1/5, Cu2 short and strongly curved caudally, and vein A lacking; forewings and hind wings each 6.2–6.5 mm long and 5.0– 5.6 mm long, respectively (n=2).</p><p>Genitalia. Abdominal segment IX short, dorsum longer than venter. Segment X composed of mesal arms and lateral arms; mesal arms long, triangular in lateral and dorsal views with subacute apices; lateral arms absent or reduced to low lobes; ventral arms completely absent. Inferior appendages each composed of main article, dorsal hook, and inner hook; basal half of main article broad in lateral and ventral views with serrate mesal and posterior margins; apical half of main article comb-like, directed mesocaudad; dorsal hook bar-like, directed caudad, slightly curved mesad, slightly shorter than main article; inner hook much shorter than dorsal hook, bar-like, curved mesad. Phallic apparatus thick, membranous in apical half with inner U-shaped sclerite, paramere absent.</p><p>Variation (Figs. 13–17). Specimens collected from all 5 Japanese localities show some variations. In the males from Ishikawa (Fig. 13) and Shizuoka (Fig. 14), the genitalia and maxillary palps are almost identical to those of males from the type locality (Fig. 12), but wing venation is different from that of the later, with the thyridial cell present in each forewing and Cu2 gently curved to marginal vein in hind wings.</p><p>In the males from Aichi (Fig. 15) and Mie (Fig. 16), lateral arms are almost completely absent (Fig. 15 F) or represented by short slender lobes (Fig. 15 D, E, Fig. 16 C), and Cu2 and A are absent in hind wings.</p><p>In a male from Shimane (Fig. 17), lateral and ventral arms are completely absent; cross vein m-cu present in each forewing.</p><p>Female (Fig. 12 H–J). Type specimens (Fig. 12 H–J): Antennal length indeterminate due to specimen condition, each scape setose, thick, long and 2 times as long as head, mesal process absent. Maxillary palps each 5-segmented and about 0.8 mm long, labial palps each 3-segmented and 0.6 mm long, all palps slender.</p><p>Wings covered with short fine setae, venation typical of other species of Lepidostomatidae; forewings and hind wings each 6.0– 6.2 mm long and 5.0– 5.3 mm long, respectively (n=2).</p><p>Lateral ends of tergite VIII broad. Lateral plates indistinct. Subgenital plate subquadrate with convex posterior margin and slightly concave anterior margin. Vaginal apparatus oval and slightly sclerotized.</p><p>Immature stages. Unknown.</p><p>Holotype: Japan, Honshu, Yamanashi: 1 male, Fujikawa-machi, Kajikazawa, Oyama-gawa, hygropetric zone, 35˚31’00”N, 138˚22’50”E, 750 m, 9.vi.1996, T. Hattori (CBM-ZI 151255).</p><p>Paratypes. 2 males, 2 females, same data as holotype (CBM-ZI 151256 – 151259).</p><p>Other specimens examined. Japan, Honshu, Yamanashi: 1 male, 7 females, same data as holotype; 6 males, 3 females, Hokuto-shi, Hakushu-cho, Ojiro-gawa, 800– 100 m, 13.vi.1992, T. Hattori; 1 male, 2 females, Hokutoshi, Hakushu-cho, Shirasu, 1170 m, 10.vi.2010, T. Hattori. Shizuoka: 1 male, 2 females, Izu-shi, Jizodo, Mt. Amagi-san, 825 m, 16.vi.1996, T. Hattori; 1 male, Yokozawa, 550 m, 4.vi.2001, T. Hattori; 1 male, Yokozawa, Mizumine, small water flow, 1100 m, 6.vi.2003, T. Hattori; 1 male, 2 females, Kawane-honcho, Senzu, Sumatakyo, 550 m, 4.vi.2007, T. Hattori. Ishikawa: 7 males, 10 females, Kashima-cho, Kogane-dani, 16.v.2000, H. Kawase. Aichi: 3 males, Shinshiro-shi, Shimoyoshida, Adera, hygropetric zone on walking trail to Adera-no-taki Fall, 7.v.2010, H. Nishimoto. Mie: 4 males, Daian-cho, Ishigure, 8–12.vi.2001, H. Morita; 1 male, Yokkaichi-shi, Mizusawa, Kanmuri-yama, 5.vii.2009, H. Morita (deposited in the personal collection of H. Morita). Shimane: 1 male, Okuizumo-cho, Mt. Sentsu-zan, Karasuyama-tozanguchi, 16–28.v.2007, M. Hayashi.</p><p>Etymology. The species is dedicated to Mr. Toshio Hattori, Shizuoka, who gifted me many valuable specimens, including specimens of this species.</p><p>Distribution (Fig. 21). Japan (central and western Honshu).</p><p>Habitat. Adults were collected from beside small streams and hygropetric zones in forests.</p><p>Japanese name. Hattori-sunatsutsu-tobikera.</p></div>	https://treatment.plazi.org/id/C213FF65FFFEF64E519FFA4AFD1DFC09	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ito, Tomiko	Ito, Tomiko (2016): The Lepidostoma coreanum Species Complex (Trichoptera, Lepidostomatidae) in the Asian Far East. Zootaxa 4061 (4): 397-417, DOI: 10.11646/zootaxa.4061.4.5
C213FF65FFFBF650519FFC64FD1AF84B.text	C213FF65FFFBF650519FFC64FD1AF84B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lepidostoma yuwanense	<div><p>Lepidostoma yuwanense sp. nov.</p><p>(Figs. 18–19, 21)</p><p>Diagnosis. Males of this species are similar to those of Lepidostoma coreanum (Kumanski &amp; Weaver 1992), but are clearly discriminated from the later by the ventral view of the main articles of the inferior appendages: subquadrate with small rectangle projections on latero-caudal corners in this species (Fig. 18 G), but semicircular without any latero-caudal projections in L. coreanum (Fig. 1 G). Females of this species are also similar to those of L. coreanum and difficult to discriminate from each other.</p><p>Male (Figs. 18–19). Type specimens (Fig. 18). Antennae about 5.5–6.0 mm long, each with scape setose, thick and long, about 1.5 times as long as head, with short mesal process near mid length. Maxillary palps extremely setose, each 2-segmented, 0.8 mm long in total, basal segment curved anterad in basal half, with tuft of very long setae apically, long apical segment emerging near middle of basal segment. Labial palps each 3-segmented, slender, 0.8 mm long in total.</p><p>Wings covered with fine setae and short scales; spindle-like setae lining Cu and encircling cubital area; forewings each with apical forks 1 and 2, discoidal cell and arculus, Cu1 and Cu2 complex in caudal area (Fig. 18 C); hind wings each with apical fork 1, vein A absent; forewings and hind wings each 5.2–5.7 mm long and 4.5– 4.5 mm long, respectively (n=3).</p><p>Genitalia. Abdominal segment IX short, dorsum slightly longer than venter. Segment X composed of mesal, lateral and ventral arms; mesal arms long, curved slightly ventrad near base and directed caudad in apical 1/3, lateral arms directed caudad and shorter than mesal arms, ventral arms long, directed caudolaterad in dorsal view; 3 pairs of arms subacute apically. Inferior appendages each composed of main article, dorsal hook, and inner hook; basal half of main article broad in lateral view, subquadrate in ventral view with serrate posterior margin and small rectangle projections at posterolateral corner; apical half of main article curved mesad, comb-shaped in lateral view; dorsal hook bar-like with spiny apex, curved mesad in ventral view; inner hook much shorter than dorsal hook, bar-like, directed laterad in ventral view. Phallic apparatus thick, membranous in apical half with inner Ushaped sclerite, paramere absent.</p><p>Variation (Fig. 19). The males collected from Okinawa-jima show some variations. The ventral arms of segment X are long, exceeding the tips of the mesal arms in lateral view. A short vein is present between C and Sc in each forewing, crossvein r-m present in each hind wing.</p><p>Female (Fig. 18 H–J). Type specimens. Antennae about 6 mm long, each with scape thick, long and 2 times as long as head, mesal process absent. Maxillary palps each 5-segmented and about 0.7 mm long, labial palps each 3- segmented and 0.4 mm long, all palps slender.</p><p>Wings covered with short fine setae, venation typical of other species of Lepidostomatidae; forewings and hind wings each 5.2–5.4 mm long and 4.5–4.8 mm long, respectively (n=2).</p><p>Lateral ends of tergite VIII broad. Lateral plates weakly sclerotized, subtriangular. Subgenital plate subquadrate slightly concave anterior margin and almost straight posterior margin with small shallow central notch. Vaginal apparatus oval and slightly sclerotized.</p><p>Immature stages. Unknown.</p><p>Holotype: Japan, Ryukyu Islands, Amami-Ooshima: 1 male, Uken-son, Yuwan, small water flow on trail to Mt. Yuwan-dake, 28˚17’22”N, 129˚17’31”E, 600 m, 9.vi.1996, T. Ito (CBM-ZI 151260).</p><p>Paratypes. 2 males, 2 females, same data as holotype (CBM-ZI 151261 – 151264).</p><p>Other specimens examined. Japan, Ryukyu Islands, Amami-Ooshima: 4 males, 3 females, same data as holotype. Okinawa-jima: 2 males, Nago-shi, Genka, hygropetric zone near Hogen-hashi, 22–24.xi.2010, T. Ito; 7 males, 2 females, same locality, 8.iv.2011, T. Ito; 1 male, same locality, 17–19.x.2014, T. Ito.</p><p>Etymology. The specific name refers to the name of the type locality.</p><p>Distribution (Fig. 21). Japan (Ryukyu Islands: Amami-Ooshima, Okinawa-jima).</p><p>Habitat. Adults were collected from beside small water flows and hygropetric zones in forests.</p><p>Japanese name. Yuwan-sunatsutsu-tobikera.</p></div>	https://treatment.plazi.org/id/C213FF65FFFBF650519FFC64FD1AF84B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ito, Tomiko	Ito, Tomiko (2016): The Lepidostoma coreanum Species Complex (Trichoptera, Lepidostomatidae) in the Asian Far East. Zootaxa 4061 (4): 397-417, DOI: 10.11646/zootaxa.4061.4.5
C213FF65FFE6F655519FF8A6FD16FE24.text	C213FF65FFE6F655519FF8A6FD16FE24.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lepidostoma ishigakiense	<div><p>Lepidostoma ishigakiense sp. nov.</p><p>(Figs. 20, 21)</p><p>Diagnosis. Males of this species are similar to those of Lepidostoma niigataense sp. nov. (Figs. 7–11), but are clearly discriminated from the later by the small rectangular projection on the laterocaudal corner the main article of each inferior appendage (Fig. 20 E, G).</p><p>Male (Fig. 20). Antennae about 5.0 mm long, each with scape setose, thick and long, about 1.5 times as long as head, short mesal process near middle. Maxillary palps setose, each 2-segmented, 0.8 mm long in total, long basal segment covered with short setae, short apical segment densely covered with long setae. Labial palps each 3- segmented, slender, 0.8 mm long in total.</p><p>Wings covered with fine setae, short scales, and spindle-like setae; spindle-like setae lining cubital area; forewings each with apical forks 1 and 2, discoidal and thyridial cells, and arculus, costa curved anteriorly, venation of caudal area complex (Fig. 20 C); hind wings each with apical fork 1, venation of caudal area complex (Fig. 20 D); forewings and hind wings each 4.2–4.6 mm long and about 4.0 mm long, respectively (n=3).</p><p>Genitalia. Abdominal segment IX short. Segment X composed of mesal and lateral arms; mesal arms elongate posteriorly, lateral arms subtriangular, about 1/2 as long as mesal arms, both pairs of arms blunt apically. Ventral arms completely absent. Inferior appendages each composed of main article, dorsal hook, and inner hook; basal half of main article broad in lateral and ventral views with serrate posterior margin and small rectangular projection at postero-lateral corner; apical half of main article bar-like, directed posteromesad with spiny apex; dorsal hook directed laterad with subacute setose apex; inner hook much shorter than dorsal hook, bar-like, directed lateroposterad in ventral view. Phallic apparatus thick, membranous in apical half, with inner U-shaped sclerite, paramere absent.</p><p>Female and immature stages. Unknown.</p><p>Holotype: Japan, Ryukyu Islands, Ishigaki-jma: 1 male, Ishigaki-shi, Shiramizu, small tributary of Naguragawa, 24˚24’46”N, 124˚09’51”E, 15 m, 17.x.2005, T. Ito (CBM-ZI 151265).</p><p>Paratypes. 2 males, same data as holotype (CBM-ZI 151266 – 151267).</p><p>Etymology. The specific name refers to the name of the type locality.</p><p>Distribution (Fig. 21). Japan (Ryukyu Islands: Ishigaki-jima). Known from only the type locality.</p><p>Habitat. Adults were collected beside a small water flow (30 cm wide, 1 cm deep) in a forest.</p><p>Japanese name. Ishigaki-sunatsutsu-tobikera.</p></div>	https://treatment.plazi.org/id/C213FF65FFE6F655519FF8A6FD16FE24	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ito, Tomiko	Ito, Tomiko (2016): The Lepidostoma coreanum Species Complex (Trichoptera, Lepidostomatidae) in the Asian Far East. Zootaxa 4061 (4): 397-417, DOI: 10.11646/zootaxa.4061.4.5
