taxonID	type	description	language	source
DDD8972326945EB1835C7E09E364EC59.taxon	type_taxon	Type species. Diploexochus echinatus Brandt, 1833, by monotypy (see Schmidt and Leistikow 2004).	en	Borja-Arrieta, Ricardo, López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Gutierrez-Estrada, Miguel, Campos-Filho, Ivanklin Soares, Navas-S., Gabriel R., Bichuette, Maria Elina (2025): From surface to caves: new species of Diploexochus Brandt, 1833 (Oniscidea, Armadillidae) from Colombia, with the description of the first troglobitic species. ZooKeys 1223: 263-288, DOI: 10.3897/zookeys.1223.133267
D66510F7C24956A2911310FF7BACEAD8.taxon	description	Figs 1, 2, 3, 4, 8 A, 13 A	en	Borja-Arrieta, Ricardo, López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Gutierrez-Estrada, Miguel, Campos-Filho, Ivanklin Soares, Navas-S., Gabriel R., Bichuette, Maria Elina (2025): From surface to caves: new species of Diploexochus Brandt, 1833 (Oniscidea, Armadillidae) from Colombia, with the description of the first troglobitic species. ZooKeys 1223: 263-288, DOI: 10.3897/zookeys.1223.133267
D66510F7C24956A2911310FF7BACEAD8.taxon	distribution	Distribution. This species is known only from its type locality in Tropical Dry Forest (TDF) and Andean forest of Sierra Nevada de Santa Marta (Fig. 1).	en	Borja-Arrieta, Ricardo, López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Gutierrez-Estrada, Miguel, Campos-Filho, Ivanklin Soares, Navas-S., Gabriel R., Bichuette, Maria Elina (2025): From surface to caves: new species of Diploexochus Brandt, 1833 (Oniscidea, Armadillidae) from Colombia, with the description of the first troglobitic species. ZooKeys 1223: 263-288, DOI: 10.3897/zookeys.1223.133267
A3B0B48A9AFE52E9A07AD8B88DC400DB.taxon	description	Figs 1, 5, 6, 7, 8 B, 13 B	en	Borja-Arrieta, Ricardo, López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Gutierrez-Estrada, Miguel, Campos-Filho, Ivanklin Soares, Navas-S., Gabriel R., Bichuette, Maria Elina (2025): From surface to caves: new species of Diploexochus Brandt, 1833 (Oniscidea, Armadillidae) from Colombia, with the description of the first troglobitic species. ZooKeys 1223: 263-288, DOI: 10.3897/zookeys.1223.133267
A3B0B48A9AFE52E9A07AD8B88DC400DB.taxon	description	Description. Maximum body length: male 8 mm, female 9 mm. Body outline as in Fig. 5 A. Color dark brown with typical muscular insertions (Figs 8 B, 13 B); upper portion of tubercles randomly depigmented; pereonite 1 epimera anterior corner, pereonites 2 – 7 paramedian portions, pereonites 3 and 4 epimera, and pleonites 3 – 5 epimera randomly depigmented. Endoantennal conglobation (Figs 5 A, 8 B, 13 B). Dorsum covered with large triangular tubercles, arranged as follows (Fig. 5 A, B): vertex of cephalon with 12 tubercles in three rows; pereonite 1 with 20 tubercles; pereonites 2 – 6 with 16 tubercles; pereonite 7 with 15 tubercles; pleonites 3 – 5 and telson with two paramedian tubercles. Pereonites 1 – 7 epimera with one line of noduli laterales per side inserted on outer surface of second tubercle of posterior row (Fig. 5 A). Dorsal surface with short and narrow semi-circular scale-setae (Fig. 5 C). Cephalon (Fig. 5 D – F) with frontal shield prominent, distinctly protruding above vertex; eyes with 20 – 21 ommatidia. Pereonites 1 – 7 epimera flattened and slightly directed outwards; pereonite 1 strongly grooved on lateral margin, inner lobe of schisma rounded, extending beyond posterior margin of outer lobe (Fig. 5 G, H), pereonite 2 with triangular and narrow ventral lobe directed outwards, not extending beyond posterior margin of epimera; pereonites 4 – 7 with oblique ventral ridge (Fig. 5 F – H). Pleonites 3 – 5 (Fig. 5 I, J) with epimera well developed, rectangular and slightly directed outwards. Telson (Fig. 5 I) with proximal part slightly broader than distal part, dorsum slightly depressed, distal margin straight. Antennula (Fig. 5 K) of three articles, proximal and distal articles subequal in length, distal article with four aesthetascs inserted sub-apically. Antenna (Fig. 5 L) short, not surpassing posterior margin of pereonite 1 when extended backwards; flagellum of two articles, distal article about twice as long as first bearing one row of two lateral aesthetascs. Mandibles with molar penicil semi-dichotomized; left mandible (Fig. 6 A) with 2 + 1 penicils, right mandible (Fig. 6 B) with 1 + 1 penicils. Maxillula (Fig. 6 C) inner endite with two stout penicils, distal margin bearing fringe of thin setae; outer endite of 4 + 6 teeth simple. Maxilla (Fig. 6 D) inner lobe rounded covered with thick setae; outer lobe rounded, three times as wide as inner lobe, covered with thin setae. Maxilliped (Fig. 6 E) basis rectangular bearing sparse setae; palp with two distinct setae on basal article; endite subrectangular, medial seta overpassing distal margin, distal margin bearing two setae. Pereopods 1 – 7 merus and carpus bearing setae on sternal margin, not sparse appearance; carpus 1 with distal setae cleft at apex; ungual seta and dactylar organ simple, both surpassing outer claw. Uropod (Fig. 6 F) protopod flattened, enlarged on basal part, distal part subrectangular, medial margin concave; exopod short inserted dorsally near medial margin bellow distinct lobe, lobe not extending beyond medial margin; endopod club-shaped bearing many short setae on distal part. Pleopods 1 – 5 exopods with monospiracular respiratory structures. Male. Pereopods 1 – 7 (Fig. 7 A, B) without particular modifications. Genital papilla as in Fig. 9 C. Pleopod 1 (Fig. 7 C) exopod triangular, as wide as long, outer and inner margin bearing many small setae, distal and proximal outer parts triangular; endopod about twice as long as exopod. Pleopod 2 (Fig. 7 D) exopod triangular, outer margin strongly concave bearing many setae; endopod slightly longer than exopod. Pleopod 3 – 5 exopods as in Fig. 7 E – G.	en	Borja-Arrieta, Ricardo, López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Gutierrez-Estrada, Miguel, Campos-Filho, Ivanklin Soares, Navas-S., Gabriel R., Bichuette, Maria Elina (2025): From surface to caves: new species of Diploexochus Brandt, 1833 (Oniscidea, Armadillidae) from Colombia, with the description of the first troglobitic species. ZooKeys 1223: 263-288, DOI: 10.3897/zookeys.1223.133267
A3B0B48A9AFE52E9A07AD8B88DC400DB.taxon	etymology	Etymology. The new species is named after the Vallenato music singer Diomedes Díaz Maestre, also known as “ El Cacique de la Junta ”.	en	Borja-Arrieta, Ricardo, López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Gutierrez-Estrada, Miguel, Campos-Filho, Ivanklin Soares, Navas-S., Gabriel R., Bichuette, Maria Elina (2025): From surface to caves: new species of Diploexochus Brandt, 1833 (Oniscidea, Armadillidae) from Colombia, with the description of the first troglobitic species. ZooKeys 1223: 263-288, DOI: 10.3897/zookeys.1223.133267
A3B0B48A9AFE52E9A07AD8B88DC400DB.taxon	distribution	Distribution. This species is known only from the type locality at Cerro Bañaderos, Hatonuevo, La Guajira, which is included into a TDF area (Fig. 1).	en	Borja-Arrieta, Ricardo, López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Gutierrez-Estrada, Miguel, Campos-Filho, Ivanklin Soares, Navas-S., Gabriel R., Bichuette, Maria Elina (2025): From surface to caves: new species of Diploexochus Brandt, 1833 (Oniscidea, Armadillidae) from Colombia, with the description of the first troglobitic species. ZooKeys 1223: 263-288, DOI: 10.3897/zookeys.1223.133267
706611444DB854F3898BA892503DEBD3.taxon	description	Figs 1, 9, 10, 11, 12, 13 C	en	Borja-Arrieta, Ricardo, López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Gutierrez-Estrada, Miguel, Campos-Filho, Ivanklin Soares, Navas-S., Gabriel R., Bichuette, Maria Elina (2025): From surface to caves: new species of Diploexochus Brandt, 1833 (Oniscidea, Armadillidae) from Colombia, with the description of the first troglobitic species. ZooKeys 1223: 263-288, DOI: 10.3897/zookeys.1223.133267
706611444DB854F3898BA892503DEBD3.taxon	description	Description. Maximum body length: male 2.7 mm, female 4.2 mm. Body outline as in Fig. 9 A. Color faintly brown; body pigments not discernible in vivo (Fig. 12 D). Endoantennal conglobation (Figs 9 A, 12 D, 13 C). Dorsum covered with large triangular tubercles, arranged as follows (Fig. 9 A, B): vertex of cephalon with 10 tubercles in two rows; pereonite 1 with 19 tubercles; pereonites 2 – 6 with 13 tubercles; pereonite 7 with 11 tubercles; pleonites 3, 4, and telson with two median tubercles. Pereonites 1 – 7 epimera with one line of noduli laterales per side inserted on outer surface of second tubercle of posterior row (Fig. 9 A). Dorsal surface densely covered with elongated semi-circular scale-setae, conferring pilous aspect (Fig. 9 C). Cephalon (Fig. 9 D – F) with prominent frontal shield, distinctly protruding above vertex; eyes with four ommatidia. Pereonites 1 – 7 epimera flattened and slightly directed outwards; pereonite 1 grooved on posterior lateral margin, inner lobe of schisma rounded and extending beyond posterior margin of outer lobe. Pereonite 2 with triangular ventral lobe rounded at apex, extending beyond posterior margin of epimera; pereonites 3 and 5 – 7 with well-marked ventral ridge (Fig. 9 A, F, G). Pleonites 3 – 5 epimera (Fig. 9 H, I) well developed, sub-rectangular, and directed outwards, distal margins rounded. Telson (Fig. 9 H) with proximal part broader than distal part, dorsum slightly depressed, distal margin sinuous. Antennula (Fig. 9 J) of three articles, proximal and distal subequal in length, distal article with five stout aesthetascs inserted sub-apically. Antenna (Fig. 9 K) short, not surpassing posterior margin of pereonite 1 when extended backward; flagellum of two articles, the distal about three times as long as first, bearing one row of two lateral aesthetascs. Mandibles with molar penicil semi-dichotomized; left mandible (Fig. 10 A) with 2 + 1 penicils, right mandible (Fig. 10 B) with 1 + 1 penicils. Maxillula (Fig. 10 C) inner endite with two stout penicils, distal margin bearing thin setae; outer endite with 4 + 6 teeth simple. Maxilla (Fig. 10 D) inner lobe rounded covered with thick setae; outer lobe rounded three times as wide as inner lobe covered with thin setae. Maxilliped basis (Fig. 10 E) rectangular, bearing sparse setae; palp with two distinct setae on basal article; endite subrectangular, medial seta surpassing distal margin, distal margin bearing one seta. Pereopods 1 – 7 merus and carpus with sparse setae on sternal margin; carpus 1 with distal seta cleft at apex; ungual seta and dactylar organ simple not surpassing outer claw. Uropod (Fig. 10 F) protopod flattened, enlarged on basal part, distal part elongated and sub-rectangular, distal margin rounded, medial margin concave with L-shaped appearance; exopod as long as endopod inserted dorsally near medial margin bellow distinct lobe, lobe not extending beyond medial margin; endopod short bearing many short setae. Pleopods 1 – 5 exopods with monospiracular respiratory structures. Male. Pereopods 1 – 7 (Fig. 11 A, B) without particular modifications. Genital papilla as in Fig. 11 C. Pleopod 1 (Fig. 11 D) exopod hour-glass shaped, twice as wide as long, inner portion rounded, outer portion triangular, distal and proximal margins narrow on middle; endopod about three times as long as exopod. Pleopod 2 (Fig. 11 E) exopod triangular, outer margin strongly concave; endopod slightly longer than exopod. Pleopod 3 – 5 exopods as in Fig. 11 F – H.	en	Borja-Arrieta, Ricardo, López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Gutierrez-Estrada, Miguel, Campos-Filho, Ivanklin Soares, Navas-S., Gabriel R., Bichuette, Maria Elina (2025): From surface to caves: new species of Diploexochus Brandt, 1833 (Oniscidea, Armadillidae) from Colombia, with the description of the first troglobitic species. ZooKeys 1223: 263-288, DOI: 10.3897/zookeys.1223.133267
706611444DB854F3898BA892503DEBD3.taxon	etymology	Etymology. Latin: troglo + bio = cave-dwelling. The new name of the species is an adjective that refers to the troglobitic category of the species.	en	Borja-Arrieta, Ricardo, López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Gutierrez-Estrada, Miguel, Campos-Filho, Ivanklin Soares, Navas-S., Gabriel R., Bichuette, Maria Elina (2025): From surface to caves: new species of Diploexochus Brandt, 1833 (Oniscidea, Armadillidae) from Colombia, with the description of the first troglobitic species. ZooKeys 1223: 263-288, DOI: 10.3897/zookeys.1223.133267
706611444DB854F3898BA892503DEBD3.taxon	distribution	Distribution. This species is known only from the type locality at Roca Madre Cave, Sucre, inserted on TDF area (Fig. 1).	en	Borja-Arrieta, Ricardo, López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Gutierrez-Estrada, Miguel, Campos-Filho, Ivanklin Soares, Navas-S., Gabriel R., Bichuette, Maria Elina (2025): From surface to caves: new species of Diploexochus Brandt, 1833 (Oniscidea, Armadillidae) from Colombia, with the description of the first troglobitic species. ZooKeys 1223: 263-288, DOI: 10.3897/zookeys.1223.133267
