taxonID	type	description	language	source
C046B71FCB2A0D33FEC85BAEFEEB8D78.taxon	materials_examined	Type species: Gracilipsodes psocopterus Sykora, 1967, by original designation. Revised generic description, males Adult: Body and wing colour yellowish brown to dark greyish brown (denuded in alcohol). Head: Antennae with broadly oval scape, about as long as eye diameter (Fig. 1 A). Frontal setal warts (f. w.) oval (Fig. 1 B); anterior setal wart (a. w.) narrowly triangular or diamond shaped (Fig. 1 B, C); anterolateral (al. w.) and posterolateral setal (pl. w.) warts oval (Fig. 1 C). Maxillary palps five-segmented (Fig. 1 A, B), labial palps three-segmented (Fig. 1 A). Thorax: Prothorax with one pair of transversely elongate pronotal setal warts (Fig. 1 C). Legs slender; tibial spur (sp.) formula 0, 1, 1 – 2, 2, 2 (e. g. Fig. 1 D – H); hind tibia with 15 – 35 blackish spines (s.) (Fig. 1 G). Forewing (Fig. 2 A) narrow, slightly widening towards rounded apex, forks 1 and 5 present; discoidal cell (dc) about as long as thyroidal cell (tc). Hind wing (Fig. 2 D) discoidal cell absent; crossvein r 1 – r 2 absent; crossvein r – m straight or convex; forks 1, 3, and 5, 1 and 5, or only 5, present; anterior edge with rasp-like, basal hairs (Fig. 2 B) and 15 – 25 hamuli present past midway (Fig. 2 C). Nygma (n): present in both wings in apical cell posterior to R 4 (Fig. 2 A, D). Male genitalia (Fig. 2 E – I): Segment IX (t. IX and s. IX) annular, laterally setose, narrowest at base of superior appendages (s. app.) or ventrally; in lateral view tergite IX (t. IX) narrower to wider than sternite IX (s. IX) in lateral view. Superior appendages setose, flattened dorsoventrally, and as long as or longer than tergum X (t. X). Tergum X with median process bifurcated or entire along its length, membranous, and bearing small spines; pair of pre-apical, lateral, sclerotized processes (pl. p.) either absent or present. Inferior appendages bipartite, setose; basal part (if. b.) produced into posteromesal lip, with small, strong spines along dorsal margin; apical part of inferior appendage (if. ap.) long, digitate, with 25 – 60 mesal spines, mesal process with between four and nine mesal spines. Harpago (h.) absent or small. Phallic apparatus simple, phallicata basally tubular, fused with phallobase, curving to various degrees ventrad along its length; pair of sclerotized, ventrally produced lateral processes present apically; phallotremal sclerite (ph. sc.) U-shaped. Inclusive species: Gracilipsodes aoupiniensis sp. nov.; Gracilipsodes aureus sp. nov.; Gracilipsodes aurorus sp. nov.; Gracilipsodes grandis sp. nov.; Gracilipsodes koghiensis sp. nov.; Gracilipsodes lanceolatus sp. nov.; Gracilipsodes robustus sp. nov.; Gracilipsodes psocopterus Sykora (1967); Gracilipsodes similis Ward (2001).	en	Malm, Tobias, Johanson, Kjell A. (2008): Revision of the New Caledonian endemic genus Gracilipsodes (Trichoptera: Leptoceridae: Grumichellini). Zoological Journal of the Linnean Society 153 (3): 425-452, DOI: 10.1111/j.1096-3642.2008.00403.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00403.x
C046B71FCB2A0D35FCDB5CD1F9218E6A.taxon	diagnosis	Diagnosis: This species is distinguished from G. lanceolatus sp. nov., G. grandis sp. nov.; G. psocopterus, and G. similis by the tibial spur formula of 2, 2, 2, and by the presence of a pair of pre-apical lateral processes of tergum X originating at the apical half of the segment. It is separated from G. aureus sp. nov. and G. robustus sp. nov. by having more hind tibial spines, in the genitalia, by the longer lateral processes of tergum X, and in the forewing, by the longer crossvein m – cu being sharply angled to M. It is separated from G. aurorus sp. nov. by lighter body colour, the slit tergum X apex, and the higher number of hind tibial and mesal spines on the apical part of the inferior appendage. It is distinguishable from G. koghiensis sp. nov. by the shorter superior appendages and shallower apical slit of tergum X. Description, male: Body and wings brown (in alcohol). Tibial spur formula: 2, 2, 2. Hind tibiae each with 26 small black spines (N = 1). Forewing (Fig. 2 A): length 7.9 mm (N = 1), apically rounded; forks 1 and 5 present; crossvein m – cu about as long as crossvein	en	Malm, Tobias, Johanson, Kjell A. (2008): Revision of the New Caledonian endemic genus Gracilipsodes (Trichoptera: Leptoceridae: Grumichellini). Zoological Journal of the Linnean Society 153 (3): 425-452, DOI: 10.1111/j.1096-3642.2008.00403.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00403.x
C046B71FCB2A0D35FCDB5CD1F9218E6A.taxon	description	2 (1) Protibiae each with one spur; pair of pre-apical, lateral processes on tergum X present (Fig. 5 C); hindwing fork 3 present (Fig. 5 B) ......................................................................................................... grandis sp. nov. – Protibiae each with either no or two spurs; pair of lateral processes on segment X absent (Figs 7 C, 9 C); hindwing fork 3 absent (Fig. 5 B) ..................................................................................................... 3 3 (2) Tibial spur formula 2, 2, 1; tergum X uniform, membranous, and tubular (Fig. 7 D, E); hindwing fork 1 absent (Fig. 7 B) ............................................................................................................... lanceolatus sp. nov. – Tibial spur formula 0, 1, 1; tergum X forming two long, straight sclerotized spines (Fig. 9 D, E); hindwing fork 1 present (Fig. 9 B) .......................................................................................................................... 4 4 (3) Inferior appendage apical part triangular in lateral view, apex truncate (Fig. 9 C) ........................................ .................................................................................................................. psocopterus, Sykora, 1967 – Inferior appendage apical part ovoid in lateral view, apex produced (Fig. 10 C) ......................................... ....................................................................................................................... similis Ward (2001) 5 (1) Pre-apical lateral processes on tergum X shorter than median process (Fig. 3 C); forewing crossvein m – cu shorter than crossvein r – m, almost at right angle to M (Fig. 3 A) ....................................................................... 6 – Pre-apical lateral processes on tergum X about as long as median process (Fig. 6 C); forewing crossvein m – cu as long as, or longer than, crossvein r – m, and at sharp angle to M (Fig. 6 A) ......................................... 7 6 (5) Superior appendages longer than tergum X; apical part of inferior appendage parallel sided in lateral view (Fig. 3 C) ..................................................................................................................... aureus sp. nov. – Superior appendages as long as tergum X; apical part of inferior appendage slightly club-shaped in lateral view (Fig. 8 C) ................................................................................................................ robustus sp. nov. 7 (5) Body dark greyish brown colour; hind tibiae each with at most 20 spines; apical part of inferior appendage with at most 30 mesal spines (Fig. 4 C) .................................................................................. aurorus sp. nov. – Body brown colour; hind tibiae each with more than 20 spines; apical part of inferior appendage with more than 35 mesal spines (Fig. 6 C) .............................................................................................................. 8 8 (7) Hindwing crossvein r – m curved (Fig. 2 D); superior appendages about as long as tergum X; tergum X shallowly slit at apex (Fig. 2 F, G) .......................................................................................... aoupiniensis sp. nov. – Hindwing crossvein r – m straight (Fig. 6 B); superior appendages longer than tergum X; tergum X deeply slit at apex (Fig. 6 D, E) .................................................................................................. koghiensis sp. nov. r – m, sharply angled with M. Hindwing (Fig. 2 D): length 5.5 mm (N = 1), apex nearly acute; forks 1, 3, and 5 present; crossvein r – m convex; costa with about 23 hamuli. Genitalia: Segment IX annular, laterally setose, narrowest at bases of superior appendages; tergite IX about as broad as sternite IX in lateral view (Fig. 2 E); anterior margins of sternite IX slightly produced anterad at mid height (Fig. 2 E). Superior appendages setose, flattened dorsoventrally, about as long as tergum X, with apices rounded in dorsal view (Fig. 2 F). Tergum X membranous; median process entire, basally wide, apex widely rounded, concave dorsally in lateral view (Fig. 2 E), apex with shallow slit in dorsal and ventral views (Fig. 2 F, G), and bearing few, small setae; pre-apical lateral processes about as long as median process, produced at apical half, sclerotized, digitate, directed posterad, slightly curving ventrad (Fig. 2 E). Inferior appendages each bipartite, setose; basal part wide in lateral view (Fig. 2 E), mesally concave, produced posteromesally into a broad, distally tapering, apically rounded lip, bearing small dorsal spines (Fig. 2 E, G); apical part of each inferior appendage digitate, mesally curved, in lateral view, parallel sided and straight, apex rounded, with 50 mesal spines (Fig. 2 E); mesal process with five strong spines (Fig. 2 F). Harpago: small, weakly sclerotized (Fig. 2 E). Phallic apparatus simple (Fig. 2 H, I), phallicata basally tubular, apex truncate, fused with phallobase, bent 25 ° ventrally at mid length; apicoventral corner with pair of sclerotized, slender lateral processes oriented posteroventrad (Fig. 2 H); phallotremal sclerite U-shaped (Fig. 2 I). Holotype (male): New Caledonia, Province Nord, Mt Aoupinié fauna reserve, 24 – 28 November 2001, Malaise trap, loc # 19 (Johanson, Pape & Viklund). Distribution: Province Nord, Mt Aoupinié. Etymology: The name aoupiniensis is derived from the type locality, Mt Aoupinié. · Figure 2. Gracilipsodes aoupiniensis sp. nov. A, forewing. B, rasp-like basal hindwing hair, detail. C, hindwing hamulus, detail. D, hindwing. E, genitalia, lateral. F, genitalia, dorsal. G, genitalia, ventral. H, phallus, lateral. I, phallus, ventral. Abbreviations of wing structures: 1, fork 1; 3, fork 3; 5, fork 5; a, anal veins; b. h., basal rasp-like hairs; cu, cubitus; dc, discoidal cell; ha., hamuli; m, media; n, nygma; r, radius; r – m, crossvein r – m, sc, subcosta; tc, thyroidal cell. Abbreviations of genitalic structures: h., harpago; if. ap., inferior appendage apical part; if. b., inferior appendage basal part; ph. sc., phallothremal sclerite; pl. p., tergum X pre-apical lateral processes; s. app., superior appendage; s. IX, sternite IX; t. IX, tergite IX; t. X, tergum X.	en	Malm, Tobias, Johanson, Kjell A. (2008): Revision of the New Caledonian endemic genus Gracilipsodes (Trichoptera: Leptoceridae: Grumichellini). Zoological Journal of the Linnean Society 153 (3): 425-452, DOI: 10.1111/j.1096-3642.2008.00403.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00403.x
C046B71FCB2E0D37FF2558BAFEC78BC5.taxon	diagnosis	Diagnosis: This species is distinguished from G. lanceolatus sp. nov., G. grandis sp. nov., G. psocopterus, and G. similis by the tibial spur formula 2, 2, 2, and the presence of a pair of pre-apical lateral processes of tergum X, originating from the apical half of the segment. It is separated from G. aurorus sp. nov., G. koghiensis sp. nov., and G. aoupiniensis sp. nov. in the genitalia, by the shorter lateral processes of tergum X, and in the forewing, by the shorter crossvein m – cu oriented at almost a right angle to M. It is distinguishable from G. robustus sp. nov. by its slightly smaller size, in the genitalia, by the longer superior appendages, and by the apical part of the inferior appendage being slender and parallel-sided. Description, male: Body golden yellowish brown. Wings greyish brown. Tibial spur formula 2, 2, 2. Hind tibiae each with 15 – 19 small black spines (N = 3). Forewing (Fig. 3 A): length 6.8 – 7.2 mm (N = 3), apically rounded; forks 1 and 5 present; crossvein m – cu about as long as or slightly shorter than crossvein r – m, and at right angle with M. Hindwing (Fig. 3 B): length 5.0 – 5.2 mm (N = 3), apex nearly acute; forks 1, 3, and 5 present; crossvein r – m convex; costa with 20 – 23 hamuli. Genitalia: Segment IX annular, laterally setose, narrowest at bases of superior appendages; tergite IX about as broad as sternite IX in lateral view (Fig. 3 C). Superior appendages setose, flattened dorsoventrally, slightly longer than tergum X, apices rounded in dorsal view (Fig. 3 D). Tergum X membranous; median process entire, basally wide, apex rounded, dorsal margin slightly concave in lateral view (Fig. 3 C), apex with shallow notch in dorsal and ventral views (Fig. 3 D, E); pre-apical lateral process shorter than median process, produced at apical half, sclerotized, digitate, directed posterolaterad, and slightly curving dorsally (Fig. 3 C). Inferior appendages each bipartite, setose; basal part wide in lateral and ventral views (Fig. 3 C, D), mesally concave, produced posteromesally into a broad, apically rounded lip, bearing small dorsal spines (Fig. 3 C, E); apical part of each inferior appendage digitate, curving mesally, in lateral view parallel-sided, apex rounded, with 30 – 40 mesal spines (Fig. 3 C); mesal process with between five and seven strong spines (Fig. 3 D). Harpago small, slender, club-shaped, apex nearly acute (Fig. 3 C). Phallic apparatus simple (Fig. 3 F, G), phallicata basally tubular, fused with phallobase, apex truncate; bent about 45 ° ventrad at mid length; apical part with pair of sclerotized, broad lateral processes, produced ventrad, curved somewhat medially, and broadly rounded at apex (Fig. 3 F); phallotremal sclerite U-shaped (Fig. 3 G). Holotype (male): New Caledonia, Province Nord, Col d’Amieu, 20 km north of Sarraméa, 306 m a. s. l., small river, 21 ° 33.877 ′ S, 165 ° 49.922 ′ E, 2 December 2001, light trap, loc # 27 (Johanson, Pape & Viklund). Paratypes: One male, same data as holotype. One male, New Caledonia, Province Sud, Sarraméa, 220 m a. s. l., 19 November 2001, light trap, loc # 10 (Johanson, Pape & Viklund); one male, New Caledonia, Province Nord, Xwê Dachava Stream, Rembai Mt, 21 ° 34.854 ′ S, 165 ° 49.478 ′ E, 317 m a. s. l., 5 – 12 January 2004, Malaise trap, loc # 108 (Johanson). Distribution: Central Grande Terre. Etymology: From Latin, aureus, golden; named for the golden, yellowish brown body colour of the species.	en	Malm, Tobias, Johanson, Kjell A. (2008): Revision of the New Caledonian endemic genus Gracilipsodes (Trichoptera: Leptoceridae: Grumichellini). Zoological Journal of the Linnean Society 153 (3): 425-452, DOI: 10.1111/j.1096-3642.2008.00403.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00403.x
C046B71FCB2E0D3AFCB35B7CFAF78C63.taxon	diagnosis	Diagnosis: This species is distinguished from G. lanceolatus sp. nov., G. grandis sp. nov., G. psocopterus, and G. similis by the tibial spur formula 2, 2, 2 and the presence of a pair of pre-apical lateral processes of tergum X, originating from the apical half of the segment. It is separated from G. aureus sp. nov. and G. robustus sp. nov. in the genitalia, by the longer pre-apical lateral processes of tergum X, and in the forewing, by a longer crossvein m – cu, oriented at a sharp angle to M. It is separated from G. koghiensis sp. nov. and G. aoupiniensis sp. nov. by having a darker body colour, by the notched tergum X apex, and by the smaller number of both hind tibial spines and mesal spines on the apical part of the inferior appendage. Description, male: Body dark greyish brown. Forewing transparent grey with pale spots. Tibial spur formula 2, 2, 2. Hind tibiae each with 15 – 20 small blackish spines (N = 27). Forewing (Fig. 4 A): length 7.0 – 7.6 mm (N = 27), apically rounded; forks 1 and 5 present; crossvein m – cu nearly as long as crossvein r – m, and sharply angled to M. Hindwing (Fig. 4 B): length 5.3 – 5.7 mm (N = 27), apex nearly acute; forks 1, 3, and 5 present; crossvein r – m convex; costa with 16 hamuli. Genitalia: Segment IX annular, laterally setose, narrowest at bases of superior appendages; in lateral view tergite IX wider than sternite IX (Fig. 4 C), central part of tergite IX posterior margin produced posteroventrally (Fig. 4 C, D), smooth, except for a few long setae in some individuals. Superior appendages setose, flattened dorsoventrally, as long as or longer than tergum X, and apices narrowly rounded in dorsal view (Fig. 4 D). Tergum X membraneous; median process entire, wide at basis, tapering to rounded apex, in ventral view with straight dorsal margin (Fig. 4 C), and apex with a wide V-shaped notch in dorsal and ventral views (Fig. 4 D, E); pre-apical lateral processes about as long as median process, produced at apical half, sclerotized, digitate, directed posterad, and slightly curved ventrad (Fig. 4 C, E). Inferior appendages each bipartite, and setose; basal part basally slender in lateral view (Fig. 4 A), wide in ventral view (Fig. 4 E), mesally concave and produced posteromesally into broad, apically rounded lip, well-sclerotized along margin, and bearing small dorsal spines (Fig. 4 C, E); apical part of each inferior appendage digitate, curving mesally; in lateral view, straight, parallel-sided, apex rounded, with 25 – 30 mesal spines (Fig. 4 C); mesal process with nine strong spines (Fig. 4 D). Harpago: small, sclerotized (Fig. 4 C). Phallic apparatus simple (Fig. 4 F, G); phallicata basally tubular, fused with phallobase, and apex truncate; bent about 45 ° ventrally at distal half; apically with pair of sclerotized, broad lateral processes, ventrally directed, tapering distally (Fig. 4 F); phallotremal sclerite U-shaped (Fig. 4 G). Holotype (male): New Caledonia, Province Nord, Mt Panié, stream at camp, 20.58167 ° S, 164.76472 ° E, 1311 m a. s. l., 9 December 2003 – 2 January 2004, Malaise trap, loc # 073 (K. A. Johanson). Paratypes: 12 males, same data as holotype. Fourteen males, ditto, except 20.58139 ° S, 164.76444 ° E, 1310 m a. s. l., loc # 074. Distribution: Province Nord, Mt Panié. Etymology: aurorus, from Latin, aurora, dawn, daybreak; referring to the dark, sclerotized basal posteromesal lip of the inferior appendage, which contrasts with the paler basal part.	en	Malm, Tobias, Johanson, Kjell A. (2008): Revision of the New Caledonian endemic genus Gracilipsodes (Trichoptera: Leptoceridae: Grumichellini). Zoological Journal of the Linnean Society 153 (3): 425-452, DOI: 10.1111/j.1096-3642.2008.00403.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00403.x
C046B71FCB230D3AFF4C5DDFFFB68F90.taxon	diagnosis	Diagnosis: This is the largest species in the genus. It is separated from the other species by the tibial spur formula 1, 1, 1 and, in the genitalia, by the pre-apical lateral processes of tergum X originating at the mid length of tergum X. Description, male: Body and wings medium brown. Tibial spur formula 1, 1, 1. Hind tibiae each with 31 – 35 small black spines (N = 3). Forewing (Fig. 5 A): length 6.5 – 9.7 mm (N = 3); apex rounded; forks 1 and 5 present; crossvein m – cu short and at a right angle to M. Hindwing (Fig. 5 B): length 4.5 – 6.8 mm (N = 3); apex rounded; forks 1, 3, and 5 present; costa with about 39 hamuli; crossvein r – m convex. Genitalia: Segment IX annular, laterally setose, narrowest at bases of superior appendages; tergite IX laterally about as wide as sternite IX (Fig. 5 C); central part of tergite IX posterior margin produced posteroventrad (Fig. 5 C, D). Superior appendages setose, flattened dorsoventrally, about as long as tergum X, apices nearly acute in dorsal view (Fig. 5 D). Tergum X membranous; median process entire, wide basally, tapering to nearly acute apex in lateral view (Fig. 5 C); apex with U-shaped notch in dorsal and ventral views (Fig. 5 D, E); pre-apical lateral processes shorter than median process, produced at mid length, sclerotized, digitate, oriented posterad (Fig. 5 C, E). Inferior appendages each bipartite, setose (Fig. 5 C); basal part wide in ventral view, mesally concave, and produced posteromesally into a broad, distally tapering, apically rounded lip, bearing small dorsal spines (Fig. 5 C, E); apical part of each inferior appendage digitate, curving mesally; in lateral view, parallel-sided, apex rounded, with about 35 mesal spines (Fig. 5 C); mesal process bearing six strong spines (Fig. 5 D). Harpago: sclerotized and slender (Fig. 5 C). Phallic apparatus simple (Fig. 5 F, G); phallicata basally tubular, fused with phallobase, apex truncate; bent about 45 ° ventrad at mid length; apex with pair of broad, sclerotized, lateral processes, being produced ventrally, curved medially, and with apices broadly rounded (Fig. 5 F, G); phallotheca extended; phallotremal sclerite U-shaped, curving anteriorly at apex (Fig. 5 G). Holotype (male): New Caledonia, Province Sud, Mt Dzumac, source stream of Ouinné River, near cross point to mountain track, 22 ° 02.439 ’ S, 166 ° 28.646 ’ E, 805 m a. s. l., 18 November – 4 December 2003, Malaise trap, loc # 029 (Johanson). Paratypes: One male, same data as holotype. One male, New Caledonia, Province Sud, Haute Yaté fauna reserve, 1760 m south of bridge Pont Perignon, 50 m upstream of bridge over stream, 22.14954 ° S, 166.701211 ° E, 180 m a. s. l., 14 December 2003 – 13 January 2004, Malaise trap, loc # 081 (Johanson). Distribution: Province Sud, southern area. Etymology: grandis, from Latin, grand, large; referring to the large size of this species in comparison with the other species within the genus.	en	Malm, Tobias, Johanson, Kjell A. (2008): Revision of the New Caledonian endemic genus Gracilipsodes (Trichoptera: Leptoceridae: Grumichellini). Zoological Journal of the Linnean Society 153 (3): 425-452, DOI: 10.1111/j.1096-3642.2008.00403.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00403.x
C046B71FCB230D3DFCCE5EAAFF058D36.taxon	diagnosis	Diagnosis: Gracilipsodes koghiensis is distinguished from G. lanceolatus sp. nov., G. grandis sp. nov., G. psocopterus, and G. similis by the tibial spur formula of 2, 2, 2, and by the presence of a pair of pre-apical lateral processes of tergum X, originating from the apical half of the segment. It is separated from G. aureus sp. nov. and G. robustus sp. nov. by hind tibia with a higher number of spines, in tergum X, in the genitalia, by the longer lateral processes, and in the forewing, by longer crossvein m – cu oriented at a sharp angle to M. It is separated from G. aurorus sp. nov. by having a paler body colour, by the higher number of hind tibial spines, by the tergum X being apically slit rather than notched, and by the apical part of the inferior appendage having a higher number of mesal spines. It is distinguishable from G. aoupiniensis sp. nov. by having longer superior appendages and, in tergum X, by the deeper apical slit. Description, male: Body brown. Forewing medium brown, with a few pale spots. Tibial spur formula 2, 2, 2. Hind tibiae each with 20 – 25 small black spines (N = 9). Forewing (Fig. 6 A): length 7.0 – 7.7 mm (N = 9), apically rounded; forks 1 and 5 present; crossvein m – cu longer than crossvein r – m, forms sharp angle with M. Hindwing (Fig. 6 B): length 4.8 – 5.5 mm (N = 9), apex narrowly rounded; forks 1, 3, and 5 present; crossvein r – m straight; costa with 23 hamuli. Genitalia: Segment IX annular, laterally setose, narrowest at ventral part; tergite IX broader than sternite IX in lateral view (Fig. 6 C), anterior margins of segment IX weakly produced anteriorly immediately below mid height (Fig. 6 C). Superior appendages setose, flattened dorsoventrally, longer than tergum X, and with apices nearly acute in dorsal view (Fig. 6 D). Tergum X membraneous; median process entire, basally wide, apex rounded, dorsal margin concave in lateral view (Fig. 6 C), apex with deep slit in dorsal and ventral views (Fig. 6 D, E); pre-apical lateral processes as long as or longer than median process, produced at apical half, sclerotized, digitate, directed posterad, and slightly curved ventrally (Fig. 6 C). Inferior appendages each bipartite, setose; basal part broad in ventral view (Fig. 6 E), mesally concave, produced posteromesally into a broad, apically rounded lip, bearing small dorsal spines (Fig. 6 C, E); apical part of each inferior appendage digitate, curving mesally; in lateral view, wide, parallel-sided, apex rounded, and bearing 50 – 60 mesal spines (Fig. 6 C); mesal process with six strong spines (Fig. 6 D). Harpago: weakly sclerotized, slender, apex nearly acute (Fig. 6 C). Phallic apparatus simple (Fig. 6 F, G); phallicata basally tubular, fused with phallobase, apex truncate; strongly bent ventrad at apical half; apical part with pair of sclerotized, slender, lateral processes, being directed ventrally and tapering distally (Fig. 6 F); phallotremal sclerite U-shaped (Fig. 6 G). Holotype (male): New Caledonia, Province Sud, Mt Koghi, 462 m a. s. l., 2 – 5 December 2001, Malaise trap, loc # 7 (Johanson, Pape & Viklund). Paratypes: One male, same data as holotype; 11 males, ditto, except 15 – 16 November 2001, net, loc # 7; one male, ditto, except 6 December 2001, light trap; one male, ditto, except c. 300 m south of Koghi Restaurant, 22.18288 ° S, 166.50490 ° E, 457 m a. s. l., 2 – 16 Novemeber 2003, Malaise trap, loc # 001 (Johanson); two males: ditto, except 16 – 26 November 2003; one male: ditto, except 22.18288 ° S, 166.50393 ° E, 447 m a. s. l., 2 – 16 November 2003, Malaise trap, loc # 002; one male, ditto, except 16 – 26 November 2003; two males, ditto, except 22.18288 ° S, 166.50245 ° E, 427 m a. s. l., 2 – 16 November 2003, Malaise trap, loc # 003; three males, ditto, except 16 – 26 November 2003; four males, ditto, except 22.18288 ° S, 166.50167 ° E, 417 m a. s. l., 16 – 26 November 2003, Malaise trap, loc # 004; one male, ditto, except c. 800 m south of Koghi Restaurant, 22.18311 ° S, 166.50564 ° E, 460 m a. s. l., 10 – 26 November 2003, Malaise trap, loc # 019; one male, ditto, except 22.18365 ° S, 166.50451 ° E, 440 m a. s. l., 11 – 26 November 2003, Malaise trap, loc # 021. Distribution: Province Sud, Mt Koghi. Etymology: The name koghiensis is derived from the type locality, Mt Koghi.	en	Malm, Tobias, Johanson, Kjell A. (2008): Revision of the New Caledonian endemic genus Gracilipsodes (Trichoptera: Leptoceridae: Grumichellini). Zoological Journal of the Linnean Society 153 (3): 425-452, DOI: 10.1111/j.1096-3642.2008.00403.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00403.x
C046B71FCB240D3FFCDB5C13FA6B8FB0.taxon	diagnosis	Diagnosis: This species is separated from the other Gracilipsodes species by having the tibial spur formula 2, 2, 1, and by having only fork 5 present in the hindwings. The genitalia are unique in having a complete tergum X lacking pre-apical lateral processes. Description, male: Body medium brown. Wings greyish brown with blackish hairs. Tibial spur formula 2, 2, 1. Hind tibiae each with 20 – 26 small blackish spines (N = 10). Forewing (Fig. 7 A): length 5.7 – 6.1 mm (N = 10); narrow, apically rounded; forks 1 and 5 present; crossvein m – cu nearly as long as crossvein r – m, forms a wide angle with M 3 + 4; crossvein r – m at sharp angle to M. Hindwing (Fig. 7 B): length 3.9 – 4.2 mm (N = 10); slender, apex nearly acute; fork 5 present; crossvein r – m straight; costa with 22 hamuli. Genitalia: Segment IX annular, laterally setose, widest at mid height; lateral part of tergite IX as wide as sternite IX (Fig. 7 C); posterior margins of sternum IX and tergum IX produced posteriorly (Fig. 7 C, D), lateral margins of sternum IX produced anteriad (Fig. 7 E). Superior appendages setose, flattened dorsoventrally, slightly longer than tergum X (Fig. 7 C), apices rounded in dorsal view (Fig. 7 D). Tergum X entire, membranous, tubular; basally and apically wide, apex rounded, dorsal margin straight in lateral view (Fig. 7 C), narrowing apically, apex with notch in dorsal and ventral views (Fig. 7 D, E); pre-apical lateral processes absent. Inferior appendages each bipartite, setose; basal part slender at base in lateral view (Fig. 7 C), mesally concave, produced posteromesally into slender, tapering, and apically rounded lip, bearing small dorsal spines (Fig. 7 C, E); apical part of each inferior appendage digitate, slender, curving mesally; in lateral view, curving slightly dorsally, widening slowly towards rounded apex, with 25 mesal spines (Fig. 7 C); mesal process with four spines (Fig. 7 D). Harpago reduced. Phallic apparatus simple (Fig. 7 F, G); phallicata basally tubular, fused with phallobase, apex truncate; curving slightly ventrally along its length; apex with pair of pre-apical, sclerotized, ventrally produced broad processes, being directed posterolaterally, and with rounded apices (Fig. 7 F); phallotremal sclerite U-shaped (Fig. 7 G). Holotype (male): New Caledonia, Province Sud, western slope of Mt Ningua, Kwé Néco Stream, at Camp Jacob, 3.7 km west-northwest of the summit of Mt Ningua, on Bouloparis-Thio Road, about 50 m upstream of road, 21 ° 43.613 ′ S, 166 ° 06.567 ′ E, 150 m a. s. l., 29 November – 12 December 2003, Malaise trap, loc # 054 (Johanson). Paratypes: Four males, same data as holotype; one male, ditto, except 12 December 2003 – 5 January 2004; 30 males, New Caledonia, Province Sud, western part of Mt Ningua, Kwé Néco Stream, 3.9 km west of summit of Mt Ningua, on Bouloparis-Thio Road, about 50 m upstream of road, 21 ° 44.359 ′ S, 166 ° 06.009 ′ E, 117 m a. s. l., 20 November 2003 – 12 December 2003, Malaise trap, loc # 035 (Johanson); one male, New Caledonia, Province Sud, western slope of Mt Ningua, Kwé Néco Stream, at Camp Jacob, 3.9 km west of summit of Mt Ningua, on Bouloparis-Thio Road, about 50 m upstream of road, 21 ° 44.083 ′ S, 166 ° 06.298 ′ E, 117 m a. s. l., 29 November 2003 – 12 December 2003, Malaise trap, loc # 053 (Johanson). Distribution: Province Sud. Etymology: lanceolatus, from Latin, lancea, spearlike; named after the slender and apically acute hindwings.	en	Malm, Tobias, Johanson, Kjell A. (2008): Revision of the New Caledonian endemic genus Gracilipsodes (Trichoptera: Leptoceridae: Grumichellini). Zoological Journal of the Linnean Society 153 (3): 425-452, DOI: 10.1111/j.1096-3642.2008.00403.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00403.x
C046B71FCB260D21FF565E89FE7C8C23.taxon	diagnosis	Diagnosis: This species is distinguished from G. lanceolatus sp. nov., G. grandis sp. nov., G. psocopterus, and G. similis by the tibial spur formula 2, 2, 2, and by the presence of a pair of pre-apical lateral processes of tergum X, originating from the apical half of the segment. It is separated from G. aurorus sp. nov., G. koghiensis sp. nov., and G. aoupiniensis sp. nov., in the genitalia, by the shorter lateral processes of tergum X, and in the forewing, by the presence of a shorter crossvein m – cu, oriented at almost a right angle to M. It is distinguishable from G. aureus sp. nov. by being slightly larger, and, in the genitalia, by the shorter superior appendages, and the thicker, slightly clubshaped apical part of the inferior appendage. Description, male: Body medium brown. Forewings brown. Tibial spur formula 2, 2, 2. Hind tibiae each with 18 – 20 small, black spines (N = 3). Forewing (Fig. 8 A): length 7.4 – 7.7 mm (N = 3); apically rounded; forks 1 and 5 present; crossvein m – cu about as long as or slightly shorter than crossvein r – m, oriented at almost a right angle with M. Hindwing (Fig. 8 B): length 5.5 mm (N = 3), apex narrowly rounded; forks 1, 3, and 5 present; crossvein r – m convex; costa with 20 hamuli. Genitalia: Segment IX annular, laterally setose; tergite IX about as broad as sternite IX in lateral view (Fig. 8 C), tergite IX posterior margin central part produced posteroventrad (Fig. 8 C, D). Superior appendages setose, flattened dorsoventrally, about as long as tergum X, apices rounded in dorsal view (Fig. 8 D). Tergum X membraneous; median process entire, basally wide, apex narrowly rounded, dorsal margin slightly concave to straight in lateral view (Fig. 8 C); apex shallowly notched in dorsal and ventral views (Fig. 8 D, E); pre-apical lateral processes shorter than median process, produced at apical half of tergite X, sclerotized, digitate, directed posterolaterally, and slightly curving dorsally (Fig. 8 C). Inferior appendages each bipartite, setose; basal part broad in ventral view (Fig. 8 E), mesally concave, produced posteromesad, apex rounded, and forming a broadly rounded lip, bearing small dorsal spines (Fig. 8 C, E); apical part of each inferior appendage digitate, curving mesally; in lateral view, slightly club-shaped, apex rounded, bearing about 40 mesal spines (Fig. 8 C); mesal process with four strong spines (Fig. 8 D). Harpago: small, slender, weakly sclerotized (Fig. 8 C). Phallic apparatus simple (Fig. 8 F, G); phallicata basally tubular, fused with phallobase, apex truncate; bent about 45 ° ventrad near mid length; apex with pair of sclerotized, lateral, ventrally produced processes, being curved somewhat medially, with rounded apices (Fig. 8 F); phallotremal sclerite U-shaped (Fig. 8 G). Holotype (male): New Caledonia, Province Sud, Mt Koghi, c. 300 m south of Koghi Restaurant, 22.18288 ° S, 166.50073 ° E, 407 m a. s. l., 2 – 16 November 2003, Malaise trap, loc # 005 (Johanson). Paratypes: Two males, same data as holotype, except 22.18288 ° S, 166.50245 ° E, 427 m a. s. l., 2 – 16 November 2003, Malaise trap, loc # 003; one male, ditto, except 22.18288 ° S, 166.50167 ° E, 417 m a. s. l., 16 – 26 November 2003, Malaise trap, loc # 004. Distribution: Province Sud, Mt Koghi. Etymology: From Latin, robustus, solid, robust; referring to the first impression of the holotype genitalia as being robust and stout.	en	Malm, Tobias, Johanson, Kjell A. (2008): Revision of the New Caledonian endemic genus Gracilipsodes (Trichoptera: Leptoceridae: Grumichellini). Zoological Journal of the Linnean Society 153 (3): 425-452, DOI: 10.1111/j.1096-3642.2008.00403.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00403.x
C046B71FCB260D21FF565E89FE7C8C23.taxon	diagnosis	Diagnosis: This species is distinguished from other species in the genus, except from G. similis, in the tibial spur formula 0, 1, 1, and by the darker body and forewing colour. It is distinguishable from G. similis by the longer superior appendages, the deeper split between the tergum X processes, and the shape of the apical part of the inferior appendage. Description, male: Body medium brown to dark greyish brown. Forewing brown to greyish brown. Tibial spur formula 0, 1, 1. Hind tibiae each with 13 – 18 spines (N = 5). Forewing (Fig. 9 A): 5.3 – 6.6 mm (N = 5); apex broadly rounded; forks 1 and 5 present; crossvein m – cu shorter than crossvein r – m, and at a right angle to M. Hindwing (Fig. 9 B): 3.9 – 4.8 mm (N = 5); apex rounded; forks 1 and 5 present; crossvein r – m straight; costa with 24 hamuli. Genitalia: Generally strongly sclerotized and dark. Segment IX annular, laterally setose; in lateral view, sternite IX forming a parallelogram, tergite IX narrower than sternite IX (Fig. 9 C), in ventral view, sternum IX median margin widely and deeply ellipsoid (Fig. 9 E). Superior appendages setose, flattened dorsoventrally, longer than tergum X, apices nearly acute in dorsal view (Fig. 9 D). Tergum X strongly sclerotized, divided into two lateral spines, connected with semipermeable membrane at basal half; in lateral view, basally wide, tapering towards truncate apex (Fig. 9 C); in ventral and dorsal view, apices slightly curving laterad (Fig. 9 D, E); pre-apical lateral processes absent. Inferior appendages each bipartite, setose; basal part broad in lateral and ventral view (Figs 9 C, E), mesally concave, produced posteromesally into a broad, apically rounded lip, bearing small dorsal spines (Fig. 9 C, E); apical part of each inferior appendage digitate, slightly curving mesally; in lateral view, rounded triangular, apex truncate, with 35 – 40 mesoventral spines (Fig. 9 C); mesal process distinct, irregular, with 10 – 15 strong spherical spines (Fig. 9 D). Harpago: small, strongly sclerotized, ovate (Fig. 9 C). Phallic apparatus simple (Fig. 9 F, G); phallicata basally tubular, fused with phallobase, apex narrow; bent strongly ventrally at midlength, distal half nearly straight, pair of sclerotized, lateral processes originate before apex, produced posteroventrad, curved slightly medially, apices narrowly rounded (Fig. 9 F, G); phallotheca extended, phallotremal sclerite U-shaped (Fig. 9 G). Material examined: One male, New Caledonia, Province Sud, Sarraméa, 207 m a. s. l., 18 – 21 November 2001, Malaise trap, loc # 12 (Johanson, Pape & Viklund); one male, New Caledonia, Province Sud, Col d’Amieu, 18 – 21 November 2001, Malaise trap, loc # 13 (Johanson, Pape & Viklund); two males, New Caledonia, Province Sud, Col d’Amieu Fauna Reserve, 319 m a. s. l., 30 November – 5 December 2001, Malaise trap, loc # 23 (Johanson, Pape & Viklund); one male, New Caledonia, Province Sud, Col d’Amieu, Xwé Ko River, at bridge on road to St Forestière, 21 ° 35.612 ′ S, 165 ° 48.241 ′ E, 368 m a. s. l., 8 January 2004, light trap, loc # 114 (Johanson); six males, New Caledonia, Province Sud, western part of Mt Ningua, Kwé Néco Stream, 3.9 km west of summit of Mt Ningua, on Bouloparis-Thio Road, about 50 m upstream of road, 21 ° 44.359 ′ S, 166 ° 06.009 ′ E, 117 m a. s. l., 20 November 2003 – 12 December 2003, Malaise trap, loc # 035 (Johanson); one male, New Caledonia, Province Nord, northern slope of Mt Rembai, Xwé Méréxwana Stream, 21 ° 33.804 ′ S, 165 ° 50.528 ′ E, 309 m a. s. l., 20 November – 12 December 2003, Malaise trap, loc # 037 (Johanson). Distribution: Central area of Grande Terre.	en	Malm, Tobias, Johanson, Kjell A. (2008): Revision of the New Caledonian endemic genus Gracilipsodes (Trichoptera: Leptoceridae: Grumichellini). Zoological Journal of the Linnean Society 153 (3): 425-452, DOI: 10.1111/j.1096-3642.2008.00403.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00403.x
C046B71FCB380D28FCA55D18F9FF8F32.taxon	diagnosis	Diagnosis: The species is distinguished from all other species in the genus, except G. psocopterus, by having the tibial spur formula 0, 1, 1, and by a darker body and forewing colour. It differs from G. psocopterus by the shorter superior appendages, by the shallower split between the tergum X processes, and by the shape of the apical part of the inferior appendage. Description, male: Body medium brown to dark greyish brown. Forewing brown to greyish brown. Tibial spur formula 0, 1, 1. Hind tibiae each with 12 – 13 small spines (N = 2). Forewing (Fig. 10 A): length 5.5 – 6.1 mm (N = 3); apex broadly rounded; forks 1 and 5 present; crossvein m – cu about as long as or slightly shorter than crossvein r – m, oriented at a sharp angle to M. Hindwing (Fig. 10 B): length 4.1 – 4.4 mm (N = 2); apex rounded; forks 1 and 5 present; crossvein r – m straight; costa with 25 – 30 hamuli. Genitalia: Generally strongly sclerotized, dark. Segment IX annular, laterally setose; in lateral view, sternite IX nearly rectangular, broad, tergite IX half as wide as sternite IX (Fig. 10 C); tergite IX posterior margin weakly produced posteroventrad in dorsal view (Fig. 10 D), in ventral view, sternum IX median margin widely and shallowly ellipsoid (Fig. 10 E). Superior appendages setose, flattened dorsoventrally, shorter than tergum X, apices nearly acute in dorsal view (Fig. 10 D). Tergum X strongly sclerotized, divided into two lateral spines connected with semipermeable membrane to distal two-thirds; in lateral view, basal part wide, tapering to rounded apex, with small ventral process basally, (Fig. 10 C); in dorsal and ventral views, apices curving laterally (Fig. 10 D, E); pre-apical lateral processes absent. Inferior appendages each bipartite, setose; basal part broad in ventral and lateral views (Fig. 10 C, E), mesally concave, produced posteromesally into a broad, apically rounded lip, bearing small dorsal spines (Fig. 10 C, E); apical part of each inferior appendage digitate, almost straight in dorsal view (Fig. 10 D); in lateral view, widening slightly at mid length, apex slender, with 40 – 50 mesoventral spines (Fig. 10 C); mesal process distinct, with 10 – 12 strong, rounded mesal spines (Fig. 10 D). Harpago: small, strongly sclerotized, ovate (Fig. 10 C). Phallic apparatus simple (Fig. 10 F, G); phallicata basally tubular, fused with phallobase, apex narrow; sharply bent ventrally at mid length, apex with a pair of sclerotized, lateral processes, being produced posteroventrad, curved somewhat medially, with apex narrowly rounded (Fig. 10 F, G); phallotremal sclerite U-shaped (Fig. 10 G). Material examined: One male, New Caledonia, Province Nord, Amoa River, 23 m a. s. l., 25 – 26 November 2001, light trap, loc # 20 (Johanson, Pape & Viklund); 13 males, New Caledonia, Province Nord, 50 m upstream of bridge on Hienghène-Tnèdo road, 3.9 km south of summit of Mt Tnèda, 2.4 km east of Tnèdo, 20 ° 43.099 ′ S, 164 ° 50.108 ′ E, 27 m a. s. l., 7 December 2003 – 1 January 2004, loc # 069 (Johanson). Distribution: Eastern parts of Province Nord. PHYLOGENETIC RESULTS In all analyses, the individual Gracilipsodes species formed monophyletic units with jackknife support of 84 – 100 % (except for G. aureus in the EF- 1 a analysis, which collapses basally of G. robustus), but the relationship among the various species differed between the genes analyzed. The gene producing the overall lowest branch support was EF- 1 a. The implied alignment of the mitochondrial ribosomal 16 S gene fragments yielded 380 homologous characters, of which 87 were parsimony informative for all 25 specimens. P distances are shown in Table 3, indicating the high similarity between G. similis and G. psocopterus, as well as for G. koghiensis and G. sp. nov. 1. The subsequent parsimony analysis of this gene alone resulted in one shortest tree (L = 252) (Fig. 11 A). We recovered Triplexa basally in the Gracilipsodes species clade consisting of (G. lanceolatus, (G. grandis, (G. psocopterus, G. similis ))). This relationship has, however, low branch support (jackknife = 67 %). This clade again forms the sister clade to the second major clade containing the three species pairs ((( G. robustus, G. aureus), (( G. aurorus, G. aoupiniensis), (G. sp. nov. 1, G. koghiensis ))). The mitochondrial COI data set consisted of 658 characters, of which 212 were parsimony informative for all 25 specimens. According to the P distances in Table 4, the highest similarity is found between G. psocopterus and G. similis. The parsimony analysis of this data set yielded two shortest trees (L = 640) (Fig. 11 B), placing Triplexa as sister to all Gracilipsodes. This position was poorly supported (jackknife = 60 %). The basal divergence within Gracilipsodes is poorly resolved, but holds two strongly supported clades (G. aureus, G. robustus) and (G. psocopterus, G. similis), both with jackknife support of 99 %. The data set of the mitochondrial COII gene fragment was one taxon shorter than for the other genes, resulting from the absence of a sequence from A. botosaneanui, and included 512 characters, of which 176 were parsimony informative. According to the P distances in Table 5, the highest similarity is found between G. similis and G. psocopterus, as well as for G. aureus and G. robustus. The parsimony analysis of this character set resulted in three shortest trees (L = 494) (Fig. 11 C), with conflicts between them assigned basally within Gracilipsodes, resulting in poor resolution at this level. Here again we found Triplexa inside Gracilipsodes as sister to [G. lanceolatus, (G. aureus, G. robustus)], but this configuration had a low jackknife support (below 50 %). As in the results of the COI dataset analysis, the results here strongly support the two clades (G. aureus, G. robustus) and (G. psocopterus, G. similis) as being monophyletic. The nuclear EF- 1 a data set excluded one specimen because of an absent sequence (the AQ 3 G. aoupiniensis), and consisted of 1099 characters, of which 194 were parsimony informative for all 24 individuals. According to Table 6, the P distances show very high sequence similarity within the species pairs G. Species set in bold are sp. nov. ‘ n. a. ’ indicates only one specimen present within the species, thus no distance span. Species set in bold are sp. nov. ‘ n. a. ’ indicates only one specimen present within the species, thus no distance span. Species set in bold are sp. nov. ‘ n. a. ’ indicates missing sequence or only one specimen present within the species, thus no distance span. Species set in bold are sp. nov. ‘ n. a. ’ indicate only one specimen present within the species, thus no distance span. aureus + G. robustus and G. psocopterus + G. similis. The parsimony analysis yielded one shortest tree (L = 440) (Fig. 11 D), where Triplexa was found to form the sister group to Gracilipsodes. The monophyly of Gracilipsodes received a high jackknife value (95 %). The individuals of G. lanceolatus were grouped together and formed the sister group to all the remaining Gracilipsodes species. The remaining species formed two major clades, one with low jackknife support (67 %) – ((( G. aureus, G. robustus), (G. aoupiniensis, (G. koghiensis, (G. aurorus, G. sp. nov. 1 )))) – and one with strong jackknife support (99 %) – (G. grandis, (G. psocopterus, G. similis). Here, all species are moderately to strongly supported, except G. aureus, which collapses basally of G. robustus. The combined data set yielded a matrix consisting of 25 individuals and 2649 characters (of which 670 were parsimony informative). The parsimony analysis resulted in a single shortest tree (L = 1839) (Fig. 12), where Triplexa formed the sister group to Gracilipsodes (as in the results of the EF- 1 a and COI analyses). The monophyly of Gracilipsodes received a jackknife support of 79 %, and the genus was divided into two major clades: (G. lanceolatus, (G. grandis, (G. psocopterus, G. similis ))) and (( G. aureus, G. robustus), (( G. koghiensis, G. sp. n. 1), (G. aoupiniensis, G. aurorus ))). Most of the nodes received strong support, except 61 % for (( G. koghiensis, G. sp. nov. 1), (G. aoupiniensis, G. aurorus )), 71 % for (G. aoupiniensis, G. aurorus), and 74 % for (G. lanceolatus, (G. grandis, (G. psocopterus, G. similis). The data-set analysis in MrModeltest (Nylander, 2004) for the separate genes gave no single model for all gene partitions, but the two genes COI and COII shared the ‘ GTR + invgamma’ model, EF- 1 a was proposed to be analyzed with ‘ GTR + gamma’, and 16 S was proposed to be analyzed with ‘ HKY + invgamma’. These models were assigned to their respective dataset partition, and were then run in MrBayes in a combined analysis. This analysis resulted in a tree topologically identical to the most parsimonious tree generated from the combined analysis (Fig. 12). All nodes received high posterior probability values (± 0.99), except the G. koghiensis + G. sp. nov. 1 clade and the G. lanceolatus – G. similis clade, with posterior probabilities of 0.91 and 0.96, respectively.	en	Malm, Tobias, Johanson, Kjell A. (2008): Revision of the New Caledonian endemic genus Gracilipsodes (Trichoptera: Leptoceridae: Grumichellini). Zoological Journal of the Linnean Society 153 (3): 425-452, DOI: 10.1111/j.1096-3642.2008.00403.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00403.x
