taxonID	type	description	language	source
C0438794D847FFA7FF017590FBF2FB05.taxon	description	Figs. 2 – 3	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D847FFA7FF017590FBF2FB05.taxon	diagnosis	Diagnosis. Head with asetose vertexal foveae small or lacking; lacking frontal fovea; antennal clubs 1 – 4 segment- ed; antennal tubercles moderately developed to lacking; postantennal notches small or lacking; postantennal cavities present from vaguely visible to as long as antennal tubercles; ventrolateral margin beneath eyes often rounded; median gular ridge narrow and low, barely visible in lateral view. Pronotum with lateral antebasal foveae punctiform or lacking; lacking median antebasal fovea. Elytra lacking basal foveae and discal stria. Abdominal tergite 1 lacking mediobasal foveae and discal carinae; ventrite 2 lacking mediobasal foveae.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D847FFA7FF017590FBF2FB05.taxon	description	Redescription [based on New Zealand members of Eupines]. Body length 1.23 – 2.10 mm, generally short and compact in dorsal view; body color reddish brown to dark brown and rarely black; head and pronotum smooth, elytra and abdomen micropunctate or smooth; lacking setae or with sparsely scattered short decumbent setae on pronotum, elytra and abdomen, average length longer than length of eyes. Head with asetose vertexal foveae (Fig. 2 A) small or lacking, often punctiform; lacking frontal fovea; frontal rostrum often moderately developed; antennal tubercles moderately developed to lacking; postantennal notches distinctively margined to lacking; postantennal cavity (Figs. 2 A – B) extend from dorsal side of head to lateral side of antennal insertions, size vary from vaguely visible to as long as antennal tubercles; eyes often well developed and prominent; antennae (Figs. 2 F – G) 9 - segmented, 10 - segmented or 11 - segmented in males and 11 - segmented in females, antennal clubs 1 – 4 segmented, with scape and pedicel elongate; ocular-mandibular carinae (Fig. 2 B) present; ventrolateral margin rounded beneath eyes or with short basal carinae; median gular ridge (Fig. 2 C) narrow and low, barely visible in lateral view, curved posteriorly to asetose gular foveae (Fig. 2 C); mouth (Figs. 2 D – E) with labial palp truncate at apex, with pair of long apical setae, mentum widest at middle, lacinia well developed, galea fused with lacinia, maxillary palpi with segment 2 slender and clavate, segment 3 trapezoidal, segment 4 large and fusiform, sensory area with stout apical setae, mandible with pair of subapical setae, labrum trapezoidal. Pronotum about as long as wide; lacking median antebasal fovea; lateral antebasal foveae punctiform or lacking; lacking antebasal sulcus and median longitudinal sulcus; setose lateral procoxal foveae (Fig. 2 J) small and often widely separated. Elytra (Fig. 3 N) slightly wider than long, longer than abdomen in dorsal view, with posterior margin often rounded; each elytron lacking basal foveae and discal carinae; sutural stria distinct to lacking. Mesoventrite (Fig. 2 K) with asetose prepectal foveae vaguely visible; setose median mesoventral foveae bifurcate at apex, often forming triangular cavity; pair of setose lateral mesoventral foveae large and nearly meet internally; setose lateral mesocoxal foveae often small. Metaventrite with setose lateral metaventral foveae small. Abdomen (Figs. 3 A, 3 G) convex, with tergite 1 about 1.5 × to 2 × longer than tergite 2; tergite 1 (Fig. 3 B) with basolateral foveae vaguely visible; lacking mediobasal fovea, discal carinae and mediobasal impression; tergites 2 – 4 (Figs. 3 C – E) lacking basolateral foveae; paratergites visible on tergites 1 – 3 and carinate on tergite 4. Abdominal ventrite 2 with basolateral foveae (Fig. 3 I) vaguely visible, lacking mediobasal fovea, ventrites 3 – 5 (Figs. 3 J – L) lacking basolateral foveae. Legs with protarsomeres 2 and 3 subequal in length, mesotarsomere 2 slightly longer than mesotarsomere 3, metatarsomere 2 about as long as metatarsomere 3. Males with antennae, protrochanters, protibiae, mesotibiae, ventrite 2 or ventrite 6 modified. Aedeagus with well-developed and often symmetrical parameres; median lobe often with two sclerites; basal bulb relatively small; lacking dorsal diaphragm. Relationships: Within the New Zealand fauna, members of Eupines are most similar to those of Simkinion Park and Pearce based on combination of the following characers: pronotum lacking a median antebasal fovea, tergite 1 lacking the mediobasal impression and ventrites 4 – 5 lacking the basolateral foveae. Members of Eupines can be readily separated from those of Simkinion by the absence of basal elytral foveae and the low median gular ridge, which is prominent in the latters. Members of Eupines are also similar to that of Eupinolus Oke in lacking the basal elytral foveae and the short abdomen. They may be separated by the presence of a small median antebasal fovea and the punctate pronotal base in members of Eupinolus, which are lacking in members of Eupines. In Australia, Chandler (2001) stated that Eupines is closely related to Storeyella Chandler and Wataranka Chandler but can be easily differentiated from both by the smooth basal pronotal area and the lack of a median antebasal pronotal fovea.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D847FFA7FF017590FBF2FB05.taxon	discussion	Comments. The lack of the basal elytral foveae, the median antebasal pronotal fovea and the basolateral foveae on tergites and ventrites, suggests that Eupines may be more derived than all other goniacerite genera in New Zealand. The subgenus Byraxis is characterized by males possessing 10 antennomeres, and males of the nominate Eupines possess 11 antennomeres: females of both subgenera have 11 - segmented antennae. Nomura & Leschen (2006), in their faunistic review of New Zeland Pselaphinae, listed nine species under the nominate subgenus in New Zealand. In the following treatment of this subgenus by Shen & Leschen (2019), three species were found based solely on female types, and one was transferred to the subgenus Byraxis. During work of the nominate subgenus in this paper, five species are revealed not members of Eupines, and one new species is described. Up to date, the subgenus Eupines comprises only one species in New Zealand, with all remaining species placed into the diverse subgenus Byraxis. In Australia it is the reverse, with the subgenus Eupines being more diverse, possibly due to incorrect subgeneric placements or insufficient study (Chandler 2001). Three species are excluded from this revision (E. (E.) grata (Sharp), E. clemens Broun and E. (E.) sternalis (Broun )). These are being described in a separate manuscript as a new genus due to the presence of the shallow antebasal pronotal sulcus, short discal elytral striae, and different genitalic characters.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D848FFA4FF01732AFB9FF8E0.taxon	description	Fig. 4; map Fig. 49	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D848FFA4FF01732AFB9FF8E0.taxon	materials_examined	Type locality: [MC] Broken River. Type depository: BMNH LT ♂, 1 PLT.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D848FFA4FF01732AFB9FF8E0.taxon	diagnosis	Diagnosis: Head with vertexal foveae punctiform; antennae 10 - segmented with 2 - segmented club; A 4, A 6, A 7, A 8 and A 10 unmodified, A 4 slightly asymmetrical, A 5 large and asymmetrical, wider than A 4 at middle and longer than A 3 + A 4, outer margin flat, A 9 large, about 4 × longer than A 8, A 10 conical, about 1.6 × longer than A 9. Abdominal ventrite 2 with pair of small discal protuberances, inserted with strongly curved apical setae; ventrite 6 with pair of median struts, about one-fourth of ventral length. Protrochanters with barely visible ventral spine, shorter than one-tenth of protrochanteral width; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D848FFA4FF01732AFB9FF8E0.taxon	description	Redescription: Body length 1.51 – 1.63 mm; body colour reddish brown to dark brown with lighter coloured elytra and legs. Head about as long as wide; frons narrowed at end, slightly shorter than half width of head at eye level; eyes well developed, with about 34 facets; vertexal foveae punctiform; lacking antennal tubercles and postantennal notches; postantennal cavities small, slightly shorter than half length of antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented club; A 4, A 6, A 7, A 8 and A 10 unmodified, A 4 slightly asymmetrical, A 5 large and asymmetrical, wider than A 4 at middle and longer than A 3 + A 4, outer margin flat, A 9 large, about 4 × longer than A 8, A 10 conical, about 1.6 × longer than A 9. Pronotum wider than long, slightly wider than head at eye level; Elytra slightly wider than long, widest near middle; humeral calli slightly prominent; sutural stria distinctive at base and vaguely visible near apex. Abdomen with tergite 1 about 1.5 × longer than tergite 2; ventrite 2 with pair of small discal protuberances, inserted with strongly curved apical setae; ventrite 6 with pair of median struts, about one-fourth of ventral length. Legs with protrochanters modified with barely visible ventral spine, shorter than one-tenth of protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.28 mm long; with elongate and thin parameres, curved downwards and twisted at apex, with two long subapical setae; median lobe with two long sclerites, slightly wider than parameres in dorsal view, left one curved inward subapically, right one bent abruptly.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D848FFA4FF01732AFB9FF8E0.taxon	discussion	Comments: Eupines (B.) anisarthra is similar to E. (B.) minuta by having A 5 and A 9 similarly modified and can be separated by the unmodified mesotibiae (a small apical spine is present in E. (B.) minuta; Fig. 31 E).	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D848FFA4FF01732AFB9FF8E0.taxon	distribution	Distribution: South Island. MC, CO. Additional material examined: 3 ♂♂: CO: 2, Carrick Range, Watts Rock, 1380 m, 31 / Oct / 1979, J. C. Watt, moss, 79 / 133; 1, JNPC, Old Man Range, Headwaters, Obelisk Ck., 1380 m, 13 / Jan / 07. Associated females (2 ♀♀): 2, CO: Carrick Range, Watts Rock, 1380 m, 31 / Oct / 1979, J. C. Watt, moss, 79 / 133.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D849FFA2FF017134FD9CFF49.taxon	description	Fig. 5; map Fig. 49	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D849FFA2FF017134FD9CFF49.taxon	materials_examined	Type locality: [MC] Mount Hutt. Type depository: BMNH LT ♂, 1 PLT.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D849FFA2FF017134FD9CFF49.taxon	diagnosis	Diagnosis: Head lacking vertexal foveae; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 6, A 7, A 8 and A 10 unmodified, A 5 asymmetrical and triangular, excavated at base, A 9 asymmetrical, about 3 × longer than A 8, inner apical margin expanded, A 10 conical, slightly impressed at base, about 1.8 × longer than A 9. Abdominal ventrite 2 with pair of small discal protuberances, inserted with strongly curved small apical setae; ventrite 6 unknown. Protrochanters with minute ventral spine, about one-ninth of protrochanteral width; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D849FFA2FF017134FD9CFF49.taxon	description	Redescription: Body length 1.43 mm; body colour reddish brown with lighter coloured tibiae and tarsi. Head wider than long; frons slightly narrowed at end, about half width of head at eye level; eyes slightly prominent in dorsal view, with about 22 facets; lacking vertexal foveae; antennal tubercles barely visible; lacking postantennal notches; postantennal cavities large, slightly longer than antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 6, A 7, A 8 and A 10 unmodified, A 5 asymmetrical and triangular, excavated at base, A 9 asymmetrical, about 3 × longer than A 8, inner apical margin expanded, A 10 conical and slightly impressed at base, about 1.8 × longer than A 9. Pronotum about as long as wide, wider than head at eye level; lacking lateral antebasal foveae. Elytra slightly wider than long, widest at apical two-thirds; humeral calli barely visible; sutural stria distinctively extending from base to apex. Abdomen with tergite 1 about 2.3 × longer than tergite 2; ventrite 2 with pair of small discal protuberances, inserted with strongly curved small apical setae; ventrite 6 unknown. Legs with protrochanters modified with minute ventral spine, about one-ninth of protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus unknown.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D849FFA2FF017134FD9CFF49.taxon	discussion	Comments: Only two type specimens are known. We did not attempt to dissect the syntypes since males of this species have distinctive antennal modifications (Fig. 5 B) that separate the species from congeners.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D849FFA2FF017134FD9CFF49.taxon	distribution	Distribution: South Island. MC.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D84FFFA1FF01779DFCB2FA24.taxon	description	Fig. 6; map Fig. 49	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D84FFFA1FF01779DFCB2FA24.taxon	diagnosis	Diagnosis: Head with vertexal foveae punctiform; antennae 10 - segmented with 3 - segmented antennal club; A 4, A 5, A 6, A 7, A 8 and A 10 unmodified, A 6 prolonged, about 1.5 × longer than A 5, A 8 slightly asymmetrical, A 9 largest and asymmetrical, strongly impressed at outer side, with laminate upper margin, A 10 conical and slightly shorter than A 9. Abdominal ventrite 2 with pair of barely visible discal protuberances, inserted with slightly curved small apical setae; ventrite 6 with pair of barely visible median struts. Protrochanters with long and acute ventral spine, about four-fifths of protrochanteral width; protibiae with medium-sized apical protuberance; mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D84FFFA1FF01779DFCB2FA24.taxon	description	Description: Body length 1.37 – 1.45 mm; body colour reddish brown with lighter coloured elytra and legs. Head wider than long; frons not narrowed at end, about three-fourths width of head at eye level; eyes well developed and prominent, with about 33 facets; vertexal foveae punctiform; antennal tubercles weak; lacking postantennal notches; postantennal cavities large, about as long as antennal tubercles in dorsal view; antennae 10 - segmented with 3 - segmented antennal club; A 4, A 5, A 6, A 7, A 8 and A 10 unmodified, A 6 prolonged, about 1.5 × longer than A 5, A 8 slightly asymmetrical, A 9 largest and asymmetrical, strongly impressed at outer side, with laminate upper margin, A 10 conical and slightly shorter than A 9. Pronotum slightly longer than wide, narrower than head at eye level; lacking lateral antebasal foveae. Elytra as long as wide, widest at apical two-thirds; humeral calli weakly developed, slightly prominent in dorsal view; sutural stria shallowly extending from base to apex. Abdomen with tergite 1 about 1.5 × longer than tergite 2; ventrite 2 with pair of barely visible discal protuberances, inserted with slightly curved small apical setae; ventrite 6 with pair of barely visible median struts. Legs with protrochanters modified with long and acute ventral spine, about four-fifths of protrochanteral width; protibiae with medium-sized apical protuberance; mesotibiae unmodified. Aedeagus about 0.23 mm long, with long parameres curved towards each other at apex, slightly sinuate, widest at about apical three-fourths, with three pairs of long subapical setae and two pairs of long apical setae; median lobe with two short and straight sclerites, curved downwards at apex, widest near base, about two-fifths length of parameres.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D84FFFA1FF01779DFCB2FA24.taxon	discussion	Comments: This species is similar to E. (B.) setifera based on the similar A 9 and apical triangular spine on the protibiae, but can be separated by possessing an enlarged A 6, much larger than A 7, while A 6 is obviously smaller than A 7 in E. (B.) setifera (Fig. 45 B).	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D84FFFA1FF01779DFCB2FA24.taxon	distribution	Distribution: North Island. AK, WO, TO.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D84FFFA1FF01779DFCB2FA24.taxon	etymology	Etymology: This specific epithet is a Latin adjective meaning ‘ short’, referring to the short sclerites of the aedeagus.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D84FFFA1FF01779DFCB2FA24.taxon	materials_examined	Type material (NZAC): Holotype. ♂, specimen entire, point-mounted: “ NEW ZEALAND, AK, Noises Is., Otata I., 7 Dec – 10 Dec 1979 // J. C. Watt & C. F. Butcher, Malaise ”. Paratypes. 18 ex.: WO: 1, Onewhero, 13 / Jan / 1985, R. A. Galbreath and S. Grant // Malaise Trap, North in Beilschmiedia tarairi forest; TO: 2, P 544 & P 547, 27 / Nov / 1951, moss, Huka Falls, Taupo, N. I.; 12, P 532 – P 543, 28 / Nov / 1951, moss, Huka Falls, Taupo, N. I.; 3, P 528 – P 530, 27 / Nov / 1951, moss, Huka Falls, Taupo, N. I.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D84CFFA1FF017277FCF4F9DF.taxon	description	Fig. 7; map Fig. 49	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D84CFFA1FF017277FCF4F9DF.taxon	diagnosis	Diagnosis: Head with vertexal foveae punctiform; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 transverse, cupulate, A 10 largest, irregularly cup-shaped, broadly excavated at ventral side. Abdominal ventrite 2 and ventrite 6 unknown. Protrochanters with long and acute ventral spine, slightly shorter than protrochanteral width; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D84CFFA1FF017277FCF4F9DF.taxon	description	Description: Body length 1.33 mm; body colour reddish brown with lighter coloured tarsi. Head slightly wider than long; frons not narrowed at end, about three-fourths width of head at eye level; eyes well developed, slightly prominent in dorsal view; vertexal foveae punctiform; antennal tubercles weakly developed; postantennal notches shallow and broad; postantennal cavities small, about half length of antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 transverse, cupulate, A 10 largest, irregularly cup-shaped, broadly excavated at ventral side. Pronotum as long as wide, slightly narrower than head at eye level; lacking lateral antebasal foveae. Elytra as long as wide, widest near middle; humeral calli barely visible in dorsal view; sutural stria shallowly extending from base to apex. Abdomen with tergite 1 about 1.6 × longer than tergite 2; ventrite 2 and ventrite 6 unknown. Legs with protrochanters modified with long and acute ventral spine, slightly shorter than protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus unknown.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D84CFFA1FF017277FCF4F9DF.taxon	discussion	Comments: Only the holotype is known. We did not attempt to dissect the holotype since the male has an irregularly cup-shaped A 10 (Figs. 7 B – C) that differentiates this species from other members of Eupines.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D84CFFA1FF017277FCF4F9DF.taxon	distribution	Distribution: North Island. NN.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D84CFFA1FF017277FCF4F9DF.taxon	etymology	Etymology: This specific epithet is a Latin noun meaning ‘ glove’, referring to the irregularly cup-shaped antennomere 10.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D84CFFA1FF017277FCF4F9DF.taxon	materials_examined	Type material (NZAC): Holotype. ♂, specimen entire, glued on card, one proleg mounted on acetate card: “ 19. xii. 1951, moss, nr. L. Omapere, N. I. P 772 [handwritten] ”.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D84DFFBFFF01733BFF4FF8E3.taxon	description	Fig. 8; map Fig. 49	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D84DFFBFFF01733BFF4FF8E3.taxon	diagnosis	Diagnosis: Head with vertexal foveae punctiform; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 greatly elongated, about five-sixths length of A 10, lateral side with two deep striae extending from base to near middle, apical margin slightly wavy, A 10 largest and generally conical, lateral side with three narrow carinae splitting one large basal impression and two small anterolateral impressions. Abdominal ventrite 2 with pair of barely visible discal protuberances, inserted with slightly curved large apical setae; ventrite 6 with pair of long and acute median struts, curved towards each other at apex, about half of ventral length. Protrochanters with long and acute ventral spine, about twice of protrochanteral width; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D84DFFBFFF01733BFF4FF8E3.taxon	description	Description: Body length 1.88 – 1.99 mm; body colour reddish brown to dark brown with lighter coloured elytra and legs. Head slightly wider than long; frons slightly narrowed at end, about half width of head at eye level; eyes well developed, conspicuously prominent in dorsal view, with about 33 facets; vertexal foveae punctiform; antennal tubercles weakly developed; postantennal notches shallow; postantennal cavities long and narrow, about two-thirds length of antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 greatly elongated and about five-sixths length of A 10, lateral side with two deep striae extending from base to near middle, apical margin slightly wavy, A 10 largest and generally conical, lateral side with three narrow carinae splitting one large basal impression and two small anterolateral impressions. Pronotum about as long as wide, slightly wider than head at eye level; with punctiform lateral antebasal foveae. Elytra slightly longer than wide, widest near apical two-thirds; humeral distinctive in dorsal view; sutural stria distinctively extending from base to apex. Abdomen with tergite 1 about twice length of tergite 2; ventrite 2 with pair of barely visible discal protuberances, inserted with slightly curved large apical setae; ventrite 6 with pair of long and acute median struts, curved towards each other at apex, about half of ventral length. Legs with protrochanters modified with long and acute ventral spine, about twice of protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.38 mm long, with long and straight parameres, widest near middle, with pair of long apical setae; median lobe with single large and horn-shaped sclerite, average width narrower than parameres.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D84DFFBFFF01733BFF4FF8E3.taxon	discussion	Comments: This species has the largest ratio of ventral spine / protrochanteral widths (Fig. 8 E) among all known Eupines in New Zealand. Distribution: North Island. BP, WN. Etymology: This specific epithet refers to the carina on the last antennomere of this species. Type material: Holotype. ♂, JNPC, specimen entire, card-mounted: “ NEW ZEALAND, WN, Waikawa Beach, 18 Mar 1990 // J. T. Nunn collection ”. Paratypes. 1 ex.: BP: 1, Waimana Valley, 20 / Feb / 1927 // A. E Brookes Collection.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D852FFBDFF017133FEE4FC69.taxon	description	Fig. 9; map Fig. 49	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D852FFBDFF017133FEE4FC69.taxon	diagnosis	Diagnosis: Head with vertexal foveae punctiform; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 longest, slightly longer than A 10, strongly impressed from middle to apex at ventral side, inserted with long spine within impression, A 10 bean-shaped, slightly impressed near base at ventral side. Abdominal ventrite 2 with single fused small discal protuberance, inserted with strongly curved minute apical setae; ventrite 6 with pair of long and acute median struts, about one-third of ventral length. Protrochanters with small ventral spine, about one-third of protrochanteral width; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D852FFBDFF017133FEE4FC69.taxon	description	Description: Body length 1.71 mm; body colour reddish brown with lighter coloured tarsi. Head wider than long; frons not narrowed at end, about three-fourths width of head at eye level; eyes moderately developed, slightly prominent in dorsal view, with about 27 facets; vertexal foveae punctiform; antennal tubercles weakly developed; lacking postantennal notches; postantennal cavities large, about as long as antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 longest, slightly longer than A 10, strongly impressed from middle to apex at ventral side, inserted with long spine within impression, A 10 bean-shaped, slightly impressed near base at ventral side. Pronotum as long as wide, about as wide as head at eye level; lacking lateral antebasal foveae. Elytra slightly longer than wide, widest at middle; humeral calli moderately developed, distinctively prominent in dorsal view; sutural stria barely visible. Abdomen with tergite 1 about 1.3 × longer than tergite 2; ventrite 2 with single fused small discal protuberance, inserted with strongly curved minute apical setae; ventrite 6 with pair of long and acute median struts, about one-third of ventral length. Legs with protrochanters modified with small ventral spine, about one-third of protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.35 mm long, with long and slender parameres, curved towards each other and bent downwards in lateral view, with three pairs of short subapical setae and two pairs of short apical setae; median lobe large, with two large median sclerites twisted together near apex, about as wide as parameres, left side of median lobe with three small sclerites in dorsal view, left one curved outwards, right two short and straight.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D852FFBDFF017133FEE4FC69.taxon	discussion	Comments: This species is characterized by having ventrite 2 modified with a single fused protuberance (Fig. 9 F), which can separate it from all congeners.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D852FFBDFF017133FEE4FC69.taxon	distribution	Distribution: North Island. ND.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D852FFBDFF017133FEE4FC69.taxon	etymology	Etymology: This specific epithet is a Latin adjective meaning ‘ grow together’, referring to the fused protuberance on ventrite 2 of this species.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D852FFBDFF017133FEE4FC69.taxon	materials_examined	Type material (NZAC): Holotype. ♂, specimen entire, glued on card, aedeagus + proleg + antenna on acetate card: “ NEW ZEALAND, ND, Puketi Forest Waipapa R. Tk., E. Branch, 30 Jan 1995 // J. Klimaszewski Sifted litter ”. Paratypes. 4 ex.: ND: 2, FMNH, Waipoua State Forest, Wairau Summit, 400 m, 27 / xi – 6 / xii / 1984, hdwd. - podocarp forest, A. Newton / M. Thayer 687 // berl., leaf & log litter forest floor; 2, FMNH, Waipoua SF., Yakas Tree Tr. 350 m, 11 – 14 / iv / 1980, broadlf. - podocarp A. Newton, M. Thayer, 710 // berl, litter at bases of Metrosideros robusta trunks.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D850FFBCFF0174BDFB51FF49.taxon	description	Fig. 10; map Fig. 49	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D850FFBCFF0174BDFB51FF49.taxon	diagnosis	Diagnosis: Head with vertexal foveae punctiform; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 8 disc-shaped, A 9 largest, slightly longer than A 10, with subapical impression at ventral side and bearing a conspicuous spine at middle, A 10 conical and slightly impressed at ventral side. Abdominal ventrite 2 with pair of barely visible discal protuberances, inserted with strongly curved small apical setae; ventrite 6 with pair of long and acute median struts, curved towards each other, about two-thirds of ventral length. Protrochanters with short and acute ventral spine, slightly shorter than one-third of protrochanteral width; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D850FFBCFF0174BDFB51FF49.taxon	description	Description: Body length 1.50 – 1.60 mm; body colour reddish brown with lighter coloured tibiae and tarsi. Head wider than long; frons not narrowed at end, about three-fourths width of head at eye level; eyes well developed, conspicuously prominent in dorsal view, with about 35 facets; vertexal foveae punctiform; antennal tubercles moderately developed; postantennal notches broad and shallow; postantennal cavities small, about half length of antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 8 disc-shaped, A 9 largest, slightly longer than A 10, with subapical impression at ventral side and bearing a conspicuous spine at middle, A 10 conical and slightly impressed at ventral side. Pronotum as long as wide, about as wide as head at eye level; lacking lateral antebasal foveae. Elytra about as long as wide, widest at middle; humeral calli and sutural stria barely visible. Abdomen with tergite 1 about twice length of tergite 2; ventrite 2 with pair of barely visible discal protuberances, inserted with strongly curved small apical setae; ventrite 6 with pair of long and acute median struts, curved towards each other, about two-thirds of ventral length. Legs with protrochanters modified with short and acute ventral spine, slightly shorter than one-third of protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.32 mm long, with slender parameres, curved towards each other, bent downwards in lateral view, with three pairs of long subapical setae and pair of short apical setae; median lobe large, with four entwined sclerites, left one straight and short, right one long and wide, slightly wider than parameres, middle two longer than outer two in dorsal view, strongly entwined together and slightly narrower than parameres.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D850FFBCFF0174BDFB51FF49.taxon	discussion	Comments: This species is most similar to E. (B.) coalita by having a similarly modified A 9 but can be separated by having A 5 transverse (Fig. 10 B) (it is longer than wide in E. (B.) coalita).	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D850FFBCFF0174BDFB51FF49.taxon	distribution	Distribution: North Island. ND, TO, BP.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D850FFBCFF0174BDFB51FF49.taxon	etymology	Etymology: This specific epithet means ‘ entwined’ in Latin, referring to the strongly entwined sclerites in aedeagus of this species.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D850FFBCFF0174BDFB51FF49.taxon	materials_examined	Type material (NZAC): Holotype. ♂, specimen entire, point-mounted: “ NEW ZEALAND, ND, Poor Knights Is., Aorangi, ridge to Oneho Hill, 17 Nov 1981 // J. C. Watt Sifted litter 81 / 141 ”. Paratypes. 10 ex.: ND: 1, Poor Knights Is., Aorangi, 11 / Nov / 1981, J. C. Watt // Beaten from Flowering Myoporum laetum; 2, Poor Knights Is., Tawhiti Rahi, 01 / Dec / 1980 G. Kuschel // Sifted litter and rotten wood on plateau, 80 / 130; 2, Poor Knights Is., Tawhiti Rahi, 02 – 10 / Dec / 1980, M. F. Tocker // Pan traps at edge of stagnant stream; 1, Poor Knights Is., Tawhiti Rahi, 04 – 10 / Dec / 1980, R. H. Kleinpaste // Malaise trap in bush; BP: 1, Kaimai Ra., 427 m, Leyland Tramline, 22 / Sep / 1981, B. M. May // litter 81 / 92; 1, Orete Forest, Te Puia Hut Bush, 15 / Sep / 1992, G. Hall & // R. C. Henderson, sifted litter 92 / 50; TO: 1, Ohakune Jubilee Park, Bush Walk, 05 / Nov / 2004, A. C. Eyles // litter & moss, bark of live tree & fallen logs; 1, Waipunga River nr. Fall, 14 / Mar / 1980, C. F. Butcher // litter and mosses 80 / 32.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D851FFBAFF0173FCFB48FF16.taxon	description	Fig. 11; map Fig. 49	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D851FFBAFF0173FCFB48FF16.taxon	materials_examined	Type locality: [AK] Hunua Range, Maketu. Type depository: BMNH LT ♂, 3 PLT; NZAC 14 PLT.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D851FFBAFF0173FCFB48FF16.taxon	diagnosis	Diagnosis: Head with vertexal foveae punctiform; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 boot-shaped with curved basal spine, A 10 bean-shaped, truncate at base in dorsal view, with bi-excavation in ventral view. Abdominal ventrite 2 with pair of large discal protuberances, inserted with strongly curved apical setae; ventrite 6 with pair of median struts, slightly shorter than one-fourth of ventral length. Protrochanters with long ventral spine, about four-sevenths of protrochanteral width; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D851FFBAFF0173FCFB48FF16.taxon	description	Redescription: Body length 1.45 – 1.56 mm; body colour reddish brown to dark brown with lighter coloured elytra and legs. Head slightly wider than long; frons slightly narrowed at end, about two-thirds width of head at eye level; eyes prominent, with about 38 facets; vertexal foveae punctiform; antennal tubercles weak; postantennal notches weak; postantennal cavities well developed, about two-thirds length of antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 boot-shaped with curved basal spine, A 10 bean-shaped, truncate at base in dorsal view, with bi-excavation in ventral view. Pronotum as long as wide, about as wide as head at eye level; lacking lateral antebasal foveae. Elytra as long as wide, widest at apical two-thirds; humeral calli vaguely visible; sutural stria distinctively extending from base to apex. Abdomen with tergite 1 about twice length of tergite 2; ventrite 2 with pair of large discal protuberances, inserted with strongly curved apical setae; ventrite 6 with pair of median struts, slightly shorter than one-fourth of ventral length. Legs with protrochanters modified with long ventral spine, about four-sevenths of protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.32 mm long, with parameres elongate and thick, curved downwards; median lobe with two long sclerites, narrower than parameres in dorsal view, both bent inwards subapically in dorsal view, with left one longer than right one.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D851FFBAFF0173FCFB48FF16.taxon	discussion	Comments: This species is most similar to E. (B.) monstrosa by having similar modifications of A 9 and A 10 and can be separated by A 10 lacking a small internal hook at the apex (Fig. 11 B).	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D851FFBAFF0173FCFB48FF16.taxon	distribution	Distribution: North Island. ND, AK, WO, CL, TO, GB, HB. Additional material examined: 23 ♂♂, ND: 3, 20 / vi / 51, moss, Ruakaka, N. I., N. Z.; AK: 1, Bethells, Matuku Reserve, 23 / Jun / 1987, R. C. Craw; CL: 1, AMNZ, Orewa Gorge, Karangahaka, 11 / 10 / 42, C. E. Clark Collection; WO: 3, JNPC, Maungatautari, Hamilton, 6 / Sep / 04, forest floor litter, collected by Sue Nunn; GB: 4, Motu river, April / 1928, A. E. Brookes Collection; To: 1, Waipunga River, nr. Falls, 14 / March / 1980, C. F. Butcher; HB: 2, Waipatiki Res., 23 / Dec / 1983, J. C. Watt. Sifted, wood mould, 83 / 144; 3, White Pine Bush, 18 / Nov / 1984, J. G. Charles. Unknown locality: 3, 2464, T. Broun Collection, A. E. Brookes Collection; 2, 2464, T. Broun. Associated females (6 ♀♀): ND: 1, 20 / vi / 51, moss, Ruakaka, N. I. N. Z.; WO: 3, JNPC, Maungatautari, Hamilton, 6 / Sep / 04, forest floor litter, collected by Sue Nunn; GB: 2, Motu river, April / 1828, A. E. Brookes Collection.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D857FFBAFF017764FB49F9D3.taxon	description	Fig. 12; map Fig. 50	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D857FFBAFF017764FB49F9D3.taxon	materials_examined	Type locality: [CL] Tairua. Type depository: BMNH HT ♂.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D857FFBAFF017764FB49F9D3.taxon	diagnosis	Diagnosis: Head with vertexal foveae asetose and small; antennae 10 - segmented with 4 - segmented antennal club; A 4, A 5, A 6, A 7 and A 10 unmodified, A 7 slightly asymmetrical, A 8 elongate, about 3 × longer than A 7, truncate at base, A 9 longest, about twice as long as A 8, concave at outer margin, concavity surrounded by strongly extended laminate margin, A 10 conical and truncate at base. Abdominal ventrite 2 with pair of small discal protuberances, inserted with slightly curved apical setae; ventrite 6 with pair of short median struts, about one-fifth of ventral length. Protrochanters with small ventral spine, about one-third of protrochanteral width; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D857FFBAFF017764FB49F9D3.taxon	description	Redescription: Body length 1.40 – 1.52 mm; body colour reddish brown to dark brown with lighter coloured legs. Head slightly wider than long; frons slightly narrowed at end, about half width of head at eye level; eyes well developed, with about 33 facets; vertexal foveae asetose and small; antennal tubercles moderately developed; lacking postantennal notches; postantennal cavities large in dorsal view, about as long as antennal tubercles in dorsal view; antennae 10 - segmented with 4 - segmented antennal club; A 4, A 5, A 6, A 7 and A 10 unmodified, A 7 slightly asymmetrical, A 8 elongate, about 3 × longer than A 7, truncate at base, A 9 longest, about twice as long as A 8, concave at outer margin, concavity surrounded by strongly extended laminate margin, A 10 conical and truncate at base. Pronotum as long as wide, about as wide as head at eye level; lacking lateral antebasal foveae. Elytra as long as wide, widest at apical two-thirds; humeral calli and sutural stria vaguely visible. Abdomen with tergite 1 about 1.5 × longer than tergite 2; ventrite 2 with pair of small discal protuberances, inserted with slightly curved apical setae; ventrite 6 with pair of short median struts, about one-fifth of ventral length. Legs with protrochanters modified with small ventral spine, about one-third of protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.23 mm long, with parameres elongate and sinuate, with three pairs of short apical setae, two pairs of long subapical setae and one pair of long median setae; median lobe with two long sclerites, about as wide as parameres in dorsal view, both curved outwards at apex, left one more sinuate than right one.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D857FFBAFF017764FB49F9D3.taxon	discussion	Comments: This is the only species known with A 8 conspicuously modified (Figs. 12 B – D).	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D857FFBAFF017764FB49F9D3.taxon	distribution	Distribution: North Island. ND, AK, CL. Additional material examined: 120 ♂♂, ND: 27, Mokohinau Is., Trig I., 27 / Feb / 1978, G. Kuschel, sifted litter, 78 / 107; 12, Mokohinau Is., Fanal I., 01 / Mar / 1978, G. Kuschel, sifted litter and ground plants, 78 / 115; 1, Mokohinau Is., Burgess I., 06 / Jan / 1984, P. D McDonald, litter, 84 / 5; 2, Hen & Chickens Is., Lady Alice I., 01 / Jan / 1982, R. Hay, litter, 82 / 4; 12, Poor Knights Is., Aorangi Crater Bay and ridge to Oneho Hill, 17 / Nov / 1981, J. C. Watt, sifted litter 81 / 141; AK: 54, Noises Is., Scott I., 25 / Feb / 1979, J. M. Cleary, M. F. Tocker, litter 79 / 31; CL: 2, Mecury Is., Korapuki I., 16 / Dec / 1987, G. Hall, sifted litter, 89 / 23; 10, AMNZ, Great Barrier Is., Little Windy Hill, 220 m, 11 / xii / 2001 – 18 / i / 2002, P. Sutton, Forest edge, Malaise trap. Associated females (201 ♀♀): ND: 73, Mokohinau Is., Trig I., 27 / Feb / 1978, G. Kuschel, sifted litter, 78 / 107; 16, Mokohinau Is., Burgess I., 06 / Jan / 1984, P. D McDonald, litter, 84 / 5; 26, Mokohinau Is., Fanal I., 01 / Mar / 1978, G. Kuschel, sifted litter and ground plants, 78 / 115; 11, Hen & Chickens Is., Lady Alice I., 01 / Jan / 1982, R. Hay litter 82 / 4; 7, Poor Knights Is. Aorangi Crater Bay and ridge to Oneho Hill, 17 / Nov / 1981, J. C. Watt, sifted litter, 81 / 141; AK: 71, Noises Is., Scott I., 25 / Feb / 1979, J. M. Cleary, M. F. Tocker, litter 79 / 31; CL: 7, Mecury Is., Korapuki I., 16 / Dec / 1987, G. Hall, sifted litter, 89 / 23.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D854FFB8FF0173A2FBB3F9D3.taxon	description	Fig. 13; map Fig. 50	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D854FFB8FF0173A2FBB3F9D3.taxon	materials_examined	Type locality: [AK] Paparoa, near Howick. Type depository: BMNH LT ♂, 1 PLT.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D854FFB8FF0173A2FBB3F9D3.taxon	diagnosis	Diagnosis: Head with vertexal foveae asetose and small; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 largest, basally concave at inner side in dorsal view, with spine in small lateral apical excavation, A 10 conical in dorsal view and flattened in lateral view, with long lateral subapical spine. Abdominal ventrite 2 with pair of small and nude discal protuberances; ventrite 6 with pair of long median struts, slightly longer than half of ventral length. Protrochanters with long ventral spine, about as long as protrochanteral width; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D854FFB8FF0173A2FBB3F9D3.taxon	description	Redescription: Body length 1.32 – 1.36 mm; body colour dark brown with lighter coloured elytra and legs. Head wider than long; frons slightly narrowed at end, about three-fifths width of head at eye level; eyes well de- veloped and prominent, with about 32 facets; vertexal foveae asetose and small; antennal tubercles barely visible; postantennal notches punctiform; postantennal cavities medium-sized in dorsal view, slightly longer than half length of antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 largest, basally concave at inner side in dorsal view, with spine in small lateral apical excavation, A 10 conical in dorsal view and flattened in lateral view, with long lateral subapical spine. Pronotum as long as wide, slightly narrower than head at eye level; lacking lateral antebasal foveae. Elytra as long as wide, widest at apical two-thirds; humeral calli moderately developed; sutural stria shallowly extending from base to apex. Abdomen with tergite 1 about 1.5 × longer than tergite 2; ventrite 2 with pair of small and nude discal protuberances; ventrite 6 with pair of long median struts, slightly longer than half of ventral length. Legs with protrochanters modified with long ventral spine, about as long as protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.30 mm long, with slender and slightly asymmetric parameres curved inwards, with pair of short apical setae; median lobe with two long sclerites, wide near base and gradually convergent towards apex, both curved inwards in dorsal view, left one longer and more curved than right one.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D854FFB8FF0173A2FBB3F9D3.taxon	discussion	Comments: This species is similar to E. (B.) petila by its similarly modified A 10. However, the strongly asymmetrical aedeagus and long median struts of ventrite 6 of E. (B.) decens (Figs. 13 G – J) can immediately separate it from E. (B.) petila.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D854FFB8FF0173A2FBB3F9D3.taxon	distribution	Distribution: North Island. AK, CL. Additional material examined: 4 ♂♂, AK: 1, Ex. Broun Collection, Hunua Ra., Maketu, Auckland, Byraxis decens Broun, T. Broun Collection, A. E. Brookes Collection; 1, Coll. A. E. Brookes, Hunua Ra., Drury, 1 / 1 / 1916, 1870, Byraxis decens Broun., Var. A. E. Brookes Collection; CL: 2, Ex. Broun Collection, Tairua, E. coast, North Is., 1870, Byraxis decens Broun, T. Broun Collection, A. E. Brookes Collection.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D85BFFB6FF017691FE44FC67.taxon	description	Fig. 14	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D85BFFB6FF017691FE44FC67.taxon	materials_examined	Type locality: New Zealand. Type depository: BMNH HT ♀.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D85BFFB6FF017691FE44FC67.taxon	discussion	Comments: Sharp (1874) described E. deformis from a single female with strangely bent protibiae and mesotibiae that, at first, he regarded were deformed. He later concluded that the tibiae were natural, and it was a valid species resembling E. dispar.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D85BFFB5FF0174B7FB78F8F6.taxon	description	Fig. 15; map Fig. 50	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D85BFFB5FF0174B7FB78F8F6.taxon	materials_examined	Type locality: [AK] Auckland. Type depository: BMNH LT ♂, 3 PLT.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D85BFFB5FF0174B7FB78F8F6.taxon	diagnosis	Diagnosis: Head with vertexal foveae punctiform; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 narrowed at base, truncate at apex, with outside surface strongly polished and slightly concave, A 10 bean-shaped, strongly concave at lateral side. Abdominal ventrite 2 with pair of slightly prominent discal protuberances, inserted with slightly curved apical setae; ventrite 6 with pair of long median struts, about two-thirds of ventral length, slightly curved inwards at apex. Protrochanters with long ventral spine, about as long as protrochanteral width; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D85BFFB5FF0174B7FB78F8F6.taxon	description	Redescription: Body length 2.00 – 2.14 mm; body colour reddish brown to dark brown with lighter coloured elytra and legs. Head about as long as wide; frons slightly narrowed at end, about three-fifths width of head at eye level; eyes well developed, with about 38 facets; vertexal foveae punctiform; antennal tubercles weak; postantennal notches punctiform; postantennal cavities long and narrow, slightly shorter than antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 narrowed at base, truncate at apex, with outside surface strongly polished and slightly concave, A 10 bean-shaped, strongly concave at lateral side. Pronotum longer than wide, slightly narrower than head at eye level; lateral antebasal foveae punctiform. Elytra about as long as wide, widest at apical three-fourths; humeral calli barely visible; sutural stria distinctively extending from base to apex. Abdomen with tergite 1 about twice length of tergite 2; ventrite 2 with pair of slightly prominent discal protuberances, inserted with slightly curved apical setae; ventrite 6 with pair of long median struts, about two-thirds of ventral length, slightly curved inwards at apex. Legs with protrochanters modified with long ventral spine, about as long as protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.36 mm long, with slender parameres, with four pairs of apical setae; median lobe with single stout and long sclerite, slightly bent and widest near middle in dorsal view.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D85BFFB5FF0174B7FB78F8F6.taxon	discussion	Comments: The lectotype and two paralectotypes of E. (B.) munroi were compared with the lectotype and one paralectotype of E. (B.) dispar, and were found to represent a single species by having identical antennal modifica- tions; the former name is here treated as the junior synonym. This species is distinguished from other Eupines by its modified A 10 and the median lobe containing a single sclerite (Figs. 15 B – J).	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D85BFFB5FF0174B7FB78F8F6.taxon	distribution	Distribution: North Island. AK, WO, TK. South Island. NN, BR, WD. Additional material examined: 28 ♂♂, AK: 1, AMNZ, Manukau Murphys Bush, litter under rimu, 29 / v / 2005, S. E. Thorpe; WO: 1, JNPC, Tirohanga Tk., Mt. Pirongia, 20 / Nov / 2005, J. T. Nunn collection; TK: 1, AMNZ, Mt. Taranaki, 520 m, N Egmont Rd., 20 – 23 / iii / 2005, J. W. Early, R. F. Gilbert; NN: 1, NW Nelson Mangarakau Sc. Res., 21 / Dec / 1992; 13, Fenian Track, carpark, Oparara, 03 / Jan / 2011, ground litter in flood zone in nikau grove; BR: 3, Greymouth, Grandjeans Tk., 10 / Nov / 2005, R. Leschen & S. Nomura, rotten logs, 42 ° 28 ' S 171 ° 12 ' E; WD: 2, Doughboy Creek, 6 KM SW Mahitahi, 05 / Feb / 1984, J. C. Watt, sifted wood mould, 84 / 21; 5, Haast River Walk, 27 / III / 2016, forest floor litter, J. Nunn; 1, NMPC, Three Mile Lagoon, 8 / xii / 2012, 3.5 km SW of Okarito, 43 ° 14.5 ' S 170 ° 7.5 ' E, M. Fikáček, J. Hájek & R. Leschen lgt. Associated females (14 ♀♀): WO: 1, Mt. Karioi near base, 11 / Oct / 1981, C. F. Butcher litter, 81 / 96; 1, JNPC, Tirohanga Tck., Mt. Pirongia, 20 / Nov / 2005, J. T. Nunn collection; TK: 1, JNPC, Potaema Walk, Mt. Egmont, 09 / 12 / 95, J. T. Nunn collection; NN: 5, Fenian Track, carpark, Oparara, 03 / Jan / 2011, ground litter flood zone in nikau grove; WD: 2, Doughboy Creek, 6 KM SW Mahitahi, 05 / Feb / 1984, J. C. Watt, sifted wood mould, 84 / 21; 4, Haast River Walk, 27 / III / 2016, forest floor litter, J. Nunn.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D858FFB3FF017146FAAFFF81.taxon	description	Fig. 16; map Fig. 50	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D858FFB3FF017146FAAFFF81.taxon	diagnosis	Diagnosis: Head lacking vertexal foveae; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 6, A 7, A 8 and A 10 unmodified, A 5 conspicuously enlarged, more than twice length of A 4, A 9 asymmetrical and largest, about 1.5 × longer than A 10, base strongly narrowed, widest at middle, A 10 conical. Abdominal ventrite 2 with pair of large discal protuberances, inserted with strongly curved small apical setae; ventrite 6 with pair of slender median struts, about three-fifths of ventral length. Protrochanters with long and acute ventral spine, about two-thirds of protrochanteral width; protibiae with large apical protuberance; mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D858FFB3FF017146FAAFFF81.taxon	description	Description: Body length 1.45 – 1.55 mm; body colour dark brown to black with lighter coloured tarsi. Head wider than long; frons slightly narrowed at end, about half width of head at eye level; eyes moderately developed, with about 24 facets; lacking vertexal foveae; antennal tubercles barely visible; lacking postantennal notches; postantennal cavities large, about 1.3 × length of antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 6, A 7, A 8 and A 10 unmodified, A 5 conspicuously enlarged, more than twice length of A 4, A 9 asymmetrical and largest, about 1.5 × longer than A 10, base strongly narrowed, widest at middle, A 10 conical. Pronotum slightly wider than long, wider than head at eye level; lacking lateral antebasal foveae. Elytra slightly longer than wide, widest at apical three-fifths; humeral calli barely visible; sutural stria shallowly extending from base to apex. Abdomen with tergite 1 about as long as tergite 2; ventrite 2 with pair of large discal protuberances, inserted with strongly curved small apical setae; ventrite 6 with pair of slender median struts, about threefifths of ventral length. Legs with protrochanters modified with long and acute ventral spine, about two-thirds of protrochanteral width; protibiae with large apical protuberance; mesotibiae unmodified. Aedeagus about 0.33 mm long, with long and large parameres, widest near apex, bent downwards in lateral view, with pair of subapical setae; median lobe with two small sclerites curved towards each other at apex, much smaller than parameres.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D858FFB3FF017146FAAFFF81.taxon	discussion	Comments: This species is similar to E. (B.) impar by having A 9 enlarged but can be separated by its modified protibiae (Fig. 16 E) (it is unmodified in E. (B.) impar).	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D858FFB3FF017146FAAFFF81.taxon	distribution	Distribution: South Island. OL, FD, SL.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D858FFB3FF017146FAAFFF81.taxon	etymology	Etymology: This species is named after the collector of the holotype, John Dugdale.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D858FFB3FF017146FAAFFF81.taxon	materials_examined	Type material (NZAC): Holotype. ♂, specimen entire, point-mounted: “ NEW ZEALAND, OL, Headlong Pk., 1524 m, South Basin, 20 Feb 1980, J. S. Dugdale // Litter 80 / 26 ”. Paratypes. 11 ex.: OL: 5, same data as holotype; FD: 1, Middle Basin, Tutoko Bench, Darran Mts. 13 / Jan / 1977 // J. S. Dugdale; 1, Middle Basin, Tutoko Bench, Darran Mts., 1524 m // 14 / Jan / 1977, J. S. Dugdale; 1, Secretary I., Mt. Grono, 29 / May / 1982, T. Haslam // Mat plants 82 / 67; SL: 3, 1 km S. Longwood Trig, 700 m, Longwood Ra., 01 / Feb / 1976 // L. L. Deitz Mats, 76 / 12.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D85EFFB2FF0177D5FE18FA4C.taxon	description	Fig. 17; map Fig. 50	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D85EFFB2FF0177D5FE18FA4C.taxon	materials_examined	Type locality: [AK] Near Howick. Type depository: BMNH LT ♂.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D85EFFB2FF0177D5FE18FA4C.taxon	diagnosis	Diagnosis: Head with vertexal foveae punctiform; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 largest, narrowed at base, widened gradually towards apex, with long and acute spine at lateral side, A 10 long and conical, slightly shorter than A 9. Abdominal ventrite 2 with pair of slightly prominent discal protuberance, inserted with strongly curved small apical setae; ventrite 6 with pair of mediumsized acute median struts, slightly shorter than one-third of ventral length. Protrochanters with long and acute ventral spine, slightly longer than protrochanteral width; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D85EFFB2FF0177D5FE18FA4C.taxon	description	Redescription: Body length 1.52 – 1.54 mm; body colour reddish brown with lighter coloured tarsi. Head slightly wider than long; frons narrowed at end, about one-third width of head at eye level; eyes moderately developed, with about 22 facets; vertexal foveae punctiform; lacking antennal tubercles; postantennal notches barely visible; postantennal cavities large, slightly shorter than antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 largest, narrowed at base, widened gradually towards apex, with long and acute spine at lateral side, A 10 long and conical, slightly shorter than A 9. Pronotum as long as wide, about as long as head at eye level; lacking lateral antebasal foveae. Elytra about as long as wide, widest at apical two-thirds; humeral calli barely visible; sutural stria weakly extending from base to apical half. Abdomen with tergite 1 about 1.5 × longer than tergite 2; ventrite 2 with pair of slightly prominent discal protuberance, inserted with strongly curved small apical setae; ventrite 6 with pair of medium-sized acute median struts, slightly shorter than one-third of ventral length. Legs with protrochanters modified with long and acute ventral spine, slightly longer than protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.30 mm long, with slender parameres, widest at middle, strongly curved inwards, with four pairs of long apical setae and pair of short subapical setae; median lobe small and fusiform, with two long sclerites curved towards each other at apex in dorsal view, widest at basal one-third, right one slightly longer than left one.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D85EFFB2FF0177D5FE18FA4C.taxon	discussion	Comments: This species is recognized by its A 9 modified with long and acute spine at the lateral side (Figs. 17 B – D).	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D85EFFB2FF0177D5FE18FA4C.taxon	distribution	Distribution: North Island. AK, CL. Additional material examined: 3 ♂♂, AK: 1, Ahuroa, 240 m, nr. Woodcocks, 5 / Jan / 1983, J. C. Watt, leaves and moss, 83 / 5; CL: 1, AMNZ, Great Barrier Island, Rasalie Bay, Benthorn Farm, 150 m, 2 / xi. – 11 / xii / 2001, P. Sutton, Boggy forest edge, Malaise trap; 1, AMNZ, Great Barrier Island, Little Windy Hill, 220 m, 26 / viii – 28 / ix / 2002, P. Sutton, forest edge, Malaise trap; GB: 1, LSAM, Urewera N. P., Ngomoko Tr., pitfall, fish & banana, 22 / Mar / 2000 C. Carlton, A. Weir, # 076.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D85FFFB1FF01729EFABEF8AB.taxon	description	Fig. 18; map Fig. 50	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D85FFFB1FF01729EFABEF8AB.taxon	diagnosis	Diagnosis: Head with vertexal foveae punctiform, surrounded by shallow impressions; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 enlarged, ventral side truncate, A 10 largest and conical, about twice length of A 9, with deep basal excavation at ventral side. Abdominal ventrite 2 with pair of small and flat discal protuberances, inserted with strongly curved long apical setae; ventrite 6 with pair of long median struts, curved towards each other, slightly longer than half of ventral length. Protrochanters with long and acute ventral spine, about as long as protrochanteral width; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D85FFFB1FF01729EFABEF8AB.taxon	description	Description: Body length 1.95 – 2.07 mm; body colour reddish brown to dark brown with lighter coloured elytra and legs. Head wider than long; frons not narrowed at end, about three-fourths width of head at eye level; eyes well developed, conspicuously prominent in dorsal view, with about 36 facets; vertexal foveae punctiform, surrounded by shallow impressions; antennal tubercles moderately developed; postantennal notches distinctive and broad; postantennal cavities large, about as long as antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 enlarged, ventral side truncate, A 10 largest and conical, about twice length of A 9, with deep basal excavation at ventral side. Pronotum about as long as wide, slightly narrower than head at eye level; lacking lateral antebasal foveae. Elytra as long as wide, widest near apical two-thirds; humeral calli weak, slightly prominent in dorsal view; sutural stria shallowly extending from base to apex. Abdomen with tergite 1 about twice length of tergite 2; ventrite 2 with pair of small and flat discal protuberances, inserted with strongly curved long apical setae; ventrite 6 with pair of long median struts, curved towards each other, slightly longer than half of ventral length. Legs with protrochanters modified with long and acute ventral spine, about as long as protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.34 mm long, with parameres elongate, widest near middle, slightly curved outwards at apex, bent downwards in lateral view, with four pairs of short subapical setae and pair of short apical setae; median lobe with single small and stout sclerite, narrower than parameres.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D85FFFB1FF01729EFABEF8AB.taxon	discussion	Comments: This species is similar to E. (B.) dispar by sharing a large body size (average length around 2.00 mm), but can be easily recognized by its conical A 10, while it is bean-shaped in E. (B.) dispar (Figs. 15 B – D).	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D85FFFB1FF01729EFABEF8AB.taxon	distribution	Distribution: North Island. TO, BP, TK, HB.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D85FFFB1FF01729EFABEF8AB.taxon	etymology	Etymology: This specific epithet is a Latin adjective meaning ‘ large’, referring to the large body size of this species.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D85FFFB1FF01729EFABEF8AB.taxon	materials_examined	Type material (NZAC): Holotype. ♂, specimen entire, point-mounted: “ NEW ZEALAND, BP, Upper Waipi- ata Karakatuwhero V. 3 Feb 1993, J. S. Dugdale // Litter 93 / 15 ”. Paratypes. 8 ex.: TO: 1, Ohakune R. C. Craw // litter, 85 / 67; 1, Kaimanawa SF., 26 / Mar / 1978, S. B. Peck, litter; 2, Opepe 27 / Mar / 1978, S. B. Peck Litter; BP: 1, Orete Forest, Te Puia Hut, 25 – 29 / Jan / 1993, R. C. Henderson // Pan trap; 1, Coll. A. E. Brookes, 26 / Dec / 1943 // leaf mould // Kaimai Range, Matamata, Waikato // A. E Brookes Collection; TK: 1, Mt. Egmont 3500 ’, 17 / Apr / 1965, litter, N. A. Walker; HB: 1, Waipukurau Lindsay Reserve, 28 / Dec / 1981, J. C. Watt // sifted litter 81 / 218.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D85CFF8FFF0170FBFB1AFB05.taxon	description	Fig. 19; map Fig. 51	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D85CFF8FFF0170FBFB1AFB05.taxon	materials_examined	Type locality: [AK] Woodhill, near the Kaipara Railway. Type depository: NZAC HT ♂.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D85CFF8FFF0170FBFB1AFB05.taxon	diagnosis	Diagnosis: Head with vertexal foveae punctiform; antennae 10 - segmented with 4 - segmented antennal club; A 4, A 5, A 6, A 7, A 8 and A 10 unmodified, A 6 and A 8 slightly asymmetrical, A 9 largest, with a deep groove at one side, forming laterally twisted lamina at apex, A 10 conical and much shorter than A 9. Abdominal ventrite 2 with pair of small discal protuberances, inserted with strongly curved small apical setae; ventrite 6 with pair of short and acute median struts, about one-tenth of ventral length. Protrochanters with long and acute ventral spine, about same to protrochanteral width; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D85CFF8FFF0170FBFB1AFB05.taxon	description	Redescription: Body length 1.33 – 1.44 mm; body colour reddish brown to dark brown with lighter coloured tibiae and tarsi. Head wider than long; frons not narrowed at end, about three-fourths width of head at eye level; eyes well developed, with about 31 facets; vertexal foveae punctiform; antennal tubercles moderately developed; postantennal notches broadly extended and meet at middle of vertex; postantennal cavities medium-sized and about two-thirds length of antennal tubercles in dorsal view; antennae 10 - segmented with 4 - segmented antennal club; A 4, A 5, A 6, A 7, A 8 and A 10 unmodified, A 6 and A 8 slightly asymmetrical, A 9 largest, with a deep groove at one side, forming laterally twisted lamina at apex, A 10 conical and much shorter than A 9. Pronotum as long as wide, slightly narrower than head at eye level; lacking lateral antebasal foveae. Elytra longer than wide, widest at about apical three-fourths; lacking humeral calli; sutural stria barely visible. Abdomen with tergite 1 about 1.2 × longer than tergite 2; ventrite 2 with pair of small discal protuberances, inserted with strongly curved small apical setae; ventrite 6 with pair of short and acute median struts, about one-tenth of ventral length. Legs with protrochanters modified with long and acute ventral spine, about same to protrochanteral width; protibiae and mesotibiae unmodi- fied. Aedeagus about 0.28 mm long, with parameres long and clavate at apex, curved inwards; median lobe with four short and stout sclerites originating from middle in dorsal view, two outside ones curved outwards, two middle ones strongly curved inwards.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D85CFF8FFF0170FBFB1AFB05.taxon	discussion	Comments: This species is widely distributed in the North Island and is also reported in Marlborough Sounds. It can be distinguished from congeners by its A 8 conspicuously transerverse and enlarged (Fig. 19 C).	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D85CFF8FFF0170FBFB1AFB05.taxon	distribution	Distribution: North Island. ND, AK, WO, CL, BP, GB, TK, HB, WN, WA. South Island. SD. Additional material examined: 90 ♂♂, ND: 1, E. Pandora Rd., Te Paki Coastal Park, 07 / Feb / 1975, A. K. Walker; 2, Mokohinau Is., Fanal I., 01 / Mar / 1978, G. Kuschel, sifted litter; 6, Poor Knights Is., Tawhiti Rahi, 10 / Sep / 1980, J. C. Watt litter; AK: 6, Lynfield, 27 / Jan / 1977, G. Kuschel, litter; 1, Noises Is., Motuhoropapa I. South Trig, 15 / Jan / 1978, L. L. Deitz, litter near pit trap; 1, Bethells Matuku Res., 26 / Jul – 20 / Aug / 1983, G. Hall and P. A. Maddison, pit traps; 3, Coll. A. E. Brookes, Sep / 1914, Titirangi Waitakere Rg., Auckland, 1476; 10, Clevedon R. A. Cumber Coll., 09 / Jul / 1958; 4, AMNZ, Lynfield, 27 / Jan / 1977, G. Kuschel; CL: 1, Little Barrier I., Te Maraeroa South side, 07 / Apr / 1984, C. T. Duval, sifted litter; 2, AMNZ, Cuvier I., West Ridge Track, 160 m, 12 / xi / 1999, J. W. Early, S. E. Thorpe; WO: 1, Waitomo, 24 / May / 1983, G. Kuschel, decayed wood and litter; 1, Mt. Pirongia SF., Rangitukia stream, 13 / Jun / 2001, R. Leschen, leaf litter, 37 ° 58 ' S 175 ° 07 ' E; BP: 2, Lottin Pt Rd., Waenga Bush, 27 / Jan / 1993, R. C. Henderson, litter; 1, Hicks Bay, 14 / Mar – 29 / Apr / 1993, J. S. Dugdale, litter; 9, Te Koau Twin Puriri, 31 / Jan / 1993, J. S. Dugdale litter; 1, Waiotuma Stm., 65 m, 28 / Jan / 1998, J. S. Dugdale, litter; 2, LUNZ, Te Koau, Twin Puiris, 15 / iii / 1993, R. M. Gilbert, J. W. M. Marris; GB: 1, East Cape, Rangiata, 17 / Mar – 30 / Apr, G. Hall, pit traps; 2, Whakaroa Scenic Reserve, 22 / Nov / 1993 – 11 / Jan / 1994, G. Hall, pit traps; 2, Kakanui, 350 m, 01 / Feb / 1993, J. S. Dugdale, litter; 2, Taikawakawa, 21 / Sep / 1992, G. Hall & R. C. Henderson, sifted litter, 92 / 58; 1, LUNZ, Kakanui, 260 m 16 / iii / 1993, J. W. M. Marris; TK: 1, White Cliffs Walkway, 5 km S. Tongaporutu, 07 / Dec / 1982, J. S. Dugdale, litter 82 / 137; HB: 13, Motuokura (Bare I.), 07 / Feb – 20 / Mar / 1995, G. Hall, pit traps; RI: 4, Coll. J. Ford., 28 / 9 / 1917, Taihape main Trunk, N. Island, A. E. Brookes Collection; WA: 1, W Shore Res., L Wairarapa, 09 / Oct / 1993; WN: 6, JNPC, Nikau Reserve, Paraparaumu, 23 / Jul / 95, J. T. Nunn collection; SD: 1, LUNZ, Titi Island, 17 / xi / 1994, J. W. M. Marris, litter ex broadleaf forest and seabird burrow; 4, JNPC, Titi Island, 03 / April / 1990, J. I. Townsend. Unknown locality: 2, 1476, T. Broun Collection, A. E. Brookes Collection. Associated females (57 ♀♀): ND: 15, Mokohinau Is., Fanal I., 01 / Mar / 1978, G. Kuschel, sifted litter; AK: 17, Lynfield, 27 / Jan / 1977, G. Kuschel, litter; 4, Coll. A. E. Brookes, Sep / 1914, Titirangi, Waitakere Rg., Auckland, 1476; 1, Bethells Matuku Res., 26 / Jul – 20 / Aug / 1983, G. Hall and P. A. Maddison, pit trap; 1, Riverhead, Forest Reserve, 30 / Aug / 1982, P. A. Maddison, litter; GB: 11, Taikawakawa, 21 / Sep / 1992, G. Hall & R. C. Henderson, sifted litter, 92 / 58; BP: 1, LUNZ, Te Koau, Twin Puiris, 15 / iii / 1993, R. M. Gilbert, J. W. M. Marris; HB: 2, Motu-okura (Bare I.), 07 / Feb – 20 / Mar / 1995, G. Hall, pit traps; SD: 5, LUNZ, Titi Island, 17 / xi / 1994, J. W. M. Marris, litter ex. broadleaf forest and seabird burrow.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D862FF8EFF017369FB91F919.taxon	description	Fig. 20; map Fig. 51	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D862FF8EFF017369FB91F919.taxon	diagnosis	Diagnosis: Head with vertexal foveae punctiform; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 6, A 7 and A 8 unmodified, A 5 greatly enlarged and asymmetrical, A 9 highly transverse and asymmetrical, ventral side broadly impressed, with ovoid lamina at end of inner side, A 10 longest and asymmetrical, irregularly shaped, with deep basal excavation at ventral side. Abdominal ventrite 2 with pair of large discal protuberances, inserted with strongly curved small apical setae; ventrite 6 unknown. Protrochanters with short ventral spine, about two-fifths of protrochanteral width; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D862FF8EFF017369FB91F919.taxon	description	Description: Body length 1.51 mm; body colour reddish brown with lighter coloured elytra, tibiae and tarsi. Head wider than long; frons slightly narrowed at end, about half width of head at eye level; eyes well developed, conspicuously prominent in dorsal view; vertexal foveae punctiform; antennal tubercles moderately developed; postantennal notches broad and shallow; postantennal cavities large, about as long as antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 6, A 7 and A 8 unmodified, A 5 greatly enlarged and asymmetrical, A 9 highly transverse and asymmetrical, ventral side broadly impressed, with ovoid lamina at end of inner side, A 10 longest and asymmetrical, irregularly shaped, with deep basal excavation at ventral side. Pronotum about as long as wide, slightly wider than head at eye level; lacking lateral antebasal foveae. Elytra as long as wide, widest at apical two-thirds; humeral calli weak, slightly prominent in dorsal view; sutural stria distinctively extending from base to apex. Abdomen with tergite 1 about 1.4 × longer than tergite 2; ventrite 2 with pair of large discal protuberances, inserted with strongly curved small apical setae; ventrite 6 unknown. Legs with protrochanters modified with short ventral spine, about two-fifths of protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.32 mm long, with long parameres bent downwards in lateral view, apex curved towards each other, nearly meet at middle, with pair of long subapical setae; median lobe with two large and stout sclerites, twisted near apex, about as wide as parameres.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D862FF8EFF017369FB91F919.taxon	discussion	Comments: This species is described from a singleton, and unfortunately, ventrite 6 was lost during dissecting. It is similar to E. (B.) platyarthra by having A 9 transverse but can be separated easily from it by its A 5 enlarged and transverse (Figs. 20 B – D), which is unmodified in E. (B.) platyarthra.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D862FF8EFF017369FB91F919.taxon	distribution	Distribution: South Island. NN.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D862FF8EFF017369FB91F919.taxon	etymology	Etymology: This species is named after one of the collectors for the holotype, Grace Hall, long time technician at NZAC who has helped significantly with Coleoptera research.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D862FF8EFF017369FB91F919.taxon	materials_examined	Type material (NZAC): Holotype. ♂, body parts mounted on acetate card: “ NEW ZEALAND, NN, Heaphy Track, Lewis Hut, 7 Nov 1999, R. Leschen, G. Hall, leaf litter 40 ° 56 ' S, 172 ° 09 ' E ”.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D863FF8CFF01716DFD00FCF9.taxon	description	Fig. 21; map Fig. 51	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D863FF8CFF01716DFD00FCF9.taxon	materials_examined	Type locality: [NN] Mount Saint Arnaud, Nelson. Type depository: BMNH LT ♂, 2 PLT; NZAC 2 PLT.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D863FF8CFF01716DFD00FCF9.taxon	diagnosis	Diagnosis: Head with vertexal foveae punctiform; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 6, A 7, A 8 and A 10 unmodified, A 5 greatly enlarged and asymmetrical, flag-shaped, dorsal side broadly concave, A 7 asymmetrical, A 8 disc-shaped and slightly asymmetrical, A 9 largest, with laminate base and boot-shaped protuberance at lateral side, forming gulf-shaped structure, A 10 conical. Abdominal ventrite 2 with pair of long and slender discal protuberances, inserted with small and strongly curved apical setae; ventrite 6 with pair of long medi- an struts, about half of ventral length. Protrochanters with long ventral spine, about three-fourths of protrochanteral width; protibiae with blunt apical protuberance; mesotibiae with apical minute triangle spine.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D863FF8CFF01716DFD00FCF9.taxon	description	Redescription: Body length 1.20 – 1.30 mm; body colour dark brown with lighter coloured tarsi. Head slightly wider than long; frons slightly narrowed at end, about half width of head at eye level; eyes slightly prominent, with about 32 facets; vertexal foveae punctiform; lacking antennal tubercles and postantennal notches; postantennal cavities barely visible in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 6, A 7, A 8 and A 10 unmodified, A 5 greatly enlarged and asymmetrical, flag-shaped, dorsal side broadly concave, A 7 asymmetrical, A 8 disc-shaped and slightly asymmetrical, A 9 largest, with laminate base and boot-shaped protuberance at lateral side, forming gulf-shaped structure, A 10 conical. Pronotum about as long as wide, wider than head at eye level; lacking lateral antebasal foveae. Elytra slightly wider than long, widest at apical three-fourths; humeral calli barely visible; sutural stria weakly extending from base to apex. Abdomen with tergite 1 about 1.2 × longer than tergite 2; ventrite 2 with pair of long and slender discal protuberances, inserted with small and strongly curved apical setae; ventrite 6 with pair of long median struts, about half of ventral length. Legs with protrochanters modified with long ventral spine, about three-fourths of protrochanteral width; protibiae with blunt apical protuberance; mesotibiae with apical minute triangle spine. Aedeagus about 0.34 mm long, with parameres wide and long, slightly curved inwards, with three pairs of apical setae; median lobe relatively small, with two sclerites much narrower than parameres, strongly curved towards each other in dorsal view, basal bulb slightly longer than two sclerites.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D863FF8CFF01716DFD00FCF9.taxon	discussion	Comments: This species can be immediately recognized by its bizarre form of A 9 (Figs. 21 B – C).	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D863FF8CFF01716DFD00FCF9.taxon	distribution	Distribution: North Island. TO. South Island. BR. Additional material examined: 8 ♂♂, TO: 3, Whanganui V., 792 m Whenuakura Flats, 11 / Oct / 1979, J. S. Dugdale, bryophytes, 79 / 94; 1, Tongariro NP., Chateau, 1000 m, 6 / Dec / 1978, G. Kuschel, Dacrydium bidwillii; NN: 1, DSC, Nelson Lakes N. P., N. slope Mt. Robert, Pinchgut Track, 950 m, 14 / xii / 84 – 6 / i / 85, Nothofagus spp. forest, A. Newton / M. Thayer, 707, flight intercept (window) trap; 1, FMNH, Nelson Lakes N. P., N. slope Mt. Robert, Pinchgut Track, 1290 m, 18 – 26 / xii / 1984, Noth. Sol. Elfin For., A. Newton / M. Thayer, 716; BR: 1, Mt. Sewell, TV station, 853 m, 22 / Nov / 1984, B. A. Holloway; 1, FMNH, Lewis Pass Nat. Res., 0.6 km S Lewis Pass, 870 m, 17 / xii / 1984 – 21 / i / 1985, Nothofagus spp. forest, A. Newton / M. Thayer, 714. Associated females (1 ♀): NN: Arnaud, T. Broun Collection, A. E. Brookes Collection.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D861FF8BFF01754DFE16F94A.taxon	description	Fig. 22; map Fig. 51	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D861FF8BFF01754DFE16F94A.taxon	materials_examined	Type locality: [BP] Tarukenga, near Rotorua. Type depository: BMNH LT ♂.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D861FF8BFF01754DFE16F94A.taxon	diagnosis	Diagnosis: Head with vertexal foveae distinct and asetose; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 cylindrical, about 3 × longer than A 8, with medium-sized spine surrounded in small anterolateral excavation, A 10 conical and slightly longer than A 9. Abdominal ventrite 2 with pair of small discal protuberances, inserted with slightly curved small apical setae; ventrite 6 with pair of slender median struts, about three-fourths of ventral length. Protrochanters with medium-sized ventral spine, about three-fifths of protrochanteral width; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D861FF8BFF01754DFE16F94A.taxon	description	Redescription: Body length 1.45 – 1.55 mm; body colour reddish brown to dark brown with lighter coloured elytra and legs. Head wider than long; frons not narrowed at end, about three-fourths width of head at eye level; eyes well developed, with about 25 facets; vertexal foveae distinct and asetose; antennal tubercles barely visible; postantennal notches weak; postantennal cavities large, about as long as antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 cylindrical, about 3 × longer than A 8, with medium-sized spine surrounded in small anterolateral excavation, A 10 conical and slightly longer than A 9. Pronotum slightly wider than long, slightly wider than head at eye level; lacking lateral antebasal foveae. Elytra about as long as wide, widest at apical three-fourths; humeral calli barely visible; sutural stria shallowly extending from base to apex. Abdomen with tergite 1 about 1.3 × longer than tergite 2; ventrite 2 with pair of small discal protuberances, inserted with slightly curved small apical setae; ventrite 6 with pair of slender median struts, about three-fourths of ventral length. Legs with protrochanters modified with medium-sized ventral spine, about three-fifths of protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.28 mm long, with parameres slightly curved inwards, widest at basal two-thirds, bearing four pairs of long apical setae and pair of short apical setae; median lobe complex, with three long sclerites, left one and middle one about as wide as parameres in dorsal view, strongly curved towards each other, right one straight and long, with pair of slightly sclerotized laminae next to three sclerites at dorsal side, with another pair of strongly sclerotized basal laminae strongly curved at ventral side, about one-third length of parameres.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D861FF8BFF01754DFE16F94A.taxon	discussion	Comments: The lectotype of E. (B.) hectori was compared with the lectotype and two paralectotypes of E. (B.) diversides (as E. (B.) diversa), and were found to represent a single species by having identical modifications of A 9 (Figs. 22 B – D), with the latter name treated as the junior synonym. Newton (2017) proposed a new name, E. (B.) diversides for the junior primary homonym, Bryaxis diversa which is now unavailable, and is also here treated as the junior synonym of E. (B.) hectori.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D861FF8BFF01754DFE16F94A.taxon	distribution	Distribution: North Island. AK, WO, CL, BP, WN. Additional material examined: 14 ♂♂, CL: 2, Rakitu I., 167 m, 07 / Jan / 1981, L. Roberts, litter, 81 / 7; 2, Coromandel, Twin Kauri Tk., 03 / Nov / 2005, R. Leschen & S. Nomura, flood debris, 37 ° 00 ' S 175 ° 38 ' E; 2, JNPC, WN, Levin L Papaitonga, 17 / XI / 2015, forest floor litter, J. Nunn; WO: 1, Collector, 14 / 3 / 1920, A. E. Brookes, Okauia Kaimai, Matamata, A. E. Brookes Collection; BP: 1, Kaimai, May / 1920, A. E. Brookes Collection; 6, Kaimai Range, Leyland Tramline, 427 m, 22 / Sep / 1981, B. M. May, litter, 81 / 92. Associated females (14 ♀♀): AK: 1, Ex. Broun Collection, Hunua Ra., Maketu, Auckland, 2735, Byraxis diversa Broun, T. Broun Collection, A. E. Brookes Collection; CL: 4, Rakitu I., 167 m, 07 / Jan / 1981, L. Roberts, litter, 81 / 7; 1, 2739, Broun; BP: 8, Top Kaimai, 16 / 12 / 35, A. E. Brookes Collection.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D867FF89FF017691FED5FC35.taxon	description	Fig. 23; map Fig. 51	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D867FF89FF017691FED5FC35.taxon	diagnosis	Diagnosis: Head lacking vertexal foveae; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7, A 8 and A 10 unmodified, A 6 and A 7 asymmetrical and subequal in size, A 8 shortened, about half length of A 7, A 9 greatly prolonged and enlarged, slightly impressed subapically at ventral side, A 10 largest and conical, about 1.4 × longer than A 9, slightly impressed at base in ventral view. Abdominal ventrite 2 with pair of minute discal protuberances, inserted with slightly curved small apical setae; ventrite 6 unknown. Protrochanters, protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D867FF89FF017691FED5FC35.taxon	description	Description: Body length 1.57 mm; body colour reddish brown with lighter coloured legs. Head wider than long; frons slightly narrowed at end, about two-fifths width of head at eye level; eyes moderately developed, with about 28 facets; lacking vertexal foveae; antennal tubercles barely visible; postantennal notches weakly demarcated; postantennal cavities large, slightly shorter than antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7, A 8 and A 10 unmodified, A 6 and A 7 asymmetrical and subequal in size, A 8 shortened, about half length of A 7, A 9 greatly prolonged and enlarged, slightly impressed subapically at ventral side, A 10 largest and conical, about 1.4 × longer than A 9, slightly impressed at base in ventral view. Pronotum as long as wide, about as wide as head at eye level; lacking lateral antebasal foveae. Elytra slightly wider than long, widest at apical two-thirds; humeral calli weak; sutural stria distinctively extending from base to apex. Abdomen with tergite 1 about 1.7 × longer than tergite 2; ventrite 2 with pair of minute discal protuberances, inserted with slightly curved small apical setae; ventrite 6 unknown. Legs with protrochanters, protibiae and mesotibiae un- modified. Aedeagus about 0.26 mm long, with long and stout parameres curved towards each other at apex, bent downwards in lateral view, widest near middle, with pair of long subapical setae and four pairs of short apical setae; median lobe lacking sclerites, with pair of fork-shaped laminae near base, lateral apical margin of median lobe sclerotized.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D867FF89FF017691FED5FC35.taxon	discussion	Comments: This is the only species within the subgenus Byraxis with protrochanters lacking a ventral spine (Fig. 23 D).	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D867FF89FF017691FED5FC35.taxon	distribution	Distribution: South Island. NN.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D867FF89FF017691FED5FC35.taxon	etymology	Etymology: This species is named after one of its collectors, Robert Hoare, lepidopterist at NZAC and assisted us with naming our new species.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D867FF89FF017691FED5FC35.taxon	materials_examined	Type material (NZAC): Holotype. ♂, specimen entire, glued on card, aedeagus + one antenna + one proleg mounted on acetate card: “ NEW ZEALAND, NN, Oparara Gorge, 19 Feb 1999, R. Leschen & R. Hoare, at light, 41 ° 13 ' S 172 ° 09 ' E ”.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D864FF89FF017499FD1FF979.taxon	description	Fig. 24; map Fig. 51	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D864FF89FF017499FD1FF979.taxon	diagnosis	Diagnosis: Head with vertexal foveae punctiform; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 6, A 7, A 8 and A 10 unmodified, A 4 asymmetrical, lateral side strongly extended, A 5 largest and somewhat beanshaped, with apex subacute and base acute in lateral view, A 9 cylindrical and elongate, about 3.5 × longer than A 8, A 10 conical, about 1.5 × longer than A 9. Abdominal ventrite 2 with pair of flat and wide discal protuberances, inserted with strongly curved large apical setae; ventrite 6 with pair of slender median struts, about half of ventral length. Protrochanters with long ventral spine, about half of protrochanteral width; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D864FF89FF017499FD1FF979.taxon	description	Description: Body length 1.41 – 1.43 mm; body colour dark brown with lighter coloured elytra and legs. Head as long as wide; frons not narrowed at end, about three-fourths width of head at eye level; eyes moderately developed, slightly prominent in dorsal view, with about 29 facets; vertexal foveae punctiform; antennal tubercles weakly developed; lacking postantennal notches; postantennal cavities long and narrow, slightly shorter than antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 6, A 7, A 8 and A 10 unmodified, A 4 asymmetrical, lateral side strongly extended, A 5 largest and somewhat bean-shaped, with apex subacute and base acute in lateral view, A 9 cylindrical and elongate, about 3.5 × longer than A 8, A 10 conical, about 1.5 × longer than A 9. Pronotum about as long as wide, wider than head at eye level; with punctiform lateral antebasal foveae. Elytra as long as wide, widest at middle; humeral calli indistinctive, slightly prominent in dorsal view; sutural stria barely visible from base to middle, shallowly extending from middle to apex. Abdomen with tergite 1 about twice length of tergite 2; ventrite 2 with pair of flat and wide discal protuberances, inserted with strongly curved large apical setae; ventrite 6 with pair of slender median struts, about half of ventral length. Legs with protrochanters modified with long ventral spine, about half of protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.28 mm long, with long and sinuate parameres, bent downwards in lateral view, apex curved towards each other, with pair of long subapical setae; median lobe with two large sclerites, base nearly meet, strongly curved from middle and entwined apically.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D864FF89FF017499FD1FF979.taxon	discussion	Comments: This species is most similar to E. (B.) minuta and E. (B.) ovalis by their similar antennal modifications but can be differentiated from both by having its protrochanters modified with long ventral spine that is otherwise barely visible (Figs. 33 D; 39 E).	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D864FF89FF017499FD1FF979.taxon	distribution	Distribution: South Island. KA.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D864FF89FF017499FD1FF979.taxon	etymology	Etymology: This species is named after HuiZhen Yu, the first author’s mother, for her unending support.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D864FF89FF017499FD1FF979.taxon	materials_examined	Type material (NZAC): Holotype. ♂, specimen entire, card-mounted: “ NEW ZEALAND, KA, Mt. Fyffe Hut, 17 Jan 2002, R. Leschen, L. Dunning, under rock, 42 ° 19.437 ' S, 173 ° 35.573 ' E, 1105 m ”. Paratypes. 1 ex.: KA: Oaro, 15 – 22 / Mar / 1980, J. W. Early, Malaise trap.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D865FF87FF01735AFD9CFFFD.taxon	description	Fig. 25; map Fig. 51	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D865FF87FF01735AFD9CFFFD.taxon	materials_examined	Type locality: [MC] Broken River, Canterbury. Type depository: BMNH LT ♂, 1 PLT.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D865FF87FF01735AFD9CFFFD.taxon	diagnosis	Diagnosis: Head with vertexal foveae punctiform; antennae 10 - segmented with 3 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 large and asymmetrical, strongly transverse and obliquely truncate at apex, with lateral acute spine, A 10 largest and conical, with basal excavation. Abdominal ventrite 2 with pair of large discal protuberances, inserted with strongly curved large apical trichomes; ventrite 6 unknown. Protrochanters with minute ventral spine, about one-tenth of protrochanteral width; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D865FF87FF01735AFD9CFFFD.taxon	description	Redescription: Body length 1.64 mm. body colour reddish brown with lighter coloured tarsi. Head slightly wider than long; frons not narrowed at end, about three-fourths width of head at eye level; eyes moderately developed, with about 31 facets; vertexal foveae punctiform; antennal tubercles weak; postantennal notches weak; postantennal cavities large, about as long as antennal tubercles in dorsal view; antennae 10 - segmented with 3 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 large and asymmetrical, strongly transverse and obliquely truncate at apex, with lateral acute spine, A 10 largest and conical, with basal excavation. Pronotum about as long as wide, wider than head at eye level; lacking lateral antebasal foveae. Elytra as long as wide, widest at about apical three-fourths; humeral calli slightly prominent; sutural stria distinctively extending from base to apex. Abdomen with tergite 1 about twice length of tergite 2; ventrite 2 with pair of large discal protuberances, inserted with strongly curved large apical trichomes; ventrite 6 unknown. Legs with protrochanters modified with minute ventral spine, about one-tenth of protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus unknown.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D865FF87FF01735AFD9CFFFD.taxon	discussion	Comments: Only two type specimens are known. We did not dissect syntypes since males of this species have distinctive A 9 modifications and two tufts of large trichomes on ventrite 2 (Figs. 25 B – C) that can immediately differentiate it from congeners.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D865FF87FF01735AFD9CFFFD.taxon	distribution	Distribution: South Island. MC.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D86AFF85FF01710BFBFCFB95.taxon	description	Fig. 26; map Fig. 52	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D86AFF85FF01710BFBFCFB95.taxon	materials_examined	Type locality: [AK] Auckland. Type depository: BMNH LT ♂, 3 PLT.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D86AFF85FF01710BFBFCFB95.taxon	diagnosis	Diagnosis: Head with vertexal foveae punctiform; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 6 slightly asymmetrical, A 9 largest, about 1.5 × longer than A 10, with apical impression at ventral side, impression about two-fifths length of A 9, inserted with a barely visible protuberance at middle, A 10 conical, with small basal impression at ventral side. Abdominal ventrite 2 with pair of barely visible discal protuberances, inserted with small apical setae; ventrite 6 with pair of long median struts, about half of ventral length. Protrochanters with a medium-sized ventral spine, about half of protrochanteral width; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D86AFF85FF01710BFBFCFB95.taxon	description	Redescription: Body length 1.38 – 1.49 mm; body colour reddish brown to dark brown with lighter coloured elytra and legs. Head wider than long; frons slightly narrowed at end, about half width of head at eye level; eyes well developed and prominent, with about 35 facets; vertexal foveae punctiform; antennal tubercles moderately developed; postantennal notches weak; postantennal cavities large, slightly shorter than antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 6 slightly asymmetrical, A 9 largest, about 1.5 × longer than A 10, with apical impression at ventral side, impression about twofifths length of A 9, inserted with a barely visible protuberance at middle, A 10 conical, with small basal impression at ventral side. Pronotum slightly wider than long, as wide as head at eye level; lateral antebasal foveae punctiform. Elytra slightly wider than long, widest at apical two-thirds; lacking humeral calli; sutural stria barely visible. Abdomen with tergite 1 slightly more than twice length of tergite 2; ventrite 2 with pair of barely visible discal protuberances, inserted with small apical setae; ventrite 6 with pair of long median struts, about half of ventral length. Legs with protrochanters modified with a medium-sized ventral spine, about half of protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.28 mm long, with slender parameres, bent near base and sinuate towards apex, curved downwards, with two pairs of apical short setae and pair of subapical long setae; median lobe bent downwards in lateral view, with two long sclerites, about as long as parameres, both sclerites curved inwards at apex in dorsal view, left one slightly shorter than right one.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D86AFF85FF01710BFBFCFB95.taxon	discussion	Comments: The holotype of E. (B.) foveatissima and the lectotype of E. (B.) sanguinea were examined and were found having the identical modifications of A 9 as the lectotype of E. (B.) impar. These names are formally synonymized here, with the fomer two names treated as junior synonyms. This species is similar to E. (B.) complector by having similar form of A 9 but can be distinguished by its A 9 with a barely visible protuberance (Fig. 26 B), which is a conspicuous apical spine in E. (B.) complector (Fig. 10 B).	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D86AFF85FF01710BFBFCFB95.taxon	distribution	Distribution: North Island. ND, AK, WO, CL, TO, BP, GB. Additional material examined: 67 ♂♂, AK: 1, 239, Clevedon; 1, Coll. A. E. Brookes, 01 / Jan / 1916, Hunua Rg., Drury, Auckland, 239; 31, Lynfield, 10 / May / 1975, G. Kuschel, litter; 4, Bethells Matuku Res., 19 / Sep – 31 / Oct / 1983, M. F. Tocker & B. G. Bennett; 1, AMNZ, Auckland Domain, 40 m, 12 / Aug / 2001, S. E. Thorpe, under rotten log; CL: 1, Little Barrier I., Pohutukawa Flat, foreshore, 03 / Oct / 1975, B. M. May, litter; 1, Little Bay, Kennedy Block, 22 / Nov / 1993, A. Larochelle, litter; WO: 1, Waharoa, 04 / Jan / 1992, P. A. Maddison, litter, 92 / 1; TO: 1, Pureora SF. Park, 610 m, 20 / Oct / 1991, B. M. May, litter; GB: 3, East Cape, Taikawakawa, 15 / Apr / 1992, G. Hall & R. C. Henderson, sifted litter, 92 / 58; 1, L. Waikaremoana Whareama Peninsula, Dec / 2004, pit traps, Station 5; 1, Gray’s Bush, Gisborne Flats, 20 / Aug / 1976, A. R. Ferguson, litter; ND: 7, Kaikohe, 1289, Apr / 1954, R. Hornabrook; BP: 5, Lottin Pt Rd., Waenga Bush, 16 / Sep / 1992, R. C. Henderson, litter; 2, Mt. Te Aroha smt., 03 / Nov / 1993, J. S. Dugdale, litter and liverworts; 2, Te Koau, Bush Walk, 243 m, 15 / Mar – 29 / Apr / 1993, J. S. Dugdale, Malaise trap; 1, Upper Waipiata Karakatuwhero V., 03 / Feb / 1993, J. S. Dugdale; 1, 243, Unknown locality: T. Broun Collection, A. E. Brookes Collection; 1, AMNZ, T. Broun Collection. Associated females (74 ♀♀): ND: 9, Kaikohe, 1289, Apr / 1954, R. Hornabrook; AK: 12, Coll. A. E. Brookes, 01 / Jan / 1916, Hunua Rg., Drury, Auckland, 239; 50, Lynfield, 10 / May / 1975, G. Kuschel, litter; NN: 3, Dun Mt., 2000 ft, 6 – 10 / 20, A. Philpott.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D868FF84FF0173F9FC8FF9E8.taxon	description	Fig. 27; map Fig. 52	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D868FF84FF0173F9FC8FF9E8.taxon	materials_examined	Type locality: [CL] Tairua. Type depository: BMNH HT ♂.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D868FF84FF0173F9FC8FF9E8.taxon	diagnosis	Diagnosis: Head with vertexal foveae punctiform; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7, A 8 and A 10 unmodified, A 9 cylindrical and flat at dorsal side, with apical spine at lateral side, A 10 conical and slightly longer than A 9. Abdominal ventrite 2 with pair of large discal protuberances, inserted with large and strongly curved apical setae; ventrite 6 with apical margin sinuate, with pair of short and acute median struts, about one-seventh of ventral length. Protrochanters with small ventral spine, about one-third of protrochanteral width; protibiae with minute spine in apical two-thirds; mesotibiae with barely visible subapical spine.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D868FF84FF0173F9FC8FF9E8.taxon	description	Redescription: Body length 1.68 – 1.72 mm; body colour reddish brown with lighter coloured tarsi. Head slightly wider than long; frons not narrowed at end, about three-fourths width of head at eye level; eyes well developed and prominent, with about 30 facets; vertexal foveae punctiform; antennal tubercles weak; postantennal notches weak; postantennal cavities medium-sized, about three-fourths width of antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7, A 8 and A 10 unmodified, A 9 cylindrical and flat at dorsal side, with apical spine at lateral side, A 10 conical and slightly longer than A 9. Pronotum wider than long, slightly wider than head at eye level; lacking lateral antebasal foveae. Elytra slightly longer than wide, widest at middle; humeral calli weak; sutural stria shallowly extending from base to apex. Abdomen with tergite 1 about 1.4 × longer than tergite 2; ventrite 2 with pair of large discal protuberances, inserted with large and strongly curved apical setae; ventrite 6 with apical margin sinuate, with pair of short and acute median struts, about oneseventh of ventral length. Legs with protrochanters modified with small ventral spine, about one-third of protrochanteral width; protibiae with minute spine in apical two-thirds; mesotibiae with barely visible subapical spine. Aedeagus about 0.31 mm long, with parameres long and wide, narrowed at base and sinuate towards apex, strongly bent downwards at apex in lateral view; median lobe small, with two small and slender sclerites, strongly curved towards each other at apex, basal bulb large, about as long as median lobe.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D868FF84FF0173F9FC8FF9E8.taxon	discussion	Comments: This species can be separated from all other Eupines by the protibiae and mesotibiae modified with small or minute spines (Figs. 27 E – F).	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D868FF84FF0173F9FC8FF9E8.taxon	distribution	Distribution: North Island. AK, CL. South Island. BR. Additional material examined: 3 ♂♂, AK: 1, Clevedon, T. Broun Collection, A. E. Brookes Collection; BR: 1, JNPC, Pororari River mouth Punakaiki, 29 / Dec / 2010, in flood debris, J. T. Nunn collection; 1, LUNZ, Okari River Cape Foulwind, N. Z., 29 / Oct / 1970, Ento. Dept. Field Trip.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D86EFF82FF017691FE20FCF9.taxon	description	Fig. 28; map Fig. 52	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D86EFF82FF017691FE20FCF9.taxon	diagnosis	Diagnosis: Head with vertexal foveae punctiform; antennae 10 - segmented with 2 - segmented antennal club; A 7, A 8 and A 10 unmodified, A 4 disc-shaped, about half length of A 3, with long spine at lateral side, A 5 largest, with large excavation at middle, outer side slightly impressed, with long and laminate basal projection, inner side impressed at apex, A 6 enlarged and strongly asymmetrical, outer side with scoop-shaped impression, A 9 cylindrical and slightly asymmetrical at apex in dorsal view, A 10 conical, about 1.5 × longer than A 9. Abdominal ventrite 2 with pair of flat and wide discal protuberances, inserted with strongly curved large apical setae, nearly meet at middle; ventrite 6 with pair of short and stout median struts, about two-fifths of ventral length. Protrochanters with minute ventral spine, about one-ninth of protrochanteral width; protibiae unmodified; mesotibiae with short subapical spine and long apical spine.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D86EFF82FF017691FE20FCF9.taxon	description	Description: Body length 1.47 – 1.49 mm; body colour dark brown with lighter coloured legs. Head wider than long; frons not narrowed at end, about three-fourths width of head at eye level; eyes moderately developed, slightly prominent in dorsal view, with about 30 facets; vertexal foveae punctiform; antennal tubercles weakly developed; lacking postantennal notches; postantennal cavities large, about as long as antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 7, A 8 and A 10 unmodified, A 4 disc-shaped, about half length of A 3, with long spine at lateral side, A 5 largest, with large excavation at middle, outer side slightly impressed, with long and laminate basal projection, inner side impressed at apex, A 6 enlarged and strongly asymmetrical, outer side with scoop-shaped impression, A 9 cylindrical and slightly asymmetrical at apex in dorsal view, A 10 conical, about 1.5 × longer than A 9. Pronotum as long as wide, wider than head at eye level; lacking lateral antebasal foveae. Elytra as long as wide, widest near middle; humeral calli weak, barely visible in dorsal view; sutural stria shallowly extending from base to apex. Abdomen with tergite 1 about 1.5 × longer than tergite 2; ventrite 2 with pair of flat and wide discal protuberances, inserted with strongly curved large apical setae, nearly meet at middle; ventrite 6 with pair of short and stout median struts, about two-fifths of ventral length. Legs with protrochanters modified with minute ventral spine, about one-ninth of protrochanteral width; protibiae unmodified; mesotibiae with short subapical spine and long apical spine. Aedeagus about 0.31 mm long, with symmetrical and large parameres, slightly sinuate, widest at middle, apex bent downwards in lateral view and curved towards each other, with two pairs of long subapical setae and pair of short subapical setae; median lobe with two highly curved sclerites, widest near base, much narrower than parameres, right one longer than left one in dorsal view.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D86EFF82FF017691FE20FCF9.taxon	discussion	Comments: This species can be distinguished from all other Eupines by having the mesotibia modified with a long apical spine and A 6 modified (Figs. 28 B – D, 28 H).	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D86EFF82FF017691FE20FCF9.taxon	distribution	Distribution: North Island. TO.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D86EFF82FF017691FE20FCF9.taxon	etymology	Etymology: This specific epithet is a Latin adjective meaning ‘ unusual’, referring to the unusual antennal form of this species.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D86EFF82FF017691FE20FCF9.taxon	materials_examined	Type material (NZAC): Holotype. ♂, specimen entire, card-mounted: “ 28. xi. 51, moss, Huka Falls, Taupo P 550 [handwritten] ”.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D86FFF81FF017530FB9DF81B.taxon	description	Fig. 29; map Fig. 52	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D86FFF81FF017530FB9DF81B.taxon	materials_examined	Type locality: [MC] Broken River, Canterbury. Type depository: BMNH HT ♂	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D86FFF81FF017530FB9DF81B.taxon	diagnosis	Diagnosis: Head with vertexal foveae punctiform; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 6, A 7, A 8 and A 10 unmodified, A 5 large and asymmetrical, bean-shaped, lateral margin laminate and curved, forming bowl-shaped excavation, A 7 asymmetrical, A 8 disc-shaped, A 9 large and asymmetrical, slightly shorter than A 10, inner side strongly excavated in dorsal view, with upper margin laminate and extended, forming gulfshaped structure, A 10 largest and conical, about 1.2 × longer than A 9. Abdominal ventrite 2 with pair of large discal protuberances slightly curving towards each other, inserted with strongly curved small apical setae; ventrite 6 with pair of short and acute median struts, about one-fourth of ventral length. Legs with protrochanters modified with small ventral spine, about one-third of protrochanteral width; protibiae and mesotibiae unmodified. Protrochanters with small ventral spine, about one-third of protrochanteral width; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D86FFF81FF017530FB9DF81B.taxon	description	Redescription: Body length 1.58 – 1.60 mm; body colour dark brown with lighter coloured legs. Head about as long as wide, frons slightly narrowed at end, about half width of head at eye level; eyes moderately developed, with about 29 facets; vertexal foveae punctiform; antennal tubercles weak; lacking postantennal notches; postantennal cavities medium-sized, about half length of antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 6, A 7, A 8 and A 10 unmodified, A 5 large and asymmetrical, bean-shaped, lateral margin laminate and curved, forming bowl-shaped excavation, A 7 asymmetrical, A 8 disc-shaped, A 9 large and asymmetrical, slightly shorter than A 10, inner side strongly excavated in dorsal view, with upper margin laminate and extended, forming gulf-shaped structure, A 10 largest and conical, about 1.2 × longer than A 9. Pronotum about as long as wide, wider than head at eye level; lacking lateral antebasal foveae. Elytra as long as wide, widest at apical two-thirds; humeral calli distinctively prominent; sutural stria distinctively extending from base to apex. Abdomen with tergite 1 about 1.8 × longer than tergite 2; ventrite 2 with pair of large discal protuberances slightly curving towards each other, inserted with strongly curved small apical setae; ventrite 6 with pair of short and acute median struts, about one-fourth of ventral length. Legs with protrochanters modified with small ventral spine, about one-third of protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.32 mm long, with sinuate parameres long and stout, widest near basal one-fourth, apex strongly bent downwards and twisted in lateral view, with three short apical setae; median lobe with two short sclerites, much shorter and narrower than parameres in dorsal view, left one distinctively curved outwards, right one point inwards.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D86FFF81FF017530FB9DF81B.taxon	discussion	Comments: The special antennal modifications (Figs. 29 B – D) can easily separate this species from all congeners.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D86FFF81FF017530FB9DF81B.taxon	distribution	Distribution: North Island. BP. South Island. NN, MC. Additional material examined: 5 ♂♂, BP: 2, BMNH, 3052. ♂, Rotoiti, 10 / 3 / 16, New Zealand, Broun Col- lection. Brit. Mus., 1922 – 482; NN: 2, FMNH, Nelson Lakes N. P., N. slope Mt. Robert, Pinchgut Track, 950 m, 14 / XII / 84 – 6 / I / 85, Nothofagus spp. forest, A. Newton / M. Thayer, 707, flight intercept (window) trap; MC: 1, BMNH, 3052, ♂, Rakaia, 9 / 8 / 1912, New Zealand, Broun Collection. Brit. Mus., 1922 – 482.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D86CFF9FFF017050FB24FFD8.taxon	description	Fig. 30; map Fig. 52	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D86CFF9FFF017050FB24FFD8.taxon	materials_examined	Type locality: New Zealand. Type depository: MNHN LT ♂; BMNH 2 PLT.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D86CFF9FFF017050FB24FFD8.taxon	diagnosis	Diagnosis: Head with vertexal foveae barely visible; antennae 10 - segmented with 4 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 7 and A 8 asymmetrical and short, A 9 with apical half impressed at dorsal side, lateral side excavated, with curved spine at upper margin of excavation, A 10 axe-shaped, with long setae around margin, broadly impressed at lateral and ventral side, strongly narrowed at base. Abdominal ventrite 2 with pair of large discal protuberances, inserted with short and wide apical setae; ventrite 6 with apical margin slightly sinuate, with pair of short median struts, slightly shorter than one-third of ventral length. Protrochanters with medium-sized ventral spine, about half of protrochanteral width; protibiae with minute apical protuberance; mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D86CFF9FFF017050FB24FFD8.taxon	description	Redescription: Body length 1.55 – 1.64 mm; body colour reddish brown to dark brown with lighter coloured elytra and legs. Head as long as wide; frons slightly narrowed at end, about half width of head at eye level; eyes well developed, with about 29 facets; vertexal foveae barely visible; antennal tubercles moderately developed; lacking postantennal notches; postantennal cavities large, slightly longer than antennal tubercles in dorsal view; antennae 10 - segmented with 4 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 7 and A 8 asymmetrical and short, A 9 with apical half impressed at dorsal side, lateral side excavated, with curved spine at upper margin of excavation, A 10 axe-shaped, with long setae around margin, broadly impressed at lateral and ventral side, strongly narrowed at base. Pronotum slightly longer than wide, about as wide as head at eye level; lacking lateral antebasal foveae. Elytra slightly longer than wide, widest at apical three-fifths; humeral calli barely prominent; sutural stria distinct and narrow, extending from base to apex. Abdomen with tergite 1 about 1.5 × longer than tergite 2; ventrite 2 with pair of large discal protuberances, inserted with short and wide apical setae; ventrite 6 with apical margin slightly sinuate, with pair of short median struts, slightly shorter than one-third of ventral length. Legs with protrochanters modified with medium-sized ventral spine, about half of protrochanteral width; protibiae with minute apical protuberance; mesotibiae unmodified. Aedeagus about 0.34 mm long, with long and stout parameres, sinuate towards apex, slightly narrowed at base, apex strongly curved inwards and twisted, forming hook-shaped end; median lobe small, with two symmetrical horn-shaped sclerites slightly curved towards each other in dorsal view, basal bulb large, nearly twice length of median lobe.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D86CFF9FFF017050FB24FFD8.taxon	discussion	Comments: The last antennal segment of E. (B.) longiceps (Figs 3.29 B – C) resembles the head of an axe, which can separate this species from all other Eupines.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D86CFF9FFF017050FB24FFD8.taxon	distribution	Distribution: North Island. WN. South Island. NC, MC, MK. Additional material examined: 69 ♂♂, WN: 16, JNPC, Waikawa Beach, 18 / Mar / 1990, J. T. Nunn Collection; NC: 9, JNPC, CC Deans Bush, Christchurch, 14 / May / 1990, J. T. Nunn Collection; 1, JNPC, Riccarton Bush, Christchurch, 23 / May / 1993, J. T. Nunn Collection; MC: 42, 13 / Oct / 1951, 21 / Jan / 1952 and 01 / Feb / 1952, Riccarton Bush, Christchurch S. I.; MK: 1, Pukaki Reserve, 08 / Jan / 2006, R. Leschen, T. Buckley, R. Hoare, tussock leaf litter, 44 ° 13 ' S 170 ° 01 ' E. Associated females (4 ♀♀): MC: 3, 13 / Oct / 1951 and 01 / Feb / 1952, Riccarton Bush, Christchurch S. I.; 1, JNPC, CC Deans Bush, Christchurch, 14 / May / 1990, J. T. Nunn Collection.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D872FF9EFF017412FE16FD89.taxon	description	Fig. 31; map Fig. 52	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D872FF9EFF017412FE16FD89.taxon	diagnosis	Diagnosis: Head with vertexal foveae punctiform; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7, A 8 and A 10 unmodified, A 9 asymmetrical and enlarged, slightly shorter than A 10, with ventromedial excavation and minute protuberance at lateral side, A 10 largest and conical. Abdominal ventrite 2 with pair of large discal protuberances close to each other, inserted with slightly curved medium-sized apical setae; ventrite 6 with pair of long and acute median struts, about one-third of ventral length. Protrochanters with short ventral spine, about two-fifths of protrochanteral width; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D872FF9EFF017412FE16FD89.taxon	description	Description: Body length 1.34 – 1.40 mm; body colour reddish brown to dark brown with lighter coloured elytra and legs. Head wider than long; frons not narrowed at end, about three-fourths width of head at eye level; eyes well developed and prominent in dorsal view, with about 34 facets; antennal tubercles moderately developed; postantennal notches broad and distinctive; vertexal foveae punctiform; postantennal cavities large, slightly shorter than antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7, A 8 and A 10 unmodified, A 9 asymmetrical and enlarged, slightly shorter than A 10, with ventromedial excavation and minute protuberance at lateral side, A 10 largest and conical. Pronotum slightly wider than long, as wide as head at eye level; lacking lateral antebasal foveae. Elytra slightly longer than wide, widest near middle; humeral calli slightly prominent in dorsal view; sutural stria weak, shallowly extending from base to apex. Abdomen with tergite 1 about 1.2 × longer than tergite 2; ventrite 2 with pair of large discal protuberances close to each other, inserted with slightly curved medium-sized apical setae; ventrite 6 with pair of long and acute median struts, about one-third of ventral length. Legs with protrochanters modified with short ventral spine, about two-fifths of protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.37 mm long, with long and stout parameres, curved towards each other at apex and bent downwards in lateral view, with three pairs of short subapical setae; median lobe with two slender sclerites, curved towards each other at apex, shorter and much narrower than parameres.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D872FF9EFF017412FE16FD89.taxon	discussion	Comments: This species can be distinguished from all other Eupines by its simple antennal modifications and ventrite 2 with pair of large discal protuberances close to each other (Figs. 31 B – D, 31 F).	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D872FF9EFF017412FE16FD89.taxon	distribution	Distribution: North Island. ND.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D872FF9EFF017412FE16FD89.taxon	etymology	Etymology: This species is named after the collector of the holotype, V. A. May	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D872FF9EFF017412FE16FD89.taxon	materials_examined	Type material (NZAC): Holotype. ♂, specimen entire, card-mounted: “ NEW ZEALAND, ND, Mangamuka Smt., 386 m, 13 Dec 1976, V. A. May Litter 76 / 109 ”. Paratypes. 1 ex.: Unknown locality: T. Broun Collection // A. E. Brookes Collection.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D873FF9DFF0175DDFE3CF9D3.taxon	description	Fig. 32; map Fig. 52	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D873FF9DFF0175DDFE3CF9D3.taxon	materials_examined	Type locality: New Zealand. Type depository: BMNH LT ♂, 1 PLT.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D873FF9DFF0175DDFE3CF9D3.taxon	diagnosis	Diagnosis: Head with vertexal foveae small and asetose; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 6, A 7 and A 8 unmodified, A 5 large and asymmetrical, about twice length of A 4, A 9 with narrow stria at middle and strongly excavated at lateral side near middle, with long apical spine at lateral side, A 10 bean-shaped, with broad basal impression at inner side, base strongly narrowed. Abdominal ventrite 2 with pair of minute protuberances, inserted with long and strongly curved apical setae; ventrite 6 with pair of long and acute median struts, slightly shorter than ventral length. Protrochanters with long ventral spine, about two-thirds of protrochanteral width; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D873FF9DFF0175DDFE3CF9D3.taxon	description	Redescription: Body length 1.60 – 1.68 mm; body colour dark brown with lighter coloured legs. Head wider than long; frons not narrowed at end, about three-fourths width of head at eye level; eyes well developed and prominent, with about 34 facets; vertexal foveae small and asetose; antennal tubercles weak; postantennal notches shallow and long; postantennal cavities large, about as long as antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 6, A 7 and A 8 unmodified, A 5 large and asymmetrical, about twice length of A 4, A 9 with narrow stria at middle and strongly excavated at lateral side near middle, with long apical spine at lateral side, A 10 bean-shaped, with broad basal impression at inner side, base strongly narrowed. Pronotum as long as wide, about as wide as head at eye level; with weak punctiform lateral antebasal foveae. Elytra about as long as wide, widest at middle; humeral calli slightly prominent; sutural stria shallowly extending from middle to apex. Abdomen with tergite 1 slightly longer than tergite 2; ventrite 2 with pair of minute protuberances, inserted with long and strongly curved apical setae; ventrite 6 with pair of long and acute median struts, slightly shorter than ventral length. Legs with protrochanters modified with long ventral spine, about two-thirds of protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.36 mm long, fin-shaped parameres narrow at base and gradually widened towards apex, apex curved inwards and slightly bent upwards in lateral view, with three pairs of long apical setae; median lobe with three slender sclerites nearly meet in dorsal view, left one and right one strongly bent near middle, middle one bifurcate near base, forming hole-shaped structure, with left sclerites pass through it.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D873FF9DFF0175DDFE3CF9D3.taxon	discussion	Comments: This species is characterized by its A 9 with a weak transverse suture, suggesting a pseudosegment (Figs. 32 B – C), a feature that is not found in congeners.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D873FF9DFF0175DDFE3CF9D3.taxon	distribution	Distribution: North Island. ND, CL. Additional material examined: 12 ♂♂, ND: 1, Tutukaka, 12 / Sep / 1980, J. C. Watt, sifted litter and nest; 5, Poor Knights Is., Aorangi Crater Bay, 11 – 16 / Nov / 1981, J. S. Dugdale, Malaise trap in coastal scrub; CL: 1, Tairua, E. coast North Is., Ex. Broun Collection, 231, T. Broun Collection, A. E. Brookes Collection; 1, Mercury Is., Korapuki I., 15 / Dec / 1987, G. Hall sifted litter; 1, Mercury Is., Green I., 23 / Dec / 1987, G. Hall, sifted litter; 1, Mercury Is., Middle I., 20 / Oct / 1983, leaf litter, I. Southey; 1, AMNZ, Greater Barrier I., Little Windy Hill, 100 m, 20 / Jan – 25 / Feb / 2003, K. Parsons; 1, Cuvier I., Wilson Peak, 100 m, 12 / Nov / 1999, J. W. Early, S. E. Thorpe close to lighthouse. Unknown locality: 1, Ex. Broun Collection, 231. Associated females (10 ♀♀): ND: 6, Tutukaka, 12 / Sep / 1980, J. C. Watt, sifted litter and nest; 1, Poor Knights Is., Aorangi Crater Bay, 11 – 16 / Nov / 1981, J. S. Dugdale Malaise trap in Coastal scrub; CL: 2, Mercury Is., Korapuki I., 15 / Dec / 1987, G. Hall, sifted litter. Unknown locality: 1, Ex. Broun Collection, 231.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D871FF9BFF017691FB26FC35.taxon	description	Fig. 33; map Fig. 53	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D871FF9BFF017691FB26FC35.taxon	diagnosis	Diagnosis: Head with vertexal foveae punctiform; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 6, A 7, A 8 and A 10 unmodified, A 5 greatly enlarged and asymmetrical, lateral side with large and shallow basal impression, A 9 cylindrical, about 5 × longer than A 8, A 10 conical, about 1.4 × longer than A 9. Abdominal ventrite 2 with pair of large and round discal protuberances, inserted with strongly curved small anterolateral setae; ventrite 6 with pair of short and acute median struts, about one-fourth of ventral length. Protrochanters with minute ventral spine, about one-seventh of protrochanteral width; protibiae unmodified; mesotibiae with minute apical spine.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D871FF9BFF017691FB26FC35.taxon	description	Description: Body length 1.41 – 1.48 mm; body colour dark brown with lighter coloured tarsi. Head as long as wide; frons greatly narrowed at end, about one-third width of head at eye level; eyes moderately developed, slightly prominent in dorsal view, with about 27 facets; vertexal foveae punctiform; antennal tubercles weak; postantennal notches distinctive and small; postantennal cavities small, about half length of antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 6, A 7, A 8 and A 10 unmodified, A 5 greatly enlarged and asymmetrical, lateral side with large and shallow basal impression, A 9 cylindrical, about 5 × longer than A 8, A 10 conical, about 1.4 × longer than A 9. Pronotum about as long as wide, wider than head at eye level; with punctiform lateral antebasal foveae. Elytra as long as wide, widest at middle; humeral calli weak, slightly prominent in dorsal view; sutural stria shallowly extending from base to apex. Abdomen with tergite 1 about twice length of tergite 2; ventrite 2 with pair of large and round discal protuberances, inserted with strongly curved small anterolateral setae; ventrite 6 with pair of short and acute median struts, about one-fourth of ventral length. Legs with protrochanters modified with minute ventral spine, about one-seventh of protrochanteral width; protibiae unmodified; mesotibiae with minute apical spine. Aedeagus about 0.32 mm long, with long and stout parameres, curved downwards in lateral view, apex slightly curved, with pair of long subapical setae; median lobe with two long and stout sclerites, slightly twisted at apex, slightly shorter and narrower than parameres.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D871FF9BFF017691FB26FC35.taxon	discussion	Comments: This species is characterized by a A 5 greatly enlarged and asymmetrical (Figs. 33 B – C).	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D871FF9BFF017691FB26FC35.taxon	distribution	Distribution: South Island. SD, NN.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D871FF9BFF017691FB26FC35.taxon	etymology	Etymology: The specific epithet refers to the minute apical spine on mesotibiae of this species.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D871FF9BFF017691FB26FC35.taxon	materials_examined	Type material (NZAC): Holotype. ♂, specimen in two parts, head + thorax glued on card, abdomen mounted on acetate card: “ NEW ZEALAND, NN, Stockton Coal Mine W 6 C dump 880 m, 13 – 20 Dec 2000 [handwritten], R. J. Toft // Rehabilitation trial Bare control plot Malaise trap, 9 [handwritten], RT ”. Paratype. 3 ex.: SD: 2, FMNH, Tennyson Inlet, west side Te Mako Bay, 125 m, 15 / xii / 1984 – 5 / i / 1985, Nothofagus-podo-hdwd., A. Newton / M. Thayer, 710; NN: 1, Mt. Arthur, 1200 m, Nelson, 13 – 20 / Nov / 1969, B. May & S. Silcock.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D876FF99FF017499FC37FFD9.taxon	description	Fig. 34; map Fig. 53	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D876FF99FF017499FC37FFD9.taxon	materials_examined	Type locality: Not indicated in the paper (Reitter 1880). Type depository: MNHN LT ♂, 1 PLT.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D876FF99FF017499FC37FFD9.taxon	diagnosis	Diagnosis: Head with vertexal foveae punctiform; antennae 10 - segmented with 3 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 7 asymmetrical, about twice length of A 8, A 8 disc-shaped, A 9 large, transverse and cupulate, with ventral impression and basolateral excavation, with hook-shaped spine at upper margin of excavation, A 10 largest, forming claw-shaped structure with A 9, base narrowed, with impression at ventral side and lateral side, gradually enlarge towards rounded apex. Abdominal ventrite 2 with pair of long discal protuberances, inserted with large and strongly curved apical setae; ventrite 6 with pair of short and acute median struts, about oneninth of ventral length. Protrochanters with long ventral spine, about two-thirds of protrochanteral width; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D876FF99FF017499FC37FFD9.taxon	description	Redescription: Body length 1.52 – 1.60 mm; body colour reddish brown to dark brown with lighter coloured elytra and legs. Head wider than long; frons slightly narrowed at end, slightly longer than half width of head at eye level; eyes well developed and prominent, with about 31 facets; vertexal foveae punctiform; antennal tubercles weak; postantennal notches distinct and rounded; postantennal cavities medium-sized and wide, about half length of antennal tubercles in dorsal view; antennae 10 - segmented with 3 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 7 asymmetrical, about twice length of A 8, A 8 disc-shaped, A 9 large, transverse and cupulate, with ventral impression and basolateral excavation, with hook-shaped spine at upper margin of excavation, A 10 largest, forming claw-shaped structure with A 9, base narrowed, with impression at ventral side and lateral side, gradually enlarge towards rounded apex. Pronotum slightly longer than wide, about as wide as head at eye level; lacking lateral antebasal foveae. Elytra about as long as wide, widest near middle; humeral calli barely visible; sutural stria weakly extending from base to apical four-fifths. Abdomen with tergite 1 slightly longer than twice length of tergite 2; ventrite 2 with pair of long discal protuberances, inserted with large and strongly curved apical setae; ventrite 6 with pair of short and acute median struts, about one-ninth of ventral length. Legs with protrochanters modified with long ventral spine, about two-thirds of protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.33 mm long, with long and stout parameres, slightly bent inwards, apex curved downwards in lateral view, bearing pair of short apical setae and pair of long subapical setae; median lobe with two asymmetrical sclerites narrower than parameres, both curved inwards, left one longer than right one in dorsal view.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D876FF99FF017499FC37FFD9.taxon	discussion	Comments: The lectotype of E. (B.) costata, the holotype of E. (B.) rudicornis and the lectotype of E. (B.) monstrosa were found representing a single species by having identical forms of A 9 and A 10 (Figs. 34 B – D), with the former two names treated as junior synonyms. This species is most similar to E. (B.) conspicua by having similar modifications of A 9 and A 10, and can be separated by A 10 with a small internal hook at its apex.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D876FF99FF017499FC37FFD9.taxon	distribution	Distribution: North Island. AK, TO, RI, WI, WA. South Island. SD, NN, BR, MC, WD, FD. Additional material examined: 56 ♂♂, AK: 1, Hunua, Maketu, T. Broun, A. E. Brooks Collection; TO: 1, JNPC, Wairuhi Sd., 01 / Sep / 193, litter, J. T. Nunn collection; RI: 1, Coll. T. R. Harris, 03 / Apr / 1924, Ohakune Main Trunk, A. E. Brooks Collection; 3, Taihape, Hautapu river, 17 / Sep / 1982, J. C. Watt litter; WI: 1, JNPC, Ashurst Domain, 10 / Aug / 1996, in earthy forest litter, J. T. Nunn collection; WA: 11, JNPC, Castlehill / Haunui area, SE Puketoi Ra., Mar / 2010, J. T. Nunn collection; SD: 1, JNPC, Essens V. Picton, 01 / Feb / 1994, Nothofagus litter, J. T. Nunn collection; NN: 1, Kohaihai River Karamea, West Coast, 30 / May / 1963, G. Kuschel; 2, Upper Maitai, 13 / Feb / 1957, E. S. Gourlay; 1, Dun Mt. 2000 ft, 6 – 10 / 20, A. Philpott; 1, Nelson, The Brook, 03 / Feb / 1973, G. Kuschel; BR: 5, Barrytown, Croesus Tk., 10 / Nov / 2005, R. Leschen & S. Nomura, litter, 42 ° 14 ' S 171 ° 19 ' E; 1, JNPC, Pororari River mouth Punakaiki, 29 / Dec / 2010, in flood debris, J. T. Nunn collection; 3, LUNZ, Pahautane Bay, 05 / Sep / 1981, R. M. Emberson, litter; WD: 4, Jackson Bay, 23 / Sep / 1979, A. K. Walker, litter moss and dead wood; 2, Haast River, 100 m, Sunny flat, 25 / Jan / 1978, G. Kuschel; 2, Ross, Mikonui Tk., 09 / Nov / 2005, R. Leschen & S. Nomura, ants in wood, 43 ° 00 ' S 170 ° 51 ' E; 4, JNPC, Haast River, Pleasant Flat, 27 / Mar / 216, J. T. Nunn collection; 4, NMPC, Mount Aspiring NP., Pleasant Flat, 27.2 km, NNE of Makarora at Mur Creek, 44 ° 08 ' S 169 ° 22.9 ' E, 120 m, 11 / xii / 2012, Fikáček, Hájek & Leschen lgt., sifting of mosses on the ground and on fallen decaying logs in the Nothofagus forest and rich shrub; MC: 12, Mt. Algidus, 20 / Sep / 1913, T. Broun, A. E. Brooks Collection; 1, Riccarton Bush, Christchurch, 13 / Oct / 1951; FD: 5, Secretary I., Gut Bay, 23 / Nov / 1983, C. F. Butcher, sifted litter; 1, Dusky Sound Seal I., 07 / Mar / 1983, C. F. Butcher, sifted litter; 2, LUNZ, Grebe V., S Arm L. Manapouri, 170 m, 04 / Feb / 1982, J. W. Early, C. A. Muir, P. T. Syrett, Sweep ferns in the forest. Unknown locality: 11, 2465, T. Broun, A. E. Brooks Collection; 1, E. S. Gourlay, 30 / Mar / 1955, Ex leaf mould; Associated females (34 ♀♀): WA: 10, JNPC, Castlehill / Haunui area. SE Puketoi Ra., Mar / 2010, J. T. Nunn collection; NN: 1, Dun Mt., 2000 ft, 6 – 10 / 20, A. Philpott; BR: 3, JNPC, Pororari River mouth Punakaiki, 29 / Dec / 2010, in flood debris, J. T. Nunn collection; WD: 14, JNPC, Haast River, Pleasant Flat, 27 / Mar / 216, J. T. Nunn collection; MC: 4, Mt. Algidus, 20 / Sep / 1913, T. Broun, A. E. Brooks Collection. Unknown locality: 2, 2465, T. Broun, A. E. Brooks Collection.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D874FF98FF01742DFF6DF865.taxon	description	Fig. 35; map Fig. 53	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D874FF98FF01742DFF6DF865.taxon	materials_examined	Type locality: [CL] Tairua. Type depository: BMNH LT ♂, 4 PLT.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D874FF98FF01742DFF6DF865.taxon	diagnosis	Diagnosis: Head with vertexal foveae punctiform; antennae 10 - segmented with 3 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 large and trapezoidal, with anterolateral impression in ventral view, with minute lateral tubercle in dorsal view, A 10 largest, bean-shaped, base obliquely truncate. Abdominal ventrite 2 with pair of median struts, inserted with strongly curved short anterolateral setae; ventrite 6 with pair of short median struts, slightly shorter than one-fourth of ventral length. Protrochanters with small ventral spine, about one-third of protrochanteral width; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D874FF98FF01742DFF6DF865.taxon	description	Redescription: Body length 1.57 – 1.63 mm; body colour reddish brown with lighter coloured tibiae and tarsi. Head slightly wider than long; frons slightly narrowed at end, about half width of head at eye level; eyes well developed, with about 29 facets; vertexal foveae punctiform; antennal tubercles weak, postantennal notches small and distinct; postantennal cavities medium-sized, about half length of antennal tubercles in dorsal view; antennae 10 - segmented with 3 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 large and trapezoidal, with anterolateral impression in ventral view, with minute lateral tubercle in dorsal view, A 10 largest, bean-shaped, base obliquely truncate. Pronotum slightly wider than long, wider than head at eye level; lacking lateral antebasal foveae. Elytra about as long as wide, widest near middle; humeral calli slightly prominent; sutural stria distinctively extending from base to apex. Abdomen with tergite 1 about 1.5 × longer than tergite 2; ventrite 2 with pair of median struts, inserted with strongly curved short anterolateral setae; ventrite 6 with pair of short median struts, slightly shorter than one-fourth of ventral length. Legs with protrochanters modified with small ventral spine, about one-third of protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.36 mm long, with parameres stout and wide, slightly sinuate, bent downwards in lateral view, apex curved towards each other, with two pairs of subapical setae, subequal in length; median lobe with two asymmetrical sclerites, strongly curved inwards, left one longer than right one in dorsal view.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D874FF98FF01742DFF6DF865.taxon	discussion	Comments: The holotype of E. (B.) forficulida was compared with the lectotype and one paralectotype of E. (B.) mundula and were found to represent a single species based on the identical forms of A 9, A 10 and ventrite 2. These names are here synonymized, with the former name treated as the junior synonym. This species is similar to E. (B.) hoarei by having similar modifications of A 9 and A 10 and can by distinguished by the protrochanter having a ventral spine, which is lacking in E. (B.) hoarei (Fig. 23 D).	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D874FF98FF01742DFF6DF865.taxon	distribution	Distribution: North Island. AK, CL, TO, BP, GB, HB, WN, WA. Additional material examined: 42 ♂♂, AK: 1, Ex. Broun Collection, Hunua Rg., Clevedon, Auckland, 239; 20, moss nr. Warkworth, 29 / Jun / 1951; CL: 1, Ex. Broun Collection, Tairua, E. coast North Is., 236; 1, Otama FP., 24 / Oct / 1993, M. C. Lariviere, litter; TO: 7, 1 km Ohakune Mtn. Rd., 19 / Nov / 1995; BP: 1, Hicks Bay, 50 m, 29 / Apr / 1993, J. S. Dugdale, litter; 1, LUNZ, Te Koau, 360 m, 26 / Oct / 1992, J. W. M. Marris; GB: 2, Te Urewera NP., Lake Waikaremoana, station 6, Apr / 2005; HB: 1, White Pine Bush, 18 / Nov / 1984, J. G. Charles, steam bank litter; WA: 2, Putangirua stm., Palliser Bay, 23 / Oct / 1995, in forest litter; WN: 1, Wellington 30 / Apr / 1950, R. Hornabrook; 4, JNPC, Greens Road, Tiritea, 02 / Jun / 2004, in rewarewa litter on stream. Associated females (3 ♀♀): AK: 2, Ex. Broun Collection, Hunua Rg., Clevedon, Auckland, 239; CL: 1, Ex. Broun Collection, Tairua, E. coast North Is., 236.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D875FF96FF0170C9FBCAFC35.taxon	description	Fig. 36; map Fig. 53	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D875FF96FF0170C9FBCAFC35.taxon	materials_examined	Type locality: [AK] Woodhill. Type depository: BMNH HT ♂.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D875FF96FF0170C9FBCAFC35.taxon	diagnosis	Diagnosis: Head with vertexal foveae small and asetose; antennae 10 - segmented with 3 - segmented antennal club; A 4, A 5, A 6, A 7, A 8 and A 10 unmodified, A 8 slightly asymmetrical, A 9 asymmetrical and broadly impressed at lateral side, with slightly extended and laminate margin in lateral view, A 10 conical, slightly longer than A 9. Abdominal ventrite 2 lacking protuberances, with two small discal tufts of setae; ventrite 6 with pair of short median struts, about one-sixth of ventral length. Protrochanters with long and acute ventral spine, about two-thirds of protrochanteral width; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D875FF96FF0170C9FBCAFC35.taxon	description	Redescription: Body length 1.39 – 1.44 mm; body colour reddish brown with lighter coloured tarsi. Head wider than long, frons not narrowed at end, about three-fourths width of head at eye level; eyes well developed and prominent, with about 26 facets; vertexal foveae small and asetose; antennal tubercles moderately developed; postantennal notches weak; postantennal cavities large, slightly shorter than antennal tubercles in dorsal view; antennae 10 - segmented with 3 - segmented antennal club; A 4, A 5, A 6, A 7, A 8 and A 10 unmodified, A 8 slightly asymmetrical, A 9 asymmetrical and broadly impressed at lateral side, with slightly extended and laminate margin in lateral view, A 10 conical, slightly longer than A 9. Pronotum about as long as wide, slightly narrower than head at eye level; lacking lateral antebasal foveae. Elytra about as long as wide, widest at apical two-thirds; humeral calli distinct and moderately prominent; sutural stria barely visible and shallowly extending from middle to apex. Abdomen with tergite 1 about 1.2 × longer than tergite 2; ventrite 2 lacking protuberances, with two small discal tufts of setae; ventrite 6 with pair of short median struts, about one-sixth of ventral length. Legs with protrochanters modified with long and acute ventral spine, about two-thirds of protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.22 mm long, with slender parameres, apex slightly curved inwards, with three pairs of subapical long setae and two pairs of long apical setae; median lobe with two helix-shaped sclerites, gradually narrowed towards apex, slightly wider than parameres in dorsal view.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D875FF96FF0170C9FBCAFC35.taxon	discussion	Comments: This is the only species within the subgenus Byraxis that lacks the protuberance on andominal ventrite 2 (Fig. 36 F), the only other species in New Zealand lacking this character is E. novem, sp. n., which has a 9 - segmented antenna.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D875FF96FF0170C9FBCAFC35.taxon	distribution	Distribution: North Island. ND, AK, CL. Additional material examined: 11 ♂♂, ND: 3, Kauri Sanct., Omahuta, 22 / Apt / 1974, G. Kuschel litter; 2, Unuwhao, 270 m, 25 / Nov / 1982, G. Kuschel, sifted litter and decayed wood; AK: 3, Noises Is., Motuhoropapa I., 14 / Dec / 1978, litter; CL: 2, AMNZ, Great Barrier I., Little Windy Hill, 220 m, 21 / Feb – 26 / Mar / 2002, P Sutton. forest edge, Malaise trap; 1, AMNZ, Ohinau I., 05 / Jul / 2000, C. McGuiness, Mahoe forest. Associated females (10 ♀♀): ND: 4, Kauri Sanct., Omahuta, 22 / Apt / 1974, G. Kuschel, litter; 4, Unuwhao, 270 m, 25 / Nov / 1982, G. Kuschel, sifted litter and decayed wood; AK: 2, Noises Is., Motuhoropapa I., 14 / Dec / 1978, litter.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D87BFF96FF017499FCE8FDB1.taxon	materials_examined	Type locality: [CL] Great Barrier Island. Type depository: BMNH LT ♀, 2 PLT.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D87BFF96FF017499FCE8FDB1.taxon	discussion	Comments: This species is based solely on female types.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D87BFF96FF017205FAF3F98D.taxon	description	Fig. 37; map Fig. 53	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D87BFF96FF017205FAF3F98D.taxon	diagnosis	Diagnosis: Head with vertexal foveae punctiform; antennae 9 - segmented with one-segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 widest at middle, base strongly concave at inner side, slightly impressed near basal two-fifths in lateral view. Abdominal ventrite 2 unmodified; ventrite 6 unknown. Protrochanters with long ventral seta, about twice of protrochanteral width, lacking ventral spine; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D87BFF96FF017205FAF3F98D.taxon	description	Description: Body length 1.33 mm; body colour dark brown with lighter coloured legs. Head wider than long; frons not narrowed at end, about three-fourths width of head at eye level; eyes moderately developed, with about 16 facets; vertexal foveae punctiform; antennal tubercles moderately developed; postantennal notches distinct and broad; postantennal cavities large, about as long as antennal tubercles in dorsal view; antennae 9 - segmented with single segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 widest at middle, base strongly concave at inner side, slightly impressed near basal two-fifths in lateral view. Pronotum slightly longer than wide, slightly narrower than head at eye level; lacking lateral antebasal foveae. Elytra as long as wide, widest near apical two-thirds; lacking humeral calli; sutural stria barely visible in dorsal view. Abdomen with tergite 1 about 1.3 × longer than tergite 2; ventrite 2 unmodified; ventrite 6 unknown. Legs with protrochanters modified with long ventral seta, about twice of protrochanteral width, lacking ventral spine; protibiae and mesotibiae unmodified. Aedeagus unknown.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D87BFF96FF017205FAF3F98D.taxon	discussion	Comments: This species has an anomalous 9 - segmented antennae (Figs. 37 B – C), which separates it from all other Eupines species. We did not dissect the singleton of this new species nor establish a subgenus for it until we can find the evidence that the two existing subgenera are monophyletic.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D87BFF96FF017205FAF3F98D.taxon	distribution	Distribution: North Island. ND.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D87BFF96FF017205FAF3F98D.taxon	etymology	Etymology: This specific epithet means ‘ nine’ in Latin, referring to the nine-segmented antennae of this species.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D87BFF96FF017205FAF3F98D.taxon	materials_examined	Type material (NZAC): Holotype. ♂, specimen entire, card-mounted: “ NEW ZEALAND, ND, Pukenui For- est, Taraire Ridge Loop, 12 Nov 2018, J. Shen // sifted leaf litter & rotten woods, 35 ° 41 ' 96 '' S, 174 ° 15 ' 90 '' E ”.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D878FF94FF017398FEBBF9CB.taxon	description	Fig. 38; map Fig. 53	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D878FF94FF017398FEBBF9CB.taxon	diagnosis	Diagnosis: Head lacking vertexal foveae; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 6, A 7, A 8 and A 10 unmodified, A 5 greatly enlarged and asymmetrical, about 3.5 × longer than A 4, A 7 slightly asymmetrical, A 8 about half length of A 7, A 9 large, about as long as A 10, base slightly impressed at dorsal side, widest near apex, A 10 conical. Abdominal ventrite 2 with pair of large discal protuberances, inserted with strongly curved small apical setae; ventrite 6 with pair of long median struts, about one-third of ventral length. Protrochanters with long and acute ventral spine, about three-fifths of protrochanteral width; protibiae with large and blunt apical protuberance; mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D878FF94FF017398FEBBF9CB.taxon	description	Description: Body length 1.42 – 1.50 mm; body colour dark brown to black with lighter coloured legs. Head wider than long; frons slightly narrowed at end, about half width of head at eye level; eyes moderately developed, with about 23 facets; lacking vertexal foveae; antennal tubercles barely visible; lacking postantennal notches; postantennal cavities long and wide, slightly longer than antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 6, A 7, A 8 and A 10 unmodified, A 5 greatly enlarged and asymmetrical, about 3.5 × longer than A 4, A 7 slightly asymmetrical, A 8 about half length of A 7, A 9 large, about as long as A 10, base slightly impressed at dorsal side, widest near apex, A 10 conical. Pronotum as long as wide, slightly wider than head at eye level; lacking lateral antebasal foveae. Elytra slightly longer than wide, widest near middle; humeral calli slightly prominent; sutural stria shallowly extending from base to apex. Abdomen with tergite 1 about 1.2 × longer than tergite 2; ventrite 2 with pair of large discal protuberances, inserted with strongly curved small apical setae; ventrite 6 with pair of long median struts, about one-third of ventral length. Legs with protrochanters modified with long and acute ventral spine, about three-fifths of protrochanteral width; protibiae with large and blunt apical protuberance; mesotibiae unmodified. Aedeagus about 0.33 mm long, with long and stout parameres nearly meet at apex, bent downwards in lateral view, with two pairs of long subapical setae and pair of short apical setae; median lobe with two slender sclerites, slightly sinuate, strongly curved towards each other at apex, much narrower than parameres.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D878FF94FF017398FEBBF9CB.taxon	discussion	Comments: This species can be distinguished from all other species by its protibiae modified with rounded apical protuberance, along with it modified A 5 (Figs. 38 B – E).	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D878FF94FF017398FEBBF9CB.taxon	distribution	Distribution: North Island. TO, TK. South Island. WD.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D878FF94FF017398FEBBF9CB.taxon	etymology	Etymology: This specific epithet is a Latin adjective meaning ‘ blunt’, referring to the blunt apical protuberance on protibiae of this species.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D878FF94FF017398FEBBF9CB.taxon	materials_examined	Type material (NZAC): Holotype. ♂, specimen entire, point-mounted: “ NEW ZEALAND, WD, Mt. Tuhua, 1067 m, E. side of L. Kaniere, 20 Nov 1984 // B. A. Holloway Moss and lichen 84 / 75 ”. Paratypes. 7 ex.: TO: 1, Whanganui V., 792 m, Whenuakura Flats, 11 / Oct / 1979, J. S. Dugdale // bryophytes, 79 / 94; TK: 3, Pouakai Hump, 1220 m, 03 / Dec / 1975, A. K. Walker // litter moss, 75 / 211; 1, W. edge Hump Plat. Pouakai Ra., 01 / Dec / 1975, J. S. Dugdale // litter, 75 / 200; 1, Holly Hut, 950 m, North Egmont, Nov / 1975, A. K. Walker // beating; WD: 1, same data as holotype.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D87EFF92FF017691FD11FF81.taxon	description	Fig. 39; map Fig. 53	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D87EFF92FF017691FD11FF81.taxon	diagnosis	Diagnosis: Head with vertexal foveae punctiform; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 6, A 7, A 8 and A 10 unmodified, A 4 asymmetrical, A 5 largest and ovoid, dorsal side broadly impressed, A 9 cylindrical and elongate, slightly asymmetrical at apex, A 10 conical, about 1.5 × longer than A 9. Abdominal ventrite 2 with pair of flat and wide discal protuberances, with strongly curved large apical setae; ventrite 6 with pair of short median struts, about one-fourth of ventral length. Protrochanters with short ventral spine, about one-seventh of protrochanteral width; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D87EFF92FF017691FD11FF81.taxon	description	Description: Body length 1.50 mm; body colour dark brown with lighter coloured elytra and tarsi. Head as long as wide; frons slightly narrowed at end, about half length of head at eye level; eyes moderately developed, slightly prominent in dorsal view, with about 31 facets; vertexal foveae punctiform; antennal tubercles weak; lacking postantennal notches; postantennal cavities large, slightly longer than antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 6, A 7, A 8 and A 10 unmodified, A 4 asymmetrical, A 5 largest and ovoid, dorsal side broadly impressed, A 9 cylindrical and elongate, slightly asymmetrical at apex, A 10 conical, about 1.5 × longer than A 9. Pronotum slightly longer than wide, wider than head at eye level, with punctiform lateral antebasal foveae. Elytra about as long as wide, widest near middle; humeral calli barely visible in dorsal view; sutural stria shallowly extending from base to apex. Abdomen with tergite 1 about 1.5 × longer than tergite 2; ventrite 2 with pair of flat and wide discal protuberances, with strongly curved large apical setae; ventrite 6 with pair of short median struts, about one-fourth of ventral length. Legs with protrochanters modified with short ventral spine, about one-seventh of protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.27 mm long, with wide and stout parameres, slightly sinuate, bent downwards in lateral view, apex slightly curved, with pair of long subapical setae; median lobe with two large sclerites slightly entwined at apex, widest near middle, much narrower than parameres.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D87EFF92FF017691FD11FF81.taxon	discussion	Comments: This species can be distinguished from all other species by its A 5 large and ovoid (Figs. 39 B – D).	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D87EFF92FF017691FD11FF81.taxon	distribution	Distribution: South Island. BR.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D87EFF92FF017691FD11FF81.taxon	etymology	Etymology: The specific epithet refers to the oval-shaped antennomere 5.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D87EFF92FF017691FD11FF81.taxon	materials_examined	Type material (NZAC): Holotype. ♂, specimen entire, glued on card, aedeagus + one antenna mounted on acetate card: “ NEW ZEALAND, BR, Sewell Peak, 15 Jan 2014, T. Buckley, M. Gimmel, R. Leschen // ant nests under rock, 810 m, 42 ° 24.497 ' S, 171 ° 20.457 ' E ”.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D87FFF91FF0177D5FD0AF849.taxon	description	Fig. 40; map Fig. 53	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D87FFF91FF0177D5FD0AF849.taxon	materials_examined	Type locality: [ND] near Whangarei Harbour. Type depository: BMNH HT ♂.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D87FFF91FF0177D5FD0AF849.taxon	diagnosis	Diagnosis: Head with vertexal foveae punctiform; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 7 and A 8 unmodified, A 6 wide and asymmetrical, much wider than A 7, A 9 asymmetrical, gradually narrow towards apex at outer side, broadly impressed from middle to apex in ventral view, A 10 conical, broadly impressed at base in ventral view and slightly longer than A 9. Abdominal ventrite 2 with pair of large and asymmetrical discal protuberances, inserted with strongly curved medium-sized apical setae; ventrite 6 large and stout, with lower median margin convex, with pair of long and stout median struts, about one-third of ventral length. Protrochanters with medium-sized ventral spine, slightly less than two-thirds of protrochanteral width; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D87FFF91FF0177D5FD0AF849.taxon	description	Redescription: Body length 1.52 – 1.59 mm; body colour reddish brown to dark brown with lighter coloured elytra, tibiae and tarsi. Head wider than long; frons slightly narrowed at end, slightly longer than half width of head at eye level; eyes slightly prominent, with about 30 facets; vertexal foveae punctiform; antennal tubercles barely visible; postantennal notches barely visible; postantennal cavities medium-sized, about two-thirds length of antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 7 and A 8 unmodified, A 6 wide and asymmetrical, much wider than A 7, A 9 asymmetrical, gradually narrow towards apex at outer side, broadly impressed from middle to apex in ventral view, A 10 conical, broadly impressed at base in ventral view and slightly longer than A 9. Pronotum slightly wider than long, about as long as head at eye level; lacking lateral antebasal foveae. Elytra about as long as wide, widest at apical two-thirds; humeral calli barely visible, sutural stria shallowly extending from base to apex. Abdomen with tergite 1 about as long as tergite 2; ventrite 2 with pair of large and asymmetrical discal protuberances, inserted with strongly curved medium-sized apical setae; ventrite 6 large and stout, with lower median margin convex, with pair of long and stout median struts, about one-third of ventral length. Legs with protrochanters modified with medium-sized ventral spine, slightly less than two-thirds of protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.44 mm long, with long and stout parameres, slightly curved towards each other at apex, strongly bent downwards in lateral view, with four pairs of short subapical setae; median lobe with two long sclerites, narrower than parameres in dorsal view, strongly curved inwards at apex, left one sinuate near middle.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D87FFF91FF0177D5FD0AF849.taxon	discussion	Comments: We examined types of E. (B.) allocera, E. (B.) sylvicola, E. (B.) rhyssarthra and E. (B.) paganus, and determined that these species are conspecific based on their identical antennal modifications and form of ventrite 2. These names are here synonymized, with the former three names treated as junior synonyms. This species can be distinguished from all other Eupines by A 6 transverse and asymmetrical (Figs. 40 B – D).	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D87FFF91FF0177D5FD0AF849.taxon	distribution	Distribution: North Island. ND, AK, CL, BP, RI. Additional material examined: 13 ♂♂, ND: 1, Omahuta SF., 04 / Feb / 1975, G. Kuschel; AK: 3, Ex. Broun Collection, near Howick, Auckland, 1645, T. Broun Collection, A. E. Brookes Collection; 1, Ex. Broun Collection, Hunua Rg., Maketu, Auckland, 1645; CL: 3, AMNZ, Great Barrier I., Mt. Hobson, 500 m, pitfall trap, 09 / Nov / – 17 / Dec / 2003, J. W. Early; BP: 3, Rurima Rocks, 21 / May / 1986, I. Mcfadden, litter; RI: 2, Taihape Hautapu River, 17 / Sep / 1982, J. C. Watt, litter. Associated females (12 ♀♀): AK: 9, Ex. Broun Collection, Hunua Rg., Maketu, Auckland, 1645. Unknown locality: 3, 1645, Broun.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D87CFFEFFF01709CFB85FF6D.taxon	description	Fig. 41; map Fig. 54	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D87CFFEFFF01709CFB85FF6D.taxon	diagnosis	Diagnosis: Head with vertexal foveae small and asetose; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 enlarged, about 3.2 × longer than A 8, ventral side truncate near apex, A 10 conical, with basal impression at ventral view. Abdominal ventrite 2 with pair of medium-sized discal protuberances, inserted with strongly curved small apical setae; ventrite 6 with pair of long and stout median struts, about one-fourth of ventral length. Protrochanters with short ventral spine, about one-third of protrochanteral width; protibiae unmodified; mesotibiae slightly convex near apex, with long tuft of subapical setae.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D87CFFEFFF01709CFB85FF6D.taxon	description	Description: Body length 1.52 – 1.62 mm; body colour reddish brown with lighter coloured legs. Head slightly wider than long; frons not narrowed at end, about three-fourths width of head at eye level; eyes well developed, conspicuously prominent in dorsal view, with about 31 facets; vertexal foveae small and asetose; antennal tubercles moderately developed; postantennal notches broad and long; postantennal cavities large and long, about as long as antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 enlarged, about 3.2 × longer than A 8, ventral side truncate near apex, A 10 conical, with basal impression at ventral view. Pronotum as long as wide, about as wide as head at eye level; lacking lateral antebasal foveae. Elytra longer than wide, widest near middle; humeral calli weak, slightly prominent in dorsal view; sutural stria shallowly extending from base to middle, barely visible from middle to apex. Abdomen with tergite 1 about as long as tergite 2; ventrite 2 with pair of medium-sized discal protuberances, inserted with strongly curved small apical setae; ventrite 6 with pair of long and stout median struts, about one-fourth of ventral length. Legs with protrochanters modified with short ventral spine, about one-third of protrochanteral width; protibiae unmodified; mesotibiae slightly convex near apex, with long tuft of subapical setae. Aedeagus about 0.34 mm long, with long and stout parameres, meet at apex, widest near middle, curved downwards in lateral view, with four pairs of subapical short setae; median lobe with two slender sclerites, slightly sinuate and meet at apex.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D87CFFEFFF01709CFB85FF6D.taxon	discussion	Comments: This species can be distinguished from all other Eupines by its mesotibiae modified with a long tuft of subapical setae (Fig. 41 E).	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D87CFFEFFF01709CFB85FF6D.taxon	distribution	Distribution: North Island. ND.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D87CFFEFFF01709CFB85FF6D.taxon	etymology	Etymology: This specific epithet means ‘ tuft’ in Latin, referring to the long tuft of subapical setae on mesotibiae.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D87CFFEFFF01709CFB85FF6D.taxon	materials_examined	Type material (NZAC): Holotype. ♂, specimen entire, card-mounted: “ NEW ZEALAND, ND, Mt. Horo- kaka, 518 m, Tangihua Ra., 16 Aug 1977, J. S. Dugdale // moss and liverworts 77 / 96 ”. Paratypes. 7 ex.: ND: 1, same data as holotype; 1, Waipoua SF., Lookout area, 29 / Oct / 1980, G. Kuschel // sifted moss, 80 / 92; 2, FMNH, Waipoua For., Te Matua Ngahere, 16 – 20 / iii / 78, 400 m, S & J. Peck, Kauri for. // Berlese moss; 2, FMNH, Waipoua SF., Yakas Tree Tr., 350 m, 11 – 14 / iv / 1980, broadlf. - podocarp A. Newton & M. Thayer, 710 // berl, litter at bases of Metrosideros robusta trunks; 1, FMNH, Omahuta For., Picnic Area near Kauri Sanctuary, 330 m, 29 / xi – 5 / xii / 1984, hdwd. - podocarp for. A. Newton / M. Thayer, 694 // flight intercept (window) trap.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D802FFEEFF0177B1FE81FDFF.taxon	description	Fig. 42; map Fig. 54	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D802FFEEFF0177B1FE81FDFF.taxon	diagnosis	Diagnosis: Head with vertexal foveae small and asetose; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 elongate, about as long as A 10, truncate at dorsal side, apical margin extended in lateral view, A 10 conical and truncate at dorsal side. Abdominal ventrite 2 with pair of small discal protuberances, inserted with slightly curved small apical setae; ventrite 6 with pair of short median struts, about one-fifth of ventral length. Protrochanters with a long and acute ventral spine, about three-fourths of protrochanteral width; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D802FFEEFF0177B1FE81FDFF.taxon	description	Description: Body length 1.30 – 1.40 mm; body colour reddish brown with lighter coloured elytra and legs. Head longer than wide; frons not narrowed at end, about three-fourths width of head at eye level; eyes moderately developed, with about 19 facets; vertexal foveae small and asetose; antennal tubercles conspicuously developed; postantennal notches broad and long; postantennal cavities large, slightly shorter than antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 elongate, about as long as A 10, truncate at dorsal side, apical margin extended in lateral view, A 10 conical and truncate at dorsal side. Pronotum about as long as wide, wider than head at eye level; lacking lateral antebasal foveae. Elytra about as long as wide, widest at apical two-thirds; humeral calli weak, slightly prominent in dorsal view; sutural stria weak, visible near base and apex. Abdomen with tergite 1 about 1.3 × longer than tergite 2; ventrite 2 with pair of small discal protuberances, inserted with slightly curved small apical setae; ventrite 6 with pair of short median struts, about one-fifth of ventral length. Legs with protrochanters modified with a long and acute ventral spine, about three-fourths of protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.35 mm long, with strongly elongated parameres, slightly sinuate, curved inwards at apex, with three pairs of long subapical setae and two pairs of long apical setae; median lobe with two long and slender sclerites, curved towards each other at apex, widest near base, shorter than parameres.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D802FFEEFF0177B1FE81FDFF.taxon	discussion	Comments: This species can be distinguished from all other Eupines by A 10 dorsally conical and truncate (Figs. 42 B – D).	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D802FFEEFF0177B1FE81FDFF.taxon	distribution	Distribution: North Island. ND, CL, BP.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D802FFEEFF0177B1FE81FDFF.taxon	etymology	Etymology: This specific epithet is derived from a Latin adjective meaning ‘ slender’, referring to the slender parameres of aedeagus of this species.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D802FFEEFF0177B1FE81FDFF.taxon	materials_examined	Type material (NZAC): Holotype. ♂, specimen entire, point-mounted: “ NEW ZEALAND, ND, Mangamuka Smt. 386 m, 13 Dec 1976, V. A. May, Litter 76 / 109 ”. Paratypes. 5 ex.: CL: 1, NEW ZEALAND, Mt. Maumaupaki, 22 / Oct / 1976, J. S. Dugdale // litter, 76 / 86; WO: 1, JNPC, Mahaukura Tck., Mt. Pirongia, 18 / Nov / 05 // J. T. Nunn Collection; TO: 1, LSAM, Mt. Ruapehu south slope, FIT site, Berlese, 25 / Mar / 2000 C. Carlton, A. Weir, # 093; BP: 1, NEW ZEALAND, Tikitiki Stream, Horohoro SF., Mamaku Plateau, 24 / Jul / 1976 // J. S. Dugdale Liverworts, mosses, litter, 76 / 58; GB: 1, LSAM, Urewera N. P., Lake Ruapane Tr. Fungusy wood, # 067, 20 / Mar / 2000, C. Carlton, A. Weir.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D803FFEEFF01724FFCE8FB78.taxon	materials_examined	Type locality: [CL] Tairua. Type depository: BMNH LT ♀, 1 PLT.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D803FFEEFF01724FFCE8FB78.taxon	discussion	Comments: This species is based solely on female types.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D803FFEDFF0173CBFD94F9AC.taxon	description	Fig. 43; map Fig. 54	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D803FFEDFF0173CBFD94F9AC.taxon	materials_examined	Type locality: [CL] Tairua. Type depository: BMNH LT ♂; NZAC 1 PLT.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D803FFEDFF0173CBFD94F9AC.taxon	diagnosis	Diagnosis: Head with vertexal foveae punctiform; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 largest and strongly transverse, with deep lateral impression and laminate margin at inner side, A 10 conical and slightly shorter than A 9, lateral side broadly impressed. Abdominal ventrite 2 with pair of minute discal protuberances, inserted with small strongly curved apical setae; ventrite 6 with pair of long median struts, about as long as ventral length. Protrochanters modified with medium-sized ventral spine, about half of protrochanteral width; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D803FFEDFF0173CBFD94F9AC.taxon	description	Redescription: Body length 1.60 mm; body colour reddish brown with lighter coloured tibiae and tarsi. Head wider than long; frons not narrowed at end, about three-fourths width of head at eye level; eyes slightly prominent, with about 29 facets; vertexal foveae punctiform; antennal tubercles weak; postantennal notches weak; postantennal cavities small, about half width of antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 largest and strongly transverse, with deep lateral impression and laminate margin at inner side, A 10 conical and slightly shorter than A 9, lateral side broadly impressed. Pronotum slightly longer than wide, about as wide as head at eye level; lacking lateral antebasal foveae. Elytra slightly longer than wide, widest at apical two-thirds; humeral calli slightly prominent; sutural stria shallowly extending from base to apex. Abdomen with tergite 1 about 1.5 × longer than tergite 2; ventrite 2 with pair of minute discal protuberances, inserted with small strongly curved apical setae; ventrite 6 with pair of long median struts, about as long as ventral length. Legs with protrochanters modified with medium-sized ventral spine, about half of protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.29 mm long, with elongate parameres slightly sinuate, longer than median lobe and basal bulb combined, slightly curved downwards at apex in lateral view, with two pairs of stout subapical setae at inner side, pair of slender subapical setae at outer side and pair of short apical setae; median lobe with two sclerites highly twisted, slightly asymmetrical, gradually narrowed towards apex, both curved inwards at apex.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D803FFEDFF0173CBFD94F9AC.taxon	discussion	Comments: This species can be distinguished from all other Eupines by A 9 strongly transverse (Figs. 43 B – D). Distribution: North Island. CL.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D801FFEBFF017691FBF0FC11.taxon	description	Fig. 44; map Fig. 54	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D801FFEBFF017691FBF0FC11.taxon	diagnosis	Diagnosis: Head lacking vertexal foveae; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 6, A 7 and A 8 unmodified, A 5 greatly enlarged and asymmetrical, about 2.5 × longer than A 4, A 8 short, about half length of A 7, A 9 largest, about 1.4 × longer than A 10, slightly impressed at base in dorsal view, widest near apex, A 10 beanshaped, slightly impressed near base in lateral side. Abdominal ventrite 2 with pair of long discal protuberances curved towards middle, inserted with strongly curved small apical setae; ventrite 6 with pair of long median struts, about half of ventral length. Protrochanters with long and acute ventral spine, about two-thirds of protrochanteral width; protibiae with large triangle apical protuberance and apical long spine; mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D801FFEBFF017691FBF0FC11.taxon	description	Description: Body length 1.59 – 1.63 mm; body colour black with lighter coloured tibiae and tarsi. Head wider than long; frons slightly narrowed at end, about half width of head at eye level; eyes weakly developed, slightly prominent in dorsal view, with about 23 facets; lacking vertexal foveae; antennal tubercles barely visible; lacking postantennal notches; postantennal cavities long and wide, slightly shorter than antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 6, A 7 and A 8 unmodified, A 5 greatly enlarged and asymmetrical, about 2.5 × longer than A 4, A 8 short, about half length of A 7, A 9 largest, about 1.4 × longer than A 10, slightly impressed at base in dorsal view, widest near apex, A 10 bean-shaped, slightly impressed near base in lateral side. Pronotum as long as wide, wider than head at eye level; lacking lateral antebasal foveae. Elytra about as long as wide, widest near middle; humeral calli weak; sutural stria shallowly extending from base to apex. Abdomen with tergite 1 about 1.5 × longer than tergite 2; ventrite 2 with pair of long discal protuberances curved towards middle, inserted with strongly curved small apical setae; ventrite 6 with pair of long median struts, about half of ventral length. Legs with protrochanters modified with long and acute ventral spine, about two-thirds of pro- trochanteral width; protibiae with large triangle apical protuberance and apical long spine; mesotibiae unmodified. Aedeagus about 0.29 mm long, with long and stout parameres curved towards each other, almost meet at apex, bent downwards in lateral view, with four pairs of small subapical setae; median lobe with two short sclerites, strongly curved at apex, much narrower than parameres.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D801FFEBFF017691FBF0FC11.taxon	discussion	Comments: This species is similar to E. (B.) dugdalei by sharing with it the dark body colour and the similar antennal modification and the aedeagal structures but differs from it by having A 9 subparallel-sided (Fig. 44 B).	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D801FFEBFF017691FBF0FC11.taxon	distribution	Distribution: South Island. SL, SI.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D801FFEBFF017691FBF0FC11.taxon	etymology	Etymology: This specific epithet refers to the protibiae of this species with long apical spine.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D801FFEBFF017691FBF0FC11.taxon	materials_examined	Type material: Holotype (NZAC). ♂, body parts mounted on acetate card: “ NEW ZEALAND, SI, Stewart I., Codfish I., North Hut, 6 Dec 1981 // B. A. Holloway, Mat plants 81 / 208 ”. Paratype (JNPC). 1 ex.: SL: Spence Burn Basin, 1150 m, Takitimu Mtns, 17 / Jan / 2000 // in tussock litter // J. T. Nunn collection.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D806FFEBFF01746AFBE8F9F9.taxon	description	Fig. 45; map Fig. 54	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D806FFEBFF01746AFBE8F9F9.taxon	materials_examined	Type locality: [WO] Mount Pirongia. Type depository: BMNH LT ♂, 1 PLT.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D806FFEBFF01746AFBE8F9F9.taxon	diagnosis	Diagnosis: Head with vertexal foveae small and asetose; antennae 10 - segmented with 4 - segmented antennal club; A 4, A 5, A 6, A 7, A 8 and A 10 unmodified, A 8 slightly asymmetrical, A 9 strongly asymmetrical and concaved at lateral side, margin of concavity laminate and extended in ventral view, A 10 conical and slightly impressed at base in ventral view. Abdominal ventrite 2 with pair of barely visible discal protuberances, inserted with slightly curved apical setae; ventrite 6 with pair of short median struts, about one-sixth of ventral length. Protrochanters with long and acute ventral spine, about two-thirds of protrochanteral width; protibiae with minute apical spine; mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D806FFEBFF01746AFBE8F9F9.taxon	description	Redescription: Body length 1.21 – 1.30 mm; body colour reddish brown to with lighter coloured tibiae and tarsi. Head wider than long; frons not narrowed at end, about three-fourths width of head at eye level; eyes well developed, with about 25 facets; vertexal foveae small and asetose; antennal tubercles moderately developed; postantennal notches distinct and broad; postantennal cavities large, about two-thirds length of antennal tubercles in dorsal view; antennae 10 - segmented with 4 - segmented antennal club; A 4, A 5, A 6, A 7, A 8 and A 10 unmodified, A 8 slightly asymmetrical, A 9 strongly asymmetrical and concaved at lateral side, margin of concavity laminate and extended in ventral view, A 10 conical and slightly impressed at base in ventral view. Pronotum as long as wide, slightly narrower than head at eye level; with punctiform lateral antebasal foveae. Elytra slightly wider than long, widest at apical two-thirds; humeral calli weak, slightly prominent; sutural stria weak, extending from basal onethird to apex. Abdomen with tergite 1 about 1.2 × longer than tergite 2; ventrite 2 with pair of barely visible discal protuberances, inserted with slightly curved apical setae; ventrite 6 with pair of short median struts, about one-sixth of ventral length. Legs with protrochanters modified with long and acute ventral spine, about two-thirds of protrochanteral width; protibiae with minute apical spine; mesotibiae unmodified. Aedeagus about 0.24 mm long, with slender parameres, slightly curved inwards at apex, slightly sinuate in dorsal view, with three pairs of long subapical setae and two pairs of long apical setae; median lobe with two short sclerites, strongly curved towards each other at apex, slightly narrower than parameres.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D806FFEBFF01746AFBE8F9F9.taxon	discussion	Comments: This species is similar to E. (B.) nemoralis by having a similar antennal modification but can be distinguished from it by its protibiae with minute apical spine (Fig. 45 F), which is lacking in E. (B.) nemoralis.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D806FFEBFF01746AFBE8F9F9.taxon	distribution	Distribution: North Island. ND, AK, WO, CL, RI. Additional material examined: 10 ♂♂, ND: 1, Paihia Opua SF., 22 / Jan / 1981, G. Kuschel; 2, Waipoua SF., Toronui Track, 30 / Oct / 1980, G. Kuschel; AK: 1, Bethells Matuku Res., 21 / Oct – 25 / Nov / 1983, G. Hall and P. A. Maddison, pit trap; 1, Waitakere Drive, 21 / Dec / 1944; CL: 2, Little Barrier I., 07 / Apr / 1984, C. T. Duval, sifted litter; 2, AMNZ, Great Barrier I., Little Windy Hill, 200 m, 21 / Nov – 13 / Dec / 2002; RI: 1, NE Ohingaiti Mangamako Rd., Paki-iti Fam, 23 / Aug / 1985, C. T. Duval, sifted litter. Associated females (16 ♀♀): ND: 3, Paihia Opua SF., 22 / Jan / 1981, G. Kuschel; 13, Waipoua SF., Toronui Track, 30 / Oct / 1980, G. Kuschel.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D807FFEAFF017390FCE8F8C8.taxon	materials_examined	Type locality: [CL] Tairua. Type depository: BMNH LT ♀, 1 PLT.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D807FFEAFF017390FCE8F8C8.taxon	discussion	Comments: This species is based solely on female types.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D807FFE8FF01711CFCEFFF6D.taxon	description	Fig. 46; map Fig. 54	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D807FFE8FF01711CFCEFFF6D.taxon	diagnosis	Diagnosis: Head with vertexal foveae punctiform; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7, A 8 and A 10 unmodified, A 8 slightly shorter than A 7, A 9 reversed cone-shaped, slightly asymmetrical, with apical spine at ventral side, A 10 conical and slightly longer than A 9. Abdominal ventrite 2 with pair of min- ute discal protuberances, inserted with strongly curved minute apical setae; ventrite 6 with pair of long and acute median struts, about half of ventral length. Protrochanters with long and acute ventral spine, about three-fourths of protrochanteral width; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D807FFE8FF01711CFCEFFF6D.taxon	description	Description: Body length 1.91 mm; body colour reddish brown with lighter coloured tibiae and tarsi. Head longer than wide; frons not narrowed at end, about three-fourths width of head at eye level; eyes moderately developed, slightly prominent in dorsal view, with about 27 facets; vertexal foveae punctiform; antennal tubercles weakly developed; lacking postantennal notches; postantennal cavities large, slightly shorter than antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7, A 8 and A 10 unmodified, A 8 slightly shorter than A 7, A 9 reversed cone-shaped, slightly asymmetrical, with apical spine at ventral side, A 10 conical and slightly longer than A 9. Pronotum longer than wide, about as wide as head at eye level; lacking lateral antebasal foveae. Elytra slightly longer than wide, widest near middle; humeral calli weak, slightly prominent in dorsal view; sutural stria barely visible. Abdomen with tergite 1 about 1.3 × longer than tergite 2; ventrite 2 with pair of minute discal protuberances, inserted with strongly curved minute apical setae; ventrite 6 with pair of long and acute median struts, about half of ventral length. Legs with protrochanters modified with long and acute ventral spine, about three-fourths of protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.36 mm long, with slender parameres curved towards each other at apex, apex bent downwards in lateral view, with five pairs of long apical setae; median lobe with two large sclerites, curved towards each other at apex, widest near middle, right one longer and much wider than left one in dorsal view.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D807FFE8FF01711CFCEFFF6D.taxon	discussion	Comments: This species is most similar to E. (B.) carinata by sharing the large body size (average length around 2.00 mm) but can be separated by having A 10 unmodified (Fig. 46 B). Distribution: North Island. GB. Etymology: This species is named after the holotype locality. Type material (NZAC): Holotype. ♂, specimen entire, glued on point, aedeagus + proleg on acetate card: “ GB, L. Waikaremoana, 19 Nov 1975, G. Kuschel [in hand] ”.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D805FFE7FF0177B1FD02FD54.taxon	description	Fig. 47; map Fig. 54	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D805FFE7FF0177B1FD02FD54.taxon	diagnosis	Diagnosis: Head with vertexal foveae punctiform; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 cylindrical and largest, about 1.5 × longer than A 10, with apical small impression surrounding small spine at ventral side, A 10 conical and slightly flattened at ventral side near base. Abdominal ventrite 2 with pair of minute discal protuberances, inserted with pair of slightly curved minute apical setae; ventrite 6 unknown. Protrochanters with short ventral spine, about two-fifths of protrochanteral width; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D805FFE7FF0177B1FD02FD54.taxon	description	Description: Body length 1.52 mm; body colour dark brown with lighter coloured tibiae and tarsi. Head wider than long; frons not narrowed at end, about three-fourths width of head at eye level; eyes well developed, conspicuously prominent in dorsal view, with about 32 facets; vertexal foveae punctiform; antennal tubercles weak; postantennal notches shallow; postantennal cavities large, slightly shorter than antennal tubercles in dorsal view; antennae 10 - segmented with 2 - segmented antennal club; A 4, A 5, A 6, A 7 and A 8 unmodified, A 9 cylindrical and largest, about 1.5 × longer than A 10, with apical small impression surrounding small spine at ventral side, A 10 conical and slightly flattened at ventral side near base. Pronotum about as long as wide, slightly wider than head at eye level; lacking lateral antebasal foveae. Elytra slightly wider than long, widest near middle; humeral calli barely visible; sutural stria barely visible near base and apex, distinctive near middle. Abdomen with tergite 1 about twice length of tergite 2; ventrite 2 with pair of minute discal protuberances, inserted with pair of slightly curved minute apical setae; ventrite 6 unknown. Legs with protrochanters modified with short ventral spine, about two-fifths of protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.33 mm long, with slender parameres asymmetrical, curved towards each other and nearly meet at apex, with two pairs of long subapical setae and three pairs of long apical setae; median lobe with two large and two small sclerites, large two slightly entwined near apex, wider and slightly shorter than parameres, small two straight.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D805FFE7FF0177B1FD02FD54.taxon	discussion	Comments: This species is described from a singleton that lacks ventrite 6. It is similar to E. (B.) hectori by having A 9 modified with a small apical spine but differs by having A 9 about twice the length of A 10 (Figs. 47 B – D).	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D805FFE7FF0177B1FD02FD54.taxon	distribution	Distribution: North Island. TO.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D805FFE7FF0177B1FD02FD54.taxon	etymology	Etymology: This species is named after the type locality.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D805FFE7FF0177B1FD02FD54.taxon	materials_examined	Type material (NZAC): Holotype. ♂, body parts on acetate card: “ NEW ZEALAND, TO, Whirinaki SF. 12 Sep 1984, C. Crowe & A. Davis, Litter 84 / 66 ”.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D80AFFE6FF0175B9FDA2FF15.taxon	description	Fig. 48; map Fig. 54	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D80AFFE6FF0175B9FDA2FF15.taxon	diagnosis	Diagnosis: Head with vertexal foveae punctiform, surrounded by shallow impressions; antennae 11 - segmented with 2 - segmented antennal clubs; A 4, A 5, A 6, A 7, A 8 and A 9 unmodified, A 10 trapezoidal, about twice length of A 9, dorsal side truncate, A 11 longest, spear-shaped, with basolateral excavation, basal margin truncate at dorsal side. Abdominal ventrite 2 unmodified; ventrite 6 with pair of long and acute median struts, about one-third of ventral length. Protrochanters with short ventral spine, about one-fifth of protrochanteral length; protibiae and mesotibiae unmodified.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D80AFFE6FF0175B9FDA2FF15.taxon	description	Description: Body length 1.25 – 1.33 mm; body colour reddish brown with lighter coloured tibiae and tarsi. Head wider than long; frons not narrowed at end, about three-fourths width of head at eye level; eyes moderately developed, prominent in dorsal view, with about 16 facets; vertexal foveae punctiform, surrounded by shallow impressions; antennal tubercles moderately developed; postantennal notches distinctive and broad; postantennal cavities large, slightly shorter than antennal tubercles in dorsal view; antennae 11 - segmented with 2 - segmented antennal clubs; A 4, A 5, A 6, A 7, A 8 and A 9 unmodified, A 10 trapezoidal, about twice length of A 9, dorsal side truncate, A 11 longest, spear-shaped, with basolateral excavation, basal margin truncate at dorsal side. Pronotum as long as wide, about as wide as head at eye level; lacking lateral antebasal foveae. Elytra slightly wider than long, widest near middle; lacking humeral calli and sutural stria. Abdomen with tergite 1 about 1.2 × longer than tergite 2; ventrite 2 unmodified; ventrite 6 with pair of long and acute median struts, about one-third of ventral length. Legs with protrochanters modified with short ventral spine, about one-fifth of protrochanteral length; protibiae and mesotibiae unmodified. Aedeagus about 0.22 mm long, with stout and symmetrical parameres, apex curved downwards in lateral view, with two pairs of long subapical setae and pair of long apical setae; median lobe with two large and symmetrical sclerites, curved towards each other in dorsal view, widest at base and narrower than parameres.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D80AFFE6FF0175B9FDA2FF15.taxon	discussion	Comments: This is the only New Zealand species that can be placed into the subgenus Eupines by having an 11 - segmented antenna. Ventrite 2 of E. (E.) undecim is unmodified, as it is in E. novum sp. n., while it is modified in all species of the subgenus Byraxis, apart from E. (B.) nemoralis (Fig. 36 F).	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D80AFFE6FF0175B9FDA2FF15.taxon	distribution	Distribution: North Island. ND.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D80AFFE6FF0175B9FDA2FF15.taxon	etymology	Etymology: This specific epithet means ‘ eleven’ in Latin, referring to the 11 - segmented antennae of this species.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D80AFFE6FF0175B9FDA2FF15.taxon	materials_examined	Type material (NZAC): Holotype. ♂, specimen entire, card-mounted: “ NEW ZEALAND, ND, Mangamuka Summit, 400 m, 13 Dec 1976, B. M. May litter 76 / 108 ”. Paratypes. 9 ex.: ND: 1, Mangamuka Summit, 400 m, 13 / Dec / 1976, V. A. May, litter, 76 / 106; 3, Mangamuka Summit, 400 m, 13 / Dec / 1976, V. A. May, litter, 76 / 109; 4, Mangamuka Gorge, 30 / Nov / 1907 // A. E. Brookes Collection; 1, Ngaiotonga Reserve, 03 / Nov / 1981, G. Kuschel // litter and decayed wood, 81 / 119.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D80BFFE6FF017098FBD1F99C.taxon	materials_examined	Type locality: [WO] Rangiriri.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D80BFFE6FF017098FBD1F99C.taxon	discussion	Comments: Type material for E. acceptus could not be located (Shen & Leschen 2019).	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D808FFE5FF01750EFEC7FB73.taxon	materials_examined	Type locality: [ND] Near Whangarei Harbour. Type depository: BMNH LT ♂, 1 PLT.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D808FFE5FF01750EFEC7FB73.taxon	discussion	Comments: Eupines (E.) nasuta has a median antebasal fovea and a punctate pronotal base, characters that are consistent with members of the genus Eupinolus. The peniculate scape, the narrow frons and the small subapical protuberance of protibiae in males also match the types of Eupinolus altulus, resulting in the new synonomy recognized above.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D808FFE5FF0176E9FD7FFCBF.taxon	description	(Figs. 11 – 12 in Shen and Leschen, 2019)	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D808FFE5FF0176E9FD7FFCBF.taxon	materials_examined	Type locality: [WO] Tuakau. Type depository: BMNH LT ♂.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D808FFE5FF0176E9FD7FFCBF.taxon	discussion	Comments: An examination of the lectotype of E. (E.) calcarata found that it has a median antebasal fovea and a punctate pronotal base, characters which are consistent with members of the genus Eupinolus Oke, thereby resulting in the new combination recognized above.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D808FFE5FF0173C2FBA4F96B.taxon	description	(Figs. 45 – 46 in Shen and Leschen, 2019)	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D808FFE5FF0173C2FBA4F96B.taxon	materials_examined	Type locality: [ND] Manaia, Whangarei Harbour. Type depository: BMNH HT ♂.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D808FFE5FF0173C2FBA4F96B.taxon	discussion	Comments: The holotype of E. (B.) ignotus has strongly narrowed elytral bases and the glabrous pronotum lacking median antebasal fovea and the antebasal sulcus that are consistent with members of the genus Gastrobothrus Broun. Here we formally transfer this species from Eupines to Gastrobothrus.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D813FFFEFF017691FC08FC45.taxon	materials_examined	Type locality: [HB] Mangawhare, Northern Wairoa. Type depository: BMNH HT ♂.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
C0438794D813FFFEFF017691FC08FC45.taxon	discussion	Comments: The holotype of E. (E.) platynota has strongly narrowed elytral bases and the glabrous pronotum lacking median antebasal fovea and antebasal sulcus that are consistent with members of the genus Gastrobothrus. Here we formally transfer this species from Eupines to Gastrobothrus.	en	Shen, Jia-Wei, Leschen, Richard A. B. (2020): Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae). Zootaxa 4777 (1): 1-84, DOI: 10.11646/zootaxa.4777.1.1
