identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
43EC867ECFDE545A83C94CB08DF980AB.text	43EC867ECFDE545A83C94CB08DF980AB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Homoeusa brevitarsis Nozaki & Maruyama 2025	<div><p>Homoeusa brevitarsis Nozaki &amp; Maruyama sp. nov.</p><p>Figs 3 G, 17 A – L, 18 (Japanese name: Nise-yamato-hirata-ariyadori)</p><p>Material examined.</p><p>Holotype • 1 ♂; “ Nipppon: Honshû / Wakayama-ken / Minabegawa-mura / 17. VII. 1999 (banana t) / leg. Maruyama-M. ” (KUM).</p><p>Paratypes. Japan: Honshu; Wakayama-ken • 1 ♂; Kimitoge Pass, Hashimoto-shi; 6. VII. 2014; T. Ito leg. (OMNH) • 1 ♀; same locality; 9. VII. 2008; T. Ito leg. (OMNH) • 1 ex.; Tsuchibotoke-tôge, Iwade-shi; 10. V. 2019; I. Matoba leg. (OMNH) • 1 ♀; same data as holotype (KUM) • 2 ♂♂, 3 ♀♀; Nishihara (alt. 180 m) [33.781, 135.413], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=135.413&amp;materialsCitation.latitude=33.781" title="Search Plazi for locations around (long 135.413/lat 33.781)">Hidakagawa-chô</a>; 11. VI. 2023; TS. Nozaki leg. LFS (KUM) .</p><p>Diagnosis.</p><p>This species is distinguished from other species of the genus by the following combination of characteristics: pronotal postero-lateral angle around 90 degrees (Fig. 17 C); metatarsi shorter than half of metatibiae (Fig. 17 F, G); apical lobe of paramere short, broad, and almost straight, length about 1 / 4 of whole paramere, apex distinctly truncate (Fig. 17 K). This species is similar to H. japonica and H. prolongata in general appearance, but it can be distinguished from these two species by its very short metatarsi and truncate apex of the parameral apical lobe.</p><p>Description.</p><p>Body (Fig. 3 G) small, broad; dorsal surface mostly moderately polished.</p><p>Head large; color brown; eyes small. Antennae (Fig. 17 A, B) short relative length of each antennal segment from first to 11 th, 2.48: 1.96: 1.39: 1.22: 1.04: 1.09: 1.00: 1.15: 1.15: 1.37: 3.26.</p><p>Thorax. Pronotum (Fig. 17 C) subrectangular, transverse (PW / PL, 1.56–1.66), widest near middle, postero-lateral angle around 90 degrees, posterior margin slightly sinuate, reddish brown, surface finely covered with setae and punctures, gently polished and slightly reticulated. Elytra (Fig. 17 C, D) slightly widened posteriad, widest at middle, then narrowed roundly, posterior margins deeply notched near lateral corners, reddish brown, surface finely covered with setae and punctures; gently polished and finely reticulated. Hind wings developed. Mesoventral process (Fig. 17 E) very narrow, with weak medial carina, forming Y-shaped, apex pointed, reaching posterior 1 / 3 of mesocoxal cavities. Metaventral process slightly produced.</p><p>Abdomen elongate, narrowed posteriad; surface covered with short setae and each posterior margin with long stout setae; finely reticulated.</p><p>Male: Metatarsi (Fig. 17 F) as long as 1 / 3 of hind tibia. 8 th sternite (Fig. 17 H) produced roundly at postero-medial margin; median lobe of aedeagus (Fig. 17 J) slender, length about 0.61 mm, apical lobe narrow, slightly curved ventrally, apex gently pointed in lateral view, apical valves elongate and mitten-shaped, dorsal arms long and sharpened apically, basal part of copulatory piece small. Paramere (Fig. 17 K) length about 0.73 mm; apical lobe of paramere short and broad, length about 1 / 4 of whole paramere, almost straight, apex distinctly truncate; four setae, b located subapically, c irregularly located more dorsally; velum elongate and narrow.</p><p>Female: Metatarsi (Fig. 17 G) slightly shorter than half of metatibia. 8 th sternite (Fig. 17 I) slightly rounded posterior margin; spermatheca (Fig. 17 L) apex of basal part slightly swollen, base largely hooklike.</p><p>Measurements (in mm) and ratios.</p><p>Body parts (N = 8): BL ≈ 2.35–2.87; AL, 0.79–0.88; HW, 0.53–0.59; PL, 0.57–0.64; PW, 0.9–0.99; EL, 0.46–0.52; EW, 0.91–1.04; HTL, 0.6–0.74; PW / PL, 1.56–1.66; AL / PL, 1.35–1.46; HTL / PL, 1.04–1.22.</p><p>Differences between sexes: HTL (male, N = 4), 0.69–0.74; HTL (female, N = 4), 0.6–0.66; HTL / PL (male, N = 4), 1.09–1.22; HTL / PL (female, N = 4), 1.04–1.09.</p><p>Distribution.</p><p>Japan (Kii peninsula, Honshu) (Fig. 18).</p><p>Symbiotic hosts.</p><p>Lasius fuliginosus species group: Lasius spathepus .</p><p>Bionomics.</p><p>Homoeusa brevitarsis sp. nov. was collected from foraging trails of host ants on the forest floor from May to July. No individual was collected using FIT. Their behavior with the host ants is unknown.</p><p>Remarks.</p><p>Although several hundred Homoeusa specimens collected from colonies of the Lasius fuliginosus species group were examined in this study, this new species is only known from the Kii Peninsula, suggesting a restricted distribution.</p><p>Etymology.</p><p>The specific name is derived from the Latin adjective, which means “ short footed ” and refers to the short tarsi of this new species.</p></div>	https://treatment.plazi.org/id/43EC867ECFDE545A83C94CB08DF980AB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Nozaki, Tsubasa;Maruyama, Munetoshi	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
0081CFB869745162881E9CA6856EE80C.text	0081CFB869745162881E9CA6856EE80C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Homoeusa gigantea Nozaki & Maruyama 2025	<div><p>Homoeusa gigantea Nozaki &amp; Maruyama sp. nov.</p><p>Figs 3 F, 15 A – I, 16 A – E (Japanese name: Ô-hirata-ariyadori)</p><p>“ Oxypodini ? sp. 1 ”: Nishida 2019: 135, fig. 10 (recorded from Saga pref.).</p><p>Material examined.</p><p>Holotype (Fig. 3 F) • 1 ♂; “ [Japan]: Fukuoka-ken, Soeda- ” / “ machi, Mt. Hikosan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.9174&amp;materialsCitation.latitude=33.4883" title="Search Plazi for locations around (long 130.9174/lat 33.4883)">Takasubaru</a> ” / “ [33.4883, 130.9174], alt. 770 ” / “ m, 24. VII. 2022, Tsubasa ” / “ NOZAKI leg. LCH 01. ” / “ 添田町英彦山鷹巣原 ” // “ Lasius ” / “ umbratus (Nylander, 1846) ” / “ アメイロケアリ ” / det. Tsubasa NOZAKI 2025 ” (KUM).</p><p>Paratype. Japan: Hokkaido • 1 ♀; Makoi, Shari-chô; 28. V. 2000; Y. Kida leg. LUM (KUM) • 2 ♀♀; Tôro lakeside, Shibecha-chô; 28. VII. 1986; S. Nomura leg. (KUM) • 2 ♀♀; Nopporo, Ebetsu-shi; 13–21. VI. 2000; Shigehisa Hori leg. (KUM) • 1 ex.; Minaminosawa, Sapporo-shi; 16–23. V. 2023; Y. Tasaku leg. FIT (HUM) • 1 ex.; Hakken-zan, Sapporo-shi; 31. V. 2002; M. Maruyama leg. (KUM) • 1 ♂; same locality; 20. VII. 2022; TS. Nozaki leg. LUU (KUM) • 1 ex.; Obihiro-no-mori (alt. 78 m) [42.8987, 143.1439], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=143.1439&amp;materialsCitation.latitude=42.8987" title="Search Plazi for locations around (long 143.1439/lat 42.8987)">Obihiro-shi</a>; 15. VII. 2022; TS. Nozaki &amp; H. Nozaki leg. LUU (KUM) • 1 ex.; Higashi-ônura, Nanae-chô; 16. VI. 1986; S. Nomura leg. (KUM) . Honshu: Iwate-ken • 1 ♂, 1 ex.; Kitayama-sansakuro, Morioka-shi; 2. V. 2020; N. Nakaya leg. LUU (IPMM) • 1 ex.; Kitayama, Morioka-shi; 23. IV – 3. V. 2024; N. Nakaya leg. FIT (IPMM) • 1 ex.; Mt. Takabora, Kamiyonai, Morioka-shi; 3. V. 2024; N. Nakaya leg. FIT (IPMM) . Gunma-ken • 1 ex.; Yubisogawa, Minakami-machi; 18. VI. 2004; M. Yasaka leg. (KUM) . Nagano-ken • 2 exs.; Odairahara, Horigane (alt. 1,100 m), Azumino-shi; 12. VII. 1998; M. Maruyama leg. LUU (KUM) • 1 ex.; Fujii, Satoyamabe [36.2513, 137.3436], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=137.3436&amp;materialsCitation.latitude=36.2513" title="Search Plazi for locations around (long 137.3436/lat 36.2513)">Matsumoto-shi</a>; 13. V. 2021; S. T. Inoue leg. LUU (KUM) • 1 ex.; Iriyamabe, Matsumoto-shi; 30. VI. 2012; M. Maruyama leg. LUU (KUM) • 1 ex.; Fujimi (alt. 1,510 m) [35.904, 138.190], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.19&amp;materialsCitation.latitude=35.904" title="Search Plazi for locations around (long 138.19/lat 35.904)">Fujimi-chô</a>; 4. VI. 2023; TS. Nozaki leg. LUM (KUM) . Gifu-ken • 1 ♀; Nabedaira-kôgen, Takayama-shi; 6. VII. 2022; TS. Nozaki leg. LUM (KUM) • 2 ♂♂, 1 ♀; Nakao (alt. 1,200 m), Takayama-shi; 5. VI. 2005; K. Kinomura leg. LUM (KUM) • 1 ex.; Iwai (alt. 830 m) [36.1320, 137.3436], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=137.3436&amp;materialsCitation.latitude=36.132" title="Search Plazi for locations around (long 137.3436/lat 36.132)">Takayama-shi</a>; 7. VII. 2022; TS. Nozaki leg. LUM (KUM) • 1 ex.; Iwase (alt. 850 m), Takayama-shi; 5. V. 2005; K. Kinomura leg. LUU (KUM) • 1 ex.; same locality; 4. VI. 2006; K. Kinomura leg. LUU (KUM) . Nara-ken • 1 ♂; Wasamata-yama (alt. 1,140 m) [34.2184, 135.9862], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=135.9862&amp;materialsCitation.latitude=34.2184" title="Search Plazi for locations around (long 135.9862/lat 34.2184)">Kamikitayama-mura</a>; 14. VI. 2023; TS. Nozaki leg. LUM (KUM) . Okayama-ken • 1 ex.; Jabuchi-no-taki, Mt. Nagi-san (alt. 579 m) [35.1567, 134.1908]; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.1908&amp;materialsCitation.latitude=35.1567" title="Search Plazi for locations around (long 134.1908/lat 35.1567)">Nagi-chô</a>; 15. V. 2022; Y. Senda leg. (EUMJ) . Hiroshima-ken • 1 ex.; Kakezu-yama, Higashiyawatahara (alt. 850–1,126 m), Kitahiroshima-chô; 27. VI – 4. VII. 2009; TA (Tatsuya). Nozaki &amp; Yoko Nozaki leg. FIT (KUM) . Shikoku: Kochi-ken • 1 ex.; Irazu-keikoku, Tsuno-chô; 14. V. 2005; TA. Miyata &amp; TO. Miyata leg. (KUM) . Kyushu: Fukuoka-ken • 2 exs.; Takasubaru, Mt. Hikosan [33.4883, 130.9174], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.9174&amp;materialsCitation.latitude=33.4883" title="Search Plazi for locations around (long 130.9174/lat 33.4883)">Soeda-machi</a>; 18. VI. 2021; TS. Nozaki leg. LUU (KUM) • 1 ♀, 1 ex.; same locality; 26. VI. 2021; TS. Nozaki leg. LUM (KUM) • 2 ♂♂; Biol. Lab. KU, Hikosan, (alt. 750 m) [33.4806, 130.9090], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.909&amp;materialsCitation.latitude=33.4806" title="Search Plazi for locations around (long 130.909/lat 33.4806)">Soeda-machi</a>; 26. V. 2021; TS. Nozaki leg. LUU (KUM) • 1 ex.; Zuibai-ji, Itoshima-shi; 8. V. 2021; T. Hashizume leg. (KUM) • 3 exs.; Mt. Shaka-dake, Kitayabe, Yame-shi; 18. V. 2017; S. Imasaka leg. FIT (KUM) . Saga-ken • 1 ex.; Kurokami-yama, Arita-chô; 27. IV. 2018; M. Nishida leg. FIT (KUM) • 1 ex.; same locality; 9. V. 2018; M. Nishida leg. FIT (KUM) . Kumamoto-ken • 1 ♂; Momiki (alt. 890 m) [32.494, 130.988], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.988&amp;materialsCitation.latitude=32.494" title="Search Plazi for locations around (long 130.988/lat 32.494)">Yatsushiro-shi</a>; 24. V. 2023; TS. Nozaki leg. LUM (KUM) • 1 ♀; Mt. Shiratomi-yama [33.4883, 130.9174], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.9174&amp;materialsCitation.latitude=33.4883" title="Search Plazi for locations around (long 130.9174/lat 33.4883)">Yatsushiro-shi</a>; 3. V. 2022; Y. Uehara leg. (KUM) . Oita-ken • 1 ♀, 3 exs.; Mt. Sobo; 17. V. 1986; S. Nomura leg. (KUM) • 1 ex.; Tsukahara, Yufu-shi; 16. V. 2014; T. Hada leg. (KUM) .</p><p>Diagnosis.</p><p>This species is distinguished from other species of the genus by the following combination of characteristics: pronotal posterolateral angle distinctly acute (Fig. 15 B); elytra distinctly widened posteriad entirely (Fig. 15 B, C); metaventral process slightly produced (Fig. 15 D); median lobe of aedeagus bulbous, length approximately 0.63 mm (Fig. 15 G). The appearance of this species is similar to H. laevigata and H. prolongata, but it can be distinguished from H. laevigata by its normal long setae of pronotum and elytra, and from H. prolongata by its larger body, aedeagus, and distinct acute posterolateral angle.</p><p>Description.</p><p>Body (Figs 3 F, 16 E) large, broad, somewhat limuloid; dorsal surface mostly moderately polished.</p><p>Head small; color yellowish brown to brown; eyes small. Antennae (Fig. 15 A) short, relative length of each antennal segment from first to 11 th, 2.91: 2.25: 1.66: 1.27: 1.18: 1.2: 1.23: 1.18: 1.00: 1.45: 4.93.</p><p>Thorax. Pronotum (Fig. 15 B) semicircular, transverse (PW / PL, 1.62–1.80), widest at posterior 1 / 3, postero-lateral angle distinctly acute, posterior margin distinctly sinuate, yellowish brown to brown, sometimes slightly transparent, surface finely covered with long setae and punctures, gently polished and slightly reticulated. Elytra (Fig. 15 B, C) distinctly widened posteriad entirely, posterior margins deeply notched near lateral corners, yellowish brown to brown, surface finely covered with long setae and punctures; gently polished and weakly reticulated. Hind wings developed. Mesoventral process (Fig. 15 D) very narrow, with indistinct medial carina, apex sharply pointed, reaching almost anterior margin of metaventrite. Metaventral process slightly produced (Fig. 15 D).</p><p>Abdomen elongate, remarkably narrowed posteriad; surface sparsely covered with short setae and each posterior margin with long stout setae; weakly reticulated.</p><p>Male: 8 th sternite (Fig. 15 E) weakly produced and angled at postero-medial margin; median lobe of aedeagus (Fig. 15 G) bulbous, length about 0.63 mm, apical lobe elogate and slightly curved ventrally, apex round in lateral view, apical valves elongate and mitten-shaped, dorsal arms short, basal part of copulatory piece large. Paramere (Fig. 15 H) length about 0.63 mm; apical lobe of paramere long, length about 1 / 3 of whole paramere, curved ventrally; four setae, b located more basally than c; velum small.</p><p>Female: 8 th sternite (Fig. 15 F) weakly produced roundly postero-medial margin; spermatheca (Fig. 15 I) S-shaped, apex of basal part slightly sinuate, base largely hooklike.</p><p>Variation.</p><p>Intraspecies variations were recognized in the apical lobe of the aedeagal median lobe. Some individuals have a small hump at the base of the ventral side and a slightly deeper basal concave. The length of the parameral apical lobe also varies slightly.</p><p>Measurements (in mm) and ratios.</p><p>Body parts (N = 10): BL ≈ 2.48–4.3; AL, 0.92–1.15; HW, 0.55–0.64; PL, 0.67–0.83; PW, 1.09–1.39; EL, 0.51–0.62; EW, 1.06–1.34; HTL, 0.61–0.75; PW / PL, 1.62–1.80; AL / PL, 1.22–1.46; HTL / PL, 0.89–1.02.</p><p>Distribution.</p><p>Japan (Hokkaido, Honshu, Shikoku, Kyushu) (Fig. 16 A).</p><p>Symbiotic hosts.</p><p>Lasius umbratus species group: L. umbratus, L. meridionalis .</p><p>Bionomics.</p><p>Homoeusa gigantea sp. nov. is rarely observed in host ants’ trails under stones or soil from April to July (Fig. 16 B, C). Some individuals were collected by FIT (N = 15). Setting baits near the ants’ foraging trails under stones (like Fig. 1 C) is effective for collecting H. gigantea sp. nov. During indoor observations, they frequently and proactively rubbed their abdomen against body of their host ant workers (Fig. 16 B, D), unlike any other species of this genus.</p><p>Remarks.</p><p>Not many specimens were collected from foraging trails without bait. This might be because of the foraging behavior of the host ant species, Lasius umbratus and L. meridionalis . Both species are members of the Lasius umbratus group (formerly called the subgenus Chthonolasius Ruzsky, 1912) considered to forage mainly underground (Donisthorpe 1927; Collingwood 1979; Seifert 1988). We also observed these Lasius species forming trails under stones or inside litter. Given this, we believe that they tend to forage mainly underground. To find Homoeusa gigantea sp. nov. in these trails naturally, first, a collector needs to find the trail. At the same time, the beetle must pass through the right stone on the long path all over the underground region. This is why a bait trap is effective. It can attract many ant workers and beetles under a particular stone. TN collected multiple individuals per bait set under a stone, suggesting that the species may potentially be more common than expected in their foraging trails.</p><p>Etymology.</p><p>The specific name is derived from the Latin adjective, which means “ huge ” and refers to the large size of this new species.</p></div>	https://treatment.plazi.org/id/0081CFB869745162881E9CA6856EE80C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Nozaki, Tsubasa;Maruyama, Munetoshi	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
DAAE202EF1DF5CEAA714ABFE50E7AF80.text	DAAE202EF1DF5CEAA714ABFE50E7AF80.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Homoeusa japonica Sharp 1874	<div><p>Homoeusa japonica Sharp, 1874</p><p>Figs 2 B, E, 3 C, 9 A – L, 10 A – H (Japanese name: Yamato-hirata-ariyadori)</p><p>Homoeusa japonica Sharp, 1874: 5 (original description); Schülke and Smetana 2015; 682 (catalogue); Maruyama et al. 2013: 25 (distribution and bionomics mentioned).</p><p>“ Homoeusa acuminata ”; Sharp 1888: 283 (recorded from “ Miyanoshita ” as H. acuminata, which is a misidentification).</p><p>Type material.</p><p>Lectotype (Fig. 1 B, E) • here designated; ♂; “ Japan. ” / G. Lewis. ” // “ Sharp Coll. ” / 1905-313 ” // “ Type ” (red round curator label) // “ SYN-TYPE ” (blue round curator label) // “ Homoeusa ” / “ japonica ” / “ type D. S. ” “ LECTOTYPE ” / “ Homoeusa ” / “ japonica ” / “ SHARP, 1874 ” / “ det. Maruyama, 2003 ” // “ NHMUK 014379775 ” (NHM).</p><p>Nontypes in the Sharp Collection.</p><p>• 1 ex.; “ Homoeusa ” / “ japonica ” / “ var. ” / “ Japan Lewis. ” // “ Japan. ” / G. Lewis. ” (written on the card which the specimen mounted) // “ Sharp Coll. ” / 1905-313 ” // “ Homoeusa ” / “ japonica ” // “ Homoeusa ” / “ japonica ” / “ SHARP, 1874 ” / “ det. Maruyama, 2003 ” // (NHM) • 1 ex.; “ Japan. ” / “ G. Lewis. ” / “ Sharp Coll. ” / 1905-313 ” // “ Homoeusa ” / “ japonica Sharp ” // “ Homoeusa ” / “ japonica ” / “ SHARP, 1874 ” / “ det. Maruyama, 2003 ” // (NHM) • 2 exs.; “ Japan. ” / “ G. Lewis. ” / “ 1910-320. ” // “ Homoeusa japonica ” // “ Homoeusa ” / “ japonica ” / “ SHARP, 1874 ” / “ det. Maruyama, 2003 ” // (NHM) • 1 ♂ “ Miyanoshita ” // “ Japan. ” / “ G. Lewis. ” / “ 1910-320. ” // “ Homoeusa ” / “ acuminata ” // “ Homoeusa ” / “ japonica ” / “ SHARP, 1874 ” / “ det. Maruyama, 2003 ” // “ Sharp (1874) ” / “ recorded as ” / ““ H. acuminata ”” / “ M. M. ” (NHM).</p><p>Additional material examined.</p><p>Japan: Honshu: Aomori-ken • 2 ♂♂; Hyakuzawa-higashi, Iwakisan [40.629, 140.327], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=140.327&amp;materialsCitation.latitude=40.629" title="Search Plazi for locations around (long 140.327/lat 40.629)">Hirosaki-shi</a>; 25. VI. 2023; TS. Nozaki leg. (KUM) . Iwate-ken • 1 ex.; Gandô-ko, Morioka-shi; 9. VI. 2019; N. Nakaya leg. LFcfF (IPMM) • 3 exs.; Takamatsu-kôen, Morioka-shi; 22. VII. 2019; N. Nakaya leg. LFcfF (IPMM) • 1 ex.; Kitayama-sansakuro, Morioka-shi; 19. V. 2019; N. Nakaya leg. LFcfF (IPMM) • 1 ex.; same locality; 5. V. 2019; N. Nakaya leg. LFcfF (IPMM) • 1 ♂, 3 exs.; Kitayama, Morioka-shi; 3. V. 2023; N. Nakaya leg. LFcfF (IPMM) . Miyagi-ken • 1 ♀; Naruko-onsen, Osaki-shi; 14–17. VI. 1999; M. Sano leg. LFO (KUM) • 1 ♂; Aoba-yama, Sendai-shi; 17. VII. 2004; K. Mizota leg. LFN (KUM) • 2 ♀♀; Toriido-yama, Komono-chô; 2. VI. 2012; N. Tsuji leg. (KUM) . Akita-ken • 1 ♂; Tamagawa, Tazawako, Semboku-shi; 12–13. VI. 1999; M. Sano leg. LFcfF (KUM) . Fukushima-ken • 1 ♂, 2 exs.; Kashi Spa., Nishigo-mura; 16. VI. 1998; M. Maruyama leg. LFcfF (KUM) • 1 ♀, 2 exs.; same locality; 17. VI. 1998; M. Maruyama leg. LFcfF (KUM) • 1 ♂, 1 ex.; Yukiwari-bashi, Nishigo-mura; 29. VII. 2000; T. Kobashi leg. (KUM) • 3 exs.; Mizuhiki, Tateiwa, Minamiaizu-machi; 12. VI. 2004; H. Kamezawa leg. LFcfF (cKam) . Ibaraki-ken • 1 ♂; Sadanami, Sekimoto-chô, Kitaibaraki-shi; 4. V. 2022; Y. Komatsuzaki leg. (KUM) • 1 ♂; Oshô-yama, Kitaibaraki-shi; 24. IV. 2022; S. Fuda leg. (KUM) • 1 ex.; Takeo-rindo (alt. 450 m), Hitachiota-shi; 3. V. 2022; Y. Komatsuzaki leg. (KUM) • 1 ♂; Asahi-tôge, Tsuchiura-shi; 8. V. 2022; Y. Komatsuzaki leg. (KUM) • 2 exs.; Nagai, Tsuchiura-shi; 27. IV. 2022; Y. Komatsuzaki leg. (KUM) . Tochigi-ken • 1 ♂, 1 ♀, 72 exs.; Ichikai-machi; 29. IV. – 2. V. 2002; S. Nagashima leg. LFcfF (KUM) . Gunma-ken • 2 ♂♂, 2 ♀♀, 4 exs.; Nakanojo Forest Park, Nakanojo-machi; 8. VI. 2001; T. Watanabe leg. (KUM) • 1 ex.; “ 4 km NW Onishidashi Rock Gdn. ”; “ 18–21. VII. 80 ” [1980]; A. &amp; Z. Smetana leg. (NMST, KUM, ex A. Smetana coll.) • 1 ex.; Sakurayama, Fujioka-shi; 3. V. 1998; S. Arai leg. (KUM) • 1 ♀, 28 exs.; Sakurayama Park, Onishi, Fujioka-shi; 9. V. 1998; S. Arai leg. LFS (KUM) • 1 ♂, 21 exs.; Mt. Sakurayama, Onishi, Fujioka-shi; 9. V. 1998; K. Toyoda leg. LFS (KUM) • 6 exs.; same locality; 18. V. 1998; K. Toyoda leg. LFS (KUM) • 5 ♂♂, 1 ♀, 18 exs.; same locality; 22. V. 1999; Shiho Arai leg. LFS (KUM) • 5 exs.; Nageishi-tôge, Fujioka-shi; 5. VI. 2001; T. Watanabe leg. (KUM) . Saitama-ken • 3 exs.; Sugiyama Jôshi, Ranzan-machi; 29. IV. 2005; K. Toyoda leg. (KUM) • 1 ex.; Mino-yama, Minano-machi; 15. VII. 2000; K. Toyoda leg. (KUM) • 10 exs.; Shioyama, Ranzan-machi; 21. VI. 1996; K. Toyoda leg. (KUM) • 1 ♂, 4 exs.; same locality; 8. VI. 1997; K. Toyoda leg. (KUM) • 2 exs.; Mt. Shioyama, Ranzan-machi; 10. V. 1998; K. Toyoda leg. LFcfF (KUM) • 1 ex.; same locality; 3. VII. 1998; K. Toyoda leg. (KUM) • 2 exs.; Shiro-yama, Kamagata, Ranzan-machi; 17. VI. 2000; K. Toyoda leg. (KUM) • 1 ex.; Near Toki-gawa, Ranzan-machi; 22. IV. 1999; K. Toyoda leg. LFcfF (KUM) • 5 exs.; Toki-gawa, Ranzan-machi; 25. IV. 1999; K. Toyoda leg. (KUM) • 2 exs.; same locality; 14. V. 2000; K. Toyoda leg. (KUM) • 1 ex.; Shôgunsawa, Ranzan-machi; 21. IV. 1999; K. Toyoda leg. (KUM) • 2 exs.; Shiroishi-tôge, Higashichichibu-mura; 9. V. 1999; K. Toyoda leg. (KUM) • 3 exs.; same locality; 20. VI. 1999; S. Arai leg. (KUM) • 2 exs.; Kuroishi Jinja, Kumai, Hatoyama-machi; 29. IV. 2000; S. Arai leg. LFS (KUM) • 1 ♂, 1 ex.; Nr. Ôno Pass, Tokigawa-machi; 5. VIII. 1998; K. Toyoda leg. LFcfF (KUM) • 1 ex.; Ushidatekubo (alt. 830 m) [35.9685, 139.1793], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=139.1793&amp;materialsCitation.latitude=35.9685" title="Search Plazi for locations around (long 139.1793/lat 35.9685)">Tokigawa-machi</a>; 21. V. 2023; S. T. Inoue &amp; N. Katsube leg. LFcfF (KUM) • 1 ex.; Hashidate, Chichibu-shi; 26. IV. 1998; K. Toyoda leg. LFS (KUM) • same locality; 26. IV. 1998; S. Arai leg. (KUM) • 1 ♂, 1 ex.; same locality; 17. VIII. 1999; K. Toyoda leg. LFN (KUM) • 1 ♀, 5 exs.; same locality; 30. VII. 2000; K. Toyoda leg. LFN (KUM) • 1 ex.; same locality; 7. VI. 2002; K. Toyoda leg. (KUM) • 1 ex.; Tenjinsha, Kita-hirasawa, Hidaka-shi; 4. VI. 1999; K. Toyoda leg. (KUM) . Chiba-ken • 11 exs.; Azeta, Sakura-shi; 20. VI. 1998; M. Maruyama leg. LFS (KUM) • 1 ♀, 4 exs.; same locality; 23–24. VI. 1998; M. Maruyama leg. LFS (KUM) • 2 exs.; same locality; 20–26. VI. 1998; M. Maruyama leg. LFS (KUM) • 2 exs.; same locality; 24. VI. 1998; M. Sano leg. (KUM) • 3 exs.; Aoba, Chiba-shi, 24. VI. 1998; M. Maruyama leg. LFS (KUM) • 6 exs.; Mt. Mifuneyama, Shôkô, Kimitsu-shi; 24. IV. 2006; H. Ono leg. LFN (KUM) • 6 exs.; same locality; 7. V. 2006; H. Ono leg. (KUM) • 1 ♀; Umegase, Daifuku-yama, Ichihara-shi, 6. V. 1998; S. Naomi leg. (KUM) • 2 exs.; Shimin-nomori Yachô-no-mori, Toyooka, Futtsu-shi; 19. IV. 2006; H. Ono leg. (KUM) . Tokyo-to • 2 exs.; Kurosawa, Oume-shi; 29. IV. 1998; U. Kurosu leg. (SPMN) • 1 ex.; Dokan-bori, Imperial Palace, Chiyoda-ku; 17. V. 2000; T. Shimada leg. (NSMT) • 3 exs.; Mt. Takao, Hachiôji-shi; 1. V. 1985; S. Kubota leg. (KUM) • 1 ex.; Mt. Takao (alt. 500 m), Hachiôji-shi; 1. VII. 1998; M. Maruyama leg. LFS (KUM) • 3 exs.; same locality; 4. VII. 1998; M. Maruyama leg. LFS (KUM) • 1 ex.; same locality; 1–4. VII. 1998; M. Maruyama leg. LFS (KUM) • 1 ♂, 2 ♀♀, 20 exs.; same locality; 4. VI. 2001; M. Maruyama leg. LFS (KUM, NHM) • 3 exs.; Mt. Takao (alt. 450 m), Hachiôji-shi; 4. VI. 2001; M. Maruyama leg. LFS (NHM) . Kanagawa-ken • 1 ♂, 1 ex.; Mt. Masukata, Kawasaki-shi; 15. VI. 1996; K. Kawada leg. (KUM) • 5 exs.; Ikuta-ryokuchi, Kawasaki-shi; 13. IV. 2002; K. Matsumoto leg. (KUM) • 4 exs.; Mikage, Tsukui, Sagamihara-shi; 18. IV. 1976; R. Kiryu leg. (SCM) • 2 ex.; same locality; 23. IV. 1977; R. Kiryu leg. (SCM) • 1 ex.; same locality; 14. IV. 1979; R. Kiryu leg. (SCM) • 1 ex.; same locality; 31. V. 1979; R. Kiryu leg. (SCM) • 2 exs.; same locality; 9. VI. 1979; R. Kiryu leg. (SCM) • 1 ex.; Nanzawa, Tsukui, Sagamihara-shi; 29. V. 1976; R. Kiryu leg. (SCM) • 4 exs.; same locality; 14. V. 1977; R. Kiryu leg. (SCM) • 2 exs.; same locality; 20. V. 1977; R. Kiryu leg. (SCM) • 1 ex.; same locality; 22. IV. 1979; R. Kiryu leg. (SCM) • 1 ex.; same locality; 24. V. 1979; R. Kiryu leg. (SCM) • 1 ex.; Miyagase, Tsukui; 9. VII. 1976; R. Kiryu leg. (SCM) • 12 exs.; Toya, Tsukui, Sagamihara-shi; 11. V. 1976; R. Kiryu leg. (SCM) • 8 exs.; same locality; 27. IV. 1976; R. Kiryu leg. (SCM) • 5 exs.; same locality; 29. IV. 1977; R. Kiryu leg. (SCM) • 1 ex.; same locality; 17. IV. 1979; R. Kiryu leg. (SCM) • 17 exs.; Kawasaki; 1. V. 1985; S. Kubota leg. (KUM) • 2 exs.; “ Trap (liver) St. 1 Y. N. S ”; Yokohama-shi; 10. V. 1988; Y. Kominami leg. (SPMN) • 3 exs.; Niiharu-shiminno-mori, Yokohama-shi; 21. V. 2001; T. Watanabe leg. LFO, LFS (KUM) • 1 ex.; Nonkendainomori, Enkaizan, Yokohama-shi; 3. V. 1991; K. Kubo leg. (SPMN) • 1 ex.; Yakunimi-yama [35.35, 139.17], Hadano-shi; 26. VI. 2014; O. Saitoh leg. LFcfF (KPMNH) • 4 exs.; Kawana, Fujisawa-shi; 14. V. 2001; T. Watanabe leg. (KUM) • 1 ♂, 15 exs.; Jimmu-ji, Zushi-shi; 20. VI. 2003; M. Maruyama leg. (KUM) • 1 ex.; Futagoyama, Zushi-shi; 15. IV. 1991; H. Watarai leg. (SPMN) • 1 ex.; Kinugasa, Yokosuka-shi; 26–27. IV. 1995; Y. Kaneko leg. (KUM) • 3 exs.; Miyanoshita, Hakone, Hakone-machi; 5. V. 1987; Y. Hirano leg. (KUM) • 1 ex.; Hakone; 1. V. 1974; Y. Hirano leg. (KUM) • 1 ex.; Miura Kurihama, Sagamihara-shi; 9. V. 1979; R. Kiryu leg. (SCM) . Yamanashi-ken • 1 ex.; Ashiyu-onsen, Ashiyu-mura; 23. V. 2001; T. Watanabe leg. (KUM) . Nagano-ken • 1 ♀, 1 ex.; Bôdaira, Shiga-kôgen (alt. 1,160 m) [36.7203, 138.4656], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.4656&amp;materialsCitation.latitude=36.7203" title="Search Plazi for locations around (long 138.4656/lat 36.7203)">Yamanouchi-machi</a>; 9. VII. 2022; TS. Nozaki leg. LFM (KUM) • 1 ♂; Fujii, Satoyamabe, Matsumoto-shi; 7. VII. 2022; TS. Nozaki leg. LFN (KUM) . Gifu-ken • 2 ♂♂; Bijo Pass, Takayama-shi, 5. VIII. 2003; K. Kinomura leg. (KUM) • 8 exs.; Ogamigo, Shôkawa (alt. 800 m), Takayama-shi; 5–6. VIII. 1998; M. Maruyama leg. LFS (KUM) • 1 ex.; Isshiki, Takayama-shi; 23. V. 2004; K. Kinomura leg. LFN (KUM) • 1 ex.; Mt. Kentô, Takayama-shi; 6. VII. 2003; K. Kinomura leg. (KUM) • 1 ♂; Neo-Mizudori, Motosu-shi; 4. VII. 2022; TS. Nozaki leg. LFN (KUM) • 1 ex.; Miyachi [Gifu-ken?]; 29. IV. 1980; Y. Ohta leg. (KUM) • 1 ex.; Yôrô Park, Yôrô-chô; 29. IV. 2004; K. Kinomura leg. (KUM) . Shizuoka-ken • 1 ex.; Kumazaka, Izu-shi; 30. V. 2022; T. Tenma leg. (KUM) . Aichi-ken • 1 ♀, 2 exs.; Jôkôji-kôen, Seto-shi; 16. IV. 2023; T. Tenma leg. (KUM) • 1 ♂, 7 exs.; Magarida, Toyooka-chô, Gamagori-shi; 4. V. 2024; T. Temma leg. LFcfF (KUM) . Kyoto-fu • 1 ♀; Amagase Forest, Uji; 8. V. 2013; T. Ito leg. (OMNH) . Osaka-fu • 1 ♂; Mt. Myoken; 2. V. 2015; T. Ito leg. (OMNH) • 1 ♀; same locality; 8. V. 2012; T. Ito leg. (OMNH) • 1 ♂; Mino-o Park; 6. VI. 2009; T. Ito leg. (OMNH) . Nara-ken • 1 ♂; Yamato-Koriyama, Yata; 18. V. 2009; T. Ito leg. (OMNH) . Wakayama-ken • 2 ♂♂, 2 ♀♀, 3 exs.; Nishihara (alt. 180 m) [33.781, 135.413], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=135.413&amp;materialsCitation.latitude=33.781" title="Search Plazi for locations around (long 135.413/lat 33.781)">Hidakagawa-chô</a>; 11. VI. 2023; TS. Nozaki leg. LFS (KUM) . Tottori-ken • 1 ♂; Mt. Hôbutsu (alt. 420 m) [35.2271, 133.4561], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=133.4561&amp;materialsCitation.latitude=35.2271" title="Search Plazi for locations around (long 133.4561/lat 35.2271)">Hino-chô</a>; 17. VI. 2024; TS. Nozaki leg. (KUM) . Shimane-ken • 1 ♂, 3 exs.; Urahikimi, Masuda-shi, Masuda-shi; 6. VI. 1998; S. Nomura leg. LFS (KUM) . Okayama-ken • 2 ♂♂, 1 ♀, 5 exs.; Ono Shine, Kawakami, Ohara-chô, Mimasaka-shi; 5. V. 2009; Y. Fujitani leg. LFS (KUM) • 1 ex.; same locality; 7. V. 2009; Y. Fujitani leg. (KUM) • 1 ex.; same locality; 15. V. 2009; Y. Fujitani leg. (KUM) • 1 ex.; same locality; 24. V. 2009; Y. Fujitani leg. (KUM) . Hiroshima-ken • 1 ♂, 3 exs.; Kakezu-yama, Higashiyawatahara (alt. 850–1,126 m), Kitahiroshima-chô; 9–18. V. 2009; TA. (Tastuya) Nozaki &amp; Y. Nozaki leg. FIT (KUM) . Yamaguchi-ken • 1 ex.; Nishimagura, Kusunoki (alt. 100 m); Ube-shi; 30. V – 1. VI. 2000; T. Kishimoto leg. (KUM) . Shikoku: Tokushima-ken • 1 ex.; Takashiro-yama, Naka-chô; 24. V. 2007; H. Fujimoto leg. (KUM) . Kagawa-ken • 1 ex.; Furodani, Miki-chô; 30. VII. 2000; K. Izawa leg. LFcfF (KUM) • 1 ex.; Usa-jinja, Nagaona, Sanuki-shi; 31. V. 2001; Maruyama et al. leg. LFS (KUM) • 1 ♂, 1 ex.; Fujio-Jinja, Nishi-uta-chô, Takamatsu-shi; 31. V. 2001; Maruyama et al. leg. LFN (KUM) • 3 exs.; same locality; 1. VI. 2001; Maruyama et al. leg. LFN (KUM) • 1 ex.; Ôtawa, Nagao, Sanuki-shi; 22. V. 2000; F. Ito leg. LFM (KUM) • 1 ♂, same locality; 7. VI. 2001; F. Ito leg. (KUM) • 2 exs.; Kamiyama, Miki-chô; 1. VI. 2001; Maruyama et al. leg. LFS (KUM) • 13 exs.; Atago-yama, Kotohira-chô; 1. VI. 2001; M. Maruyama leg. LFS (KUM) • 1 ♀, 7 exs.; Ôtaki-san, Shionoe-chô, Takamatsu-shi; 2. VI. 2001; Maruyama et al. leg. (KUM) • 1 ex.; Daisen-zan, Kotonami-chô; 1. VI. 2001; Maruyama et al. leg. LFM (KUM) . Ehime-ken • 1 ex.; Higashino, Matsuyama-shi; 30. V. 2015; Y. Hisasue leg. (KUM) • 3 ♀♀; same locality; 30. V. 2015 (KUM) • same locality; 22. IV. 2017; R. Koga leg. (EUMJ) . Kochi-ken • 1 ♀; Ashizuri-misaki (alt. 120 m) [32.748, 133.0136], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=133.0136&amp;materialsCitation.latitude=32.748" title="Search Plazi for locations around (long 133.0136/lat 32.748)">Tosashimizu-shi</a>; 21. V. 2024; TS. Nozaki leg. (KUM) . Kyushu: Fukuoka-ken • 1 ex.; Mt. Inunaki, Miyawaka-shi; 22. V. 1980; T. Gotô leg. (KUM) • 1 ex.; Mt. Tachibana; 6–9. V. 1996; T. Ueno, Y. Higashiura &amp; H. Goto leg. FIT (cSch) • 1 ex.; same locality; 10–14. V. 1996; T. Ueno, Y. Higashiura &amp; H. Goto leg. FIT (cSch) • 1 ♀; Takasubaru Highland, Mt Hikosan, Soeda-machi; 26. VI. 2022; S. G. (Shugo) Inoue leg. LFcfF (KUM) • 2 exs.; Biol. Lab. Mt. Hikosan, Soeda-machi; 7. VI. 1993; S. Nomura leg. LFcfF (KUM) • 2 ♂♂, 1 ♀; Biol. Lab. KU, Hikosan, (alt. 750 m) [33.48, 130.90], Soeda-machi; 22. V. 2022; TS. Nozaki leg. LFcfF (KUM) • 2 ♂♂, 1 ♀; same locality; 16. V. 2023; TS. Nozaki leg. LFcfF (KUM) . Saga-ken • 1 ♀, 1 ex.; Mt. Kurokami-yama, Arita-chô; 28. IV. 2019; M. Nishida leg. LFcfF (KUM) • 2 exs.; same locality; 15. V. 2019; M. Nishida leg. (KUM) • 1 ♂; Mt. Kurokami-yama (alt. 400 m), Arita-chô; 9. V. 2021; Y. Hisasue leg. LFcfF (KUM) • 1 ex.; Tosen-zan, Ureshino-shi; 29. IV. 2018; M. Nishida leg. FIT (KUM) . Nagasaki-ken • 1 ♂; Tanuki-no-o, Omura-shi; 5. V. 2018; M. Nishida leg. (KUM) . Oita-ken • 2 ♀♀; Oike, Kurodake, Yufu-shi; 17. VI. 2021; TS. Nozaki leg. (KUM) . Miyazaki-ken • 1 ♂; Ômoridake forest road (alt. 600 m) [32.0435, 131.1973], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=131.1973&amp;materialsCitation.latitude=32.0435" title="Search Plazi for locations around (long 131.1973/lat 32.0435)">Aya-machi</a>; 12. VI. 2024; TS. Nozaki &amp; T. Kinoda leg. LFS (KUM) .</p><p>Diagnosis.</p><p>This species is distinguished from the other species of the genus by the following combination of characteristics: pronotal posterolateral angle around 90 degrees (Fig. 9 C); elytra distinctly widened posteriad entirely (Fig. 9 C); apical lobe of paramere long, length about half of whole paramere, curved ventrally, apex simply round (Fig. 9 I, J). This species is quite similar to H. longicornis in its general appearance; however, the round apex of the parameral apical lobe can separate it.</p><p>Redescription.</p><p>Body (Figs 3 C, 10 H) medium-sized, broad, stout; dorsal surface mostly moderately polished.</p><p>Head large; color reddish brown to dark brown; eyes large. Antennae (Fig. 9 A, B) long; relative length of each antennal segment from first to 11 th, 2.32: 1.48: 1.19: 1.07: 1.05: 1.01: 1.00: 1.01: 1.25: 1.4: 2.88.</p><p>Thorax. Pronotum (Fig. 9 C) subrectangular, transverse (PW / PL, 1.59–1.71), widest near middle, postero-lateral angle around 90 degrees, posterior margin slightly sinuate, reddish yellow to reddish brown, surface finely covered with setae and punctures, gently polished and slightly reticulated. Elytra (Fig. 9 C, D) distinctly widened posteriad entirely, posterior margins shallowly notched near lateral corners, reddish yellow to reddish brown, surface finely covered with setae and punctures; gently polished and weakly reticulated. Hind wings developed. Mesoventral process (Fig. 9 E) very narrow, with weak medial carina, forming Y-shaped, apex sharply pointed, reaching posterior 1 / 3 of mesocoxal cavities. Metaventral process weakly produced.</p><p>Abdomen elongate, narrowed posteriad; surface covered with short setae and each posterior margin with long stout setae; finely reticulated.</p><p>Male: 8 th sternite (Fig. 9 F) rounded at posterior margin; median lobe of aedeagus (Fig. 9 H) slender, length about 0.55 mm, apical lobe narrow and elongate, slightly curved dorsally, apex gently pointed in lateral view, apical valves simple and elongate, without dorsal arms, basal part of copulatory piece very small. Paramere (Fig. 9 I, J) length about 0.65 mm; apical lobe of paramere very long, length about half of whole paramere, curved ventrally, slightly dilated apicad, apex simply round; four setae, a far longer than others, b located most apically, velum short and suboval.</p><p>Female: 8 th sternite (Fig. 9 G) truncated or slightly rounded at posterior margin; spermatheca (Fig. 9 K, L) S-shaped, base of apical part not swollen, base largely hooklike.</p><p>Variations.</p><p>Geographical variations were found in the parameral apical lobe. The populations from Chugoku, Shikoku, and Kyushu had a less curved apical lobe, with a finely dentate dorsal margin (Figs 9 I, 10 A), whereas the population from the eastern part of Honshu had a more curved apical lobe with a smooth dorsal margin (Figs 9 J, 10 B – D). The distribution of each population is shown in Fig. 10 E.</p><p>Measurements (in mm) and ratios.</p><p>Body parts (N = 19): BL ≈ 2.14–3.27; AL, 0.81–1.03; HW, 0.51–0.66; PL, 0.48–0.63; PW, 0.78–1.03; EL, 0.42–0.51; EW, 0.85–1.02; HTL, 0.52–0.67; PW / PL, 1.59–1.71; AL / PL, 1.36–1.86; HTL / PL, 0.94–1.15.</p><p>Distribution.</p><p>Japan (Honshu, Shikoku, Kyushu) (Fig. 10 E).</p><p>Symbiotic hosts.</p><p>Lasius fuliginosus species group: L. cf. fuliginosus, L. morisitai, L. nipponensis, L. orientalis, L. spathepus .</p><p>Bionomics.</p><p>Homoeusa japonica can be observed in Lasius fuliginosus species group foraging trails on the forest floor from April to August (Fig. 10 F). Only a few individuals were collected using FIT (N = 7). They frequently raised the tip of their abdomen toward approaching host ant workers, both in the field and indoors (Fig. 10 G).</p><p>Remarks.</p><p>According to the original description (Sharp 1874), the type locality was assumed to be Nagasaki City, Nagasaki Prefecture, Kyushu. This species was rarely collected from the trails of L. nipponensis and L. orientalis .</p><p>The differences in the parameres between the two populations were significant; however, no other differences were found. They may have been distributed separately because they were never collected from the same locality. We conclude that they can be reasonably treated as intraspecific populations. This species tends to occur at lower altitudes than H. longicornis does.</p></div>	https://treatment.plazi.org/id/DAAE202EF1DF5CEAA714ABFE50E7AF80	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Nozaki, Tsubasa;Maruyama, Munetoshi	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
A1D83E522C585107A6EAC74C71EAF852.text	A1D83E522C585107A6EAC74C71EAF852.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Homoeusa Kraatz 1856	<div><p>Genus Homoeusa Kraatz, 1856</p><p>Homoeusa Kraatz, 1856: 76.</p><p>Myrmobiota Casey, 1893: 594.</p><p>Soliusa Casey, 1900: 53</p><p>Note.</p><p>See Nozaki and Maruyama (2022 a) for a detailed synonymic list and the diagnosis of this genus.</p><p>Remarks.</p><p>No apparent sexual dimorphism other than in the genitalia or terminalia has been reported in this genus. However, based on extensive examination of several species, females were often found to have shorter and denser stout setae on the middle posterior margin of the 8 th sternite than males did (Fig. 9 F, G). Moreover, there are more or less interspecific variation in the marginal shape of the 8 th sternite on each sex. Therefore, we provide illustrations and descriptions of this characteristic for all the species, including the well-described species in our previous papers (Nozaki and Maruyama 2022 a, 2022 b), Homoeusa rufescens and H. ovata .</p><p>To measure the antennae, we provided the width and aspect ratio of each segment in the previous paper. However, they usually have elliptical cross-sections, which can cause instability during measurement. Therefore, we provided the relative length of each segment instead.</p><p>The males have various parameral apical lobe shapes and arrangements of the four setae across species. Although strict homology is not available for these four setae, for the convenience of a comparative description, we named them as Sawada (1970 b) did for H. prolongata . Each seta typically has the following characteristics in Homoeusa (e. g., Fig. 13 H): a, longest and distributed at the most basal position; b, medium in length and distributed dorsally; c, medium in length and distributed ventrally; d, shortest, thinnest, and distributed at the most apical position.</p></div>	https://treatment.plazi.org/id/A1D83E522C585107A6EAC74C71EAF852	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Nozaki, Tsubasa;Maruyama, Munetoshi	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
1C8E87D60BD15CAA97F75475F86EDEA9.text	1C8E87D60BD15CAA97F75475F86EDEA9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Homoeusa laevigata Sharp 1888	<div><p>Homoeusa laevigata Sharp, 1888</p><p>Figs 2 A, D, 3 B, 7 A – I, 8 A – H (Japanese name: Tsuya-hirata-ariyadori)</p><p>Homoeusa laevigata Sharp, 1888: 283 (original description); Schülke and Smetana 2015; 682 (catalogue); Maruyama et al. 2013: 24 (distribution and bionomics mentioned).</p><p>Type material.</p><p>Holotype (Fig. 2 A, D) • ♂; “ Homoeusa ” / “ laevigata ” / “ Type ” / “ D. S. ” (written on the card which the specimen mounted) // “ Seba ” / “ 30. y. ’ 81. ” // “ Type ” (red round curator label) // “ HOLOTYPE ” / “ Homoeusa ” / “ laevigata ” / “ SHARP, 1888 ” / “ det. MARUYAMA, 2003 ” // “ NHMUK 014379778 ” (dissected by MM) (NHM).</p><p>Additional material examined.</p><p>Japan: Honshu: Miyagi-ken • 2 exs.; Aoba-yama, Sendai-shi; 10. VII. 2004; K. Mizota leg. LFN (KUM) • 3 ♂♂, 1 ♀, 7 exs.; same locality; 17. VII. 2004 K. Mizota leg. LFN (KUM) • 2 ♂♂; Aoba Forest Greenery [38.2577, 140.8255], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=140.8255&amp;materialsCitation.latitude=38.2577" title="Search Plazi for locations around (long 140.8255/lat 38.2577)">Sendai-shi</a>; 12. VIII. 2024; TS. Nozaki leg. LFN (KUM) . Fukushima-ken • 2 ♀♀; Kuraizaka-toge, Nishiaizu-machi; 18. VI. 2022; K. Haga leg. LFN (KUM) • 1 ♂; Ueda dam, Kaneyama-machi; 25. VI. 2023; K. Haga leg. LFN (KUM) . Ibaraki-ken • 1 ♂, 1 ex.; Mt. Yamizo, Daigo-machi; 4. V. 2022; Y. Komatsuzaki leg. (KUM) . Saitama-ken • 1 ex.; Hashidate riv. (alt. 500 m), Chichibu-shi; 17. VII. 1999; K. Toyoda leg. LFN (KUM) • 2 ♂♂, 1 ♀, 3 exs.; Hashidate, Chichibu-shi; 17. VII. 1999; K. Toyoda leg. LFN (KUM) • 2 ♂♂, 1 ♀, 12 exs.; same locality; 30. VII. 1999; K. Toyoda leg. LFN (KUM) • 1 ♂, 1 ex.; same locality; 30. VII. 2000; (NHM) . Yamanashi-ken • 2 exs.; Kissawa [35.7219, 138.5376], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.5376&amp;materialsCitation.latitude=35.7219" title="Search Plazi for locations around (long 138.5376/lat 35.7219)">Kai-shi</a>; 31. V. 2025; M. Maruyama leg. LFN (KUM) • 1 ex.; Gozaishi-kôsen (alt. 900 m); Nirasaki-shi; 25. VII. 2002; S. Nomura leg. LFN (KUM) . Nagano-ken • 1 ♂, 3 ♀♀, 3 exs.; Mt. Iwatakeyama, Hakuba-mura; 16. VI. 2001; T. Watanabe leg. LFN (KUM) • 1 ♂, 1 ♀, 6 exs.; Midarebashi, Chikuhoku-mura; 21. VII. 1999; T. Watanabe leg. (KUM) • 1 ex.; Shiozawa [36.3268, 138.5983], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.5983&amp;materialsCitation.latitude=36.3268" title="Search Plazi for locations around (long 138.5983/lat 36.3268)">Karuizawa-machi</a>; 5. VI. 2024; K. Goino leg. LFN (KUM) • 1 ex.; Nagakura, Karuizawa-machi; 12. V. 2024; S. Sakurai leg. LFN (KUM) • 1 ♂, 1 ♀, 5 exs.; Jôyama, Matsumoto-shi; 12–13. VII. 1998; M. Maruyama leg. LFN (KUM) • 1 ♂, 5 exs.; Satoyamabe [36.2457, 138.0096], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.0096&amp;materialsCitation.latitude=36.2457" title="Search Plazi for locations around (long 138.0096/lat 36.2457)">Matsumoto-shi</a>; 12. V. 2021; TS. Nozaki leg. LFN (KUM) • 1 ex.; same locality; 30. VII. 2021 TS. Nozaki leg. LFN (KUM) • 1 ex.; Kamimaki [35.85, 137.98], Ina-shi; 1–4. VII. 2021; T. Hashizume leg. LFN (KUM) • 1 ex.; same locality 26. VI. 2022; I. Hashizume leg. (KUM) .</p><p>Diagnosis.</p><p>This species is distinguished from other species of the genus by the following combination of characteristics: body blackish, polished; pronotal postero-lateral angle acute (Fig. 7 B, C); elytra distinctly widened posteriad entirely (Fig. 7 B); pronotum and elytra covered with shortened setae (Fig. 7 B, C); metaventral process hardly produced; male 8 th sternite (Fig. 7 E) slightly emarginated at the middle of the posterior margin; median lobe of aedeagus bulbous; basal part of copulatory piece large. This species is similar to H. gigantea sp. nov. in body size and shape but can be distinguished by its blackish color and shortened setae on the pronotum and elytra.</p><p>Redescription.</p><p>Body (Figs 3 B, 8 G, H) medium-sized, broad, somewhat limuloid; dorsal surface mostly polished.</p><p>Head small; color blackish brown; eyes small. Antennae (Fig. 7 A) short, gently clubbed; relative length of each antennal segment from first to 11 th, 3.2: 1.95: 1.48: 1.18: 1.14: 1.00: 1.00: 1.11: 1.43: 1.64: 4.5.</p><p>Thorax. Pronotum (Fig. 7 B) subrectangular, transverse (PW / PL, 1.53–1.57), widest at posterior 1 / 3, postero-lateral angle distinctly acute, posterior margin distinctly sinuate, blackish brown, surface finely covered with shortened setae and punctures, polished and hardly reticulated. Elytra (Fig. 7 B, C) distinctly widened posteriad entirely, posterior margins shallowly notched near lateral corners, blackish brown, surface finely covered with shortened setae and punctures, polished and hardly reticulated. Hind wings developed. Mesoventral process (Fig. 7 D) very narrow, with indistinct medial carina, apex sharply pointed, reaching middle of mesocoxal cavities. Metaventral process hardly produced.</p><p>Abdomen elongate, narrowed posteriad; surface covered with very short setae and each posterior margin with long stout setae; slightly reticulated.</p><p>Male: 8 th sternite (Fig. 7 E) slightly emarginated at postero-median margin; median lobe of aedeagus (Fig. 7 G) bulbous, length about 0.60 mm, apical lobe thick and almost straight, apex gently pointed in lateral view, apical valves broad and mitten-shaped, dorsal arms very short, basal part of copulatory piece large. Paramere (Fig. 7 H) length about 0.68 mm; apical lobe of paramere long and narrow, length about 1 / 3 of whole paramere, slightly curved ventrally; four setae, normally arranged, velum normal.</p><p>Female: 8 th sternite (Fig. 7 F) slightly rounded at posterior margin; spermatheca (Fig. 7 I, J) S-shaped, apex of basal part weakly swollen.</p><p>Variations.</p><p>In some individuals, the parameral apical lobe is irregularly sharpened apicad. The arrangement of the four setae varies slightly.</p><p>Measurements (in mm) and ratios.</p><p>Body parts (N = 10): BL ≈ 2.44–3.96; AL, 0.91–1.01; HW, 0.52–0.64; PL, 0.6–0.75; PW, 0.94–1.15; EL, 0.48–0.56; EW, 0.96–1.16; HTL, 0.6–0.72; PW / PL, 1.53–1.57; AL / PL, 1.34–1.51; HTL / PL, 0.96–1.03.</p><p>Differences between sexes: AL / PL (male, N = 5), 1.43–1.51; AL / PL (female, N = 5), 1.34–1.42.</p><p>Distribution.</p><p>Japan (Honshu) (Fig. 8 A).</p><p>Symbiotic hosts.</p><p>Lasius fuliginosus species group: L. nipponensis .</p><p>Bionomics.</p><p>Homoeusa laevigata can be observed in Lasius nipponensis foraging trails on forest floors or at the base of trunks (Fig. 8 B) from May to August. No individuals were collected using FIT.</p><p>This species appears to be more integrated into its host colony than other Homoeusa species. Trophallactic behavior with the host ant worker has been observed around nests and foraging trails (Maruyama et al. 2013). In addition, near a colony, TN observed them licking the workers’ legs and bodies as the worker ants groomed each other. In the indoor observations, they frequently raised the tip of their abdomen toward host ant workers (Fig. 8 C), grooming (Fig. 8 E), and displaying trophallaxis (Fig. 8 D, F). Unexpectedly, they were also observed eating food by themselves indoors.</p><p>Remarks.</p><p>The presently known distribution of H. laevigata (Fig. 8 A) is restricted to the east of Hida Mountains in central Honshu. TN examined the collection by Mr. K. Kinomura from some colonies of L. nipponensis in Takayama-shi, Gifu-ken, and surveyed a single colony of the ant in June in Motosu-shi, Gifu-ken. These surveys did not identify H. laevigata west of the Hida Mountains. Considering the absence of this species with Lasius species other than L. nipponensis, and their behavior toward host ant workers, this species could be a specialist for this ant species.</p></div>	https://treatment.plazi.org/id/1C8E87D60BD15CAA97F75475F86EDEA9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Nozaki, Tsubasa;Maruyama, Munetoshi	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
4B546179050E5C999F044986E90FD707.text	4B546179050E5C999F044986E90FD707.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Homoeusa longicornis Sharp 1888	<div><p>Homoeusa longicornis Sharp, 1888</p><p>Figs 2 C, F, 3 D, 11 A – K, 12 A – F (Japanese name: Higenaga-hirata-ariyadori)</p><p>Homoeusa longicornis Sharp, 1888: 283 (original description); Schülke and Smetana 2015; 682 (catalogue); Maruyama et al. 2013: 25 (distribution and bionomics mentioned).</p><p>Type material.</p><p>Holotype (Fig. 2 C, F) • ♂; “ Homoeusa ” / “ longicornis ” / “ D. S. Type ” (written on the card which the specimen mounted) // “ Japan. ” / “ G. Lewis. ” / “ 1910–320 ” // “ Sapporo. ” / “ 5. VIII. – 16. VIII. 80. ” // “ Type ” (red round curator label) // “ HOLOTYPE ” / “ Homoeusa ” / “ longicornis ” / “ SHARP, 1888 ” / “ det. MARUYAMA, 2003 ” // “ NHMUK 014379776 ” (dissected by MM) (NHM).</p><p>Additional material examined.</p><p>Japan: Hokkaido • 2 exs.; Hebinuma, Teshio-gawa, Teshio-chô; 9. VII. 1992; S. Hori leg. (KUM) • 2 ♂♂; Mukôengaru, Engaru-chô; 25. V. 2000; Y. Kida leg. (KUM) • 3 exs.; Kanayama, Makoi, Shari-chô; 24. VI. 2002; Y. Kida leg. (KUM) • 3 exs.; Taihei, Maruseppu, Engaru-chô; 28. V. 2000; Y. Kida leg. LFcfF (KUM) • 22 exs.; same locality; 29–31. V. 2000; Y. Kida leg. LFcfF (KUM) • 11 exs.; same locality; 2–3. VI. 2000; Y. Kida leg. LFcfF (KUM) • 1 ex.; same locality; 13. VI. 2000; Y. Kida leg. (KUM) • 6 ♂♂, 5 ♀♀, 288 exs.; same locality; 16–17. VI. 2000; Y. Kida leg. LFcfF (KUM) • 1 ♂, 7 exs.; same locality; 19–21. VIII. 2000; Y. Kida leg. LFcfF (KUM) • 2 ♂♂, 1 ♀, 17 exs.; same locality; 25. VIII. 2000; Y. Kida leg. LFcfF (KUM) • 1 ex.; Memanbetsu, Ôzora-chô; 7. VIII. 2018; Y. Yoshimura leg. LFcfF (KUM) • 1 ex.; Kashiwa-gaoka-kôen, Bihoro-chô, 23. VI. 2000; Y. Yamaga leg. (KUM) • 1 ex.; Mitomi-shizenkôen, Bihoro-chô; 13. VI. 2000; Y. Yamaga leg. (KUM) • 2 exs.; Attoko, Nemuro-shi; 30. VI. 1996; Y. Hirama leg. (KUM) • 5 exs.; Ôsawaguchi, Nopporo-shinrin-kôen, Ebetsu-shi; 2. VI. 1999; M. Maruyama leg. (KUM) • 26 exs.; Ôsawaguchi, Nopporo-shizenkôen, Ebetsu-shi; 16–19. VI. 2001; S. Hori leg. (KUM) • 2 exs.; Mt. Maruyama, Sapporo-shi; 6. VI. 1998; M. Maruyama leg. (KUM) • 2 exs.; Tomanbetsu, Nopporo, Ebetsu-shi; 29. V. 2002; M. Maruyama leg. LFcfF (KUM) • 2 exs.; Shibumbersu, Nopporo-shinrin-kôen, Ebetsu-shi; 11. V. 2000; M. Maruyama leg. LFcfF (KUM) • 1 ex.; Mt. Moiwa, Sapporo-shi, 16. VI. 1983; Y. Hirano leg. (KUM) • 10 exs.; Hitsujigaoka, Sapporo-shi; 18. V. 2000; M. Maruyama leg. LFS (KUM) • 24 exs.; Kannon-zawa, Sapporo-shi; 1. VI. 2002; M. Maruyama leg. LFcfF (KUM) • 1 ♂, 2 ♀♀, 24 exs.; same locality; 31. VI. 2002; M. Maruyama leg. LFcfF (KUM) • 2 exs.; Hyakumatsuzawa, Sapporo-shi; 8. VI. 1998; M. Maruyama leg. LFcfF (KUM) • 4 exs.; Hakken-zan, Sapporo-shi; 31. V. 2002; M. Maruyama leg. LFcfF &amp; LU (KUM) • 3 exs.; same locality; 1. VI. 2002; M. Maruyama leg. LFcfF &amp; LU (KUM) • 1 ex.; Yûtoku, Ôtaki-mura, Date-shi; 24. VIII. 2001; M. Maruyama leg. (KUM) . Honshu: Aomori-ken • 1 ex.; Yokonagare-tôge (alt. 300 m) [41.264, 141.314], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.314&amp;materialsCitation.latitude=41.264" title="Search Plazi for locations around (long 141.314/lat 41.264)">Higashidori-mura</a>; 26. VI. 2023; TS. Nozaki leg. LFcfF (KUM) • 1 ♂, 1 ♀, 1 ex.; Kuromori (alt. 280 m) [40.677, 140.191], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=140.191&amp;materialsCitation.latitude=40.677" title="Search Plazi for locations around (long 140.191/lat 40.677)">Ajigasawa-machi</a>; 24. VI. 2023; TS. Nozaki leg. LFcfF (KUM) • 1 ♀; Yunosawa, Tokiwano (alt. 490 m) [40.633, 140.268], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=140.268&amp;materialsCitation.latitude=40.633" title="Search Plazi for locations around (long 140.268/lat 40.633)">Hirosaki-shi</a>; 25. VI. 2023; TS. Nozaki leg. LFcfF (KUM) . Miyagi-ken • 1 ♀; Toriido-yama, Komono-chô; 2. VI. 2012; N. Tsuji leg. (KUM) . Yamagata-ken • 1 ♂, 1 ex.; Senjuin, Yamadera (alt. 300–320 m) [38.319, 140.448], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=140.448&amp;materialsCitation.latitude=38.319" title="Search Plazi for locations around (long 140.448/lat 38.319)">Yamagata-shi</a>; 15. VII. 2024; K. Mizota leg. LFcfF (KUM) . Fukushima-ken • 4 exs.; Kashi Spa., Nishigo-mura; 16. VI. 1998; M. Maruyama leg. LFcfF (KUM) • 1 ♀, 3 exs.; same locality; 17. VI. 1998; M. Maruyama leg. LFcfF (KUM) • 1 ex.; Yukiwari-bashi, Nishigo-mura; 29. VII. 2000; M. Maruyama leg. (KUM) . Ibaraki-ken • 1 ex.; Sadanami, Sekimoto-chô, Kitaibaraki-shi; 4. V. 2022; Y. Komatsuzaki leg. (KUM) . Gunma-ken • 1 ex.; Tanigawa Spa, Minakami-chô; 7. VI. 2001; T. Watanabe leg. (KUM) • 1 ex.; Near Shima, Nakanojo-machi; 12. VI. 1986; T. Katsumoto leg. (SCM) • 2 ♂♂, 1 ♀, 4 exs.; Nakanojo Forest Park, Nakanojo-machi; 8. VI. 2001; T. Watanabe leg. (KUM) • 10 exs.; Sakurayama Park, Onishi, Fujioka-shi; 9. V. 1998; Shiho Arai leg. LFS (KUM) • 4 exs.; Mt. Sakurayama, Onishi, Fujioka-shi; 9. V. 1998; K. Toyoda leg. LFS (KUM) • 3 exs.; same locality; 18. V. 1998; K. Toyoda leg. LFS (KUM) • 2 ♂♂, 3 ♀♀, 3 exs.; same locality; 22. V. 1999; Shiho Arai leg. LFS (KUM) • 1 ex.; Shiozawa-toge, Fujioka-shi, 6. IX. 2001; T. Watanabe leg. (KUM) . Saitama-ken • 1 ex.; Mt. Sakurayama, Onishi; 2. V. 2009; K. Arai leg. (KUM) • 2 exs.; Mt. Kamafuse yama, Fu-uppu (alt. 570 m), Yorii-machi; 22. IV. 2006; K. Arai leg. LFcfF (KUM) • 1 ex.; Chichibu-kôgen, Higashichichibu-mura; 3. VII. 1999; K. Toyoda leg. (KUM) • 6 exs.; Kasayama Sancho, Koshigoe, Ogawa-machi; 30. IV. 2005; K. Arai leg. LFS (KUM) • 2 exs.; Karibazaka Pass [35.9683, 139.1816], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=139.1816&amp;materialsCitation.latitude=35.9683" title="Search Plazi for locations around (long 139.1816/lat 35.9683)">Tokigawa-machi</a>; 24. V. 2022; TS. Nozaki leg. LFcfF (KUM) • 2 exs.; Hashidate, Chichibu-shi; 26. IV. 1998; K. Toyoda leg. LFS (KUM) • 1 ex.; same locality; 26. IV. 1998; S. Arai leg. (KUM) • 3 exs.; same locality; 7. VI. 2002; K. Toyoda leg. LFM (KUM) • 1 ex.; Koakazawa, Irikawa, Mt. Hakutai-san, Ootaki (alt. 1,100 m), Chichibu-shi; 7–8. VI. 2008; K. Arai leg. (KUM) . Tokyo-to • 4 exs.; Mt. Takao, Hachiôji-shi; 1. V. 1985; S. Kubota leg. (KUM) • 2 exs.; Mt. Takaozan, Hachiôji-shi; 26. IV. 1986; A. Izumi leg. LFS (KUM) • 2 exs.; Mt. Takao, Hachiôji-shi; 1. VII. 1998; M. Maruyama leg. LFS (KUM) • 1 ex.; same locality; 4. VII. 1998; M. Maruyama leg. LFS (KUM) • 1 ex.; Mt. Takao (alt. 500 m), Hachiôji-shi; 1–4. VII. 1998; M. Maruyama leg. (KUM) • 1 ♂, 1 ♀, 12 exs.; Mt. Takao (alt. 450 m), Hachiôji-shi; 4. VI. 2001; M. Maruyama leg. (KUM) • 1 ♂, 1 ex.; same locality, Hachiôji-shi; 4. VI. 2001; M. Maruyama leg. LFS (NHM) . Kanagawa-ken • 2 exs.; Hokizawa, Nishi-tanzawa; 25–26. VI. 1983; Y. Hirano leg. (KUM) • 1 ex.; Minami-Ashigara-shi; 11. II. 1975; Y. Hirano leg. (KUM) • 2 exs.; Mt. Daiyûzan, Minami-Ashigara-shi; 28. VI. 1987; A. Izumi leg. LFcfF (KUM) • 1 ex.; Daiyuzan, Hakone; 26. V. 1986; Y. Shibata leg. (KUM) • 1 ex.; Daiyuzan, Ashigara; 21. V. 1989; Y. Shibata leg. (KUM) • 2 exs.; Hakone; 4. V. 1987; M. Tao leg. (KUM) . Niigata-ken • 1 ex.; Ochinomizu, Tokamachi-shi; 19. IV. 1998; T. Kishimoto leg. LFcfF (KUM) • 1 ex.; Foot of Mt. Uedafuji, Yuzawa-machi; 18. IV. 1998; H. Yoshitomi leg. (KUM) • 1 ex.; Yuzawa-machi; 20–22. VI. 2002; S. Nagashima leg. LFcfF (KUM) . Yamanashi-ken • 1 ♂; Gozaishi-kôsen (alt. 900 m), Nirasaki-shi; 25. VII. 2002; S. Nomura leg. LFN (KUM) • 3 exs.; Ashiyu-onsen, Ashiyu-mura; 23. V. 2001; T. Watanabe leg. (KUM) • 1 ex.; same locality; 24. V. 2001; T. Watanabe leg. (KUM) . Nagano-ken • 1 ex.; Midarebashi, Chikuhoku-mura; 21. VII. 1999; T. Watanabe leg. (KUM) • 1 ♂, 1 ex.; Ôkubo, Komoro-shi; 29–30. IV. 2001; H. Kamezawa leg. LFcfF (cKam) . Gifu-ken • 2 ♂♂; Bijo Pass, Takayama-shi; 5. VIII. 2003; K. Kinomura leg. (KUM) • 1 ♂, 1 ex.; Hirugano Bunsuirei Park, Gujo-shi; 10. VI. 2013; K. Kinomura leg. (KUM) • 1 ♂; Mt. Kentô, Takayama-shi; 6. VII. 2003; K. Kinomura leg. (KUM) • 2 ♂♂; Yôrô Park, Yôrô-chô; 29. IV. 2004; K. Kinomura leg. (KUM) . Shizuoka-ken • 2 exs.; Utsuno, Fujinomiya; 30. V. 2000; T. Watanabe leg. Fish feed Bait, pinned with a Phidole ant det. by K. Matuura (KUM) . Aichi-ken • 1 ♀; Jôkôji-kôen, Seto-shi; 16. IV. 2023; T. Tenma leg. (KUM) . Kyoto-fu • 1 ♂, 1 ex.; Tenno-zan, Oyamazaki-chô; 16. IV. 2008; T. Ito leg. (OMNH) • 2 exs.; same locality; 31. V. 2008; T. Ito leg. (OMNH) • 1 ♀; Amagase Forest, Uji-shi; 8. V. 2013; T. Ito leg. (OMNH) • 1 ex.; Otokoyama, Yawata-shi; 11. V. 2014; T. Ito leg. (OMNH) • 1 ♂; same locality; 14. VI. 2008; T. Ito leg. (OMNH) . Osaka-fu • 1 ♂; Mt. Mikusa, Nose-chô; 21. VI. 2008; T. Ito leg. (OMNH) • 1 ♂; Mino-o Park, Mino-shi; 28. V. 2008; T. Ito leg. (OMNH) . Hyogo-ken • 2 ♂♂, 1 ♀, 1 ex.; Dojo, Kobe-shi; 13. IV. 2009; T. Ito leg. (OMNH) . Nara-ken • 1 ♀; Yamato-Koriyama, Yata; 9. VII. 2009; T. Ito leg. (OMNH) • 1 ♂; same locality; 18. V. 2009; T. Ito leg. (OMNH) . Wakayama-ken • 1 ♂; Kimitoge Pass, Hashimoto-shi; 1. VI. 2009; T. Ito leg. (OMNH) • 1 ♂; same locality; 27. IV. 2014; T. Ito leg. (OMNH) . Tottori-ken • 1 ex.; Mt. Daisen, Daisen-chô; 3–5. VI. 1980; S. Naomi leg. (KUM) . Shimane-ken • 1 ♂; Urahikimi, Masuda-shi, Masuda-shi; 6. VI. 1998; S. Nomura leg. LFS (KUM) . Hiroshima-ken • 1 ♂; Mt. Iwahi (alt. 1,110 m) [35.0712, 133.2192], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=133.2192&amp;materialsCitation.latitude=35.0712" title="Search Plazi for locations around (long 133.2192/lat 35.0712)">Shobara-shi</a>; 16. VI. 2024; TS. Nozaki leg. LFcfF (KUM) . Shikoku: Tokushima-ken • 2 exs.; Takashiro-yama, Naka-chô; 24. V. 2007; H. Fujimoto leg. LFS (KUM) . Kagawa-ken • 1 ex.; Ôtawa, Nagao, Sanuki-shi; 22. V. 2000; F. Ito leg. (KUM) • 2 exs.; Daisen-zan, Kotonami-chô; 1. VI. 2001; M. Maruyama et al. leg. LFN (KUM) • 1 ex.; same locality; 3. V. 2004; H. Fujimoto leg. LFcfF (KUM) . Ehime-ken • 1 ♂, 1 ♀; Mt. Ishiduchi, Saijo-shi; 29. VI. 2014; Y. Hisasue leg. (KUM) . Kochi-ken • 1 ♂, 1 ♀; Hokigamine (alt. 500 m) [33.6719, 133.6846], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=133.6846&amp;materialsCitation.latitude=33.6719" title="Search Plazi for locations around (long 133.6846/lat 33.6719)">Kami-shi</a>; 11. V. 2023; TS. Nozaki leg. LFS (KUM) .</p><p>Diagnosis.</p><p>This species is distinguished from the other species of the genus by the following combination of characteristics: pronotal posterolateral angle around 90 degrees (Fig. 11 C); elytra distinctly widened entirely posteriad (Fig. 11 C, D); apical lobe of paramere very long, length over half of whole paramere, curved ventrally, apex blade-shaped (Fig. 11 I). This species is quite similar to H. japonica in its general appearance but can be separated by the blade-like apex of its parameral apical lobe.</p><p>Redescription. Body (Figs 3 D, 12 F) large, broad, stout; dorsal surface mostly moderately polished.</p><p>Head large; color reddish brown to dark brown; eyes small. Antennae (Fig. 11 A, B) very long.</p><p>Thorax. Pronotum (Fig. 11 C) subrectangular, transverse (PW / PL, 1.53–1.64), widest anterior 1 / 3, postero-lateral angle around 90 degrees, posterior margin slightly sinuate, reddish yellow to reddish brown, surface finely covered with setae and punctures, gently polished and slightly reticulated. Elytra (Fig. 11 C, D) distinctly widened posteriad entirely, posterior margins shallowly notched near lateral corners, reddish yellow to reddish brown, surface finely covered with setae and punctures; gently polished and weakly reticulated. Hind wings developed. Mesoventral process (Fig. 11 E) very narrow, with weak medial carina, forming Y-shaped, apex sharply pointed, reaching posterior 1 / 3 of mesocoxal cavities. Metaventral process weakly produced.</p><p>Abdomen elongate, narrowed posteriad; surface covered with short setae and each posterior margin with long stout setae; finely reticulated.</p><p>Male: Relative length of each antennal segment from first to 11 th, 1.66: 1.13: 1.16: 1.07: 1.00: 1.04: 1.00: 1.06: 1.01: 1.13: 2.25. 8 th sternite (Fig. 11 F) weakly rounded at posterior margin; median lobe of aedeagus (Fig. 11 H) slender, length about 0.63 mm, apical lobe elongated, ventral side slightly humped at middle, apex gently pointed in lateral view, apical valves simple and very long, without dorsal arms, basal part of the copulatory piece very small. Paramere (Fig. 11 I) length about 0.79 mm; apical lobe of paramere extremely long, length over half of whole paramere, curved ventrally, dilated apicad, apex blade-shaped; four setae, a far longer than others, b located most apically, velum elongate, broad and produced apically.</p><p>Female: Relative length of each antennal segment from first to 11 th, 2.37: 1.57: 1.29: 1.09: 1.06: 1.00: 1.06: 1.08: 1.17: 1.40: 2.88. 8 th sternite (Fig. 11 G) truncated or slightly rounded at posterior margin; spermatheca (Fig. 11 J, K) S-shaped, apex of basal part not swollen, base largely hooklike.</p><p>Variation.</p><p>Apical lobe of aedeagal median lobe varies on the degree of dorsally curve, intension of middle hump ventral side, and depth of basal concave (Fig. 12 A – C) independently of geographical locations.</p><p>Measurements (in mm) and ratios.</p><p>Body parts (N = 12): BL ≈ 2.58–3.54; AL, 0.98–1.3; HW, 0.59–0.7; PL, 0.59–0.72; PW, 0.97–1.13; EL, 0.5–0.59; EW, 0.94–1.13; HTL, 0.65–0.8; PW / PL, 1.53–1.64; AL / PL, 1.5–1.91; HTL / PL, 0.98–1.13.</p><p>Differences between sexes: AL / PL (male, N = 5), 1.82–1.91; AL / PL (female, N = 7), 1.5–1.65; HTL / PL (male, N = 5), 1.05–1.13; HTL / PL (female, N = 7), 0.98–1.04.</p><p>Distribution.</p><p>Japan (Hokkaido, Honshu, Shikoku) (Fig. 12 D).</p><p>Symbiotic hosts.</p><p>Lasius fuliginosus species group: L. cf. fuliginosus, L. morisitai, L. nipponensis, L. orientalis, L. spathepus .</p><p>Bionomics.</p><p>Homoeusa longicornis can be observed in Lasius fuliginosus species group foraging trails on the forest floor from April to August (Fig. 12 E). To the best of our knowledge, no individual was collected using FIT. The beetles were observed to be frequently raising the tip of their abdomen when approaching the host ant workers in the field and indoors.</p><p>Remarks.</p><p>This species is rarely collected from the trails of L. nipponensis and L. orientalis . This species is very similar to H. japonica in terms of its habitus (Fig. 12 F) and bionomics. H. longicornis tends to occur at higher altitudes than that of H. japonica . However, both species frequently co-occur at moderate altitudes in Honshu and Shikoku, even within the same colony of host ants. Precisely distinguishing between the two species is challenging without examining male genitalia. The pronotum length can be helpful; that of H. longicornis is usually longer than 0.65 mm, whereas that of H. japonica is shorter than 0.65 mm, but some exceptions remain in both species.</p></div>	https://treatment.plazi.org/id/4B546179050E5C999F044986E90FD707	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Nozaki, Tsubasa;Maruyama, Munetoshi	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
49EEBEA60488591F94B6A0D4266766A7.text	49EEBEA60488591F94B6A0D4266766A7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Homoeusa ovata Nozaki & Maruyama 2022	<div><p>Homoeusa ovata Nozaki &amp; Maruyama, 2022</p><p>Fig. 6 D – F, H (Japanese name: Hime-hirata-ariyadori)</p><p>Homoeusa ovata Nozaki and Maruyama 2022 b: 1 (provideing depository); Nozaki and Maruyama 2022 a: 48 (detailed description but unavailable).</p><p>“ Homoeusa sp.: ” Maruyama et al. 2013: 24 (distribution and bionomics mentioned).</p><p>Additional material from Japan.</p><p>Hokkaido • 1 ex.; Mt. Moiwa (alt. 520 m), Sapporo-shi; 27. VII. 1991; A. Smetana leg. (NMST, ex A. Smetana coll.) • 5 exs.; Memanbetsu, Ôzora-chô; 7. VIII. 2018; Y. Yoshimura leg. LFcfF (KUM) . Honshu: Aomori-ken • 1 ex.; Yokonagare-tôge (alt. 300 m) [41.264, 141.314], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.314&amp;materialsCitation.latitude=41.264" title="Search Plazi for locations around (long 141.314/lat 41.264)">Higashidori-mura</a>; 26. VI. 2023; TS. Nozaki leg. (KUM) . Iwate-ken • 8 exs.; Gandô-ko, Morioka-shi; 9. VI. 2019; N. Nakaya leg. LFcfF (IPMM) • 1 ex.; Kitayama-sansakuro, Morioka-shi; 31. VII. 2019; N. Nakaya leg. LFcfF (IPMM) . Yamagata-ken • 1 ex.; Senjuin, Yamadera (alt. 300–320 m) [38.319, 140.448]; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=140.448&amp;materialsCitation.latitude=38.319" title="Search Plazi for locations around (long 140.448/lat 38.319)">Yamagata-shi</a>; 15. VII. 2024; K. Mizota leg. LFcfF (KUM) . Ibaraki-ken • 2 exs.; Asahi-tôge, Tsuchiura-shi; 8. V. 2022; Y. Komatsuzaki leg. (KUM) . Saitama-ken • 1 ex.; Ushidatekubo (alt. 830 m) [35.9685, 139.1793], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=139.1793&amp;materialsCitation.latitude=35.9685" title="Search Plazi for locations around (long 139.1793/lat 35.9685)">Tokigawa-machi</a>; 21. V. 2023; S. T. (Shota) Inoue &amp; N. Katsube leg. LFcfF (KUM) . Kanagawa-ken • 1 ex.; Kowakien, Mt. Hakone, Hakone-chô; 22. V. 1963; R. Kiryu leg. (SCM) • 1 ex.; Toya, Tsukui, Sagamihara-shi; 11. IV. 1976; R. Kiryu leg. (SCM) • 2 exs.; Nanzawa, Tsukui, Sagamihara-shi; 14. IV. 1977; R. Kiryu leg. (SCM) . Niigata-ken • 2 exs.; Sado Daira Ski Resort, Shinbo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.35&amp;materialsCitation.latitude=38.076" title="Search Plazi for locations around (long 138.35/lat 38.076)">Kanai</a> (alt. 390 m) [38.076, 138.350], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.35&amp;materialsCitation.latitude=38.076" title="Search Plazi for locations around (long 138.35/lat 38.076)">Sado-shi</a>; 19. VI. 2023; TS. Nozaki &amp; W. R. Liang leg. LFN (KUM) • 1 ex.; Sado Daira Ski Resort, Shinbo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.35&amp;materialsCitation.latitude=38.075" title="Search Plazi for locations around (long 138.35/lat 38.075)">Kanai</a> (alt. 380 m) [38.075, 138.350], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.35&amp;materialsCitation.latitude=38.075" title="Search Plazi for locations around (long 138.35/lat 38.075)">Sado-shi</a>; 20. VI. 2023; TS. Nozaki &amp; W. R. Liang leg. LFO (KUM) • 2 exs.; Sado Daira Ski Resort, Shinbo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.355&amp;materialsCitation.latitude=38.075" title="Search Plazi for locations around (long 138.355/lat 38.075)">Kanai</a> (alt. 380 m) [38.075, 138.355], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.355&amp;materialsCitation.latitude=38.075" title="Search Plazi for locations around (long 138.355/lat 38.075)">Sado-shi</a>; 20. VI. 2023; TS. Nozaki &amp; W. R. Liang leg. LFcfF (KUM) . Nagano-ken • 1 ex.; Bôdaira, Shiga-kôgen (alt. 1,160 m) [36.7203, 138.4656], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.4656&amp;materialsCitation.latitude=36.7203" title="Search Plazi for locations around (long 138.4656/lat 36.7203)">Yamanouchi-machi</a>; 9. VII. 2022; TS. Nozaki leg. LFM (KUM) • 2 exs.; Satoyamabe [36.2457, 138.0096], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.0096&amp;materialsCitation.latitude=36.2457" title="Search Plazi for locations around (long 138.0096/lat 36.2457)">Matsumoto-shi</a>; 12. V. 2021; TS. Nozaki leg. LFN (KUM) . Gifu-ken • 2 exs.; Mt. Kentô, Takayama-shi; 6. VII. 2003; K. Kinomura leg. (KUM) • 2 exs.; Bijo Pass, Takayama-shi; 5. VIII. 2003; K. Kinomura leg. (KUM) • 1 ex.; Isshiki, Takayama-shi; 23. V. 2004; K. Kinomura leg. (KUM) • 1 ex.; Nabedaira-kôgen, Takayama-shi; 30. VI. 2013; K. Kinomura leg. (KUM) • 2 exs.; Hirugano Bunsuirei Park, Gujo-shi; 10. VI. 2013; K. Kinomura leg. (KUM) • 1 ex.; Yôrô Park, Yôrô-chô; 29. IV. 2004; K. Kinomura leg. (KUM) • 2 exs.; Mitahora, Gifu-shi; 10. VI. 2013; K. Kinomura leg. (KUM) . Shizuoka-ken • 2 exs.; Kumazaka, Izu-shi; 30. V. 2022; T. Tenma leg. (KUM) . Aichi-fu • 2 exs.; Jôkôji-kôen, Seto-shi, 4. V. 2023; T. Tenma leg. (KUM) . Osaka-fu • 3 exs.; Sakurai [34.8828, 135.6525], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=135.6525&amp;materialsCitation.latitude=34.8828" title="Search Plazi for locations around (long 135.6525/lat 34.8828)">Shimamoto-chô</a>; 9. V. 2021; Y. Takatani leg. LFS (KUM) . Shikoku: Kochi-ken • 1 ex.; Ashizuri-misaki (alt. 120 m) [32.7480, 133.0136], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=133.0136&amp;materialsCitation.latitude=32.748" title="Search Plazi for locations around (long 133.0136/lat 32.748)">Tosashimizu-shi</a>; 21. V. 2024; TS. Nozaki leg. (KUM) . Kyushu: Miyazaki-ken • 1 ex.; Ômoridake forest road (alt. 600 m) [32.0435, 131.1973], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=131.1973&amp;materialsCitation.latitude=32.0435" title="Search Plazi for locations around (long 131.1973/lat 32.0435)">Aya-machi</a>; 12. VI. 2024; TS. Nozaki &amp; T. Kinoda leg. LFS (KUM) .</p><p>Diagnosis.</p><p>See Nozaki and Maruyama (2022 a).</p><p>Additional description.</p><p>Relative length of each antennal segment from first to 11 th, 2.31: 1.98: 1.48: 1.29: 1.17: 1.00: 1.14: 1.14: 1.26: 1.4: 3.36. Hindwings developed.</p><p>Male. 8 th sternite (Fig. 6 D) gently round at posterior margin; apical lobe of paramere (Fig. 6 F), setae b and c irregularly distributed at the acute corner on dorsal side.</p><p>Female. 8 th sternite (Fig. 6 E) gently round at posterior margin.</p><p>Distribution.</p><p>Russia (Primorskyi krai), Korea, Japan (Hokkaido, Honshu, Shikoku, Kyushu) (Fig. 6 H).</p><p>Symbiotic hosts.</p><p>Lasius fuji, L. cf. fuliginosus, Lasius morisitai, L. nipponensis, L. orientalis, L. spathepus .</p></div>	https://treatment.plazi.org/id/49EEBEA60488591F94B6A0D4266766A7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Nozaki, Tsubasa;Maruyama, Munetoshi	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
B8B9331D8F9F55778EA30E260A3174A6.text	B8B9331D8F9F55778EA30E260A3174A6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Homoeusa prolongata Sawada 1970	<div><p>Homoeusa prolongata Sawada, 1970</p><p>Figs 3 E, 13 A – I, 14 A – D (Japanese name: Hirata-ariyadori)</p><p>Homoeusa prolongata Sawada, 1970 b: 57, fig. 14 (original description); Schülke and Smetana 2015; 682 (catalogue); Maruyama et al. 2013: 26 (distribution and bionomics mentioned).</p><p>Type material.</p><p>Not examined.</p><p>Additional material examined.</p><p>Japan: Hokkaido • 2 exs.; Near Hime-numa, Oshidomariwannai (alt. 130 m), Rishiri-chô; 5. VI. 2024; T. Yamauchi leg. (OUAVM) • 1 ex.; “ Horokanai Butokamabetsu Riv ” (alt. 300 m); 3. VIII. 1991; A. Smetana leg. (NMST, ex A. Smetana coll.) • 1 ex.; Kanayama, Makoi, Shari-chô; 24. VI. 2002; Y. Kida leg. (KUM) • 1 ♂, 1 ex.; Teine-yama [43.0836, 141.2024], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.2024&amp;materialsCitation.latitude=43.0836" title="Search Plazi for locations around (long 141.2024/lat 43.0836)">Sapporo-shi</a>; 23. V. 2020; Y. Tasaku leg. (KUM) • 1 ♂; Ôsawaguchi, Nopporo-shizenkôen, Ebetsu-shi; 16–19. VI. 2001; S. Hori leg. (KUM) • 2 exs.; Kannon-zawa, Sapporo-shi; 19. V. 2021; Y. Tasaku leg. (KUM) • 2 exs.; Hakken-zan, Sapporo-shi; 1. VI. 2002; M. Maruyama leg. (KUM) • 1 ♀; Mt. Kariba, Shiribeshi; 12. VI. 1986; S. Nomura leg. (KUM) . Honshu: Aomori-ken • 1 ex.; Iwaki river riverbed, Kawazura (alt. 10 m) [40.694, 140.455], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=140.455&amp;materialsCitation.latitude=40.694" title="Search Plazi for locations around (long 140.455/lat 40.694)">Itayanagi-machi</a>; 28. VI. 2023; TS. Nozaki leg. LNS (KUM) . Iwate-ken • 1 ♀, 1 ex.; Kitayama, Morioka-shi; 23. IV – 3. V. 2024; N. Nakaya leg. FIT (IPMM) • 1 ex.; Mt. Takabora, Kamiyonai, Morioka-shi; 21. IV – 3. V. 2024; N. Nakaya leg. FIT (IPMM) . Akita-ken • 1 ex.; Kawara-bo, Mt. Hachimine, Hanamaki-shi; 22–24. VI. 1980; S. Naomi leg. (KUM) . Fukushima-ken • 1 ex.; Zenjin, Mt. Chokai, Yurihonjo-shi; 19–20. VI. 1980; S. Naomi leg. (KUM) • 1 ex.; Yokomuki, Mt. Adatara, Inawashiro-machi; 10. VII. 1985; S. Nomura leg. (KUM) • 1 ex.; Kozodaira, Hinoemata-mura; 26. VII. 1996; S. Naomi leg. (KUM) • 2 exs.; Nanairi, Hinoemata-mura; 24. VII. 1990; T. Kishimoto leg. (KUM) . Tochigi-ken • 1 ex.; Mt. Nasu; 3. VI. 1994; S. Naomi leg. (KUM) • 2 exs.; Chuzenji, Nikko-shi; 28–30. VI. 1982; S. Naomi leg. (KUM) • 1 ex.; same locality; 24. VII. 1985; S. Nomura leg. (KUM) • 1 ex.; “ Nikko N. P. Lake Chuzenjiko ” (alt. 1,280 m); “ 14. VII. 80 ” [1980]; A. &amp; Z. Smetana leg. (NMST, ex A. Smetana coll.) • 1 ex.; “ Nikko N. P. Senjugahara ” (alt. 1400 m); “ 15. VII. 80 ” [1980]; A. &amp; Z. Smetana leg. (KUM, ex A. Smetana coll.) • 1 ex.; Nagaishi Forest Road, Ashikaga-shi; 27. V. 1990; H. Ohkawa leg. (KUM) • 1 ♂; Watarase-yûsuichi, Fujioka-machi; 16. IV. 2000; H. Ôkawa leg. (KUM) . Gunma-ken • 1 ex.; Mikuni-tôge, Minakami-machi; 27. VI. 1997; S. Naomi leg. (KUM) • 1 ex.; Hôshi-onsen, Minakami-machi; 26. VI. 1997; S. Naomi leg. (KUM) • 1 ex.; Amemi-rindou, Minakami-machi; 4. V. 1998; Shiho Arai leg. (KUM) • 1 ♂, 1 ex.; Sakurayama, Fujioka-shi; 9. V. 1998; Shiho Arai leg. (KUM) • 1 ex.; same locality; 22. V. 1999; K. Toyoda leg. LNH (KUM) • 1 ♀, 3 exs.; same locality; 22. V. 1999; Shiho Arai leg. LNH (KUM) • 1 ex.; same locality; 27. V. 2000; Shiho Arai leg. LNH (KUM) . Saitama-ken • 1 ex.; Mt. Jômine-san, Minano-machi; 29. V. 1999; Shiho Arai leg. LNH (KUM) • 1 ex.; Chichibu-kôgen, Higashichichibu-mura; 5. V. 2000; Shiho Arai leg. (KUM) • 1 ex.; Sugaya, Ranzan-machi; 14. VI. 1994; K. Toyoda leg. LNH (KUM) • 1 ex.; same locality; 16. VI. 1994; K. Toyoda leg. (KUM) • 1 ex.; Iwadono, Higashimatsuyama-shi; 3. V. 2005; K. Toyoda leg. LNJ (KUM) • 1 ex.; Near Tsumasaka-Toge, Hanno-shi; 20. VI. 1998; K. Toyoda leg. (KUM) • 1 ♂; Kawaura-keikoku (alt. 600 m), Chichibu-shi; 20. VI. 1998; Shiho Arai leg. (KUM) • 1 ♀; same locality; 27. VI. 1998; Shiho Arai leg. (KUM) • 1 ex.; Kaminaguri, Hanno-shi; 23. V. 1998; Shiho Arai leg. LNH (KUM) • 1 ex.; Sumiyairi-rindô, Hanno-shi; 3. VI. 2000; Shiho Arai leg. LNH (KUM) . Chiba-ken • 2 exs.; Ashida (alt. 10 m), Narita-shi; 17–25. VII. 2014; R Nakamura leg. FIT (KUM) • 11 exs.; Okuno, Ichihara-shi; 12–14. VII. 2000; S. Hatsushiba &amp; T. Shimada leg. (KUM) • 1 ex.; same locality; 16–18. V. 2001; S. Hatsushiba &amp; T. Shimada leg. (KUM) • 1 ex.; same locality; 12–14. V. 2001; S. Hatsushiba &amp; T. Shimada leg. (KUM) • 1 ex.; Mt. Gundari, Ichinomiya-machi; 12. IV. 1997; S. Nomura leg. (KUM) • 1 ex.; Umegase, Daifuku-yama, Ichihara-shi; 6. V. 1998; S. Naomi leg. (KUM) • 1 ex.; Amatsu (alt. 70 m) [35.1406, 140.1512], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=140.1512&amp;materialsCitation.latitude=35.1406" title="Search Plazi for locations around (long 140.1512/lat 35.1406)">Kamogawa-shi</a>; 3. V. 2016; S. Sejima leg. Litter (KUM) . Tokyo-to • 1 ex.; Ônita, Ome-shi; 20. V. 1991; T. Kishimoto leg. (KUM) • 1 ex.; Oyamada-chô, Machida-shi; 19. V. 1992; T. Kishimoto leg. (KUM) • 1 ex.; Machida-shi; 28. V. 1981; T. Niizato leg. (KUM) . Kanagawa-ken • 3 exs.; Nanzawa, Tsukui, Sagamihara-shi; 6. V. 1977; R. Kiryu leg. (SCM) • 1 ex.; Miyagase, Aikô; 19. IV. 1977; R. Kiryu leg. (SCM) • 2 exs.; Kozukue, Yokohama-shi; 21. IV. 1994; H. Sato leg. (KUM) • 1 ♂; Niiharutyô 、 Midori-ku [35.50, 139.51], Yokohama-shi; 25. IV. 2012; O. Saitoh leg. (KPMNH) • 3 exs.; Trap St. 1. Park / “ Y. N. S ”, Yokohama-shi; 8. VI. 1988; Y. Kominami leg. (SPMN) • 1 ex.; Yakunimi-yama [35.35, 139.17], Hadano-shi; 27. VI. 2014; O. Saitoh leg. (KPMNH) • 1 ex.; Kinugasa, Yokosuka-shi; 26–27. IV. 1995; S. Tsuboyama leg. (KUM) . Niigata-ken • 1 ♂; Kita-ikari (alt. 760 m) [38.1500, 138.4049], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.4049&amp;materialsCitation.latitude=38.15" title="Search Plazi for locations around (long 138.4049/lat 38.15)">Sado-shi</a>; 18. VI. 2023; TS. Nozaki leg. (KUM) • 3 exs.; Mt. Tadara-mine, Ryotsu (alt. 810 m) [38.13, 138.38]; Sado-shi; 11. VI. 2012; M. Yoshida leg. (KUM) • 2 exs.; Yokoyama, Sado-shi; 24–25. V. 1995; T. Kishimoto leg. (KUM) • 1 ex.; Kaikake-onsen, Yuzawa-machi; 23. VI. 1996; K. Haga leg. Leaf litter in Beech forest (KUM) • 1 ex.; Sasagamine, Myoko-shi; 14–15. VI. 1980; S. Naomi leg. (KUM) • 1 ex.; Suginosawa (alt. 950 m) [36.8519, 138.1399], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.1399&amp;materialsCitation.latitude=36.8519" title="Search Plazi for locations around (long 138.1399/lat 36.8519)">Myoko-shi</a>; 3. VI. 2018; S. Sejima leg. Litter (KUM) . Fukui-ken • 1 ex.; Mt. Houonji-san, Katsuyama-shi; 31. V. 1989; Kouichi Matsumoto leg. (KUM) • 1 ex.; Mt. Gongen, Echizen-shi; 22. V. 1981; H. Sasaji leg. (KUM) . Yamanashi-ken • 1 ♀; Daibosatsu-rei [35.73, 138.82], Koshu-shi; 9. VI. 2019; O. Saitoh leg. (KPMNH) • 1 ex.; Ryomata-goya, Noro-gawa riv., Minami-alps-shi; 3. VIII. 1996; K. Haga leg. Fallen branch (KUM) • 5 ♂♂, 1 ♀, 2 exs.; Karumizu-rindo (alt. 1600 m), Narusawa-mura; 22–29. VI. 2011; T. Watanabe leg. FIT (KUM) • 1 ex.; same locality; 29. VI – 6. VII. 2011; T. Watanabe leg. FIT (KUM) • 1 ex.; same locality; 27. VI – 4. VII. 2012; T. Watanabe leg. FIT (KUM) • 1 ex.; same locality; 25. VII – 1. VIII. 2012; T. Watanabe leg. FIT (KUM) • 2 exs.; Subashiri-guchi, Fujisan (alt. 1,400 m), Oyama-chô; 15–20. V. 2011; T. Watanabe leg. FIT (KUM) • 1 ex.; Subashiri, Fujisan 1 - gome (alt. 1,400 m), Oyama-chô; 20–26. V. 2011; T. Watanabe leg. FIT (KUM) . Nagano-ken • 2 exs.; Shiga-kôgen (alt. 1,380 m) [36.728, 138.478], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.478&amp;materialsCitation.latitude=36.728" title="Search Plazi for locations around (long 138.478/lat 36.728)">Yamanouchi-machi</a>; 9. VII. 2022; TS. Nozaki leg. Litter (KUM) • 1 ex.; same locality; 9. VII. 2022; TS. Nozaki leg. LNH (KUM) • 1 ♂; Mizunashiike, Shiga-kôgen, Yamanouchi-machi; 9. VII. 2022; TS. Nozaki leg. (KUM) • 1 ex.; Satoyamabe, Matsumoto-shi; 4. VI. 2009; T. Komatsu leg. LNJ (KUM) • 1 ♂; Tobira, Matsumoto-shi; 5. VI. 2011; M. Maruyama et al. leg. (KUM) • 1 ♀; Kohigashi, Chino-shi; 19. VI. 2011; M. Maruyama et al. leg. (KUM) • 1 ♂; Kannon-daira, Yatsuga-take; 11. VII. 2000; H. Hoshina leg. Beating (KUM) . Shizuoka-ken • 1 ex.; Subashiri-guchi, Fujisan (alt. 1,400 m), Oyama-chô; 15–22. VI. 2011; T. Watanabe leg. FIT (KUM) • 2 exs.; Hiekawa, Izu-shi; 2. VI. 1983; T. Tahira leg. (KUM) . Kyoto-fu • 1 ex.; Mt. Yôrô-san, Maizuru-shi; 25–26. V. 2000; K. Mizuno leg. (KUM) . Hyogo-ken • 1 ex.; Mt. Hyonosen; 5. VI. 1984; S. Nomura leg. (KUM) • 1 ex.; Mt. Mimuro-yama; 7. VI. 1997; H. Hoshina leg. (KUM) . Nara-ken • 1 ex.; Nara; “ 11. VIII. 80 ” [1980]; A. &amp; Z. Smetana leg. (KUM, ex A. Smetana coll.) • 1 ex.; Nishitani, Yoshino-chô; 18–19. V. 1995; Y. Nakahara leg. Bait trap (KUM) • 2 exs.; Mt. Nehan-dake (alt. 1,140 m), Shimokitayama-mura; 27. VI. 1998; T. Kishimoto leg. (KUM) . Wakayama-ken • 1 ex.; Hirai, Kozagawa-chô; 15. VIII. 1999; M. Maruyama leg. LNJ (KUM) • 1 ex.; same locality; 16. VII. 1999; M. Maruyama leg. LNN (KUM) . Tottori-ken • 1 ex.; Hokidaisen, Daisen-chô; 3. VI. 1980; S. Naomi leg. (KUM) • 3 exs.; Mt. Daisen, Daisen-chô; 3–5. VI. 1980; S. Naomi leg. (KUM) • 2 exs.; Ônarubara-kôgen (alt. 770 m), Kôfu-chô; 18–24. V. 2008; A. Watanabe leg. FIT (KUM) • 3 exs.; same locality; 7–23. VI. 2008; A. Watanabe leg. FIT (KUM) • 1 ex.; same locality; 24. V – 1. VI. 2008; A. Watanabe leg. FIT (KUM) . Shimane-ken • 1 ♀; Dangyô-no-taki, Dôgo Is., Okinoshima-chô; 5–16. VI. 2008; T. Shimada leg. (KUM) . Okayama-ken • 1 ex.; Kanehira, Bicchu-chô (alt. 300 m), Takahashi-shi; 22. V – 2. VI. 2010; A. Watanabe leg. FIT (KUM) • 1 ex.; same locality, Takahashi-shi; 2–19. VI. 2010; A. Watanabe leg. FIT (KUM) . Hiroshima-ken • 1 ♂; Mt. Tate-eboshi (alt. 1,180 m) [35.0527, 133.0696], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=133.0696&amp;materialsCitation.latitude=35.0527" title="Search Plazi for locations around (long 133.0696/lat 35.0527)">Shobara-shi</a>; 15. VI. 2024; TS. Nozaki leg. (KUM) • 1 ex.; Kakezu-yama, Higashiyawatahara (alt. 850–1,126 m), Kitahiroshima-chô; 30. IV – 9. V. 2009; TA. Nozaki &amp; Y. Nozaki leg. FIT (KUM) • 2 exs.; same locality; 9–18. V. 2009; TA. Nozaki &amp; Y. Nozaki leg. FIT (KUM) • 1 ex.; same locality; 30. V – 13. VI. 2009; TA. Nozaki &amp; Y. Nozaki leg. FIT (KUM) • 5 exs.; same locality; 13–27. VI. 2009; TA. Nozaki &amp; Y. Nozaki leg. FIT (KUM) • 1 ex.; same locality; 27. VI – 4. VII. 2009; TA. Nozaki &amp; Y. Nozaki leg. FIT (KUM) • 1 ♂, 1 ex.; same locality; 4–18. VII. 2009; TA. Nozaki &amp; Y. Nozaki leg. FIT (KUM) . Yamaguchi-ken • 1 ex.; Nodani, Tokuji-chô; 21. IV. 2004; Y. Fujitani leg. (KUM) . Shikoku: Tokushima-ken • 1 ex.; Meoto-ike, Tsurugi-chô; 30. VI. 2006; H. Fujimoto leg. (KUM) . Kagawa-ken • 2 exs.; Nagara-dam, Ayakami-chô; 5. V. 2003; S. Nagashima leg. in litter (KUM) • 2 exs.; Shôzaka, Sogisho-higashi, Ayagawa-chô; 9–12. IV. 2009; H. Fujimoto leg. FIT (KUM) • 1 ex.; same locality; 12–18. IV. 2009; H. Fujimoto leg. FIT (KUM) • 1 ex.; same locality; 22–30. V. 2009; H. Fujimoto leg. FIT (KUM) • 1 ex.; same locality; 25. IV – 2. V. 2010; H. Fujimoto leg. FIT (KUM) . Kochi-ken • 1 ex.; Mt. Tsutsujô-zan (alt. 1,600 m), Ino-chô; 30. VI. 2007; TA. (Takasuke) Miyata &amp; TO. (Toshie) Miyata leg. (KUM) • 1 ex.; Ekawa, Nishitosa (alt. 120 m) [33.217, 131.784], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=131.784&amp;materialsCitation.latitude=33.217" title="Search Plazi for locations around (long 131.784/lat 33.217)">Shimanto-shi</a>; 12. V. 2023; TS. Nozaki leg. (KUM) . Kyushu: Fukuoka-ken • 1 ♂; Mt. Inunaki, Miyawaka-shi; 22. V. 1980; T. Gotô leg. (KUM) • 1 ex.; Mt. Tachibana; 1. VI. 1978; K. Yamagishi leg. (KUM) • 1 ex.; same locality; 5. V. 1979; K. Yamagishi leg. (KUM) • 6 exs.; same locality; 10–14. V. 1996; T. Ueno, Y. Higashiura &amp; H. Goto leg. FIT (cSch) • 4 exs.; same locality; 15–17. V. 1996; T. Ueno &amp; H. Goto leg. FIT (cSch) • 1 ♀, 2 exs.; Mt. Shiouji-yama, Onojo-shi; 8. V. 2013; K. Kido leg. FIT (KUM) • 1 ex.; same locality; 12. V. 2013; K. Kido leg. FIT (KUM) • 1 ex.; same locality; 18. V. 2013; K. Kido leg. FIT (KUM) • 2 exs.; Mt. Hikosan; 13. VI. 1966; K. Takeno leg. (KUM) • 1 ♂, 1 ♀, 4 exs.; Seinen-no-ie, Hikosan (alt. 800 m), Soeda-machi; 22. V. 2011; M. Maruyama &amp; T. Kanao leg. LNJ (KUM) • 2 exs.; Kusenbu-yama; 24. V. 2014; K. Kido leg. FIT (KUM) • 1 ex.; Mt. Miyomizu-yama, Miyama-shi; 16. V. 1997; T. Ishikawa leg. (KUM) . Saga-ken • 1 ♂; Mt. Ten-zan (alt. 1040 m) [33.339, 130.143], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.143&amp;materialsCitation.latitude=33.339" title="Search Plazi for locations around (long 130.143/lat 33.339)">Karatsu-shi</a>; 29. VI. 2022; S. T. Inoue &amp; Y. Hisasue leg. LNJ (KUM) • 1 ex.; Mt. Tenzan (alt. 1,000 m) [33.340, 130.142], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.142&amp;materialsCitation.latitude=33.34" title="Search Plazi for locations around (long 130.142/lat 33.34)">Saga-shi</a>; 18. VI. 2022; Y. Hisasue leg. (KUM) • 2 exs.; Kurinoki-tôge, Mt. Kunimi-yama; 15. V. 2019; M. Nishida leg. FIT (KUM) • 1 ex.; Mt. Kurokami-yama, Arita-chô; 18. V. 2018; M. Nishida leg. FIT (KUM) • 2 exs.; Mt. Mifune, Takeo-shi; 27. V. 1984; S. Naomi leg. (KUM) • 1 ex.; Tosen-zan (alt. 350 m), Ureshino-shi; 19. V. 1997; S. Nomura leg. (KUM) • 1 ex.; Tosen-zan, Ureshino-shi; 20. V. 2018; M. Nishida leg. FIT (KUM) • 2 exs.; Mt. Tara, Tara-chô; 29. IV. 1985; S. Naomi leg. (KUM) . Nagasaki-ken • 1 ♀; Mt. Mitake, Tsushima Is., Tsushima-shi; 31. V. 1988; S. Nomura leg. (KUM) • 2 exs.; Tanuki-no-o, Ômura-shi; 8. V. 2019; M. Nishida leg. FIT (KUM) • 1 ex.; Mt. Yake, Shimabara-shi; 21. IV. 1977; Imasaka leg. (KUM) . Oita-ken • 1 ex.; Mt. Takasaki-yama, Beppu-shi; 9. V. 2013; T. Miyake leg. FIT (KUM) . Kagoshima-ken • 1 ex.; Mt. Kurinodake, Kurino-chô; 29. VI. 1995; S. Onoda leg. (SPMN) • 1 ♂; Nageishidaira, Yakushima Is. (alt. 1,825 m) [30.322, 130.512], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.512&amp;materialsCitation.latitude=30.322" title="Search Plazi for locations around (long 130.512/lat 30.322)">Yakushima-chô</a>; 18. VIII. 2021; S. Inoue leg. (KUM) .</p><p>Diagnosis.</p><p>This species is distinguished from other species of the genus by the following combination of characteristics: a slightly produced metaventral process (Fig. 13 D), median lobe of aedeagus bulbous with a length of approximately 0.47 mm (Fig. 13 G). This species is similar to H. ovata and H. japonica, but it can be distinguished by a more flattened and semicircular pronotum, less produced metaventral process, and a bulbous median lobe of the male aedeagus.</p><p>Redescription.</p><p>Body (Figs 3 E, 14 D) small, broad, somewhat limuloid; dorsal surface mostly moderately polished.</p><p>Head large; color yellowish brown to dark brown; eyes large. Antennae (Fig. 13 A) short; relative length of each antennal segment from first to 11 th, 2.39: 1.78: 1.14: 1.17: 1.00: 1.01: 1.10: 1.15: 1.20: 1.45: 3.94.</p><p>Thorax. Pronotum (Fig. 13 B) semicircular, transverse (PW / PL, 1.51–1.65), widest at posterior 1 / 3, postero-lateral angle acute to around 90 degrees, posterior margin slightly sinuate, yellowish brown to dark brown, surface finely covered with setae and punctures, gently polished and slightly reticulated. Elytra (Fig. 13 B, C) slightly widened posteriad, widest at posterior 1 / 3, then narrowed roundly, posterior margins deeply notched near lateral corners, yellowish brown to dark brown, surface finely covered with setae and punctures; gently polished and weakly reticulated.</p><p>Hind wings developed. Mesoventral process (Fig. 13 D) very narrow, with indistinct medial carina, apex sharply pointed, reaching almost anterior margin of mesocoxal cavities. Metaventral process slightly produced.</p><p>Abdomen elongate, narrowed posteriad; surface sparsely covered with short setae and each posterior margin with long stout setae; finely reticulated.</p><p>Male: 8 th sternite (Fig. 13 E) weakly produced and angled at postero-medial margin; median lobe of aedeagus (Fig. 13 G) bulbous, length about 0.47 mm, apical lobe short and slightly curved ventrally, apex round in lateral view, apical valves elongate and mitten-shaped, dorsal arms short, basal part of copulatory piece large. Paramere (Fig. 13 H) length about 0.47 mm; apical lobe of paramere long, length about 1 / 3 of whole paramere, curved ventrally; four setae, normally arranged, b located a bit more basally than c, velum small.</p><p>Female: 8 th sternite (Fig. 13 F) weakly produced roundly postero-medial margin; spermatheca (Figs 13 I, J) S- or C-shaped, apex of basal part slightly sinuate, base largely hooklike.</p><p>Variation.</p><p>Intraspecies variations are recognized in the apical lobe of the aedeagal median lobe. Some individuals have a small hump at the base of the ventral side and a slightly deeper basal concave. The length of the parameral apical lobe also varies slightly.</p><p>Measurements (in mm) and ratios.</p><p>Body parts (N = 13): BL ≈ 1.9–2.78; AL, 0.67–0.81; HW, 0.43–0.52; PL, 0.46–0.61; PW, 0.73–0.92; EL, 0.37–0.51; EW, 0.73–0.98; HTL, 0.4–0.48; PW / PL, 1.51–1.65; AL / PL, 1.22–1.53; HTL / PL, 0.8–0.9.</p><p>Distribution.</p><p>Japan (Hokkaido, Honshu, Shikoku, Kyushu) (Fig. 14 A).</p><p>Symbiotic hosts.</p><p>Lasius niger species group: L. hayashi, L. japonicus, L. sakagamii, L. productus .</p><p>Bionomics.</p><p>This species was collected from the nests of host ants under stones and in decaying wood and leaf litter near the entrances to these colonies from April to August. Many individuals were collected using FIT (N = 69). Bait traps set near the nest entrances were also effective in collecting H. prolongata . During indoor observations, H. prolongata was frequently observed, raising the tip of its abdomen toward approaching host ant workers, and sometimes just stopping (Fig. 14 B, C).</p><p>Remarks.</p><p>The type specimens are currently not available. Sawada (1970 a, 1970 b) described 25 aleocharine species from Shiga Heights, Nagano Prefecture, Honshu, including H. prolongata . According to Takuto Hashizume, who specializes in the taxonomy of homalotines and their relatives, he could not access these type specimens by contacting the National Museum of Nature and Science, Tsukuba (“ National Science Museum Tokyo ”), which is designated in papers as a deposit, or by searching the Osaka Museum of Natural History, Osaka, Japan. However, the original descriptions and drawings of H. prolongata are sufficiently informative for species-level taxonomy. Therefore, we conclude that a neotype designation is not necessary currently.</p><p>Maruyama et al. (2013) reported that some individuals similar to H. prolongata were found in trails of the Lasius fuliginosus group. However, we did not find any specimens of H. prolongata mounted together with L. fuliginosus . H. ovata may have been misrecognized as H. prolongata because some individuals of H. ovata resemble H. prolongata in terms of body size and general habitus.</p></div>	https://treatment.plazi.org/id/B8B9331D8F9F55778EA30E260A3174A6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Nozaki, Tsubasa;Maruyama, Munetoshi	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
E7F592F5B108547CA51BF5BB3E8692DA.text	E7F592F5B108547CA51BF5BB3E8692DA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Homoeusa rufescens (Sharp 1874)	<div><p>Homoeusa rufescens (Sharp, 1874)</p><p>Fig. 6 A – C, G (Japanese name: Hoso-hirata-ariyadori)</p><p>Thiasophila rufescens Sharp, 1874: 5; Fenyes 1919: 393 (catalogue); Bernhauer and Scheerpeltz 1926: 771 (catalogue); Smetana 2004: 488 (catalogue).</p><p>Homoeusa rufescens: Maruyama and Zerche 2014: 17 (mentioning actual generic affiliation); Schülke and Smetana 2015: 682 (catalogue); Nozaki and Maruyama 2022 a: 42 (the combination confirmed).</p><p>Additional material.</p><p>Japan: Honshu: Ibaraki-ken • 2 exs.; Asahi-tôge, Tsuchiura-shi; 8. V. 2022; Y. Komatsuzaki leg. (KUM) • 1 ex.; Nagai, Tsuchiura-shi; 27. IV. 2022; Y. Komatsuzaki leg. (KUM) . Chiba-ken • 5 exs.; Shimin-nomori Yachô-no-mori, Toyooka, Futtsu-shi; 10. V. 2007; H. Ono leg. (KUM) . Tokyo-to • 1 ex.; Mt. Takao (alt. 450 m), Hachiôji-shi; 4. VI. 2001; M. Maruyama leg. LFS (NHM) . Kanagawa-ken • 1 ex.; Nonkendainomori, Enkaizan, Yokohama-shi; 3. V. 1991; K. Kubo leg. (SPMN) . Nagano-ken • 2 exs.; Fuji, Satoyamabe, Matsumoto-shi; 21. V. 2004; T. Komatsu leg. (KUM) • 1 ex.; Fuji, Satoyamabe, Matsumoto-shi; 9. VI. 2004; T. Komatsu leg. (KUM) . Gifu-ken • 3 exs.; Mitahora, Gifu-shi, 10. VI. 2013; K. Kinomura leg. (KUM) • 2 exs.; Hirugano Bunsuirei Park, Gujo-shi; 10. VI. 2013; K. Kinomura leg. (KUM) . Shizuoka-ken • 2 exs.; Kumazaka, Izu-shi, 30. V. 2022; T. Tenma leg. (KUM) . Aichi-ken • 1 ex.; Jôkôji-kôen, Seto-shi; 4. V. 2023; T. Tenma leg. (KUM) . Osaka-fu • 3 exs.; Sakurai [34.8828, 135.6525], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=135.6525&amp;materialsCitation.latitude=34.8828" title="Search Plazi for locations around (long 135.6525/lat 34.8828)">Shimamoto-chô</a>; 9. V. 2021; Y. Takatani leg. LFS (KUM) . Shikoku: Kagawa-ken • 1 ex.; Shirahige-jinja, Ayagami-chô; 25. IV. 2004; H. Fujimoto leg. LFM (KUM) . Kochi-ken • 2 exs.; Ashizuri-misaki (alt. 120 m) [32.7480, 133.0136], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=133.0136&amp;materialsCitation.latitude=32.748" title="Search Plazi for locations around (long 133.0136/lat 32.748)">Tosashimizu-shi</a>; 21. V. 2024; TS. Nozaki leg. (KUM) . Kyushu: Fukuoka-ken • 2 exs.; Bio. Exp. Center of Kyushu Univ., Mt. Hikosan, Soeda-machi; 24. IV. 2021; N. Tsuji leg. (KUM) • 1 ♂; Biol. Lab. KU, Hikosan (alt. 750 m), Soeda-machi; 22. V. 2011; M. Maruyama &amp; T. Kanao leg. (KUM) . Miyazaki-ken • 1 ex.; Ômoridake forest road (alt. 600 m) [32.0435, 131.1973], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=131.1973&amp;materialsCitation.latitude=32.0435" title="Search Plazi for locations around (long 131.1973/lat 32.0435)">Aya-machi</a>; 12. VI. 2024; TS. Nozaki &amp; T. Kinoda leg. LFS (KUM) .</p><p>Diagnosis.</p><p>See Nozaki and Maruyama (2022 a).</p><p>Additional description.</p><p>Relative length of each antennal segment from first to 11 th, 2.27: 1.63: 1.37: 1.27: 1.02: 1.02: 1.00: 1.05: 1.07: 1.44: 3.34. Hind wings developed.</p><p>Male. 8 th sternite (Fig. 6 A) weakly round at posterior margin; four setae of parameral apical lobe (Fig. 6 C) normally arranged.</p><p>Female. 8 th sternite (Fig. 6 B) gently round at posterior margin.</p><p>Distribution.</p><p>Japan (Honshu, Shikoku, Kyushu) (Fig. 6 G).</p><p>Symbiotic hosts.</p><p>Lasius cf. fuliginosus, L. morishitai, L. nipponensis, L. spathepus .</p></div>	https://treatment.plazi.org/id/E7F592F5B108547CA51BF5BB3E8692DA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Nozaki, Tsubasa;Maruyama, Munetoshi	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
41E2F413A9C851B5B401F51195E80F50.text	41E2F413A9C851B5B401F51195E80F50.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Homoeusa translucida Nozaki & Maruyama 2025	<div><p>Homoeusa translucida Nozaki &amp; Maruyama sp. nov.</p><p>Figs 3 A, 4 A – I, 5 A, B Japanese name: Sukashi-hirata-ariyadori</p><p>“ Homoeusa sp. 4 ”: Saitoh et al. 2017: 33, fig. 65 (recorded from Kanagawa-ken).</p><p>“ Oxypodini ? sp. 2 ”: Nishida 2019: 135, fig. 11 (recorded from Saga-ken).</p><p>Material examined.</p><p>Holotype (Fig. 1 A) • ♂; “ Mt. Minoyama, Minano, ” / “ Saitama, Japan, alt. 500 - ” / “ 560 m; MC:5439-00-69. ” / “ 21. V. 1999 M. UCHIDA ” (KUM) . Paratypes. Japan: Hokkaido • 1 ♀; Nopporo Forest Park, Sapporo-shi; 24. V. &amp; 5. VI. 2008; Tomáš Lackner leg. (NHMW) . Honshu: Kanagawa-ken • 1 ex.; Yakunimi-yama, Hadano-shi; 10. V. 2013; O. Saitoh leg. (KPMNH) • 1 ♀; same locality; 24. V. 2014; O. Saitoh leg. FIT (KPMNH) • 1 ♂; same locality; 8. VI. 2014; O. Saitoh leg. FIT (KPMNH) . Shimane-ken • 1 ex.; Urahikimi, Masuda-shi; 6. VI. 1998; S. Nomura leg. (KUM) . Okayama-ken • 1 ex.; Nagi-yama, Nagi-chô; 17. V. 2005; W. Suzuki leg. (KUM) . Shikoku: Ehime-ken • 1 ex.; Mt. Sara-ga-mine, Toon-shi; 13. VII. 2008; Y. Yamauchi leg. (EUMJ) . Kochi-ken • 1 ex.; Kajigamori, Ôtoyo-chô; 20. V. 1951; M. Miyamoto leg. (EUMJ) . Kyushu: Fukuoka-ken • 1 ex.; Takasubaru, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.9174&amp;materialsCitation.latitude=33.4883" title="Search Plazi for locations around (long 130.9174/lat 33.4883)">Mt. Hikosan</a> [33.4883, 130.9174], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.9174&amp;materialsCitation.latitude=33.4883" title="Search Plazi for locations around (long 130.9174/lat 33.4883)">Soeda-machi</a>; 27. VI. 2021; TS. (Tsubasa) Nozaki leg. LUM (KUM) • 1 ex.; Bio. lab. KU. Mt. Hikosan, Soeda-machi; 28. III – 27. IV. 2023; J. Abe leg. Malaise trap (KUM) . Saga-ken • 1 ♂; Mt. Seira, Imari-shi; 28. V. 1984; S. Nomura leg. (KUM) • 1 ♂; Kurokami-yama, Arita-cho; 18. V. 2018; M. Nishida leg. FIT (KUM) .</p><p>Diagnosis.</p><p>This species can be easily distinguished from the other species of the genus by the following combination of characteristics: body color yellowish orange; pronotum strongly transverse (PW / PL, 1.84–2.05), postero-lateral angle distinctly acute (Fig. 4 B); metaventral process weakly produced (Fig. 4 D); pronotum and elytra sparsely covered with modified stout setae (Fig. 4 B, C); median lobe of aedeagus apical lobe straight, apex round and slightly humped in lateral view, apical valves very large and cheliform (Fig. 4 G); spermatheca (Fig. 4 I) L-shaped, basal part very long. This species is slightly similar to H. gigantea sp. nov. in color but differs in its pronotal and setal shapes.</p><p>Description.</p><p>Body (Figs 3 A, 5 B) small, broad; dorsal surface mostly polished.</p><p>Head large; color yellowish orange; eyes large. Antennae (Fig. 4 A) long; relative length of each antennal segment from first to 11 th, 2.48: 1.96: 1.39: 1.22: 1.04: 1.09: 1.00: 1.15: 1.15: 1.37: 3.26.</p><p>Thorax. Pronotum (Fig. 4 B) subrectangular, strongly transverse (PW / PL, 1.84–2.05), widest near middle, postero-lateral angle distinctly acute, posterior margin distinctly sinuate, color yellowish orange, subtransparent, surface sparsely covered with modified stout setae and punctures, gently polished without reticulation. Elytra (Fig. 4 B, C) slightly widened posteriad, widest at middle, then narrowed roundly, posterior margins shallowly notched near lateral corners, color yellowish orange, subtransparent, surface sparsely covered with modified stout setae and punctures, gently polished without reticulation. Hind wings developed. Mesoventral process (Fig. 4 D) narrow, with weak medial carina, apex gently pointed, reaching posterior 1 / 3 of mesocoxal cavities, Metaventral process weakly produced.</p><p>Abdomen elongate, narrowed posteriad; surface sparsely covered with short setae and each posterior margin with long stout setae; covered with fine vertical furrows.</p><p>Male: 8 th sternite (Fig. 4 E) widely protruding at middle of posterior margin; median lobe of aedeagus (Fig. 4 G) elongate, length about 0.57 mm, apical lobe straight, apex round and slightly humped in lateral view, apical valves large and cheliform, dorsal arms boot-shaped; basal part of copulatory piece very large and having claw-shaped projection, basal corner acutely pointed. Paramere (Fig. 4 H) length about 0.59 mm; apical lobe of paramere long, length over 1 / 3 of whole paramere, curved ventrally, slightly dilated apicad; four setae, c located a little more apically than b; velum narrow and membranous.</p><p>Female: 8 th sternite (Fig. 4 F) slightly protruding at the middle of posterior margin; spermatheca (Fig. 4 I) L-shaped, basal part very long, apex of basal part not swollen, base almost straight and only slightly swollen.</p><p>Measurements (in mm) and ratios.</p><p>Body parts (N = 10): BL ≈ 2.13–2.91; AL, 0.93–1.07; HW, 0.51–0.57; PL, 0.5–0.59; PW, 1–1.13; EL, 0.47–0.52; EW, 0.98–1.16; HTL, 0.69–0.76; PW / PL, 1.84–2.05; AL / PL, 1.69–1.95; HTL / PL, 1.29–1.46.</p><p>Distribution.</p><p>Japan (Honshu, Shikoku, Kyushu) (Fig. 5 A).</p><p>Symbiotic hosts.</p><p>Lasius umbratus species group: L. meridionalis (only single example).</p><p>Bionomics.</p><p>Two individuals were collected by FIT, and a single individual was collected by the malaise trap. The behavior with the host ants is unknown.</p><p>Remarks.</p><p>This species is relatively rare, and most specimens have no information regarding their host ants. However, a single specimen (Fig. 5 B) was collected with L. meridionalis using a bait trap set by TN under the stone where the ant trail was located.</p><p>Etymology.</p><p>The specific name is derived from the Latin adjective, which means “ transparent ” and refers to the character of the pronotum of this new species.</p></div>	https://treatment.plazi.org/id/41E2F413A9C851B5B401F51195E80F50	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Nozaki, Tsubasa;Maruyama, Munetoshi	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
