taxonID	type	description	language	source
C75D8643CF33545DAD7AA33600D4D34C.taxon	description	Fig. 3, Table 2	en	Shen, Deming, Zhou, Haiying, Wang, Chaoyang, Xu, Jun, Xu, Shi, Zhong, Yifeng, Wang, Zhenyu, Cai, Hao, Wu, Yahui, Wei, Meicai, Li, Yankuo (2025): A new species of Boulenophrys (Anura, Megophryidae) from northern Jiangxi, China. Zoosystematics and Evolution 101 (4): 2251-2266, DOI: 10.3897/zse.101.166556
C75D8643CF33545DAD7AA33600D4D34C.taxon	etymology	Etymology. The specific epithet nebulosa (feminine of Latin nebulosus) refers to the frequently mist-shrouded montane forests above 600 m on Mt. Lushan, Jiangxi Province, China. This name not only reflects the species’ habitat preference but also alludes to the region’s famous “ Cloud & Fog Tea ” (云雾茶, yún wù chá) speciality. The English vernacular name “ Yunwu Horned Toad ” is here proposed, with the corresponding Chinese name “ 云雾角蟾 ” (Yúnwù Jiǎochán).	en	Shen, Deming, Zhou, Haiying, Wang, Chaoyang, Xu, Jun, Xu, Shi, Zhong, Yifeng, Wang, Zhenyu, Cai, Hao, Wu, Yahui, Wei, Meicai, Li, Yankuo (2025): A new species of Boulenophrys (Anura, Megophryidae) from northern Jiangxi, China. Zoosystematics and Evolution 101 (4): 2251-2266, DOI: 10.3897/zse.101.166556
C75D8643CF33545DAD7AA33600D4D34C.taxon	diagnosis	Diagnosis. This species can be distinguished from its congeners by the combination of following morphological characteristics: (1) adult males, SVL 31.8 – 32.3 mm (n = 3); adult females, SVL 35.0 – 37.3 mm (n = 4); (2) canthus rostralis well developed, tongue not notched posteriorly; (3) tympanum distinct; (4) vomerine ridges weak; vomerine teeth present; (5) outer margin of the upper eyelid bears a small, prominent horn-like tubercle; supratympanic fold distinct, narrow, curving posteroventrally to above the arm; (6) relative finger lengths I <II <IV <III; indistinct subarticular tubercle on base of each finger; (7) heels meeting when hindlimbs folded; tibiotarsal articulation reaching center of eye when leg stretched forward; (8) toes with narrow fringes and only rudimentary webbing; (9) rough dorsal skin, a V-shaped, X-shaped or Y-shaped ridge is present on central back of trunk, dorsal surface bears short, irregular and weak ridges, with scattered tubercles and granules, dorsolateral ridges absent; several tubercles on flanks and dorsal hindlimbs; ventral skin smooth with granules; (10) dorsal surface yellowish brown; hollow brown triangular marking between eyes; brown V-shaped, X-shaped, or Y-shaped marking on central back of trunk; dorsal limbs and digits with dark brown transverse bands; and (11) dense nuptial spines on dorsal bases of fingers I and II in breeding adult males, subgular vocal sac present in males.	en	Shen, Deming, Zhou, Haiying, Wang, Chaoyang, Xu, Jun, Xu, Shi, Zhong, Yifeng, Wang, Zhenyu, Cai, Hao, Wu, Yahui, Wei, Meicai, Li, Yankuo (2025): A new species of Boulenophrys (Anura, Megophryidae) from northern Jiangxi, China. Zoosystematics and Evolution 101 (4): 2251-2266, DOI: 10.3897/zse.101.166556
C75D8643CF33545DAD7AA33600D4D34C.taxon	description	Description of holotype. JXNU 21042833, adult male, body size small, SVL 32.2 mm; head width slightly smaller than head length, HDW / HDL 0.91; snout rounded in dorsal view, projecting, sloping backward to mouth in profile, protruding well beyond margin of lower jaw; top of head flat; eyes large, ED 0.35 of HDL, pupil vertical, ovoid in natural state; nostril obliquely ovoid; canthus rostralis well developed; loreal region slightly oblique; internasal distance larger than interorbital distance; tympanic region oblique, tympanum moderate in size, margin clear, upper margin in contact with supratympanic fold, TD / ED 0.41; large ovoid choanae at base of maxilla; vomerine ridge weak, vomerine teeth present, maxillary teeth present; margin of tongue rounded, not notched distally; presence of subgular vocal sac. Forearm length 0.24 of SVL, hand 0.22 of SVL; webbing absent between fingers, lateral fringes absent, relative finger length I <II <IV <III; tips of fingers slightly dilated, round; indistinct subarticular tubercles present; inner metacarpal tubercle observably enlarged, outer one indistinct; single nuptial pad bearing nuptial spines present on dorsal surface of first and second fingers, respectively. Hindlimbs short, heels meeting when flexed hindlimbs are held at right angles to the body axis; tibiotarsal articulation reaching the center of the eye when the hindlimb is stretched forward along the body; tibiofibular length 0.45 of SVL, foot length 0.39 of SVL; relative toe length I <II <V <III <IV; tips of toes rounded and slightly dilated; toes with narrow lateral fringes and rudimentary webbing; subarticular tubercles indistinct at the base of each toe; inner metatarsal tubercle long and ovoid; outer metatarsal tubercle absent. Dorsal skin smooth with granules; a discontinuous Y-shaped ridge present on the central dorsum of trunk, discontinuous longitudinal ridges present laterally; several rounded tubercles present on the flanks and dorsal surfaces of the hindlimbs; a prominent, small horn-like tubercle located on the upper eyelid; supratympanic fold distinct, extending from the posterior corner of the eye to above the insertion of the forelimb; tympanum distinct; ventral skin smooth to weakly granular, with spinose tubercles surrounding the cloacal region; pectoral gland small and distinct, located closer to the axilla; a single large femoral gland present on the posterior surface of the thigh, situated approximately midway between the knee and the cloaca.	en	Shen, Deming, Zhou, Haiying, Wang, Chaoyang, Xu, Jun, Xu, Shi, Zhong, Yifeng, Wang, Zhenyu, Cai, Hao, Wu, Yahui, Wei, Meicai, Li, Yankuo (2025): A new species of Boulenophrys (Anura, Megophryidae) from northern Jiangxi, China. Zoosystematics and Evolution 101 (4): 2251-2266, DOI: 10.3897/zse.101.166556
C75D8643CF33545DAD7AA33600D4D34C.taxon	description	Tadpole description. The tadpole was confirmed as Boulenophrys nebulosa sp. nov. by molecular phylogenetic analyses. The following tadpole description is based on the specimen JXNU 21110702 at Gosner Stage 29 (Fig. 5). Body depressed and longitudinally oval; TOL 28.0 mm; body width longer than body height, BW / BH 1.3; eyes dorsolateral, pupils round; nostril near eyes; spiracle on the left side of the body and distinct; tail approximately 2.3 times as long as snout-vent length; tail height greater than body height, TAH / BH 1.2; dorsal fin arising just behind tail base, highest near mid-length, tapering gradually to bluntly pointed tip; oral disk terminal; lips expanded, directed upward, forming umbelliform structure; lateral corners of oral disc fold medially, turning upward when not fully extended. In preservative, the background color of the head and trunk fades from brown to dark brown, interspersed with dark-brown chromatophores, and becomes progressively paler toward the tail tip. Areas lacking white pigmentation contain fewer melanocytes. The belly bears dense white spots, especially concentrated on the chest, while the lateral surfaces of the tail are covered with yellowish-brown spots. BH 2.7; BW 3.6; IOD 2.5; MW 1.6; SL 1.8; SS 3.2; SVL 8.5; TAH 3.3; TAL 19.5; TBW 1.4; TOL 28.0.	en	Shen, Deming, Zhou, Haiying, Wang, Chaoyang, Xu, Jun, Xu, Shi, Zhong, Yifeng, Wang, Zhenyu, Cai, Hao, Wu, Yahui, Wei, Meicai, Li, Yankuo (2025): A new species of Boulenophrys (Anura, Megophryidae) from northern Jiangxi, China. Zoosystematics and Evolution 101 (4): 2251-2266, DOI: 10.3897/zse.101.166556
C75D8643CF33545DAD7AA33600D4D34C.taxon	distribution	Distribution and habits. Boulenophrys nebulosa sp. nov. is currently known only from its type locality, Mt. Lushan, Jiujiang City, Jiangxi Province, China. The species occurs at elevations of 600 – 1,200 m, primarily in mixed coniferous and broadleaved forests (Fig. 7). Individuals are typically found on the forest floor or under leaf litter near montane streams, as well as along moist roadside drainage channels. Calling males were observed from May to July, all of which were observed calling beneath accumulations of coniferous leaf litter. Females were occasionally observed in drainage ditches lacking flowing water. Tadpoles are observed throughout the year and coexist with the tadpoles of Quasipaa spinosa and B. lushanensis in some aquatic habitats.	en	Shen, Deming, Zhou, Haiying, Wang, Chaoyang, Xu, Jun, Xu, Shi, Zhong, Yifeng, Wang, Zhenyu, Cai, Hao, Wu, Yahui, Wei, Meicai, Li, Yankuo (2025): A new species of Boulenophrys (Anura, Megophryidae) from northern Jiangxi, China. Zoosystematics and Evolution 101 (4): 2251-2266, DOI: 10.3897/zse.101.166556
