taxonID	type	description	language	source
C604E518AE531D72FD9C827F6AFCA365.taxon	type_taxon	Type genus: Mesitius Spinola, 1851.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE531D72FD9C827F6AFCA365.taxon	diagnosis	Diagnosis Head and mesosoma usually densely foveolate; malar space projected; contour of eye protruding; presence of anterior depression of occiput; ventral half of mesoccipital carina angled. Median pronotal sulcus usually present. Presence of space between tegula and mesoscutum; notauli developed, usually converging posteriorly. Presence of metapostnotal depression; metapostnotal-propodeal carina usually present and converging posteriorly; paraspiracular carina usually present, when absent then also PPP absent. Second metasomal segment distinctly longer than other segments and at least half as long as metasoma in dorsal view. Male genitalia with: genital ring projected; gonostipes and basivolsella fused; harpe bilobate apically.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE541D75FD9F83A869D4A3E3.taxon	diagnosis	Diagnosis Flagellomeres with setae dense and short (shorter than half-width of flagellomere); frons and dorsal area of pronotum densely foveolate. Median pronotal sulcus present. Median mesonotal sulcus usually absent (♀). Male hypopygium longer than wide with posterior branches broad and short; basal part of hypopygium with lateral and anterior margin incurved. Male genitalia with dorsal harpe S-shaped.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE541D75FD9F83A869D4A3E3.taxon	biology_ecology	Host Larva of Cryptocephalus sinaita moricei Pic, 1908 (Coleoptera, Chrysomelidae) (Argaman 2003).	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE541D75FD9F83A869D4A3E3.taxon	distribution	Distribution Afrotropical, Oriental and Palaearctic Regions.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE551D79FD8E80DD6A24A443.taxon	description	urn: lsid: zoobank. org: act: A 0405 FAC- 324 B- 49 FA-A 5 F 7 - 5796 DECE 9 FD 0 Fig. 2	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE551D79FD8E80DD6A24A443.taxon	diagnosis	Diagnosis This new species can be recognized by its having flagellomeres with dense setae shorter than one-third of width of flagellomere; median pronotal sulcus present and not foveolate; median mesonotal sulcus present near posterior margin of mesoscutum; PPP equal to or slightly longer than its width at base; hypopygium longer than wide with anterior margin broad and posterior branches short; genitalia with dorsal harpe narrower than ventral harpe. This new species is similar to H. neomoczari (Barbosa & Azevedo, 2018) in having flagellomere I less than 2.0 × length of flagellomere II. However, it can be distinguished by having median pronotal sulcus not foveolate (distinctly foveolate in H. neomoczari); median mesonotal sulcus present (absent in H. neomoczari); PPP with apical margin rounded in lateral view (acute in H. neomoczari).	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE551D79FD8E80DD6A24A443.taxon	etymology	Etymology The specific epithet ‘ curtisulcus ’ is a combination of the Latin words ‘ curtus ’ (‘ short’) and ‘ sulcus ’ (‘ groove’) referring to the median mesonotal sulcus of this species present on the posterior third of the mesoscutum.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE551D79FD8E80DD6A24A443.taxon	materials_examined	Material examined Holotype CHINA – Shandong • ♂; Laoshan; 3 Aug. 1995; Ping Cai leg.; ZJUH 958281. Paratype (1 ♂) CHINA – Shandong • 1 ♂; Laoshan; 4 Aug. 1995; Junhua He leg.; ZJUH 955913.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE551D79FD8E80DD6A24A443.taxon	description	Description Male MEASUREMENTS (n = 2). Body length 4.66 – 5.02 mm; fore wing length 2.65 – 3.15 mm; LH 1.07 – 1.09 × WH; WF 1.67 – 1.74 × LE; LE 0.99 – 1.02 × DEV; POL 1.62 – 1.70 × AOL; OOL 0.95 – 1.01 × WOT; DPV 1.90 – 2.14 × DAO; median length of pronotum 0.53 – 0.54 × its width along posterior margin; length of hypopygium 1.73 – 1.79 × its width; length of posterior branches of hypopygium 0.13 – 0.16 × length of hypopygium. COLOURATION. Black. Mandible dark brown, teeth brown; maxillary and labial palpi brown. Antenna brown, scape blackish brown. Legs brown, coxae and femora dark brown, tarsi yellowish brown. Fore wing hyaline, light brown; veins and pterostigma brown. Trochanters, tibiae apically and tarsi yellowish brown. Mesosoma blackish brown. Metasomal segments brown apically. PUBESCENCE. Body with short setae, denser on ventral surface. Antenna with short appressed setae, 0.25 × width of flagellomere (Fig. 2 C). Eye with sparse short setae, nearly 4.0 × diameter of an ommatidium. Wings with short, dense brown setae. Legs with dense, short setae. Metasoma with setae longer than setae on head and mesosoma; T 1 with setae laterally, T 2 with basal glabrous triangular area dorsally. HEAD. Longer than wide, LH 1.07 × WH. Mandible with four apical teeth, ventralmost one longest. Clypeus trilobite, medio-clypeal lobe with anterior margin rounded (Fig. 2 B); medio-clypeal carina present, not extending into frons, arched in lateral view. Antenna thin, flagellomeres cylindrical, length of flagellomere III 1.48 × length of flagellomere II. Frons coriaceous and densely foveolate (Fig. 2 B); WF 1.74 × LE. Contour of eye distinctly protruding in antero-dorsal view (Fig. 2 B); LE 1.02 × DEV. Anterior ocellus partly anterior to imaginary top line of eyes in antero-dorsal view; POL 1.62 × AOL, OOL 1.01 × WOT, DPV 1.90 × DAO. Vertex coriaceous and densely foveolate, sides of head behind eyes rounded, vertex crest protruding. Gena coriaceous and densely foveolate, ventral surface with shallower and sparser foveae, medioccipito-genal suture present. Occipital carina present, occiput coriaceous. MESOSOMA. Dorsal pronotal area coriaceous and densely foveolate, median length 0.54 × width along posterior pronotal margin; anterolateral corner rounded; median pronotal sulcus complete, not foveolate (Fig. 2 D); lateral pronotal area areolate; cervical pronotal area coarse. Mesoscutum coriaceous with dense punctures, elevated posteriorly; median mesonotal sulcus present on posterior third of mesonotum (Fig. 2 D); notauli complete and converging posteriorly (Fig. 2 D); parapsidal signum shallow and absent near anterior margin. Mesoscutellum coriaceous and foveolate; mesoscutum-mesoscutellar sulcus present, laterally dilated (Fig. 2 D); medio-longitudinal sulcus absent. Dorsal surface of MPC elevated; median length of dorsal surface of MPC equal to its half-width; metapostnotum strongly areolate, metapostnotal median carina complete; metapostnotal-propodeal carina complete, distinctly oblique anteriorly and converging posteriorly; dorsal propodeal area striate; paraspiracular and lateral marginal carinae of MPC complete; PPP present, 0.21 × median length of dorsal surface of MPC, apical margin of PPP rounded in lateral view; transverse posterior carina of MPC complete; propodeal declivity areolate, median and lateral carinae complete; lateral surface of MPC areolate, sublateral carina of MPC not extending to PPP. Propleuron coriaceous and densely foveolate. Mesopleuron coriaceous, densely foveolate; subalar impression present; posterior oblique sulcus of mesopleuron foveolate; postepicnemial sulcus present; ventral surface foveolate, with short carina near mesocoxa, medio-longitudinal sulcus present on apical half. WINGS. Fore wing with 2 r-rs & Rs 2 v slightly curved apically; posterior margin of pterostigma straight, prestigma absent (Fig. 2 F); Cu 2 v nebulous. Hind wing with three distal hamuli, distance between 1 st and 2 nd hamuli 1.43 × distance between 2 nd and 3 rd hamuli. METASOMA. Metasomal terga shiny, with punctures; T 1 with tiny sparse punctures, medio-longitudinal sulcus present at basal third of T 1; T 2 with dense punctures, coriaceous basally (Fig. 2 E); basal half of T 3 – 4 coriaceous. Metasomal sterna shiny, with variably sized punctures; S 1 with dense punctures. Length of hypopygium 1.79 × its width; anterior margin incurved (Fig. 2 G); posterior margin incurved, posterior branches broad and short, with long setae, length of branches 0.16 × length of hypopygium. MALE GENITALIA. Harpe bilobate, dorsal harpe S-shaped (Fig. 2 H); ventral harpe broader than dorsal harpe, with long setae apically (Fig. 2 I). Cuspis bilobate, with long setae; digitus bent and papillate apically, with setae basally; aedeagus bottle-shaped (Fig. 2 H). Female Unknown.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE551D79FD8E80DD6A24A443.taxon	distribution	Distribution China (Shandong) (Fig. 20).	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE551D79FD8E80DD6A24A443.taxon	discussion	Remarks The type specimens of this species were identified as Sulcomesitius borneoensis Móczár, 1976 by Xu et al. (2003) because of the presence of a median mesonotal impression on the mesoscutum near the posterior margin. However, some males of the genus Heterocoelia also have a median mesonotal sulcus on the mesoscutum (Azevedo et al. 2018; Barbosa et al. 2022), and the shape of hypopygium is now used as an important character for the generic identification of the males. After an examination of the male hypopygium, we included these specimens in the genus Heterocoelia rather than Sulcomesitius because of the longer than wide hypopygium with posterior branches broad and short. In Heterocoelia they cannot be associated with a named species; therefore, the species is here described as H. curtisulcus sp. nov.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE581D7BFE4D83BF6823A0B4.taxon	description	Fig. 3	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE581D7BFE4D83BF6823A0B4.taxon	materials_examined	Material examined Holotype CHINA – Shandong • ♂; Laoshan; 5 Aug. 1995; Ping Cai leg.; ZJUH 958428.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE581D7BFE4D83BF6823A0B4.taxon	description	Re-description Male MEASUREMENTS. Body length 3.61 mm. Fore wing length 2.22 mm. COLOURATION. Black. Mandible brown, teeth yellowish brown; maxillary and labial palpi brown. Antenna brown, scape blackish brown. Wings hyaline; fore wing light brown, veins and pterostigma light brown. Legs dark brown, tarsi yellowish brown. Metasomal segments black with posterior margin brown. PUBESCENCE. Body with short setae, denser on ventral surface. Antenna with short suberect setae, length of setae one-third width of flagellomere (Fig. 3 C). Eye with sparse, short setae, nearly 4.0 × diameter of an ommatidium. Wings with short, dense brown setae. Legs with dense, short setae. Metasoma with sparse setae; T 1 with setae laterally, T 2 with basal glabrous triangular area dorsally. HEAD. Length equal to width, LH 1.01 × WH. Clypeus trilobite, medio-clypeal lobe with anterior margin slightly acute medially (Fig. 3 B); medio-clypeal carina present, not extending into frons, arched in lateral view. Antenna thin, flagellomeres cylindrical, length of flagellomere III 1.69 × length of flagellomere II. Frons coriaceous and densely foveolate, frontal line shallow and extending to anterior ocellus (Fig. 3 B); WF 1.69 × LE. Contour of eye protruding in antero-dorsal view (Fig. 3 B); LE 1.16 × DEV. Anterior ocellus partly anterior to imaginary top line of eyes in antero-dorsal view; POL 1.57 × AOL, OOL 1.03 × WOT, DPV 1.63 × DAO. Vertex coriaceous and densely foveolate, sides of head behind eyes rounded, vertex crest protruding. Gena coriaceous and densely foveolate, ventral surface coriaceous; medioccipito-genal suture present. Occipital carina complete; occiput coriaceous. MESOSOMA. Dorsal pronotal area coriaceous and densely foveolate, median length 0.49 × width along posterior pronotal margin; anterolateral corner rounded; median pronotal sulcus foveolate (Fig. 3 D); lateral pronotal area areolate; cervical pronotal area coarse. Mesoscutum coriaceous with punctures, elevated posteriorly; median mesonotal sulcus absent (Fig. 3 D); notauli complete and converging posteriorly (Fig. 3 D); parapsidal signum shallow, absent near anterior margin. Mesoscutellum coriaceous and sparsely foveolate (Fig. 3 D); mesoscutum-mesoscutellar sulcus present, laterally dilated (Fig. 3 D); medio-longitudinal sulcus absent. Dorsal surface of MPC elevated; median length of dorsal surface of MPC 1.16 × its half-width; metapostnotum strongly areolate, metapostnotal median carina complete (Fig. 3 E); metapostnotal-propodeal carina complete, distinctly oblique anteriorly and converging posteriorly; dorsal propodeal area striate; paraspiracular and lateral marginal carinae of MPC complete; PPP present, 0.19 × median length of dorsal surface of MPC (Fig. 3 E); transverse posterior carina of MPC complete; propodeal declivity areolate, median and lateral carinae complete; lateral surface of MPC areolate. Propleuron coriaceous, propleural epicoxal sulcus foveolate. Mesopleuron coriaceous and densely foveolate, posterior area transversely striate; subalar impression present; posterior oblique sulcus of mesopleuron foveolate; postepicnemial sulcus present; ventral surface coriaceous, foveolate near mesocoxa; anterior mesofurcal pit present. WINGS. Fore wing with 2 r-rs & Rs 2 v slightly curved apically; posterior margin of pterostigma straight (Fig. 3 G); Cu 2 v nebulous. Hind wing with three distal hamuli, distance between 1 st and 2 nd hamuli longer than distance between 2 nd and 3 rd hamuli. METASOMA. Metasomal terga shiny with punctures; T 1 with tiny sparse punctures, medio-longitudinal sulcus present at basal one-third; T 2 with punctures (Fig. 3 F); T 3 – 4 coriaceous basally. Metasomal sterna shiny, with variably sized punctures; S 1 irregularly striate, metapostnotal median carina weak. Length of hypopygium 1.84 × its width; anterior margin incurved (Fig. 3 H); posterior margin bilobate, posterior branches broad and short with long setae, length of posterior branches 0.14 × length of hypopygium. MALE GENITALIA. Harpe bilobate, dorsal harpe S-shaped (Fig. 3 I); ventral harpe broader than dorsal harpe, with long setae apically (Fig. 3 J). Cuspis bilobate with long setae; digitus bent and papillate apically, with setae basally; aedeagus bottle-shaped (Fig. 3 I). Female Unknown.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE581D7BFE4D83BF6823A0B4.taxon	distribution	Distribution China (Shandong) (Fig. 20).	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE581D7BFE4D83BF6823A0B4.taxon	discussion	Remarks This species was included in the genus Sulcomesitius by Xu et al. (2003) because it has the anterior corners of the clypeus obtuse laterally. However, the generic identification of the males is primarily based on the shape of the hypopygium (Argaman 2003; Azevedo et al. 2018). We examined the types of this species and transfer it from Sulcomesitius to Heterocoelia because it has the hypopygium longer than wide with the posterior branches 0.14 × the length of the hypopygium, the dorsal harpe narrower than the ventral harpe and the antennal setae short.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE5A1D7CFE4187CE6A9AA222.taxon	description	Fig. 4	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE5A1D7CFE4187CE6A9AA222.taxon	materials_examined	Material examined Holotype CHINA – Fujian • ♀; Zhangzhou; Apr. 1987; Naiquan Lin leg.; ZJUH 984787. Other material (4 ♀♀) CHINA – Fujian • 1 ♀; Fuzhou, Jinshan; Apr. 1987; Naiquan Lin leg.; ZJUH 202300001 • 1 ♀; Shouning; 14 Jul. 1987; Changming Liu leg.; ZJUH 9611394 • 1 ♀; Fuzhou; 13 Aug. 1989; Changming Liu leg.; ZJUH 966313 • 1 ♀; Fuzhou; 14 Aug. 1990; Changming Liu leg.; ZJUH 966320.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE5A1D7CFE4187CE6A9AA222.taxon	description	Re-description Female MEASUREMENTS (n = 5). Body length 5.42 – 6.88 mm; fore wing length 2.85 – 3.20 mm; LH 1.06 – 1.08 × WH; WF 1.42 – 1.57 × LE; LE 1.35 – 1.43 × DEV; POL 1.22 – 1.34 × AOL; OOL 0.90 – 1.04 × WOT; median length of pronotum 0.48 – 0.53 × its width along posterior margin; median length of dorsal surface of MPC 0.71 – 0.83 × its half-width; length of PPP 0.68 – 0.72 × median length of dorsal surface of MPC. COLOURATION. Black. Mandible brown; maxillary and labial palpi brown, lighter ventrally. Antenna black; scape, pedicel and flagellomeres I – II brown. Legs brown, procoxa partly dark brown. Fore wing hyaline, brown with longitudinal colourless band medially; veins yellowish brown to brown, pterostigma yellowish brown. Posterior margin of T 1 – 2 and S 2 – 3, apical half of T 3 and metasomal segments 4 – 7 yellowish brown. PUBESCENCE. Body with short, dense light yellowish-brown setae. Antenna with dense appressed setae, sparser on scape (Fig. 4 C). Eye with sparse erect setae, 3.0 × diameter of an ommatidium. Wings with short, dense brown setae. Setae of metasoma longer than setae on head and mesosoma; T 1 with setae laterally, T 2 with basal glabrous triangular area dorsally. HEAD. Longer than wide, LH 1.06 × WH. Mandible with four apical teeth, ventralmost one longest. Clypeus trilobite; medio-clypeal lobe with anterior margin slightly acute medially (Fig. 4 B); medio-clypeal carina present, not extending into frons, arched in lateral view. Antenna thickened, ratio between length and width of flagellomeres IV – VII are 0.87, 0.71, 0.78 and 0.95 respectively; pedicel shorter than flagellomere I, length of pedicel 0.74 × length of flagellomere I. Frons coriaceous and densely foveolate; WF 1.57 × LE (Fig. 4 B). Contour of eye protruding in antero-dorsal view (Fig. 4 B); LE 0.36 × LH; LE 1.35 × DEV. Anterior ocellus crossing imaginary top line of eyes in antero-dorsal view; POL 1.23 × AOL, OOL 1.04 × WOT, DPV 2.73 × DAO. Vertex coriaceous and densely foveolate; sides of head behind eyes rounded, vertex crest protruding; occipital carina complete. Gena coriaceous and densely foveolate; medioccipito-genal suture and hypostomal sulcus present. MESOSOMA. Dorsal pronotal area coriaceous and densely foveolate, median length 0.53 × width along posterior pronotal margin (Fig. 4 D); anterolateral corner weakly angled; median pronotal sulcus foveolate; lateral pronotal area obliquely striate; cervical pronotal area coarse. Mesoscutum distinctly elevated on apical two-thirds, coriaceous with dense punctures; median mesonotal sulcus absent, only impressed on apical half (Fig. 4 E); notauli complete and converging posteriorly; parapsidal signum shallow, absent near anterior margin. Mesoscutellum coriaceous with dense punctures; mesoscutum-mesoscutellar sulcus present, laterally dilated (Fig. 4 E); medio-longitudinal sulcus absent. Dorsal surface of MPC elevated; median length of dorsal surface of MPC 0.78 × its half-width; metapostnotum strongly areolate, metapostnotal median carina complete (Fig. 4 E); metapostnotal-propodeal carina distinctly oblique anteriorly and converging posteriorly; dorsal propodeal area transversely striate; paraspiracular and lateral marginal carinae of MPC complete, lateral marginal carina nearly straight; PPP long, 0.69 × median length of dorsal surface of MPC, not bent upward in dorsal view; transverse posterior carina of MPC complete; propodeal declivity areolate, median and lateral carinae complete; anterior metapleural area transversely striate; lateral surface of MPC irregularly striate, areolate near submarginal carina of MPC. Propleuron coriaceous. Mesopleuron coriaceous, densely foveolate; subalar impression present; posterior oblique sulcus of mesopleuron present and foveolate; postepicnemial sulcus weak. WINGS. Fore wing with 2 r-rs & Rs 2 v slightly curved apically; posterior margin of pterostigma straight, prestigma absent (Fig. 4 G); Cu 2 v nebulous. Hind wing with three distal hamuli, distance between 1 st and 2 nd hamuli longer than distance between 2 nd and 3 rd hamuli. METASOMA. Metasomal terga shiny with punctures; T 1 with sparse punctures, medio-longitudinal sulcus present at basal one-third; T 2 with dense punctures, coriaceous basally (Fig. 4 F); T 3 – 4 basal half coriaceous. Metasomal sterna shiny with variably sized punctures; S 1 irregularly striate. Male Unknown.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE5A1D7CFE4187CE6A9AA222.taxon	distribution	Distribution China (Fujian) (Fig. 20).	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE5A1D7CFE4187CE6A9AA222.taxon	discussion	Remarks The examination of the holotype of S. punctulata Xu, He & Terayama, 2003 as well as other specimens showed that the mesoscutum is only impressed near the posterior margin rather than having a median mesonotal sulcus, and the antenna is sometimes remarkably thickened medially, which are the main diagnostic features of female Sulcomesitius. Females of Heterocoelia, however, have at most the mesoscutum shallowly and broadly impressed (Móczár 1984), the antenna slightly thickened and the pronotal median sulcus weak. Therefore, we transfer S. punctulata from the genus Sulcomesitius to the genus Heterocoelia.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE5D1D7FFDB585506ADFA1C9.taxon	description	urn: lsid: zoobank. org: act: 0 D 4365 A 9 - 6 ED 5 - 4203 - 9104 - A 5702 C 7171 AF Fig. 5	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE5D1D7FFDB585506ADFA1C9.taxon	diagnosis	Diagnosis This new species can be recognized by having the antenna thickened, the median pronotal sulcus foveolate, the anterior margin of PPP rounded in lateral view and the hind wing with three dorsal hamuli. This species is similar to H. varicolor sp. nov. by having the legs partly dark brown. However, it can be distinguished from it because it has the following characteristics: distance between 1 st and 2 nd hamuli 2.0 × distance between 2 nd and 3 rd hamuli (1.5 × in H. varicolor sp. nov.); apical margin of PPP rounded in lateral view (truncate in H. varicolor sp. nov.).	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE5D1D7FFDB585506ADFA1C9.taxon	etymology	Etymology The specific epithet ‘ remota ’ is Latin for ‘ removed’, referring to the distance between the 1 st and 2 nd distal hamuli being distinctly longer than the distance between the 2 nd and 3 rd distal hamuli of the hind wing.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE5D1D7FFDB585506ADFA1C9.taxon	materials_examined	Material examined Holotype CHINA – Yunnan • ♀; Naban River Watershed National Nature Reserve, Naban Tea Factory; [22.15843 ˚ N, 100.66487 ˚ E]; alt. 732 m; Flight Interception Trap; 16 May 2009; Lingzeng Meng leg.; IZCAS IOZ (E) 2059129.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE5D1D7FFDB585506ADFA1C9.taxon	description	Description Female MEASUREMENTS. Body length 6.09 mm. Fore wing length 2.67 mm. COLOURATION. Black. Mandible yellowish brown. Antenna dark brown, yellowish brown ventrally; scape, pedicel and flagellomere I reddish brown. Fore wing hyaline, brown with longitudinal light yellowish-brown band near pterostigma; veins yellowish brown to brown, pterostigma yellowish brown. Legs dark brown, trochanters, femora apically, tibiae and tarsi reddish brown. T 5 brown, T 6 – 7 yellowish brown. PUBESCENCE. Body with short, dense light yellowish-brown setae. Antenna with dense appressed setae, sparser on scape. Eye with sparse erect setae, 3.0 × diameter of an ommatidium. Wings with short, dense brown setae. Setae of metasoma longer than setae of head and mesosoma; T 1 with setae laterally, T 2 with basal glabrous triangular area dorsally. HEAD. Longer than wide, LH 1.07 × WH. Mandible with four apical teeth, ventralmost one longest. Clypeus trilobite, medio-clypeal lobe with anterior margin acute medially (Fig. 5 B); medio-clypeal carina present, not extending into frons, arched in lateral view. Antenna slightly thickened. Frons coriaceous and densely foveolate; frontal line weak (Fig. 5 B); WF 1.36 × LE. Contour of eye protruding in antero-dorsal view; LE 0.39 × LH; LE 1.74 × DEV. Anterior ocellus crossing imaginary top line of eyes in antero-dorsal view; POL 1.211 × AOL, OOL 1.09 × WOT, DPV 1.74 × DAO. Vertex coriaceous and densely foveolate; sides of head behind eyes converging posteriorly, vertex crest protruding. Occipital carina complete, occiput coriaceous. Gena coriaceous and densely foveolate, shallower foveolate ventrally; medioccipito-genal suture and hypostomal sulcus present. MESOSOMA. Dorsal pronotal area coriaceous, densely foveolate; median length 0.43 × width along posterior margin; anterolateral corner rounded; median pronotal sulcus foveolate (Fig. 5 C); lateral pronotal area obliquely striate; cervical pronotal area coarse. Mesoscutum distinctly elevated on apical two-thirds, coriaceous with dense punctures; notauli complete and converging posteriorly; parapsidal signum shallow, absent near anterior margin. Mesoscutellum coriaceous and foveolate with punctures intermediately; mesoscutum-mesoscutellar sulcus present, laterally dilated; medio-longitudinal sulcus absent (Fig. 5 D). Dorsal surface of MPC elevated; median length of dorsal surface of MPC 0.81 × its half-width; metapostnotum strongly areolate, metapostnotal median carina complete (Fig. 5 D); metapostnotal-propodeal carina complete, distinctly oblique anteriorly and converging posteriorly; dorsal propodeal area transversely striate; paraspiracular and lateral marginal carinae of MPC complete, lateral marginal carina nearly straight (Fig. 5 D); PPP long, 0.48 × median length of dorsal surface of MPC, apex rounded in lateral view; transverse posterior carina of MPC complete; propodeal declivity areolate, median and lateral carinae complete; anterior metapleural area transversely striate; lateral surface of MPC transversely striate. Propleuron coriaceous, foveolate near procoxa. Mesopleuron coriaceous, densely foveolate; subalar impression present; posterior oblique sulcus of mesopleuron present and foveolate; anterior mesofurcal pit present. WINGS. Fore wing with 2 r-rs & Rs 2 v slightly curved apically; posterior margin of pterostigma straight, prestigma absent (Fig. 5 F); Cu 2 v nebulous. Hind wing with three distal hamuli, distance between 1 st and 2 nd distal hamuli 2.0 × distance between 2 nd and 3 rd distal hamuli. METASOMA. Metasomal terga shiny with punctures; T 1 with sparse punctures, medio-longitudinal sulcus present on basal one-fourth; T 2 with dense punctures, coriaceous basally (Fig. 5 E); T 3 – 4 basal half coriaceous. Metasomal sterna shiny, with variably sized punctures. Male Unknown.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE5D1D7FFDB585506ADFA1C9.taxon	distribution	Distribution China (Yunnan) (Fig. 20).	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE401D63FDBE801F6ADFA261.taxon	description	urn: lsid: zoobank. org: act: 8 F 078314 - F 9 E 1 - 4 FAC- 8 E 57 - 6 FA 509 C 0869 A Fig. 6	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE401D63FDBE801F6ADFA261.taxon	diagnosis	Diagnosis This new species can be recognized from other species of this genus by having the foveae on the dorsal pronotal area sparser and shallower than those on the head, the length of the PPP slightly longer than its width at the base, the anterior margin of the PPP rounded in lateral view, the legs reddish brown and the hind wing with three distal hamuli.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE401D63FDBE801F6ADFA261.taxon	etymology	Etymology The specific epithet ‘ unicolor ’ is a combination of the Latin words ‘ uni- ’ and ‘ color ’ which refers to the unicoloured reddish-brown legs of this species.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE401D63FDBE801F6ADFA261.taxon	materials_examined	Material examined Holotype CHINA – Yunnan • ♀; Naban River Watershed National Nature Reserve, Mandian; 26 Apr. 2009; Lingzeng Meng leg.; IZCAS IOZ (E) 2059143.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE401D63FDBE801F6ADFA261.taxon	description	Description Female MEASUREMENTS. Body length 6.80 mm. Fore wing length 2.78 mm. COLOURATION. Black. Mandible light brown, teeth brown; maxillary and labial palpi brown, lighter ventrally. Antenna black; scape light brown, pedicel and flagellomere I brown. Fore wing hyaline, brown with longitudinal colourless band medially; veins dark brown, pterostigma yellowish brown. Legs light brown. T 4 – 7 brown or light brown. PUBESCENCE. Body with short, dense light yellowish-brown setae. Antenna with dense appressed setae, sparser on scape. Eye with sparse erect setae, 3.0 × diameter of an ommatidium. Wings with short dense brown setae. Setae of metasoma longer than setae of head and mesosoma; T 1 with setae laterally, T 2 with basal glabrous triangular area dorsally. HEAD. Longer than wide, LH 1.08 × WH. Mandible with four sharp teeth, ventralmost one longest. Clypeus trilobite; medio-clypeal lobe with anterior margin rounded (Fig. 6 B); medio-clypeal carina present, not extending into frons, arched in lateral view. Antenna thickened, ratio between length and width of flagellomeres II – V 0.92, 0.87, 0.92 and 1.04, respectively; pedicel longer than flagellomere I, length of pedicel 1.06 × length of flagellomere I. Frons coriaceous and densely foveolate; WF 1.39 × LE. Contour of eye protruding in antero-dorsal view (Fig. 6 B); LE 0.39 × LH; LE 1.46 × DEV. Anterior ocellus crossing imaginary top line of eyes in antero-dorsal view; POL 1.32 × AOL, OOL 0.84 × WOT, DPV 2.73 × DAO. Vertex coriaceous and densely foveolate; sides of head behind eyes rounded, vertex crest protruding. Occipital carina complete, occiput coriaceous. Gena coriaceous and densely foveolate, shallower foveolate ventrally; medioccipito-genal suture and hypostomal sulcus present. MESOSOMA. Dorsal pronotal area coriaceous and shallowly foveolate, median length 0.52 × width along posterior pronotal margin; anterolateral corner weakly angled (Fig. 6 B); median pronotal sulcus foveolate; lateral pronotal area areolate; cervical pronotal area coarse. Mesoscutum distinctly elevated on apical two-thirds, coriaceous with dense punctures; median mesonotal sulcus absent, only impressed in middle of apical half (Fig. 6 E); notauli complete and converging posteriorly; parapsidal signum shallow, absent near anterior margin. Mesoscutellum coriaceous and foveolate, with punctures intermediately; mesoscutum-mesoscutellar sulcus present, laterally dilated (Fig. 6 E); medio-longitudinal sulcus absent. Dorsal surface of MPC elevated; median length of dorsal surface of MPC 0.74 × its half-width; metapostnotum strongly areolate, metapostnotal median carina complete (Fig. 6 F); metapostnotal-propodeal carina complete, distinctly oblique anteriorly and converging posteriorly; dorsal propodeal area transversely striate; paraspiracular and lateral marginal carinae of MPC complete, lateral marginal carinae nearly straight; PPP long, length of PPP 0.45 × median length of dorsal surface of MPC, apical margin of PPP rounded in lateral view (Fig. 6 F); transverse posterior carina of MPC complete; propodeal declivity areolate, median and lateral carinae complete; anterior metapleural area transversely striate; lateral surface of MPC irregularly striate, areolate near submarginal carina of MPC. Propleuron coriaceous. Mesopleuron coriaceous, densely foveolate; subalar impression present; posterior oblique sulcus of mesopleuron present and foveolate; postepicnemial sulcus weak. WINGS. Fore wing with 2 r-rs & Rs 2 v slightly curved apically; posterior margin of pterostigma straight, prestigma absent (Fig. 6 H); Cu 2 v nebulous. Hind wing with three dorsal hamuli, distance between 1 st and 2 nd distal hamuli 1.54 × distance between 2 nd and 3 rd distal hamuli. METASOMA. Metasomal terga shiny with punctures; T 1 with sparse punctures, medio-longitudinal sulcus present on basal third of T 1; T 2 with dense punctures, coriaceous basally (Fig. 6 G); basal half of T 3 – 4 coriaceous. Metasomal sterna shiny with variably sized punctures; S 1 irregularly striate. Male Unknown.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE401D63FDBE801F6ADFA261.taxon	distribution	Distribution China (Yunnan) (Fig. 20).	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE421D62FDBA85686ADFA1E0.taxon	description	urn: lsid: zoobank. org: act: F 083 F 8 DC- 6699 - 46 AE- 801 B-FAFF 42634300 Fig. 7	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE421D62FDBA85686ADFA1E0.taxon	diagnosis	Diagnosis This new species can be recognized by having the median pronotal sulcus foveolate, the anterior margin of the PPP truncate in lateral view and the hind wing with three distal hamuli. This species is similar to H. remota sp. nov. for having the legs partly dark brown. It can be distinguished from the latter by having the hind wing with the distance between the 1 st and 2 nd distal hamuli 1.5 × the distance between 2 nd and 3 rd distal hamuli (2.0 in H. remota) and the apex of the PPP is truncate in lateral view (rounded in H. remota).	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE421D62FDBA85686ADFA1E0.taxon	etymology	Etymology The specific epithet is a combination of ‘ varius ’ (Latin for ‘ different’) and ‘ color ’ (Latin for ‘ tint’), and refers to the variably coloured legs of this species.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE421D62FDBA85686ADFA1E0.taxon	materials_examined	Material examined Holotype CHINA – Yunnan • ♀; Naban River Watershed National Nature Reserve, Mengsong Town, Benggangxinzhai Plantation; [22.18418 ˚ N, 100.64986 ˚ E]; alt. 797 m; 12 May 2008; Trap; Lingzeng Meng leg.; IZCAS IOZ (E) 2059142.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE421D62FDBA85686ADFA1E0.taxon	description	Description Female MEASUREMENTS. Body length 6.19 mm. Fore wing length 3.04 mm. COLOURATION. Black. Mandible reddish brown. Antenna dark brown, yellowish brown ventrally; scape, pedicel and flagellomere I reddish brown. Fore wing hyaline, brown with longitudinal light yellowish band medially; veins yellowish brown or brown, pterostigma yellowish brown. Legs dark brown, trochanters, femora apically, tibiae and tarsi reddish brown. T 4 brown with posterior margin yellowish brown; T 5 – 7 yellowish brown. PUBESCENCE. Body with short dense light yellowish-brown setae. Antenna with dense appressed setae, sparser on scape (Fig. 7 C). Eye with sparse erect setae, 3.0 × diameter of an ommatidium. Wings with short dense brown setae. Setae of metasoma longer than setae of head and mesosoma; T 1 with setae laterally, T 2 with basal glabrous triangular area dorsally. HEAD. Longer than wide, LH 1.06 × WH. Clypeus trilobite; medio-clypeal lobe with anterior margin slightly acute medially; medio-clypeal carina present, not extending into frons, arched in lateral view. Antenna thickened. Frons coriaceous and densely foveolate (Fig. 7 B); WF 1.36 × LE. Contour of eye slightly protruding in antero-dorsal view; LE 0.40 × LH; LE 1.56 × DEV. Anterior ocellus crossing imaginary top line of eyes in antero-dorsal view; POL 0.96 × AOL, OOL 0.97 × WOT, DPV 2.21 × DAO. Vertex coriaceous and densely foveolate; sides of head behind eyes nearly straight, converging posteriorly; vertex crest protruding. Gena coriaceous and densely foveolate. Occipital carina complete. MESOSOMA. Dorsal pronotal area coriaceous, densely foveolate; median length 0.44 × width along posterior pronotal margin (Fig. 7 D); anterolateral corner rounded; median pronotal sulcus incomplete; lateral pronotal area obliquely striate; cervical pronotal area coarse. Mesoscutum distinctly elevated on apical two-thirds, coriaceous with dense punctures (Fig. 7 E); median mesonotal sulcus absent, impressed in middle of apical half (Fig. 7 E); notauli complete and converging posteriorly; parapsidal signum shallow, absent near anterior margin. Mesoscutellum coriaceous, foveolate with puncture intermediately; mesoscutum-mesoscutellar sulcus present, laterally dilated (Fig. 7 E); medio-longitudinal sulcus absent. Dorsal surface of MPC elevated; median length of dorsal surface of MPC 0.89 × its half-width; metapostnotum strongly areolate, metapostnotal median carina complete (Fig. 7 F); metapostnotal-propodeal carina complete, distinctly oblique anteriorly and converging posteriorly; dorsal propodeal area transversely striate; paraspiracular and lateral marginal carinae of MPC complete, lateral marginal carina nearly straight (Fig. 7 G); PPP long, 0.47 × median length of dorsal surface of MPC, apex truncate in lateral view; transverse posterior carina of MPC complete; propodeal declivity areolate, median and lateral carinae complete; anterior metapleural area transversely striate; lateral surface of MPC transversely striate, areolate near submarginal carina of MPC. Mesopleuron coriaceous, densely foveolate; subalar impression present; posterior oblique sulcus of mesopleuron present and foveolate. WINGS. Fore wing with 2 r-rs & Rs 2 v slightly curved apically; posterior margin of pterostigma straight, prestigma absent (Fig. 7 H); Cu 2 v nebulous. Hind wing with three distal hamuli, distance between 1 st and 2 nd distal hamuli 1.50 × distance between 2 nd and 3 rd distal hamuli. METASOMA. Metasomal terga shiny with punctures; T 1 with sparse punctures, medio-longitudinal sulcus present at basal one-fourth; T 2 with dense punctures, coriaceous basally (Fig. 7 G); T 3 – 4 basal half coriaceous. Metasomal sterna shiny with variably sized punctures; S 1 irregularly striate. Male Unknown.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE421D62FDBA85686ADFA1E0.taxon	distribution	Distribution China (Yunnan) (Fig. 20).	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE451D64FD91801E6964A5A8.taxon	diagnosis	Diagnosis Male antenna with erect setae longer than width of flagellomere. Head and pronotum foveolate. PPP present. Hypopygium with posterior branches broad and short, one-quarter length of hypopygium (Fig. 8 G). Male genitalia with dorsal harpe filamentary (Fig. 8 H), aedeagus usually with apical sickle process.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE451D64FD91801E6964A5A8.taxon	biology_ecology	Host Unknown.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE451D64FD91801E6964A5A8.taxon	distribution	Distribution Afrotropical and Oriental Regions.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE451D69FD8285806970A449.taxon	description	urn: lsid: zoobank. org: act: 7 E 226425 - 0 DEA- 43 DE-A 965 - 1663 A 167 B 2 B 1 Fig. 8	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE451D69FD8285806970A449.taxon	diagnosis	Diagnosis This new species can be recognized by having the antennal setae longer than the width of the flagellomeres; median pronotal sulcus present and foveolate; median mesonotal sulcus absent, at most with a long fovea near the posterior margin; PPP short, basal width nearly equal to its length; the hypopygium longer than wide, with the posterior branches short (Fig. 8 G); genitalia with dorsal harpe filamentary, and narrower than ventral harpe. This species is similar to I. exilisulcus sp. nov. by sharing the simple aedeagus without a sickle-shaped process apically. However, I. afoveatus sp. nov. can be distinguished by having the mesoscutum and mesoscutellum not foveolate (more or less foveolate in I. exilisulcus); the mesoscutellum without a medio-longitudinal suture (present in I. exilisulcus); the basal width of PPP nearly equal to its length (two-thirds of length in I. exilisulcus); T 2 with sparse punctures (denser in I. exilisulcus).	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE451D69FD8285806970A449.taxon	etymology	Etymology The specific epithet ‘ afoveatus ’ is a combination of the Latin words ‘ a ’ (for ‘ without’) and ‘ foveatus ’ and refers to the absence of foveae on the mesoscutum and mesoscutellum.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE451D69FD8285806970A449.taxon	materials_examined	Material examined Holotype CHINA – Fujian • ♂; Minqing, Xiongjiang Town; 13 – 17 Jul. 2005; Zaifu Xu leg.; ZJUH 200609178. Paratype (1 ♂) CHINA – Hainan • 1 ♂; Bawangling; 7 – 11 Jul. 2006; Liqiong Weng leg.; ZJUH 200700151.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE451D69FD8285806970A449.taxon	description	Description Male MEASUREMENTS (n = 2). WF 1.78 – 1.83 × LE; LE 0.99 – 1.05 × DEV; POL 1.27 – 1.55 × AOL; OOL 0.87 – 0.92 × WOT; DPV 1.51 – 1.56 × DAO; length of hypopygium 1.41 – 1.53 × its width; length of posterior branches of hypopygium 0.18 – 0.19 × length of hypopygium. COLOURATION. Black, metasoma mainly brown, T 3 yellowish brown. Mandible yellowish brown; maxillary and labial palpi brown. Antenna brown, scape dark brown. Legs brown, trochanters, femora apically, tibiae apically, and tarsi yellowish brown. Fore wing hyaline; veins and pterostigma pale yellowish brown. PUBESCENCE. Body with short sparse setae. Antenna with erect setae, nearly equal to width of flagellomere (Fig. 8 B). Eye with sparse short setae, nearly 3.0 × diameter of an ommatidium. Wings with short, dense brown setae. T 1 nearly glabrous; T 2 with sparse setae laterally. HEAD. Slightly shorter than wide, LH 0.98 × WH. Mandible with three apical teeth, ventralmost one largest. Clypeus trilobite; medio-clypeal lobe with apical margin acute medially; medio-clypeal carina present, not extending into frons, arched in lateral view. Antenna thin, flagellomeres cylindrical. Frons coriaceous and densely foveolate, frontal line weak (Fig. 8 A); WF 1.83 × LE. Eye small, LE 0.35 × LH; contour of eye distinctly protruding in antero-dorsal view (Fig. 8 A); LE 0.99 × DEV. Anterior ocellus partly anterior to imaginary top line of eyes in antero-dorsal view; POL 1.27 × AOL, OOL 0.87 × WOT, DPV 1.51 × DAO. Vertex coriaceous and densely foveolate, sides of head behind eyes distinctly converging posteriorly (Fig. 8 A). Occipital carina present, vertex crest nearly straight. MESOSOMA. Dorsal pronotal area coriaceous and densely foveolate, median length 0.33 × width of posterior pronotal margin, anterolateral corner rounded; median pronotal sulcus foveolate; lateral pronotal area obliquely striate; cervical pronotal area coarse (Fig. 8 C). Mesoscutum coriaceous and sparsely foveolate; median mesonotal sulcus absent; notauli complete and converging posteriorly (Fig. 8 C); parapsidal signum shallow and absent near anterior margin; lateral area of mesoscutum depressed anteriorly (Fig. 8 C). Mesoscutellum weakly coriaceous with shallow punctures; mesoscutum-mesoscutellar sulcus present, laterally dilated (Fig. 8 C). Dorsal surface of MPC elevated; median length of dorsal surface of MPC equal to its half-width (Fig. 8 D); metapostnotum strongly areolate, metapostnotal median carina complete, slightly tortuous (Fig. 8 D); metapostnotal-propodeal carina complete, slightly oblique anteriorly and converging posteriorly; dorsal propodeal area densely striate; paraspiracular carinae of MPC complete; lateral marginal carina of MPC complete and protruding (Fig. 8 D); PPP short, 0.1 × median length of dorsal surface of MPC, apex acute and bent upward in lateral view; transverse posterior carina of MPC complete; propodeal declivity coriaceous, areolate near transverse posterior carina, median and lateral carinae complete; lateral surface of MPC sparsely striate, areolate near submarginal carina of the MPC. Propleuron coriaceous. Mesopleuron coriaceous, densely and shallowly foveolate; subalar impression present; posterior oblique sulcus of mesopleuron present and foveolate. WINGS. Fore wing with 2 r-rs & Rs 2 v slightly curved apically (Fig. 8 F); posterior margin of pterostigma nearly straight, prestigma absent; Cu 2 v absent. Hind wing with three distal hamuli. METASOMA. Metasomal terga shiny with punctures; T 1 with sparse punctures, medio-longitudinal sulcus present on anterior third of T 1; T 2 with sparse punctures, weakly coriaceous basally (Fig. 8 E). Metasomal sterna shiny with variably sized punctures; S 1 coriaceous. Length of hypopygium 1.4 × its width; anterior stalk with anterior margin straight (Fig. 8 G); posterior margin bilobate (Fig. 8 G), posterior branches broad and short with long setae, 0.2 × length of hypopygium. MALE GENITALIA. Harpe bilobate, dorsal harpe filamentary and straight (Fig. 8 H); ventral harpe distinctly broader than dorsal harpe and with long setae apically (Fig. 8 I). Cuspis bilobate with long setae; digitus bent and papillate apically, with setae basally; aedeagus bottle-shaped with apical margin rounded (Fig. 8 H). Female Unknown.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE451D69FD8285806970A449.taxon	distribution	Distribution China (Fujian, Hainan) (Fig. 20).	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE481D6AFD888388697DA795.taxon	description	urn: lsid: zoobank. org: act: 233 C 0 A 49 - 2466 - 44 BA-A 7 B 9 - 433819 F 1 EF 0 D Fig. 9	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE481D6AFD888388697DA795.taxon	diagnosis	Diagnosis This new species can be recognized by having the antennal setae longer than the width of the flagellomere; median pronotal sulcus present and not distinctly foveolate; median mesonotal sulcus absent, at most with a fovea or foveolate sulcus near posterior margin; the length of PPP longer than its width at the base; the hypopygium longer than wide, with posterior branches short; the genitalia with the dorsal harpe filamentary and narrower than the ventral harpe. This species is similar to I. afoveatus sp. nov. by sharing the simple aedeagus (without a sickle-shaped process apically). However, it can be distinguished as follows: the mesoscutellum has a medio-longitudinal suture, the PPP is longer than its width at the base and T 2 has denser punctures than in I. afoveatus.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE481D6AFD888388697DA795.taxon	etymology	Etymology The specific epithet ‘ exilisulcus ’ is a combination of the Latin words ‘ exilis ’ (‘ weak’) and ‘ sulcus ’ (‘ groove’) and refers to the mesoscutellum of this species having a weak medio-longitudinal suture.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE481D6AFD888388697DA795.taxon	materials_examined	Material examined Holotype CHINA – Hainan • ♂; Bawangling; 7 – 11 Jul. 2006; Liqiong Weng leg.; ZJUH 200700110. Paratypes (9 ♂♂) CHINA – Hainan: 1 ♂; same collection data as holotype; ZJUH 200700135 • 6 ♂♂; Bawangling; 7 – 11 Jul. 2006; Jingxian Liu leg.; ZJUH 200700012, 200700028, 200700040, 200700041, 200700075, 200700084 • 1 ♂; Jianfengling; 12 – 15 Jul. 2006; Tianfei Chen leg.; ZJUH 200700786. – Yunnan • 1 ♂; Mt. Gaoligong; 20 – 21 Jul. 2006; Jie Zeng leg.; ZJUH 200701080.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE481D6AFD888388697DA795.taxon	description	Description Male MEASUREMENTS (n = 10). Body length 2.85 – 4.01 mm; fore wing length 1.81 – 2.49 mm; LH 0.95 – 1.03 × WH; WF 1.75 – 1.91 × LE; LE 0.95 – 1.11 × DEV; POL 1.44 – 1.68 × AOL; OOL 0.86 – 0.99 × WOT; DPV 1.65 – 2.3 × DAO; median length of pronotum 0.42 – 0.48 × its width along posterior margin; length of MPC 0.93 – 1.12 × its half-width; length of PPP 0.19 – 0.27 × median length of dorsal surface of MPC; length of hypopygium 1.28 – 1.51 × its width; length of hypopygial posterior branches 0.21 – 0.26 × length of hypopygium. COLOURATION. Black. Mandible brown, teeth yellowish brown; maxillary and labial palpi brown. Antenna yellowish brown, scape brown. Legs brown, trochanters, tibiae apically, and tarsi yellowish brown. Prothorax and mesothorax dark brown. Fore wing hyaline, light brown with longitudinal colourless band medially; veins and pterostigma pale light brown. Metasoma mainly brown, partly dark brown. PUBESCENCE. Body with short setae. Antenna with erect setae, nearly equal to width of flagellomere (Fig. 9 C). Eye with short setae, nearly 4.0 × diameter of an ommatidium. Wings with short dense brown setae. T 1 nearly glabrous, T 2 with sparse setae laterally. HEAD. Slightly shorter than wide, LH 0.95 × WH. Mandible with three apical teeth, ventral larger than upper two. Clypeus trilobite; medio-clypeal lobe with anterior margin acute medially; medio-clypeal carina present, not extending into frons, arched in lateral view. Antenna thin, flagellomeres cylindrical. Frons coriaceous and densely foveolate, frontal line weak (Fig. 9 B); WF 1.79 × LE. Eye small, LE 0.35 × LH; contour of eye distinctly protruding in antero-dorsal view; LE 1.06 × DEV. Anterior ocellus partly anterior to imaginary top line of eyes in antero-dorsal view (Fig. 9 B); POL 1.60 × AOL, OOL 0.98 × WOT, DPV 2.23 × DAO. Vertex coriaceous and densely foveolate, sides of head behind eyes distinctly converging posteriorly, vertex crest nearly straight. Gena coriaceous and densely foveolate, ventral surface with fovea shallower and sparser, medioccipito-genal suture present. Occipital carina present. MESOSOMA. Dorsal pronotal area coriaceous and densely foveolate, median length 0.33 × width along posterior pronotal margin; anterolateral corner rounded; median pronotal sulcus nearly complete (Fig. 9 D); lateral pronotal area obliquely striate; cervical pronotal area coarse. Mesoscutum elevated, coriaceous and shallowly foveolate; median mesonotal sulcus absent; notauli complete and converging posteriorly; parapsidal signum shallow and absent near anterior margin; lateral area of mesoscutum depressed anteriorly (Fig. 9 D). Mesoscutellum coriaceous, distinctly foveolate; mesoscutum-mesoscutellar sulcus present, laterally dilated; medio-longitudinal suture present (Fig. 9 D). Dorsal surface of MPC elevated; median length of dorsal surface of MPC equal to its half-width; metapostnotum strongly areolate, metapostnotal median carina complete; metapostnotal-propodeal carina complete, distinctly oblique anteriorly and converging posteriorly; dorsal propodeal area striate; paraspiracular and lateral marginal carinae of MPC complete, lateral marginal carina nearly straight; PPP present, 0.27 × median length of dorsal surface of MPC, apex acute and not bent upward in lateral view (Fig. 9 E); transverse posterior carina of MPC complete; propodeal declivity weakly striate, areolate near transverse posterior carina, median and lateral carinae complete; lateral surface of MPC irregularly striate, areolate near submarginal carina of MPC. Propleuron coriaceous. Mesopleuron coriaceous, densely foveolate; subalar impression present; posterior oblique sulcus of mesopleuron present and foveolate; ventral surface without fovea, with short carina near mesocoxa; anterior mesofurcal pit and mesodiscrimen present. WINGS. Fore wing with 2 r-rs & Rs 2 v slightly curved apically; posterior margin of pterostigma straight, prestigma absent; Cu 2 v absent. Hind wing with three distal hamuli. METASOMA. Metasomal terga shiny with punctures; T 1 with sparse punctures, medio-longitudinal sulcus present at basal one-third; T 2 with punctures, coriaceous basally (Fig. 9 F). Metasomal sterna shiny with variably sized punctures; S 1 irregularly striate. Length of hypopygium 1.28 × width; anterior stalk narrowed with anterior margin straight (Fig. 9 H); posterior margin bilobate, posterior branches broad and short with long setae, 0.23 × length of hypopygium. MALE GENITALIA. Harpe bilobate, dorsal harpe filamentary and straight (Fig. 9 I); ventral harpe distinctly broader than dorsal harpe (Fig. 9 J), with long setae apically. Cuspis bilobate with long setae; digitus bent and papillate apically, with setae basally; aedeagus bottle-shaped, apical margin acute medially (Fig. 9 I). Female Unknown.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE481D6AFD888388697DA795.taxon	distribution	Distribution China (Hainan, Yunnan) (Fig. 20).	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE4B1D6AFDC7856B692FA3E6.taxon	materials_examined	Material examined None in this study.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE4B1D6AFDC7856B692FA3E6.taxon	distribution	Distribution China (Hong Kong) (Fig. 20).	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE4B1D6AFD89831469DFA21E.taxon	type_taxon	Type species: Metrionotus pappi Móczár, 1970: 202.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE4B1D6AFD89831469DFA21E.taxon	diagnosis	Diagnosis Malar space convergent anteriorly. Male antenna with flagellomeres short. Mesoscutum without median mesonotal sulcus. PPP weak. Male hypopygium wider than long, anterior median stalk of hypopygium acute apically, posterior branches short. Male genitalia with dorsal harpe club-shaped and as wide as ventral harpe.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE4B1D6AFD89831469DFA21E.taxon	biology_ecology	Host Unknown.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE4B1D6AFD89831469DFA21E.taxon	distribution	Distribution Afrotropical, Oriental, and Palaearctic Regions.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE4B1D6DFD93870369DFA7E9.taxon	diagnosis	Diagnosis Body foveolate. Anterior ocellus usually entirely anterior to imaginary top line of eyes in antero-dorsal view. Dorsal pronotal area with median pronotal sulcus; dorsal pronotal area usually with anterolateral corner weakly projected. Mesoscutum with median mesonotal sulcus usually developed. PPP usually robust. Male hypopygium longer than wide, basal part of hypopygium with lateral and anterior margin incurved; posterior branches lobose and long. Male genitalia with dorsal harpe usually S-shaped, ventral harpe narrower than dorsal one.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE4B1D6DFD93870369DFA7E9.taxon	biology_ecology	Host Unknown.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE4B1D6DFD93870369DFA7E9.taxon	distribution	Distribution Afrotropical, Oriental, and Palaearctic Regions.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE4C1D6EFD9E84686ADFA4AC.taxon	description	urn: lsid: zoobank. org: act: 1 E 90 D 2 B 8 - 2938 - 4 D 0 B-BEAA- 5 CE 53006645 D Fig. 10	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE4C1D6EFD9E84686ADFA4AC.taxon	diagnosis	Diagnosis This new species can be recognized by having the median pronotal sulcus developed, the PPP distinctly longer than its width at the base, the apical margin of the PPP distinctly acute in dorsal view and the hind wing with four distal hamuli. This species is similar to S. impressus Xu, He & Terayama, 2003. However, the new species can be distinguished by having the following characteristics: length of eye equal to width of frons (WF 1.33 × LE in S. impressus), legs partly dark brown (reddish brown in S. impressus), and T 4 – 5 dark brown (brown or reddish brown in S. impressus), punctures on lateral surface of T 2 as dense as on dorsal surface (denser on lateral surface than on dorsal surface in S. impressus).	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE4C1D6EFD9E84686ADFA4AC.taxon	etymology	Etymology The specific epithet is a combination of ‘ angustus ’ (Latin for ‘ narrow’) and ‘ frons ’, referring to the comparatively narrow frons.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE4C1D6EFD9E84686ADFA4AC.taxon	materials_examined	Material examined Holotype CHINA – Yunnan • ♀; Xishuangbanna, Naban River Watershed National Nature Reserve; [22.12961 ˚ N, 100.66612 ˚ E]; alt. 746 m; 6 Jun. 2008; A. Weigel leg.; IZCAS IOZ (E) 2059138.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE4C1D6EFD9E84686ADFA4AC.taxon	description	Description Female MEASUREMENTS (holotype, Fig. 10 A). Body length 7.06 mm. Fore wing length 3.16 mm. COLOURATION. Body black. Mandible brown; maxillary palp and labial palp brown. Antenna black, yellowish brown ventrally; scape, pedicel, and flagellomeres I – II brown. Fore wing bi-banded, veins brown or yellowish brown, pterostigma yellowish brown. Legs dark brown; trochanters, tibiae and tarsi yellowish brown. T 1 – 3 with posterior margin yellowish brown, metasomal segments 3 – 5 dark brown with posterior margin yellowish brown, metasomal segments 6 – 7 yellowish brown. PUBESCENCE. Body with short setae; maxillary palp, labial palp and antenna with dense appressed setae (Fig. 10 C). Metasomal segments with setae longer than setae of head and mesosoma; T 1 with sparse setae posterolaterally; T 2 densely setose posteriorly and laterally with basal triangular area glabrous. Wings with dense, short yellowish-brown setae. HEAD. Longer than wide, LH 1.07 × WH. Mandible with 4 apical teeth, ventralmost one sharp and largest. Clypeus trilobite; medio-clypeal lobe with anterior margin acute medially (Fig. 10 B); medio-clypeal carina not extending into frons, arched in lateral view. Antenna distinctly thickened, at least flagellomeres II – IX shorter than wide. Frons coriaceous and densely foveolate (Fig. 10 B); WF 1.01 × LE. Contour of eye protruding in antero-dorsal view, LE 0.46 × LH, LE 2.08 × DEV. Anterior ocellus entirely anterior to imaginary top line of eyes; POL 1.33 × AOL, OOL 0.93 × WOT, DPV 2.36 × DAO. Sides of head behind eyes rounded and slightly converging posteriorly. Vertex coriaceous and densely foveolate, vertex crest slightly protruding. Malar space 1.21 × distance between posterior ocelli and vertex crest. Gena coriaceous, densely foveolate, with shallow foveolae ventrally. Occipital carina complete, occiput coriaceous with shallow punctures. Medioccipito-genal suture present. Hypostomal sulcus present. MESOSOMA. Dorsal pronotal area coriaceous and densely foveolate, median length 0.51 × width of posterior pronotal margin (Fig. 10 D); median pronotal sulcus present; lateral pronotal area obliquely strigate; cervical pronotal area coarse. Mesoscutum with notauli complete, parapsidal signum shallow but complete; lateral area of mesoscutum coriaceous with shallow punctures and depressed anteriorly. Mesoscutellum coriaceous with punctures, densely foveolate; mesoscutum-mesoscutellar sulcus present, laterally dilated. Median length of dorsal surface of MPC 1.04 × its half-width; metapostnotum strongly areolate, metapostnotal median carina complete, metapostnotal-propodeal carina complete and converging posteriorly; dorsal propodeal area transversely striate; paraspiracular and lateral marginal carinae of MPC complete and parallel (Fig. 10 E); length of PPP 0.32 × median length of dorsal surface of MPC (Fig. 10 E); propodeal declivity areolate, median and lateral carinae complete; lateral surface of MPC areolate, anterior metapleural area transversely striate. Propleuron coriaceous and shallowly foveolate. Mesopleuron coriaceous and densely foveolate; subalar impression weak; posterior oblique sulcus of mesopleuron present and foveolate. WINGS. Fore wing with 2 r-rs & Rs 2 v slightly curved apically, 0.80 × Sc + R 2 v; posterior margin of pterostigma straight, prestigma absent (Fig. 10 G); Cu 2 v present as yellowish-brown trace. Hind wing with four distal hamuli. METASOMA. Metasomal terga shiny with punctures; T 1 with tiny, sparse punctures; T 2 with dense punctures separated by 1 – 5 × their diameter, posterior margin without punctures (Fig. 10 F); T 3 – 5 with three semi-circular impressions on posterior margin. Metasomal sterna shiny with punctures; S 1 irregularly striate, metapostnotal median carina weak; S 2 – 3 with dense punctures separated by less than 2.0 × their diameter. Male Unknown.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE4C1D6EFD9E84686ADFA4AC.taxon	distribution	Distribution China (Yunnan) (Fig. 20).	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE4F1D53FE4C83D569FAA291.taxon	description	Fig. 11	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE4F1D53FE4C83D569FAA291.taxon	diagnosis	Diagnosis Sulcomesitius breviculus (Xu, He & Terayama, 2003) can be distinguished from other species of the genus Sulcomesitius by having the anterolateral corner of the pronotum rounded, the length of PPP at most equal to its basal width, the hypopygium distinctly narrower anteriorly, and the dorsal harpe of the genitalia 8 - shaped (Fig. 11 I).	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE4F1D53FE4C83D569FAA291.taxon	materials_examined	Material examined Holotype CHINA – Zhejiang • ♂; Kaihua County, Gutian Mountain; 22 Jul. 1992; Hong Wu leg.; ZJUH 949066. Paratypes (2 ♂♂) CHINA – Zhejiang • 1 ♂; same collection data as for holotype; ZJUH 949082 • 1 ♂; same collection data as for preceding but 17 Jul. 1992; Yun Ma leg.; ZJUH 923783. Other material (18 ♂♂) CHINA – Zhejiang • 1 ♂; same collection data as for holotype but 18 Jul. 1992; Yun Ma leg.; ZJUH 923914 • 3 ♂♂; same collection data as for preceding but Xuexin Chen leg.; ZJUH 923359, 923443, 923551 • 1 ♂; same collection data as for preceding but 19 Aug. 2003; Wuqing Fan leg.; ZJUH 20047497 • 2 ♂♂; same collection data as for preceding but 2 Jul. 2005; Min Shi leg.; ZJUH 200601769, 200601849 • 7 ♂♂; same collection data as for preceding but 3 Jul. 2005; Qiong Wu leg.; ZJUH 200602041, 200602046, 200602047, 200602048, 200602053, 200602065, 200602103 • 1 ♂; Longquan City, Fengyang Mountain; 7 Aug. 2003; Xiaoxia Yu leg.; ZJUH 20034610 • 1 ♂; Linan County, Qingliang Mountain; 10 Aug. 2005; Hongying Zhang leg.; ZJUH 200603355 • 1 ♂; Linan County, Qianmutian Glacier; 25 Jul. 2011; Shengnan Song leg.; ZJUH 201503630. – Yunnan • 1 ♂; Tengchong City, Gaoligong Mountain; 20 – 21 Jul. 2006; Jie Zeng leg.; ZJUH 200701076.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE4F1D53FE4C83D569FAA291.taxon	description	Re-description Male MEASUREMENTS (n = 21). Body length 3.71 – 5.32 mm; length of fore wing 2.19 – 3.27 mm; LH 0.96 – 1.03 × WH; WF 1.21 – 1.41 × LE; LE 0.42 – 0.47 × LH; LE 1.44 – 1.82 × DEV; POL 1.54 – 2.0 × AOL; OOL 0.76 – 0.93 × WOT; DPV 0.99 – 1.66 × DAO; malar space 0.86 – 1.73 × distance between posterior ocelli and vertex crest; median length of pronotum 0.38 – 0.44 × width of pronotum along posterior margin; median length of dorsal surface of MPC 0.9 – 1.0 × its half-width; length of PPP 0.12 – 0.22 × median length of dorsal surface of MPC; length of 2 r-rs & Rs 2 v 0.85 – 1.08 × length of Sc + R 2 v; length of posterior branches of hypopygium 0.42 – 0.53 × length of hypopygium. COLOURATION. Body dark bronw to black. Antenna yellowish brown, darker apically. Fore wing light brown, slightly darker on apical half; veins brown or yellowish brown, pterostigma yellowish brown. Legs yellowish brown, coxae and femora brown. Metasoma brown to dark brown, anterior margin of T 2 with pair of yellowish-brown semicircular spots. PUBESCENCE. Body with short setae. Maxillary and labial palpi with dense, short setae. Antenna with short suberect setae, length of setae 0.25 × width of flagellomere II (Fig. 11 C). Eye with sparse short setae, nearly 5.0 × diameter of an ommatidium. Wings with short dense brown setae. Metasomal segments with setae longer than setae of head and mesosoma, T 1 with sparse setae posterolaterally, T 2 with basal triangular area glabrous. HEAD. Slightly longer than wide, LH 1.04 × WH. Clypeus trilobite; medio-clypeal lobe with anterior margin acute medially; medio-clypeal carina not extending into frons, arched in lateral view. Antenna slender, length of flagellomere II 1.88 × its width, 0.75 × length of flagellomere I. Frons coriaceous and densely foveolate (Fig. 11 B); WF 1.34 × LE. Contour of eye protruding in antero-dorsal view; LE 0.43 × LH, LE 1.58 × DEV. Anterior ocellus partly anterior to imaginary top line of eyes; POL 1.57 × AOL, OOL 0.93 × WOT, DPV 1.27 × DAO. Vertex coriaceous and densely foveolate, sides of head behind eyes rounded and slightly converging posteriorly, vertex crest protruding. Malar space 0.89 × distance between posterior ocelli and vertex crest. Gena coriaceous, densely foveolate, with shallow foveolae ventrally. Occipital carina complete, occiput coriaceous with shallow punctures. Medioccipito-genal suture present. MESOSOMA. Dorsal pronotal area coriaceous and densely foveolate, median length of dorsal pronotal area 0.42 × width along posterior pronotal margin; anterolateral corner rounded; median pronotal sulcus present (Fig. 11 D); lateral pronotal area obliquely striate; cervical pronotal area coarse. Mesoscutum elevated, coriaceous with shallow punctures; median mesonotal sulcus absent, with a fovea near posterior margin; basal half with medio-line carinate (Fig. 11 D); notauli complete and converging posteriorly, parapsidal signum shallow and absent near anterior margin. Mesoscutellum slightly elevated, coriaceous, densely foveolate with punctures; mesoscutum-mesoscutellar sulcus present, laterally slightly dilated (Fig. 11 E). Median length of dorsal surface of MPC 0.93 × its half-width; metapostnotum strongly areolate, metapostnotal median carina complete, metapostnotal-propodeal carina complete and converging posteriorly (Fig. 11 E); dorsal propodeal area transversely striate; paraspiracular and lateral marginal carinae of MPC complete; length of PPP 0.15 × median length of dorsal surface of MPC; transverse posterior carina of MPC complete; propodeal declivity areolate, median and lateral carinae complete; anterior metapleural area shiny, weakly striate; lateral surface of MPC with submarginal carina and posterior carina, areolate. Propleuron coriaceous, propleural epicoxal sulcus present. Mesopleuron coriaceous and densely foveolate; subalar impression weak; posterior oblique sulcus of mesopleuron present and foveolate; postepicnemial sulcus present; ventral surface of mesopleuron coriaceous and irregularly foveolate, area between mesocoxa with three large foveae; anterior mesofurcal pit present and deep. WINGS. Fore wing with 2 r-rs & Rs 2 v slightly curved apically, 1.12 × Sc + R 2 v; posterior margin of pterostigma protruding, prestigma absent (Fig. 11 G); Cu 2 v present as yellowish-brown trace. Hind wing with four distal hamuli. METASOMA. Metasomal terga shiny with punctures; T 1 with tiny sparse punctures, medio-longitudinal sulcus present on basal one-third; T 2 with sparse punctures separated by more than their own diameter; basal half of T 3 – 4 weakly coriaceous with punctures posteriorly. Metasomal sterna shiny, with variably sized punctures; S 1 irregularly striate, metapostnotal median carina weak; S 2 with dense, variably sized punctures. Hypopygium longer than wide; basal part of hypopygium with lateral margin distinctly incurved (Fig. 11 H); posterior branches distinctly narrower than median notch, inner margin with long setae, length of posterior branches 0.45 × length of hypopygium. MALE GENITALIA. Harpe bilobate, basal half of dorsal harpe filamentary, apical half 8 - shaped (Fig. 11 I), longer than ventral harpe. Inner margin of gonostipe with series of sharp parallel carinae (Fig. 11 J). Cuspis with apical margin incurved; digitus bent and papillate apically, with setae basally; aedeagus bottle-shaped (Fig. 11 I). Female Unknown.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE4F1D53FE4C83D569FAA291.taxon	distribution	Distribution China (Zhejiang, Yunnan) (Fig. 20).	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE4F1D53FE4C83D569FAA291.taxon	discussion	Remarks This species was included by Xu et al. (2003) in the genus Heterocoelia because it has the anterior corners of the clypeus rectangular laterally, the mesoscutum without a median mesonotal sulcus and the apex of the PPP rounded in lateral view. However, the generic identification of the males is mainly based on the shape of the hypopygium (Argaman 2003; Azevedo et al. 2018). After studying the males, we transfer H. breviculus from the genus Heterocoelia to the genus Sulcomesitius because the hypopygium of this species is longer than wide, with the posterior branches lobose and 0.45 × the length of the hypopygium (posterior branches short and broad in Heterocoelia); the head and pronotum densely foveolate; the pronotum with a median pronotal sulcus and PPP present (although comparatively short compared to other species of Sulcomesitius).	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE721D53FE2285E06F76A05C.taxon	materials_examined	Material examined None in this study.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE721D53FE2285E06F76A05C.taxon	distribution	Distribution China (Hong Kong, Taiwan) (Fig. 20), Burma and Mediterranean Subregion.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE721D54FE3C87A6695EA4C1.taxon	description	Fig. 12	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE721D54FE3C87A6695EA4C1.taxon	materials_examined	Material examined Holotype CHINA – Guangxi • ♀; Longsheng County, Huaping Nature Reserve; [25.39 ˚ N, 109.55 ˚ E]; 23 Jun. 1982; Junhua He leg.; ZJUH 823323. Other material (1 ♀) CHINA – Guizhou • 1 ♀; Leigong Mountain, Xiaodanjiang; 4 – 5 Jun. 2005; Jingxian Liu leg.; ZJUH 20059386.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE721D54FE3C87A6695EA4C1.taxon	description	Re-description Female MEASUREMENTS (n = 2). Body length 6.35 – 6.38 mm; length of fore wing 3.02 – 3.57 mm; LH 1.03 – 1.04 × WH; WF 1.32 – 1.33 × LE; LE 1.63 – 1.77 × DEV; POL 1.32 – 1.34 × AOL; OOL 0.79 – 0.81 × WOT; DPV 2.02 – 2.11 × DAO; malar space 1.07 – 1.47 × distance between posterior ocelli and vertex crest; median length of dorsal surface of MPC 0.90 – 0.92 × its half-width; length of PPP 0.45 – 0.46 × median length of dorsal surface of MPC. COLOURATION. Body black. Mandible brown, maxillary palpi and labial palpi brown. Antenna black, yellowish brown ventrally, scape, pedicel and flagellomeres I – II brown. Fore wing bi-banded, veins brown, pterostigma yellowish brown. Legs brown. T 2 posteriorly and T 3 – 4 yellowish brown. PUBESCENCE. Body with short setae; maxillary palp, labial palp and antenna with dense appressed setae (Fig. 12 C). Eye with sparse erect setae, 5.0 × diameter of an ommatidium. Setae of metasoma longer than setae of head and mesosoma; T 1 with sparse setae posterolaterally; T 2 with dense setae, basal triangular area glabrous. Wings with dense short yellowish-brown setae. HEAD. Length nearly equal to width, LH 1.03 × WH. Mandible with four apical teeth, ventralmost largest with apex sharp. Clypeus trilobite; medio-clypeal lobe with anterior margin acute medially; medio-clypeal carina not extending into frons, arched in lateral view. Antenna distinctly thickened, at least flagellomeres II – IX shorter than wide. Frons coriaceous and densely foveolate (Fig. 12 B); WF 1.33 × LE. Contour of eye distinctly protruding in antero-dorsal view, LE 0.41 × LH, LE 1.63 × DEV. Anterior ocellus entirely anterior to imaginary top line of eyes; POL 1.34 × AOL, OOL 0.79 × WOT, DPV 2.11 × DAO. Sides of head behind eyes rounded and slightly converging posteriorly. Vertex coriaceous and densely foveolate, vertex crest slightly protruding. Malar space 1.2 × distance between posterior ocelli and vertex crest. Gena coriaceous, densely foveolate. Occipital carina complete, occiput coriaceous with shallow punctures. Medioccipito-genal suture present. Hypostomal sulcus present. MESOSOMA. Dorsal pronotal area coriaceous and densely foveolate, median length 0.57 × width along posterior pronotal margin; median pronotal sulcus present (Fig. 12 D); lateral pronotal area obliquely striate; cervical pronotal area coarse. Mesoscutum distinctly elevated, coriaceous with dense punctures (Fig. 12 E); median mesonotal sulcus incomplete, present on posterior one-third (Fig. 12 E); notauli complete, incurved posteriorly (Fig. 12 E); parapsidal signum incomplete, absent on anterior one-third; lateral area of mesoscutum depressed anteriorly. Mesoscutellum coriaceous, densely foveolate; mesoscutum-mesoscutellar sulcus present, laterally dilated (Fig. 12 E). Median length of dorsal surface of MPC 0.90 × its half-width; metapostnotum strongly areolate, metapostnotal median carina complete, metapostnotal-propodeal carina complete and converging posteriorly; dorsal propodeal area transversely striate; paraspiracular and lateral marginal carina of MPC complete, lateral marginal carinae straight (Fig. 12 F); propodeal declivity areolate, median and lateral carinae complete; length of PPP 0.45 × median length of dorsal surface of MPC. Lateral surface of MPC areolate, anterior metapleural area transversely striate. Propleuron coriaceous and foveolate. Mesopleuron coriaceous and densely foveolate; subalar impression weak; posterior oblique sulcus of mesopleuron present and foveolate; postepicnemial sulcus weak; ventral surface of mesopleuron coriaceous and shallowly foveolate. WINGS. Fore wing with 2 r-rs & Rs 2 v slightly curved apically, 0.79 × length of Sc + R 2 v; posterior margin of pterostigma straight, prestigma absent (Fig. 12 H); Cu 2 v present as yellowish-brown trace. Hind wing with four distal hamuli. METASOMA. Metasomal terga shiny with punctures; T 1 with tiny sparse punctures; T 2 with dense, variably sized punctures, posterior margin without punctures (Fig. 12 G); T 3 – 5 with three semicircular impressions on posterior margin. Metasomal sterna shiny with punctures; S 1 irregularly striate, metapostnotal median carina weak; S 2 – 3 with dense punctures separated by less than 2.0 × their diameter. Male Unknown.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE721D54FE3C87A6695EA4C1.taxon	distribution	Distribution China (Guangxi, Guizhou) (Fig. 20).	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE751D59FD9D82306ADFA79E.taxon	description	urn: lsid: zoobank. org: act: 5 C 37 C 810 - F 7 CA- 4185 - ABC 3 - CF 0 D 39400001 Fig. 13	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE751D59FD9D82306ADFA79E.taxon	diagnosis	Diagnosis This new species can be recognized by its having the median pronotal sulcus present and not distinctly foveolate, the median mesonotal sulcus of the mesoscutum absent, the apex of the PPP acute in lateral view and the hypopygium of the male with the anterior margin broad and the posterior branches long and lobose. This species can be separated from other species of the genus Sulcomesitius by having the antenna with short proclinate setae; dorsal pronotal area with anterolateral corner weakly projected; median mesonotal sulcus of mesoscutum absent; hypopygium with posterior branches wider than median notch.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE751D59FD9D82306ADFA79E.taxon	etymology	Etymology The specific epithet is a combination of ‘ latus ’ (Latin for ‘ wide’) and ‘ bilobatus ’ (Latin for ‘ bilobed’) because both posterior branches of the hypopygium are wider than its posterior median notch.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE751D59FD9D82306ADFA79E.taxon	materials_examined	Material examined Holotype CHINA – Yunnan • ♂; Yingjiang, Tongbiguan; 23 Jun. 2005; Jingxian Liu leg.; ZJUH 200609145.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE751D59FD9D82306ADFA79E.taxon	description	Description Male MEASUREMENTS. Body length 4.4 mm. Fore wing length 2.8 mm. COLOURATION. Body black. Mandible brown, apical half yellowish brown, teeth brown; maxillary and labial palpi yellowish brown. Antenna brown, yellowish brown ventrally. Fore wing light brown, veins brown or yellowish brown, pterostigma yellowish brown. Legs brown, trochanters, tibiae apically and tarsi yellowish brown. T 1 partly dark brown, lateral and posterior margin of T 2 dark brown, T 3 – 4 brown with black basally, T 5 – 6 brown; S 1 black, S 2 – 4 brown posteriorly, S 5 – 6 brown. PUBESCENCE. Body with short setae. Maxillary palp and labial palp with dense, short setae. Antenna with short, proclinate setae, 0.25 × width of flagellomere II (Fig. 13 C). Eye with sparse short setae, nearly 4.0 × diameter of an ommatidium. Wings with short, dense yellowish-brown setae. Setae of metasoma longer than setae of head and mesosoma, T 1 with sparse setae posterolaterally, T 2 with basal triangular area glabrous. HEAD. Length equal to width, LH 1.0 × WH. Clypeus trilobite; medio-clypeal lobe with apical margin rounded; medio-clypeal carina not extending into frons, arched in lateral view. Antenna slender, length of flagellomere III 1.85 × length of flagellomere II. Frons coriaceous and densely foveolate (Fig. 13 B); LE 0.42 × LH, WF 1.31 × LE. Contour of eye distinctly protruding in antero-dorsal view (Fig. 13 B); LE 1.46 × DEV. Anterior ocellus partly anterior to imaginary top line of eyes; POL 1.36 × AOL, OOL 0.94 × WOT, DPV 1.38 × DAO. Vertex coriaceous and densely foveolate, sides of head behind eyes rounded and slightly converging posteriorly, vertex crest slightly protruding. Malar space 1.0 × distance between posterior ocelli and vertex crest. Gena coriaceous, densely foveolate, with shallow foveolae ventrally. Occipital carina complete, occiput coriaceous with shallow punctures. Medioccipito-genal suture present. MESOSOMA. Dorsal pronotal area coriaceous and densely foveolate, median length 0.36 × width along posterior pronotal margin, anterolateral corner weakly projected; median pronotal sulcus present (Fig. 13 D); lateral pronotal area obliquely strigate; cervical pronotal area coarse. Mesoscutum elevated, coriaceous with shallow punctures; median mesonotal sulcus absent (Fig. 13 E); notauli complete and converging posteriorly, parapsidal signum shallow and absent near anterior margin. Mesoscutellum weakly coriaceous with shallow punctures, apical two-thirds foveolate; mesoscutum-mesoscutellar sulcus present, laterally dilated (Fig. 13 E). Median length of dorsal surface of MPC 0.95 × its half-width; metapostnotum strongly areolate, metapostnotal median carina complete, metapostnotal-propodeal carina complete and converging posteriorly; dorsal propodeal area transversely striate; paraspiracular and lateral marginal carinae of MPC complete; length of PPP 0.29 × median length of dorsal surface of MPC (Fig. 13 E); transverse posterior carina of MPC complete; propodeal declivity weakly areolate, median and lateral carinae complete; anterior metapleural area transversely striate; lateral surface of MPC with submarginal carina and posterior carina, anterior half of lateral surface of MPC transversely striate, posterior half of lateral surface of MPC areolate. Propleuron coriaceous, foveolate near procoxa, propleural epicoxal sulcus present. Mesopleuron coriaceous and densely foveolate; subalar impression weak; posterior oblique sulcus of mesopleuron present and foveolate; postepicnemial sulcus present; ventral surface of mesopleuron coriaceous with punctures, area between mesocoxa with large foveola; anterior mesofurcal pit present. WINGS. Fore wing with 2 r-rs & Rs 2 v slightly curved apically, 0.85 × length of Sc + R 2 v; posterior margin of pterostigma protruding, prestigma absent (Fig. 13 G); Cu 2 v present as yellowish-brown trace. Hind wing with three distal hamuli, distance between 1 st and 2 nd hamuli longer than distance between 2 nd and 3 rd hamuli. METASOMA. Metasomal terga shiny with punctures; T 1 with tiny sparse punctures, medio-longitudinal sulcus present on basal one-third; T 2 with punctures denser than on T 1, posterior margin with scattered punctures. Metasomal sterna shiny, with variably sized punctures; S 1 irregularly striate, metapostnotal median carina weak. Hypopygium longer than wide; lateral margin of basal part of hypopygium slightly incurved, anterior margin incurved (Fig. 13 H); posterior branches wider than median notch, 0.41 × length of hypopygium, with long setae apically. MALE GENITALIA. Harpe bilobate, dorsal harpe S-shaped and longer than ventral harpe (Fig. 13 I). Cuspis with apical margin incurved (Fig. 13 J); digitus bent and papillate apically, with setae basally; aedeagus bottle-shaped (Fig. 13 I). Female Unknown.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE751D59FD9D82306ADFA79E.taxon	distribution	Distribution China (Yunnan) (Fig. 20).	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE781D5AFD8C80E46ADAA416.taxon	description	urn: lsid: zoobank. org: act: 44 B 36209 - 219 F- 46 AD- 9526 - 3 CA 3 F 58 AC 114 Fig. 14	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE781D5AFD8C80E46ADAA416.taxon	diagnosis	Diagnosis This new species can be recognized by having the median pronotal sulcus weak, mesoscutum with median impression near posterior margin, PPP distinctly longer than its width at base with apex acute in lateral view, length of posterior branches of the male hypopygium 0.51 × length of hypopygium and dorsal harpe of genitalia longer than ventral harpe. This species is similar to S. zhaoi (Xu & He, 2006). However, the new species can be distinguished by having the mesoscutum shallowly punctured (densely punctured in S. zhaoi), the PPP distinctly longer than its width at the base (nearly equal to its width at the base in S. zhaoi), the apex of the PPP acute in dorsal view (blunt in S. zhaoi), and T 2 with shallower and sparser punctures than in S. zhaoi.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE781D5AFD8C80E46ADAA416.taxon	etymology	Etymology The specific epithet is derived from ‘ sparsus ’ (Latin for ‘ scattered’), referring to the sparse punctures of T 2.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE781D5AFD8C80E46ADAA416.taxon	materials_examined	Material examined Holotype CHINA – Hainan • ♂; Baisha County, Yinggeling; 1 – 2 May 2008; Jingxian Liu leg.; ZJUH 200800040. Paratypes (3 ♂♂) CHINA – Hainan • 1 ♂; Bawangling; 7 – 11 Jul. 2006; Jingxian Liu leg.; ZJUH 200700063 • 1 ♂; Baisha, Nankai County; 29 Apr. 2008; Jingxian Liu leg.; ZJUH 200800018 • 1 ♂; Baisha County, Yinggeling; 1 – 2 May 2008; Jingxian Liu leg.; ZJUH 200800030.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE781D5AFD8C80E46ADAA416.taxon	description	Description Male MEASUREMENTS (n = 4). Body length 3.42 – 3.81 mm; length of fore wing 2.02 – 2.47 mm; LH 1.0 – 1.01 × WH; WF 1.39 – 1.54 × LE; LE 0.39 – 0.41 × LH; LE 1.23 – 1.38 × DEV; POL 1.44 – 1.53 × AOL; OOL 0.83 – 1.0 × WOT; DPV 1.17 – 1.72 × DAO; malar space 1.11 – 1.25 × distance between posterior ocelli and vertex crest; median length of dorsal surface of MPC 0.92 – 0.95 × its half-width; length of PPP 0.38 – 0.42 × median length of dorsal surface of MPC; length of 2 r-rs & Rs 2 v 0.81 – 0.91 × length of Sc + R 2 v; length of posterior branches 0.5 – 0.53 × length of hypopygium. COLOURATION. Body black or dark brown. Mandible yellowish brown, teeth brown; maxillary and labial palpi brown. Antenna brown, darker apically and lighter ventrally. Eye brown or dark brown. MPC with lateral margin brown, posterior projection brown apically. Fore wing light brown, darker on apical half; veins dark brown or brown, pterostigma brown. Legs dark brown, trochanters, femora apically, tibiae apically and tarsi brown, protarsus yellowish brown. T 1 with posterior and lateral margin dark brown, posterior margin of T 2 and T 3 brown, T 4 brown, posterior margin of T 4 and T 5 yellowish brown; S 2 – 4 yellowish brown apically. PUBESCENCE. Body with short setae. Maxillary and labial palpi with dense, short setae. Antenna with short, suberect yellowish-brown setae, 0.35 × width of flagellomere II (Fig. 14 C). Eye with sparse, short setae, nearly 5.0 × diameter of an ommatidium. Wings with short, dense yellowish-brown setae. Setae of metasoma longer than setae of head and mesosoma, T 1 with sparse setae posterolaterally, T 2 with basal triangular area glabrous. HEAD. Length equal to width, LH 1.0 × WH. Clypeus trilobite; medio-clypeal lobe with apical margin slightly acute medially (Fig. 14 B); medio-clypeal carina not extending into frons, arched in lateral view. Antenna slender, length of flagellomere III 2.0 × length of flagellomere II. Frons coriaceous and densely foveolate (Fig. 14 B); WF 1.46 × LE. Contour of eye slightly protruding in antero-dorsal view (Fig. 14 B); LE 0.40 × LH, LE 1.38 × DEV. Anterior ocellus partly anterior to imaginary top line of eyes; POL 1.52 × AOL, OOL 1.0 × WOT, DPV 1.58 × DAO. Vertex coriaceous and densely foveolate, sides of head behind eyes rounded and slightly converging posteriorly, vertex crest slightly protruding. Malar space 0.89 × distance between posterior ocelli and vertex crest. Gena coriaceous, densely foveolate. Occipital carina complete. Medioccipito-genal suture present. MESOSOMA. Dorsal pronotal area coriaceous and densely foveolate, median length 0.49 × width along posterior pronotal margin, anterolateral corner weakly projected; median pronotal sulcus weak (Fig. 14 D); lateral pronotal area obliquely striate; cervical pronotal area coarse. Mesoscutum distinctly elevated, coriaceous, with variably sized shallow punctures; median mesonotal sulcus absent, with median impression on apical half (Fig. 14 E); notauli complete and converging posteriorly, parapsidal signum shallow and absent near anterior margin; lateral area of mesoscutum depressed anteriorly. Mesoscutellum weakly coriaceous with shallow punctures, shallowly foveolate; mesoscutum-mesoscutellar sulcus present, laterally dilated (Fig. 14 E). Median length of dorsal surface of MPC less than its half-width (0.92); metapostnotum strongly areolate, metapostnotal median carina complete, metapostnotal-propodeal carina complete and converging posteriorly; dorsal propodeal area transversely striate; paraspiracular and lateral marginal carinae of MPC complete, lateral marginal carina nearly straight (Fig. 14 E); length of PPP 0.38 × median length of dorsal surface of MPC (Fig. 14 E); transverse posterior carina of MPC complete; propodeal declivity weakly areolate, median and lateral carinae complete; anterior metapleural area transversely striate; lateral surface of MPC with submarginal carina and posterior carina, irregularly striate. Mesopleuron coriaceous and densely foveolate; posterior oblique sulcus of mesopleuron present and foveolate; postepicnemial sulcus present; ventral surface of mesopleuron coriaceous, with irregular-sized punctures, area between mesocoxa with large foveae; postepicnemial sulcus; anterior mesofurcal pit present. WINGS. Fore wing with 2 r-rs & Rs 2 v slightly curved apically, 0.92 × as long as Sc + R 2 v; posterior margin of pterostigma protruding, prestigma absent (Fig. 14 G); Cu 2 v present as yellowish-brown trace. Hind wing with three distal hamuli, distance between 1 st and 2 nd distal hamuli longer than distance between 2 nd and 3 rd distal hamuli. METASOMA. Metasomal terga shiny, with punctures; T 1 with tiny sparse punctures, medio-longitudinal sulcus present at basal one-third; T 2 with dense punctures separated less than 3.0 × their diameter, posterior margin with scattered punctures. Metasomal sterna shiny, with variably sized punctures; S 1 irregularly striate, metapostnotal median carina weak; S 2 with dense, variably sized punctures. Hypopygium longer than wide; basal part of hypopygium with lateral and anterior margin incurved (Fig. 14 H); posterior branches 0.51 × length of hypopygium, distinctly narrower than median notch, with long setae apically. MALE GENITALIA. Harpe bilobate, dorsal harpe S-shaped and longer than ventral harpe (Fig. 14 I). Cuspis with apical margin incurved; digitus bent and papillate apically, with setae basally (Fig. 14 J); aedeagus bottle-shaped (Fig. 14 I). Female Unknown.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE781D5AFD8C80E46ADAA416.taxon	distribution	Distribution China (Hainan) (Fig. 20).	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE7B1D5FFE2E836B69B2A517.taxon	description	Fig. 15	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE7B1D5FFE2E836B69B2A517.taxon	materials_examined	Material examined Holotype CHINA – Fujian • ♂; Fuzhou; 19 Sept. 1990; Xiufu Zhao leg.; ZJUH 967700. Paratype (1 ♂) CHINA – Fujian • 1 ♂; Fuzhou; 13 Aug. 1989; Changming Liu leg.; ZJUH 966312. Other material (6 ♂♂) CHINA – Fujian • ♂; holotype of Heterocoelia sinensis; Fuzhou; 3 Jul. 1990; Naiquan Lin leg.; ZJUH 967530 • 1 ♂; Fuzhou, Jin Mountain; 30 May 2001; Naiquan Lin leg.; ZJUH 202300002 • 1 ♂; Minqing, Xiongjiang Town; 13 – 17 Jul. 2005; Zaifu Xu leg.; ZJUH 200609182. – Hainan • 2 ♂♂; Bawangling; 7 – 11 Jul. 2006; Jingxian Liu leg.; ZJUH 200700036, 200700078 • 1 ♂; Baisha County, Yinggeling; 1 – 2 May 2008; Jingxian Liu leg.; ZJUH 200800043. Re-description Male MEASUREMENTS (n = 8). TL 4.84 – 6.21 mm; length of fore wing 2.46 – 3.29 mm; LH 1.0 – 1.03 × WH; WF 1.53 – 1.75 × LE; LE 1.06 – 1.33 × DEV; POL 1.33 – 1.61 × AOL; OOL 0.92 – 0.99 × WOT; DPV 1.55 – 2.39 × DAO; malar space 1.0 – 1.38 × distance between posterior ocelli and vertex crest; median length of dorsal surface of MPC 0.73 – 0.86 × its half-width; length of PPP 0.37 – 0.61 × median length of dorsal surface of MPC; 2 r-rs & Rs 2 v 0.71 – 0.96 × as long as Sc + R 2 v; length of posterior branches of hypopygium 0.45 – 0.5 × length of hypopygium. COLOURATION. Body black. Mandible yellowish brown or brown, apical half yellowish brown; maxillary and labial palpi brown or dark brown. Antenna dark brown with ventral surface brown. Eye brown or dark brown. Fore wing light brown, darker on apical half; veins dark brown or brown, pterostigma brown. Legs dark brown or blackish brown, trochanters, femora apically, tibiae and tarsi brown and yellowish brown. MPC with lateral margin brown, PPP brown apically. T 1 with lateral and posterior margin dark brown, S 2 apically and S 3 posteriorly brown, T 4 – 5 brown with posterior margin yellowish brown. PUBESCENCE. Body with short setae. Maxillary and labial palpi with dense, short setae. Antenna with short proclinate yellowish-brown setae, 0.22 × width of flagellomere II (Fig. 15 C). Eye with sparse, short setae, nearly 5.0 × diameter of an ommatidium. Wings with short, dense yellowish-brown setae. Setae of metasoma longer than setae of head and mesosoma, T 1 with sparse setae posterolaterally, T 2 with basal triangular area glabrous. HEAD. Slightly longer than wide, LH 1.03 × WH. Clypeus trilobite; medio-clypeal lobe with apical margin acute medially; medio-clypeal carina not extending into frons, arched in lateral view. Antenna slender, length of flagellomere III 2.0 × its width. Frons coriaceous and densely foveolate (Fig. 15 B); WF 1.67 × LE. Contour of eye protruding in antero-dorsal view (Fig. 15 B); LE 0.35 – 0.39 × LH; LE 1.09 × DEV. Anterior ocellus entirely anterior to imaginary top line of eyes; POL 1.36 × AOL, OOL 0.98 × WOT, DPV 2.39 × DAO. Vertex coriaceous and densely foveolate, sides of head behind eyes rounded and slightly converging posteriorly, vertex crest slightly protruding. Malar space 1.22 × distance between posterior ocelli and vertex crest. Gena coriaceous, densely foveolate and shallower ventrally. Occipital carina complete, occiput coriaceous with shallow punctures. Medioccipito-genal suture present. MESOSOMA. Dorsal pronotal area coriaceous and densely foveolate, median length 0.45 × width along posterior pronotal margin, anterolateral corner projected; median pronotal sulcus weak (Fig. 15 D); lateral pronotal area obliquely striate; cervical pronotal area coarse. Mesoscutum elevated, coriaceous with shallow punctures; median mesonotal sulcus absent (Fig. 15 D); notauli complete and converging posteriorly, parapsidal signum shallow and absent near anterior margin. Mesoscutellum weakly coriaceous with shallow punctures, apical two-thirds foveolate; mesoscutum-mesoscutellar sulcus present, laterally dilated (Fig. 15 E); median line slightly elevated. Median length of dorsal surface of MPC 0.86 × its half-width; metapostnotum strongly areolate, metapostnotal median carina complete, metapostnotal-propodeal carina complete and converging posteriorly; dorsal propodeal area transversely striate; paraspiracular and lateral marginal carinae of MPC complete; length of PPP 0.37 × median length of dorsal surface of MPC (Fig. 15 E); transverse posterior carina of MPC complete; propodeal declivity weakly areolate, median and lateral carinae complete; anterior metapleural area transversely striate; lateral surface of MPC with submarginal carina and posterior carina, irregularly striate. Propleuron coriaceous, shallowly foveolate, propleural epicoxal sulcus present. Mesopleuron coriaceous and densely foveolate; subalar impression weak; posterior oblique sulcus of mesopleuron present and foveolate; postepicnemial sulcus present; ventral surface of mesopleuron coriaceous, with irregular-sized punctures, area between mesocoxae with large foveae; anterior mesofurcal pit present. WINGS. Fore wing with 2 r-rs & Rs 2 v slightly curved apically, 0.81 × as long as Sc + R 2 v; posterior margin of pterostigma protruding, prestigma absent (Fig. 15 G); Cu 2 v present as yellowish-brown trace. Hind wing with three distal hamuli, distance between 1 st and 2 nd distal hamuli longer than distance between 2 nd and 3 rd distal hamuli. METASOMA. Metasomal terga shiny, with punctures; T 1 with tiny, sparse punctures, medio-longitudinal sulcus present at basal one-third; T 2 with dense punctures separated by less than 3.0 × their diameter, posterior margin with scattered punctures (Fig. 15 F). Metasomal sterna shiny, with variably sized punctures; S 1 irregularly striate, metapostnotal median carina weak; S 2 with dense, variably sized punctures. Hypopygium longer than wide; lateral and anterior margin of basal part of hypopygium incurved (Fig. 15 H); posterior branches 0.5 × length of hypopygium, distinctly narrower than median notch, with long setae apically. MALE GENITALIA. Harpe bilobate, dorsal harpe S-shaped and longer than ventral harpe (Fig. 15 I). Cuspis with apical margin incurved; digitus bent and papillate apically, with setae basally (Fig. 15 J); aedeagus bottle-shaped (Fig. 15 I). Female Unknown.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE7B1D5FFE2E836B69B2A517.taxon	distribution	Distribution Oriental Region (China: Fujian, Hainan) (Fig. 20).	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE7B1D5FFE2E836B69B2A517.taxon	discussion	Remarks After an examination of the type series of Heterocoelia zhaoi Xu & He, 2006 and H. sinensis Xu & He, 2006, we transfer both taxa to the genus Sulcomesitius because their head and pronotum are densely foveolate; the hypopygium of the male is longer than wide, with the posterior branches long, at least 0.45 × the length of the hypopygium; the dorsal harpe of the genitalia is S-shaped. Heterocoelia sinensis used to be distinguished from H. zhaoi by having the length of PPP 0.60 × the median length of the dorsal surface of MPC (0.4 × in H. zhaoi) and T 2 densely and deeply punctate throughout, with uniform punctures (broadly granulate basally, distinctly punctate posteriorly, with uniform punctures in H. zhaoi) (Xu & He 2006). Our re-examination of the holotypes of both taxa suggests their conspecificity since both holotypes have the PPP 0.50 × the median length of the dorsal surface of MPC. As for the difference in sculpture of T 2, this may be an artefact; therefore, it is insufficient to retain H. sinensis as a valid species.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE7E1D5FFD8D826A6964A3F1.taxon	diagnosis	Diagnosis Head and pronotum densely foveolate. Malar space usually as long as VOL. Male antenna with suberect setae, nearly equal to half-width of flagellomere; anterior ocellus of male usually entirely anterior to the imaginary top line of eyes in antero-dorsal view. Dorsal pronotal area usually with anterolateral corner distinctly projected. T 2 usually with scattered punctures. Hypopygium of male with posterior branches long and slender, lateral and anterior margin of basal part of hypopygium distinctly incurved. Hind wing with at least four distal hamuli.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE7E1D5FFD8D826A6964A3F1.taxon	biology_ecology	Host Unknown.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE7E1D5FFD8D826A6964A3F1.taxon	distribution	Distribution Afrotropical and Oriental Regions.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE7F1D40FDAE801F6AC0A2E4.taxon	description	urn: lsid: zoobank. org: act: 686744 DD-BF 0 C- 4087 - 91 C 9 - 380 D 8 B 877806 Fig. 16	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE7F1D40FDAE801F6AC0A2E4.taxon	diagnosis	Diagnosis This new species is similar to Z. cambodianus (Móczár, 1976) in having the anterolateral corner of the dorsal pronotal area distinctly projected and the PPP shorter than half-length of the dorsal surface of MPC. However, it can be distinguished by having the length of PPP nearly equal to the width at its base and the apical margin of PPP acute in lateral view (rounded in Z. cambodianus); the pterostigma with the posterior margin straight (protruding in Z. cambodianus); the posterior branches of the hypopygium 0.36 × the length of the hypopygium (0.48 × the length of the hypopygium in Z. cambodianus).	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE7F1D40FDAE801F6AC0A2E4.taxon	etymology	Etymology The specific epithet refers to the short and acute PPP in lateral view; ‘ acutus ’ is Latin for ‘ sharp’.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE7F1D40FDAE801F6AC0A2E4.taxon	materials_examined	Material examined Holotype CHINA – Sichuan • ♂; Panzhihua, Ertan; 30 Jul. 2006; Zhilei Gao leg.; ZJUH 200611246.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE7F1D40FDAE801F6AC0A2E4.taxon	description	Description Male MEASUREMENTS. Body length 6.21 mm. Length of forewing 3.57 mm. COLOURATION. Black. Mandible brown. Antenna brown, darker apically. Fore wing hyaline, light brown; veins yellowish brown or brown, pterostigma yellowish brown. Legs dark brown, trochanters, femora apically, tibiae and tarsi brown. T 1 posteriorly, T 2 apical half and T 4 – 5 yellowish brown. PUBESCENCE. Body with short setae. Maxillary and labial palpi with dense, short setae. Antenna with short, suberect setae, 0.16 × width of flagellomere II (Fig. 16 C). Eye with sparse, short setae, nearly 6.0 × diameter of an ommatidium. Wings with short, dense brown setae. Setae of metasoma longer than setae of head and mesosoma, T 1 nearly glabrous, T 2 with basal triangular area glabrous. HEAD. Shorter than wide, LH 0.94 × WH. Clypeus trilobite, anterior margin of medio-clypeal lobe acute medially; medio-clypeal carina not extending into frons, arched in lateral view. Antenna slender. Frons coriaceous and densely foveolate, median frontal line absent (Fig. 16 B); WF 1.59 × LE. Contour of eye protruding in antero-dorsal view (Fig. 16 B); LE 0.41 × LH, LE 1.27 × DEV. Anterior ocellus partly anterior to imaginary top line of eyes; POL 1.58 × AOL, OOL 0.98 × WOT, DPV 2.32 × DAO. Vertex coriaceous and densely foveolate, sides of head behind eyes nearly parallel, vertex crest protruding. Gena coriaceous, densely foveolate, with shallow foveolae ventrally. Occipital carina complete. MESOSOMA. Dorsal pronotal area coriaceous and foveolate, median length 0.49 × width along posterior pronotal margin, anterolateral corner distinctly projected; median pronotal sulcus present (Fig. 16 D); lateral pronotal area obliquely striate; cervical pronotal area coarse. Mesoscutum elevated, coriaceous with dense punctures; median mesonotal sulcus present on apical one-third, foveolate (Fig. 16 E); notauli complete and converging posteriorly, parapsidal signum shallow, absent near anterior margin. Mesoscutellum coriaceous and foveolate; mesoscutum-mesoscutellar sulcus arched, laterally dilated (Fig. 16 E). Median length of dorsal surface of MPC equal to its half-width; metapostnotum strongly areolate, metapostnotal median carina complete, metapostnotal-propodeal carina complete and converging posteriorly (Fig. 16 E); dorsal propodeal area transversely striate; paraspiracular and lateral marginal carinae of MPC complete; length of PPP 0.32 × median length of dorsal surface of MPC (Fig. 16 E); transverse posterior carina of MPC complete; propodeal declivity weakly transversely striate, areolate near posterior marginal carina, median and lateral carinae complete; anterior metapleural weakly striate; lateral surface of MPC areolate, submarginal and posterior carinae present. Mesopleuron coriaceous and densely foveolate; subalar impression weak; posterior oblique sulcus of mesopleuron present and foveolate; postepicnemial sulcus present; ventral surface of mesopleuron coriaceous and shallowly foveolate; anterior mesofurcal pit present and deep. WINGS. Fore wing with 2 r-rs & Rs 2 v slightly curved apically, 1.12 × as long as Sc + R 2 v; posterior margin of pterostigma protruding, prestigma absent (Fig. 16 G); Cu 2 v present as light yellowish-brown trace. Hind wing with four distal hamuli. METASOMA. Metasomal terga shiny, with punctures; T 1 with shiny, sparse punctures, medio-longitudinal sulcus present at basal one-third; T 2 with dense punctures. Metasomal sterna shiny, with variably sized punctures; S 1 irregularly striate, metapostnotal median carina weak; S 2 with dense, variably sized punctures. Length of hypopygium 2.18 × its width; lateral and anterior margin of basal part of hypopygium distinctly incurved (Fig. 16 H); posterior branches 0.36 × length of hypopygium, distinctly narrower than median notch; apical margin of posterior branches and bottom of posterior median notch with long setae. MALE GENITALIA. Harpe bilobate, dorsal harpe filamentary (Fig. 16 I), ventral harpe finger-like, with long setae (Fig. 16 J). Cuspis with apical margin incurved; digitus bent and papillate apically, with setae basally; aedeagus bottle-shaped (Fig. 16 I). Female Unknown.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE7F1D40FDAE801F6AC0A2E4.taxon	distribution	Distribution China (Sichuan) (Fig. 20).	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE611D45FDF0841E6F10A2BE.taxon	description	Fig. 17	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE611D45FDF0841E6F10A2BE.taxon	materials_examined	Material examined (8 ♂♂) CHINA – Henan • 1 ♂; Baotianman; 13 – 15 Jul. 1998; Xuexin Chen leg.; ZJUH 991803. – Zhejiang • 1 ♂; paratype of Sulcomesitius rectus; Qingyuan; 26 – 27 Jul. 1985; Quancong Wu leg.; ZJUH 851898 • 1 ♂; Tianmu Mountain; 27 Jul. 1988; Ying Qian leg.; ZJUH 940245 • 1 ♂; Songyang; 27 Jun. 1994; Hanlin Chen leg.; ZJUH 954190 • 1 ♂; Xitianmu Mountain; 28 Jul. 2003; Xiaoxia Yu leg.; ZJUH 20039255. – Fujian • 1 ♂; Wuyi Mountain; 20 Jul. 1985; Naiquan Lin leg.; ZJUH 968094. – Guangdong • 1 ♂; Xinfeng, Yunjin Mountain; 19 Jun. 2002; Zaifu Xu leg.; ZJUH 20029190. – Guangxi • 1 ♂; holotype of Sulcomesitius rectus; Liuzhou; 9 Jun. 1982; Junhua He leg.; ZJUH 822502. Re-description Male MEASUREMENTS (n = 8). LH 0.93 – 1.0 × WH; WF 1.36 – 1.48 × LE; LE 0.41 – 0.45 × LH; LE 1.36 – 1.75 × DEV; POL 1.58 – 1.97 × AOL; OOL 0.82 – 1.03 × WOT; DPV 0.63 – 1.72 × DAO; malar space 1.52 – 2.08 × distance between posterior ocelli and vertex crest; median length of dorsal pronotal area 0.4 – 0.49 × width along posterior pronotal margin; median length of dorsal surface of MPC 0.96 – 1.05 × its half-width; length of PPP 0.29 – 0.33 × median length of dorsal surface of MPC; length of posterior branches of hypopygium 0.42 – 0.5 × length of hypopygium. COLOURATION. Body black. Mandible yellowish brown, teeth reddish brown; maxillary and labial palpi yellowish brown. Antenna brown, darker apically. Fore wing hyaline, yellowish brown; veins yellowish brown, pterostigma light brown. Legs brown, coxae dark brown. T 1 dark brown, T 2 black with posterior and lateral margin brown; T 3 – 4 brown. PUBESCENCE. Body with short setae. Maxillary and labial palpi with dense, short setae. Antenna with short, suberect setae, 0.47 × width of flagellomere II (Fig. 17 C). Eye with sparse, short setae, nearly 5.0 × diameter of an ommatidium. Wings with short, dense brown setae. Setae of metasoma longer than setae of head and mesosoma, T 1 nearly glabrous, T 2 with basal triangular area glabrous. HEAD. Shorter than wide, LH 0.96 × WH. Clypeus trilobite, medio-clypeal lobe with apical margin acute medially; medio-clypeal carina not extending into frons, arched in lateral view. Antenna slender. Frons coriaceous and densely foveolate, median frontal line absent (Fig. 17 B); WF 1.48 × LE. Contour of eye protruding in antero-dorsal view; LE 0.43 × LH, LE 1.59 × DEV. Anterior ocellus partly anterior to imaginary top line of eyes; POL 1.58 × AOL, OOL 0.95 × WOT, DPV 1.38 × DAO. Vertex coriaceous and densely foveolate, sides of head behind eyes nearly parallel, vertex crest protruding. Gena coriaceous, densely foveolate, with shallow foveolae ventrally. Occipital carina complete. Medioccipito-genal suture present. MESOSOMA. Dorsal pronotal area coriaceous and densely foveolate, median length 0.47 × width along posterior pronotal margin, anterolateral corner projected; median pronotal sulcus foveolate (Fig. 17 D); lateral pronotal area obliquely striate; cervical pronotal area coarse. Mesoscutum elevated, coriaceous, with dense punctures; median mesonotal sulcus fovea-like, present near posterior margin (Fig. 17 E); notauli complete and converging posteriorly, parapsidal signum shallow and absent near anterior margin. Mesoscutellum slightly elevated, coriaceous and foveolate; mesoscutum-mesoscutellar sulcus present, laterally dilated. Median length of dorsal surface of MPC nearly equal to its half-width; metapostnotum strongly areolate, metapostnotal median carina complete, metapostnotal-propodeal carina complete and converging posteriorly; dorsal propodeal area transversely striate; paraspiracular and lateral marginal carinae of MPC complete (Fig. 17 F); length of PPP 0.29 × median length of dorsal surface of MPC; transverse posterior carina of MPC complete; propodeal declivity areolate, median and lateral carinae complete; anterior metapleural area shiny, weakly striate; lateral surface of MPC areolate, submarginal and posterior carinae present. Propleuron coriaceous, foveolate near procoxa. Mesopleuron coriaceous and densely foveolate; subalar impression weak; posterior oblique sulcus of mesopleuron present and foveolate; postepicnemial sulcus present; ventral surface of mesopleuron coriaceous and shallowly foveolate; anterior mesofurcal pit present and deep. WINGS. Fore wing with 2 r-rs & Rs 2 v slightly curved apically, 1.12 × length of Sc + R 2 v; posterior margin of pterostigma protruding, prestigma absent (Fig. 17 H); Cu 2 v present as light yellowish-brown trace. Hind wing with four distal hamuli. METASOMA. Metasomal terga shiny, with punctures; T 1 with shiny sparse punctures, medio-longitudinal sulcus present at basal one-third; T 2 with punctures (Fig. 17 G). Metasomal sterna shiny, with variably sized punctures; S 1 irregularly striate, metapostnotal median carina weak; S 2 with dense, variably sized punctures. Hypopygium longer than wide; lateral and anterior margin of basal part of hypopygium distinctly incurved (Fig. 17 I); length of posterior branches 0.48 × length of hypopygium, distinctly narrower than median notch; apical margin of posterior branches and bottom of posterior median notch with long setae. MALE GENITALIA. Harpe bilobate, dorsal harpe filamentary (Fig. 17 J), ventral harpe finger-like, with long setae (Fig. 17 K). Cuspis with apical margin incurved; digitus bent and papillate apically, with setae basally; aedeagus bottle-shaped (Fig. 17 J). Female Unknown.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE611D45FDF0841E6F10A2BE.taxon	distribution	Distribution China (Henan, Zhejiang, Fujian, Guangdong, Guangxi) (Fig. 20), Cambodia, Laos.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE611D45FDF0841E6F10A2BE.taxon	discussion	Remarks The specimens (numbered ZJUH 822502, 851898) belonging to the type series of Sulcomesitius rectus Xu, He & Terayama, 2003, the specimen numbered 954190 being identified as S. vechti Móczár, 1979 and the specimens numbered 991803 and 940245 as S. laoensis Móczár, 1976 were included in the genus Sulcomesitius by Xu et al. (2003) because of the obtuse anterior corners of the clypeus laterally, the mesoscutum with median mesonotal sulcus being well developed and the PPP long and with a sharp apex in dorsal view, whereas the shape of the hypopygium, which is the main character for the generic identification of the males, was not examined. After re-examination of all the specimens mentioned above, we transferred them to the species Zimankos cambodianus under the genus Zimankos because all of them have the hypopygium with the posterior branches filamentary (lobose in Sulcomesitius) (Fig. 17 I), the dorsal harpe of the genitalia is filamentary and distinctly narrower than the ventral harpe (S-shaped or 8 - shaped, at least as wide as the ventral harpe in Sulcomesitius) and the dorsal pronotal area has the anterolateral corner projected (weakly projected in Sulcomesitius).	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE641D46FDBC85C46ACEA786.taxon	description	urn: lsid: zoobank. org: act: 92 E 51160 - 3 B 20 - 4 E 76 - 9 E 16 - 47014 EB 1 FD 77 Fig. 18	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE641D46FDBC85C46ACEA786.taxon	diagnosis	Diagnosis This new species is similar to Z. cambodianus in having the PPP with the apical margin rounded in lateral view. However, it can be distinguished from the latter by having the hind wing with five distal hamuli (four in Z. cambodianus).	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE641D46FDBC85C46ACEA786.taxon	etymology	Etymology The specific epithet is derived from the hamuli (‘ hamulus ’ is Latin for ‘ small hook’), because members of this species have five distal hamuli on the hind wing.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE641D46FDBC85C46ACEA786.taxon	materials_examined	Material examined Holotype CHINA – Zhejiang • ♂; Longquan, Fengyang Mountain; 4 Aug. 2008; Jingxian Liu leg.; ZJUH 200801198.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE641D46FDBC85C46ACEA786.taxon	description	Description Male MEASUREMENTS. Body length 5.48 mm. Length of forewing 3.26 mm. COLOURATION. Body black. Mandible yellowish brown, teeth brown. Antenna brown, darker apically. Fore wing hyaline, brown; veins and pterostigma dark brown. Legs dark brown, trochanters, femora apically and tibiae brown, tarsi yellowish brown. Lateral and posterior margin of T 2 brown; apical half of T 3 and T 4 – 5 brown. PUBESCENCE. Body with short setae. Maxillary and labial palpi with dense, short setae. Antenna with short setae, length of setae 0.19 × width of flagellomere II (Fig. 18 C). Eye with sparse, short setae, nearly 5.0 × diameter of an ommatidium. Wings with short, dense brown setae. Setae of metasoma longer than setae of head and mesosoma, T 1 nearly glabrous, T 2 with basal triangular area glabrous. HEAD. Shorter than wide, LH 0.95 × WH. Clypeus trilobite, medio-clypeal lobe with apical margin acute medially; medio-clypeal carina not extending into frons, arched in lateral view. Antenna slender. Frons coriaceous and densely foveolate, median frontal line weak (Fig. 18 B); WF 1.54 × LE. Contour of eye protruding in antero-dorsal view (Fig. 18 B); LE 0.41 × LH; LE 1.41 × DEV. Anterior ocellus partly anterior to imaginary top line of eyes; POL 1.57 × AOL, OOL 0.85 × WOT, DPV 1.56 × DAO. Vertex coriaceous and densely foveolate, sides of head behind eyes nearly parallel, vertex crest protruding. Gena coriaceous, densely foveolate, with shallow foveolae ventrally. Occipital carina complete. Medioccipito-genal suture present. MESOSOMA. Dorsal pronotal area coriaceous and densely foveolate, median length 0.46 × width along posterior pronotal margin, anterolateral corner angled; median pronotal sulcus nearly complete (Fig. 18 D); lateral pronotal area obliquely striate; cervical pronotal area coarse. Mesoscutum elevated, coriaceous with dense punctures; median mesonotal sulcus present on apical one-third and foveolate (Fig. 18 E); notauli complete and converging posteriorly, parapsidal signum absent near anterior margin. Mesoscutellum elevated, coriaceous and foveolate; mesoscutum-mesoscutellar sulcus arched, laterally slightly dilated (Fig. 18 E). Median length of dorsal surface of MPC 0.95 × half-width; metapostnotum strongly areolate, metapostnotal median carina complete, metapostnotal-propodeal carina complete and converging posteriorly (Fig. 18 F); dorsal propodeal area transversely striate; paraspiracular and lateral marginal carinae of MPC complete; length of PPP 0.32 × median length of dorsal surface of MPC (Fig. 18 F); transverse posterior carina of MPC complete; propodeal declivity areolate, median and lateral carinae complete; anterior metapleural area shiny, weakly striate; lateral surface of MPC obliquely striate, submarginal and posterior carinae present. Mesopleuron coriaceous and densely foveolate; subalar impression weak; posterior oblique sulcus of mesopleuron present and foveolate; postepicnemial sulcus present; ventral surface of mesopleuron coriaceous and shallowly foveolate; anterior mesofurcal pit present and deep. WINGS. Fore wing with 2 r-rs & Rs 2 v slightly curved apically; posterior margin of pterostigma straight, prestigma absent (Fig. 18 H); Cu 2 v present as light yellowish-brown trace. Hind wing with five hamuli. METASOMA. Metasomal terga shiny, with punctures; T 1 with shiny, sparse punctures, medio-longitudinal sulcus present at basal one-third; T 2 with dense punctures. Metasomal sterna shiny, with variably sized punctures (Fig. 18 G); S 1 irregularly striate, metapostnotal median carina weak; S 2 with dense, variably sized punctures. Hypopygium longer than wide, length 2.1 × its width; lateral and anterior margin of basal part of hypopygium distinctly incurved (Fig. 18 I); posterior branches 0.48 × length of hypopygium, distinctly narrower than median notch; apical margin of posterior branches and bottom of posterior median notch with long setae. MALE GENITALIA. Harpe bilobate, dorsal harpe filamentary (Fig. 18 J), ventral harpe finger-like, with long setae (Fig. 18 K). Cuspis with apical margin incurved; digitus bent and papillate apically, with setae basally; aedeagus bottle-shaped (Fig. 18 J). Female Unknown.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE641D46FDBC85C46ACEA786.taxon	distribution	Distribution China (Zhejiang) (Fig. 20).	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE671D48FDE680CA6FACA04B.taxon	description	Fig. 19	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE671D48FDE680CA6FACA04B.taxon	materials_examined	Material examined (27 ♂♂) CHINA – Fujian • 1 ♂; Chongan, Qilichang; 25 Jul. 1985; Minghui Liu leg.; ZJUH 202300003 • 1 ♂; Minqing, Xiongjiang; 13 – 17 Jul. 2005; Zaifu Xu leg.; ZJUH 200609183. – Hainan • 3 ♂♂; Bawangling; 7 – 11 Jul. 2006; Jingxian Liu leg.; ZJUH 200700006, 200700086, 200700129 • 4 ♂♂; Jianfengling; 12 – 15 Jul. 2006; Liqiong Weng leg.; ZJUH 200700404, 200700405, 200700471, 200700503 • 5 ♂♂; Jianfengling; 12 – 15 Jul. 2006; Wenyong Zhang leg.; ZJUH 200700651, 200700653, 200700657, 200700660, 200700673 • 1 ♂; Jianfengling; 12 – 15 Jul. 2006; Feitian Chen leg.; ZJUH 200700712 • 1 ♂; Diaoluoshan; 16 – 17 Jul. 2006; Wenyong Zhang leg.; ZJUH 200701506 • 1 ♂; Diaoluoshan; 16 – 17 Jul. 2006; Tianfei Chen leg.; ZJUH 200701562 • 1 ♂; Wuzhishan, Suiman; 16 – 18 May 2007; Jiezeng leg.; ZJUH 201800023 • 4 ♂♂; Yinggeling; 28 May – 3 Jun. 2007; Liqiong Weng leg.; ZJUH 200706916, 200706929, 200706936, 200706938 • 2 ♂♂; Baisha, Nankai County; 20 Jun. 2008; Jingxian Liu leg.; ZJUH 200800021, 200800025. – Guangxi • 1 ♂; Guilin, Maoershan; 2 – 10 Aug. 2005; Binxiao leg.; ZJUH 200609382. – Yunnan • 1 ♂; Yingjiang, Tongbiguan; 23 Jun. 2005; Jingxian Liu leg.; ZJUH 200609151 • 1 ♂; Gaoligong Mountain; 20 – 21 Jul. 2006; Jie Zeng leg.; ZJUH 200701075. Re-description	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE671D48FDE680CA6FACA04B.taxon	description	Male MEASUREMENTS (n = 27). LH 0.95 – 0.99 × WH; WF 1.16 – 1.45 × LE; LE 0.43 – 0.49 × LH; LE 1.46 – 2.12 × DEV; POL 1.49 – 1.87 × AOL; OOL 0.69 – 0.91 × WOT; DPV 0.78 – 1.57 × DAO; median length of dorsal pronotal area 0.43 – 0.51 × width along posterior pronotal margin; median length of dorsal surface of MPC 1.07 – 1.21 × its half-width; length of PPP 0.21 – 0.31 × median length of dorsal surface of MPC; length of hypopygium 1.90 – 2.42 × its width; length of posterior branches 0.39 – 0.52 × length of hypopygium. COLOURATION. Body black. Mandible brown; maxillary and labial palpi brown. Antenna brown, darker apically. Fore wing hyaline, yellowish brown; veins yellowish brown to brown, pterostigma yellowish brown. Legs brown, trochanters, tibiae and tarsi yellowish brown. T 1 dark brown, T 2 black with posterior and lateral margin brown; T 3 – 4 brown. PUBESCENCE. Body with short setae. Maxillary and labial palpi with dense, short setae. Antenna with short, suberect setae, 0.33 × width of flagellomere I (Fig. 19 C). Eye with sparse, short setae, nearly 5.0 × diameter of an ommatidium. Wings with short, dense brown setae. Setae of metasoma longer than setae of head and mesosoma, T 1 nearly glabrous, T 2 with basal triangular area glabrous. HEAD. Slightly shorter than wide, LH 0.97 × WH. Clypeus trilobite, medio-clypeal lobe with anterior margin acute medially; medio-clypeal carina not extending into frons, arched in lateral view. Antenna slender. Frons coriaceous and densely foveolate, median frontal line absent (Fig. 19 B); WF 1.16 × LE. Contour of eye protruding in antero-dorsal view; LE 0.49 × LH, LE 2.12 × DEV. Anterior ocellus partly anterior to imaginary top line of eyes in antero-dorsal view; POL 1.52 × AOL, OOL 0.83 × WOT, DPV 0.93 × DAO. Vertex coriaceous and densely foveolate, sides of head behind eyes rounded, vertex crest protruding. Gena coriaceous, densely foveolate, with shallow foveolae ventrally. Occipital carina complete. Medioccipito-genal suture present. MESOSOMA. Dorsal pronotal area coriaceous, with shallower and bigger foveae than on head, median length of dorsal pronotal area 0.44 × width along posterior pronotal margin, anterolateral corner projected; median pronotal sulcus present and weakly foveolate (Fig. 19 D); lateral pronotal area obliquely striate; cervical pronotal area coarse. Mesoscutum elevated, coriaceous, with punctures; median mesonotal sulcus present as a big fovea near posterior margin (Fig. 19 D); notauli complete and converging posteriorly, parapsidal signum shallow and absent near anterior margin. Mesoscutellum coriaceous and foveolate; mesoscutum-mesoscutellar sulcus present, laterally dilated (Fig. 19 E). Median length of dorsal surface of MPC 1.15 × half-width; metapostnotum strongly areolate, metapostnotal median carina complete, metapostnotal-propodeal carina complete and converging posteriorly; dorsal propodeal area transversely striate; paraspiracular and lateral marginal carinae of MPC complete; length of PPP 0.26 × as long as median length of dorsal surface of MPC, transverse posterior carina of MPC complete (Fig. 19 E); propodeal declivity weakly areolate, median and lateral carinae complete; anterior metapleural area shiny, weakly striate; lateral surface of MPC areolate, submarginal and posterior carinae present. Propleuron coriaceous, foveolate near procoxa. Mesopleuron coriaceous and densely foveolate; subalar impression weak; posterior oblique sulcus of mesopleuron absent; postepicnemial sulcus present; ventral surface of mesopleuron coriaceous and shallowly foveolate; anterior mesofurcal pit present and deep. WINGS. Fore wing with 2 r-rs & Rs 2 v slightly curved apically; posterior margin of pterostigma protruding, prestigma absent (Fig. 19 G); Cu 2 v present as light yellowish-brown trace. Hind wing with four distal hamuli. METASOMA. Metasomal terga shiny, with punctures; T 1 with shiny, sparse punctures, medio-longitudinal sulcus present at basal one-third; T 2 with punctures (Fig. 19 G). Metasomal sterna shiny, with variably sized punctures; S 1 irregularly striate, metapostnotal median carina weak; S 2 with dense, variably sized punctures. Hypopygium longer than wide; lateral and anterior margin of basal part of hypopygium distinctly incurved (Fig. 19 H); posterior branches 0.39 × as long as length of hypopygium, distinctly narrower than median notch; apical margin of posterior branches and bottom of posterior median notch with long setae. MALE GENITALIA. Harpe bilobate, dorsal harpe filamentary (Fig. 19 I), ventral harpe finger-like, with long setae (Fig. 19 J). Cuspis with apical margin incurved; digitus bent and papillate apically, with setae basally; aedeagus bottle-shaped (Fig. 19 I). Female Unknown.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
C604E518AE671D48FDE680CA6FACA04B.taxon	distribution	Distribution China (Fujian, Hainan, Guangxi, Yunnan) (Fig. 20), Vietnam. New record for China.	en	Wang, Chunhong, He, Junhua, Chen, Xuexin (2024): Revision of subfamily Mesitiinae (Hymenoptera, Bethylidae) from China. European Journal of Taxonomy 960 (1): 1-61, DOI: 10.5852/ejt.2024.960.2679, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2679/12407
