taxonID	type	description	language	source
C60DAE358637D832D8B7FB26BA9CFE07.taxon	description	(Figs 4 – 15)	en	Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins, Solodovnikov, Alexey (2024): Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae). Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1): 1-42, DOI: 10.1093/zoolinnean/zlad143, URL: https://doi.org/10.1093/zoolinnean/zlad143
C60DAE358637D832D8B7FB26BA9CFE07.taxon	type_taxon	Type species: Loncovilius semiflavus (Fairmaire and Germain 1862).	en	Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins, Solodovnikov, Alexey (2024): Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae). Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1): 1-42, DOI: 10.1093/zoolinnean/zlad143, URL: https://doi.org/10.1093/zoolinnean/zlad143
C60DAE358637D832D8B7FB26BA9CFE07.taxon	diagnosis	Diagnosis: Loncovilius can be distinguished from all other genera in Amblyopinini based on the combination of the following characters: frontoclypeal (epistomal) suture complete (Fig. 7 C – E); basisternum flat, with few punctures, without pair of macrosetae; with rounded flexible postcoxal hypomeral extension or process; two short empodial setae on each tarsomere 5; ventral side of meso- and metatarsi in males and females with pale adhesive setae (Fig. 7 F); abdomen without protergal glands; all tergites without posterior transverse basal carina (PTBC); lateral tergal sclerites IX at least slightly longer than tergite VIII. Free-living; females distinctly larger than males.	en	Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins, Solodovnikov, Alexey (2024): Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae). Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1): 1-42, DOI: 10.1093/zoolinnean/zlad143, URL: https://doi.org/10.1093/zoolinnean/zlad143
C60DAE358637D832D8B7FB26BA9CFE07.taxon	description	Description: Frontoclypeal (epistomal) suture complete; neck constriction fully developed; neck moderately wide; nuchal ridge latero-ventrally extended anteriad beyond level of postgenal ridge, merged with, or coming very close to, infraorbital ridge; dorsal basal ridge absent; ventral basal ridge extending more or less parallel to ventral portion of postoccipital suture; postgenal ridge partially or completely missing; no extra ridge anterior to postgenal ridge; postmandibular ridge short; infraorbital ridge incomplete; frontoclypeal, anterior frontal, parocular, posterior frontal, and basal punctures present; supra-antennal and interocular punctures absent. To avoid future confusion with the terminology of the setiferous punctures of the head in ventral view we decided to follow that of figure 1 of Smetana and Davies (2000) except for its ' genal seta' which we rename here as postmandibular setiferous puncture (PMP) in order to not confuse it with the ' genal puncture' of Brunke et al. (2019) which is located in the anterodorsal genal region. PMP is found in the anteroventral genal region, in the area delimited by the margin of the eye, the base of the mandible, and the maxilla, generally associated with the anterior part of the postmandibular ridge when present. In Loncovilius, the PMP is present, in the germaini group it is well developed (Fig. 8 E), and in the rest of the groups, it is reduced (Fig. 8 F). Gula with distinct transverse basal impression; gular sutures (gs) separated from each other. Antennal insertions situated at equal distance to frontoclypeus and to eye (except L. germaini with antennal insertions situated closer to frontoclypeus than to eye); antennae relatively long, all antennomeres elongate, antennomere 1 shorter than or subequal to antennomeres 2 and 3 combined, antennomeres 4 to 11 with tomentose pubescence, males with antennomere 11 (a 11) distinctly longer than antennomere (a 10) (a 11: a 10 ratio> 1.5), this character varies in females. Mandibles in dorsal view slightly straight with curved base and apex; right mandible with proximal tooth and distal tooth with varying shape; teeth of left mandible vary. In germaini group, on right mandible, distal tooth bifid, space between proximal and distal teeth narrow and smooth. In edwardsianus, lividipennis, semiflavus, and variabilis groups, on right mandible, distal tooth not bifid, space between proximal and distal teeth wide and rough; in germaini group left mandible only with proximal, not bifid tooth; in edwardsianus, lividipennis, semiflavus, and variabilis groups left mandible with proximal and distal teeth not bifid, the latter short and truncated, space between teeth rough, in lateral view right and left mandibles deflexed ventrally at an angle of 25 – 30 ° c. one-fourth to one-third distance from base to apex; ligula small and entire; labrum with transparent apical membrane; maxillary palpomere 4 (mp 4, apical) more or less subconical, with evenly narrowed apex, as long as or longer than palpomere 2; preapical maxillary palpomere 3 only weakly wider than apical palpomere, about as long as mp 4; labial palpomere 2 with sparse brushes of setae. Ŋorao: Chaetotaxy of pronotum varies between species groups. Paired punctures in dorsal series (PPDS) may be present in different combinations; sublateral setiferous punctures (SLSP) defined here as punctures found in the area between PPDS, anterior marginal row of setiferous punctures (AMRSP), large lateral setiferous puncture (LLSP) and large posterior setiferous puncture (LPSP). Loncovilius carlsbergi sp. nov. has the most complete chaetotaxy set where all types of setiferous punctures occur (Fig. 9 A). Pronotum and prosternum not fused, pronotosternal suture complete even in procoxal cavity; postcoxal hypomeral translucent process present, rounded or triangular; sternacostal ridge not protruding, not joining with superior line of hypomeron; basisternum (bs) flat with few punctures, triangular, its lateral arms narrowed subapically, without pair of macrosetae, bs distinctly longer than furcasternum (fs) (bs / fs ratio> 1.5). Elytra without humeral spines or spine-like setae; mesoscutellum setose without posterior scutellar ridge or subbasal ridge; hind wings with veins CuA and MP 4 fused in one vein, vein MP 3 absent; metascutellum with mid-longitudinal suture; apex of intercoxal process in mesothorax rounded or broadly pointed, forming obtuse angle; anterior margin of mesoventrite confluent with distinctly oblique sternopleural suture; posterior border of mesocoxal cavities complete; mesometasternal suture present. Tarsal formula 5 - 5 - 5; protibiae cylindrical to slightly broadened apically, ventrally with a middle long spine directed anteriad; protarsomeres 1 – 3 distinctly wider than long, bilobed, and ventrally with pale adhesive setae; ventral side of meso- and metatarsomeres 1 – 4 in males and females with pale adhesive setae; pale adhesive setae on meso- and metatarsi with terminal plate (Fig. 8 B) or without (Fig. 8 C); all legs with a pair of empodial setae distinctly shorter than claws; procoxae with internal and external ridges present, internal not running along external ridge, ending distinctly before or nearly joining external ridge; metacoxae with no more than four spines on posterior surface; mesotarsal segments with pale adhesive setae. Abdomen: Protergal glands absent; tergites III – VI with anterior transverse basal carina (ATBC) laterally straight continuing to paratergites, tergite VII with ATBC not continuing to paratergite, tergites VII and VIII with ATBC descending laterally without reaching tergite margin (Fig. 9 D); all tergites without PTBC; anterior margin of tergites and sternites III – VI entire; tergite VII with white fringe along posterior edge; tergite IX in males at least slightly longer than tergite VIII; apical margin of male sternite VIII with medial emargination; basal portion of male sternite IX asymmetrical; paramere fused to median lobe only at base and very closely appressed to median lobe along entire length, often of complex and diverse shape, with minute pale sensory peg setae or usual peg setae; internal sac often with visible sclerites; ovipositor with two paired gonocoxites and minute styli.	en	Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins, Solodovnikov, Alexey (2024): Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae). Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1): 1-42, DOI: 10.1093/zoolinnean/zlad143, URL: https://doi.org/10.1093/zoolinnean/zlad143
C60DAE358637D832D8B7FB26BA9CFE07.taxon	distribution	Distribution and habitat: The genus is only known from Chile and Argentina; in Chile from the regions of Araucanía, Aysén, Los Lagos, Los Ríos, Magallanes and Chilean Antarctica, Maule, Ñuble, O’Higgins, and from Santiago Metropolitan; in Argentina from the provinces of Chubut, Neuquén and Río Negro (Fig. 4). The genus occurs in the area from lowlands to 2600 m elevation in the mountains, with an average annual rainfall of between 500 and 3000 mm and an average annual temperature of between 1 ° C and 12 ° C. It is confined to various types of forests in the Chilean Matorral, Magellanic subpolar, and Valdivian temperate forest ecoregions. It has been collected using Malaise and flight intercept traps, sifting moss, dead wood, and leaf litter, beating and directly from flowers of the families Cunoniaceae, Ericaceae, Loranthaceae, Myrtaceae, and Winteraceae.	en	Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins, Solodovnikov, Alexey (2024): Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae). Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1): 1-42, DOI: 10.1093/zoolinnean/zlad143, URL: https://doi.org/10.1093/zoolinnean/zlad143
C60DAE358637D832D8B7FB26BA9CFE07.taxon	discussion	Remarks: The monophyly of Loncovilius is supported by two unique synapomorphies: females with pale adhesive setae present at mesotarsal segments 2 – 4 [(character 69, state 1; reversed condition in variabilis and edwardsianus groups (character 69, state 0)]; tergite IX at least slightly longer than tergite VIII (ratio up to 1.1, character 92, state 1); and by five homoplastic synapomorphies: eyes small [(eye / head length ratio more than three-tenths but less than half, character 37, state 1; in semiflavus group and L. impunctus eyes of medium size (eyes / head length ratio more than half but less than three-quarters, character 37, state 2)]; metascutellar mid-longitudinal suture well developed (character 82, state 0); protergal glands absent (character 84, state 0); female sternite VIII with medial emargination at apical margin [(character 99, state 1; reversed condition in variabilis and edwardsianus groups (character 100, state 0)]; paramere (s) with black sensory peg setae [(character 105, state 1; reversed condition in edwardsianus group (character 105, state 0)].	en	Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins, Solodovnikov, Alexey (2024): Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae). Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1): 1-42, DOI: 10.1093/zoolinnean/zlad143, URL: https://doi.org/10.1093/zoolinnean/zlad143
C60DAE358625D832D915FD87BDEFFC26.taxon	diagnosis	Diagnosis: Head transverse, with coarse non-setiferous punctures; nuchal ridge complete dorsally, nuchal and infraorbital ridges joined; postmandibular ridge distinctly separated from eye margin and well-developed PMP (Fig. 8 E; Brunke et al. 2019: fig. 1); right mandible with distal tooth bifid; left mandible only with proximal, non-bifid tooth. Pronotum transverse, outline evenly curved. Tergites III and IV with two patches of punctures of moderate size and density, with a middle impunctate line (Fig. 9 A); ovipositor, its second gonocoxite without macroseta medially (Fig. 9 F).	en	Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins, Solodovnikov, Alexey (2024): Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae). Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1): 1-42, DOI: 10.1093/zoolinnean/zlad143, URL: https://doi.org/10.1093/zoolinnean/zlad143
C60DAE358625D833D893FC49BFEDF9A4.taxon	description	(Figs 5 A, 9 A, 10 A – C, 12 A, 13 A, 14 A) Germain 1903: 412 (as Philonthus cribripennis, original description); Bernhauer and Schubert 1914: 332 (as Philonthus chilensis, replacement name); Scheerpeltz 1933: 1344 (as Philonthus germaini, replacement name); Coiffait and Sáiz 1966: 406 (as Loncovilius (Lienturius) germaini, characters); Coiffait and Sáiz 1968: 365 [as Loncovilius (Lienturius) germaini; checklist]; Sáiz 1971: 385 [as Loncovilius (Lienturius) germaini; listed without new data]; Herman 2001 a: 26 (as Loncovilius chilensis, nomenclature); Herman 2001 b: 3083 (as Loncovilius chilensis, catalogue); Newton 2017: 22 (as Loncovilius germaini, nomenclature). Material eoamined: Supporting Information, File S 7. Diagnosis: Head dorsally without microsculpture; gula without isodiametric microsculpture in the middle; females with a 11 distinctly longer than a 10 (a 11: a 10 ratio <1.5). Elytra, including epipleura, evenly setose and with several coarse non-setiferous punctures. Protibiae sexually dimorphic, with laterodorsal row of thick spines only present in females. Tergite VIII with medial apical emargination only in females. Aedeagus as in Figure 10 A – C. Description: Measurements ♂ [min – max (average); N = 6]: FBL = 2.9 – 3.44 (3.19); TL = 5.6 – 6.25 (6.01); HW = 0.87 – 1.01 (0.94); HL = 0.76 – 0.86 (0.82); HW / HL = 1.12 – 1.2 (1.14); PW = 1.11 – 1.29 (1.21); PL = 0.97 – 1.18 (1.08); PW / PL = 1.1 – 1.14 (1.12); EW = 1.4 – 1.64 (1.52); EL = 1.16 – 1.4 (1.3); PW / HW = 1.26 – 1.33 (1.29). Measurements ♀ [min – max (average); N = 6]: FBL = 3.36 – 3.87 (3.6); TL = 7.31 – 7.77 (7.54); HW = 0.96 – 1.12 (1.03); HL = 0.85 – 0.95 (0.91); HW / HL = 1.11 – 1.18 (1.13); PW = 1.25 – 1.48 (1.34); PL = 1.13 – 1.28 (1.2); PW / PL = 1.1 – 1.16 (1.12); EW = 1.5 – 1.88 (1.68); EL = 1.38 – 1.64 (1.49); PW / HW = 1.26 – 1.33 (1.3). Head black to brown-black; mouthparts, antennae, pronotum, legs, disc of elytra, and abdominal segments dark reddish-brown; epipleura, abdominal tergites, and sternites with posterior margin light reddish-brown. Head transverse; dorsally and ventrally glossy with few micropunctures, dorsally without microsculpture, with coarse non-setiferous punctures; posterior angles indistinct with few setiferous punctures of medium and small size. Eyes small (EYL / HL = x = 0.46), from 1.4 to 1.86 times longer than the length of temples (in lateral view); distance between eyes about 1.68 times as long as length of the eye in males and 1.8 times in females. Antennal insertions situated closer to frontoclypeus than to eye; antennomeres 2 and 3 subequal in length; antennomeres 5 to 10 subequal in length; antennomere 11 from 1.53 to 1.69 times longer than antennomere 10 in males and from 1.5 to 1.6 in females. Basal and parocular punctures usually single; posterior frontal puncture located anterior to temporal puncture; no small setose punctures between frontoclypeal punctures and anterior frontal punctures; ventral basal ridge rather straight in L-shape, almost united with gular sutures; postgenal ridge absent; postmandibular ridge distinctly separated from eye margin; well-developed PMP; nuchal ridge present laterally and dorsally, fused with infraorbital ridge; gula without microsculpture medially, gular sutures moderately separated. Mandibles with dorsolateral groove; labrum with subrectangular sclerotized region and entire apical margin; mentum with seta beta only; penultimate labial palpomere markedly dilated apicad, apical palpomere distinctly narrower than penultimate and more or less subconical. Pronotum transverse, convex, evenly curved, usually with one PPDS with additional paired punctures adjacent to AMRSP (APP), without SLSP, with few coarse non-setiferous punctures; flexible postcoxal hypomeral extension (process) interrupted by inferior line. Elytra evenly setose and with several coarse non-setiferous punctures. Mesosternumwithfourmacrosetaearrangedinarowmedially. In both sexes, metatarsomeres 2 – 4 with pale adhesive setae, without terminal plate; metatarsomere 1 shorter than 5, metatarsomere 4 ventrally with apical margin slightly sinuate. Protibiae sexually dimorphic, with laterodorsal row of thick spines only in females. In males, mesotarsomeres 1 – 4 with pale adhesive setae, mesotarsomeres 1 – 3 with terminal plate. In females, mesotarsomeres 2 – 4 with pale adhesive setae, without terminal plate. Abdominal tergites III and IV with two patches of punctures of moderate size and density, with middle impunctate line (Fig. 9 A); in females tergite VIII emarginate medio-apically. Sternite III with basal transverse carina converging evenly at an obtuse angle; lateral tergal sclerites IX not dorsoventrally or laterally flattened. Male sternite VIII with deep V-shaped emargination medially (Fig. 14 A); female sternite VIII with U-shaped emargination medially; male sternite IX with obtuse emargination medially, its basal portion shorter than distal portion; male tergite X subtruncate medially (Fig. 12 A); female tergite X with U-shaped emargination medially (Fig. 13 A); ovipositor, its second gonocoxite without macrosetae medially (Fig. 9 F). Aedeagus. As in Figure 10 A – C; its total length ~ 1.21. Distribution and habitat: Loncovilius germaini is known only from Chile, from the Araucanía, Aysén, and Los Lagos regions where it occurs in the eastern portion of the Valdivian temperate forest ecoregion, from 750 to 1450 m of elevation. It has been collected by sifting leaf litter and moss, also by using Malaise and window traps, as well as carrion and dung baited pitfall traps. Remarks: Loncovilius germaini was described as Philonthus cribripennis based on an unspecified number of syntypes from an uncertain type locality in Chile. This name was found to be preoccupied and replaced by another preoccupied name P. chilensis Bernhauer and Schubert 1914; the latter was replaced again by P. germaini Scheerpeltz, 1933. Coiffiat and Sáiz (1966, 1968) and Sáiz (1971) treated L. germaini as a member of the subgenus Lienturius of the genus Loncovilius without adding much data about morphology or distribution of this species, for which even the aedeagus hitherto remained not illustrated. Coiffait and Sáiz (1966) mentioned that the species was known from a female holotype kept in the collection of the Natural History Museum of Chile. At the same time, Camousseight (1980) did not list this species among the type material in Germain’s collection in this museum. We did not find any material that could be interpreted as types of L. germaini. Given the uncertainty about type material in the original description and the catalogue by Camousseight (1980), the type status of the female listed by Coiffait and Sáiz (1966) as ‘ holotype’ must be verified in the future. For now, we have matched the identity of this species based on the description by Coiffait and Sáiz (1966). As explained by Newton (2017), Herman’s (2001 a) interpretation of both names as replacements for only P. cribripennis and his resurrection of P. chilensis as the valid name for this species was incorrect.	en	Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins, Solodovnikov, Alexey (2024): Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae). Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1): 1-42, DOI: 10.1093/zoolinnean/zlad143, URL: https://doi.org/10.1093/zoolinnean/zlad143
C60DAE358624D830D979F9C9BE3BF9C5.taxon	description	(Figs 5 B, 7 C, 10 D – F, 12 B, 13 B, 14 B) Zoobank registration: urn: lsid: zoobank. org: act: 4 DCCE 49 B- 5875 - 4 A 73 - 9 DA 6 - 203 BC 4455 E 20. Type material. Holotype: male, mounted, with genitalia in a separate microvial, with labels as follows: Wellington Is. Pto. Eden [- 49.11, - 74.40] 4. XII 1958 / CHILE Fr Kuschel / HOLOTYPE Loncovilius hammondi des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Deposited at NHM. Paratypes: Wellington Is. Pto. Eden [- 49.11, - 74.40] 4. XII. 1958 / CHILE Fr Kuschel [4 ♂ 8 ♀ NHM; 1 ♂ NHMD ex. NHM; 1 ♂ CZUG ex. NHM]; Nothofagus forest / CHILE HE 6. Puerto Eden Isla Wellington [- 49.11, - 74.40] 49 ° S I 200 ft. 2. xii. 1958 [1 ♀ MNNC ex. NHM]; Chiloé S. Pedro [- 43.32, - 73.72] 10. XI. 1958 / CHILE Fr Kuschel / Flowers of Pernettya mucronata / Qued. sp. 8 / ‘ Loncovilius’ cf germaini Scheerp. P. M. Hammond det. 1980 [1 ♂ NHM]; CHILE: Cautín Pr. Volcán Villarrica [- 39.36, - 71.94], 1120 m, site 654 15 - 29. xii. 1982 Noth. Dombeyi-Saxe-gothea with Drimys A. Newton and M. Thayer / window trap [1 ♂ 1 ♀ FMNH]. All paratypes are supplied with the label: PARATYPE Loncovilius hammondi des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Diagnosis: Head dorsally with microsculpture of transverse waves; gula with isodiametric microsculpture in the middle; females with a 11 distinctly longer than a 10 (a 11: a 10 ratio> 1.5). Elytra reddish-brown, disc with coarse non-setiferous punctures and only a few rows of setiferous punctures, epipleura evenly setose. Protibiae not sexually dimorphic, in both sexes with laterodorsal row of thick spines. Tergite VIII with medial apical emargination in both sexes. Aedeagus as in Figure 10 D – F. Description: Measurements ♂ [min – max (average); N = 6]: FBL = 3.5 – 3.91 (3.67); TL = 6.7 – 7.31 (7.01); HW = 1.06 – 1.16 (1.11); HL = 0.94 – 1.02 (0.97); HW / HL = 1.12 – 1.16 (1.14); PW = 1.36 – 1.51 (1.43); PL = 1.16 – 1.26 (1.21); PW / PL = 1.17 – 1.2 (1.19); EW = 1.65 – 1.82 (1.76); EL = 1.38 – 1.63 (1.49); PW / HW = 1.29 – 1.3 (1.29). Measurements ♀ [min – max (average); N = 6]: FBL = 3.88 – 4.22 (4.04); TL = 7.06 – 8.88 (7.81); HW = 1.13 – 1.21 (1.18); HL = 0.98 – 1.07 (1.03); HW / HL = 1.13 – 1.19 (1.15); PW = 1.5 – 1.67 (1.58); PL = 1.23 – 1.37 (1.31); PW / PL = 1.18 – 1.22 (1.20); EW = 1.9 – 2.3 (2.05); EL = 1.67 – 1.78 (1.7); PW / HW = 1.31 – 1.38 (1.33). Head black to brown-black; most of antennae, pronotum, mouthparts, elytra, legs, and abdomen reddish-brown; pronotum, apical antennomeres, tibiae, and disc of abdominal segments darker. Head transverse; dorsally and ventrally glossy with few micropunctures and microsculpture of transverse waves, with coarse non-setiferous punctures; posterior angles indistinct with few setiferous punctures of medium and small size. Eyes small (EYL / HL = x = 0.47), from 1.3 to 1.8 times longer than temples (in lateral view); distance between eyes about 1.7 times as long as length of eye in males and 1.68 times in females. Antennal insertions situated at equal distance from frontoclypeus and from eye; antennomeres 2 and 3 usually subequal in length; antennomeres 4 to 6 gradually reduced in length; antennomeres 7 to 10 subequal in length; antennomere 11 is from 1.53 to 1.69 times as long as antennomere 10 in males and from 1.63 to 1.71 times in females. Basal and parocular punctures usually single; posterior frontal puncture located anterior to temporal puncture; without small setiferous punctures between frontoclypeal punctures and anterior frontal punctures; ventral basal ridge rather straight in L-shape, almost united with gular sutures; postgenal ridge absent; postmandibular ridge distinctly separated from eye margin; well-developed PMP; nuchal ridge present laterally and dorsally, fused with infraorbital ridge; gula with microsculpture medially, gular sutures moderately separated. Mandibles with dorsolateral groove; labrum with subrectangular sclerotized region and emarginate apical margin; mentum with seta beta only; penultimate labial palpomere markedly dilated apicad, apical palpomere distinctly narrower than previous and more or less subconical. Pronotum transverse, convex, evenly curved, with one PPDS and with an APP, without SLSP, with several coarse non-setiferous punctures; flexible postcoxal hypomeral extension (process) interrupted by inferior line. Elytra with epipleura evenly setiferous punctate; disc with only a few rows of setiferous punctures and with several coarse non-setiferous punctures. Mesosternum with four macrosetae arranged in a row medially. Protibiae not sexually dimorphic, with laterodorsal row of thick spines in both sexes. In males, mesotarsomeres 1 – 4 with pale adhesive setae, mesotarsomeres 1 – 3 with terminal plate. In females, mesotarsomeres 2 – 4 with pale adhesive setae, without terminal plate. In both sexes, metatarsomeres 2 – 4 with pale adhesive setae, without terminal plate; metatarsomere 1 shorter than 5, metatarsomere 4 ventrally with apical margin slightly sinuate. Abdominal tergites III and IV with two patches of punctures of moderate size and density, with middle impunctate line; tergite VIII emarginate medio-apically. Sternum III with basal transverse carina strongly arcuate; lateral tergal sclerites IX laterally flattened only in males. Male sternite VIII with a broad V-shaped emargination medially (Fig. 14 B); female sternite VIII with a U-shaped emargination medially; male sternite IX with an obtuse emargination medially, its basal portion shorter than distal portion; male tergite X truncate medially (Fig. 12 B); female tergite X truncate to slightly arcuate medially (Fig. 13 B); ovipositor, its second gonocoxite without macrosetae medially. Aedeagus. As in Figure 10 D – F; its total length ~ 1.21. Etymology: The species epithet isa patronym in recognition of the late Peter. M. Hammond, a prominent coleopterist who also conducted some work on this genus, which was unpublished but which we were able to trace through his labels. Distribution and habitat: Loncovilius hammondi is known only from Chile, from the Araucanía, Los Lagos, as well as from Magallanes and Chilean Antarctica regions. Known distribution consists of three widely separated areas, two of them confined to the Valdivian temperate forest and one to the Magellanic subpolar forest. It has been collected using window traps, and directly from flowers of Pernetya mucronata (Ericaceae).	en	Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins, Solodovnikov, Alexey (2024): Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae). Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1): 1-42, DOI: 10.1093/zoolinnean/zlad143, URL: https://doi.org/10.1093/zoolinnean/zlad143
C60DAE358627D831D94FF9ECBDF2FABE.taxon	description	(Figs 5 C, 10 G, H, 12 C, 13 C, 14 C) Zoobank registration: urn: lsid: zoobank. org: act: 45463 DC 0 - 4 ECA- 4 B 24 - 8220 - 24008 B 5 BE 531. Type material. Holotype: male, mounted, with genitalia in a separate microvial, with labels as follows: CHILE: Cherquenco [~ - 38.70, - 72.01] VII- 1954 L. Pena, leg. / ♂ / FMNH-INS 0000 024 367 / HOLOTYPE Loncovilius impunctus des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Deposited at FMNH. Paratypes: CHILE: Cherquenco [~ - 38.70, - 72.01] VII- 1954 L. Pena, leg. / ♀ / FMNH-INS 0000 024 368 - 369 [2 ♀ FMNH]; CHILE: Malleco Prov. 20 km E Manzanar [- 38.44, - 71.51] 1100 m 19 - 21. xii. 1976 H. F. Howden / CNCI L 2004 - 28 / ♀ / FMNH-INS 0000 024 465 [1 ♀ CNC]; Chile: Tolhuaco [~ - 38.23, - 71.71] 11. I. 62 / Coll. L. Peña / ♂ / FMNH-INS 0000 024 364 – 366 [2 ♂ FMNH; 1 ♀ MNNC ex. FMNH]. All paratypes are supplied with the label: PARATYPE Loncovilius impunctus des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022 Diagnosis: Head dorsally with meshed microsculpture; gula with isodiametric microsculpture in the middle; females with antennomere 11 (a 11) distinctly longer than antennomere 10 (a 10; a 11: a 10 ratio> 1.5). Elytra reddish-brown, disc glossy with only a few rows of setiferous punctures, epipleura evenly setose. Protibiae not sexually dimorphic, in both sexes with laterodorsal row of thick spines. Tergite VIII in both sexes with medial apical emargination. Aedeagus as in Figure 10 G, H. Description: Measurements ♂ [min – max (average); N = 3]: FBL = 3.35 – 3.59 (3.48); HW = 1.01 – 1.1 (1.07); HL = 0.85 – 0.93 (0.89); HW / HL = 1.18 – 1.21 (1.19); PW = 1.4 – 1.44 (1.42); PL = 1.2 – 1.25 (1.23); PW / PL = 1.14 – 1.17 (1.15); EW = 1.67 – 1.7 (1.68); EL = 1.3 – 1.44 (1.36); PW / HW = 1.28 – 1.39 (1.33). Measurements ♀ [min – max (average); N = 4]: FBL = 3.71 – 3.93 (3.81); TL = 7.5 – 7.53 (7.52); HW = 1.17 – 1.18 (1.18); HL = 0.93 – 0.97 (0.95); HW / HL = 1.22 – 1.26 (1.24); PW = 1.54 – 1.59 (1.56); PL = 1.29 – 1.37 (1.32); PW / PL = 1.16 – 1.19 (1.18); EW = 1.82 – 1.9 (1.87); EL = 1.46 – 1.59 (1.54); PW / HW = 1.32 – 1.35 (1.33). Head, mouthparts, and legs black to brown-black; pronotum, antennae, elytra, and abdomen dark reddish-brown, elytra lighter. Head transverse; dorsally and ventrally glossy with few micropunctures and microsculpture of transverse meshes, with coarse non-setiferous punctures; posterior angles indistinct with few setiferous punctures of medium and small size. Eyes of medium size (EYL / HL = x = 0.52), from 1.51 to 1.64 times longer than temples (in lateral view); distance between eyes around 1.8 times the length of eye. Antennal insertions situated at equal distance from frontoclypeus and from eye; antennomeres 2 and 3 subequal in length; antennomeres 4 to 10 gradually reduced in length; a 11 from 1.54 to 1.75 times longer than a 10 in males and from 1.54 to 1.68 in females. Basal and parocular punctures usually single; posterior frontal puncture located anterior to temporal puncture; without small setose punctures between frontoclypeal and anterior frontal punctures; ventral basal ridge rather straight in L-shape, almost united with gular sutures; postgenal ridge absent; postmandibular ridge distinctly separated from eye margin; well-developed PMP; nuchal ridge present laterally and dorsally, fused with infraorbital ridge; gula with microsculpture medially, gular sutures moderately separated. Mandibles with dorsolateral groove; labrum with subrectangular sclerotized region and emarginate apical margin; mentum with seta beta only; penultimate labial palpomere markedly dilated apicad, apical palpomere distinctly narrower than penultimate and more or less subconical. Pronotum transverse, convex, evenly curved, with one PPDS and an APP, without SLSP, with few coarse non-setiferous punctures; flexible postcoxal hypomeral extension (process) interrupted by inferior line. Elytra with epipleura evenly setose; disc with only a few rows of setiferous punctures. Mesosternum with four macrosetae arranged in a row medially. Protibiae not sexually dimorphic, with laterodorsal row of thick spines in both sexes. In males, mesotarsomeres 1 – 4 with pale adhesive setae, mesotarsomeres 1 – 3 with terminal plate. In females, mesotarsomeres 2 – 4 with pale adhesive setae, without terminal plate. In both sexes, metatarsomeres 2 – 4 with pale adhesive setae, without terminal plate; metatarsomere 1 shorter than 5, metatarsomere 4 ventrallywith apical margin slightly sinuate. Abdominal tergites III and IV with two patches of punctures of moderate size and density, with middle impunctate line; tergite VIII emarginate medio-apically. Sternite III with basal transverse carina strongly arcuate; lateral tergal sclerites IX laterally flattened only in males. Male sternite VIII with broad V-shaped emargination medially (Fig. 14 C); female sternite VIII with U-shaped emargination medially; male sternite IX with obtuse emargination medially, its basal portion shorter than distal portion; male tergite X truncate medially (Fig. 12 C); female tergite X truncate to slightly arcuate medially (Fig. 13 C); ovipositor, its second gonocoxite without macrosetae medially. Aedeagus as in Figure 10 G, H; its total length 1.2. Median lobe bulky, in parameral view with truncated apex; internal sac with pair of long I-shaped copulatory sclerites. Paramere slightly produced over apex of median lobe, wider than median lobe, its sides converging into M-shape apex, with dark peg setae, in lateral view almost straight and slightly concave apically. Etymology: The species epithet is derived from the Latin prefix ‘ im ’ (not) and adjective ‘ punctate ’ (dotted), referring to the disc of pronotum and elytra mostly impunctate. Distribution and habitat: Loncovilius impunctus is known from the Araucanía Region in Chile where it is confined to Valdivian temperate forest ecoregion. It is unknown how it was collected.	en	Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins, Solodovnikov, Alexey (2024): Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae). Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1): 1-42, DOI: 10.1093/zoolinnean/zlad143, URL: https://doi.org/10.1093/zoolinnean/zlad143
C60DAE358626D831D906FAD1BCD4FF3D.taxon	diagnosis	Diagnosis: Head usually elongate, without coarse non-setiferous punctures; nuchal ridge complete dorsally, joined with infraorbital ridge; gula with isodiametric microsculpture in the middle; postmandibular ridge located close to eye margin (Fig. 8 F); PMP (usually reduced); females with a 11 distinctly longer than a 10 (a 11: a 10 ratio> 1.5). Pronotum slightly wider than long, its outline evenly curved. Elytra with even but sparse setiferous punctation on disc and epipleura. Protibiae not sexually dimorphic, in both sexes without laterodorsal row of thick spines. Mesotarsomeres 1 – 4 not sexually dimorphic, in both sexes with pale adhesive setae and mesotarsomeres 1 – 3 in both sexes with terminal plate. Tergites III and IV with several rows of fine punctures of moderate density, without sizeable impunctate area near to posterior margin (Fig. 9 B); tergite VIII only slightly emarginate medio-apically in females; ovipositor, its second gonocoxite with one macroseta medially. Aedeagus as in Figure 10 I – K.	en	Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins, Solodovnikov, Alexey (2024): Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae). Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1): 1-42, DOI: 10.1093/zoolinnean/zlad143, URL: https://doi.org/10.1093/zoolinnean/zlad143
C60DAE358626D837DBBFFEB6BE22FB93.taxon	description	(Figs 1 C, 5 D, 9 B, 10 I – K, 12 D, 13 D, 14 D) Zoobank registration: urn: lsid: zoobank. org: act: 6008 BBBE- 2 EF 2 - 4459 - 8 ACD- 5 AA 4 E 242 AA 4 A. Type material. Holotype: male, pinned, with genitalia in a separate microvial, with labels as follows: CHILE: [Los Lagos, Hualaihué] Palena Pr.: Austral Highway, km 62.9 (6.7 km S Contao turnoff), 230 m, 41 ° 51.155 ʹ S, 72 ° 42.175 ʹ W [- 41.85, - 72.70], 23 - 25. i. 1997, young secondary Valdivian rainforest; / beating flowers Tepulia stipularis (myrtaceae) [Myrtaceae] A. Newton and M. Thayer FIELD MUSEUM NAT. HIST. / HOLOTYPE Loncovilius variabilis des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Deposited at FMNH. Paratypes: CHILE: [Araucanía, Vilcún] Cautín Temuco, Cherquenco [~ - 38.69, - 72.00] I-II. 1978 L. E. Peña leg. FIELD MUS. NAT. HIST. [3 ♂ 1 ♀ FMNH]; CHILE: Reg. XI [Aysén] Queulat N. P. [~ - 44.38, - 72.25] P. M. Hammond / Below base 1 - 10. xii. 03 shrub flowers / NHM / [5 ♂ 4 ♀ NHM; 1 ♂ 1 ♀ NHMD ex. NHM; 1 ♀ CZUG ex. NHM]; [same data as above except:] Glacier Path 21. xi- 12. xii. 03 / Malaise trap [1 ♀ NHM]; [same data as above except:] Base area Beating [or] Base Camp Area 21. xi- 12. xii. 03 [8 ♂ 7 ♀ NHM]; CHILE: [Los Lagos, Hualaihué] Palena Pr.: Austral Highway, km 62.9 (6.7 km S Contao turnoff), 230 m, 41 ° 51.15 ’ S, 72 ° 42.17 ’ W [- 41.85, - 72.70], 23 - 25. i. 1997, young secondary Valdivian rainforest; / beating flowers Tepulia stipularis (myrtaceae) [Myrtaceae] A. Newton and M. Thayer FIELD MUSEUM NAT. HIST [5 ♂ FMNH]; Chile [Los Lagos] Llanquihue Lago Chapo [- 41.40, - 72.52] febrero 1985 leg. J. Zuñiga [1 ♂ 3 ♀ FMNH]; HORNOHUINCO SW. Lago Chapo Lianquihue [Los Lagos, Llanquihue, - 41.40, - 72.52] 21,24 - Dic. 1972 Coll: L. E. Pena [1 ♂ 1 ♀ FMNH]; [CHILE: Los Lagos] Rio Gol-Gol Osorno Prov. [~ - 40.66, - 72.18] / II- 8 - 57 Peña / CHILE [green label] / CNCI L 2004 - 28 / ♂ / FMNH-INS 0000 024 263 - 265 [3 ♂ CNC]; CHILE: Chiloé Pr., Isla Chiloé, Compu Alto [- 42.87, - 73.70], 18. i. 1998, T. Cekalovic leg. FIELD MUS. NAT. HIST. / FMNH-INS 0000 024 165 - 167 [1 ♂ FMNH; 1 ♂ 1 ♀ MNNC ex. FMNH]; CHILE: Prov. Chiloé: Isla Chiloé, Compu Alto [- 42.87, - 73.70] 18 - I- 1998 T. Cekalovic CHIL 1 C 98 001 / [Barcode] SM 0722732 KUNHM-ENT [1 ♂ SEMC]; [CHILE: Los Lagos, Puerto Montt] Prov. Llanquihue Correntoso [~ - 41.45, - 72.66] 22 - Enero- 1969 Coll: L. E. Pena [2 ♂ FMNH]; CHILE: [Los Lagos, Chonchi] Chiloé Pr.: Miraflores, road to (0.6 km W Hwy 5), 130 m, 42 ° 46.73 ʹ S 73 ° 47.71 ʹ W [- 42.7788, - 73.7953] 17. i. 1997, secondary Valdivian rainforest: / beating flowering Cald-cluvia paniculata [Caldcluvia paniculata] (Cunoniaceae) A. Newton and M. Thayer 994 FIELD MUS. NAT. HIST. [1 ♂ 1 ♀ FMNH]; [CHILE: Los Lagos, Hualaihué] 11 Chile X, peninsula Huequi under logs termas, Porcelana 42 ° 27 ʹ 30.8 ʹʹ S 72 ° 27 ʹ 15.0 ʹʹ W [- 42.4585, - 72.4541] 40 m R. Vila leg. 13. xii. 2014 / Loncovilius sp det. V. Assing 2015 / V. Assing Coll. J Jenkins Shaw det. 2016 [yellow label] [1 ♂ NHMW]; CHILE: [Los Lagos, Puyehue National Park] Osorno Prov. Parque Nac. Puyehue, 4.1 km E Anticura [- 40.66, - 72.12], 430 m, trap site 662 19 - 26. xii. 1982 Valdivian rainfor. A. Newton and M. Thayer / Screen- sweeping / Loncovilius sp. 7 det. A. Newton 1984 / ♀ / FMNH-INS 0000 024 163 [1 ♀ FMNH]; CHILE: [Los Lagos, Puyehue National Park] Osorno Prov., Puyehue Nat. Pk. Aguas Calientes [- 40.73, - 72.30], 500 m, 20. XII. 1984 - 8. II. 1985 / FMHD # 85 - 930 Derumbes for. Trail, S. andJ. Peck, P # 85 - 45, FIT FIELD MUSEUM NAT HIST / ♀ / FMNH-INS 0000 024 169 [1 ♂ FMNH]; CHILE: [Los Ríos] Valdivia Prov., 26 km SE Panguipulli [- 39.75, - 72.15] 300 m, 16. XII. 1984 - 11. II. 1985 / FMHD # 85 - 921, Nothofagus remnant S. andJ. Peck, P # 85 - 34, carrion trap FIELD MUSEUM NAT HIST / ♀ / FMNH-INS 0000 024 168 [1 ♀ FMNH]; CHILE: [Los Ríos] Prov. Valdivia: Parque Oncol (Cruce Sendero Tepual) Rio Cruces [- 39.72, - 73.27] 8 - I- 2001 T Cekalovic CHIL 1 CO 1 005 / [Barcode] SM 0723095 KUNHM-ENT [1 ♂ SEMC]; [CHILE: Los Ríos] 85 / Süd-Chile 31.1.90 Puerto Puyehue [~ - 40.65, - 72.33] leg. G. H. Schwabe [2 ♂ 1 ♀ SDEI]; [CHILE: Magallanes and Chilean Antarctica] Welling Is. Pto. Eden [- 49.11, - 74.40] 13. XII. 1958 / CHILE Fr Kuschel / Quedius nigroflavus Tottenham TYPE [manuscript name] / NHM [1 ♂ NHM]; [CHILE: Magallanes and Chilean Antarctica] Welling Is. Pto. Eden [- 49.11, - 74.40] 13. XII. 1958 / CHILE Fr Kuschel / Quedius nigroflavus Tottenham PARATYPE [manuscript name] / NHM [3 ♂ 2 ♀ NHM]; [CHILE: Magallanes and Chilean Antarctica] / Paratype / Welling Is. Pto. Eden [- 49.11, - 74.40] 30. XI. 1958 / C. E. Tottenham collection. B. M. 1974 - 587 / CHILE Fr Kuschel / nigroflavus Tot [manuscript name] / NHM / PARATYPE Loncovilius variabilis des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2023 / NHMUK 015025170 [1 ♀ NHM]; [CHILE: Magallanes and Chilean Antarctica] Welling Is. Pto. Eden [- 49.11, - 74.40] 13. XII. 1958 / CHILE Fr Kuschel / Qued. Sp. 5 / Loncovilius sp. Nov. P. M. Hammond det. 1980 / NHM [Blue label] / NHMD 917069 [1 ♀ NHM]; CHILE: Santiago [Santiago Metropolitan]: El Alfalfal [~ - 33.51, - 70.20] I. 1977 L. E. Peña, leg. FIELD MUS. NAT. HIST. / FMNH-INS 0000 024 157 – 162 [1 ♂ 2 ♀ FMNH; 1 ♂ 1 ♀ NHMD ex. FMNH; 1 ♂ CZUG ex. FMNH; * this is potentially a labelling error. The L. variabilis model shown in Figure 3 does not include this occurrence. The sets of variables and the calibration results that do include this occurrence can be consulted in the Supporting Information, Table S 1, and in the Supporting Information, File S 6]; [CHILE: unknown locality and date] Chili. / Chili [Green round label] / Sharp Coll 1905 - 313. / Quedius nigroflavus Tottenham PARATYPE [manuscript name] / NHM [1 ♀ NHM]; [CHILE: unknown locality and date] Chili Sharp Coll. / Quedius nigroflavus Brnh (Typ.) [manuscript name] / BRIT. MUSEUM DON. ARROW / B. Typ. ohne Flg. d streifen / Chicago NHMus M. Bernhauer Collection / Bernhauer MS name? Current genus: Loncovilius teste A. Newton 2000 [1 ♂ FMNH]; [CHILE: unknown locality and date] 53793 / Reed / Chili / Fry Coll. 1905.100 / Chicago NHMus M. Bernhauer Collection / ♂ / FMNH-INS 0000 024 171 [1 ♂ FMNH]. All paratypes with label: PARATYPE Loncovilius variabilis des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Diagnosis: Same as variabilis group diagnosis. Description: Measurements ♂ [min – max (average); N = 10]: FBL = 2.79 – 3.19 (2.98); TL = 5.12 – 6.25 (5.61); HW = 0.72 – 0.8 (0.77); HL = 0.74 – 0.83 (0.79); HW / HL = 0.95 – 1 (0.97); PW = 0.95 – 1.05 (1); PL = 0.88 – 0.98 (0.93); PW / PL = 1.02 – 1.09 (1.07); EW = 1.31 – 1.54 (1.38); EL = 1.17 – 1.43 (1.25); PW / HW = 1.23 – 1.33 (1.30). Measurements ♀ [min – max (average); N = 10]: FBL = 3.05 – 3.53 (3.33); TL = 5.5 – 7.44 (6.4); HW = 0.79 – 0.86 (0.83); HL = 0.82 – 0.9 (0.85); HW / HL = 0.94 – 1 (0.97); PW = 1.05 – 1.17 (1.10); PL = 1 – 1.08 (1.04); PW / PL = 1.03 – 1.09 (1.06); EW = 1.36 – 1.74 (1.56); EL = 1.22 – 1.55 (1.43); PW / HW = 1.29 – 1.39 (1.34). Head, pronotum, and elytra black to brown-black, head darker; antennae, mouthparts, and legs yellowish-brown; abdominal tergites and sternites variable in coloration, from dark reddish-brown to brown-black with pale posterior margin. Head from about as wide as long to, usually, slightly longer than wide; dorsally and ventrally glossy with few micropunctures and very fine microsculpture of transverse waves, without coarse non-setiferous punctures; posterior angles indistinct with sparse setiferous punctures of medium and small size. Eyes small (EYL / HL = x = 0.47), from 1.03 to 1.29 times as long as temples (in lateral view); distance between eyes about 1.25 times as long as length of eye in males and 1.34 times in females. Antennomere 3 slightly longer than antennomere 2; antennomeres 4 to 7 subequal in length; antennomeres 8 to 10 subequal in length; a 11 1.53 - 1.72 times as long as a 10 in males and 1.47 - 1.70 in females. Basal puncture usually double; parocular punctures usually single; posterior frontal puncture located anterior to temporal puncture; no small setiferous punctures between frontoclypeal and anterior frontal punctures; ventral basal ridge more or less straight in checkmark shape, almost united with gular sutures; postgenal ridge reduced to small trace near gular sutures; postmandibular ridge reaching close to eye margin; PMP greatly reduced; nuchal ridge present laterally and dorsally, fused with infraorbital ridge; gula with isodiametric microsculpture medially, gular sutures moderately and equally separated in both sexes. Mandibles without dorsolateral groove; labrum with sub-rectangular sclerotized region and entire to slightly notched apical margin; mentum with seta alpha only; penultimate labial palpomere only weakly dilated apicad, subequal in width with more or less subconical to fusiform apical palpomere. Pronotum slightly wider than long, convex, evenly curved, with two PPDS, without APP, with one SLSP; flexible postcoxal hypomeral extension (process) not interrupted by inferior line. Elytra evenly punctate, interspaces from about as long as, to 5 or even 10 times as long as, diameter of punctures. Mesosternum with four macrosetae arranged in a row medially. Protibiae not sexually dimorphic, without laterodorsal row of thick spines in both sexes. Mesotarsomeres 1 – 4 not sexually dimorphic, in both sexes with pale adhesive setae and mesotarsomeres 1 – 3 in both sexes with terminal plate. Metatarsomeres 2 – 4 with pale adhesive setae, without terminal plate; metatarsomere 1 shorter than 5, metatarsomere 4 ventrally with apical margin bilobed. Abdominal tergites III and IV with several rows of sparse and fine punctures of moderate density, without sizeable impunctate area; tergite VIII only slightly emarginate medio-apically in females. Sternite III with basal transverse carina strongly arcuate; lateral tergal sclerites IX not dorsoventrally or laterally flattened. Male sternite VIII with broad and shallow U-shaped emargination medially (Fig. 14 D); female sternite VIII without emargination; male sternite IX without emargination medially (apex subtruncate), its basal portion moderately longer than distal portion; male tergite X with anterior margin subtruncate (Fig. 12 D); female tergite X apically truncate (Fig. 13 D); ovipositor, its second gonocoxite with one macroseta medially. Aedeagus as in Figure 10 I – K; its total length ~ 0.95. Median lobe rod-like, apex not emarginate; internal sac with median vertical sclerite and pair of short I-shaped copulatory sclerites. Paramere slightly produced over apex of median lobe; wider than median lobe; converging into a subrounded apex, slightly projected in the middle; with dark peg setae; in lateral view paramere almost straight, slightly concave apically. Etymology: The species epithet is the Latin adjective ‘ variabilis ’ (variable) and refers to a notable intraspecific morphological variation of this taxon. This variation can occur between individuals from the same locality, e. g. it can be completely blackishbrown forms and others with a reddish-brown abdomen. The head punctation also varies, and loss or duplication of paraocular and basal punctures is common. Distribution and habitat: Loncovilius variabilis is known only from Chile, from the following regions: Araucanía, Aysén, Los Lagos, Los Ríos, Magallanes, and Chilean Antarctica, and from Santiago Metropolitan. The species occurs from lowlands to 1300 m of elevation in the mountains; confined to various types of forests in the Magellanic subpolar and Valdivian temperate forest ecoregions. It has been collected using Malaise traps, and directly from flowers, e. g. from Tepualia stipularis (Myrtaceae) and Caldcluvia paniculata (Cunoniaceae).	en	Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins, Solodovnikov, Alexey (2024): Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae). Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1): 1-42, DOI: 10.1093/zoolinnean/zlad143, URL: https://doi.org/10.1093/zoolinnean/zlad143
C60DAE358620D837D92CFB3CBED1F9A7.taxon	diagnosis	Diagnosis: Head elongate, without coarse non-setiferous punctures; nuchal and infraorbital ridges not joined; gula with isodiametric microsculpture in the middle; postmandibular ridge located close to eye margin; PMP strongly reduced or rarely absent; females with a 11 slightly longer than a 10 (a 11: a 10 ratio <1.5). Pronotum without SLSP. Tergites III and IV with only a few rows of coarse, sparse punctures, with large impunctate area near apical margin (Fig. 9 C); tergite VIII without medial apical emargination; ovipositor, its second gonocoxite with three or more macrosetae medially (Fig. 9 H).	en	Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins, Solodovnikov, Alexey (2024): Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae). Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1): 1-42, DOI: 10.1093/zoolinnean/zlad143, URL: https://doi.org/10.1093/zoolinnean/zlad143
C60DAE358620D834D971F935BA3EFA3B.taxon	description	(Figs 6 B, 7 F, 10 O – Q, 12 F, 13 F, 14 F) Zoobank registration: urn: lsid: zoobank. org: act: 9199958 E- 5 D 1 F- 4 C 33 - 9 DE 8 - 4 FBDDBFBFD 72. Type material. Holotype: male, mounted, with labels as follows: CHILE: Malleco Prov. 6.5 Km E Malalcahuello 1080 m, trap site 651 13 - 31. xii. 1982 North. Dombeyi with Chusquea A. Newton and M. Thayer / window trap / Loncovilius Sp. 5 Solodovnikov det. 2002 / L. cantharoides J. Jenkins Shaw det. 2016 / NHMD 916904 / HOLOTYPE Loncovilius cantharoides des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Deposited at FMNM. Paratypes: CHILE: Malleco Prov. 6.5 km E Malalcahuello [- 38.46, - 71.51], 1080 m, trap site 651, 13 - 31. xii. 1982 Noth. dombeyi [Nothofagus dombeyi] with Chusquea A. Newton and M. Thayer / window trap / FMNH-INS 0000 023 458 – 463, 465 – 466, 467 – 471, 472 – 474, 476 – 477, 478, 481, 484 – 485 [13 ♂ 9 ♀ FMNH]; CHILE: Malleco Prov., 40 km W [40 km E] Curacautin [- 38.44, - 71.50], 1500 m, 12. XII. 1984 - 16. II. 1985 / FMHD # 85 - 905, Nothofagus - Araucaria, S. Peck, P # 85 - 19, Malaise FIELD MUSEUM NAT HIST / FMNH-INS 0000 023 490 – 532, 602 – 613 [35 ♂ 24 ♀ FMNH]; CHILE: Malleco Prov. 14 km E Malalcahuello [- 38.43, - 71.48] 1570 m, trap site 649, 13 - 31. xii. 1982 Noth. pumilio-Araucaria f. [Nothofagus pumilio - Araucaria forest], A. Newton and M. Thayer / window trap 649 / FMNH-INS 0000 023 475, 478, 480, 482, 486 – 489 [3 ♂ 5 ♀ FMNH]; CHILE: Malleco Prov. 12 km E Malacahuello [- 38.43, - 71.49] 1350 m, trap site 650 13 - 31. xii. 1982 Noth. dombeyi-Araucaria f. A. Newton and M. Thayer / carrion trap (squid) / ♂ / FMNH-INS 0000 023 595 [1 ♂ FMNH]; CHILE, MallecoProv. 20 km E Manzanar [- 38.44, - 71.51] 1100 m 19 - 21. xii. 1976 H. F. Howden / Beating / CNCI L 2004 - 28 / ♂ / FMNH-INS 0000 023 593 - 594 [2 ♂ CNC]; CHILE: Malleco: Las Raices (W of) [- 38.54, - 71.45], 21 - 22. XII. 1976 L. E. Peña leg. FIELD MUSE. NAT. HIST. / FMNH-INS 0000 023 596 [2 ♂ FMNH]; [CHILE: Araucanía] Rio Blanco Malleco Prov. [- 39.10, - 71.61] / I- 27 - 59 Pena / CNC 928331 - 928333 [1 ♂ 2 ♀ CNC]; [CHILE: Araucanía] Cabrerias 1100 metro Cord. Nahuelbuta [- 37.71, - 73.03] 18. I. 1977 / Loncovilius sp. det. Newton 2000 / ♂ / FMNH-INS 0000 023 577 [1 ♂ 2 ♀ FMNH]; CHILE: [Araucanía] Malleco: Cord. Nahuelbuta, Cabreria [- 37.71, - 73.03] 1100 m, 14. XII. 1976 L. E. Peña leg. FIELD MUS. NAT. HIST. / FMNH-INS 0000 023 585 - 590, 592, 600 [6 ♂ 2 ♀ FMNH]; Chile [Araucanía, Curacautín]: P. N. Lonquimay, 23 / 12 / 2013, 38.44 ° S – 71.51 ° W [- 38.4400, - 71.5100], litter layer small stream [1 ♀ TSC]; Chile [Araucanía, Pucón]: Palguin, 05 / 12 / 2013, 39.43 ° S – 71.79 ° W [- 39.4300, - 71.7900], litter layer / Loncovilius n. sp. [1 ♂ 1 ♀ TSC]; [CHILE: Araucanía, Melipeuco] ♂ / Chile, La Araucania (IX Región), 85 km E Temuco, 26 km NNE Melipeuco, Parque Nacional Corguillío, NE coast of Laguna Conguillío, 38 ° 38.24 ʹ S, W 71 ° 36.74 ʹ W [- 38.6291, - 71.6203], 1200 m V. I. Gusarov 9. xii. 1999 / Loncovilius sp. n. A. Solodovnikov det. 2004 [1 ♂ NHMO]; CHILE: Nuble [Ñuble] Prov., 72 km SE Chillan, Trancas nr Termas [- 36.90, - 71.44], 1700 m, 6. XII. 1984 - 19. II. 1985 / FMNHD # 85 - 893, Nothofagus forest, S. Peck, P # 85 - 8 FIT FIELD MUSEUM NAT HIST / FMNH-INS 0000 023 578 – 584 [3 ♂ 4 ♀ FMNH]; ARGENTINA, Rio Negro 8: Lago Nahuel Huapi Puerto Blest [- 41.02, - 71.81], 770 m 4. xii. 1978 Mision Cientifica Danesa [pale orange label 2 ♂ 1 ♀ NHMD; 1 ♂ 1 ♀ MNNC ex. NHMD; 1 ♂ 1 ♀ CZUG ex. NHMD]; [ARGENTINA: same locality, only the date different (16. xii. 1978) 1 ♂ 3 ♀ NHMD; 1 ♂ 1 ♀ NHM ex. NHMD]; Argentina, Rio Negro Prov., Bariloche forest, Nothofagus antartica, unburnt, 41 ° 21 ʹ 9 ʹʹ S 71 ° 37 ʹ 34.5 ʹʹ W, [- 41.352500, - 71.626250] 823 ± 150 m, ABCONF 17 P. Sackmann / Argentina, Patagonia ABCONF 17 Paula Sackmann leg. / Loncovilius sp. ♀ V. I. Gusarov det. 2002 [1 ♀ NHMO]; ARGENTINA, Neuquén: Cerro Malo, 6 km N Pucara, Jan. 22, 1972 L. Herman 874 / AMNH 1489 [pale orange label] / FMNH-INS 0000 023 597 – 599 [1 ♂ 2 ♀ FMNH]. All paratypes with label: PARATYPE Loncovilius cantharoides des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Diagnosis: Nuchal ridge with small portion absent dorsally. Pronotum outline evenly curved. Elytra yellow or yellowish-brown with brown-black mark around mesoscutellum and suture. Protibiae sexually dimorphic, with laterodorsal row of thick spines only females. Aedeagus as in Figure 10 O – Q. Description: Measurements ♂ [min – max (average); N = 5]: FBL = 3.98 – 4.34 (4.22); TL = 7.18 – 8.4 (7.59); HW = 0.93 – 1.08 (1); HL = 1.03 – 1.23 (1.16); HW / HL = 0.88 – 0.91 (0.89); PW = 1.28 – 1.36 (1.32); PL = 1.25 – 1.36 (1.31); PW / PL = 1 – 1.04 (1.02); EW = 1.74 – 1.92 (1.84); EL = 1.59 – 1.81 (1.7); PW / HW = 1.26 – 1.28 (1.27). Measurements ♀ [min – max (average); N = 5]: FBL = 4.43 – 4.67 (4.55); TL = 8.13 – 8.47 (8.3); HW = 1.18 – 1.22 (1.20); HL = 1.25 – 1.33 (1.29); HW / HL = 0.92 – 0.94 (0.93); PW = 1.44 – 1.49 (1.47); PL = 1.4 – 1.44 (1.42); PW / PL = 1.03 – 1.03 (1.03); EW = 2 – 2.1 (2.05); EL = 1.78 – 1.9 (1.84); PW / HW = 4.43 – 4.67 (4.55). Head and pronotum brown-black with slight greenish, bluish, or golden overtones, with yellow margins; antennae, mouthparts, and legs yellow or yellowish-brown; elytra yellow with brown-black mark around mesoscutellum and suture; tergites and sternites dark reddish-brown with pale posterior margin. Head elongate; dorsally and ventrally dull with metallic iridescence; with micropunctures and microsculpture of transverse isodiametric meshes, without coarse non-setiferous punctures; posterior angles indistinct with several setiferous punctures of medium and small size. Eyes small (EYL / HL = x = 0.44), from 1.05 to 1.11 times as long as temples (in lateral view); distance between eyes about 1.24 times as long as length of eye in both sexes. Antennomere 3 moderately longer than antennomere 2; antennomeres 4 and 5 subequal in length; antennomeres 8 to 10 subequal in length; a 11 from 1.52 to 1.7 times long as a 10 in males and from 1.4 to 1.48 in females. Basal and parocular punctures usually double; posterior frontal puncture located anterior to temporal puncture; with small setiferous punctures between frontoclypeal and anterior frontal punctures: ventral basal ridge in checkmark shape, almost united with gular sutures; postgenal ridge reduced; postmandibular ridge reaching close to eye margin; PMP greatly reduced; nuchal ridge present laterally and a little dorsally, not fused with infraorbital ridge; gula with conspicuous isodiametric microsculpture medially, gular sutures widely and equally separated in both sexes. Mandibles without dorsolateral groove; labrum with trapezoidal sclerotized region and entire apical margin; mentum with seta alpha only; penultimate labial palpomere only weakly dilated apicad, subequal in width with more or less subconical apical palpomere. Pronotum about as wide as long, convex, evenly curved, with two PPDS, without APP, without SLSP; flexible postcoxal hypomeral extension (process) not interrupted by inferior line. Elytra evenly punctate, interspaces from about as long as, to four or five times as long as, diameter of punctures. Mesosternum with four macrosetae arranged in a row medially. Protibiae sexually dimorphic, with laterodorsal row of thick spines only females. In males, mesotarsomeres 1 – 4 with pale adhesive setae, mesotarsomeres 1 – 3 with terminal plate. In females, mesotarsomeres 1 – 4 with pale adhesive setae, without terminal plate. In both sexes, metatarsomeres 2 – 4 with pale adhesive setae, without terminal plate; metatarsomere 1 shorter than 5, metatarsomere 4 ventrally with apical margin bilobed. Abdominal tergites III and IV with only a few rows of coarse, sparse punctures, with large, impunctate area; tergite VIII not emarginate medio-apically. Sternite III with basal transverse carina evenly converging at obtuse angle; lateral tergal sclerites IX not dorsoventrally or laterally flattened. Male sternite VIII with U-shaped emargination medially (Fig. 14 F); female sternite VIII without emargination; male sternite IX without emargination medially, with rounded apex, its basal portion shorter than distal portion; male tergite X with anterior margin subtruncate (Fig. 12 F); female tergite X medio-apically distinctly subacute to acute (Fig. 13 F); ovipositor, its second gonocoxite with three macrosetae medially. Aedeagus as in Figure 10 O – Q; its total length 1.25 – 1.31. Median lobe rod-like; internal sac with pair of long inverted L-shaped copulatory sclerites. Paramere distinctly produced over apex of median lobe; wider than median lobe; converging to a subrectangular apex; with small translucent peg setae; in lateral view almost straight, curved away from median lobe apically. Etymology: The species epithet is derived from the Greek κανθαΡίς (kantharís; soldier beetles) and suffix - ειδής (- eid ḗ s; like), as the habitus resembles some species of the beetle family Cantharidae. Distribution and habitat: Loncovilius cantharoides is known from Chile, from the Araucanía and Ñuble regions, and from the westernmost portions of the Neuquén and Río Negro provinces of Argentina. The species occurs from 800 to 1600 m of elevation; reported in the Araucaria - Nothofagus forests from the central and north-central portions of the Valdivian temperate forest ecoregion. It has been collected using Malaise, window, carrion and dung-baited pitfall traps, and sifting leaf litter.	en	Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins, Solodovnikov, Alexey (2024): Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae). Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1): 1-42, DOI: 10.1093/zoolinnean/zlad143, URL: https://doi.org/10.1093/zoolinnean/zlad143
C60DAE358623D80ADBD3FA56BEF8FE79.taxon	description	(Figs 1 A, 6 A, 10 L – N, 12 E, 13 E, 14 E) = Loncovilius edwardsi (Bernhauer, 1935) nec Quedius edwardsi Sharp, 1886. Bernhauer, 1935: 92 (as Quedius edwardsi, original description); Korge, 1963 (as Quedius edwardsianus, replacement name); Coiffait and Sáiz 1966: 413 (Quedius edwardsianus, redescription without new material or data); Coiffait and Sáiz, 1968: 365 (listed as Quedius edwardsianus, without new data); Newton 2017: 28 (as Loncovilius edwardsianus, moved from Quedius to Loncovilius). Type material. Lectotype: (here designated): male, pinned, with genitalia in a separate microvial, with labels as follows: S. Chile: Llanquihue prov. F. and M. Edwards. B. M. 1927 – 63 / Casa Pangue [- 41.04, - 71.87] 4 - 10. xii. 1926. / Bernhauer det. [unreadable] / Chicago NHM M. Bernhauer collection / Quedius edwardsi Brh. n. sp. / Edwardsi Bernhauer Quedius / LECTOTYPE Loncovilius edwardsianus (Korge, 1963) des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Deposited at FMNH. Paralectotype: male, pinned, with genitalia attached to the card, with labels as follows: S. Chile: Llanquihue prov. F. and M. Edwards. B. M. 1927 – 63 / Casa Pangue [- 41.04, - 71.87] 4 - 10. xii. 1926. / Quedius edwardsi Bernh n. sp. / LECTOTYPE C. E. Tottenham of Q. edwardsi Bernh. Nec Sharp / Quedius ultor n. n. Tottenham / edwardsi repl. By edwardsianus Korge 1963 M. K. Thayer det. 2005 / NHMUK 015024785 / PARALECTOTYPE Loncovilius edwardsianus (Korge, 1963) des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2023. Deposited at NHM. Other material eoamined: Supporting Information, File S 7. Diagnosis: Nuchal ridge absent dorsally. Pronotum outline hexagonal. Elytra yellowish-brown to brown-black with greenish, bluish, or purplish iridescence. Protibiae without sexual dimorphism, both males and females without laterodorsal row of thick spines. Aedeagus as in Figure 10 L – N. Description: Measurements ♂ [min – max (average); N = 6]: FBL = 3.95 – 4.21 (4.1); TL = 7.92 – 8 (7.95); HW = 1 – 1.11 (1.05); HL = 1.13 – 1.25 (1.19); HW / HL = 0.83 – 0.92 (0.88); PW = 1.15 – 1.24 (1.21); PL = 1.15 – 1.25 (1.21); PW / PL = 0.95 – 1.03 (1); EW = 1.76 – 1.9 (1.81); EL = 1.67 – 1.71 (1.69); PW / HW = 1.11 – 1.21 (1.15). Measurements ♀ [min – max (average); N = 3]: FBL = 4.47 – 4.76 (4.59); TL = 7.32 – 8.56 (7.76); HW = 1.16 – 1.21 (1.18); HL = 1.3 – 1.36 (1.33); HW / HL = 0.87 – 0.9 (0.89); PW = 1.33 – 1.5 (1.4); PL = 1.31 – 1.44 (1.37); PW / PL = 1.01 – 1.04 (1.02); EW = 1.98 – 2.13 (2.06); EL = 1.83 – 1.96 (1.9); PW / HW = 1.15 – 1.24 (1.18). Head and pronotum dark brown to black with metallic blue overtones; antennae, mouthparts, pronotum at margins and legs yellowish-brown; elytra yellowish-brown to brown-black with greenish, bluish, or purplish iridescence, sometimes darker around mesocutellum and at suture; abdominal tergites and sternites brown-black or dark reddish-brown with pale posterior margin. Head elongate; dorsally and ventrally with iridescence and with micropunctures and microsculpture of transverse isodiametric meshes, without coarse non-setiferous punctures; posterior angles indistinct with moderate setiferous punctures of medium and small-size. Eyes small (EYL / HL = x = 0.44), from 1.18 to 1.46 times as long as temples (in lateral view) in males and 0.84 to 1.04 in females; distance between eyes about 1.13 times as long as length of eye in males and 1.44 times in females. Antennomere 3 moderately longer than antennomere 2; antennomeres 4 to 7 subequal in length; antennomeres 8 to 10 subequal in length; a 11 from 1.52 to 1.75 times as long as a 10 in males and from 1.23 to 1.4 in females. Basal and parocular punctures usually double; posterior frontal puncture located anterior to temporal puncture; without small setose punctures between frontoclypeal and anterior frontal punctures; ventral basal ridge more or less straight in checkmark shape, ending distinctly far from gular sutures; postgenal ridge reduced; postmandibular ridge reaching close to eye margin; PMP greatly reduced; nuchal ridge present only laterally, not fused with infraorbital ridge; gula with conspicuous isodiametric microsculpture medially, gular sutures moderately and equally separated in both sexes. Mandibles without dorsolateral groove; labrum with trapezoidal sclerotized region and entire apical margin; mentum with seta alpha only; penultimate labial palpomere only weakly dilated apicad, subequal in width with more or less subconical apical palpomere. Pronotum about as wide as long, convex, hexagonal, with two PPDS, without APP, without SLSP; flexible postcoxal hypomeral extension (process) not interrupted by inferior line. Elytra evenly punctate, interspaces from about as long as, to two or even up to six times as long as, diameter of punctures. Mesosternum with six macrosetae arranged in a row medially. Metasternum with several macrosetae. Protibiae not sexually dimorphic, without laterodorsal row of thick spines in both sexes. In males, mesotarsomeres 1 – 4 with pale adhesive setae, mesotarsomeres 1 – 3 with terminal plate. In females, mesotarsomeres 1 – 4 with pale adhesive setae, without terminal plate. In both sexes, metatarsomeres 2 – 4 with pale adhesive setae, without terminal plate; metatarsomere 1 shorter than 5, metatarsomere 4 ventrally with apical margin bilobed. Abdominal tergites III and IV with only a few rows of coarse, sparse punctures, with large impunctate area; tergite VIII not emarginate medio-apically. Sternite III with basal transverse carina usually evenly converging at an obtuse angle; lateral tergal sclerites IX not dorsoventrally or laterally flattened. Male sternite VIII with U-shaped emargination medially (Fig. 14 E); female sternite VIII without emargination; male sternite IX without emargination medially, with rounded apex, its basal portion shorter than distal portion; male tergite X apically subtruncate (Fig. 12 E); female tergite X with apical margin distinctly subacute to acute (Fig. 13 E); ovipositor, its second gonocoxite with three macrosetae medially. Aedeagus as in Figure 10 L – N; its total length 1.25 – 1.31. Median lobe rod-like, its apex in parameral view slightly V-shaped, emarginate, internal sac with pair of long inverted L-shaped copulatory sclerites. Paramere distinctly produced over apex of median lobe; wider than median lobe; converging to a round apex; with small translucent peg setae; in lateral view almost straight, curved away from median lobe apically. Distribution and habitat: Loncovilius edwardsianus is known from Chile, from the Araucanía, Aysén, Biobío, Los Lagos, and Los Ríos regions. In addition, it occurs in the westernmost portions of the Chubut and Río Negro provinces of Argentina. The species occurs from 200 to 1200 m elevation; it is reported from wet sclerophyll and open Nothofagus forests along the Valdivian temperate forest ecoregion. It has been collected by sifting leaf litter and moss. Remarks: Loncovilius edwardsianus was described under the preoccupied name Quedius edwardsi based on an unspecified number of syntypes from Casa Pangue in Llanquihue Province in Southern Chile. Korge (1963) replaced the name with Q. edwardsianus and, interestingly, this species was treated in the genus Quedius by Coiffait and Sáiz (1966, 1968) who neither studied the type material nor reported any new material. Presumably, all mentioned authors knew this species only from literature because even Sáiz (1971) did not mention it in the fauna of Chile. The species was transferred to the genus Loncovilius much later by Newton (2017). To fix the identity of the species, a male from the Bernhauer’s collection at the FMNH, which is clearly a syntype, is here designated as a lectotype of L. edwardsianus.	en	Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins, Solodovnikov, Alexey (2024): Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae). Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1): 1-42, DOI: 10.1093/zoolinnean/zlad143, URL: https://doi.org/10.1093/zoolinnean/zlad143
C60DAE35861DD80AD90CFE10BDBCFC36.taxon	diagnosis	Diagnosis: Head about as wide as long, without coarse non-setiferous punctures; nuchal ridge absent dorsally, nuchal and infraorbital ridges joined; gula without isodiametric microsculpture in the middle; postmandibular ridge located close to eye margin; PMP (usually reduced); females with antennomere 11 distinctly longer than antennomere 10 (a 11: a 10 ratio> 1.5). Pronotum outline evenly curved, with one SLSP. Protibiae not sexually dimorphic, in both sexes without laterodorsal row of thick spines. Tergites III and IV with only a few rows of coarse punctures of moderate density, with large impunctate area near apical margin (Fig. 9 D); tergite VIII with medial apical emargination in both sexes; ovipositor, its second gonocoxite with one macroseta medially (Fig. 9 G).	en	Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins, Solodovnikov, Alexey (2024): Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae). Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1): 1-42, DOI: 10.1093/zoolinnean/zlad143, URL: https://doi.org/10.1093/zoolinnean/zlad143
C60DAE35861DD80BD95BFC44BE97FCB5.taxon	description	(Figs 6 D, 11 D – F, 12 H, 13 H, 14 D) Zoobank registration: urn: lsid: zoobank. org: act: B 426 E 064 - 0842 - 4911 - A 48 A-C 7 A 73 A 73 B 5 E 0. Type material. Holotype: male, pinned, with genitalia in a separate microvial, with labels as follows: CHILE: Valdivia: Purulón [- 39.47, - 72.69], 10. I. 1986 L. E. Peña, leg. FIELD MUS. NAT. HIST. / ♂ / FMNH-INS 0000 023 665 / HOLOTYPE Loncovilius barclayi des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Deposited at FMNH. Paratypes: CHILE: Valdivia: Purulón [- 39.47, - 72.69], 10. I. 1986 L. E. Peña, leg. FIELD MUS. NAT. HIST. / ♂ / FMNH-INS 0000 023 667 / Loncovilius barclayi J. L. Reyes-Hernández [1 ♀ FMNH]; CHILE: Valdivia: Purulón [- 39.47, - 72.69], 10. I. 1986 L. E. Peña, leg. FIELD MUS. NAT. HIST. / ♂ / FMNH-INS 0000 023 666 / Loncovilius barclayi J. L. Reyes-Hernández [1 ♂ NHM ex. FMNH]; Nahuelbuta IX Region-Chile, 20 - I- 1993, Coll. T Curkovic / Loncovilius? semiflavus det. Newton 2000 [1 ♂]; [CHILE:] COLL. CERDA MNHN CHILE / Nahuelbuta Parque [- 37.80, - 73.01] 22.1.1981. P. [illegible] B / Santiago Mus [MNNC 1 ♂]; CHILE, IX r., 25 - 31. i. 2005, P. N. Nahuelbula [Araucanía, Nahuelbuta National Park] 37 ° 48 ʹ 530 73 ° 00 ʹ 954 [- 37.80, - 73.01], 1200 m, Sv Bíly lgt. / NHMD 916914 [1 ♂ NMPC]; [CHILE: no exact location] col. Fairmaire / 1251 / Quedius semiflavus n. sp. / FMNH-INS 0000 024 190 [1 ♂ MNNC ex. FMNH]. All paratypes with label: PARATYPE Loncovilius barclayi des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Diagnosis: Elytra yellow or yellowish-brown with black marks around mesoscutellum and on suture. Male tergite X trilobed (Fig. 12 H); female tergite X deeply emarginate medio-apically (Fig. 13 H). Aedeagus as in Figure 11 D – F. Description: Measurements ♂ [min – max (average); N = 6]: FBL = 2.42 – 3.86 (3.56); TL = 7.13 – 7.5 (7.32); HW = 0.9 – 0.98 (0.95); HL = 0.9 – 0.98 (0.95); HW / HL = 0.99 – 1.03 (1); PW = 1.19 – 1.31 (1.25); PL = 1.25 – 1.3 (1.28); PW / PL = 0.93 – 1.02 (0.98); EW = 1.54 – 1.75 (1.64); EL = 1.45 – 1.65 (1.56); PW / HW = 1.28 – 1.34 (1.31). Measurements ♀ (N = 1): FBL = 4.03; TL = 7.5; HW = 1.05; HL = 1; HW / HL = 1.05; PW = 1.38; PL = 1.38; PW / PL = 1; EW = 1.7; EL = 1.65; PW / HW = 1.31. Head and pronotum yellow-orange; antennae, mouthparts, and legs yellow or yellowish-brown; elytra yellow or yellowish-brown with dark mark around mesoscutellum and on suture; meso- and metasternum, all coxae, femora and abdomen dark reddish-brown; abdominal tergites and sternites with pale posterior margin. Head from slightly wider than long, to slightly longer than wide; dorsally and ventrally glossy with few micropunctures and very fine microsculpture of transverse waves, without coarse non-setiferous punctures; posterior angles indistinct with sparse setiferous punctures of medium and small-size. Eyes medium size (EYL / HL = x = 0.52), from 1.4 to 1.55 times longer than length of temples (in lateral view); distance between eyes about equal to length of eye. Antennomeres 2 and 3 subequal in length; antennomeres 4 to 6 subequal in length; antennomeres 6 to 10 gradually reduced in length; a 11 1.75 to 1.88 times as long as a 10 in males and 1.69 in females. Basal and parocular punctures usually single; posterior frontal puncture located anterior to temporal puncture; ventral basal ridge more or less straight in checkmark shape, almost united with gular sutures; postgenal ridge reduced; postmandibular ridge reaching close to eye margin; PMP greatly reduced; nuchal ridge present only laterally, fused with infraorbital ridge; gular sutures moderately separated. Mandibles without dorsolateral groove; labrum with trapezoidal sclerotized region and entire apical margin; mentum with seta alpha only; penultimate labial palpomere only weakly dilated apicad, subequal in width with more or less subconical apical palpomere. Pronotum about as wide as long, strongly convex, evenly curved, with two PPDS, without APP, with one SLSP; flexible postcoxal hypomeral extension (or process) not interrupted by inferior line. Elytra evenly punctate, interspaces from about as long as, to two or four times as long as, diameter of punctures. Mesosternum with four macrosetae arranged in a row medially. Protibiae not sexually dimorphic, without laterodorsal row of thick spines in both sexes. In males, mesotarsomeres 1 – 4 with pale adhesive setae, mesotarsomeres 1 – 3 with terminal plate. In females, mesotarsomeres 2 – 4 with pale adhesive setae without terminal plate. In both sexes, metatarsomeres 2 – 4 with pale adhesive setae, without terminal plate; metatarsomere 1 shorter than 5, metatarsomere 4 ventrally with apical margin straight or slightly sinuate. Abdominal tergites III and IV with only a few rows of coarse punctures of moderate density, with large impunctate area; tergite VIII distinctly emarginate medio-apically. Sternite III with basal transverse carina descending medially at a sharp point; lateral tergal sclerites IX not dorsoventrally or laterally flattened. Male sternite VIII with strong, V-shaped emargination medially (Fig. 14 H); female sternite VIII with weak apical medial emargination; male sternite IX not emarginate medio-apically, its basal portion distinctly longer than distal portion; male tergite X trilobed (Fig. 12 H); female tergite X deeply emarginate medio-apically (Fig. 13 H); ovipositor, its second gonocoxite with one macroseta medially (Fig 9 G). Aedeagus as in Figure 11 D – F, its length ~ 1.46; median lobe rod-like, in parameral view apex deeply emarginate; internal sac with pair of long I-shaped copulatory sclerites. Paramere slightly produced over apex of median lobe, wider than median lobe, its sides converging into a deeply emarginate apex, with dark peg setae, in lateral view almost straight and slightly concave apically. Etymology: The species epithet is a patronym in recognition of Maxwell V. L. Barclay, in particular for his earlier mentoring of J. J. S .. Distribution and habitat: Loncovilius barclayi is known only from Chile, from the Araucanía and Los Ríos regions, in the north-central portion of the Valdivian temperate forest ecoregion. It is unknown how it was collected.	en	Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins, Solodovnikov, Alexey (2024): Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae). Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1): 1-42, DOI: 10.1093/zoolinnean/zlad143, URL: https://doi.org/10.1093/zoolinnean/zlad143
C60DAE35861CD808D8FEFCDCBECAF946.taxon	description	(Figs 6 C, 7 D, 9 D, G, 11 A – C, 12 G, 13 G, 14 G) Fairmaire and Germain 1862: 428 (original description as Quedius semiflavus); Gemminger and Harold 1868: 570 (as synonym of Quedius lividipennis); Bernhauer and Schubert 1916: 434 (as Quedius semiflavus, catalogue); Germain 1903: 443 (as Loncovilius semiflavus, characters); Coiffait and Sáiz 1966: 411 (as Loncovilius s. s. semiflavus; characters); Coiffait and Sáiz, 1968: 365 (as Loncovilius s. s. semiflavus, checklist); Sáiz 1971: 385 (as Loncovilius s. s. semiflavus; additional material); Herman 2001 b: 3083 (as Loncovilius semiflavus; catalog). Type material. Lectotype: male, pinned, with genitalia in a separate microvial, with labels as follows: Coll. et det A. Fauvel Quedius semiflavus Fairm and Germ. R. I. S. c. N. B. 17.479 / Concepcion / Lectotype [Red label] / Loncovilius (s. s.) semiflavus 1964 det. Coiff. et Sáiz / J. L. Reyes-Hernández det. 2022 [RBINS]. Paralectotypes: Coll. et det. A. Fauvel, Quedius semiflavus Fairm and Germ., R. I. Sc. N. B. 17.479 / paralectotype [red paper] / Loncovilius semiflavus, 1964 det. Coiff. et Sáiz / J. L. Reyes-Hernández det. 2022; concepcion / semiflavus Fairm. Germ. / R. I. Sc. N. B. 17.479, Coll. et det. A. Fauvel / paralectotype [red paper] / Loncovilius semiflavus, 1964 det. Coiff. et Saiz / J. L. Reyes-Hernández det. 2022 [RBINS]. Other material eoamined: Supporting Information, File S 7. Diagnosis: Elytra translucent yellow or yellowish-brown. Male tergite X with broad U-shape emargination medially (Fig. 12 G); female tergite X slightly emarginate medio-apically (Fig. 13 G). Aedeagus as in Figure 11 A – C. Description: Measurements ♂ [min – max (average); N = 10]: FBL = 3.82 – 4.19 (4.02); TL = 6.25 – 6.88 (6.54); HW = 1 – 1.05 (1.03); HL = 0.98 – 1.09 (1.02); HW / HL = 0.96 – 1.04 (1.01); PW = 1.28 – 1.44 (1.35); PL = 1.28 – 1.43 (1.35); PW / PL = 0.99 – 1.01 (1); EW = 1.66 – 1.8 (1.74); EL = 1.5 – 1.8 (1.65); PW / HW = 1.27 – 1.37 (1.32). Measurements ♀ [min – max (average); N = 10]: FBL = 4.1 – 4.45 (4.29); TL = 7.2 – 8.6 (7.77); HW = 1.08 – 1.15 (1.1); HL = 1.04 – 1.13 (1.06); HW / HL = 1 – 1.06 (1.04); PW = 1.4 – 1.5 (1.47); PL = 1.38 – 1.5 (1.46); PW / PL = 0.98 – 1.05 (1.01); EW = 1.85 – 1.9 (1.88); EL = 1.67 – 1.82 (1.77); PW / HW = 1.27 – 1.39 (1.34). Head and pronotum yellow-orange or reddish-brown, usually females darker than males; elytra translucent yellow; antennae, mouthparts, and legs yellow or yellowish-brown; mesosternum, metasternum, metacoxa and abdomen dark reddish-brown; abdominal segments with pale posterior margin. Head from slightly wider than long, to slightly longer than wide; dorsally and ventrally glossy with few micropunctures and very fine microsculpture of transverse waves, without coarse non-setiferous punctures; posterior angles indistinct with sparse setiferous punctures of medium and small size. Eyes medium size (EYL / HL = x = 0.5), from 1.2 to 1.4 times as long as temples (in lateral view); distance between eyes about 1.2 times as long as length of eye in males and 1.39 times in females. Antennomeres 2 and 3 subequal in length; antennomeres 4 and 5 subequal in length; antennomeres 5 to 10 gradually becoming shorter apicad; a 11 is from 1.88 to 2 times as long as a 10 in males and from 1.6 to 1.8 times in females. Basal puncture single; parocular puncture usually single too, distinctly separated from eye margin; posterior frontal puncture located anterior to temporal puncture; ventral basal ridge rather straight, almost united with gular sutures; postgenal ridge reduced; postmandibular ridge reaching close to eye margin; PMP greatly reduced; nuchal ridge distinct only laterally, nuchal and infraorbital ridges fused; gular sutures moderately and equally separated in both sexes. Mandibles without dorsolateral groove; labrum with trapezoidal sclerotized region and entire apical margin; mentum with seta alpha only; penultimate labial palpomere only weakly dilated apicad, subequal in width with more or less subconical apical palpomere. Pronotum about as wide as long, strongly convex, evenly curved at sides, with two PPDS, without APP, with one SLSP; flexible postcoxal hypomeral extension (process) not interrupted by inferior line. Elytra evenly punctate, interspaces from about as long as, to three times as long as, diameter of punctures. Mesosternum with three or four macrosetae arranged in a row medially. Protibiae not sexually dimorphic, without laterodorsal row of thick spines in both sexes. In males, mesotarsomeres 1 – 4 with pale adhesive setae, mesotarsomeres 1 – 3 with terminal plate. In females, mesotarsomeres 2 – 4 with pale adhesive setae without terminal plate. In both sexes, metatarsomeres 2 – 4 with pale adhesive setae, without terminal plate; metatarsomere 1 shorter than 5, metatarsomere 4 ventrally with apical margin straight or slightly sinuate. Abdominal tergites III and IV with only a few rows of coarse punctures of moderate density, with large impunctate area; tergite VIII distinctly emarginate medio-apically. Sternite III with basal transverse carina descending medially at a sharp point; lateral tergal sclerites IX not dorsoventrally or laterally flattened. Male sterniteVIIIwithstrong (Fig. 14 G), V-shapedemarginationmedially; female sternite VIII with weak apical medial emargination; male sternite IX with arcuate emargination medially, its basal portion distinctly longer than distal portion; male tergite X with broad U-shape emargination medially (Fig. 12 G); female tergite X slightly emarginate medio-apically (Fig. 13 G); ovipositor, its second gonocoxite with one macroseta medially. Aedeagus as in Figure 11 A – C, its length 1.21; median lobe rod-like with moderately emarginate apex; paramere slightly produced over apex of median lobe, wider than median lobe, its sides converging in a trapezoidal slightly emarginate apex, with small translucent ‘ peg setae’, in lateral view almost straight and slightly concave apically. Distribution and habitat: Loncovilius semiflavus is known only from Chile, from the Biobío, Maule, Ñuble, and Santiago Metropolitan regions where it occurs from 200 to 1200 m of elevation. Its distribution is confined to the northern portions of the Valdivian temperate forest ecoregion. It has been collected by sifting leaf litter and beating the forest vegetation and flowers, e. g. from Drimys winteri (Winteraceae). Remarks: Lonvcovilius semiflavus was described as a species of Quedius based on an unspecified number of specimens from ‘ Concepcion’ (Concepción city in Central Chile) (Fairmaire and Germain 1862). Gemminger and Harold (1868) listed it as a synonym of Quedius lividipennis Fairmaire and Germain, 1862, but Germain (1903) considered both species valid when erecting a new genus Loncovilius for them. Coiffait and Sáiz (1966) examined three specimens from the collection of A. Fauvel in Brussels that they considered syntypes of L. semiflavus and designated one male as a lectotype. Morphology and labels of these three specimens match the original description and the type locality, respectively. According to Camousseight (1980), the type material of the species described by Philibert Germain is kept at the National Museum of Chile in Santiago. However, even though Camousseight (1980) listed type material for L. lividipennis (see below that species) and some other Amblyopinini as being deposited there, he did not mention any types of L. semiflavus. Presumably, they could have disappeared, as some other material did during a period when this collection was neglected (Camousseight, 1980), while the material from RBINS studied by Coiffait and Sáiz (1966) are syntypes that P. Germain could have shared with A. Fauvel, a prominent colleague of that time. Based on that material, Coiffait and Sáiz (1966) illustrated the aedeagus of L. semiflavus and, together with L. lividipennis, treated it as a member of the subgenus Loncovilius s. s.. Apart from the types, a specimen from La Jaula in Curicó Province of Chile listed in Sáiz (1971), was the only additional material reported for this species prior to our revision.	en	Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins, Solodovnikov, Alexey (2024): Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae). Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1): 1-42, DOI: 10.1093/zoolinnean/zlad143, URL: https://doi.org/10.1093/zoolinnean/zlad143
C60DAE35861FD808D90AF96BBA7DFE79.taxon	diagnosis	Diagnosis: Head about as wide as long, without coarse non-setiferous punctures; nuchal ridge complete dorsally, connected with infraorbital ridge; gula without isodiametric microsculpture in the middle; posterior frontal puncture located posterior to temporal puncture; postmandibular ridge located close to eye margin; PMP (usually reduced); females with antennomere 11 distinctly longer than antennomere 10 (a 11: a 10 ratio> 1.5). Pronotum outline evenly curved. Tergites III and IV with several rows of punctures of moderate size and density, without a sizeable impunctate area near to apical margin (Fig. 9 E); tergite VIII with medial apical emargination in both sexes; ovipositor, its second gonocoxite with one macroseta medially.	en	Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins, Solodovnikov, Alexey (2024): Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae). Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1): 1-42, DOI: 10.1093/zoolinnean/zlad143, URL: https://doi.org/10.1093/zoolinnean/zlad143
C60DAE35861FD809DB8AFE10BBDFFE67.taxon	description	(Figs 7 B, E, 8 A, 11 I, J, 12 J, 13 J, 14 J) Zoobank registration: urn: lsid: zoobank. org: act: C 2 A 7 BAAE- 3 D 40 - 4 ED 6 - 9 C 8 D- 09602 F 86 B 9 B 3. Type material. Holotype: male, pinned, with labels as follows: [CHILE: Magallanes and Chilean Antarctica, Puerto Edén] Wellington Is. Pto. Eden [- 49.11, - 74.40] 4. XII 1958 / CHILE Fr Kuschel / NHM [blue label] / NHMD 916967 / HOLOTYPE Loncovilius carlsbergi des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Deposited at NHM. Paratypes: [CHILE: Magallanes and Chilean Antarctica, Puerto Edén] Wellington Is. Pto. Eden 4. XII. 1958 / CHILE Fr Kuschel / NHM [blue label] [2 ♂ 6 ♀ NHM; 1 ♀ NHMD ex. NHM; 1 ♀ CZUG ex. NHM; 1 ♂ FMNH ex. NHM]; [same data as above except:] 20. XI. 1958 [5 ♂ 1 ♀ NHM; 1 ♀ FMNH ex. NHM]; [CHILE: Magallanes and Chilean Antarctica] Nothofagus antarctica / CHILE HE I. Puerto Eden, Isla Wellington [- 49.126, - 74.40] 490 S 40 ft. 29. xi. 1958 / NHM [1 ♀ NHM]; [same data as above except:] CHILE HE 6. [1 ♀ NHM]; [CHILE: Magallanes and Chilean Antarctica] S. CHILE Pto. Edén Wellington Is. [- 49.126, - 74.40] Dec. 7 - 15, ’ 62 PJDarlinton / FIELD MUSEUM ex MCZ (retained duplic.) / Loncovilius lividipennis (F + G) det. A. Newton 1993 [1 ♂ FMNH]; [CHILE: Magallanes and Chilean Antarctica] Nothofagus antarctica / CHILE Pena. Munoz, HM 3. Gamero. 20 – 40 ft. [- 52.33, - 73.53, 12 m] 27. xii. 1958 / NHM [3 ♂ 2 ♀ NHM; 1 ♂ 1 ♀ MNNC ex. NHM]; [CHILE: Magallanes and Chilean Antarctica] Munoz Gamero [- 52.33, - 73.53] 27. XII. 1958 / CHILE Fr. Kuschel / Shore / NHM [6 ♂ 3 ♀ NHM; 1 ♂ NHMD ex. NHM]; [CHILE: Magallanes and Chilean Antarctica] P. Voillians [Puerto Williams, - 54.93, - 67.619] 10.1 - 1959 / CHILE Fr Kuschel / 4 / NHM [1 ♂ CZUG ex. NHM]. All paratypes with label: PARATYPE Loncovilius carlsbergi des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Diagnosis: Pronotum with APP. Elytra yellowish-brown. Sternite III with basal transverse carina evenly converging at an obtuse angle. Aedeagus as in Figure 11 I, J. Description: Measurements ♂ [min – max (average); N = 6]: FBL = 3.5 – 3.95 (3.69); TL = 6.47 – 7.17 (6.71); HW = 0.87 – 0.95 (0.9); HL = 0.89 – 0.95 (0.91); HW / HL = 0.95 – 1.03 (0.99); PW = 1.19 – 1.3 (1.25); PL = 1.11 – 1.2 (1.16); PW / PL = 1.03 – 1.13 (1.08); EW = 1.75 – 1.86 (1.81); EL = 1.46 – 1.8 (1.61); PW / HW = 1.36 – 1.44 (1.38). Measurements ♀ [min – max (average); N = 6]: FBL = 3.73 – 3.94 (3.85); TL = 6.82 – 8.54 (7.92); HW = 0.91 – 0.96 (0.94); HL = 0.93 – 0.97 (0.95); HW / HL = 0.98 – 1 (0.99); PW = 1.2 – 1.36 (1.28); PL = 1.2 – 1.26 (1.23); PW / PL = 1 – 1.09 (1.04); EW = 1.71 – 1.96 (1.84); EL = 1.59 – 1.74 (1.67); PW / HW = 1.32 – 1.42 (1.36). Head, mandibles, legs, last antennomeres, and last palpomeres dark-brown; first palpomeres and antennomeres, and elytra yellow; pronotum yellowish-brown; tergites and sternites with pale posterior margin. Head from slightly wider than long, to slightly longer than wide; dorsally and ventrally glossy with few micropunctures and microsculpture of transverse waves, without coarse non-setiferous punctures; posterior angles indistinct with several setiferous punctures of medium and small-size. Eyes small size (EYL / HL = x = 0.45), from 1.27 to 1.35 times as long as temples (in lateral view) in males and from 1 to 1.16 times in females; distance between eyes about 1.24 times as long as length of eye in males and 1.41 times in females. Antennomeres 2 and 3 subequal in length; antennomeres 4 to 7 subequal in length; antennomeres 8 antennomeres 10 subequal in length; a 11 from 1.62 to 2 times as long as a 10 in males and from 1.56 to 1.69 in females. Basal and parocular punctures usually single; posterior frontal puncture located posterior to temporal puncture; without small setose punctures between frontoclypeal punctures and anterior frontal punctures; ventral basal ridge rather straight, ends distinctly far from gular sutures; postgenal ridge present; postmandibular ridge reaching close to eye margin; PMP greatly reduced; nuchal ridge present laterally and dorsally, fused with infraorbital ridge; gula without microsculpture medially, gular sutures moderately separated in males and widely in females. Mandibles without dorsolateral groove; labrum with trapezoidal sclerotized region and entire apical margin; mentum with seta alpha only; penultimate labial palpomere markedly dilated apicad, apical palpomere distinctly narrower than previous and more or less subconical. Pronotum slightly transverse, convex, evenly curved, with two PPDS, with an APP, with one SLSP; flexible postcoxal hypomeral extension (process) not interrupted by inferior line. Elytra evenly punctate, interspaces from about as long as, to three or five times as long as, diameter of punctures. Mesosternum with five macrosetae arranged in two rows medially. Protibiae sexually dimorphic, with laterodorsal row of thick spines only in females. In males, mesotarsomeres 1 – 4 with pale adhesive setae, mesotarsomeres 1 – 3 with terminal plate. In females, mesotarsomeres 2 – 4 with pale adhesive setae without terminal plate. In both sexes, metatarsomeres 2 – 4 with pale adhesive setae, without terminal plate; metatarsomere 1 moderately longer than 5, metatarsomere 4 ventrally with apical margin bilobed. Abdominal tergites III and IV with several rows of punctures of moderate size and density, without a sizeable impunctate area; tergite VIII emarginate medio-apically. Sternite III with basal transverse carina evenly converging at an obtuse angle; lateral tergal sclerites IX not dorsoventrally or laterally flattened. Male sternite VIII with rectangular V-shaped emargination medially (Fig. 14 J); female sternite VIII with arcuate emargination medially; male sternite IX with obtuse emargination medially, its basal portion moderately longer than distal portion; male tergite X with U-shaped emargination medially (Fig. 12 J); female tergite X with U-shaped emargination medially (Fig. 13 J); ovipositor, its second gonocoxite with one macroseta medially. Aedeagus as in Figure 11 I, J; its total length 1.35 – 1.45. Etymology: The species epithet is in honour of the Denmarkbased Carlsberg beer brewery, whose generous grants and beverages have facilitated much research in the Solodovnikov Lab. In addition, the male aedeagus resembles a bottle opener. Distribution and habitat: Loncovilius carlsbergi is known only from the Magallanes and Chilean Antarctica Region of Chile, where it occurs in lowlands confined to Magellanic subpolar forest ecoregions. It is unknown how it was collected.	en	Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins, Solodovnikov, Alexey (2024): Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae). Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1): 1-42, DOI: 10.1093/zoolinnean/zlad143, URL: https://doi.org/10.1093/zoolinnean/zlad143
C60DAE35861ED80EDB33FE0EBAF8FDA5.taxon	description	(Figs 7 A, 9 E, 11 G, H, 12 I, 13 I, 14 I) Fairmaire and Germain 1862: 430 (as Philonthus lividipennis, original description); Fauvel, 1866: 338 (as Quedius lividipennis, additional characters); Bernhauer and Schubert 1916: 427 (as Quedius lividipennis, catalogue); Scheerpeltz, 1972: 23 [as Quedius (Loncovilius) lividipennis; additional material]; Germain 1903: 443 (as Loncovilius lividipennis, characters); Coiffait and Sáiz 1966: 409 (as Loncovilius s. s. lividipennis; characters; additional material); Coiffait and Sáiz, 1968: 365 (as Loncovilius s. s. lividipennis, checklist); Sáiz 1971: 385 (as Loncovilius s. s. lividipennis; additional material); Herman 2001 b: 3083 (as Loncovilius lividipennis, catalogue); Jenkins Shaw et al., 2020: 337 (as Loncovilius nr. semiflavus; phylogeny). Material eoamined: Supporting Information, File S 7. Diagnosis: Pronotum very rarely with APP. Elytra yellow or yellowish-brown. Sternite III with basal transverse carina sharply pointed medioapically. Aedeagus as in Figure 11 G, H. Description: Measurements ♂ [min – max (average); N = 10]: FBL = 3.09 – 3.36 (3.22); TL = 5.15 – 6.86 (6.20); HW = 0.79 – 0.84 (0.81); HL = 0.78 – 0.84 (0.81); HW / HL = 0.98 – 1.02 (1); PW = 1.05 – 1.17 (1.11); PL = 0.97 – 1.08 (1.02); PW / PL = 1.08 – 1.11 (1.09); EW = 1.38 – 1.84 (1.56); EL = 1.33 – 1.44 (1.4); PW / HW = 1.31 – 1.42 (1.37). Measurements ♀ [min – max (average); N = 10]: FBL = 3.13 – 3.68 (3.42); TL = 5.87 – 6.68 (6.29); HW = 0.78 – 0.89 (0.85); HL = 0.8 – 0.9 (0.85); HW / HL = 0.98 – 1.02 (1); PW = 0.95 – 1.24 (1.14); PL = 0.89 – 1.16 (1.06); PW / PL = 1.04 – 1.11 (1.08); EW = 1.54 – 1.82 (1.64); EL = 1.33 – 1.63 (1.5); PW / HW = 1.1 – 1.41 (1.35). Most of antennae, mouthparts, pronotum, elytra, legs yellow or yellowish-brown; last three or five antennomeres and pronotum sometimes darker; head, mandibles, pro-, meso-, and metasternum, metacoxal, and abdomen reddish-brown; tergites and sternites with pale posterior margin. Head from slightly wider than long, to slightly longer than wide; dorsally and ventrally glossy with few micropunctures and microsculpture of transverse waves, without coarse non-setiferous punctures; posterior angles indistinct with several setiferous punctures of medium and small size. Eyes small (EYL / HL = x = 0.47), from 1.07 to 1.44 times as long as temples (in lateral view) in males and 1.06 to 1.25 in females; distance between eyes about 1.16 times as long as length of eye in males and 1.35 times in females. Antennomeres 2 and 3 subequal in length; antennomeres 4 to 7 subequal in length; antennomeres 8 to 10 subequal in length; a 11 from 1.88 to 2.2 times long as a 10 in males and from 1.72 to 1.94 in females. Basal and parocular punctures usually single; posterior frontal puncture located posterior to temporal puncture; without small setose punctures between frontoclypeal punctures and anterior frontal punctures; ventral basal ridge rather straight, almost united with gular sutures; postgenal ridge present; postmandibular ridge reaching close to eye margin; PMP greatly reduced; nuchal ridge present laterally and dorsally, fused with infraorbital ridge; gula without microsculpture medially, gular sutures moderately separated. Mandibles without dorsolateral groove; labrum with trapezoidal sclerotized region and entire apical margin; mentum with seta alpha only; penultimate labial palpomere markedly dilated apicad, apical palpomere distinctly narrower than previous and more or less subconical. Pronotum slightly transverse, convex, evenly curved, with two PPDS, usually without APP, with one SLSP; flexible postcoxal hypomeral extension (process) not interrupted by inferior line. Elytra evenly punctate, interspaces from about as long as, to one or three times as long as, diameter of punctures. Mesosternum with seven macrosetae arranged in two rows medially. Protibiae usually sexually dimorphic, with laterodorsal row of thick spines only in females; rarely in males too. In males, mesotarsomeres 1 – 4 with pale adhesive setae, mesotarsomeres 1 – 3 with terminal plate. In females, mesotarsomeres 2 – 4 with pale adhesive setae without terminal plate. In both sexes, metatarsomeres 2 – 4 with pale adhesive setae, without terminal plate; metatarsomere 1 moderately longer than 5, metatarsomere 4 ventrally with apical margin straight or slightly sinuate. Abdominal tergites III and IV with several rows of punctures of moderate size and density, without a sizeable impunctate area; tergite VIII emarginate medio-apically. Sternite III with basal transverse carina descending medially at a sharp point; lateral tergal sclerites IX not dorsoventrally or laterally flattened. Male sternite VIII with deep sharp V-shaped emargination medially (Fig. 14 I); female sternite VIII with arcuate emargination medially; male sternite IX with V-shaped emargination medially, its basal portion moderately longer than distal portion; male tergite X with deep U-shaped emargination medially (Fig. 12 I); female tergite X with V-shaped emargination medially (Fig. 13 I); ovipositor, its second gonocoxite with one macroseta medially. Aedeagus as in Figure 11 G, H; its total length 1.05 – 1.15. Distribution and habitat: Loncovilius lividipennis is known from Chile and Argentina. In Chile, it occurs in the Araucanía, Biobío, Los Lagos, Los Ríos, and Ñuble regions. In Argentina, it is confined to the westernmost portions of the Chubut, Neuquén, and Río Negro provinces. The species occurs from 550 to 1600 m elevation; it is reported in the Araucaria - Nothofagus forests from central and north-central portions of the Valdivian temperate forest ecoregion. It has been collected using Malaise and window traps, beating the forest vegetation, attracted to ultraviolet lights, sifted from moss and leaf litter, and directly from male cones of Araucaria araucana and flowers, e. g. from Desmaria mutabilis (Loranthaceae) on Nothofagus sp. Remarks: This species was described in the genus Philonthus (Fairmaire and Germain 1862: 430) based on an unspecified number of syntypes from the Andean forests near Chillan, a city in central Chile. Fauvel (1866) moved the species to the genus Quedius, while Germain (1903) to his genus Loncovilius. Scheerpeltz (1972), who treated Loncovilius at the rank of a subgenus of Quedius, listed significant new material for this species. Coiffait and Sáiz (1966) provided the species redescription and first illustration of the aedeagus based on the non-type material that comes from localities other than ‘ Chillan’. Sáiz (1971) listed a few more specimens. Camousseight (1980) lists a ‘ paratype N 2267 – 2272 ’ of Philonthus lividipennis from Germain’s collection at the National Museum of Chile in Santiago, which is apparently a syntype. Coiffait and Sáiz (1966) denoted one of the specimens they examined as a neotype of L. lividipennis. Although we were unable to examine any syntypes of L. lividipennis, we assume that the currently accepted concept of this species is correct, because the only other externally similar species, L. carlsbergi sp. nov., is known only further south, from the Magellanic subpolar forests ecoregion.	en	Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins, Solodovnikov, Alexey (2024): Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae). Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1): 1-42, DOI: 10.1093/zoolinnean/zlad143, URL: https://doi.org/10.1093/zoolinnean/zlad143
C60DAE358619D80FDBE6FDCCBEA6FD47.taxon	description	(Fig. 15 A) Apoquedius Scheerpeltz, 1972: 24, 25 (original description; as subgenus of Quedius; included species: aeneipennis and piciformis); Herman 2001 a: 24 (synonym of Loncovilius); Newton, 2022 (synonym of Loncovilius (Lienturius). Type genus: Apoquedius aeneipennis (Fairmaire and Germain, 1862) comb. nov. Included species: Apoquedius aeneipennis (Fairmaire and Germain, 1862) comb. nov., A. piciformis (Bernhauer, 1912) comb. nov. Diagnosis: Apoquedius can be diagnosed based on the head with frontoclypeal suture distinct only laterally; wide neck; paraocular punctures present; apical segment of maxillary palpi fusiform; apical segment of labial palpi fusiform, widest at the middle portion; apical labial palpomere distinctly longer than penultimate labial palpomere; flexible postcoxal hypomeral extension or process elongated and thin; two short empodial setae on each tarsal 5 segment; metacoxae with more than four spines on the posterior surface; abdomen with apical row of setae at the anterior margin of tergites and sternites III – VI; anterior margin of tergites III – VI distinctly crenulate. Apoquedius is easily distinguished from Loncovilius by the shape of the maxillary palps, which are fusiform; the shape of the postcoxal process which is thin and elongated; ventral side of meso- and metatarsi in males and females without pale adhesive setae; presence of protergal glands; presence of PTBC in tergites III to V; the most apical row of setae at the apical margin of tergites and sternites III to VI giving a crenulated appearance. Distribution: As of now, Apoquedius is restricted to Chile and Argentina. Remarks: The monophyly of Apoquedius is supported by a unique synapomorphy: flexible postcoxal hypomeral extension or process elongated and thin (character 58, state 1); and 11 homoplastic synapomorphies: head with frontoclypeal suture distinct only laterally (character 1, state 1); infraorbital ridge (IOR) not merged with postgenal ridge (PGR) (character 17, state 1); apical segment of maxillary palpi more or less fusiform, widest at its middle portion (character 29, state 0); apical segment of labial palpi more or less fusiform, widest at the middle portion (character 32, state 0); apical labial palpomere distinctly longer than penultimate labial palpomere (character 33, state 1); two parocular punctures on each side (character 41, state 1); basisternum from about as long as to moderately longer than furcasternum (character 46, state 0); protibiae, apical tibial spur distinctly outside tibial margin (character 66, state 1); more than four spines on the posterior surface of metacoxae (character 73, state 1); mesothorax with sternopleural (anapleural) suture transverse, or nearly transverse (very slightly oblique) (character 79, state 0); abdomen with apical row of setae at the anterior margin of tergites and sternites III – VI and anterior margin of tergites III – VI distinctly crenulate (character 90, state 1). Several undescribed species from southern South America and one described species from Tasmania likely belong to Apoquedius pending further examination.	en	Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins, Solodovnikov, Alexey (2024): Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae). Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1): 1-42, DOI: 10.1093/zoolinnean/zlad143, URL: https://doi.org/10.1093/zoolinnean/zlad143
C60DAE358618D80FD8CEFD6EBBB0FE79.taxon	description	(Fig. 15 B) Coiffait and Sáiz 1966: 403 [original description; as subgenus of Loncovilius; species included: aeneipennis (as heeri), discoideus, germaini, and leiocephalus]; Sáiz 1969: 9 (subgenus of Lienturius; notes); Sáiz 1971: 382, 383 (subgenus of Lienturius; characters; Chile); Newton, 2022 [synonym of Quedius (Apoquedius) Scheerpeltz, 1972]. Type species: Lienturius leiocephalus (Solier, 1849) comb. nov. Diagnosis: The phylogenetic relationship of Lienturius to the rest of the Amblyopinini genera remains unknown, so it is likely that the diagnosis of this genus will change as we wait for the results of our pending phylogenomic research. For the moment the only species of this genus is distinguished based on the combination of the following characters: head with three or more paraocular punctures on each side (not in a row), of which at least one is close to the eye margin and one is far from the eye margin and the remaining paraocular punctures, posterior frontal puncture anterior to temporal puncture; flexible postcoxal hypomeral extension (process) rounded and not interrupted by inferior line; elytra without humeral spines or spine-like setae; two short empodial setae on each tarsomere 5; tergites III – VI with PTBC, sternite III with basal transverse carina converging abruptly at an acute angle with rounded tip. Lienturius is easily distinguished from Loncovilius by the absence of a complete frontoclypeal suture; ventral side of meso- and metatarsi in males and females without pale adhesive setae; presence of protergal glands; presence of PTBC in tergites III to VI. Distribution: Only known from the Southern zone of Chile. Remarks: The new combination implemented here [Lienturius leiocephalus (Solier, 1849)] is supported by the phylogenetic analysis based on eight homoplasious synapomorphies: right mandible with proximal tooth and distal tooth, distal tooth not bifid, space between proximal and distal teeth narrow and smooth (character 24, state 2); labrum, shape of anterior margin (only sclerotized part, excluding apical membrane where present) entire, without emargination (character 28, state 1); three or more paraocular punctures (character 41, state 2); posterior frontal puncture anterior to temporal puncture (character 42, state 0); pronotum with SLSP (character 55, state 1); protarsi with tarsomeres 1 – 3 as long as wide (character 63, state 0); tergite VI with PTBC (character 89, state 1); sternite III with basal transverse carina converging abruptly at an acute angle with rounded tip (character 99, state 3).	en	Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins, Solodovnikov, Alexey (2024): Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae). Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1): 1-42, DOI: 10.1093/zoolinnean/zlad143, URL: https://doi.org/10.1093/zoolinnean/zlad143
C60DAE358618D80CDB06FE1CBEC6FA5B.taxon	description	(Fig. 15 C) Sphingoquedius Bernhauer 1941: 27 (original description; species included: strandi); Klimaszewski et al. (1996): 151 (Sphingoquedius; endemic to New Zealand); Solodovnikov and Schomann 2009: 34 (included in phylogenetic analysis); Jenkins Shaw and Solodovnikov (2016): 22 (absent on Lord Howe Island); Solodovnikov and Brunke (2016): 43 (new combination); Solodovnikov and Jenkins Shaw (2017): 318 (notes on relationship); Jenkins Shaw et al. 2017: 714 (included in phylogenetic analysis); Jenkins Shaw et al. 2020 a: 45 (notes on relationship); Jenkins Shaw et al. 2020 b: 2 (description of new fossil species). Type species: Sphingoquedius strandi Bernhauer, 1941. Included species: Sphingoquedius brevis (Sáiz 1971) comb. nov., S. discoideus (Fairmaire and Germain, 1862) comb. nov., S. meto Jenkins Shaw et al., 2020 b, S. nanus (Sáiz, 1971) comb. nov., S. novaezeelandiae (Duvivier, 1883), S. strandi Bernhauer, 1941. Diagnosis: The diagnosis of Sphingoquedius is likely to change pending phylogenomic results; however, these species can be tentatively placed in Sphingoquedius based on the large eyes and presence of patches of pale radiating setae on abdominal tergites III and IV. American Sphingoquedius are easily distinguished from Loncovilius by empodial setae as long as the tarsal claws; ventral side of meso- and metatarsi in males and females without pale adhesive setae; presence of protergal glands; presence of patches of pale radiating setae on abdominal tergites; sternite III with basal transverse carina converges abruptly at an acute angle with rounded tip (projection sides invaginated). Distribution: After the taxonomic changes implemented here, Sphingoquedius occurs in New Zealand and southern South America. A single fossil species (Sphingoquedius meto) is known from the earliest Miocene of New Zealand (Jenkins Shaw et al. 2020 b). Remarks: The new combinations implemented here [Sphingoquedius nanus (Sáiz, 1971); Sphingoquedius brevis (Sáiz, 1971); Sphingoquedius discoideus (Fairmaire and Germain, 1862)] are supported by the phylogenetic analysis where species of Sphingoquedius are united based on two unique synapomorphies: empodial setae longer or at least as long as tarsal claws (character 62, state 1; empodial setae missing in S. strandi); abdomen with tergites III and IV bearing patches of pale radiating setae (character 93, state 2; also see: Jenkins Shaw et al. 2020 b: fig. 2.3); and a single homoplasious synapomorphy: postmandibular ridge located close to eye margin (character 16, state 1). Numerous described and undescribed species from New Zealand and Australia may also belong to Sphingoquedius and will be dealt with separately pending further study. Even though the generic placement of the new combinations implemented here may change further, their placement in Sphingoquedius is supported by the phylogenetic analysis and can at least be regarded as a step in the right direction.	en	Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins, Solodovnikov, Alexey (2024): Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae). Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1): 1-42, DOI: 10.1093/zoolinnean/zlad143, URL: https://doi.org/10.1093/zoolinnean/zlad143
