taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
C63C340FB7314A6D1292F9AE6EEB95B1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359956/files/figure.png	https://doi.org/10.5281/zenodo.17359956	Figure 17. Hypoctoninae Pocock, 1899: pedipalp tibia (manus), retrolateral aspect. (A) Thelyphonellus amazonicus (Butler, 1972), ♂(MPEG [URO] 9). (B) Wounaan vanegasae (Giupponi and Vasconcelos, 2008), comb. n., ♂(MCZ 64455). (C) Yekuana venezolensis (Haupt, 2009), comb. n., holotype ♂(ZMB 48289). Abbreviations: bl, basal lobe; dl, distal lobe. Scale bars = 0.5 mm (A), 1 mm (B,C).	Figure 17. Hypoctoninae Pocock, 1899: pedipalp tibia (manus), retrolateral aspect. (A) Thelyphonellus amazonicus (Butler, 1972), ♂(MPEG [URO] 9). (B) Wounaan vanegasae (Giupponi and Vasconcelos, 2008), comb. n., ♂(MCZ 64455). (C) Yekuana venezolensis (Haupt, 2009), comb. n., holotype ♂(ZMB 48289). Abbreviations: bl, basal lobe; dl, distal lobe. Scale bars = 0.5 mm (A), 1 mm (B,C).	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB7314A6D1292F9AE6EEB95B1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359918/files/figure.png	https://doi.org/10.5281/zenodo.17359918	Figure 9. Hypoctoninae Pocock, 1899: pedipalp tibia (manus), retrolateral aspect (A) and opisthosoma, lateral aspect (B), illustrating surface macro- and microsculpture. (A) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 0.25 mm (A), 0.5 mm (B).	Figure 9. Hypoctoninae Pocock, 1899: pedipalp tibia (manus), retrolateral aspect (A) and opisthosoma, lateral aspect (B), illustrating surface macro- and microsculpture. (A) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 0.25 mm (A), 0.5 mm (B).	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB7314A6D1292F9AE6EEB95B1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359946/files/figure.png	https://doi.org/10.5281/zenodo.17359946	Figure 13. Hypoctoninae Pocock, 1899: opisthosomal flagellum (whip), dorsal (A,B) and ventral (C) aspects. (A,B) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831), proximal segments (A) and segments V to IX (B). (C) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050), complete flagellum. Scale bars = 1 mm (A,B), 2 mm (C).	Figure 13. Hypoctoninae Pocock, 1899: opisthosomal flagellum (whip), dorsal (A,B) and ventral (C) aspects. (A,B) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831), proximal segments (A) and segments V to IX (B). (C) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050), complete flagellum. Scale bars = 1 mm (A,B), 2 mm (C).	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB7314A6D1292F9AE6EEB95B1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359906/files/figure.png	https://doi.org/10.5281/zenodo.17359906	Figure 7. Hypoctoninae Pocock, 1899: pedipalp trochanter (A,B), femur (C,D), and patella (E,F), dorsal aspect. (A,C,E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D,F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Abbreviations: ap, apophysis; ppd, principal prodorsal tubercle; pv, proventral tubercle. Scale bars = 1 mm.	Figure 7. Hypoctoninae Pocock, 1899: pedipalp trochanter (A,B), femur (C,D), and patella (E,F), dorsal aspect. (A,C,E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D,F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Abbreviations: ap, apophysis; ppd, principal prodorsal tubercle; pv, proventral tubercle. Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB7314A6D1292F9AE6EEB95B1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359916/files/figure.png	https://doi.org/10.5281/zenodo.17359916	Figure 8. Hypoctoninae Pocock, 1899: pedipalp tibia (manus), retrolateral (A,B), retrodorsal (C,D), and proventral (E,F) aspects. (A,C,E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D,F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Abbreviations: ap, apophysis; bl, basal lobe; dl, distal lobe; pvd, proventral distal tubercle. Scale bars = 1 mm.	Figure 8. Hypoctoninae Pocock, 1899: pedipalp tibia (manus), retrolateral (A,B), retrodorsal (C,D), and proventral (E,F) aspects. (A,C,E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D,F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Abbreviations: ap, apophysis; bl, basal lobe; dl, distal lobe; pvd, proventral distal tubercle. Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB7314A6D1292F9AE6EEB95B1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359883/files/figure.png	https://doi.org/10.5281/zenodo.17359883	Figure 4. Hypoctoninae Pocock, 1899: carapace, dorsal aspect (A,B) and coxosternal region, ventral aspect (C,D). (A,C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm.	Figure 4. Hypoctoninae Pocock, 1899: carapace, dorsal aspect (A,B) and coxosternal region, ventral aspect (C,D). (A,C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB7314A6D1292F9AE6EEB95B1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359875/files/figure.png	https://doi.org/10.5281/zenodo.17359875	Figure 2. Map of the Caribbean and northern South America, plotting known locality records of Neotropical whip-scorpions (Hypoctoninae Pocock, 1899), based on material examined and published records. Abbreviations: Bra, Brazil; Col, Colombia; DRe, Dominican Republic; Ecu, Ecuador; FGu, French Guiana; Guy, Guyana; Hai, Haiti; Pan, Panama; Per, Peru; PRi, Puerto Rico; Sur, Suriname; Ven, Venezuela.	Figure 2. Map of the Caribbean and northern South America, plotting known locality records of Neotropical whip-scorpions (Hypoctoninae Pocock, 1899), based on material examined and published records. Abbreviations: Bra, Brazil; Col, Colombia; DRe, Dominican Republic; Ecu, Ecuador; FGu, French Guiana; Guy, Guyana; Hai, Haiti; Pan, Panama; Per, Peru; PRi, Puerto Rico; Sur, Suriname; Ven, Venezuela.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB7314A6D1292F9AE6EEB95B1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359873/files/figure.png	https://doi.org/10.5281/zenodo.17359873	Figure 1. Single most parsimonious tree (L = 53) obtained by analysis of 45 morphological characteristics for ten species of whip-scorpions (Hypoctoninae Pocock, 1899), via implicit enumeration with equal weights. Morphological synapomorphies (using ACCTRAN optimization) illustrated on each branch, with numbers above indicating characteristics and numbers below indicating character states. Solid (black) circles indicate uniquely derived character states, and empty (white) circles indicate parallel derivations. Measure of support (Goodman–Bremer) for branches indicated at nodes. Tree consistency index (CI): 0.87; tree retention index (RI): 0.85. Refer to Appendix B for character matrix and Appendix C for list of characters.	Figure 1. Single most parsimonious tree (L = 53) obtained by analysis of 45 morphological characteristics for ten species of whip-scorpions (Hypoctoninae Pocock, 1899), via implicit enumeration with equal weights. Morphological synapomorphies (using ACCTRAN optimization) illustrated on each branch, with numbers above indicating characteristics and numbers below indicating character states. Solid (black) circles indicate uniquely derived character states, and empty (white) circles indicate parallel derivations. Measure of support (Goodman–Bremer) for branches indicated at nodes. Tree consistency index (CI): 0.87; tree retention index (RI): 0.85. Refer to Appendix B for character matrix and Appendix C for list of characters.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB7314A6D1292F9AE6EEB95B1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359895/files/figure.png	https://doi.org/10.5281/zenodo.17359895	Figure 5. Hypoctoninae Pocock, 1899: opisthosoma, dorsal (A,B) and ventral (C,D) aspects. (A,C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm.	Figure 5. Hypoctoninae Pocock, 1899: opisthosoma, dorsal (A,B) and ventral (C,D) aspects. (A,C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB7314A6D1292F9AE6EEB95B1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359935/files/figure.png	https://doi.org/10.5281/zenodo.17359935	Figure 11. Hypoctoninae Pocock, 1899: dissected sternite II (genital) illustrating internal surface with sclerites and gonopods, dorsal (A,B), posterodorsal (C,D), and dorsal, slightly posterior (E) aspects. (A,C,E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Abbreviations: Fi, fistula; LaM, lamina medialis; LoD, lobus dorsalis; LoL1, lobus lateralis primus; LoL2, lobus lateralis secundus; Me, mensa. Scale bars = 1 mm.	Figure 11. Hypoctoninae Pocock, 1899: dissected sternite II (genital) illustrating internal surface with sclerites and gonopods, dorsal (A,B), posterodorsal (C,D), and dorsal, slightly posterior (E) aspects. (A,C,E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Abbreviations: Fi, fistula; LaM, lamina medialis; LoD, lobus dorsalis; LoL1, lobus lateralis primus; LoL2, lobus lateralis secundus; Me, mensa. Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB7314A6D1292F9AE6EEB95B1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359954/files/figure.png	https://doi.org/10.5281/zenodo.17359954	Figure 16. Hypoctoninae Pocock, 1899: coxosternal region, ventral aspect. (A) Etienneus africanus (Hentschel, 1899), ♀(AMCC [LP 4654]). (B) Ravilops wetherbeei (Armas, 2002), ♀(AMCC [LP 3342]). (C) Thelyphonellus amazonicus (Butler, 1972), ♀(MPEG [URO] 7). (D) Wounaan vanegasae (Giupponi and Vasconcelos, 2008), comb. n., ♂(MCZ 64455). (E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Markedly sclerotized parts of the median sternum (mesosternum) indicated in black. Scale bars = 1 mm.	Figure 16. Hypoctoninae Pocock, 1899: coxosternal region, ventral aspect. (A) Etienneus africanus (Hentschel, 1899), ♀(AMCC [LP 4654]). (B) Ravilops wetherbeei (Armas, 2002), ♀(AMCC [LP 3342]). (C) Thelyphonellus amazonicus (Butler, 1972), ♀(MPEG [URO] 7). (D) Wounaan vanegasae (Giupponi and Vasconcelos, 2008), comb. n., ♂(MCZ 64455). (E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Markedly sclerotized parts of the median sternum (mesosternum) indicated in black. Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB7314A6D1292F9AE6EEB95B1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359880/files/figure.png	https://doi.org/10.5281/zenodo.17359880	Figure 3. Hypoctoninae Pocock, 1899: habitus, dorsal aspect. (A) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bar = 5 mm.	Figure 3. Hypoctoninae Pocock, 1899: habitus, dorsal aspect. (A) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bar = 5 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB7314A6D1292F9AE6EEB95B1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359903/files/figure.png	https://doi.org/10.5281/zenodo.17359903	Figure 6. Hypoctoninae Pocock, 1899: chelicera, prolateral (A,C) and retrolateral (B,D) aspects. (A,B) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (C,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 0.5 mm.	Figure 6. Hypoctoninae Pocock, 1899: chelicera, prolateral (A,C) and retrolateral (B,D) aspects. (A,B) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (C,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 0.5 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB7334A661292FF5D690B9581.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359873/files/figure.png	https://doi.org/10.5281/zenodo.17359873	Figure 1. Single most parsimonious tree (L = 53) obtained by analysis of 45 morphological characteristics for ten species of whip-scorpions (Hypoctoninae Pocock, 1899), via implicit enumeration with equal weights. Morphological synapomorphies (using ACCTRAN optimization) illustrated on each branch, with numbers above indicating characteristics and numbers below indicating character states. Solid (black) circles indicate uniquely derived character states, and empty (white) circles indicate parallel derivations. Measure of support (Goodman–Bremer) for branches indicated at nodes. Tree consistency index (CI): 0.87; tree retention index (RI): 0.85. Refer to Appendix B for character matrix and Appendix C for list of characters.	Figure 1. Single most parsimonious tree (L = 53) obtained by analysis of 45 morphological characteristics for ten species of whip-scorpions (Hypoctoninae Pocock, 1899), via implicit enumeration with equal weights. Morphological synapomorphies (using ACCTRAN optimization) illustrated on each branch, with numbers above indicating characteristics and numbers below indicating character states. Solid (black) circles indicate uniquely derived character states, and empty (white) circles indicate parallel derivations. Measure of support (Goodman–Bremer) for branches indicated at nodes. Tree consistency index (CI): 0.87; tree retention index (RI): 0.85. Refer to Appendix B for character matrix and Appendix C for list of characters.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB7334A661292FF5D690B9581.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359875/files/figure.png	https://doi.org/10.5281/zenodo.17359875	Figure 2. Map of the Caribbean and northern South America, plotting known locality records of Neotropical whip-scorpions (Hypoctoninae Pocock, 1899), based on material examined and published records. Abbreviations: Bra, Brazil; Col, Colombia; DRe, Dominican Republic; Ecu, Ecuador; FGu, French Guiana; Guy, Guyana; Hai, Haiti; Pan, Panama; Per, Peru; PRi, Puerto Rico; Sur, Suriname; Ven, Venezuela.	Figure 2. Map of the Caribbean and northern South America, plotting known locality records of Neotropical whip-scorpions (Hypoctoninae Pocock, 1899), based on material examined and published records. Abbreviations: Bra, Brazil; Col, Colombia; DRe, Dominican Republic; Ecu, Ecuador; FGu, French Guiana; Guy, Guyana; Hai, Haiti; Pan, Panama; Per, Peru; PRi, Puerto Rico; Sur, Suriname; Ven, Venezuela.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB7334A661292FF5D690B9581.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359880/files/figure.png	https://doi.org/10.5281/zenodo.17359880	Figure 3. Hypoctoninae Pocock, 1899: habitus, dorsal aspect. (A) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bar = 5 mm.	Figure 3. Hypoctoninae Pocock, 1899: habitus, dorsal aspect. (A) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bar = 5 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB7334A661292FF5D690B9581.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359883/files/figure.png	https://doi.org/10.5281/zenodo.17359883	Figure 4. Hypoctoninae Pocock, 1899: carapace, dorsal aspect (A,B) and coxosternal region, ventral aspect (C,D). (A,C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm.	Figure 4. Hypoctoninae Pocock, 1899: carapace, dorsal aspect (A,B) and coxosternal region, ventral aspect (C,D). (A,C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB7334A661292FF5D690B9581.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359895/files/figure.png	https://doi.org/10.5281/zenodo.17359895	Figure 5. Hypoctoninae Pocock, 1899: opisthosoma, dorsal (A,B) and ventral (C,D) aspects. (A,C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm.	Figure 5. Hypoctoninae Pocock, 1899: opisthosoma, dorsal (A,B) and ventral (C,D) aspects. (A,C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB7334A661292FF5D690B9581.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359903/files/figure.png	https://doi.org/10.5281/zenodo.17359903	Figure 6. Hypoctoninae Pocock, 1899: chelicera, prolateral (A,C) and retrolateral (B,D) aspects. (A,B) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (C,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 0.5 mm.	Figure 6. Hypoctoninae Pocock, 1899: chelicera, prolateral (A,C) and retrolateral (B,D) aspects. (A,B) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (C,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 0.5 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB7334A661292FF5D690B9581.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359906/files/figure.png	https://doi.org/10.5281/zenodo.17359906	Figure 7. Hypoctoninae Pocock, 1899: pedipalp trochanter (A,B), femur (C,D), and patella (E,F), dorsal aspect. (A,C,E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D,F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Abbreviations: ap, apophysis; ppd, principal prodorsal tubercle; pv, proventral tubercle. Scale bars = 1 mm.	Figure 7. Hypoctoninae Pocock, 1899: pedipalp trochanter (A,B), femur (C,D), and patella (E,F), dorsal aspect. (A,C,E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D,F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Abbreviations: ap, apophysis; ppd, principal prodorsal tubercle; pv, proventral tubercle. Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB7334A661292FF5D690B9581.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359916/files/figure.png	https://doi.org/10.5281/zenodo.17359916	Figure 8. Hypoctoninae Pocock, 1899: pedipalp tibia (manus), retrolateral (A,B), retrodorsal (C,D), and proventral (E,F) aspects. (A,C,E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D,F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Abbreviations: ap, apophysis; bl, basal lobe; dl, distal lobe; pvd, proventral distal tubercle. Scale bars = 1 mm.	Figure 8. Hypoctoninae Pocock, 1899: pedipalp tibia (manus), retrolateral (A,B), retrodorsal (C,D), and proventral (E,F) aspects. (A,C,E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D,F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Abbreviations: ap, apophysis; bl, basal lobe; dl, distal lobe; pvd, proventral distal tubercle. Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB7334A661292FF5D690B9581.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359918/files/figure.png	https://doi.org/10.5281/zenodo.17359918	Figure 9. Hypoctoninae Pocock, 1899: pedipalp tibia (manus), retrolateral aspect (A) and opisthosoma, lateral aspect (B), illustrating surface macro- and microsculpture. (A) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 0.25 mm (A), 0.5 mm (B).	Figure 9. Hypoctoninae Pocock, 1899: pedipalp tibia (manus), retrolateral aspect (A) and opisthosoma, lateral aspect (B), illustrating surface macro- and microsculpture. (A) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 0.25 mm (A), 0.5 mm (B).	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB7334A661292FF5D690B9581.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359929/files/figure.png	https://doi.org/10.5281/zenodo.17359929	Figure 10. Hypoctoninae Pocock, 1899: leg I tarsus (A,B) and closeup (C,D), retroventral (A,C), prolateral (B), and retrodorsal (D) aspects. (A,C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm (A,B), 0.5 mm (C,D).	Figure 10. Hypoctoninae Pocock, 1899: leg I tarsus (A,B) and closeup (C,D), retroventral (A,C), prolateral (B), and retrodorsal (D) aspects. (A,C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm (A,B), 0.5 mm (C,D).	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB7334A661292FF5D690B9581.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359935/files/figure.png	https://doi.org/10.5281/zenodo.17359935	Figure 11. Hypoctoninae Pocock, 1899: dissected sternite II (genital) illustrating internal surface with sclerites and gonopods, dorsal (A,B), posterodorsal (C,D), and dorsal, slightly posterior (E) aspects. (A,C,E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Abbreviations: Fi, fistula; LaM, lamina medialis; LoD, lobus dorsalis; LoL1, lobus lateralis primus; LoL2, lobus lateralis secundus; Me, mensa. Scale bars = 1 mm.	Figure 11. Hypoctoninae Pocock, 1899: dissected sternite II (genital) illustrating internal surface with sclerites and gonopods, dorsal (A,B), posterodorsal (C,D), and dorsal, slightly posterior (E) aspects. (A,C,E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Abbreviations: Fi, fistula; LaM, lamina medialis; LoD, lobus dorsalis; LoL1, lobus lateralis primus; LoL2, lobus lateralis secundus; Me, mensa. Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB7334A661292FF5D690B9581.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359942/files/figure.png	https://doi.org/10.5281/zenodo.17359942	Figure 12. Hypoctoninae Pocock, 1899: opisthosoma, pygidium and base of flagellum (whip), dorsal (A,D), lateral (B,E), and ventral (C,F) aspects. (A–C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (D–F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm.	Figure 12. Hypoctoninae Pocock, 1899: opisthosoma, pygidium and base of flagellum (whip), dorsal (A,D), lateral (B,E), and ventral (C,F) aspects. (A–C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (D–F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB7334A661292FF5D690B9581.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359946/files/figure.png	https://doi.org/10.5281/zenodo.17359946	Figure 13. Hypoctoninae Pocock, 1899: opisthosomal flagellum (whip), dorsal (A,B) and ventral (C) aspects. (A,B) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831), proximal segments (A) and segments V to IX (B). (C) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050), complete flagellum. Scale bars = 1 mm (A,B), 2 mm (C).	Figure 13. Hypoctoninae Pocock, 1899: opisthosomal flagellum (whip), dorsal (A,B) and ventral (C) aspects. (A,B) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831), proximal segments (A) and segments V to IX (B). (C) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050), complete flagellum. Scale bars = 1 mm (A,B), 2 mm (C).	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB7334A661292FF5D690B9581.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359954/files/figure.png	https://doi.org/10.5281/zenodo.17359954	Figure 16. Hypoctoninae Pocock, 1899: coxosternal region, ventral aspect. (A) Etienneus africanus (Hentschel, 1899), ♀(AMCC [LP 4654]). (B) Ravilops wetherbeei (Armas, 2002), ♀(AMCC [LP 3342]). (C) Thelyphonellus amazonicus (Butler, 1972), ♀(MPEG [URO] 7). (D) Wounaan vanegasae (Giupponi and Vasconcelos, 2008), comb. n., ♂(MCZ 64455). (E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Markedly sclerotized parts of the median sternum (mesosternum) indicated in black. Scale bars = 1 mm.	Figure 16. Hypoctoninae Pocock, 1899: coxosternal region, ventral aspect. (A) Etienneus africanus (Hentschel, 1899), ♀(AMCC [LP 4654]). (B) Ravilops wetherbeei (Armas, 2002), ♀(AMCC [LP 3342]). (C) Thelyphonellus amazonicus (Butler, 1972), ♀(MPEG [URO] 7). (D) Wounaan vanegasae (Giupponi and Vasconcelos, 2008), comb. n., ♂(MCZ 64455). (E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Markedly sclerotized parts of the median sternum (mesosternum) indicated in black. Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB7334A661292FF5D690B9581.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359956/files/figure.png	https://doi.org/10.5281/zenodo.17359956	Figure 17. Hypoctoninae Pocock, 1899: pedipalp tibia (manus), retrolateral aspect. (A) Thelyphonellus amazonicus (Butler, 1972), ♂(MPEG [URO] 9). (B) Wounaan vanegasae (Giupponi and Vasconcelos, 2008), comb. n., ♂(MCZ 64455). (C) Yekuana venezolensis (Haupt, 2009), comb. n., holotype ♂(ZMB 48289). Abbreviations: bl, basal lobe; dl, distal lobe. Scale bars = 0.5 mm (A), 1 mm (B,C).	Figure 17. Hypoctoninae Pocock, 1899: pedipalp tibia (manus), retrolateral aspect. (A) Thelyphonellus amazonicus (Butler, 1972), ♂(MPEG [URO] 9). (B) Wounaan vanegasae (Giupponi and Vasconcelos, 2008), comb. n., ♂(MCZ 64455). (C) Yekuana venezolensis (Haupt, 2009), comb. n., holotype ♂(ZMB 48289). Abbreviations: bl, basal lobe; dl, distal lobe. Scale bars = 0.5 mm (A), 1 mm (B,C).	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB7334A661292FF5D690B9581.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359960/files/figure.png	https://doi.org/10.5281/zenodo.17359960	Figure 18. Hypoctoninae Pocock, 1899: dissected sternite II (genital), dorsal aspect, illustrating internal surface with spermathecae. (A) Ravilops wetherbeei (Armas, 2002), ♀(AMCC [LP 3342]). (B) Thelyphonellus amazonicus (Butler, 1972), ♀(MPEG [URO] 13). (C) Wounaan vanegasae (Giupponi and Vasconcelos, 2008), comb. n., ♀(IMCN 9931). Scale bars = 0.5 mm.	Figure 18. Hypoctoninae Pocock, 1899: dissected sternite II (genital), dorsal aspect, illustrating internal surface with spermathecae. (A) Ravilops wetherbeei (Armas, 2002), ♀(AMCC [LP 3342]). (B) Thelyphonellus amazonicus (Butler, 1972), ♀(MPEG [URO] 13). (C) Wounaan vanegasae (Giupponi and Vasconcelos, 2008), comb. n., ♀(IMCN 9931). Scale bars = 0.5 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB7334A661292FF5D690B9581.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359962/files/figure.png	https://doi.org/10.5281/zenodo.17359962	Figure 19. Hypoctoninae Pocock, 1899: dissected sternite II (genital), dorsal aspect, illustrating internal surface with spermathecae. (A,B) Thelyphonellus amazonicus (Butler, 1972), ♀(SMF 30076), Serra do Navio, Brazil (A); ♀(MPEG [URO] 13), Jari Celulose S.A., Brazil (B). (C) Thelyphonellus ruschii Weygoldt, 1979, paratype ♀(BMNH, SMF). (D) Wounaan vanegasae (Giupponi and Vasconcelos, 2008), comb. n., ♀(IMCN 9931). (A,C) Redrawn from Weygoldt (1979: 112, abbs. 7,9) [48]. Scale bars = 0.5 mm.	Figure 19. Hypoctoninae Pocock, 1899: dissected sternite II (genital), dorsal aspect, illustrating internal surface with spermathecae. (A,B) Thelyphonellus amazonicus (Butler, 1972), ♀(SMF 30076), Serra do Navio, Brazil (A); ♀(MPEG [URO] 13), Jari Celulose S.A., Brazil (B). (C) Thelyphonellus ruschii Weygoldt, 1979, paratype ♀(BMNH, SMF). (D) Wounaan vanegasae (Giupponi and Vasconcelos, 2008), comb. n., ♀(IMCN 9931). (A,C) Redrawn from Weygoldt (1979: 112, abbs. 7,9) [48]. Scale bars = 0.5 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB73A4A7C1292F9AD6FAD95DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359873/files/figure.png	https://doi.org/10.5281/zenodo.17359873	Figure 1. Single most parsimonious tree (L = 53) obtained by analysis of 45 morphological characteristics for ten species of whip-scorpions (Hypoctoninae Pocock, 1899), via implicit enumeration with equal weights. Morphological synapomorphies (using ACCTRAN optimization) illustrated on each branch, with numbers above indicating characteristics and numbers below indicating character states. Solid (black) circles indicate uniquely derived character states, and empty (white) circles indicate parallel derivations. Measure of support (Goodman–Bremer) for branches indicated at nodes. Tree consistency index (CI): 0.87; tree retention index (RI): 0.85. Refer to Appendix B for character matrix and Appendix C for list of characters.	Figure 1. Single most parsimonious tree (L = 53) obtained by analysis of 45 morphological characteristics for ten species of whip-scorpions (Hypoctoninae Pocock, 1899), via implicit enumeration with equal weights. Morphological synapomorphies (using ACCTRAN optimization) illustrated on each branch, with numbers above indicating characteristics and numbers below indicating character states. Solid (black) circles indicate uniquely derived character states, and empty (white) circles indicate parallel derivations. Measure of support (Goodman–Bremer) for branches indicated at nodes. Tree consistency index (CI): 0.87; tree retention index (RI): 0.85. Refer to Appendix B for character matrix and Appendix C for list of characters.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB73A4A7C1292F9AD6FAD95DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359875/files/figure.png	https://doi.org/10.5281/zenodo.17359875	Figure 2. Map of the Caribbean and northern South America, plotting known locality records of Neotropical whip-scorpions (Hypoctoninae Pocock, 1899), based on material examined and published records. Abbreviations: Bra, Brazil; Col, Colombia; DRe, Dominican Republic; Ecu, Ecuador; FGu, French Guiana; Guy, Guyana; Hai, Haiti; Pan, Panama; Per, Peru; PRi, Puerto Rico; Sur, Suriname; Ven, Venezuela.	Figure 2. Map of the Caribbean and northern South America, plotting known locality records of Neotropical whip-scorpions (Hypoctoninae Pocock, 1899), based on material examined and published records. Abbreviations: Bra, Brazil; Col, Colombia; DRe, Dominican Republic; Ecu, Ecuador; FGu, French Guiana; Guy, Guyana; Hai, Haiti; Pan, Panama; Per, Peru; PRi, Puerto Rico; Sur, Suriname; Ven, Venezuela.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB73A4A7C1292F9AD6FAD95DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359880/files/figure.png	https://doi.org/10.5281/zenodo.17359880	Figure 3. Hypoctoninae Pocock, 1899: habitus, dorsal aspect. (A) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bar = 5 mm.	Figure 3. Hypoctoninae Pocock, 1899: habitus, dorsal aspect. (A) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bar = 5 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB73A4A7C1292F9AD6FAD95DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359883/files/figure.png	https://doi.org/10.5281/zenodo.17359883	Figure 4. Hypoctoninae Pocock, 1899: carapace, dorsal aspect (A,B) and coxosternal region, ventral aspect (C,D). (A,C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm.	Figure 4. Hypoctoninae Pocock, 1899: carapace, dorsal aspect (A,B) and coxosternal region, ventral aspect (C,D). (A,C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB73A4A7C1292F9AD6FAD95DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359895/files/figure.png	https://doi.org/10.5281/zenodo.17359895	Figure 5. Hypoctoninae Pocock, 1899: opisthosoma, dorsal (A,B) and ventral (C,D) aspects. (A,C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm.	Figure 5. Hypoctoninae Pocock, 1899: opisthosoma, dorsal (A,B) and ventral (C,D) aspects. (A,C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB73A4A7C1292F9AD6FAD95DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359903/files/figure.png	https://doi.org/10.5281/zenodo.17359903	Figure 6. Hypoctoninae Pocock, 1899: chelicera, prolateral (A,C) and retrolateral (B,D) aspects. (A,B) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (C,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 0.5 mm.	Figure 6. Hypoctoninae Pocock, 1899: chelicera, prolateral (A,C) and retrolateral (B,D) aspects. (A,B) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (C,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 0.5 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB73A4A7C1292F9AD6FAD95DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359906/files/figure.png	https://doi.org/10.5281/zenodo.17359906	Figure 7. Hypoctoninae Pocock, 1899: pedipalp trochanter (A,B), femur (C,D), and patella (E,F), dorsal aspect. (A,C,E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D,F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Abbreviations: ap, apophysis; ppd, principal prodorsal tubercle; pv, proventral tubercle. Scale bars = 1 mm.	Figure 7. Hypoctoninae Pocock, 1899: pedipalp trochanter (A,B), femur (C,D), and patella (E,F), dorsal aspect. (A,C,E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D,F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Abbreviations: ap, apophysis; ppd, principal prodorsal tubercle; pv, proventral tubercle. Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB73A4A7C1292F9AD6FAD95DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359916/files/figure.png	https://doi.org/10.5281/zenodo.17359916	Figure 8. Hypoctoninae Pocock, 1899: pedipalp tibia (manus), retrolateral (A,B), retrodorsal (C,D), and proventral (E,F) aspects. (A,C,E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D,F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Abbreviations: ap, apophysis; bl, basal lobe; dl, distal lobe; pvd, proventral distal tubercle. Scale bars = 1 mm.	Figure 8. Hypoctoninae Pocock, 1899: pedipalp tibia (manus), retrolateral (A,B), retrodorsal (C,D), and proventral (E,F) aspects. (A,C,E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D,F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Abbreviations: ap, apophysis; bl, basal lobe; dl, distal lobe; pvd, proventral distal tubercle. Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB73A4A7C1292F9AD6FAD95DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359918/files/figure.png	https://doi.org/10.5281/zenodo.17359918	Figure 9. Hypoctoninae Pocock, 1899: pedipalp tibia (manus), retrolateral aspect (A) and opisthosoma, lateral aspect (B), illustrating surface macro- and microsculpture. (A) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 0.25 mm (A), 0.5 mm (B).	Figure 9. Hypoctoninae Pocock, 1899: pedipalp tibia (manus), retrolateral aspect (A) and opisthosoma, lateral aspect (B), illustrating surface macro- and microsculpture. (A) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 0.25 mm (A), 0.5 mm (B).	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB73A4A7C1292F9AD6FAD95DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359929/files/figure.png	https://doi.org/10.5281/zenodo.17359929	Figure 10. Hypoctoninae Pocock, 1899: leg I tarsus (A,B) and closeup (C,D), retroventral (A,C), prolateral (B), and retrodorsal (D) aspects. (A,C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm (A,B), 0.5 mm (C,D).	Figure 10. Hypoctoninae Pocock, 1899: leg I tarsus (A,B) and closeup (C,D), retroventral (A,C), prolateral (B), and retrodorsal (D) aspects. (A,C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm (A,B), 0.5 mm (C,D).	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB73A4A7C1292F9AD6FAD95DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359935/files/figure.png	https://doi.org/10.5281/zenodo.17359935	Figure 11. Hypoctoninae Pocock, 1899: dissected sternite II (genital) illustrating internal surface with sclerites and gonopods, dorsal (A,B), posterodorsal (C,D), and dorsal, slightly posterior (E) aspects. (A,C,E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Abbreviations: Fi, fistula; LaM, lamina medialis; LoD, lobus dorsalis; LoL1, lobus lateralis primus; LoL2, lobus lateralis secundus; Me, mensa. Scale bars = 1 mm.	Figure 11. Hypoctoninae Pocock, 1899: dissected sternite II (genital) illustrating internal surface with sclerites and gonopods, dorsal (A,B), posterodorsal (C,D), and dorsal, slightly posterior (E) aspects. (A,C,E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Abbreviations: Fi, fistula; LaM, lamina medialis; LoD, lobus dorsalis; LoL1, lobus lateralis primus; LoL2, lobus lateralis secundus; Me, mensa. Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB73A4A7C1292F9AD6FAD95DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359942/files/figure.png	https://doi.org/10.5281/zenodo.17359942	Figure 12. Hypoctoninae Pocock, 1899: opisthosoma, pygidium and base of flagellum (whip), dorsal (A,D), lateral (B,E), and ventral (C,F) aspects. (A–C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (D–F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm.	Figure 12. Hypoctoninae Pocock, 1899: opisthosoma, pygidium and base of flagellum (whip), dorsal (A,D), lateral (B,E), and ventral (C,F) aspects. (A–C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (D–F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB73A4A7C1292F9AD6FAD95DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359946/files/figure.png	https://doi.org/10.5281/zenodo.17359946	Figure 13. Hypoctoninae Pocock, 1899: opisthosomal flagellum (whip), dorsal (A,B) and ventral (C) aspects. (A,B) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831), proximal segments (A) and segments V to IX (B). (C) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050), complete flagellum. Scale bars = 1 mm (A,B), 2 mm (C).	Figure 13. Hypoctoninae Pocock, 1899: opisthosomal flagellum (whip), dorsal (A,B) and ventral (C) aspects. (A,B) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831), proximal segments (A) and segments V to IX (B). (C) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050), complete flagellum. Scale bars = 1 mm (A,B), 2 mm (C).	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB73A4A7C1292F9AD6FAD95DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359954/files/figure.png	https://doi.org/10.5281/zenodo.17359954	Figure 16. Hypoctoninae Pocock, 1899: coxosternal region, ventral aspect. (A) Etienneus africanus (Hentschel, 1899), ♀(AMCC [LP 4654]). (B) Ravilops wetherbeei (Armas, 2002), ♀(AMCC [LP 3342]). (C) Thelyphonellus amazonicus (Butler, 1972), ♀(MPEG [URO] 7). (D) Wounaan vanegasae (Giupponi and Vasconcelos, 2008), comb. n., ♂(MCZ 64455). (E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Markedly sclerotized parts of the median sternum (mesosternum) indicated in black. Scale bars = 1 mm.	Figure 16. Hypoctoninae Pocock, 1899: coxosternal region, ventral aspect. (A) Etienneus africanus (Hentschel, 1899), ♀(AMCC [LP 4654]). (B) Ravilops wetherbeei (Armas, 2002), ♀(AMCC [LP 3342]). (C) Thelyphonellus amazonicus (Butler, 1972), ♀(MPEG [URO] 7). (D) Wounaan vanegasae (Giupponi and Vasconcelos, 2008), comb. n., ♂(MCZ 64455). (E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Markedly sclerotized parts of the median sternum (mesosternum) indicated in black. Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB73A4A7C1292F9AD6FAD95DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359956/files/figure.png	https://doi.org/10.5281/zenodo.17359956	Figure 17. Hypoctoninae Pocock, 1899: pedipalp tibia (manus), retrolateral aspect. (A) Thelyphonellus amazonicus (Butler, 1972), ♂(MPEG [URO] 9). (B) Wounaan vanegasae (Giupponi and Vasconcelos, 2008), comb. n., ♂(MCZ 64455). (C) Yekuana venezolensis (Haupt, 2009), comb. n., holotype ♂(ZMB 48289). Abbreviations: bl, basal lobe; dl, distal lobe. Scale bars = 0.5 mm (A), 1 mm (B,C).	Figure 17. Hypoctoninae Pocock, 1899: pedipalp tibia (manus), retrolateral aspect. (A) Thelyphonellus amazonicus (Butler, 1972), ♂(MPEG [URO] 9). (B) Wounaan vanegasae (Giupponi and Vasconcelos, 2008), comb. n., ♂(MCZ 64455). (C) Yekuana venezolensis (Haupt, 2009), comb. n., holotype ♂(ZMB 48289). Abbreviations: bl, basal lobe; dl, distal lobe. Scale bars = 0.5 mm (A), 1 mm (B,C).	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB73A4A7C1292F9AD6FAD95DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359952/files/figure.png	https://doi.org/10.5281/zenodo.17359952	Figure 15. Hypoctoninae Pocock, 1899: lateral (sinistral) ocular tubercle. (A) Thelyphonellus amazonicus (Butler, 1972), ♂(MPEG [URO] 9). (B) Etienneus africanus (Hentschel, 1899), ♀(AMCC [LP 4654]). Abbreviations: Ao, anterior, Mo, median, and Po, posterior (peripheral) ocelli; ado, anterodorsal, and pvo, posteroventral (central) ocelli. Scale bars = 0.25 mm.	Figure 15. Hypoctoninae Pocock, 1899: lateral (sinistral) ocular tubercle. (A) Thelyphonellus amazonicus (Butler, 1972), ♂(MPEG [URO] 9). (B) Etienneus africanus (Hentschel, 1899), ♀(AMCC [LP 4654]). Abbreviations: Ao, anterior, Mo, median, and Po, posterior (peripheral) ocelli; ado, anterodorsal, and pvo, posteroventral (central) ocelli. Scale bars = 0.25 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72C4A711292FB146E6A9124.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359873/files/figure.png	https://doi.org/10.5281/zenodo.17359873	Figure 1. Single most parsimonious tree (L = 53) obtained by analysis of 45 morphological characteristics for ten species of whip-scorpions (Hypoctoninae Pocock, 1899), via implicit enumeration with equal weights. Morphological synapomorphies (using ACCTRAN optimization) illustrated on each branch, with numbers above indicating characteristics and numbers below indicating character states. Solid (black) circles indicate uniquely derived character states, and empty (white) circles indicate parallel derivations. Measure of support (Goodman–Bremer) for branches indicated at nodes. Tree consistency index (CI): 0.87; tree retention index (RI): 0.85. Refer to Appendix B for character matrix and Appendix C for list of characters.	Figure 1. Single most parsimonious tree (L = 53) obtained by analysis of 45 morphological characteristics for ten species of whip-scorpions (Hypoctoninae Pocock, 1899), via implicit enumeration with equal weights. Morphological synapomorphies (using ACCTRAN optimization) illustrated on each branch, with numbers above indicating characteristics and numbers below indicating character states. Solid (black) circles indicate uniquely derived character states, and empty (white) circles indicate parallel derivations. Measure of support (Goodman–Bremer) for branches indicated at nodes. Tree consistency index (CI): 0.87; tree retention index (RI): 0.85. Refer to Appendix B for character matrix and Appendix C for list of characters.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72C4A711292FB146E6A9124.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359875/files/figure.png	https://doi.org/10.5281/zenodo.17359875	Figure 2. Map of the Caribbean and northern South America, plotting known locality records of Neotropical whip-scorpions (Hypoctoninae Pocock, 1899), based on material examined and published records. Abbreviations: Bra, Brazil; Col, Colombia; DRe, Dominican Republic; Ecu, Ecuador; FGu, French Guiana; Guy, Guyana; Hai, Haiti; Pan, Panama; Per, Peru; PRi, Puerto Rico; Sur, Suriname; Ven, Venezuela.	Figure 2. Map of the Caribbean and northern South America, plotting known locality records of Neotropical whip-scorpions (Hypoctoninae Pocock, 1899), based on material examined and published records. Abbreviations: Bra, Brazil; Col, Colombia; DRe, Dominican Republic; Ecu, Ecuador; FGu, French Guiana; Guy, Guyana; Hai, Haiti; Pan, Panama; Per, Peru; PRi, Puerto Rico; Sur, Suriname; Ven, Venezuela.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72C4A711292FB146E6A9124.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359880/files/figure.png	https://doi.org/10.5281/zenodo.17359880	Figure 3. Hypoctoninae Pocock, 1899: habitus, dorsal aspect. (A) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bar = 5 mm.	Figure 3. Hypoctoninae Pocock, 1899: habitus, dorsal aspect. (A) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bar = 5 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72C4A711292FB146E6A9124.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359883/files/figure.png	https://doi.org/10.5281/zenodo.17359883	Figure 4. Hypoctoninae Pocock, 1899: carapace, dorsal aspect (A,B) and coxosternal region, ventral aspect (C,D). (A,C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm.	Figure 4. Hypoctoninae Pocock, 1899: carapace, dorsal aspect (A,B) and coxosternal region, ventral aspect (C,D). (A,C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72C4A711292FB146E6A9124.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359895/files/figure.png	https://doi.org/10.5281/zenodo.17359895	Figure 5. Hypoctoninae Pocock, 1899: opisthosoma, dorsal (A,B) and ventral (C,D) aspects. (A,C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm.	Figure 5. Hypoctoninae Pocock, 1899: opisthosoma, dorsal (A,B) and ventral (C,D) aspects. (A,C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72C4A711292FB146E6A9124.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359903/files/figure.png	https://doi.org/10.5281/zenodo.17359903	Figure 6. Hypoctoninae Pocock, 1899: chelicera, prolateral (A,C) and retrolateral (B,D) aspects. (A,B) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (C,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 0.5 mm.	Figure 6. Hypoctoninae Pocock, 1899: chelicera, prolateral (A,C) and retrolateral (B,D) aspects. (A,B) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (C,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 0.5 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72C4A711292FB146E6A9124.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359906/files/figure.png	https://doi.org/10.5281/zenodo.17359906	Figure 7. Hypoctoninae Pocock, 1899: pedipalp trochanter (A,B), femur (C,D), and patella (E,F), dorsal aspect. (A,C,E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D,F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Abbreviations: ap, apophysis; ppd, principal prodorsal tubercle; pv, proventral tubercle. Scale bars = 1 mm.	Figure 7. Hypoctoninae Pocock, 1899: pedipalp trochanter (A,B), femur (C,D), and patella (E,F), dorsal aspect. (A,C,E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D,F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Abbreviations: ap, apophysis; ppd, principal prodorsal tubercle; pv, proventral tubercle. Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72C4A711292FB146E6A9124.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359916/files/figure.png	https://doi.org/10.5281/zenodo.17359916	Figure 8. Hypoctoninae Pocock, 1899: pedipalp tibia (manus), retrolateral (A,B), retrodorsal (C,D), and proventral (E,F) aspects. (A,C,E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D,F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Abbreviations: ap, apophysis; bl, basal lobe; dl, distal lobe; pvd, proventral distal tubercle. Scale bars = 1 mm.	Figure 8. Hypoctoninae Pocock, 1899: pedipalp tibia (manus), retrolateral (A,B), retrodorsal (C,D), and proventral (E,F) aspects. (A,C,E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D,F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Abbreviations: ap, apophysis; bl, basal lobe; dl, distal lobe; pvd, proventral distal tubercle. Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72C4A711292FB146E6A9124.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359918/files/figure.png	https://doi.org/10.5281/zenodo.17359918	Figure 9. Hypoctoninae Pocock, 1899: pedipalp tibia (manus), retrolateral aspect (A) and opisthosoma, lateral aspect (B), illustrating surface macro- and microsculpture. (A) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 0.25 mm (A), 0.5 mm (B).	Figure 9. Hypoctoninae Pocock, 1899: pedipalp tibia (manus), retrolateral aspect (A) and opisthosoma, lateral aspect (B), illustrating surface macro- and microsculpture. (A) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 0.25 mm (A), 0.5 mm (B).	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72C4A711292FB146E6A9124.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359929/files/figure.png	https://doi.org/10.5281/zenodo.17359929	Figure 10. Hypoctoninae Pocock, 1899: leg I tarsus (A,B) and closeup (C,D), retroventral (A,C), prolateral (B), and retrodorsal (D) aspects. (A,C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm (A,B), 0.5 mm (C,D).	Figure 10. Hypoctoninae Pocock, 1899: leg I tarsus (A,B) and closeup (C,D), retroventral (A,C), prolateral (B), and retrodorsal (D) aspects. (A,C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm (A,B), 0.5 mm (C,D).	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72C4A711292FB146E6A9124.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359935/files/figure.png	https://doi.org/10.5281/zenodo.17359935	Figure 11. Hypoctoninae Pocock, 1899: dissected sternite II (genital) illustrating internal surface with sclerites and gonopods, dorsal (A,B), posterodorsal (C,D), and dorsal, slightly posterior (E) aspects. (A,C,E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Abbreviations: Fi, fistula; LaM, lamina medialis; LoD, lobus dorsalis; LoL1, lobus lateralis primus; LoL2, lobus lateralis secundus; Me, mensa. Scale bars = 1 mm.	Figure 11. Hypoctoninae Pocock, 1899: dissected sternite II (genital) illustrating internal surface with sclerites and gonopods, dorsal (A,B), posterodorsal (C,D), and dorsal, slightly posterior (E) aspects. (A,C,E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Abbreviations: Fi, fistula; LaM, lamina medialis; LoD, lobus dorsalis; LoL1, lobus lateralis primus; LoL2, lobus lateralis secundus; Me, mensa. Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72C4A711292FB146E6A9124.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359942/files/figure.png	https://doi.org/10.5281/zenodo.17359942	Figure 12. Hypoctoninae Pocock, 1899: opisthosoma, pygidium and base of flagellum (whip), dorsal (A,D), lateral (B,E), and ventral (C,F) aspects. (A–C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (D–F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm.	Figure 12. Hypoctoninae Pocock, 1899: opisthosoma, pygidium and base of flagellum (whip), dorsal (A,D), lateral (B,E), and ventral (C,F) aspects. (A–C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (D–F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72C4A711292FB146E6A9124.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359946/files/figure.png	https://doi.org/10.5281/zenodo.17359946	Figure 13. Hypoctoninae Pocock, 1899: opisthosomal flagellum (whip), dorsal (A,B) and ventral (C) aspects. (A,B) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831), proximal segments (A) and segments V to IX (B). (C) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050), complete flagellum. Scale bars = 1 mm (A,B), 2 mm (C).	Figure 13. Hypoctoninae Pocock, 1899: opisthosomal flagellum (whip), dorsal (A,B) and ventral (C) aspects. (A,B) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831), proximal segments (A) and segments V to IX (B). (C) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050), complete flagellum. Scale bars = 1 mm (A,B), 2 mm (C).	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72C4A711292FB146E6A9124.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359948/files/figure.png	https://doi.org/10.5281/zenodo.17359948	Figure 14. Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050), hemispermatophore, ectal aspect, soft tissues removed (A), and ental aspect (B,C), soft tissues not removed. Scale bar = 1 mm.	Figure 14. Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050), hemispermatophore, ectal aspect, soft tissues removed (A), and ental aspect (B,C), soft tissues not removed. Scale bar = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72C4A711292FB146E6A9124.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359954/files/figure.png	https://doi.org/10.5281/zenodo.17359954	Figure 16. Hypoctoninae Pocock, 1899: coxosternal region, ventral aspect. (A) Etienneus africanus (Hentschel, 1899), ♀(AMCC [LP 4654]). (B) Ravilops wetherbeei (Armas, 2002), ♀(AMCC [LP 3342]). (C) Thelyphonellus amazonicus (Butler, 1972), ♀(MPEG [URO] 7). (D) Wounaan vanegasae (Giupponi and Vasconcelos, 2008), comb. n., ♂(MCZ 64455). (E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Markedly sclerotized parts of the median sternum (mesosternum) indicated in black. Scale bars = 1 mm.	Figure 16. Hypoctoninae Pocock, 1899: coxosternal region, ventral aspect. (A) Etienneus africanus (Hentschel, 1899), ♀(AMCC [LP 4654]). (B) Ravilops wetherbeei (Armas, 2002), ♀(AMCC [LP 3342]). (C) Thelyphonellus amazonicus (Butler, 1972), ♀(MPEG [URO] 7). (D) Wounaan vanegasae (Giupponi and Vasconcelos, 2008), comb. n., ♂(MCZ 64455). (E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Markedly sclerotized parts of the median sternum (mesosternum) indicated in black. Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72C4A711292FB146E6A9124.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359956/files/figure.png	https://doi.org/10.5281/zenodo.17359956	Figure 17. Hypoctoninae Pocock, 1899: pedipalp tibia (manus), retrolateral aspect. (A) Thelyphonellus amazonicus (Butler, 1972), ♂(MPEG [URO] 9). (B) Wounaan vanegasae (Giupponi and Vasconcelos, 2008), comb. n., ♂(MCZ 64455). (C) Yekuana venezolensis (Haupt, 2009), comb. n., holotype ♂(ZMB 48289). Abbreviations: bl, basal lobe; dl, distal lobe. Scale bars = 0.5 mm (A), 1 mm (B,C).	Figure 17. Hypoctoninae Pocock, 1899: pedipalp tibia (manus), retrolateral aspect. (A) Thelyphonellus amazonicus (Butler, 1972), ♂(MPEG [URO] 9). (B) Wounaan vanegasae (Giupponi and Vasconcelos, 2008), comb. n., ♂(MCZ 64455). (C) Yekuana venezolensis (Haupt, 2009), comb. n., holotype ♂(ZMB 48289). Abbreviations: bl, basal lobe; dl, distal lobe. Scale bars = 0.5 mm (A), 1 mm (B,C).	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72E4A771292FC2B6F2E966B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359873/files/figure.png	https://doi.org/10.5281/zenodo.17359873	Figure 1. Single most parsimonious tree (L = 53) obtained by analysis of 45 morphological characteristics for ten species of whip-scorpions (Hypoctoninae Pocock, 1899), via implicit enumeration with equal weights. Morphological synapomorphies (using ACCTRAN optimization) illustrated on each branch, with numbers above indicating characteristics and numbers below indicating character states. Solid (black) circles indicate uniquely derived character states, and empty (white) circles indicate parallel derivations. Measure of support (Goodman–Bremer) for branches indicated at nodes. Tree consistency index (CI): 0.87; tree retention index (RI): 0.85. Refer to Appendix B for character matrix and Appendix C for list of characters.	Figure 1. Single most parsimonious tree (L = 53) obtained by analysis of 45 morphological characteristics for ten species of whip-scorpions (Hypoctoninae Pocock, 1899), via implicit enumeration with equal weights. Morphological synapomorphies (using ACCTRAN optimization) illustrated on each branch, with numbers above indicating characteristics and numbers below indicating character states. Solid (black) circles indicate uniquely derived character states, and empty (white) circles indicate parallel derivations. Measure of support (Goodman–Bremer) for branches indicated at nodes. Tree consistency index (CI): 0.87; tree retention index (RI): 0.85. Refer to Appendix B for character matrix and Appendix C for list of characters.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72E4A771292FC2B6F2E966B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359875/files/figure.png	https://doi.org/10.5281/zenodo.17359875	Figure 2. Map of the Caribbean and northern South America, plotting known locality records of Neotropical whip-scorpions (Hypoctoninae Pocock, 1899), based on material examined and published records. Abbreviations: Bra, Brazil; Col, Colombia; DRe, Dominican Republic; Ecu, Ecuador; FGu, French Guiana; Guy, Guyana; Hai, Haiti; Pan, Panama; Per, Peru; PRi, Puerto Rico; Sur, Suriname; Ven, Venezuela.	Figure 2. Map of the Caribbean and northern South America, plotting known locality records of Neotropical whip-scorpions (Hypoctoninae Pocock, 1899), based on material examined and published records. Abbreviations: Bra, Brazil; Col, Colombia; DRe, Dominican Republic; Ecu, Ecuador; FGu, French Guiana; Guy, Guyana; Hai, Haiti; Pan, Panama; Per, Peru; PRi, Puerto Rico; Sur, Suriname; Ven, Venezuela.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72E4A771292FC2B6F2E966B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359880/files/figure.png	https://doi.org/10.5281/zenodo.17359880	Figure 3. Hypoctoninae Pocock, 1899: habitus, dorsal aspect. (A) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bar = 5 mm.	Figure 3. Hypoctoninae Pocock, 1899: habitus, dorsal aspect. (A) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bar = 5 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72E4A771292FC2B6F2E966B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359883/files/figure.png	https://doi.org/10.5281/zenodo.17359883	Figure 4. Hypoctoninae Pocock, 1899: carapace, dorsal aspect (A,B) and coxosternal region, ventral aspect (C,D). (A,C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm.	Figure 4. Hypoctoninae Pocock, 1899: carapace, dorsal aspect (A,B) and coxosternal region, ventral aspect (C,D). (A,C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72E4A771292FC2B6F2E966B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359895/files/figure.png	https://doi.org/10.5281/zenodo.17359895	Figure 5. Hypoctoninae Pocock, 1899: opisthosoma, dorsal (A,B) and ventral (C,D) aspects. (A,C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm.	Figure 5. Hypoctoninae Pocock, 1899: opisthosoma, dorsal (A,B) and ventral (C,D) aspects. (A,C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72E4A771292FC2B6F2E966B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359903/files/figure.png	https://doi.org/10.5281/zenodo.17359903	Figure 6. Hypoctoninae Pocock, 1899: chelicera, prolateral (A,C) and retrolateral (B,D) aspects. (A,B) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (C,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 0.5 mm.	Figure 6. Hypoctoninae Pocock, 1899: chelicera, prolateral (A,C) and retrolateral (B,D) aspects. (A,B) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (C,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 0.5 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72E4A771292FC2B6F2E966B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359906/files/figure.png	https://doi.org/10.5281/zenodo.17359906	Figure 7. Hypoctoninae Pocock, 1899: pedipalp trochanter (A,B), femur (C,D), and patella (E,F), dorsal aspect. (A,C,E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D,F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Abbreviations: ap, apophysis; ppd, principal prodorsal tubercle; pv, proventral tubercle. Scale bars = 1 mm.	Figure 7. Hypoctoninae Pocock, 1899: pedipalp trochanter (A,B), femur (C,D), and patella (E,F), dorsal aspect. (A,C,E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D,F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Abbreviations: ap, apophysis; ppd, principal prodorsal tubercle; pv, proventral tubercle. Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72E4A771292FC2B6F2E966B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359916/files/figure.png	https://doi.org/10.5281/zenodo.17359916	Figure 8. Hypoctoninae Pocock, 1899: pedipalp tibia (manus), retrolateral (A,B), retrodorsal (C,D), and proventral (E,F) aspects. (A,C,E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D,F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Abbreviations: ap, apophysis; bl, basal lobe; dl, distal lobe; pvd, proventral distal tubercle. Scale bars = 1 mm.	Figure 8. Hypoctoninae Pocock, 1899: pedipalp tibia (manus), retrolateral (A,B), retrodorsal (C,D), and proventral (E,F) aspects. (A,C,E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D,F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Abbreviations: ap, apophysis; bl, basal lobe; dl, distal lobe; pvd, proventral distal tubercle. Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72E4A771292FC2B6F2E966B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359918/files/figure.png	https://doi.org/10.5281/zenodo.17359918	Figure 9. Hypoctoninae Pocock, 1899: pedipalp tibia (manus), retrolateral aspect (A) and opisthosoma, lateral aspect (B), illustrating surface macro- and microsculpture. (A) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 0.25 mm (A), 0.5 mm (B).	Figure 9. Hypoctoninae Pocock, 1899: pedipalp tibia (manus), retrolateral aspect (A) and opisthosoma, lateral aspect (B), illustrating surface macro- and microsculpture. (A) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 0.25 mm (A), 0.5 mm (B).	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72E4A771292FC2B6F2E966B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359929/files/figure.png	https://doi.org/10.5281/zenodo.17359929	Figure 10. Hypoctoninae Pocock, 1899: leg I tarsus (A,B) and closeup (C,D), retroventral (A,C), prolateral (B), and retrodorsal (D) aspects. (A,C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm (A,B), 0.5 mm (C,D).	Figure 10. Hypoctoninae Pocock, 1899: leg I tarsus (A,B) and closeup (C,D), retroventral (A,C), prolateral (B), and retrodorsal (D) aspects. (A,C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm (A,B), 0.5 mm (C,D).	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72E4A771292FC2B6F2E966B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359935/files/figure.png	https://doi.org/10.5281/zenodo.17359935	Figure 11. Hypoctoninae Pocock, 1899: dissected sternite II (genital) illustrating internal surface with sclerites and gonopods, dorsal (A,B), posterodorsal (C,D), and dorsal, slightly posterior (E) aspects. (A,C,E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Abbreviations: Fi, fistula; LaM, lamina medialis; LoD, lobus dorsalis; LoL1, lobus lateralis primus; LoL2, lobus lateralis secundus; Me, mensa. Scale bars = 1 mm.	Figure 11. Hypoctoninae Pocock, 1899: dissected sternite II (genital) illustrating internal surface with sclerites and gonopods, dorsal (A,B), posterodorsal (C,D), and dorsal, slightly posterior (E) aspects. (A,C,E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (B,D) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Abbreviations: Fi, fistula; LaM, lamina medialis; LoD, lobus dorsalis; LoL1, lobus lateralis primus; LoL2, lobus lateralis secundus; Me, mensa. Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72E4A771292FC2B6F2E966B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359942/files/figure.png	https://doi.org/10.5281/zenodo.17359942	Figure 12. Hypoctoninae Pocock, 1899: opisthosoma, pygidium and base of flagellum (whip), dorsal (A,D), lateral (B,E), and ventral (C,F) aspects. (A–C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (D–F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm.	Figure 12. Hypoctoninae Pocock, 1899: opisthosoma, pygidium and base of flagellum (whip), dorsal (A,D), lateral (B,E), and ventral (C,F) aspects. (A–C) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (D–F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72E4A771292FC2B6F2E966B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359946/files/figure.png	https://doi.org/10.5281/zenodo.17359946	Figure 13. Hypoctoninae Pocock, 1899: opisthosomal flagellum (whip), dorsal (A,B) and ventral (C) aspects. (A,B) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831), proximal segments (A) and segments V to IX (B). (C) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050), complete flagellum. Scale bars = 1 mm (A,B), 2 mm (C).	Figure 13. Hypoctoninae Pocock, 1899: opisthosomal flagellum (whip), dorsal (A,B) and ventral (C) aspects. (A,B) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831), proximal segments (A) and segments V to IX (B). (C) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050), complete flagellum. Scale bars = 1 mm (A,B), 2 mm (C).	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72E4A771292FC2B6F2E966B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359948/files/figure.png	https://doi.org/10.5281/zenodo.17359948	Figure 14. Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050), hemispermatophore, ectal aspect, soft tissues removed (A), and ental aspect (B,C), soft tissues not removed. Scale bar = 1 mm.	Figure 14. Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050), hemispermatophore, ectal aspect, soft tissues removed (A), and ental aspect (B,C), soft tissues not removed. Scale bar = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72E4A771292FC2B6F2E966B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359954/files/figure.png	https://doi.org/10.5281/zenodo.17359954	Figure 16. Hypoctoninae Pocock, 1899: coxosternal region, ventral aspect. (A) Etienneus africanus (Hentschel, 1899), ♀(AMCC [LP 4654]). (B) Ravilops wetherbeei (Armas, 2002), ♀(AMCC [LP 3342]). (C) Thelyphonellus amazonicus (Butler, 1972), ♀(MPEG [URO] 7). (D) Wounaan vanegasae (Giupponi and Vasconcelos, 2008), comb. n., ♂(MCZ 64455). (E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Markedly sclerotized parts of the median sternum (mesosternum) indicated in black. Scale bars = 1 mm.	Figure 16. Hypoctoninae Pocock, 1899: coxosternal region, ventral aspect. (A) Etienneus africanus (Hentschel, 1899), ♀(AMCC [LP 4654]). (B) Ravilops wetherbeei (Armas, 2002), ♀(AMCC [LP 3342]). (C) Thelyphonellus amazonicus (Butler, 1972), ♀(MPEG [URO] 7). (D) Wounaan vanegasae (Giupponi and Vasconcelos, 2008), comb. n., ♂(MCZ 64455). (E) Wounaan yarigui, gen. et sp. n., holotype ♂(IAvH I 2831). (F) Yekuana wanadi, gen. et sp. n., holotype ♂(AMNH IZC 325050). Markedly sclerotized parts of the median sternum (mesosternum) indicated in black. Scale bars = 1 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
C63C340FB72E4A771292FC2B6F2E966B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17359952/files/figure.png	https://doi.org/10.5281/zenodo.17359952	Figure 15. Hypoctoninae Pocock, 1899: lateral (sinistral) ocular tubercle. (A) Thelyphonellus amazonicus (Butler, 1972), ♂(MPEG [URO] 9). (B) Etienneus africanus (Hentschel, 1899), ♀(AMCC [LP 4654]). Abbreviations: Ao, anterior, Mo, median, and Po, posterior (peripheral) ocelli; ado, anterodorsal, and pvo, posteroventral (central) ocelli. Scale bars = 0.25 mm.	Figure 15. Hypoctoninae Pocock, 1899: lateral (sinistral) ocular tubercle. (A) Thelyphonellus amazonicus (Butler, 1972), ♂(MPEG [URO] 9). (B) Etienneus africanus (Hentschel, 1899), ♀(AMCC [LP 4654]). Abbreviations: Ao, anterior, Mo, median, and Po, posterior (peripheral) ocelli; ado, anterodorsal, and pvo, posteroventral (central) ocelli. Scale bars = 0.25 mm.	2024-09-30	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo		Zenodo	biologists	Botero-Trujillo, Ricardo;Moreno-González, Jairo A.;Prendini, Lorenzo			
