taxonID	type	description	language	source
C563080BFFD03A7DFF21697EF3EBF8F2.taxon	materials_examined	Type species by monotypy, L. erichsonii Rosenhauer	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFD03A7DFF21697EF3EBF8F2.taxon	description	Description (based on Leschen and Beutel 2010). Total length 1.25 – 3.04 mm. Body elongate, dorsally slightly convex and ventrally flattened. Surface glabrous with vestiture of erect, suberect, or decumbent setae; foveae on head (between the antennal insertion and the eye, see Lawrence and Hlavac 1979), pronotum, and ventrites (figs 24 – 28), lacking tubercles or well-developed ridges; punctation consisting of large macropunctures dorsally (figs 22, 23, 31, 33) and smaller micropunctures dorsally and ventrally; surfaces with microsculpture (mostly apparent in dissections using compound microscope, see fig 33). Color variable. Head slightly declined and not srongly prognathous, not abruptly constricted to form a neck; temples, transverse occipital ridge and stridulatory files absent. Antennal insertions more or less exposed in dorsal view; antennal grooves absent. Compound eyes finely facetted; interfacetal setae present; two ocelli present (absent in some species). Mandible bidentate without well-developed mola and prostheca. Corporotentorium without median process. Prothorax quadrate to transverse. Pronotum with lateral carinae complete, weakly to strongly explanate, with or without a distinct bead and not dentate along entire margin. Procoxal cavities open. Protrochantin exposed with cavities strongly transverse and externally open, but narrowly so by acute hypomeral processes. Elytra with 10 punctate rows or striae, plus scutellary striole; suture not deflected at apex; epipleuron present in apical third (ending at level of metaventrite). Mesoventrite with paired procoxal rests (fig 24). Mesocoxae conical. Tibial apices with an apical comb of spines; tarsomeres 1 – 3 lobed below. Abdominal ventrites 1 and 2 connate (fig 28), lateral pores and canals present. Ovipositor short with short styli.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFD03A7DFF21697EF3EBF8F2.taxon	discussion	Comments. Species of Laricobius can be distinguished from other members of Derodontidae by the vestiture of long setae, lobed tarsomeres, open procoxal cavities, presence of a scutellary striole and 10 rows of punctures on the eyltra. A key to the genera is provided in Lawrence and Hlavac (1979). A few characters in Leschen and Beutel (2010) require updating as follows. Ocelli, which are a diagnostic feature of Derodontidae, have recently been recorded as absent in a few species of Laricobius (Montgomery et al. 2011; see also C 10, below in cladistic analysis). Franz (1958 b) referred to the ocelli of Laricobius as “ dorsal lobes ” because they did not have a sense organ visible in serial sections. True ocelli are confirmed by the innervation of the optic nerve with the ocellus which was been demonstrated by Leschen and Beutel (2004) for Derodontus LeConte. The mesoventrite has paired procoxal rests (not absent, as stated in Leschen and Beutel 2010). The ostium of the ejaculatory duct in the male genitalia of all derodontids is large and striate (figs 29, 30), and the distribution of this character in Coleoptera is unknown, though it does not appear in Cucujiformia which I have examined. Cuticular features of adult derodontids and their potential function as chemical defense organs were discussed by Lawrence and Hlavac (1979). Laricobius has a range of cuticular features that include microsculpture (fig 33), micropunctures, macropunctures (figs 21 – 23, 31 – 33), and foveae (or pits; figs 24 – 27, 29). The difference between foveae and macropunctures is one of comparable size and depth of penetration of the integument and while micropunctures have a limited depth, I restrict the term foveae to those deeper invaginations at the four corners of the pronotum (seen more clearly in lateral view), on the ventrites and pleura, and the postantennal region of the head (see Lawrence and Hlavac 1979). Usually, these larger invaginations have white secretions, and in the postantennal fovea there are setae (see fig 6 of Lawrence and Hlavac 1979). The macropunctures of the elytra have been called “ window punctures, ” by Lawrence and Hlavac (1979) and these do not appear to have the annulation or multiple rings as seen in other foveae (compare figs 24 and 33). Franz (1958 b) discussed the arrangement of the macropunctures (= pits) on the head of L. erichsonii and discovered that the number is variable and in remaining species these cephalic macropunctures may be connected by a weak to well-developed furrow, which may also vary within some species. By contrast, the foveature (number and position), though, is consistent among the material I have dissected. Various characters are critical for the identification of Laricobius species, and these are presented in the diagnoses and keys for each species. Coloration of the body and appendages is useful and in the diagnoses and cladistic analysis I have referred to color value (light or pale versus dark) while in the descriptions the color is detailed. Elytral striae of Asian species without an explanate pronotum tend to have several macropunctures posteriorly confluent, especially in striae rows 1 – 4 and / or the punctures are contained within deep grooves. Stria 1 may also have punctures that are posteriorly confluent in species that have an explanate pronotum. Detailed life history studies and descriptions of immature stages exist for two Laricobius species (i. e., Zilahi- Balogh et al. 2006) and like other members of the family (Lawrence & Hlavac 1979; Crowson 1980; Lawrence 1982; Leschen & Beutel 2009), larvae and adults occur together in the same habitat. The foremost work on Laricobius biology and morphology was done by Franz (1958 a, b) for L. erichsonii, and his studies should be consulted for a detailed account of the life history of this species. Predatory behavior of Laricobius is thought to be derived in the family (Leschen 2000) from more primitive associations with fungi where the Holarctic genus Derodontus is found on homobasidiomycte fungi (Lawrence & Hlavac 1979; Leschen 1994, 2002; Dodelin 2004) and the south temperate genus Nothoderodontus Crowson, sister taxon to Laricobius, is associated with sooty molds (Lawrence 1985). Larvae have been described and keyed by several authors (Franz 1958; Fukuda 1963, Lawrence & Hlavac 1979; Lawrence 1991; Lawrence et al. 1999 b; Zilahi-Balogh et al. 2006). The classification and phylogenetic relationships of Derodontidae, and within it, were summarized by Leschen and Beutel (2010). The family was removed from Bostrichiformia (Lawrence & Newton 1995) and is now included in Derodontiformia Lawrence et al. (2010) along with Nosodendridae and Jacobsoniidae. Larval and adult morphology suggests that derodontids may be the sister taxon to Nosodendridae (Beutel 1996; Ge et al. 2007), but this is an open question as phylogenetic results are mixed (Caterino et al. 2002; Beutel & Leschen 2005; Ge et al. 2007; Hunt et al. 2007). Though only four genera, there are three subfamilies in Derodontidae: Peltasticinae LeConte contains the single genus Peltastica Mannerheim, Derodontinae LeConte contains Derodontus, and Laricobiinae Mulsant & Rey contains Laricobius and Nothoderodontus. Relationships among the genera have been proposed by Crowson (1959, 1980), Fukuda (1963), and Lawrence & Hlavac (1979). Based on larval characters, Fukuda (1963) divided the family into the Laricobius and Derodontus groups, the latter consisting of Pelatistica and a clade comprising Derodontus and Nothoderodontus, largely in agreement with Crowson (1955, 1959; but see 1981). Lawrence & Hlavac (1979) placed Peltastica at the base of their two preferred trees, with uncertain relationships among Derodontus, Laricobius, and Nothoderodontus. In the recent cladistic analysis using adult and larval characters, Ge et al. (2007) showed that Peltastica is the sister taxon to the remaining Derodontidae.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFD63A7BFF216FD4F180FAC6.taxon	description	(figs 1, 34)	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFD63A7BFF216FD4F180FAC6.taxon	diagnosis	Diagnosis. Body bicolored. Head dark; ocelli present. Prothorax transverse with well developed lateral carina and sides explanate. Pronotum dark; posterior tooth present. Scutellum dark. Surface of elytra incurvate; elytral punctures not confluent. Epipleura at base dark. Ventrites mostly dark. Femora dark. Tibiae light. Aedeagus without median carina on phallobase; medium lobe acute; apices of parameres without an internal ridge.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFD63A7BFF216FD4F180FAC6.taxon	description	Description. Length 1.25 – 1.50 mm (x = 1.34, n = 5). Body bicolored, with head dark brown to black; pronotum, hypomeron, elytra caramel brown to dark tan, scutellum black, epipleuron at base dark brown to black, prosternum darker than hypomeron, ventrites dark brown, lighter brown on the tip of the abdominal ventrites; antenna tan with AI darker, palpi tan, femora dark brown with apices lighter, tibiae and tarsi light brown to tan. Head with ocelli; u-shaped furrow present; interocular distance about 4 x the width of the eye; macropunctures irregular to ovate, present in central area of frons; micropunctures coarse; setae erect, moderately elongate with an average length as long as 2 / 3 the width of the eye. Antenna with length of A 3 about equal to A 4, ratios 1.7: 1.4: 1.2: 1.3: 1.4: 1.1: 1.2: 1.0: 1.4: 1.4: 1.7; A 11 strongly asymmetrical. Prothorax transverse, dorsoventrally flattened, and widest at middle, sides parallel and straight in apical half and strongly converging posteriorly, and not constricted apically; about 0.80 x as long as wide (pronotal length / greatest pronotal width = 0.75 – 0.88, x = 0.80); depth = 0.17 – 0.36 mm (x = 0.27); pronotum laterally broadly explanate, lateral carina with a distinct sharp edge; anterior angle right and subrounded, not forming a tooth; posterior angle indistinct and broadly rounded and with a short tooth; foveae visible in dorsal view; macropunctures on disc of variable in size, narrowly to well separated or contiguous, micropunctures coarse; setae decumbent to erect, long and greater than width of eye. Elytra about 3.30 x as long as wide (elytral length / greatest elytral width = 3.13 – 3.57, x = 3.30) and 3.17 x as long as pronotum (elytral length / pronotal length = 2.59 – 3.29, x = 3.17); surface incurvate at basal third; macropunctures not coalescing to form grooves (apart from those of striae 1 along the suture at apical third), separated by ½ to 1 puncture diameter; micropunctures fine; microsculpture not visible; setae suberect and about as long as length of the eye. Aedeagus with phallobase transverse, about 1 / 3 the length of the median lobe, median carina absent; median lobe projecting slightly beyond apices of parameres, relatively narrow with its greatest width subequal to width of paramere, apex acute, median groove or carina absent, ostium subapical; parameres slender and acute, short setae present on apices, internal subapical ridge absent.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFD63A7BFF216FD4F180FAC6.taxon	discussion	Comments. Laricobius baoxingensis is the only species of the genus with a subparallel-sided explanate margin (fig 1). The pronotal shape is somewhat similar to L. taiwanensis, but can easily be distinguished from it by the coloration of the head and elytra and presence of ocelli. The holotype in IZAS was not examined. The dissected male for this study was not part of the type series and the illustrations published in Háva (2009 a, Fig. 3; 2009 c, Fig. 4) of the genitalia do not match the specimens here. The pinned specimens of the type series were covered with a residue making some of the cuticular and setal characters difficult to observe. The spiculum gastrale was not observed. Hosts. Plant: Tsuga chinensis (Franch) Pritzel ex Diels Adelgid: Adelges tsugae. (Annand)	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFD63A7BFF216FD4F180FAC6.taxon	biology_ecology	Biology. Habitat (Zilahi-Balogh & Jelínek, 2007).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFD63A7BFF216FD4F180FAC6.taxon	distribution	Distribution. China: Sichuan.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFD63A7BFF216FD4F180FAC6.taxon	materials_examined	Type material examined. PARATYPES: 4, NiBa Gorge Forestry Station, near Qiagi village, Baoxing, Sichuan, China 5 – 8 IV 2002 coll. G. Zilahi-Balogh / T. McAvoy / Host: Adelges, Tsuga chinensis Lat. 30 ° 41 ‘ 44 “ N; 102 ° 41 ‘ 44 “ E Elev. 8899 ft. / 2 (hw in pencil) / Laricobius sp. n. Jelínek & Zilahi-Balogh / Laricobius baoxingensis PT (hw on red label) (1, HNHM, 1, NMPC, 2, NMNH; note that the sequence of label data is different from Zilahi- Balogh & Jelínek 2007). Additional material examined. 1, NiBa Gorge Forestry Station near Qiagi Village, Baoxing, Sichuan, China, 5 – 8 IV 2002, coll. G. Zilahi-Balogh / T. McAvoy, Laricobius sp. n. baoxingensis Jelínek & Zilahi-Balogh, 1 (VTRC); 1, IV 2005, Niba Gou, W Liu (VTRC); CHINA: Sichuan, Nibagou Jiaoqi Baoxing, 6 May 2007, Zhang G K from hemlock (IZAS).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFD73A78FF216F9AF3AEFDA9.taxon	description	(fig 2)	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFD73A78FF216F9AF3AEFDA9.taxon	diagnosis	Diagnosis. Body bicolored. Head dark; ocelli present. Prothorax quadrate without well developed lateral carina, sides not strongly explanate. Pronotum dark; posterior tooth absent. Scutellum dark. Surface of elytra incurvate; elytral punctures posteriorly confluent. Epipleura dark. Ventrites dark. Femora dark. Tibiae dark.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFD73A78FF216F9AF3AEFDA9.taxon	description	Description. Length 3.04 mm. Body bicolored, mostly black, head, prothorax, scutellum, sides and apices of the elytra, and venter black, central disc of elytra red-brown; antenna tan with AI darker and club darker, palpi tan, femora and tibiae black, extreme base of tibiae and tarsi dark brown. Head with ocelli; u-shaped furrow absent; interocular distance about 2.5 x the width of the eye; macropunctures distinctly ovate, somewhat shallowly impressed at middle and more impressed on the frons, scattered and present at the center of the frons; micropunctures coarse; setation short and decumbent, length shorter than 1 / 2 the width of the eye. Antenna with length of A 3 about equal to A 4, ratios 1.7: 1.5: 1.2: 1.3: 1.4: 1.1: 1.2: 1.0: 1.4: 1.3: 2.2; A 11 strongly asymmetrical. Prothorax quadrate and highly convex, widest at middle, sides weakly constricted apically; about 0.81 x as long as wide (pronotal length / greatest pronotal width = 0.81); depth = 0.44 mm; pronotum not explanate, lateral carina without a distinct sharp edge; anterior angle acute, deflected, and forming a large lateroventrally directed tooth (the angle formed between the anterior margin and lateral carina, excluding the tooth, is about 65 °); posterior angle indistinct and broadly rounded and without a short tooth; anterior and posterior foveae visible only in lateral view; macropunctures ovate, deep, well separated to contiguous; micropunctures coarse; setae decumbent short and about half the width of eye. Elytra about 2.95 x as long as wide (elytral length / greatest elytral width = 2.95) and 3.47 x as long as pronotum (elytral length / pronotal length = 3.47); surface incurvate at basal third; macropunctures, especially of stria 1, coalescing posteriorly to form grooves, separated by an average of ½ to 1 puncture diameter; micropunctures fine; microsculpture not visible; setae suberect and shorter than half length of the eye.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFD73A78FF216F9AF3AEFDA9.taxon	discussion	Comments. Laricobius bicolor is the only species with bicolored elytra lacking an explanate pronotum. This species was described based on a single female.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFD73A78FF216F9AF3AEFDA9.taxon	distribution	Distribution. China: Sichuan.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFD73A78FF216F9AF3AEFDA9.taxon	materials_examined	Type material examined. HOLOTYPE (APUC). 1 female (card mounted and dissected), China: W Sichuan, Ya´an Prefecture, Tianquan Co., E Erlang Shan Pass / 2900 m, 22. vi. 1999 29.52.36 N, 102.17.82 E. leg A. Pütz / Sammlung Andreas Pütz Eisenhüttenstadt (yellow label) / Laricobius sp. n. (hw) det. A. Pütz 2006 (hw) / HOLO- TYPE Ƥ Laricobius bicolor sp. n. (hw) Jirí Háva 2007 (red label).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFD43A78FF2168AAF315F82C.taxon	description	(figs 3)	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFD43A78FF2168AAF315F82C.taxon	diagnosis	Diagnosis. Body bicolored. Head dark; ocelli present. Prothorax transverse with well developed lateral carina and sides explanate. Pronotum light; posterior tooth absent. Scutellum dark. Surface of elytra not incurvate; elytral punctures not confluent. Epipleura light. Ventrites dark. Femora light. Tibiae light.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFD43A78FF2168AAF315F82C.taxon	description	Description. Length 2.56 mm. Body bicolored, mostly dark tan above, head, scutellum, and ventrites black, hypomeron and epipleuron tan; antenna and legs tan with AI darker. Dorsal surfaces subglabrous. Head with ocelli; u-shaped furrow present; interocular distance about 4 x the width of the eye; macropunctures ovate or fused, small to lineate and shallowly impressed, absent from central area of frons; micropunctures coarse; setation short, erect to suberect, length about half the width of the eye. Antenna with length of A 3 about equal to A 4, ratios 2.1: 2.1: 1.4: 1.4: 1.4: 1.3: 1.2: 1.0: 1.5: 1.6: 2.3; A 11 not strongly asymmetrical. Prothorax transverse and dorsoventrally flattened, widest at middle, sides gradually narrowing anteriorly and more steeply convergent posteriorly; about 0.72 x as long as wide (pronotal length / greatest pronotal width = 0.72); depth = 0.24 mm; pronotum laterally broadly explanate with well developed lateral carina with a distinct sharp edge; anterior angle about 65 ° and not forming a tooth (may be obscured by setae); posterior angle indistinct and broadly rounded and without a short tooth; foveae visible in dorsal view; macropunctures ovate, deep, and well-separated; micropunctures coarse; setae erect and elongate, longer than half the width of eye. Elytra about 3.30 x as long as wide (elytral length / greatest elytral width = 3.30) and 3.41 x as long as pronotum (elytral length / pronotal length = 3.41); surface not incurvate at basal third; macropunctures not coalescing posteriorly to form grooves, separated by an average of 1 puncture diameter; micropunctures fine; microsculpture not visible; vestiture biseriate, mostly consisting of suberect setae, with scattered erect setae, elongate and about 2 / 3 the length of the eye.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFD43A78FF2168AAF315F82C.taxon	discussion	Comments. Among the species that have an explanate pronotal margin, L. caucasicus is one of three species that has the head darker than the pronotum. It can be distinguished from L. laticollis and L. baoxingensis by the shape of the pronotum. Rost (1893) mentions in the description that L. caucasicus that there is a shallow impression before the middle of the elytra; this feature is lacking in the specimen here examined. Háva (2006) listed the type deposition of Rost’s specimen (s) of L. caucasicus as questionably in the MFNB, and queries for types to Berlin and Dresden where some of the Rost material was eventually deposited, either by direct deposition or by exchange (Horn et al. 1990), did not yield specimens. A single specimen that is provisionally identified as L. caucasicus based on the original description and matching the distribution recorded in Rost (1893) is described here. I decided not to name this specimen as a neotype because the abdomen and hind legs of the female specimen (N. Havill, pers. com.) were removed for DNA work prior to this study. Hosts. Plant: Abies nordmanniana (Steven) Spach.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFD43A78FF2168AAF315F82C.taxon	distribution	Distribution. Georgia: Guria.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFD43A78FF2168AAF315F82C.taxon	materials_examined	Material examined. 1, GEORGIA: Guria Province Bakhmaro 9 June 2007 Coll. M. Kenis ex. Abies nordmanniana, Laricobius caucasicus?, Sample: Havill 07 - 66 (YPM)	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDA3A76FF216A8EF478FA6C.taxon	description	(figs 4, 35)	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDA3A76FF216A8EF478FA6C.taxon	diagnosis	Diagnosis. Body unicolored. Head dark; ocelli present. Prothorax quadrate without well developed lateral carina, sides not strongly explanate. Pronotum dark; posterior tooth absent. Scutellum dark. Surface of elytra incurvate; elytral punctures posteriorly confluent. Epipleura dark. Ventrites dark. Femora dark. Tibiae dark. Aedeagus with median carina on phallobase; medium lobe subacute; apices of parameres without an internal ridge.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDA3A76FF216A8EF478FA6C.taxon	description	Description. Length 2.54. Body unicolored black; antenna with AI black, funicle light tan, club brown; palpi tan, extreme bases of tibiae and tarsi brown. Head with ocelli; u-shaped furrow indistinct; interocular distance about 2.5 x the width of the eye; macropunctures ovate and shallowly impressed but slghtly more impressed on frons; micropunctures coarse; setation very short and suberect, length shorter than 1 / 3 the width of the eye. Antenna with length of A 3 about equal to A 4, ratios 1.9: 1.5: 1.2: 1.3: 1.4: 1.3: 1.3: 1.0: 1.6: 1.6: 2.2; A 11 strongly asymmetrical. Prothorax quadrate and highly convex, widest at middle, sides sinuate; about 0.81 x as long as wide (pronotal length / greatest pronotal width = 0.81); depth = 0.36; pronotum not laterally explanate, lateral carina without a distinct sharp edge; anterior angle acute, deflected, and forming a short laterally directed tooth (the angle formed between the anterior margin and lateral carina, excluding the tooth is about 60 °); posterior angle indistinct and broadly rounded and without a short tooth; anterior foveae visible only in lateral view, posterior fovea visible dorsally; macropunctures ovate, deep, anteriorly contiguous or well-separated; micropunctures coarse; setae short, erect, and about 1 / 3 the width of eye. Elytra about 3.16 x as long as wide (elytral length / greatest elytral width = 3.16) and 3.38 x as long as pronotum (elytral length / pronotal length = 3.38); surface incurvate at basal third; macropunctures, especially of striae 1 – 4, coalescing posteriorly to form grooves, separated by an average of ½ to 1 puncture diameter, but those of striae 1 – 3 fusing at incurvature; micropunctures sparse; microsculpture not visible; setae suberect and short, about 1 / 3 length of the eye. Aedeagus with phallobase subquadrate, about 2 / 3 the length of the median lobe, median carina present; median lobe projecting slightly beyond apices of parameres, apex subacute, median groove present; parameres broad and acute with a distinct subapical tooth, moderately short setae present on apices. Spiculum gastrale apically broad.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDA3A76FF216A8EF478FA6C.taxon	discussion	Comments. Among the species that have the elytral punctures posteriorly coalescing, L. daliensis is the only species with dark tibiae that is the same color as the femora. This species is known from a single specimen. The aedeagus of the type specimen was drawn and observed from its original placement on the card with the specimen, therefore some of the aedeagal characters were not observed (the width of the median lobe, the placement of ostium and presence of internal subapical ridge).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDA3A76FF216A8EF478FA6C.taxon	distribution	Distribution. China: Yunnan.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDA3A76FF216A8EF478FA6C.taxon	materials_examined	Type material examined. HOLOTYPE (APUC). 1 male (card mounted and dissected), China, Yunnan {CH 07 - 03}, Dali Bai Auton. Pref., Diancang Shan W Dali, 25 ° 41´49 ´´ N, 100 ° 06´24 ´´ E, 2970 m, sifted at rock edges and under small shrubs, 28. V. 2007, leg. A. Pütz / Laricobius n. sp. 3 (hw) det. A. Pütz 2007 (hw) / HOLO- TYPE 3 Laricobius daliensis sp. n. Jiří Háva 2009 (red label with thin black border).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDA3A74FF216FF7F160FD19.taxon	description	(figs 5, 22, 24, 26, 31, 36)	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDA3A74FF216FF7F160FD19.taxon	diagnosis	Diagnosis. Body bicolored or unicolored. Head dark; ocelli present. Prothorax transverse with well developed lateral carina and sides explanate. Pronotum dark; posterior tooth usually present. Scutellum dark. Surface of elytra usually not incurvate; elytral punctures not confluent. Epipleura at base dark. Ventrites dark. Femora dark. Tibiae light. Aedeagus without median carina on phallobase; medium lobe acute; apices of parameres without an internal ridge.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDA3A74FF216FF7F160FD19.taxon	description	Description. Length 2.36 – 2.78 mm (x = 2.55, n = 16). Body unicolored black (in southern European forms) or more typically bicolored with head, prothorax, scutellum, epipleura, and ventrites light brown to black, elytra along sides to apex (covering striae 7 or 8 – 10) and suture, and femora light brown to black, or sutural stripes distinct only to basal 1 / 3, with the broad central disc red brown; antenna reddish brown to light tan with AI and club often slightly darker, palpi, tibiae, and tarsi reddish brown to tan (tibiae lighter than femora). Head with ocelli; u-shaped furrow present; interocular distance about 4 x the width of the eye; macropunctures weakly to strongly impressed, ovate or fused and lineate, sometimes present at the center of the frons; micropunctures coarse; setation elongate, mostly erect to suberect, average length about 2 / 3 the width of the eye. Antenna with length of A 3 about equal to A 4, ratios 2.1: 1.6: 1.6: 1.4: 1.3: 1.2: 1.1: 1.0: 1.4: 1.6: 2.1; A 11 not strongly asymmetrical. Prothorax transverse and dorsoventrally flattened, widest at middle, sides weakly sinuate or unevenly convex, with margin converging more sharply posteriorly than anteriorly; about 0.72 x as long as wide (pronotal length / greatest pronotal width = 0.72 – 0.80, x = 0.84); depth = 0.26 – 0.36 mm (x = 0.31); pronotum laterally broadly explanate, lateral carina with a distinct sharp edge; anterior angle about 65 ° and forming a tooth; posterior angle indistinct and broadly rounded and often with a short tooth; foveae visible in dorsal view; macropunctures ovate, deep, usually well-separated, may sometimesbe contiguous; micropunctures coarse; setae erect and elongate, about as long as width of eye. Elytra about 3.30 x as long as wide (elytral length / greatest elytral width = 3.14 – 3.58, x = 3.30) and 3.17 x as long as pronotum (elytral length / pronotal length = 2.96 – 3.29, x = 3.17); surface typically not incurvate (present in some specimen, but usually very weak) at basal third; macropunctures not coalescing posteriorly to form grooves (apart from those of striae 1 along the suture at the apical 1 / 3 in some specimens), separated by an average of 1 / 2 puncture diameter; micropunctures fine; microsculpture present or absent; vestiture mostly consisting of suberect setae with scattered erect setae, elongate and average length about 2 / 3 the length of the eye. Aedeagus with phallobase transverse, about 1 / 2 the length of the median lobe, median carina absent; median lobe projecting slightly beyond apices of parameres, relatively broad with its greatest width larger than the width the parameres, apex acute, median groove or carina absent, ostium subapical; parameres moderately broad and subacute, long and short setae present on apices, internal subapical ridge absent. Spiculum gastrale apically acute and narrowed.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDA3A74FF216FF7F160FD19.taxon	discussion	Comments. Laricobius erichsonii can be distinguished from other members having an explanate pronotal margin by the bicolored elytra, anterior lateral margin forming a tooth, and the tibiae pale, lighter than femora. The last character, in combination with the large elytral punctures that are uniform in size will separate L. erichsonii from the similar looking L. rubidus. Gorham (1878) referred to a species in the Fry collection from France as possibly identical to “ lituratus, Costa ” and a “ new species ” supposedly listed in Chevrolat (1874: 28). Neither the species name “ lituratus ” nor the genus Laricobius was in Chevrolat (1874) cited by Gorham (1878) and the page referred to concerns Orthoptera. The Gorham reference was mistaken by Háva (2007) who incorrectly listed L. lituratus Gorham, 1878: 156 as a synonymn of L. erichsonii; the former name originally described in Salpingus Illiger is now recognized as Lissodema lituratum (A. Costa, 1847: 158; see Pollock and Löbl (2008 )). The type specimens of L. erichsonii were listed questionably in the MNHN by Háva (2009) but were obtained from MFNB. Pic (1895: 88) named a variety from Southern Europe, the subspecies niger, that has black elytra and stronger punctation than the typical bicolored forms elsewhere. Háva (2009) listed the type depository of L. erichsonii niger as MNHN but specimens were not located there. Black specimens were examined from Switzerland and France, but the series are mixed with very dark forms, most with consistent elytral color patterns as seen in typical forms and there was no marked difference in the punctation among these specimens. I have examined no recent specimens of L. erichsonii which was introduced to North America (Nova Scotia, New Brunswick, Newfoundland, Washington and Oregon, see Clark & Brown 1958, 1960, Hatch 1962, Lawrence & Hlavac 1979, and Majka 2007) after first being reported in Maine (Lovell 1915). This observation was followed up by collection queries to North American museums and literature searches (systematic, biocontrol, and faunistics). These data corroborate evidence that L. erichsonii was last observed in North America in 1974 and all vouchers of field recovery one or more years following release in eastern North America were L. rubidus (M. Montgomery, pers. com.). Hosts (Franz 1958 a). Plants: Larix decidua Mill., Pinus cembra, L., Pinus mugo Turra, Pinus sylvestris L. Adelgids: Adelges nusslini (Bomer), Adelges piceae (Ratzeburg) preferred host according to Franz 1958 a), Pineus pineoides (Cholodkovsky), Pineus strobi (Hartig).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDA3A74FF216FF7F160FD19.taxon	biology_ecology	Biology. Natural history (Franz 1958 a, b; Clark & Brown 1958).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDA3A74FF216FF7F160FD19.taxon	distribution	Distribution (Háva 2007). Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, France, Great Britain, Germany, Hungary, Liechtenstein, Italy, The Netherlands, Poland, Romania, Slovakia, Switzerland, Serbia and Montenegro.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDA3A74FF216FF7F160FD19.taxon	materials_examined	Type material examined. LECTOTYPE (MFNB). 1 (unknown sex, point mounted on very short point), 17843 / HIST. - Coll. (Coleoptera) Nr. 17843 Laricobius Erichsonii Rosenh. Tyrol. Rosenhauer Zool. Mus. Berlin (black bordered white label) / Erichsonii Rosenhauer * Tyrol. Rosenh. (elegant hw on black bordered faded white label) / SYNTYPUS Laricobius erichsonii Rosenhauer; 1846 labelled by MNHUB 2009 / LECTOTYPE Laricobius erichsonii Rosenhauer, desig, R. Leschen, 2009. PARALECTOTYPES (MFNB). 3, (unknown sex, point mounted on very short point), HIST. - Coll. (Coleoptera) Nr. 17843 Laricobius Erichsonii Rosenh. Tyrol. Rosenhauer Zool. Mus. Berlin (black bordered white label) / SYNTYPUS Laricobius erichsonii Rosenhauer; 1846 labelled by MNHUB 2009 / PARALECTOTYPE Laricobius erichsonii Rosenhauer, desig, R. Leschen, 2009. Additional material examined. EUROPE: Austria: 2, Zirbitzkogel, Eduard Knirsch (FMNH). Czech Republic: 3, Kostelec n. Boh. 7.53.; ex coll. Pfeffer (NMPC). England: 1, Shrubland Estate (ES) TM 15., 31 / V, C. Johnson (NZAC). France: 3, G. Serullaz, Modane, Laric. erichsonii vr niger, Coll. A. Méquignon (MHNG); 13, Alp. Hte Prov.; Col. d'Allos, 2150 m; 12. IX. 1982 Löbl (MHNG); 1, Gard, Mt. Aigual, 1300 m, Col de Serreyrede, 6. IV. 1980, Löbl (MHNG). Germany: 14 Süd-Tyrol, Reittter (1, CAS; 7, FMNH; 6 NMPC); 2, Hanau (RASC); 2, Limburg, Suffrian (MFNB); 1, Ex. Adelges, Black Forest, 4. 1969 (NZAC). Italy: 1, Torino, Sentriéne, 2150 m. 28. VIII. 67, A. Comellini (MHNG). Sweden: 3, Sweden (NMNH). Swizerland: 2, Grisons, s / Samnaun, (2050 m) 26. VIII. 68, Cl. Besuchet (MHNG); 1, Ti, Gambarogna, 1600.6 / 38, Toumayeff (MHNG); 1, Grisons, Val Poschiavo, 2050 m, s / Cavaione 27. VIII, C. Besuchet 83 (MHNG); Zuoz, Eng. Gr, Coll. 9 / 74 1640, Toumayeff (MHNG); 5, Enga din; Helena Pavlov; кол. РЭО (Елены Павл.) (RASC). NORTH AMERICA (all from lab colonies): Canada: Fredricton, New Brunswick, 1954 (CNC); 3, Vancouver, 1964 (CNC). United States: Bent Creek Lab, NC; 5 - 17 - 60, G. D. Amman (YPM).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFD83A72FF21695AF47FFEEA.taxon	description	(figs 7, 37)	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFD83A72FF21695AF47FFEEA.taxon	diagnosis	Diagnosis. Body bicolored. Head dark; ocelli present. Prothorax transverse without well developed lateral carina, sides not strongly explanate. Pronotum light; posterior tooth absent. Scutellum dark. Surface of elytra incurvate; elytral punctures posteriorly confluent. Epipleura dark. Ventrites dark. Femora light. Tibiae light. Aedeagus with medium lobe subacute.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFD83A72FF21695AF47FFEEA.taxon	description	Description. Length 2.64 mm. Body bicolored, mostly dark tan dorsally and the hypomera, head, venter, scutellum, and epipleura chocolate brown; antenna with AI dark tan, funicle light tan, club dark tan; palpi light tan, femora, tibiae and tarsi tan, with the metafemora brown to apical 1 / 3. Head with ocelli; u-shaped furrow absent; interocular distance about 2 x the width of the eye; macropunctures ovate and deeply impressed and present at the center of the frons; micropunctures coarse; setation short, length shorter than 1 / 3 the width of the eye. Antenna with length of A 3 about equal to A 4, ratios 2.0: 1.7: 1.3: 1.4: 1.5: 1.3: 1.2: 1.0: 1.6: 1.5: 2.2; A 11 strongly asymmetrical. Prothorax slightly transverse and moderately convex, widest at middle, sides sinuate, but not constricted greatly; about 0.79 x as long as wide (pronotal length / greatest pronotal width = 0.79); depth = 0.32; pronotum not laterally explanate, lateral carina without a distinct sharp edge; anterior angle acute, deflected, and forming a short laterally directed tooth (the angle formed between the anterior margin and lateral carina, excluding the tooth is about 90 °); posterior angle indistinct and broadly rounded and without a short tooth; anterior foveae visible only in lateral view, posterior foveae visible in dorsal view; macropunctures ovate, deep, and well-separated, micropunctures present but sparse; setae short, and about 1 / 2 the width of eye. Elytra about 2.76 x as long as wide (elytral length / greatest elytral width = 2.76) and 3.40 x as long as pronotum (elytral length / pronotal length = 3.40); surface incurvate at basal third; macropunctures, especially of striae 1 – 4, coalescing posteriorly to form grooves, separated by an average of ½ of a puncture diameter; micropunctures fine; microsculpture not visible; setae suberect and short, about 1 / 3 length of the eye. Aedeagus with phallobase slightly transverse and rectangular, about 2 / 3 the length of the median lobe; median lobe projecting significantly beyond apices of parameres, relatively narrow, apex subacute; parameres broad and apically blunt, apices curving towards midline and ending in a sharp point, asetose. Spiculum gastrale apically broad.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFD83A72FF21695AF47FFEEA.taxon	discussion	Comments. Laricobius incognatus can be distinguished from other species having posteriorly confluent elytral punctures by its bicolored body and uniformly colored elytra; the relatively more transverse prothorax and black scutellum will also separate this species from the similar-looking L. mirabilis. This species was described from two male specimens from the Shennongjia Mountains and I examined the holotype which was covered with a residue making the cuticular and setal characters difficult to assess (the setae were stuck flat against the cuticle). The genitalia was not cleared and was drawn from the card mount, so some of the characters were not examined (the width of the median lobe, presence of median carina, and the placement of ostium and presence of internal subapical ridge).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFD83A72FF21695AF47FFEEA.taxon	distribution	Distribution. China: Hubei.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFD83A72FF21695AF47FFEEA.taxon	materials_examined	Type material examined. HOLOTYPE (JHPC). 1, male (card mounted and dissected, right metatarsus missing), China, W Hubei, 10. – 14. VI. DASHENNONGJIA mts. 31.5 N 110.3 E, 2100 – 2900 m Jaroslav Turna leg., 2002 / HOLOTYPE Laricobius incognatus sp. n. Jiři Háva 2009 (name hw, red label).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDE3A72FF216B76F0C0F99B.taxon	description	(figs 8, 38)	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDE3A72FF216B76F0C0F99B.taxon	diagnosis	Diagnosis. Body unicolored. Head dark; ocelli present. Prothorax quadrate without well developed lateral carina, sides not strongly explanate. Pronotum dark; posterior tooth absent. Scutellum dark. Surface of elytra incurvate; elytral punctures posteriorly confluent. Epipleura dark. Ventrites dark. Femora light. Tibiae light. Aedeagus with medium lobe acute.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDE3A72FF216B76F0C0F99B.taxon	description	Description. Length 2.40 mm. Body unicolored chocolate brown; antenna with AI dark tan, funicle light tan, club dark tan; palpi light tan; legs unicolored tan. Head with ocelli; u-shaped furrow absent; interocular distance about 2 x the width of the eye; macropunctures ovate and deeply impressed, scattered and present at center of frons; micropunctures fine; setation short, decumbent, length about 1 / 3 the width of the eye. Antenna with length of A 3 distinctly shorter than A 4, ratios 2.0: 2.0: 1.0: 1.4: 1.6: 1.2: 1.6: 1.2: 1.8: 2.0: 2.4; A 11 strongly asymmetrical. Prothorax quadrate and convex, widest at middle, sides sinuate; about 0.8 x as long as wide (pronotal length / greatest pronotal width = 0.8); depth = 0.5 mm; pronotum not laterally explanate, lateral carina poorly developed; anterior angle acute and deflected, forming a short tooth (the angle formed between the anterior margin and lateral carina, excluding the tooth, is about 90 °); posterior angle indistinct and broadly rounded and without a short tooth; anterior foveae visible only in lateral view, posterior foveae visible in dorsal view; macropunctures ovate, deep, and wellseparated, micropunctures fine; setae short, decumbent and about 1 / 3 to 1 / 2 the width of eye. Elytra about 1.43 x as long as wide (elytral length / greatest elytral width = 1.43) and 2.8 x as long as pronotum (elytral length / pronotal length = 2.80); surface incurvate at basal third; macropunctures, especially of striae 1 – 4, coalescing posteriorly to form grooves, separated by an average of ½ of a puncture diameter; micropunctures fine; microsculpture not visible; setae suberect, short, about 1 / 3 length of the eye. Aedeagus with phallobase slightly transverse and rectangular, about 2 / 3 the length of the median lobe; median lobe projecting significantly beyond apices of parameres, apex acute and triangulate with a dorsal carina; parameres broad and apically blunt, and anvil-shaped with laterally directed tooth, subapical setae present and moderately elongate. Spiculum gastrale apically broad.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDE3A72FF216B76F0C0F99B.taxon	discussion	Comments. Laricobius jizu can be distinguished from the similar-looking species L. daliensis and L. loebli by the unicolored, tan legs. This species is known from a single male specimen. The genitalia were drawn from the carded specimen and the width of the median lobe, presence of median carina, the placement of ostium and presence of internal subapical ridge were not observed. Hosts. Plant: Tsuga.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDE3A72FF216B76F0C0F99B.taxon	distribution	Distribution. CHINA: Sichuan, Yunnan.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDE3A72FF216B76F0C0F99B.taxon	materials_examined	Type material examined. HOLOTYPE (NHMB): YUNNAN, 30 May – 3 Jun JIZU MTS., 1983 25.58 N 100.21 E, Bolm lgt., 2800 / HOLOTYPE 3 Laricobius jizu sp. n. J. Háva det. 2009 (red label with narrow black bor- der). Additional material examined. CHINA Sichuan, Goudingshan Danba, 19 Oct 2006, Lu, W H, from hemlock (IZAS).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDE3A73FF216CD9F200F961.taxon	description	(figs 9, 39)	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDE3A73FF216CD9F200F961.taxon	diagnosis	Diagnosis. Body bicolored. Head dark; ocelli absent. Prothorax transverse with well developed lateral carina and sides explanate. Pronotum dark; posterior tooth present. Scutellum dark. Surface of elytra incurvate; elytral punctures not confluent. Epipleura at base dark. Ventrites dark. Femora dark. Tibiae light. Aedeagus without median carina on phallobase; medium lobe subacute; apices of parameres without an internal ridge.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDE3A73FF216CD9F200F961.taxon	description	Description. Length 2.36 – 2.80 mm (x = 2.59, n = 8). Body bicolored with head, prothorax, scutellum, elytra along sides to level of abdominal ventrite 2 (covering striae 9 – 10, sometimes striae 8) and suture along scutellary striole (sometimes to apex or absent), epipleuron, and venter chocolate brown; elytra reddish brown to dark tan; antenna reddish brown to dark brown with AI darker (not the club), palpi, femora, and tibiae dark brown, tarsi lighter. Head without ocelli; u-shaped furrow present, but very shallow and broad; interocular distance about 4 x the width of the eye; macropunctures rounded distinct to indistinct and not rounded, fusing sometimes to form irregular trenches; micropunctures fine to coarse; setae erect, moderately elongate with an average length as long as 2 / 3 the width of the eye. Antenna with length of A 3 about equal to A 4, ratios 1.7: 1.4: 1.3: 1.2: 1.2: 1.0: 1.1: 1.0: 1.2: 1.2: 1.7; A 11 not strongly asymmetrical. Prothorax slightly transverse and rectangular, dorsoventrally flattened, and widest at basal third, sides converging anteriorly from widest point and weakly convex posteriorly; about 0.81 x as long as wide (pronotal length / greatest pronotal width = 0.76 – 0.87, x = 0.81); depth = 0.24 – 0.40 mm (x = 0.33); pronotum laterally broadly explanate, lateral carina with a distinct sharp edge; anterior angle 90 º to about 80 ° and subacute, not forming a tooth; posterior angle distinct and marked by a short tooth; foveae visible in dorsal view; macropunctures on disc indistinct, shallowly to well-impressed and well-separated; micropunctures coarse; setae erect, long and 2 / 3 or equal to width of eye. Elytra about 3.32 x as long as wide (elytral length / greatest elytral width = 2.97 – 3.79, x = 3.32) and 3.15 x as long as pronotum (elytral length / pronotal length = 3.06 – 3.25, x = 3.15); surface weakly incurvate at basal third; macropunctures not coalescing to form grooves, separated by 1 / 2 puncture diameter; micropunctures fine; microsculpture not visible; setae suberect and long, about 2 / 3 length of the eye. Aedeagus with phallobase quadrate, about 1 / 2 the length of the median lobe, median carina absent; median lobe projecting slightly beyond apices of parameres, narrow with its greatest width larger than the width of the parameres, apex subacute, median groove or carina absent, ostium subapical; parameres slender and acute, very short setae present on apices (visible under high magnification), internal subapical ridge absent. Spiculum gastrale apically narrow.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDE3A73FF216CD9F200F961.taxon	discussion	Comments. Laricobius kangdingensis can be distinguished from most species by the lack of ocelli. It can be reliably distinguished from the other ocelli-lacking species by its genitalia and distribution. The holotype (IZAS) was not examined, but several paratypes were available for study. The sequence of information on the label data of the type specimens I examined were not as they appear in Zilahi-Balogh & Jelínek (2007). Hosts. Plant: Tsuga chinensis, Tsuga dumosa (D. Don) Eichler. Adelgid: Adelges tsugae.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDE3A73FF216CD9F200F961.taxon	biology_ecology	Biology. Habitat and rearing under quarantine (Zilahi-Balogh & Jelínek, 2007).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDE3A73FF216CD9F200F961.taxon	distribution	Distribution. China: Sichuan.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDE3A73FF216CD9F200F961.taxon	materials_examined	Type material examined. PARATYPES. 12 IV 2002 Luoxiba Deng, Kangding Co. Sichuan Prov. China, coll. T. McAvoy / G. Zilahi-Balogh / Host: Adelges infested Tsuga chinenis [sic] Lat / long: 30 º 03 ‘ 00 “ N / 102 º 02 ‘ 00 “ elev: 2840 m / Paratype 5 Ƥ (5 Ƥ hw) / / Laricobius kangdingensis PT (hw on red label) (1, NMPC, 2, NMNH). Additional material examined. 4, 12 IV 2002, Luoxiba Deng, Kangding Co. Sichuan Prov, China; coll: T. McAvoy / G. Zilahi-Balogh, Host: Adelges infested Tsuga chinensis, Lat / long: 30 º 03 ' 00 '' N / 102 ° 02 ' 00 " E, elev: 2840 m, Laricobius sp. n. kangdingensis Zilahi-Balogh & Jelínek (VTRC); 3, Houditong, Ningshan Co., Shaanxi, 17 Oct. 2002, G Yu (YPM); 3, Sichuan Province, Danba County, Donggu town, Maoniugou, 1 December 2007, Zhang Dekui, Havill Sample # 08 - 267 (YPM); 2, same but Havill Sample # 08 - 269 (YPM); 5, same but 30.61 N 101.76 E, 2710 m, Havill sample 08 - (YPM); 2, Sichuan Province, Lixian, Duergou; downtown Dagou, 31.58 N 102.95 E, 2670 m, 21 August 2008, Coll.: Feng Bo, Havill sample 08 - 268 (YPM); 1, Sichuan, Kuiyonggou Danba, 15 Dec 2006, Lu W H, from hemlock (IZAS); 1, same but 10 Feb 2007 (IZAS); 1, Sichuan, Dagou Lixian, 10 May 2006, Xiao N P, from hemlock (IZAS); 1, same but 22 Sep 2006, Zhou J H, from hemlock (IZAS). Lab colony: 2, USA Virginia Blacksburg, VA Tech, From lab colony, 2004, Coll.: H. Gatton (VTRC); 6, CHINA, Sichuan Province, Laboratory reared, 11 October 2007, J. Zhou, Havill Sample # 08 - 266 (YPM); 1, CHINA, from Lab, Liu Y, 18 Apr 2007, from hemlock (IZAS).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDF3A70FF216CF2F51EFA49.taxon	description	(figs 10, 40)	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDF3A70FF216CF2F51EFA49.taxon	diagnosis	Diagnosis. Body bicolored. Head dark; ocelli present. Prothorax transverse with well developed lateral carina and sides explanate. Pronotum dark; posterior tooth absent. Scutellum dark. Surface of elytra not incurvate; elytral punctures not confluent. Epipleura at base dark. Ventrites dark. Femora light. Tibiae light. Aedeagus without median carina on phallobase; medium lobe subacute; apices of parameres without an internal ridge.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDF3A70FF216CF2F51EFA49.taxon	description	Description. Length 2.80 mm. Body bicolored with head, prothorax, scutellum, elytra along sides to level above abdominal ventrite 2 (covering stria 10), epipleuron, and venter chocolate brown; elytra reddish brown to dark tan; antenna reddish brown to dark brown with AI and club darker, legs reddish brown to dark tan. Head with ocelli; u-shaped furrow present; interocular distance about 4 x the width of the eye; macropunctures rounded, distinct to indistinct and not rounded, fusing sometimes to form irregular trenches and present at the center of the frons; micropunctures coarse; setae erect, moderately elongate with an average length as long as 2 / 3 the width of the eye. Antenna with length of A 3 about equal to A 4, ratios 1.9: 1.6: 1.4: 1.3: 1.3: 1.2: 1.1: 1.0: 1.5: 1.6: 2.4; A 11 not strongly asymmetrical. Prothorax transverse and rectangular, dorsoventrally flattened, and widest at middle, sides evenly rounded; about 0.73 x as long as wide (pronotal length / greatest pronotal width = 0.73); depth = 0.42; pronotum laterally broadly explanate, lateral carina with a distinct sharp edge; anterior angle variable (90 º to about 60 ° and rounded), not forming a tooth; posterior angle indistinct, short tooth absent at the posterior of the lateral margin; foveae visible dorsally; macropunctures on disc distinct, well-impressed and well-separated; micropunctures coarse; setae erect, long with average length 2 / 3 or equal to width of eye. Elytra about 3.48 x as long as wide (elytral length / greatest elytral width = 3.48) and 3.37 x as long as pronotum (elytral length / pronotal length = 3.37); surface not incurvate at basal third; macropunctures not coalescing to form grooves, separated by an average of 1 / 2 puncture diameter; micropunctures fine; microsculpture visible; setae suberect to erect, moderately long averaging about 2 / 3 the length of the eye. Aedeagus with phallobase quadrate, about 1 / 2 the length of the median lobe, median carina absent; median lobe not projecting beyond apices of parameres, narrow with its greatest width less than the width of the parameres, apex subacute and converging apically at about apical 1 / 3, median groove or carina absent, ostium subapical; parameres broad and subacute, short setae present on apices and along inner margin, internal subapical ridge absent. Spiculum gastrale apically narrow.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDF3A70FF216CF2F51EFA49.taxon	discussion	Comments. Among the ocelli-bearing species with an explanate pronotal margin, L. kovalevi can be distinguished by the black head and pronotum and lighter elytra. The posterior pronotal tooth is absent in L. kovalevi, which is present in L. rubidus, while the femora and tibiae are light which can distinguish L. kovalevi from L. erichsonii. I had two paratypes in relatively poor condition from ZMUM for study and only the male genitalia were dissected from one of these. A single measured specimen fell outside of the published size range of the species (2.1 – 2.5 mm). At the time this manuscript was in review, additional specimens of this species from Japan were sent sent to N. Havill (USDA) and myself by S. Shiyake (OMNH), and based on dissection, this species is confirmed as a L. kovalevi. Hosts. Plant: Pinus pumila (Pall.) Regel.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDF3A70FF216CF2F51EFA49.taxon	distribution	Distribution. Russia: Far East. Japan: Honshū.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDF3A70FF216CF2F51EFA49.taxon	materials_examined	Type material examined. PARATYPES (ZMUM): 2, ИАГАД. обп. ЛГ – 19 Стоковое, 11 KM ЮВ КУЛУ, 14 0 6 1976, Л. Глумкова, Paratypus Laricobius kovalevi Nikitsky (red label with name hw). Additional material examined. 5, Tekari-dake, Senzu, Honkawane-cho, Haibara-gun, Shizuoka Pref., Japan, 24. September, 2001, Y. Tahira leg., altitude 2,540 m, on Pinus pumila (2, OMNH; 2, NZAC; 1, YPM); 1, Mt. Hakkoda, Aomori Pref., Japan, 4 October 2007, S. Shiyake leg., altitude 1,520 m, on Pinus pumila (1, YPM).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDC3A71FF216C0AF5B6F916.taxon	description	(figs 11, 41)	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDC3A71FF216C0AF5B6F916.taxon	diagnosis	Diagnosis. Body bicolored. Head dark; ocelli present. Prothorax transverse with well developed lateral carina and sides explanate. Pronotum light; posterior tooth present or absent. Scutellum usually dark. Surface of elytra weakly incurvate; elytral punctures not confluent. Epipleura dark. Ventrites dark. Femora light. Tibiae light. Aedeagus without median carina on phallobase; medium lobe apiculate; apices of parameres without an internal ridge.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDC3A71FF216C0AF5B6F916.taxon	description	Description. Length 2.30 – 2.76 mm (x = 2.52, n = 15). Body bicolored with head, scutellum (usually), epipleuron, and venter chocolate brown; pronotum, elytra, and hypomeron light brown or tan; antenna light brown or tan with AI usually darker, legs reddish light brown or tan, bases and apices of the femora and tibiae may be darker. Head with ocelli; u-shaped furrow present; interocular distance about 4 x the width of the eye; macropunctures rounded and distinct, sometimes fusing and sometimes present at the center of the frons; micropunctures coarse; setae erect, moderately elongate with an average length as long as 2 / 3 the width of the eye. Antenna with length of A 3 about equal to A 4, ratios 2.1: 1.6: 1.3: 1.4: 1.4: 1.1: 1.1: 1.0: 1.6: 1.4: 2.1; A 11 not strongly asymmetrical. Prothorax transverse and rectangular, dorsoventrally flattened, and widest at middle, sides converging anteriorly from widest point and weakly convex posteriorly; about 0.68 x as long as wide (pronotal length / greatest pronotal width = 0.63 – 0.71, x = 0.68); depth = 0.26 – 0.40 mm (x = 0.32); pronotum laterally explanate, lateral carina with a distinct sharp edge; anterior angle about 60 ° and subacute, not forming a tooth; foveae visible in dorsal view; posterior angle indistinct or distinct with short tooth present or absent; macropunctures on disc distinct, well-impressed and wellseparated or contiguous at the anterior middle of disc, micropunctures coarse; setae suberect, short and average length about 1 / 2 width of eye. Elytra about 3.22 x as long as wide (elytral length / greatest elytral width = 3.00 – 3.47, x = 3.22) and 3.35 x as long as pronotum (elytral length / pronotal length = 3.20 – 3.54, x = 3.35); surface weakly incurvate at basal third; macropunctures not coalescing to form grooves (apart from those of striae 1 along the suture near apex in some specimens), separated by an average of 1 / 2 puncture diameter; micropunctures fine; microsculpture visible on some specimens; setae suberect, short and an average of about 1 / 2 the length of the eye. Aedeagus with phallobase transverse, about 1 / 2 the length of the median lobe, median carina absent; median lobe projecting slightly beyond apices of parameres, moderately broad with its greatest width subequal to parameres, sides gradually narrowing posteriorly with the apex apiculate, median groove or carina absent; ostium subapical; parameres broad and rounded apically, short setae present on apices and along inner margin, internal subapical ridge absent. Spiculum gastrale apically narrowed.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDC3A71FF216C0AF5B6F916.taxon	discussion	Comments. Among the species having an explanate pronotal margin, L. laticollis is one of three species with the head dark and pronotum light. It can be distinguished from L. caucasicus by having an incurvate elytral surface and from L. baoxingensis by the shape of the pronotum. Fall (1916) had 14 specimens, taken at the University of Washington Campus February and April, but I have only examined specimens collected in February 1911. The single specimen labelled as “ type ” is here designated as the lectotype, this does not have a black scutellum; the remaining syntypes are mixed. Hosts. Plants: Pseudotsuga menzeseii (Mirb.) Franco, Tsuga heterophylla (Raf.) Sarg. Adelgid: Adelges cooleyi (Gillette) (M. Montgomery, pers. com.).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDC3A71FF216C0AF5B6F916.taxon	distribution	Distribution. United States: California, Idaho, Oregon, Washington. Canada: Alberta, British Columbia.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDC3A71FF216C0AF5B6F916.taxon	materials_examined	Type material examined. 7 syntypes (all pointed MCZ): 1, Seattle, II- 12 - 11 (hw) / H. C. FALL COLLEC- TION / W. T. / TYPE laticollis. (name hw) / M. C. Z. Type 24993 (red label, number hw) / Aug – Dec 2005 MCZ Image Database (bordered label with camera icon) / LECTOTYPE, designated by R. Leschen 2009; 1, W. T. / Univ. of W. Campus. Seattle. IV – 6 – 12 (hw) / H. C. FALL COLLECTION / Laricobius laticollis Fall. (hw); 2, Seattle Wash. (hw) / H. C. FALL COLLECTION; 1, W. T. / Seattle, Campus Univ. of W., II- 12 - 11 (hw) / H. C. FALL COLLEC- TION; 1, W. T. / Seattle, II- 12 - 11 (hw) / H. C. FALL COLLECTION; 1, “ Campus ” (hw), Seattle, Wash (printed) II- 12 - 11 T. C. K. (hw) / H. C. FALL COLLECTION. Additional material examined. CANADA. Alberta: 1, Truner V., Alta. VI. 1.1933, Owen Bryant (CAS); 20, Ghost Dam, Alberta, several dates, BF and JL Carr (CNC); 2, Calgary, 1973, BF and JL Carr (CNC); 1, Coleman, Alberta (CNC). British Columbia: 2, Salmon Arm, 3. V. 35, Hugh Leech, 10351; 1, same but 30. IV. 1930 (CAS); 1, same but 30. IV. 1933 on Douglas fir boughs, 23. III. 34, beating Douglas fir branches, E. S. Ross Collection (CAS); 1, Sanca, 22. IV. 1933 G. Stace Smith, from swift stream (1, CAS; 4, CNC). UNITED STATES. California: 8, Fieldbrook, Humboldt Co., Mar. 27, ’ 38, B. P. Bliven No. 801 (CAS); Humbolt Co., v. 3.17, F. W. Nunenmacher (FMNH). Idaho: 1, Moscow, v- 28 1950, N. M. Downie (FMNH); 1, same but v- 20, 1950 (FMNH); 1, Troy, v- 14, 1950, N. M. Downie (FMNH); 1, Deary, v- 14, 1950, N. M. Downie (FMNH); 1, Moscow Mts, II. 10.1934, I. W. Bales (FMNH). Oregon: 7, Milton, Ore, G. H. Nelson, IV- 3 - 49, G. W. Green collection (CAS). Washington: 1, Fort Lewis, Pierce Co. V- 3.1946, P. H. arnaudf (CAS); 2, Seattle, VII- 13 on Douglas Fir, Van Dyke Collection (CAS); 1, Seattle, Wash. III- 7 - 13, sweeping Douglas fir (CAS); 3, Grand Coulee, Washington, BF and JL Carr (CNC); 11, King Co., Seattle; 17 - 24 February 2007, D. McDonald, Havill Samples # 07 - 01, 07 - 01 A, 07 - 01 B, 07 - 01.1 (YPM).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDD3A6EFF216D2AF2A2FA81.taxon	description	(figs 12, 42, 50)	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDD3A6EFF216D2AF2A2FA81.taxon	diagnosis	Diagnosis. Body unicolored. Head dark; ocelli present. Prothorax quadrate without well developed lateral carina, sides not strongly explanate. Pronotum dark; posterior tooth absent. Scutellum dark. Surface of elytra not incurvate; elytral punctures posteriorly confluent. Epipleura dark. Ventrites dark. Meso- and metafemora dark (profemora light). Tibiae light. Aedeagus with median carina on phallobase; medium lobe acute; apices of parameres without an internal ridge.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDD3A6EFF216D2AF2A2FA81.taxon	description	Description. Length 2.36 – 2.86 mm (x = 2.56, n = 3). Body unicolored chocolate brown; antenna with AI dark tan, funicle light tan, club dark tan; palpi light tan; legs bicolored with prolegs light brown or tan, with apex of tibiae darker; midleg light to dark brown with femora darker at apical 1 / 3 or 2 / 3 darker, tibiae slightly darker and tarsi lighter; hindleg with femora brown, tibiae slightly lighter and tarsi light brown or tan. Head with ocelli; u-shaped furrow absent; interocular distance about 2 x the width of the eye; punctures ovate and deeply impressed, present at the center of the frons; micropunctures moderately coarse; setation short, decumbent, length about 1 / 3 the width of the eye. Antenna with length of A 3 shorter than A 4, ratios 1.7: 1.4: 1.0: 1.4: 1.4: 1.2: 1.2: 1.0: 1.3: 1.3: 1.8; A 11 strongly asymmetrical. Prothorax quadrate and convex, widest at middle, sides sinuate; about 0.81 x as long as wide (pronotal length / greatest pronotal width = 0.78 – 0.84, x = 0.81); depth = 0.30 – 0.40 mm (x = 0.35); pronotum not laterally explanate, lateral carina without a distinct sharp edge and reduced to a narrow bead; anterior angle acute and deflected, forming a short tooth (the angle formed between the anterior margin and lateral carina, excluding the tooth, is about 90 °); posterior angle indistinct and broadly rounded and without a short tooth; anterior foveae visible only in lateral view, posterior foveae visible in dorsal view; macropunctures ovate, deep, and well-separated, micropunctures moderately coarse; setae short, decumbent to suberect, and about 1 / 3 to 1 / 2 the width of eye. Elytra about 2.98 x as long as wide (elytral length / greatest elytral width = 2.65 – 3.23, x = 2.98) and 3.17 x as long as pronotum (elytral length / pronotal length = 2.90 – 3.33, x = 3.17); surface not incurvate at basal third; macropunctures, especially of striae 1 – 5, coalescing posteriorly to form grooves, separated by an average of ½ of a puncture diameter; micropunctures fine; microsculpture not visible; setae suberect, short, about 1 / 3 the length of the eye. Aedeagus with phallobase slightly transverse and rectangular, about 1 / 2 the length of the median lobe, median carina present; median lobe projecting significantly beyond apices of parameres, relatively narrow with its greatest length narrower than the parameres, apex acute and attenuate, median carina or groove absent, placement of ostium basal; parameres moderately broad and apically blunt, and anvil-shaped with laterally directed tooth, apical setae present and moderately elongate, internal subapical ridge absent. Spiculum gastrale apically broad.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDD3A6EFF216D2AF2A2FA81.taxon	discussion	Comments. Among species with coalescent elytral punctuation L. loebli is the only species with the coloration of the prolegs different from the middle and hindlegs. This species was described based on a single female.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDD3A6EFF216D2AF2A2FA81.taxon	distribution	Distribution. Nepal.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFDD3A6EFF216D2AF2A2FA81.taxon	materials_examined	Type material examined. HOLOTYPE (MHNG). 1 female (card mounted and dissected), NEPAL: distr. Kathmandu: Phulcoki 2500 m, 28 – 29. IV. 84 Löbl – Smetana / Laricobius sp. n. (hw) det. Löbl 19 / HOLOTYPE Ƥ Laricobius loebli sp. n. J. Háva & J. Jelínek det. 2000 (red label). Additional material examined. 1, 525 NEPAL: Solukhumbu Distr. Hinku Dranka Khola Bridge, 2000 m, 18 – 19 V. 1997 leg. W. SCHAWALLER (SMNS); 2, 564 NEPAL: Dailekh Distr. Dailekh to Mabuchin Pass, 2300 m, 3. – 4. VI. 1998 leg. W. SCHAWALLER (SMNS).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC23A6EFF216FD2F20DF864.taxon	diagnosis	Diagnosis (based on Nikitsky & Lafer, 1992). Length 2.1 mm. Body rusty brown, with the thorax a little darker ventrally, elytra bicolored with a black-brown or black stripe along the suture; antennae yellow-brown, legs partially darkened; dorsal vestiture erect. Ocelli present but difficult to distinguished from the rusty color of the head. Prothorax transverse (longer than wide), sides almost evenly convex and converging from middle, with anterior angles rounded and posterior angles “ visible ”. Elytra elongate (1.5 – 1.6 X longer than wide).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC23A6EFF216FD2F20DF864.taxon	discussion	Comments. No specimen of this species was examined. The description of the species was based on a single male specimen originally deposited in the ZMUM. It was subsequently lost in mail prior to this study (Nikitsky, personal communication). The diagnosis above was based on the figure of the head and prothorax and the key in Nikitsky and Lafer (1992), I provide the diagnosis above that will help to distinguish this species from L. kovalevi and L. sahlbergi.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC23A6EFF216FD2F20DF864.taxon	distribution	Distribution. Russia: Far East.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC03A6CFF216A8EF4EBFA14.taxon	description	(fig 13)	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC03A6CFF216A8EF4EBFA14.taxon	diagnosis	Diagnosis. Body unicolored. Head light; ocelli present. Prothorax quadrate without well developed lateral carina, sides not strongly explanate. Pronotum light; posterior tooth absent. Scutellum light. Surface of elytra incurvate; elytral punctures posteriorly confluent. Epipleura light. Ventrites dark. Femora light. Tibiae light. Aedeagus with medium lobe acute.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC03A6CFF216A8EF4EBFA14.taxon	description	Description. Length 2.86 mm. Body unicolored, mostly light brown or tan dorsally, prosternum light brown, thoracic and abdominal ventrites dark brown; antenna (with A 1 slightly darker), palpi, and legs light tan. Head with ocelli; u-shaped furrow absent; interocular distance about 2 x the width of the eye; macropunctures ovate and deeply impressed, scattered and present at center of frons; micropunctures coarse; microsculpture visible; setation short, length shorter than 1 / 3 the width of the eye. Antenna with length of A 3 shorter than A 4, ratios 1.6: 1.3: 1.0: 1.3: 1.5: 1.4: 1.5: 1.0: 1.4: 1.3: 1.8; A 11 strongly asymmetrical. Prothorax quadrate and convex, widest just antemedially, sides sinuate; about 0.82 x as long as wide (pronotal length / greatest pronotal width = 0.82); depth = 0.40 mm; pronotum not laterally explanate, lateral carina poorly developed and reduced to a narrow bead; anterior angle acute, deflected, and forming a short laterally directed tooth (the angle formed between the anterior margin and lateral carina, excluding the tooth, is about 90 °); anterior foveae visible in lateral view only, posterior foveae visible dorsally; posterior angle indistinct and broadly rounded and without a short tooth; macropunctures ovate, deep, and well-separated; micropunctures coarse; microsculpture visible; setae short, and about 1 / 2 the width of eye. Elytra about 3.23 x as long as wide (elytral length / greatest elytral width = 3.23) and 3.33 x as long as pronotum (elytral length / pronotal length = 3.33); surface weakly incurvate at basal third; macropunctures, especially of striae 1 – 2, coalescing posteriorly to form grooves, separated by an average of 1 / 3 of a puncture diameter; micropunctures fine; setae decumbent and short, about 1 / 2 length of the eye. Aedeagus (based on Háva 2009 a) with phallobase slightly transverse and rectangular, about 2 / 3 the length of the median lobe, median lobe projecting significantly beyond apices of parameres, relatively broad and apically arrowshape-d with its greatest width about that of the parameres, apex acute; parameres broad and apically acute, internal subapical ridge absent.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC03A6CFF216A8EF4EBFA14.taxon	discussion	Comments. Laricobius mirabilis is the only species with coalescent elytral punctuation that is unicolorous tan in color. It was described based on a single female, which was covered with a thin clear residue making some of the characters difficult to observe. The male genitalia was drawn by Háva (2009 a), presumably from an additional specimen that was recorded by Háva (2009 b), and was not made available for study.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC03A6CFF216A8EF4EBFA14.taxon	distribution	Distribution. China: Shaanxi, Yunnan.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC03A6CFF216A8EF4EBFA14.taxon	materials_examined	Type material examined. HOLOTYPE (NMPC). 1 female (card mounted and dissected), CHINA, Shangxi prov. Qing Ling Shan Mts., track Hou Zen Zi vill. to Taibai Shan, 3500 m, / 2 – 4 July 1998, alpina meadows, Z. Jindra, O. Šafránek, et M. Trýzna leg. / HOLOTYPE (red label) / Laricobius mirabilis sp. n. J. Háva & J. Jelínek det. 1999 / Mus. Nat. Pragae 66130 (number handwritten) Inv. (red label). Material not examined (see Háva 2009 b). China, N-Yunnan [C 2005 - 05 A], Diqing Tibet. Aut. Pref., Zhongdian Co., Xue Shan near lake 23 km S Zhongdian, 3895 m / 27 ° 37.1´N, 99 ° 38.5´E, devast mixed forest, meadows, lake border leaf litter, dead wood, sifted, 6. vi. 2005, M. Schülke leg., 1 Ƥ, J. Háva det. (SMNS).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC03A6DFF216C2FF250F96E.taxon	description	(figs 14, 29, 30, 43)	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC03A6DFF216C2FF250F96E.taxon	diagnosis	Diagnosis. Body bicolored. Head dark; ocelli absent. Prothorax transverse with well developed lateral carina and sides explanate. Pronotum dark; posterior tooth absent. Scutellum dark. Surface of elytra usually incurvate; elytral punctures not confluent. Epipleura dark. Ventrites dark. Femora dark. Tibiae light. Aedeagus with median carina on phallobase; medium lobe acute; apices of parameres without an internal ridge.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC03A6DFF216C2FF250F96E.taxon	description	Description. Length 2.40 – 2.84 mm (x = 2.58, n = 9). Body bicolored with head, prothorax, scutellum, elytra along sides to level of abdominal ventrite 3 (usually covering striae 7 – 10) and usually along suture and scutellary striole (but may be reduced to the area surrounding the scutellum), epipleuron, venter, and usually the femora chocolate brown; elytra orange-red to dark or reddish tan; antenna reddish brown to dark brown with AI and club usually darker, palpi, and tibiae light to reddish brown, tarsi lighter. Head without ocelli; u-shaped furrow present; interocular distance about 4 x the width of the eye; macropunctures rounded and distinct to indistinct and shallowly impressed, scattered and present at the center of dronsf; micropunctures coarse; setae erect, moderately elongate with an average length as long as 2 / 3 to full width of the eye. Antenna with length of A 3 about equal to A 4, ratios 1.8: 1.5: 1.3: 1.2: 1.2: 1.1: 1.1: 1.0: 1.2: 1.3: 1.9; A 11 not strongly asymmetrical. Prothorax transverse and rectangular, dorsoventrally flattened, and widest at middle, sides weakly convex, converging anteriorly from widest point and weakly convex or convergent posteriorly; about 0.74 x as long as wide (pronotal length / greatest pronotal width = 0.67 – 0.77, x = 0.74); depth = 0.28 – 0.44 mm (x = 0.37); pronotum laterally broadly explanate, lateral carina with a distinct sharp edge; anterior angle 90 º to about 80 ° and subacute, not forming a tooth; posterior angle indistinct, rounded, and without a short tooth; foveae visible in dorsal view; macropunctures on disc distinct, usually weaklyimpressed and well-separated; micropunctures strong; setae erect, moderately long and 2 / 3 to full width of eye. Elytra about 3.41 x as long as wide (elytral length / greatest elytral width = 3.13 – 3.73, x = 3.41) and 3.39 x as long as pronotum (elytral length / pronotal length = 3.13 – 3.71, x = 3.39); surface weakly incurvate at basal third; macropunctures not coalescing to form grooves, separated by 1 / 2 puncture diameter; micropunctures fine; microsculpture not visible; setae suberect to subdecumbent, short and on average about 2 / 3 length of the eye. Aedeagus with phallobase slightly elongate (longer than wide), about 2 / 3 the length of the median lobe, median carina present; median lobe projecting very slightly beyond apices of parameres, relatively narrow with its greatest width larger than the width of the parameres, apex acute and attenuate, median groove and carina absent, ostium subapical; parameres slender and acute, very short setae present on apices and along inner margin, internal subapical ridge absent. Spiculum gastrale apically narrow.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC03A6DFF216C2FF250F96E.taxon	discussion	Comments. Laricobius naganoensis can be distinguished from most species by the lack of ocelli. It can be reliably distinguished from the remaining species that lack ocelli by having pale tibiae and the acute median lobe of the male genitalia. Laricobius naganoensis can also be distinguished from these species, as well as the sympatric species L. osakensis, by the more vivid orange-red coloration of the elytra. This new species is based on specimens collected from relatively high altitude areas in Nagano Prefecture where it is sympatric with L. osakensis. Hosts. Plant: Tsuga diversifolia (Maxim.) Mast. Adelgid: Adelges tsugae.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC03A6DFF216C2FF250F96E.taxon	distribution	Distribution. Japan: Honshū.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC03A6DFF216C2FF250F96E.taxon	materials_examined	Type material examined. HOLOTYPE (OMNH): JAPAN Nagano Prefecture Ootaki Mt Ontake 35.86938 N 137.51064 E 2050 m 29 October 2009 A. Lamb S. Shiyake / Beaten from Tsuga diversifolia Havill Sample # 09 - 215.7 / HOLOTYPE Laricobius naganoensis design. R. Leschen 2010. PARATYPES: 1, JAPAN Nagano Prefecture Matsumoto Mt. Norikura 15 October 2008 Coll.: S. Shiyake A. Lamb Beaten from Tsuga diversifolia male Havill sample 08 - 291.1 (genitalia mounted separately on microslide, YPM); 1, JAPAN Nagano Prefecture Matsumoto Mt. Norikura 36.12 N 137.59 E 2030 m 28 October 2009 Coll.: A. Lamb S. Shiyake Laricobius sp. Female Beaten from Tsuga diversifolia Havill sample 09 - 214.2 (OMNH); 1, JAPAN Nagano Prefecture Ootaki Mt Ontake 35.87 N 137.51 E 2050 m 29 October 2009 Coll.: A. Lamb, S. Shiyake; Laricobius sp. Female Beaten from Tsuga diversifolia Havill sample 09 - 215.2 (OMNH); 5, JAPAN Nagano Prefecture Ootaki Mt Ontake 35.86938 N 137.51064 E 2050 m 29 October 2009 A. Lamb S. Shiyake Beaten from Tsuga diversifolia Havill Sample # 09 - 215 (2, OMNH; 1, YPM; 1 BMNH; 1 NMNH); 1, JAPAN; Nagano Prefecture Shiga-kogen 17 October 2008 Coll. S. Shiyake A. Lamb Laricobius osakensis Beaten from Tsuga diversifolia male Havill Sample # 08 - (OMNH); 1, JAPAN Nagano Prefecture Matsumoto Mt. Norikura 15 October 2008 Coll.: S. Shiyake A. Lamb Beaten from Tsuga diversifolia female Havill sample # 08 - 291.2 (OMNH); 1, JAPAN Nagano Prefecture Ootaki Mt Ontake 35.87 N 137.51 E 2050 m 29 October 2009 Coll. A. Lamb S. Shiyake Laricobius sp. Male Beaten from Tsuga (OMNH).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC13A68FF216CF2F25AFC1A.taxon	description	(figs 15, 44)	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC13A68FF216CF2F25AFC1A.taxon	diagnosis	Diagnosis. Body unicolored. Head dark; ocelli present. Prothorax transverse with well-developed lateral carina and sides explanate. Pronotum dark; posterior tooth present. Scutellum dark. Surface of elytra not incurvate; elytral punctures not confluent. Epipleura dark. Ventrites dark. Femora dark. Tibiae dark. Aedeagus with median carina on phallobase; medium lobe subacute; apices of parameres with an internal ridge.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC13A68FF216CF2F25AFC1A.taxon	description	Description. Length 2.02 – 2.73 mm (x = 2.49, n = 16). Body unicolored black; palpi and tarsi red or dark brown, tibiae black or dark brown and lighter at near the joints. Head with ocelli; u-shaped furrow present but shallow; interocular distance about 4 x the width of the eye; macropunctures rounded and distinct to indistinct and shallowly impressed and absent from the center of frons; micropunctures very coarse, especially posteriorly; setae erect, moderately elongate with an average length as long as 2 / 3 to full width of the eye. Antenna with length of A 3 about equal to A 4, ratios 1.7: 1.6: 1.6: 1.5: 1.5: 1.1: 1.2: 1.0: 1.4: 1.4: 1.9; A 11 generally not strongly asymmetrical. Prothorax transverse and rectangular, dorsoventrally flattened, and widest at middle, sides weakly convex, converging anteriorly from widest point and weakly convex or convergent posteriorly; about 0.76 x as long as wide (pronotal length / greatest pronotal width = 0.70 – 0.85, x = 0.76); depth = 0.28 – 0.44 mm (x = 0.34); pronotum laterally broadly explanate, lateral carina with a distinct sharp edge; anterior angle 90 º to about 60 ° and acute, not forming a tooth; posterior angle distinct and marked by a tooth which may be reduced; foveae visible dorsally; macropunctures on disc distinct to poorly defined, and shallowly impressed, well-separated; micropunctures coarse to moderately coarse; setae erect, moderately long and 2 / 3 to full width of eye. Elytra about 3.20 x as long as wide (elytral length / greatest elytral width = 2.91 – 3.56, x = 3.20) and 3.35 x as long as pronotum (elytral length / pronotal length = 2.81 – 3.64, x = 3.35); surface weakly incurvate at basal third; macropunctures not coalescing to form grooves (apart from those of striae 1 along the suture near apex), separated by 1 / 2 puncture diameter; micropunctures fine; microsculpture not visible; setae mostly suberect with scattered erect setae, elongate and on average about 2 / 3 to full length of the eye. Aedeagus with phallobase slightly transverse (wider than long), about 2 / 3 the length of the median lobe, median carina present; median lobe projecting beyond apices of parameres, relatively broad with its greatest width larger than the width of the parameres, gradually tapering apically with apex subacute, median carina or groove absent; ostium subapical; parameres moderately slender, and acute, short setae present on apices and along inner margin, internal subapical ridge present. Spiculum gastrale apically narrow and acute.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC13A68FF216CF2F25AFC1A.taxon	discussion	Comments. Laricobius nigrinus is the only unicolorous black species with a well-developed explanate pronotal margin. It can be distinguished from unicolorous L. sahlbergi and melanistic L. erichsonii by the dark femora and tibiae. Fender (1945) clearly states that the holotype, allotype, and five paratypes were collected from Bear Springs, Oregon, June 20, 1904, and one paratype from Creston, British Columbia, collected by G. Stace Smith October 29, 1933. The type material in the CAS and MCZ from Oregon are of different dates and the date given in the original description is probably a combination of a typo of “ 1904 ” and a simple error of “ 10 ” instead of “ 20 ” (the BC type was not recovered). Despite these and other inconsistencies among the labels, I consider that those below were part of the type series. After studies to determine its efficacy as a biocontrol agent for A. tsugae (Zilahi-Balogh et al. 2002 a, b), L. nigrinus was introduced the mid-Atlantic states in 2003 (Zilahi-Balogh et al. 2006) and is now widespread throughout eastern North America where hybridization between L. nigrinus and the native L. rubidus has been detected using microsatellite markers (Havill et al. in press). The external morphology of the hybrid individuals matches L. rubidus. Hosts. Plants: Larix sp., Larix occidentalis Nutt., Pinus monticola Douglas ex D. Don, Tsuga heterophylla. Adelgids: Adelges tsugae, Adelges lariciatus (Patch) Pineus sp.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC13A68FF216CF2F25AFC1A.taxon	biology_ecology	Biology. Natural history, larvae, morphology (Zilahi-Balogh et al. 2002 a, b, 2003, 2006).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC13A68FF216CF2F25AFC1A.taxon	distribution	Distribution. United States: Alaska, California, Idaho, Oregon, Washington, Wyoming (* intoduced to eastern North America and now in Delaware, Georgia, Kentucky, Maine, Massachusettes, Maryland, North Carolina, New Hampshire, New Jersey, New York, Pennsylvania, Tennessee, Vermont, Virginia, West Virginia. Canada: Alberta, British Columbia).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC13A68FF216CF2F25AFC1A.taxon	materials_examined	Type material examined. HOLOTYPE (CAS): BearSpringsOre; VI- 10 - 1940 [hw]; KM & DM Fender / Laricobius nigrinus Fender [hw], HOLOTYPE / California Academy of Sciences; Type No. 5527 [hw] / [hw label] Holotype [vertical with red highlight] Laricobius nigrinus Fender. PARATYPES: BearSpringsOre; VI- 10 - 1940 [hw]; KM & DM Fender / Laricobius nigrinus Fender [hw], ALLOTYPE / [hw label] Allotype [vertical w red highlight] Laricobius nigrinus Fender / Collection of the CALIFORNIA ACADEMY OF THE SCIENCES; San Francisco, California / [2 nd hw label] Allotype [vertical w red highlight], No. 5528, Laricobius nigrinus Fender (CAS). 1, Bear Springs Ore, 01 - 10. 1940, KM & DM Fender (printed and hw date) / Paratype (red label, hw) / Laricobius nigrinus Fender (red label, hw) / C. A. Frost Collection 1962 (MCZ). 1, same but 10 / 6 / 1940 (MCZ). Additional material examined. CANADA. Alberta: 11, Ghost Dam, Alberta (CNC). British Columbia: 4, Victoria, 20 – 21 Oct 1994, leg. L. M. Humble, FSQF- 94 - 3052 (VTRC); Saanichton, 12 IV 1991, T. heterophylla, L. Humble, CPFP Seed Orchard (VTRC); 5, Victoria, ex Tsugae heterophylla, Adelges tsugae (CNC); Vancouver, 3. IV. 1931, H. B. Leech, Blaisdell Collection (CAS); 1, Vancouver (CNC); 3, Victoria, 2 II 2002, A. Lamb, Tsuga heterophylla, Adelges tsugae (VTRC); 1, near Vernon, 10 May 2007, G. Zilahi-Balogh, Late instar collected in May, reared in lab to adult, ex. Larix oxidentalis (VTRC); 1, Vernon, Kalamalka Seed Orchard, ex larch cone hit by adelgids, collected as larva 7 May, live adult recovered 27 Aug 2007, J. E. Corrigan (VTRC). UNITED STATES. Alaska: 1, Whitehorse, Yukon Territory (NMNH). California: 1, Teh. Co., California (UNHD). Idaho: 13, Bonner Co. Sandpoint, 8 Oct 2008, D. Mausel, beating western hemlock with A. tsugae (VTRC); 2, Moscow (MCZ); 1, Kootenai County, Coeur d'Alene, Couger Gulch, 9 March 2007, D. Mausel, ex. Tsuga heterophylla, Sample: Havill 07 - 03.3 (PBNH); 9, same but Sample: Havill 07 - 03.1 (PNHM); 8, same but Sample: Havill 07 - 04 (PNHM); 3, Latah County, Moscow, 9 November 2007, Coll. S. Cool, D. Mausel, ex. Pinus monticola, Sample: Havill 07 - 134 (PNHM); 1, same but Sample: Havill 07 - 134.3 (PNHM). Oregon: 6, Oregon, Portland, Grant Park, 23 January 2006, Coll. D. Ross, Havill Sample # 06 - 119 (PNHM); 1, Polk Co. Bethel Heights, 45 ° 02.750 ' N, 123 ° 08.000 ' W, 26. X. 2007, Coll. Glen Kohler, Sample: Havill 07 - 135 (PNHM). Washington: 54, King Co. Seattle, 12 Oct 2008, D. Mausel, beating western hemlock with A. tsugae, 77 m, 47 º 39 ' 58.62 N, 122 º 20 ' 37.69 W (VTRC); 1, Grand Coulee (CNC); 7, King County, Seattle, 18 January 2007, coll. D. Mausel, C. Jubb, ex. Tsuga heterophylla; Sample: Havill 07 - 02 (PNHM); 1, same but Sample: Havill 07 - 02.2 (PBNH); 18, same but Sample: Havill 07 - 138 (PBNH); King Co.; Seattle; Marymoore Park, 7 April 2008, Coll. R. C. McDonald, ex. Tsuga heterphyllia, Sample 08 - 54.1 (VTRC); 4, Thurston Co. Olympia, 26 October 2007, Coll. G. Kohler, ex. Tsuga heterophylla, Sample: Havill 07 - 136 (PNHM); 1, same but Sample: Havill 07 - 136.6 (PNHM); 1, same but Sample: Havill 07 - 136.7 (PNHM).; Longmire Spgs VII- 12 - 19, Blaisdell Collection (CAS). Wyoming: 1, Saratoga, (NMNH). Lab reared: Virginia: 23, Blacksburg, VA; 10 IV 2003 (VTRC); 4, Washington, King Co., F 1 reared in VA Tech Lab, June 2006, Coll. D. Mausel, Havill Sample # 06 - 115 (PNHM).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC43A69FF216E26F45CFAC6.taxon	description	(figs 16, 43, 51)	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC43A69FF216E26F45CFAC6.taxon	diagnosis	Diagnosis. Body bicolored. Head dark; ocelli absent. Prothorax transverse with well-developed lateral carina and sides explanate. Pronotum dark; posterior tooth absent. Scutellum dark. Surface of elytra incurvate; elytral punctures not confluent. Epipleura at base dark. Ventrites dark. Tibiae and femora dark. Aedeagus with median carina on phallobase; medium lobe subacute; apices of parameres without an internal ridge.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC43A69FF216E26F45CFAC6.taxon	description	Description. Length 2.18 – 2.82 mm (x = 2.50, n = 15). Body bicolored with head, prothorax, scutellum, elytra along sides to level of abdominal ventrite 3 (usually covering striae 7 – 10) and suture along scutellary striole, epipleuron, venter, and usually the femora brown to black; elytra dark to light tan; antenna tan with AI dark tan and club usually darker than funicle but lighter than A 1, palpi, and tarsi tan, femora and tibiae dark tan or brown. Head without ocelli; u-shaped furrow present, but very weak or absent; interocular distance about 4 x the width of the eye; macropunctures rounded and distinct to indistinct and shallowly impressed; micropunctures coarse; setae erect, moderately elongate with an average length as long as 2 / 3 the width of the eye. Antenna with length of A 3 about equal to A 4, ratios 2.0: 1.7: 1.4: 1.5: 1.5: 1.1: 1.2: 1.0: 1.4: 1.5: 2.1; A 11 not strongly asymmetrical. Prothorax transverse and rectangular, dorsoventrally flattened, and widest at middle, sides weakly convex, converging anteriorly from widest point and weakly convex or convergent posteriorly; about 0.77 x as long as wide (pronotal length / greatest pronotal width = 0.68 – 0.87, x = 0.77); depth = 0.25 – 0.38 mm (x = 0.32); pronotum laterally broadly explanate, lateral carina with a distinct sharp edge; anterior angle 90 º to about 80 ° and subacute, not forming a tooth; posterior angle distinct, rounded, and without a short tooth; foveae visible dorsally; macropunctures on disc distinct, usually weakly-impressed and well-separated; micropunctures coarse; setae erect, moderately long and 2 / 3 to full width of eye. Elytra about 3.24 x as long as wide (elytral length / greatest elytral width = 2.81 – 3.53, x = 3.24) and 3.20 x as long as pronotum (elytral length / pronotal length = 2.81 – 3.61, x = 3.20); surface weakly incurvate at basal third; macropunctures not coalescing to form grooves, separated by 1 / 2 puncture diameter; micropunctures sparse; microsculpture visible in some specimens (but usually absent); setae suberect to subdecumbent, short and on average about 1 / 3 to 2 / 3 length of the eye. Aedeagus with phallobase transverse (wider than long), just over 2 / 3 the length of the median lobe, median carina present; median lobe projecting slightly beyond apices of parameres, relatively broad with its greatest width larger than the width of the parameres, apex subacute, median carina or groove absent, ostium subapical; parameres narrow, slender and acute, very short setae present on apices and not along inner margin, internal subapical ridge absent. Spiculum gastrale apically narrow.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC43A69FF216E26F45CFAC6.taxon	discussion	Comments. Laricobius osakensis lacks ocelli and can be distinguished from other species with this character by the absence of a well-developed posterior tooth on the pronotal margin. The dark tibiae and red-brown rather than vivid orangish elytral coloration distinguish it from L. naganoensis: but to confirm the species collected in higher altitudes of Nagano Japan, dissection of males is recommended. Hosts. Plants: Tsuga diversifolia, Tsuga sieboldii Carr. Adelgids: Adelges tsugae.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC43A69FF216E26F45CFAC6.taxon	biology_ecology	Biology. Natural history notes (Montgomery et al. 2011).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC43A69FF216E26F45CFAC6.taxon	distribution	Distribution. Japan: Honshū, Shikoku.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC43A69FF216E26F45CFAC6.taxon	materials_examined	Type material examined. PARATYPES. 14, JAPAN, Gunma Prefecture, Katashina, Mt. Nikko-Shirane, 36.81287 N, 139.34099 E, 1550 m, 31 October 2009, Coll.: A. Lamb & S. Shiyake, Havill Sample # 09 - 219 (2, NZAC; 2, OMNH; 10, YPM); 1 JAPAN, Nagano Prefecture, Shiga-kogen, 17 October 2008, Coll.: S. Shiyake & A. Lamb, Havill Sample # 08 - 289.2 (YPM); 1, JAPAN, Nagano Prefecture, Matsumoto, Mt. Norikura, 36.11419 N, 137.61401 E, 1590 m, 28 October 2009, Coll.: A. Lamb & S. Shiyake, Havill Sample # 09 - 213 (YPM); 3, JAPAN, Nagano Prefecture, Matsumoto, Mt. Norikura, 36.12203 N, 137.58749 E, 2030 m, 28 October 2009, Coll.: A. Lamb & S. Shiyake, Havill Sample # 09 - 214 (YPM); 3, JAPAN, Nagano Prefecture, Ootaki, Mt. Ontake, 35.86938 N, 137.51064 E, 2050 m, 29 October 2009, Coll.: A. Lamb, & S. Shiyake, Havill Sample # 09 - 215 (YPM); 4, JAPAN, Nagano Prefecture, Yamanouchi, Shiga-kogan, 36.68213 N, 138.50032 E, 1720 m, 30 October 2009, Coll.: A. Lamb & S. Shiyake, Havill Sample # 09 - 218 (YPM); 4, JAPAN, Osaka Prefecture, Kawachi Nagano, Site # 1 Kita-san, 34 º 24.150 ' N, 135 º 33.761 ' E, 231 m, 27 Feb 2006, Coll.: T. McAvoy & A. Lamb, Virginia Tech nos. 147 – 150 (YPM); 3, JAPAN, Tochigi Prefecture, Nikko, Konsei Pass, 36.81883 N, 139.39461 E, 2020 m, 1 November 2009, Coll.: A. Lamb & S. Shiyake, Havill Sample # 09 - 216 (YPM); 4, JAPAN, Tochigi Prefecture, Nikko, Nikko-Yumoto Spa, 36.80325 N, 139.4203 E, 1485 m, 31 October 2009, Coll.: A. Lamb & S. Shiyake, Havill Sample # 09 - 22 (YPM). Additional material examined. JAPAN. 2, various localities in Osaka Prefecture and Shikoku (Kansai area?), January 2008, A. Lamb, Havill Sample # 08 - 263.1 (PNHM); 2, JAPAN, Osaka Prefecture, Kobe Municipal Arboretum, 34 ° 44.445 ' N, 135 ° 10.566 ' E, 2 - III- 2006 – 7 - III- 2006, Coll.: T. McAvoy, A. Lamb & S. Shigehiko, Virginia Tech nos. 177 – 178 (YPM). UNITED STATES. Virginia (lab reared): 6, Nov. 2007, M. Beck, F 2 originally from Osaka Pre., Virginia Tech nos. 171 – 176; 1, Virginia Tech rearing facility, 2007 – 2008, A. Lamb, Havill Sample # 08 - 264 (YPM); 2, same but October to November 2006, A. Lamb, F 1 generation; emerged as adults in Oct. to Nov. 2006, parents from various sites in Japan; Honshu, Havill Sample # 06 - 117 (YPM).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC53A67FF216F9AF1D8FC89.taxon	description	(figs 17, 24, 25, 32, 46)	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC53A67FF216F9AF1D8FC89.taxon	diagnosis	Diagnosis. Body bicolored. Head dark; ocelli present. Prothorax transverse with well-developed lateral carina and sides explanate. Pronotum dark; posterior tooth usually present. Scutellum dark. Surface of elytra incurvate; elytral punctures not confluent. Epipleura at base dark. Ventrites dark. Femora dark. Tibiae dark. Aedeagus without median carina on phallobase; medium lobe subacute; apices of parameres with an internal ridge.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC53A67FF216F9AF1D8FC89.taxon	description	Description. Length 2.03 – 2.50 mm (x = 2.32, n = 16). Body bicolored, with head, prothorax, scutellum, and ventrites light brown to black, elytra along sides to apex (covering striae 7 – 10 or 8 – 10) and suture dark brown to black, or sutural stripes distinct and present only to basal 1 / 3 or less, with the broad central disc dark tan to deep red brown; antenna light to dark brown with AI and club sometimes darker, rarely the funicle dark brown to black, palpi, and tarsi reddish brown to tan, femora and tibiae light to chocolate brown (tibiae same color as femora). Head with ocelli; u-shaped furrow present but weakly impressed; interocular distance about 4 x the width of the eye; macropunctures weakly to strongly impressed, ovate; micropunctures coarse; setation elongate, erect, average length about 2 / 3 to full width of the eye. Antenna with length of A 3 about equal to A 4, ratios 1.9: 1.7: 1.4: 1.3: 1.4: 1.2: 1.2: 1.0: 1.3: 1.4: 1.8; A 11 not strongly asymmetrical. Prothorax transverse and dorsoventrally flattened, widest at middle, sides weakly sinuate or unevenly convex, with margin converging more sharply posteriorly than anteriorly; about 0.75 x as long as wide (pronotal length / greatest pronotal width = 0.67 – 0.81, x = 0.75); depth = 0.25 – 0.39 mm (x = 0.30); pronotum laterally broadly explanate, lateral carina with a distinct sharp edge; anterior angle variable from 90 º to about 65 °, acute but not forming a tooth; posterior angle distinct and marked by a short tooth that may be weakly formed; foveae visible in dorsal view; macropunctures ovate, deep, usually wellseparated but sometimes contiguous; micropunctures coarse; setae erect and elongate, length about 2 / 3 to full width of eye. Elytra about 3.22 x as long as wide (elytral length / greatest elytral width = 2.75 – 3.65, x = 3.22) and 3.25 x as long as pronotum (elytral length / pronotal length = 3.00 – 3.67, x = 3.25); surface strongly to weakly incurvate at basal third; macropunctures not coalescing posteriorly to form grooves (apart from those of striae 1 along the suture at the apical 1 / 3 in some specimens), separated by an average of 1 / 2 puncture diameter; micropunctures fine; microsculpture visible or not; vestiture mostly consisting of suberect setae with scattered erect setae, elongate and average length about 2 / 3 to full width of eye. Aedeagus with phallobase transverse (wider than long), about 2 / 3 the length of the median lobe, median carina absent; median lobe projecting beyond apices of parameres, relatively broad with its greatest width larger than the width of the parameres, gradually tapering apically with apex subacute, median carina or groove absent, ostium subapical; parameres moderately broad, apex angulate, short setae present on apices and absent along inner margin, internal subapical ridge present. Spiculum gastrale apically acute.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC53A67FF216F9AF1D8FC89.taxon	discussion	Comments. Laricobius rubidus can be distinguished from other bicolored species with ocelli and explanate pronotal margins (especially the similar looking L. erichsonii) by the dark-colored tibiae which is the same color as the femora, by the punctures on the head shallowly impressed, the anterior angle not forming a tooth, and most easily by the aedeagal characters. The internal subapical ridge on the parameres is shared only with the species L. nigrinus. Preliminary genetic data suggest that L. nigrinus and L. rubidus can successfully reproduce in the field (Havill et al. in press). Host plants and distributions were recently included in the study by Majka (2007). Hosts. Plants: Abies fraseri (Pursh) Poir., Abies balsamaea (L.) Mill., Pinus strobus L., Pinus banksiana Lamb., Tsuga canadensis (L.) Carriére, Picea rubens Sarg. Adelgids: Adelges tsugae, Adelges piceae (Ratzeburg), Pineus strobi.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC53A67FF216F9AF1D8FC89.taxon	biology_ecology	Biology. Natural history (Clark & Brown 1960, Lawrence & Hlavac 1979).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC53A67FF216F9AF1D8FC89.taxon	distribution	Distribution. Canada: New Brunswick, Nova Scotia, Ontario, Quebec. United States: Connecticut, Georgia, Maine, Maryland, Massachusettes, New Hampshire, New York, North Carolina, Pennsylvania, Rhode Island, Tennessee, Vermont, West Virginia.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC53A67FF216F9AF1D8FC89.taxon	materials_examined	Type material examined. Holotype (MCZ). 1, pointed, D. C / Laricobius rubidus Lec. (hw) / erichsonii 2 (hw) / M. C. Z. Type 32369 (red label, number hw) / LARICOBIUS RUBIDUS LEC. Det. J. F. Lawrence (bordered det. label, name hw) / Holotype (red label) / Jan – Jul 2005 MCZ Image Database (bordered label with camera icon). Additional material examined. CANADA. New Brunswick: 1 (CNC). Ontario: 1, Constance B., 16. v 1933, W. J. Brown (CAS); 2, Ottawa (CAS); 5, Toronto, R. J. Crew (CAS); 24 (CNC). Quebec: 4, Aylmer, Que., 1. V 1933, W. J. Brown / [hw label] Det. 1934 W. J. B. (CAS); 1, same but 18 - v- 1934 (CAS); 1, same but 9. v 1932 (CAS); 1, same but 1. v 1933 (CAS); 1, Aylmer, 12. v 1932, W. J. Brown (CAS); 2, Lanoraie, 13 iii 77, F. Liard, 1992 Acc. Z- 18,343 (FMNH). UNITED STATES. Connecticut: 1, New Haven Co., New Haven, 30 March 1994, Coll.: S. Lyon (USDA); same but 19 April 1994 (USDA); 6, New Haven Co, Lake Whitney, 31 X 1997, M. Montgomery, W. Lu (USDA); 1, same but 14 vii 1993 (USDA); 1, Hamden, end of Ingram St., Water Co. property, 27 March 2007, Coll.: N. Havill, ex. Tsuga canadensis, Sample: Havill 07 – 14 (YPM); 1, Hamden, USDA Forest Service Northern Research Stn., 51 Mill Pond Rd., 17 April 2008, Coll.: N. Havill, A. Serafin, ex. Pinus strobus, Sample 08 - 57.1 (YPM). Maryland: 1, Allegany County, Rocky Gap, 3 May 2006, Coll.: D. Mausel, ex. Tsuga canadensis, larva collected. reared in lab, UTM 4397022 N, 0699934 E, Sample: Havill 06 - 126 (YPM). Massachusettes: 1 (CAS); 1, Holyoke, Mt. Tom State Park, 5 May 2008, Coll.: J. Biroscak, ex. Pinus strobus, Sample 08 - 71.1 (YPM). Michigan: 1 (CAS). Minnesota: 1, on Pinus strobes (CNC). New Hampshire: 53, (UNHD). New York: 1 (CNC). North Carolina: 1, Watauga County, Holloway Gap, 16 April 2006, Coll.: D. Mausel, ex. Tsuga canadensis, larva collected. reared in lab, UTM 4000032 N, 0431471 E, Sample: Havill 06 - 122.3 (YPM); 1, same but Sample: Havill 06 - 122.4 (YPM); 3, Watauga County, Fosco, Holloway Mountain Road, 17 April 2008, Coll.: M. E. Montgomey, R. C. McDonald, ex. Pinus strobus; 36.14 N: 81.76 W, Sample 08 - 61 (YPM); 1, same but Sample 08 - 61.1 (YPM); 2, Yancey County, Locust Creek, 16 April 2006, Coll.: D. Mausel, ex. Tsuga canadensis, larva collected. reared in lab, UTM 3964523 N, 0390540 E, Sample: Havill 06 - 123 (YPM); 1, same but Sample: Havill 06 - 124.1 (YPM). Pennsylvania: 1, Cumberland Co., Hogestown, 12 April 1994, Coll.: S. Lyon (USDA); 3, Huntingdon Co., Rothrock State Forest, 4 May 2006, Coll.: D. Mausel, ex. Tsuga canadensis, larva collected. reared in lab, UTM 4504040 N, 0268595 E, Sample: Havill 06 - 128 (YPM). Tenessee: 7, Sevier Co., Smokies Nat. Park, Laurel Creek, 9 April 2006, Coll.: D. Mausel, ex. Tsuga canadensis, larva collected. reared in lab; UTM 3943370 N, 0250526 E, Sample: Havill 06 - 125 (YPM); 1, same but Sample: Havill 06 - 125.5 (YPM); 1, same but Sample: Havill 06 - 125.6 (YPM). Virginia: 6, Montgomery Co. Blacksburg, 6 - 30 IV 2001, coll. G. Zilahi-Balogh, Host tree: Tsuga canadensis, Host: Adelges tsugae, Habitat: plantation <10 yr. mixed, w / Abies fraseri, Pinus strobus (VTRC); 3, Montgomery Co. Prices Fork, 6 – 30 IV 2001, G. Zilahi-Balogh, Host: Tsuga canadensis infested w / Adelges tsugae (VTRC); 7, Smyth Co., Hurricane Camp, 20 April 2006, Coll.: D. Mausel, ex. Tsuga canadensis, larva collected. reared in lab, UTM 4064121 N; 0456464 E, Sample: Havill 06 - 129 (YPM); 6, Montgomery Co. Kentland Farm, 28 April 2006, Coll.: D. Mausel, ex. Tsuga canadensis, larva collected. reared in lab, UTM 4118054 N, 0536469 E, Sample: Havill 06 - 132 (YPM); 1, Bland Co., Lick Creek, 18 April 2006, Coll.: D. Mausel, ex. Tsuga canadensis, larva collected. reared in lab, UTM 4096147 N; 0461976 E, Sample: Havill 06 - 130 (YPM); 8, Giles Co., North Fork, 15 April 2006, Coll.: D. Mausel, ex. Tsuga canadensis, larva collected. reared in lab, UTM 4144202 N, 0542872 E, Sample: Havill 06 - 133 (YPM); 1, same but Sample: Havill 06 - 133.9 (YPM); 7, Giles Co., Big Stony, 15 April 2006, Coll.: D. Mausel, ex. Tsuga canadensis, larva collected, reared in lab, UTM 4141138 N, 0538027 E, Sample: Havill 06 - 135 (YPM); 1, Grayson Co., Highland Trail, 24 April 2006, Coll.: D. Mausel, ex. Tsuga canadensis, larva collected, reared in lab, UTM 4060842 N, 0453808 E; Sample: Havill 06 - 134 (YPM). Vermont: 1 (CNC). West Virginia: 15, Pocahontas Co., Seneca State Forest, 27 April 2006, Coll.: D. Mausel, ex. Tsuga conadensis, larva collected. reared in lab, UTM 4239369 N, 0594329 E; Sample: Havill 06 - 120 (YPM); 1, Pocahontas Co., Watoga State Park, 27 April 2006, Coll.: D. Mausel, ex. Tsuga canadensis, larva collected. reared in lab, UTM 4217462 N, 0579409 E, Sample: Havill 06 - 121.10 (YPM); 20, but Sample: Havill 06 - 121 (YPM). Laboratory reared: 3, Blacksburg, VA; 10 IV 2003 (VTRC).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFCB3A64FF2169D7F341FC1A.taxon	description	(figs 18, 47)	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFCB3A64FF2169D7F341FC1A.taxon	diagnosis	Diagnosis. Body unicolored. Head dark; ocelli present. Prothorax transverse with well-developed lateral carina and sides explanate. Pronotum dark; posterior tooth absent. Scutellum dark. Surface of elytra incurvate; elytral punctures not confluent. Epipleura dark. Ventrites dark. Femora dark. Tibiae light. Aedeagus without median carina on phallobase; medium lobe subapiculate; apices of parameres without an internal ridge.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFCB3A64FF2169D7F341FC1A.taxon	description	Description. Length 2.58 – 2.72 mm (x = 2.63, n = 4). Body unicolored dark brown; palpi, tibiae, and tarsi tan or yellow brown, antennae yellow with A 1 darker and club yellow brown. Head with ocelli; u-shaped furrow present; interocular distance about 4 x the width of the eye; macropunctures distinct to indistinct, impressed; micropunctures coarse; setae erect, moderately short with an average length as long as 1 / 2 width of the eye. Antenna with length of A 3 about equal to A 4, ratios 1.8: 1.5: 1.3: 1.2: 1.2: 1.0: 1.1: 1.0: 1.3: 1.4: 2.4; A 11 not strongly asymmetrical. Prothorax transverse and rectangular, dorsoventrally flattened, and widest at middle, sides weakly convex, converging anteriorly from widest point and more steeply convergent posteriorly; about 0.73 x as long as wide (pronotal length / greatest pronotal width = 0.71 – 0.76, x = 0.73); depth = 0.30 – 0.40 mm (x = 0.35); pronotum laterally broadly explanate, lateral carina with a distinct sharp edge; anterior angle about 60 ° and acute, not forming a tooth; posterior angle distinct and not marked by a short tooth; foveae visible dorsally; macropunctures on disc distinct, well-impressed, and well-separated; micropunctures coarse; setae erect, moderately long and the average length 2 / 3 width of eye. Elytra about 3.17 x as long as wide (elytral length / greatest elytral width = 3.00 – 3.30, x = 3.17) and 3.32 x as long as pronotum (elytral length / pronotal length = 3.19 – 3.47, x = 3.32); surface weakly incurvate at basal third; macropunctures not coalescing to form grooves, separated by an average of 1 puncture diameter; micropunctures fine; microsculpture not visible; setae suberect and erect, short and on average about 1 / 2 the length of the eye. Aedeagus with phallobase transverse (wider than long), about 2 / 3 the length of the median lobe, median carina absent; median lobe not projecting beyond apices of parameres, relatively broad with its greatest width subequal to the width of the parameres, gradually tapering apically with apex subapiculate, median carina or groove absent, ostium subapical; parameres broad, apices rounded, short apical and subapical setae present, internal subapical ridge absent. Spiculum gastrale apically narrow or acute.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFCB3A64FF2169D7F341FC1A.taxon	discussion	Comments. Laricobius sahlbergi is the only unicolorous dark brown species with a well-developed explanate pronotal margin. It can be distinguished from unicolorous L. nigrinus by its pale femora and from melanistic L. erichsonii by the absence of posterior pronotal teeth and the acute median lobe of the male. I obtained four specimens from the HNMH, not all having the same labelling. Reitter (1883) gave a size range, so it is likely he had a series (only one specimen had a label bearing the type locality) and all the HNMH specimens, including one from the E. Frivaldszky collection, are considered part of the syntype series. The locality abbreviated on the lectotype label (Fatjanovsk.) may be the modified German translation of the Russian place name listed as Fatjanowskaja in Andree (1896). This may also be written as Fatianowskaya in older literature, but in modern maps and literature it is written as Fatianowo (or Fatianovo which means the same as Fatianovskaya) in Yaroslavl Oblast in European Russia with the coordinates 57.01 ’. 45.64 ” N, 38.54 ’ 49.77 ” E (A. Solodovnikov, pers. com.). Nikitsky (1992) listed the species range as middle Siberia in Krusnayerskykry (= Krasnoyarskij of Krasnoyarskij Kraj; A. Solodovnikov, pers. com.).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFCB3A64FF2169D7F341FC1A.taxon	distribution	Distribution. Russia: East Siberia. China: Mongolia.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFCB3A64FF2169D7F341FC1A.taxon	materials_examined	Type material examined. LECTOTYPE (HNHM). 1 female (card mounted and dissected), Fatjanovsk. / J. Sahlb. / Laricobius sahlbergi m. (hw) / Coll. Reitter / Holotypus (red) Laricobius sahlbergi 1883 Reitter (name and date hw on white label with red border) / Laricobius sahlbergi Reitter det. Nikitsky, 1992 (hw) / Laricobius Ƥ sahlbergi Rtt. Jirí Háva det. 0 3 (hw and printed) / LECTOTYPE Laricobius sahlbergi Reitter desig, R. Leschen, 2009. PARALECTOTYPES (HNHM). 1, male (card mounted and dissected, genitalia in glycerin pinned below specimen), Sibir. arct. / Paratypus (red) Laricobius sahlbergi 1883 Reitter (name and date hw on white label with red bor- der) / Coll. Reitter / PARALECTOTYPE Laricobius sahlbergi Reitter desig, R. Leschen, 2009. 1, sex unknown (card mounted), Sibir. arct. / J. Sahlb. / Paratypus (red) Laricobius sahlbergi 1883 Reitter (name and date hw on white label with red border) / Coll. Reitter / PARALECTOTYPE Laricobius sahlbergi Reitter desig, R. Leschen, 2009. 1, sex unknown (card mounted), Sibir. arct. / J. Sahlb. / L. Sahlbergi (hw) coll. E. Frivaldszky / 660 b 5 (hw on small folded black bordered label) / Sahlbergi Reitt Sibir. arct. (hw) / PARALECTOTYPE Laricobius sahlbergi Reitter desig, R. Leschen, 2009 Additional material examined. CHINA. 1, Mongolia, Central aimak Chentej Gebirge 25. IV. 1971 leg. Zs. Peregi / Laricobius sahlbergi Reitt. V. Richter det. 1973 (HNHM).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC83A65FF216E23F4CDFEEA.taxon	description	(fig 19)	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC83A65FF216E23F4CDFEEA.taxon	diagnosis	Diagnosis. Body bicolored. Head light; ocelli present. Prothorax transverse without well-developed lateral carina, sides not strongly explanate. Pronotum light; posterior tooth present. Scutellum dark. Surface of elytra incurvate; elytral punctures posteriorly confluent. Epipleura dark. Ventrites dark. Femora dark. Tibiae light.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC83A65FF216E23F4CDFEEA.taxon	description	Description. Length 2.24 mm. Body bicolored, mostly yellowish tan apart from the meso- and metaventritesthat are completely black, the abdominal ventrites which are infuscate, and most of the elytra which is mostly black to the apical 1 / 3 along suture, then expanding obliquely to apex to level of stria 5; antenna with AI and club slightly darker. Head with ocelli; u-shaped furrow absent; interocular distance about 2 x the width of the eye; macropunctures ovate and well-impressed, scattered and present at center of frons; micropunctures moderately coarse; setation short, decumbent, length about 1 / 3 the width of the eye. Antenna with length of A 3 about equal to A 4, ratios 2.1: 1.8: 1.7: 1.7: 1.6: 1.2: 1.2: 1.0: 1.4: 1.3: 2.2; A 11 strongly asymmetrical. Prothorax transverse and moderately convex, widest at apical 1 / 3, sides sinuate, and strongly convergent posteriorly; about 0.78 x as long as wide (pronotal length / greatest pronotal width = 0.78); depth = 0.32 mm; pronotum not laterally explanate, lateral carina narrow and reduced to a bead; anterior angle deflected, forming a short ridge and not dentate (the angle formed between the anterior margin and lateral carina, excluding the ridge, is about 60 °); posterior angle indistinct and broadly rounded and with a very short tooth; anterior foveae visible only in lateral view, posterior foveae visible in dorsal view; macropunctures ovate, deep, and generally well-separated; micropunctures moderately coarse; setae short, decumbent, and about 1 / 2 the width of eye. Elytra about 2.80 x as long as wide (elytral length / greatest elytral width = 2.80) and 3.00 x as long as pronotum (elytral length / pronotal length = 3.00); surface incurvate at basal third; macropunctures, especially of striae 1 – 3, coalescing posteriorly to form grooves, separated by an average of 2 / 3 of a puncture diameter; micropunctures fine; microsculpture not visible; setae suberect to subdecumbent, short, about 1 / 2 length of the eye.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC83A65FF216E23F4CDFEEA.taxon	discussion	Comments. Laricobius schawalleri is a distinctive species within the group with confluent elytral punctures by having a transverse prothorax and the apical and posteromedial portions of the elytra yellowish tan. This species was described based on a single female.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC83A65FF216E23F4CDFEEA.taxon	distribution	Distribution. Nepal.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC83A65FF216E23F4CDFEEA.taxon	materials_examined	Type material examined. HOLOTYPE (SMNS). 1 (card mounted and dissected), 564 NEPAL: Dailekh Distr. Dailekh to Mabuchin Pass, 2500 m, 3. – 4. VI. 1998 leg. W. SCHAWALLER / Laricobius sp. (hw) det. SCHA- WALLER 1999 / HOLOTYPE Laricobius schawalleri sp. n. J. Jelínek & J. Háva 2000 (red label).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC93A65FF216B76F17BF840.taxon	description	(figs 20, 48)	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC93A65FF216B76F17BF840.taxon	diagnosis	Diagnosis. Body bicolored. Head dark; ocelli absent. Prothorax transverse with well-developed lateral carina and sides explanate. Pronotum dark; posterior tooth present. Scutellum dark. Surface of elytra usually not incurvate; elytral punctures not confluent. Epipleura dark. Ventrites dark. Femora dark. Tibiae usually dark. Aedeagus with median carina on phallobase; medium lobe subacute; apices of parameres without an internal ridge.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC93A65FF216B76F17BF840.taxon	description	Description. Length 1.58 – 2.54 mm (x = 2.67, n = 15). Body bicolored with head, prothorax, scutellum, elytra along sides to level of abdominal ventrite 5 (usually extending to stria 4 and covering striae 5 – 10), along suture (though maybealit reduced or absent), epipleuron, and venter black or chocolate brown; elytra dark red brown; antenna with A 1 dark brown, funicle yellow brown, and club red-brown; palpi and legs dark brown, tarsi brown, but lighter in some specimens. Head without ocelli; u-shaped furrow present; interocular distance about 4 x the width of the eye; macropunctures rounded and distinct, impressed and sometimes fused into trenches and set into depressions; micropunctures moderately coarse; setae erect, elongate with an average length as long as 2 / 3 to full width of the eye. Antenna with length of A 3 about equal to A 4, ratios 2.1: 1.4: 1.1: 1.2: 1.3: 1.1: 1.1: 1.0: 1.3: 1.2: 1.9; A 11 not strongly asymmetrical. Prothorax transverse and rectangular, dorsoventrally flattened, and widest at middle, sides weakly convex, converging anteriorly from widest point and weakly convex, subparallel anteriorly and convergent posteriorly; about 0.79 x as long as wide (pronotal length / greatest pronotal width = 0.75 – 0.83, x = 0.79); depth = 0.16 – 0.38 mm (x = 0.29); pronotum laterally broadly explanate, lateral carina with a distinct sharp edge; anterior angle 90 º to about 80 ° and subacute, not forming a tooth; posterior angle distinct, rounded, and with a short tooth; foveae visible in dorsal view; macropunctures on disc distinct, well-impressed and well-separated; micropunctures coarse; setae erect, moderately long with length 2 / 3 to full width of eye. Elytra about 2.91 x as long as wide (elytral length / greatest elytral width = 1.86 – 3.71, x = 2.91) and 2.92 x as long as pronotum (elytral length / pronotal length = 1.73 – 3.30, x = 2.12); surface not or weakly-incurvate at basal third; macropunctures not coalescing to form grooves, separated on an average by 1 / 2 puncture diameter; micropunctures fine; microsculpture not visible; setae suberect to decumbent, short and on average about 2 / 3 to full length of the eye. Aedeagus with phallobase transverse (wider than long), 2 / 3 the length of the median lobe, median carina present; median lobe projecting slightly beyond apices of parameres, somewhat narrow with its greatest width equal to width of the parameres at their basal 1 / 3, apex subacute, median groove or carina absent, ostium subapical; parameres, slender and narrow, apically acute, very short setae present apically and along inner margin, internal subapical ridge absent. Spiculum gastrale narrow.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC93A65FF216B76F17BF840.taxon	discussion	Comments. There are no discernable ocelli in this species, though the color of the cuticle that surrounds the deeper cephalic macropunctures may be lighter. Among the species that lack ocelli, L. taiwanensis has the sides of the prothorax weakly convex, which will distinguish this species from the others. Having the dark tibiae will distinguish L. taiwanensis from L. naganoensis, the presence of of the posterior pronotal tooth will distinguish L. taiwanensis from L. osakensis, and the dark red coloration and presence of a median carina on the basal piece will distinguish L. taiwanensis from L. kangdingensis. The holotype (NMNS) was not examined, but several paratypes were, belonging to the same series. Hosts. Plants: Tsuga chinensis. Adelgid: Adelges tsugae.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC93A65FF216B76F17BF840.taxon	biology_ecology	Biology. Habitat and rearing (Yu & Montgomery 2007).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC93A65FF216B76F17BF840.taxon	distribution	Distribution. Taiwan: Kao Hsiung.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFC93A65FF216B76F17BF840.taxon	materials_examined	Type material examined. PARATYPES: 21 (10, BMNH [with all original labels hw]; 4, NMNH; 3, YPM; 4, USDA, one each with Havill Sample # 08 - 50, Havill Sample # 08 - 51), TAIWAN Kao Hsiung Co. Road 20, 2 Km west of Ya Kou Hotel 26 May 1994 S. T. Murphy / PARATYPE Laricobius taiwanensis, Yu and Montgomery (yellow green label).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFCF3A5DFF216CE5F16FFDC5.taxon	description	(figs 21, 49)	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFCF3A5DFF216CE5F16FFDC5.taxon	diagnosis	Diagnosis. Body bicolored. Head dark; ocelli present. Prothorax quadrate without well-developed lateral carina, sides not strongly explanate. Pronotum dark; posterior tooth absent. Scutellum dark. Surface of elytra incurvate; elytral punctures not posteriorly confluent. Epipleura dark. Ventrites dark. Femora light. Tibiae light. Aedeagus with medium lobe acute; apices of parameres without an internal ridge.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFCF3A5DFF216CE5F16FFDC5.taxon	description	Description. Length 2.00. Body bicolored with chocolate brown head, prothorax, ventrites, scutellum, and flanks of elytra to striae 7 and remaining areas and appendages light tan; antenna with AI and club darker. Head with ocelli; u-shaped fovea absent; interocular distance about 2 x the width of the eye; macropunctures ovate and well impressed; micropunctures coarse; setation short and decumbent, length about 1 / 3 the width of the eye. Antenna with length of A 3 less than A 4, ratios 1.8: 1.7: 1.0: 1.5: 1.7: 1.5: 1.3: 1.1: 1.7: 1.7: 2.1; A 11 strongly asymmetrical. Prothorax quadrate and highly convex, widest at middle, sides sinuate; about 0.84 x as long as wide (pronotal length / greatest pronotal width = 0.84); depth = 0.45; pronotum not laterally explanate, lateral carina reduced to a narrow bead; anterior angle acute, deflected, and forming a short laterally directed tooth (the angle formed between the anterior margin and lateral carina, excluding the tooth, is about 90 °); posterior angle indistinct and broadly rounded and without a short tooth; anterior foveae visible only in lateral view, posterior foveae visible in dorsal view; macropunctures ovate, deep, and well-separated; micropunctures moderately coarse; setae short, decumbent, and lengths about 1 / 3 the width of eye. Elytra about 1.6 x as long as wide (elytral length / greatest elytral width = 1.6) and 3.40 x as long as pronotum (elytral length / pronotal length = 3.40); surface incurvate at basal third; macropunctures of striae 1 coalescing posteriorly to form grooves, separated by an average of 1 puncture diameter; micropunctures sparse; microsculpture not visible; setae suberect and short, about 1 / 3 length of the eye. Aedeagus with phallobase transverse, about 2 / 3 the length of the median lobe; median lobe projecting beyond apices of parameres, apex acute, median groove present; parameres broad, apically acute and anvil-shaped with a distinct subapical tooth, moderately short setae present subapically, internal subapical ridge absent. Spiculum gastrale apically broad.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFCF3A5DFF216CE5F16FFDC5.taxon	discussion	Comments. Among the species without explanate pronotal margins, L. wittmeri is the only member of the group that lacks confluent elytral punctures (apart from the sutural striae). This species was described from two specimens (one male and one female), and only the male type was examined, the other was not made available for study. The Holotype is subteneral, and not fully pigmented. An additional specimen that was completely teneral was located in the CNC. The previously extracted genitalia of the Holotype was distorted and the width of the median lobe, presence of the median carina, ostium, and presence of internal subapical ridge were not observed.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFCF3A5DFF216CE5F16FFDC5.taxon	distribution	Distribution. Nepal.	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
C563080BFFCF3A5DFF216CE5F16FFDC5.taxon	materials_examined	Type material examined. HOLOTYPE (NHMB). O Nepal 1980 W. Wittmer / Mumbug, O Makalu 3500 m, 9.6. / HOLOTYPE 3 Laricobius wittmeri sp. n. J. Háva det. 2009 (red label with narrow black border). Additional material examined. Nepal: 1, 27 ° 58 ' N. 85 ° 00 ' E. Mal. Tr. 2, 11, 200 ', 28 May 1967, Can. Nepal Exped. (CNC).	en	Leschen, Richard A. B. (2011): World review of Laricobius (Coleoptera: Derodontidae). Zootaxa 2908: 1-44, DOI: 10.5281/zenodo.201491
