taxonID	type	description	language	source
C4290E04F40CFFEF118602D55A8A3D40.taxon	type_taxon	Type species In describing Scatonomus, Erichson (1835) included four species in the genus – namely, S. viridis, S. fasciculatus, S. barbatus and S. chalybaeus – but he did not fix a type species. Despite S. viridis being the first species described and the only one illustrated by Erichson, he did not refer to it as the type species of the genus. However, in the course of our literature review, we found a taxonomic remark in Westwood (1847) where the author referred to S. viridis as ‘ the typical species’ of Scatonomus. According to article 69 of ICZN (1999), Westwood’s publication must be accepted as a valid designation of the type species of the genus.	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F403FFE0111D06E158423B9F.taxon	description	(Figure 1 (a), 2 (a), 4 (a) and 5 (a ))	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F403FFE711F10555593D3B52.taxon	diagnosis	Diagnosis Scatonomus barbatus and S. fasciculatus are closely related species. Both species have body longer than 9 mm; dorsal surface shiny, almost smooth; genal edge not distinctly angulate laterally (Figure 3 (a, d )); and clypeal edge bidentate, with deep lateral emarginations filled with a tuft of long setae (Figure 3 (a, d )). However, S. barbatus may be distinguished from S. fasciculatus by the following combination of characters: median clypeal teeth shorter than the lateral tuft of setae (Figure 3 (a )); anterior edge of gena rounded (Figure 3 (a )); and dorsum with green reflections (Figure 1 (a )).	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F403FFE711F10555593D3B52.taxon	description	Redescription Male: length: 9 – 11 mm. Dorsum black with green reflections (Figure 1 (a )). Head: densely punctate, slightly wrinkled medially (Figure 3 (a )). Anterior and posterior edges narrowly margined. Clypeo-genal and fronto-clypeal sutures arched, moderately impressed, not carinate. Clypeal median teeth upturned, separated by a deep U-shaped emargination (Figure 3 (a )). Anterior portion of clypeus strongly concave medially, surface of concavity less punctate than remaining portions of head. Edges of clypeus and gena densely coated with rows of long erect setae. Antennal club globose, densely coated with long setae. Thorax: pronotum shiny, sparsely punctate, mid-longitudinal sulcus feebly impressed on disc. Lateral edge of hypomera with a row of indistinct setose punctures, setae short. Transverse carina of hypomera almost complete, not reaching lateral edge. Surface of mesepimeron and mesanepisternum with green reflections, finely striate. Anterior and lateral lobes of metaventrite with inconspicuous microgranulation; lateral lobes also with shallow punctures. Metaventral disc densely puncticulate. Elytra: striae thin, shallow, indistinctly punctate; striae on disc 7 times narrower than third interstria. Seventh stria not reaching the basal and apical portions of elytra. Eighth stria weakly impressed, visible at the apical half. Ninth stria weakly impressed near elytral base. Interstriae flat, surface shiny and sparsely punctate. Legs [ventral surface]: profemora densely punctate, surface between punctures smooth. Mesofemur and metafemur sparsely punctate. External edge of protibia with 3 distinct teeth separated by U-shaped emarginations. Outer edge of mesotibia and metatibia with a row of sparse setae and small teeth throughout their length. Abdomen: ventrite VI 1.5 times as long as ventrite V. Pygidium: 1 / 3 wider than long. Disc punctate, punctures spaced by 3 times their diameter. Genital capsule: paramera apex divided into apical and lateral lobes by a deep L-shaped emargination. Apical and lateral lobes slightly acute (Figure 5 (a )). Female: differs from male by the abdominal ventrite VI twice as long as V. Disc of pygidium with punctures spaced by 2 times their diameter; apical edge of pygidium slightly angulate.	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F406FFE511AF002B584B3DC6.taxon	description	(Figures 1 (b), 2 (a), 3 (b) and 6 (d ))	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F407FFE411FA074C58833A1E.taxon	description	(Figures 1 (c), 2 (b), 3 (c), 4 (h), 5 (b) and 6 (c ))	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F407FFFB115C07F458923D48.taxon	diagnosis	Diagnosis Scatonomus chalybaeus resembles S. fasciculatus and S. barbatus in having an elongated body, larger than 9 mm, and by lacking coarse and ocellate punctures on pronotum and elytra. However, S. chalybaeus is distinguished by having clypeal edge with a pair of cylindrical and short teeth, bounded by shallow lateral emarginations, not filled with a tuft of setae (Figure 2 (b) and 3 (c )).	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F407FFFB115C07F458923D48.taxon	description	Redescription Male: length: 10 – 13 mm. Dorsum black, usually with dark blue or green reflections (Figure 1 (c )). Head: dorsal surface transversely wrinkled, finely punctate medially (Figure 3 (c )). Anterior edge narrowly margined. Anterior genal edge and lateral clypeal edge straight (Figure 3 (c )). Clypeo-genal suture wide, straight, distinctly impressed. Fronto-clypeal suture ill defined between eyes. Clypeus and frons impressed medially. Inner edge of dorsal portion of eyes straight. Ventral surface of clypeus and gena filled with short setae. Each median tooth with a tubercle on ventral surface. Antennal club coated with sparse and long setae. Thorax: pronotum convex, lacking anteromedian impressions or knobs. Disc shiny, slightly wrinkled and finely punctate. Mid- longitudinal sulcus of pronotum inconspicuous. Transverse carina of hypomera incomplete, not reaching the lateral edge. Surface of mesepimeron and mesanepisternum black, finely striate. Elytra: s triae thin, shallow, indistinctly punctate; striae on disc at least 10 times narrower than the third interstria. Lateral edge of striae finely carinate. Seventh stria absent at the basal one-seventh of elytra. Eighth stria present only at apical two-thirds. Ninth stria present at the apical half of elytra. Interstriae almost flat, shiny and finely punctate. Legs [ventral surface]: profemora punctate on anterior half; posterior half with setose punctures and microgranulation throughout. Outer edge of protibia with 3 acute teeth, basal tooth smaller than median and apical teeth. Mesofemur and metafemur finely punctate. Outer edge of mesotibia with row of erect setae and small teeth. Abdomen: abdominal ventrites II – VI with row of small tubercles along posterior edge. Ventrite V as long as VI. Pygidium: 1 / 3 wider than long, disc shiny, with well-defined microgranulation. Genital capsule: paramera apex divided into apical and lateral lobes by shallow V-shaped emargination. Apical lobe rounded. Lateral lobe truncate, slightly wider than long (Figure 5 (b )). Female: differs from male in abdominal ventrites II – VI smooth along posterior edge. Ventrite V slightly convex medially, with half length of ventrite VI. Pygidial disc shiny, sparsely punctate.	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F419FFFA11F6013058633CC2.taxon	description	(Figures 1 (d), 3 (d), 4 (f), 5 (c) and 6 (a ))	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F419FFF911E0019A5A653C57.taxon	diagnosis	Diagnosis Scatonomus fasciculatus and S. barbatus are closely related. Both species have body longer than 9 mm; dorsal surface shiny, almost smooth; genal edge not distinctly angulate laterally (Figure 3 (a, d )); and clypeal edge bidentate, with deep lateral emarginations filled with a tuft of long setae (Figure 3 (a, d )). However, S. fasciculatus can be distinguished by the following combination of characters: clypeal median teeth elongated, approximately as long as the lateral tuft of setae (Figure 3 (d )); anterior edge of gena slightly straight (Figure 3 (d), arrow); and dorsum with blue or purple reflections (Figure 1 (d )).	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F419FFF911E0019A5A653C57.taxon	description	Redescription Male: length: 9 – 13 mm. Dorsum with black, blue or purple reflections (Figure 1 (d )). Head: densely punctate, slightly wrinkled medially (Figure 3 (d )). Anterior and posterior edges narrowly margined. Clypeo-genal and clypeo-frontal sutures carinate, arched. Clypeal median teeth upturned, separated by U-shaped emargination (Figure 3 (d )). Anterior portion of clypeus strongly concave medially, surface less punctate than remaining portions of head. Clypeal and genal edges with rows of long erect setae. Antennal club globose, densely coated with long setae. Thorax: pronotum shiny, puncticulate, slightly wrinkled. Anteromedian portion of pronotal disc longitudinally wrinkled, and with welldefined microgranulation. Mid-longitudinal sulcus feebly impressed on pronotal disc. Transverse carina of hypomera almost complete, not reaching lateral edge. Anterior portion of hypomera with dense long setae; lateral edge with row of coarse setose punctures, setae short. Surface of mesepimeron filled with dense and fine striae. Surface of mesanepisternum with well-defined microgranulation. Anterior and lateral lobes of metaventrite with well-defined microgranulation, lacking shallow punctures. Disc flat, densely puncticulate. Elytra: striae thin, shallow, indistinctly punctate; width of striae on disc 7 times narrower than the third interstria. Seventh and eighth striae effaced or absent at elytral base. Ninth stria present, weakly impressed at elytral base. Tenth stria inconspicuous. Interstriae plane, shiny, densely punctate, slightly wrinkled. Legs [ventral surface]: profemora densely punctate, surface between punctures smooth. Mesofemora and metafemor sparsely punctate. Outer edge of protibia with 3 distinct teeth separated by deep U-shaped emarginations. Outer edge of mesotibiae and metatibiae with row of short setae and small teeth. Abdomen: posterior half of abdominal ventrites II – VI convex at transverse midline. Ventrite V slightly depressed medially, as long as ventrite VI. Pygidium: 1 / 3 wider than long. Surface shiny, with punctures spaced by 2 times their diameter. Genital capsule: paramera apex divided into apical and lateral lobes by deep Lshaped emargination. Apical lobe rounded; lateral lobe slightly acute (Figure 5 (c )). Female: differs from male by the abdominal ventrite VI twice longer than V. Disc of pygidium with punctures spaced by 2 times their diameter; apical edge of pygidium slightly angulate.	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F41BFFF811830171584A3C1D.taxon	description	(Figures 1 (e), 3 (e), 5 (d) and 6 (b ))	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F41CFFFF11D500C858433D9A.taxon	description	(Figures 1 (f), 3 (f), 5 (e) and 6 (c ))	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F41CFFFE11C507155F2D3A78.taxon	diagnosis	Diagnosis Scatonomus insignis is similar to S. foresti sp. nov. in having the following combination of characters: body length> 13 mm; clypeal edge with two median teeth bounded by lateral emarginations (Figure 3 (e, f); and dorsum coarsely punctate with dark blue to purple reflections (Figure 1 (e, f )). However, S. insignis may be distinguished by the clypeal edge widely margined; clypeal median teeth rounded, separated by a V-shaped emargination (Figure 3 (f )); antennal club densely coated with yellow setae, antennomeres elongate, slightly pointed at apex; clypeal teeth prolonged beyond the remainder clypeal margin (Figure 3 (f )); pronotal disc densely puncticulate, slightly wrinkled; and elytral interstriae densely punctate.	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F41CFFFE11C507155F2D3A78.taxon	description	Redescription Male: Length: 13 – 14.5 mm. Dorsum black, with dark blue or purple reflections (Figure 1 (f )). Head: surface wrinkled and densely punctate; punctures less impressed near anterior edge (Figure 3 (f )). Clypeo-genal suture straight, finely sulcate. Fronto-clypeal suture inconspicuous between eyes. Anterior clypeal edge with a row of dorsal orange setae. Clypeus transversely elevated posteriorly. Ventral surface of clypeus densely setose; setae near the anterior edge visible in dorsal view. Ventral clypeal process produced into a transverse and arched carina. Thorax: pronotum with an anteromedian knob and a pair of shallow impressions positioned more posteriorly to the knob. Pronotal disc densely puncticulate; mid-longitudinal sulcus present on posterior half. Transverse carina of hypomera incomplete, not reaching the lateral edge. Posterior portion of hypomera finely striate, bearing sparse setae along lateral edge. Surface of mesepimeron and mesanepisternum with green reflections, finely striate and shallowly punctate. Anterior lobe of metaventrite smooth. Lateral lobes of metaventrite finely striate and sparsely punctate. Metaventral disc slightly convex, punctate, with shallow mid-longitudinal sulcus posteriorly. Elytra: striae thin, shallow, indistinctly punctate; width of striae on disc at least 10 times narrower than third interstria. Lateral edge of striae slightly carinate. Seventh and eighth striae absent at the basal one-fifth of elytra. Ninth stria carinate on its apical half and narrowly sulcate on remaining parts. Legs [ventral surface]: profemora puncticulate; posterior portion with setose punctures, setae long. Outer edge of protibiae with 2 distinct teeth separated by deep Ushaped emargination; basal tooth inconspicuous. Outer edge of mesotibiae and metatibiae with a distinct row of small teeth. Inner edge of mesotibiae and metatibiae carinate, eventually with few setae on apical third. Abdomen: ventrites II – IV with a row of coarse punctures along posterior portion. Ventrite V slightly shorter than VI at medial portion. Pygidium: disc sparsely puncticulate. Genital capsule: paramera apex divided into apical and lateral lobes by shallow V-shaped emargination. Apical lobe rounded. Lateral lobe truncate, approximately as wide as long; vertical portion of the edge slightly curved inward medially (Figure 5 (e )). Female: differs from male by having the ventral surface of ventrites II – IV with microgranulation on posterior portion, ventrites V – VI sparsely punctate. Disc of pygidium densely puncticulate.	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F41EFFFD11D1027858823F0A.taxon	description	(Figures 1 (g), 3 (g), 4 (b, e, g), 5 (f) and 6 (b ))	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F41EFFFC11C202C25A343F05.taxon	diagnosis	Diagnosis Scatonomus janssensi seems to be closely related to S. viridis viridis and S. mitzae sp. nov., as the three have the clypeal edge with one central emargination, lacking conspicuous teeth (Figure 3 (g, h, l )). However, S. janssensi is distinguished from these related species by the presence of a strongly upturned and widely margined clypeal edge (Figure 3 (g) and 4 (g )); dorsal surface of clypeus divided by anteromedian longitudinal carina (Figure 4 (g), black arrow); posterior portion of clypeus with broad and transverse carina (Figure 4 (g), white arrow); and pronotal disc densely coated with ocellate and fine punctures (Figure 4 (e )).	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F41EFFFC11C202C25A343F05.taxon	description	Redescription Male: Length: 11 – 12 mm. Dorsum black, with green or yellow reflections (Figure 1 (g )). Head: surface coarsely punctate (Figure 3 (g )). Clypeo-genal suture sulcate, almost straight (Figure 3 (g )). Clypeal margination with dense row of orange setae on dorsal portion. Fronto-clypeal suture inconspicuous between eyes. Anterior clypeal and genal edges with a row of short ventral setae, visible in dorsal view. Ventral portion of clypeus and gena with setose punctures. Ventral clypeal process without transverse carina. Antennal club globose, antennomeres enlarged, with sparse yellow setae. Thorax: surface of pronotum with dense ocellate and fine punctures. Pronotal disc with mid-longitudinal sulcus wider medially, lacking ocellate punctures. Transverse carina of hypomera complete, forming a small tooth at the lateral edge. Anterior portion of hypomera densely coated with setose punctures, setae denser on anterior portion and along the lateral edge; posterior portion with row of setae close to posterior edge. Surface of mesepimeron and mesanepisternum with green reflections, surface finely striate and sparsely coated with short setae. Anterior lobe of metaventrite with well-defined microgranulation. Metaventral disc flat, punctate, with shallow mid-longitudinal sulcus. Lateral lobes of metaventrite with well-defined microgranulation and shallow punctures. Elytra: striae thin, indistinctly punctate; on the disc, striae 3 times narrower than the interstriae. Interstriae surface slightly wrinkled, densely punctate and with well-defined microgranulation. Eighth stria as wide as seventh, both striae absent close to elytral base. Ninth stria finely carinate at the basal one-third, not reaching elytral base. Tenth stria present on basal third of elytra. Legs [ventral surface]: profemora densely puncticulate, slightly wrinkled. Mesofemora and metafemora puncticulate. Posterior half of profemora with setose punctures, setae short. Outer edge of protibiae with 3 teeth; apical and median teeth distinct, separated by U-shaped emargination; basal teeth almost inconspicuous. Outer carina of mesotibiae and metatibiae with indistinct teeth and short setae. Abdomen: posterior half of ventrites II – IV with row of shallow punctures. Pygidium: surface barely shiny, puncticulate, surface between punctures smooth near basal edge. Genital capsule: paramera apex divided into apical and lateral lobes by shallow V-shaped emargination. Apical lobe rounded. Lateral lobe truncate, slightly wider than long (Figure 5 (f )). Female: differs from male by having the posterior half of ventrites II – IV lacking punctures. Pygidial disc barely shiny, densely puncticulate, space between punctures with well-defined microgranulation near basal margination.	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F41FFFF3118007D15AA33AE7.taxon	description	(Figures 1 (h), 3 (h), 4 (a, d), 5 (g) and 6 (a ))	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F41FFFF3118007D15AA33AE7.taxon	etymology	Etymology Named in honour of Mileva ‘ Mitza’ Meric Einstein. Mileva was a talented Serbian physicist. She was Albert Einstein’s first wife, and her contributions to his groundbreaking science are hotly discussed.	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F41FFFF3118007D15AA33AE7.taxon	diagnosis	Diagnosis Scatonomus mitzae sp. nov. seems to be closely related to S. viridis viridis and S. janssensi by having a single central emargination on clypeal edge, without teeth or lateral emarginations (Figure 3 (g, h, l )). However, S. mitzae sp. nov. is distinguished by having a deep Ushaped clypeal emargination (Figure 3 (h )). Antennal club densely coated with long setae. Elytral striae thin (Figure 4 (a), arrow); width of striae on disc at least 10 times narrower than the third interstria.	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F41FFFF3118007D15AA33AE7.taxon	description	Description Male: Length: 15.0 – 15.4 mm. Body black, with greenish or yellowish reflections (Figure 1 (h )). Head: surface wrinkled and densely punctate (Figure 3 (h )). Posterior edge narrowly margined medially (Figure 3 (h )). Clypeo-genal suture sulcate, almost straight. Frontoclypeal suture inconspicuous between eyes. Clypeus transversely elevated at posterior portion. Frons almost flat. Clypeal margination upturned. Ventral clypeal process produced into a small tubercle. Antennal club globose. Thorax: pronotal disc with a shallow mid-longitudinal sulcus. Anteromedial portion of the pronotal disc with a low central knob. Transverse carina of hypomera complete, reaching the lateral edge. Anterior portion of hypomera densely coated with setae; lateral edge with a row of setae, visible in dorsal view; posterior portion with short setae near margin of procoxae. Surface of mesepimeron and mesanepisternum with green reflections; finely striate. Meso-metaventral suture straight medially. Anterior lobe of metaventrite with well-defined microgranulation. Metaventral disc flat, punctate, with a shallow longitudinal sulcus medially. Lateral lobes of metaventrite with well-defined microgranulation and shallow punctures. Elytra: striae thin, ill-impressed punctures throughout their length (Figure 4 (a )). Seventh stria absent at the basal one-seventh of elytra. Eighth stria carinate, effaced along anterior half. Ninth stria finely carinate, visible at the apical portion of elytra. Tenth stria absent. Interstriae with microgranulated surface (Figure 4 (a )). Legs [ventral view]: anterior half of profemora densely puncticulate; posterior half with setose punctures. Outer edge of protibia with three weakly defined teeth, shallowly emarginate between teeth. Outer edge of mesotibia and metatibia with a row of small teeth at its basal two-thirds. Outer carina of mesotibia and metatibia with a row of well-defined teeth and setae. Inner margin of metatibia carinate, except by the apical third. Abdomen: surface of ventrites punctate; lateral portion of ventrites I – IV with well-defined microgranulation. Posterior half of ventrites II – IV with a row of shallow punctures. Ventrite V slightly shorter than VI at medial portion. Pygidium: 1 / 3 wider than long. Surface densely puncticulate. Genital capsule: paramera apex divided into apical and lateral lobes by a deep V-shaped emargination. Apical lobe rounded; lateral lobe truncate, approximately as wide as long (Figure 5 (g )). Female: unknown.	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F411FFF2114B03A55A363B20.taxon	diagnosis	Diagnosis Scatonomus barbatus, S. canhedoae sp. nov., S. fasciculatus and S. paulosawayai have deep lateral emarginations filled with setae at the clypeal edge (Figure 3 (a, b, d, i )). However, S. paulosawayai has the clypeal edge lacking median teeth and the median portion between the lateral emarginations with rectangular shape (Figure 3 (i )), dorsum with green or blue reflections (Figure 1 (i )), head and pronotum wrinkled and elytral interstriae opaque, with well-defined microgranulation.	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F411FFF2114B03A55A363B20.taxon	description	Redescription Male: unknown. Female: length: 13 mm. Dorsum with black, green or blue reflections (Figure 1 (i )). Head: surface wrinkled (Figure 3 (i )). Clypeo-genal and fronto-clypeal sutures wide, sulcate. Anterior clypeal and genal edges with rows of ventral, orange setae (Figure 3 (i )). Antennal club filled with short setae. Thorax: pronotal surface wrinkled and puncticulate. Pronotal disc slightly elevated at the anteromedian portion. Mid-longitudinal sulcus on pronotal disc filled with fine punctures. Transverse carina of hypomera complete, reaching the lateral edge. Anterior and lateral edges of hypomera densely coated with setae. Surface of mesepimeron and mesanepisternum with green reflections, finely striate. Disc of metaventrite flat, punctate, with shallow longitudinal sulcus medially. Lateral lobes of metaventrite with well-defined microgranulation and sparse shallow punctures. Elytra: striae thin, punctures shallowly impressed. Seventh stria almost reaching the elytral base. Eighth stria absent at anterior one-fifth. Ninth stria carinate, inconspicuous near basal portion of elytra. Interstriae punctate, with well-defined microgranulation. Legs [ventral surface]: profemora puncticulate, posterior edge with setose punctures near trochanter. Outer edge of protibiae with two acute teeth, apical tooth longer than median and basal teeth. Meso- and metafemora finely punctate. Outer carina of mesotibiae and metatibiae with row of small teeth. Inner edge of mesotibiae and metatibiae carinate, except by its apical one-third. Abdomen: posterior half of ventrites II – IV slightly depressed. Ventrite V 3 times shorter than ventrite VI. Pygidium: shiny and puncticulate.	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F411FFF2112103C3582D3EAE.taxon	description	(Figures 1 (i), 3 (i) and 6 (d ))	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F412FFF111E403C3588A3EAE.taxon	description	(Figures 1 (j, k), 3 (j, k), 4 (c, i, j), 5 (h) and 6 (d ))	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F412FFF0110E03A55A013E9F.taxon	diagnosis	Diagnosis Scatonomus thalassinus resembles S. insignis and S. mitzae sp. nov. in having clypeal edge with 3 emarginations (Figure 3 (f, h, j, k )) and dorsum with coarse punctures (Figure 1 (f, h, j, k )). However, S. thalassinus is distinguished from these two species by the dorsum with green metallic colouration (Figure 1 (j, k )); clypeal edge moderately margined with median teeth acute, separated by a deep V-shaped emargination (some specimens have clypeal edge widely margined with margination produced into 4 rounded teeth, separated by Ushaped emarginations; see taxonomic remarks) (Figure 3 (j, k); abdominal ventrites II – IV usually with a row of wrinkles on posterior edge (Figure 4 (i )).	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F412FFF0110E03A55A013E9F.taxon	description	Redescription Male: Length: 10 – 13 mm. Dorsum metallic green, usually with red or yellow reflections on head and pronotum (Figure 1 (j, k )). Head: dorsal surface barely shiny, with dense coarse punctures, except on anterior portion of clypeus with sparse punctures. Fronto-clypeal junction smooth medially (Figure 3 (j, k )). Clypeal edge distinctly margined (Figure 3 (j, k )). Clypeo-genal suture sulcate, slightly curved outward on posterior half. Ventral clypeal process transverse, produced into an arched carina. Anterior clypeal and genal edges with a row of ventral setae, visible in dorsal view. Thorax: pronotal disc coarsely punctate, lacking wrinkles; mid-longitudinal sulcus smooth, broader medially. Transverse carina of hypomera incomplete, not reaching lateral edge. Anterior portion of hypomera finely striate, with erect yellow setae; setae on lateral edge visible in dorsal view. Posterior portion of hypomera smooth. Lateral edge of hypomera with sparse setae. Surface of mesepimeron and mesanepisternum coarsely punctate, slightly microsculptured. Surface of disc and anterior lobe of metaventrite shallowly punctate; lateral lobes with ill-defined microgranulation and sparse coarse punctures. Elytra: striae thin, ill-impressed, inconspicuously punctate; on disc at least 10 times narrower than third interstria. Lateral edges of striae finely carinate. Seventh and eighth striae absent on basal one-fifth of elytra. Eighth stria carinate. Ninth stria visible only at the posterior half of elytra. Tenth stria absent. Interstriae shiny, filled with dense shallow punctures; in some individuals interstriae barely shiny with well-defined microgranulation. Legs [ventral surface]: profemora puncticulate. Posterior edge of profemora with setose punctures near trochanter. Meso- and metafemora finely punctate. Outer edge of protibiae with 3 teeth, apical and median teeth well developed, separated by a deep U-shaped emargination; basal teeth small, separated from median tooth by shallow V-shaped emargination. Outer edge of meso- and metatibiae with row of small teeth on basal two-thirds; apical third carinate. Inner edge of meso- and metatibiae carinate, except by the apical third. Abdomen: ventrite V approximately 2 times shorter than ventrites II – IV and VI at medial portion. Ventrites V – VI flat medially. Ventrites II – IV wrinkled on posterior edge. Pygidium: surface barely shiny, with sparse punctures and well-defined microgranulation. Genital capsule: paramera apex divided into apical and lateral lobes by shallow V-shaped emargination. Apical lobe rounded. Lateral lobe widely rounded (Figure 5 (h )). Female: differs from the male by abdominal ventrites II – IV only finely punctate on posterior half, and ventrites V – VI convex medially.	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F414FFF611C303B759933EB8.taxon	diagnosis	Diagnosis Scatonomus viridis viridis, S. janssensi and S. mitzae sp. nov. are distinguished from other species of the genus by having a single central emargination on clypeal edge, without teeth or lateral emarginations (Figure 3 (g, h, l )). Scatonomus viridis viridis is distinguished from S. janssensi by the clypeal edge rounded, projected forward, neither widely margined nor strongly upturned (Figure 3 (l )); and dorsal surface of clypeus lacking anteromedian longitudinal carina. In comparison with S. mitzae sp. nov., S. viridis viridis differs by having a shallow central clypeal emargination (Figure 3 (l )); antenal club coated with very short setae; elytral striae slightly narrowed, lateral margins distinctly carinate, width of striae on disc at least 7 times narrower than the third interstria.	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F414FFF611C303B759933EB8.taxon	description	Redescription Male: length: 11 – 12 mm. Dorsum black, with greenish or reddish reflections (Figure 1 (l )). Head: densely punctate, wrinkled (Figure 3 (l )). Anterior edge narrowly margined (Figure 3 (l )). Clypeo-genal suture sulcate, almost straight. Fronto-clypeal suture weakly defined between eyes. Clypeus with a transverse elevation at posterior portion. Two shallow impressions on interocular space. Anterior clypeal and genal edges with a row of sparse ventral setae. Ventral surface of clypeus densely punctate, wrinkled and glabrous. Thorax: surface of pronotal disc wrinkled, densely puncticulate. Lateral portions of pronotal disc with ocellate punctures. Anteromedian portion of pronotum with a knob and a pair of shallow impressions positioned posteriorly to the knob. Transversal carina of hypomera complete, reaching the lateral edge. Surface of mesepimeron and mesanepisternum with green or dark blue reflections, finely striate. Mesepimeron with sparse and short setae. Anterior lobe of metaventrite with well-defined microgranulation. Metasternal disc flat, punctate, with a distinct mid- longitudinal sulcus. Lateral lobes of metaventrite with welldefined microgranulation and shallow punctures. Elytra: striae shallow, feebly punctate. Seventh stria absent at the basal one-seventh of elytra. Eighth stria carinate, interrupted on basal half, absent near the anterior portion of elytra. Ninth stria carinate, present only at the apical fifth of elytra, punctate on its apical half. Tenth stria carinate, present at medial portion of elytra. Interstriae with punctate and microgranulated surface. Legs [ventral surface]: profemora densely puncticulate, surface between punctures slightly wrinkled. Mesofemur and metafemur sparsely punctate. Outer edge of protibia with 3 indistinct teeth separated by feeble U-shaped emarginations. Outer edge of mesotibia and metatibia with a row of coarse teeth. Outer carina of mesotibia and metatibia with weakly defined teeth, glabrous. Inner edge of metatibia carinate, except by apical onethird that bears few setae. Abdomen: ventrites sparsely punctate, ventrites I – V with welldefined microgranulation on anterior half. Posterior half of ventrites II – IV with a row of coarse, transversely elongated punctures. Ventrite V slightly shorter than ventrites II – IV and 2 times shorter than ventrite VI. Pygidium: shiny, slightly wrinkled, densely puncticulate. Genital capsule: paramera apex divided into apical and lateral lobes by a deep Vshaped emargination. Apical lobe slightly acute. Lateral lobe truncate, slightly wider than long (Figure 5 (j )). Female: differs from male by posterior half of ventrites II – IV lacking row of coarse, transversally elongated punctures.	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F414FFF711FC03C358453EB9.taxon	description	(Figures 1 (l), 3 (l), 5 (i) and 6 (a ))	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F415FFF610C001EB589D3A13.taxon	description	Pereira and Martínez (1959) described S. viridis walzi based on three females from Contrera, Paraguay. According to the original description, the subspecies may be distinguished from S. viridis viridis by the clypeus with a shallower central emargination, shorter clypeal teeth and sparser microsculpture on the head, pronotum and elytra. During the course of our revision, we could examine the female holotype of S. viridis walzi only through a low-quality photo of the dorsal habitus of the specimen. Pereira and Martínez (1959) stated that one paratype of their subspecies was deposited in the MZSP collection, but we could not find this specimen during our visit to the museum. Due to the limited material available for a more comprehensive comparative study of the subspecies comprising S. viridis, we opted to maintain the current taxonomic arrangement, in which S. viridis is divided into two subspecies, until more material becomes available for future studies.	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
C4290E04F416FFF510C0020F5A5C3D1F.taxon	description	In their original description, Vulcano and Pereira (1973) described (and illustrated) S. lauropalui with three teeth on the apical third of the protibial edge, with all teeth directed forward. The authors used these characteristics to assign the new species to the genus Scatonomus. However, after the examination of the holotype, we agree that the protibial teeth are positioned at the apical half of the external edge, and only the apical tooth is directed forward. This finding means that this species belongs to Anomiopus instead of Scatonomus. Therefore, we propose the new combination Anomiopus lauropalui (Vulcano and Pereira 1973). Within Anomiopus, this species is easily distinguished from any other by two tufts of setae on the interocular space (Vulcano and Pereira 1973, Fig 2). Canhedo (2006) divided the species of Anomiopus into three species groups, and A. lauropalui best fits the diagnosis of the group smaragdinus, in which the species can be arranged based mainly on the absence of a tooth transversely inserted on the dorso-lateral surface of metatibiae, and by having at least the first metatarsomere emarginate at the apex (Canhedo 2006).	en	Valois, Marcely C., Vaz-de-Mello, Fernando Z., Silva, Fernando A. B. (2020): A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 54 (17 - 18): 1081-1114, DOI: 10.1080/00222933.2020.1781947, URL: http://dx.doi.org/10.1080/00222933.2020.1781947
