taxonID	type	description	language	source
CB52563B9D59CA48FF7CFF62FBD8FF40.taxon	description	Figs 1 – 80	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D59CA48FF7CFF62FBD8FF40.taxon	materials_examined	Type species. Hesperia aquilina Plötz, 1882, by designation of ICZN (1967). Taxonomic history. The genus Artines was described by Godman (1901), who designated Thracides aepitus Geyer, 1832 as the type-species, and provided figures of the venation of the forewing and the male genitalia of a specimen that he thought belonged to this species. However, based on the illustrations provided, this specimen corresponds to Hesperia aquilina Plötz, 1882. Godman (1901) further provided the description of what he considered a new species, naming it Artines atizies, and illustrating the genitalia and the ventral surface of the wings. However, this species is a synonym of Thracides aepitus, as interpreted by Evans (1955). Thus, Godman (1901) had two species, H. aquilina, mistakenly identified by him as T. aepitus and T. aepitus, mistakenly interpreted by him as a new species. The type species of Artines was selected on the basis of an identification error, which led Hemming (1964, 1965) to suggest the alteration of its type species to Hesperia aquilina, an alteration later formalized by the ICZN (1967). In addition to Godman (1901), the authors who provided descriptions of species in Artines were Mabille (1904), Draudt (1923), Evans (1955), Mielke (1968, 1992), Steinhauser & Austin (1993) and Dolibaina et al. (2015). The others provided citations in catalogs and taxonomic comments.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D59CA48FF7CFF62FBD8FF40.taxon	diagnosis	Diagnosis. Artines differs from the other genera of Moncini by the third segment of the labial palpus long, erect and pointed; antenna longer than 1 / 2 length of costa; nudum with 11 to 12 segments, only on apiculus; male with two androconial organs: a sagitated brand on base of CuA 1 – CuA 2 of forewing (absent in the “ aepitus group ”) (Fig. 3) and a tuft of elongated scales along 2 A on dorsal hind wing (Fig. 4); ventral forewing with a violet sheen at apical and costal areas; ventral hind wing with a central black spot at distal end of the discal cell (absent only in A. bipunctata O. Mielke, 1968), distally surrounded by six blue to violet discal spots between Rs – CuA 2, which are externally limited by an arched postdiscal band, from lower 1 / 2 of Sc + R 1 – Rs to upper 1 / 2 of CuA 2 – 2 A; aedeagus with a ventral projection; opening of ejaculatory bulb at the near-dorsal proximal margin; opening of the aedeagus distal; cornuti present; lamella postvaginalis bifid; ostium at the central portion of the sterigma, ventrally projected by folds of the lamellae ante and postvaginalis; ductus bursae long and narrow, first folded in spiral and distally arched to the left.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D59CA48FF7CFF62FBD8FF40.taxon	description	Description. Male. Head: dorsally dark brown, with some white and elongated scales at center of vertex, on transfrontal and fronto-clypeal sutures and on the paraocular area; gena white. Labial palpus brown, densely speckled with white scales, darker at apex; first segment short; second segment twice as long as first; third segment about 2 / 3 of second, erect and pointed. Antenna 1 / 2 length of costa, brown dorsally, white ventrally at base of antennomeres and club, cream at apiculus; club 2 / 3 of the shaft lengh; nudum with 11 to 12 segments, only on apiculus. Thorax: dorsally dark brown; ventrally grayish. Legs externally brown, internally white, somewhat cream on tibiae and tarsi; foretibia with epiphysis; midtibia with spines and a pair of apical spurs; hind tibia with a pair of median and apical spurs. Forewing length: 12 – 15.5 mm. Forewing triangular; anterior margin slightly convex; apex obtuse and rounded; outer margin convex, more projected at the end of M 2 and M 3; tornus rounded; anal margin straight. Hindwing rounded; anterior margin convex; apex rounded; outer margin convex, slightly concave from CuA 2 to near tornus; tornus rounded; anal margin straight. Dorsal forewing: ground color dark brown with yellow scales scattered on costal area and proximal 1 / 3 of CuA 2 – 2 A; two groups of white spots: one apical, consisting of three opaque and minute spots in R 3 – R 4, R 4 – R 5 and R 5 – M 1, another discal, consisting of two larger, opaque to semi-hyaline spots, one smaller at the end of the proximal 1 / 3 of M 3 – CuA 1, the other below dci in CuA 1 – CuA 2; there may be a small, diffuse yellowish spot in CuA 2 – 2 A; brand, when present, sagitated, located at the base of CuA 1 – CuA 2 and with two distal projections, the upper one longer than the lower one (Figs 3, 21 – 26); fringe light tan, darker at end of veins. Dorsal hind wing: ground color as on the dorsal forewing; tufts of elongated scales along 2 A (Fig. 4); fringes white to light brown. Ventral forewing: ground color as on dorsal surface, paler on anal area, sometimes with large, diffuse white spot on distal 1 / 2 of CuA 2 – 2 A; costal area violet with gray to yellow scattered scales; three to four bluish white apical spots in R 2 – R 3, R 3 – R 4, R 4 – R 5 and R 5 – M 1, first one displaced to base; discal spots as on dorsal surface; apical area with two violet bands separated by a ground color stripe, one submarginal between R 4 – CuA 2 (less evident in CuA 1 – CuA 2) and another apical between R 3 – CuA 2; marginal line dark brown: fringes as on dorsal surface. Ventral hind wing: ground color as on ventral forewing, appearing violet and with grayish scales scattered on basal 1 / 2, between costal margin and upper 1 / 2 of CuA 2 – 2 A, and between 3 A and anal margin; paler area between lower 1 / 2 of CuA 2 – 2 A and 3 A; apical portion of discal cell dark brown with yellow to orange scales; discal area with central black spot separated from discal cell by bluish band on dcm and dci (absent in A. bipunctata); six blue to violet discal spots between Sc + R 1 – CuA 2 surrounded by a ground color border, the spot in Sc + R 1 – Rs shifted to base; postdiscal band yellow or gray from lower 1 / 2 of Sc + R 1 – Rs to the upper 1 / 2 of CuA 2 – 2 A, limited distally by a stripe with the same color as ground, and externally to this, a violet submarginal band between Rs and anterior 1 / 3 of CuA 2 – 2 A; marginal line as on forewing; fringes as on dorsal surface. Abdomen: dorsally dark brown, white to grayish ventrally, with median dark brown stripe. Genitalia: tegumen more or less quadrate to rectangular; ventral projection of tegumen fused with dorsal projection of saccus, assuming that the boundaries between these structures are at the angle between them. Fenestra absent. Saccus ranging from shorter to longer than tegumen + uncus. Uncus more or less quadrate, rectangular or tapered, simple or bifid. Gnathos divided, not exceeding uncus and with a central membranous area in lateral view. Valva of variable shape; costa narrow, continuous with or separated from ampulla; harpe simple or bifid, dorsally fused or separated from ampulla. Aedeagus with a ventral projection; opening of ejaculatory bulb at dorso-proximal end, sometimes separated from the anterior margin by a short distance; opening of aedeagus distal; cornuti as numerous spines, either over sclerotized plate or directly over membrane (sclerotized plate absent in A. bipunctata). Fultura inferior in dorsal view formed by a pair of stems, one on each side of aedeagus, connected by membranous area (Figs 27 K – 35 K), in lateral view with anterior end simple or bifid. Female. Forewing length: 13.5 – 14 mm. Similar to male, differing in the absence of androconial organs (Figs 3 – 4). Genitalia: tergum VIII with complete spiracular opening (except in A. panama Medeiros & Carneiro sp. nov.). Region between seventh sternite and lamella antevaginalis membranous or sclerotized. Sterigma completely or partially sclerotized; lamella antevaginalis developed, with or without proximo-ventral folds; lamella postvaginalis bifid or with disto-lateral projections, lateral folds represented by a pair of flaps, projected ventral and medially (Fig. 61 A – B). Bursa copulatrix globular; ostium at central portion of sterigma, ventrally projected by folds of the lamellae ante and postvaginalis, surrounded by membranous or sclerotized area; ductus bursae long, membranous or with basal portion sclerotized, first folded in spiral and distally arched to the left, signa represented by two lateral stripes of small spines along the ductus bursae. Papilla analis rectangular, wider than long.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D5FCA4BFF7CFAB3FAE6FF6C.taxon	description	Figs 1, 4 – 13, 27 – 35, 42 – 48, 54 – 60, 66 – 72, 78 – 79 Description. Male and female: forewing with opaque and reduced discal spots; ventral forewing with white, broad and diffuse spot on distal 1 / 2 of CuA 2 – 2 A; ventral forewing with the discal area yellow, containing central black spot, postdiscal band yellow, limited distally by a dark brown not wavy stripe. Male: no brand; saccus triangular; uncus simple and wide, with or without disto-lateral lobes; arms of gnathos with not pointed apex; harpe and ampulla not fused; harpe bifid (except in A. solange Medeiros & O. Mielke sp. nov.), with ventral process of variable size, wide or narrow and cylindrical, and the dorsal process with conical and strongly sclerotized spines (Fig. 27 E); aedeagus with the disto-dorsal expansion triangular (when vesica is not everted it is difficult to observe); ventral projection of aedeagus simple, narrow and long, directed parallel to the longitudinal axis (Fig. 27 J); cornutus as a sclerotized plate on distal end of vesica, containing elongate and setae-like spines, sometimes very thick as in A. bamba Medeiros & O. Mielke sp. nov. (Figs 42 – 48); fultura inferior straight in lateral view, with proximal end bifid (Figs 27 L – 35 L). Female: externally similar to male; region between seventh sternite and lamella antevaginalis sclerotized, forming a rounded plate that covers the opening of the ostium ventrally; lamella postvaginalis completely sclerotized, ranging from more or less triangular to rectangular; lamella antevaginalis containing proximo-ventral folds (Fig. 54 B); lamella postvaginalis bifid, with lateral fold present or absent, strongly sclerotized, proximo-lateral expansions simple or bilobed and with variable length (Fig. 54 B); ductus bursae membranous or with small, irregular and strongly sclerotized plates. Included species: Artines aepitus (Geyer, 1832), Artines panama Medeiros & Carneiro sp. nov., Artines solange Medeiros & O. Mielke sp. nov., Artines delfos Medeiros & Casagrande sp. nov., Artines maya A. Warren & Dolibaina, 2015, Artines litoralis Medeiros & Dolibaina sp. nov., Artines rica Steinhauser & Austin, 1993, Artines liege Medeiros, O. Mielke & Casagrande sp. nov. and Artines bamba Medeiros & O. Mielke sp. nov.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D5DCA44FF7CFA1BFB80F82C.taxon	description	Figs 5, 27, 42, 54, 66, 78	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D5DCA44FF7CFA1BFB80F82C.taxon	description	Cobalus aepitus; Herrich-Schäffer, 1869. Corr. - Blatt zool. - min. Ver. Regensburg 23: 203. Phlebodes aepitus; Butler, 1870. Ent. monthly Mag. 7: 93. Pamphila aepitus; Kirby, 1871. Syn. Cat. Diurn. Lep., p. 604. Hesperia aepitus; Kirby *, 1879. Cat. Diurn. Lep. Hewitson, p. 221. — Plötz, 1882. Stett. ent. Ztg. 44: 34.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D5DCA44FF7CFA1BFB80F82C.taxon	description	(no genus) atizies; Beattie, 1976. Rhop. Direct., p. 68, 88. Artines epitus [sic]; Gernaat; Beckles & Andel, 2012. Butt. Suriname, p. 603. Taxonomic history. Thracides aepitus Geyer, 1832 was described based on an unstated number of males from “ Java ”. Later, the species was relocated to Nisoniades Hübner, [1819] (Westood 1852), Cobalus Hübner, [1819] (Herrich-Schäffer 1869), Phlebodes Hübner, [1819] (Butler 1870), Pamphila Fabricius, 1807 (Kirby 1871) and Hesperia Fabricius, 1793 (Kirby 1879). The type locality published by Geyer (1832) was corrected by Evans (1949) to South America.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D5DCA44FF7CFA1BFB80F82C.taxon	diagnosis	Diagnosis. Artines aepitus differs from the other species of the “ aepitus group ” in the following combination of characters: ventral process of harpe wide, longer than tegumen + uncus and with distal 1 / 3 angled; dorsal process of harpe on proximal margin; sterigma triangular; lamella antevaginalis with triangular proximo-ventral folds; lamella postvaginalis with weakly developed proximo-ventral expansions, slightly bilobed.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D5DCA44FF7CFA1BFB80F82C.taxon	description	Redescription. Head: antenna with nudum bearing 12 segments. Forewing length: 14 – 15 mm. Dorsal forewing: two or three apical spots, spot in R 5 – M 1 slightly displaced to base; discal spot in M 3 – CuA 1 rounded and spot in CuA 1 – CuA 2 arched; spot in CuA 2 – 2 A present or absent. Ventral forewing (Fig. 5 B): three apical white to blue spots, spot in R 2 – R 3 not very conspicuous; discal spots as on dorsal surface, but more conspicuous; apical area with yellow and gray scales concentrated along veins, between R 3 and CuA 2. Ventral hind wing (Fig. 5 B): apical portion of discal cell with few yellow scales; central black spot longer than wide; blue discal spots rounded. Genitalia (Figs 27 A – L, 42): tegumen more or less quadrate, with proximal margin straight and distal margin convex and slightly rounded (Fig. 27 C), in lateral view, shorter than length of gnathos; ventral arm of tegumen longer than dorsal arm of saccus (Fig. 27 A). Saccus as long as tegumen + uncus, base with the same size as lateral margin (Fig. 27 B). Uncus, in dorsal view, rectangular, about one and a half times longer than wide, shorter than tegumen and without disto-lateral projections, distal margin approximately straight (Fig. 27 C). Gnathos as in the other species of the group (Figs 27 A, 27 D). Valva three and a half times longer than wide; sacculus about 1 / 2 width of valva; costa narrow, not contiguous with ampulla; ampulla elongated, shorter than costa, distally directed ventrally; ventral process of harpe wide, longer than tegumen + uncus, distal 1 / 3 angular, apical setae thick and long, in dorsal view narrow and slightly arched inwards; dorsal process of harpe on proximal margin, triangular in dorsal view and narrower than ventral process (Fig. 27 E – F). Aedeagus shorter than valva, moderately curved dorsally; ventral projection distally pointed and slightly curved ventrally (Fig. 27 G – J); posterior end of cornutus broad and triangular, with thin and straight spines (Fig. 42). Female (Fig. 5 C – D). Forewing length: 13.5 – 14 mm. Similar to male. Genitalia (Figs 54 A – B, 66): tergum VIII with complete spiracular opening. Sterigma triangular; lamella antevaginalis with proximo-ventral folds triangular and short; lamella postvaginalis without lateral fold, distal projections narrow, separated medially by a shallow and narrow indentation, proximo-lateral projections weakly developed and bilobed. Ostium narrow, surrounded by membranous areas; ductus bursae without sclerotized plates. Variation. The white spots of forewing vary in shape and size. The discal spots are always present and the apical spots vary from one to two.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D5DCA44FF7CFA1BFB80F82C.taxon	discussion	Comments. The male genitalia of A. aepitus are similar to A. panama sp. nov., differing in the ventral process of harpe with distal 1 / 3 angled and dorsal process on proximal margin. Kirby (1879) recorded this species from the states of Pará and Rio de Janeiro, Brazil. Since this species is from the Amazon, the record from Rio de Janeiro may be correspond to A. litoralis sp. nov.. The record from Pará cannot be confirmed, since species similar to A. aepitus occur in the Amazon. According to Godman (1901), A. atizies occurs in Panama and Espírito Santo, Brazil and according to Evans (1955) it occurs in Pernambuco, Brazil. The record from Panama most likely corresponds to A. panama sp. nov., whereas the records from Brazil may correspond to A. litoralis sp. nov. The other authors who mention the species as occurring in Panama probably followed the identification of the authors cited above (Mabille 1904; Draudt 1923; Bell 1946; Lewis 1973; Garwood & Lehman 2013). The records from Peru need confirmation, since other species with similar wing pattern occur in that region (Bell 1946; Evans 1955; Lamas 1994; Lamas et al. [1997]; Garwood et al. 2007, 2009). Geographic distribution (Fig. 78). The species occurs along the Amazon Rainforest, with records concentrated to the north and west, extending to Trinidad & Tobago. VENEZUELA — Monagas: Jusepín. TRINIDAD & TOBAGO (Godman 1901) — Arima and Diego Martin (Cock 2011). Couva / Tabaquite / Talparo: Caparo and Las Lomas (Cock 2011). Port of Spain (Kaye 1904). Sangre Grande: Cumaca (Cock 2011). GUYANA — Alto Takutu- Alto Essequibo (Godman 1901). SURINAME (Cock 2014) — Brokopondo: Paramaribo (Gernaat; Beckles & Van Andel 2012). Marowijne: Moengo (Williams & Bell 1931). Para (de Jong 1983). FRENCH GUIANA — St-Laurentdu-Maroni: Saül. BRAZIL — Pará. Roraima: Alto Alegre. ECUADOR — Napo (Murray 2000). Imbabura (Evans 1955). PERU — Huánuco: Tingo Maria.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D5DCA44FF7CFA1BFB80F82C.taxon	description	Temporal distribution. This species occurs in February, April, June, July, August and October. According to the bibliography it also occurs in May (Williams & Bell 1931), January, March and December (Cock 2011).	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D5DCA44FF7CFA1BFB80F82C.taxon	biology_ecology	Host plant. Unknown.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D5DCA44FF7CFA1BFB80F82C.taxon	etymology	Etymology. Not informed by the author.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D5DCA44FF7CFA1BFB80F82C.taxon	materials_examined	Type material. Syntypes of Thracides aepitus were not found in the ZMHU, where other Geyer types are deposited. The absence of primary types associated to the great morphological similarity of A. aepitus with the other species of the genus has led to several misidentification which unable the recognition of the diversity of the group. Thus, in order to ensure the stability and correct identification of this name, a male neotype for Thracides aepitus is here designated, referred by Evans (1955) as Artines aepitus. This specimen, illustrated by Warren et al. (2018), is deposited at NHMUK and has the following labels: / Type H. T. / Type / Takutu R [iver], Brit. Guiana. H. Whitely. / ♂ / Sp. figured. / Sp. figured. / Photo’d / B. C. A. Lep. Rhop. Artines atizies, Godm. / Godman-Salvin Coll. 1913. - 2. / 919 /. Two other labels will be added: / Neotypus / Neotypus Thracides aepitus Geyer, 1832 Medeiros, Dolibaina, Carneiro, O. Mielke & Casagrande det. 2018 /. The material studied by Godman (1901) of Artines atizies is deposited in the NHMUK. Given that this name has been proposed for individuals of both sexes from numerous countries, probably including more than one species (see comments section), a lectotype is here designated in order to ensure its stability and correct identification. The syntype male from Guyana, interpreted by Evans (1955) as the type of this taxon, is here referred to as the lectotype, while the other specimens studied by the author are not considered paralectotypes. This specimen is the same as the above-named neotype of Thracides aepitus, to which the following labels will be added: / Lectotypus / Lectotypus Artines atizies Godman, 1901 Medeiros, Dolibaina, Carneiro, O. Mielke & Casagrande det. 2018 /. The neotype and lectotype labels will be sent to the curator of the NHMUK. Examined material. The neotype and lectotype designated above and the following specimens: VENEZUELA — Monagas: Jusepín, 1 male, 1 female, 1. X. 1965, F. Fernandes & C. J. Rosales leg., OM 29.417, OM 30.125 (OM). FRENCH GUIANA — St-Laurent-du-Maroni: Saül, 1 male, 15. VIII. 2015, no collector, DZ 41.873 (DZUP). BRA- ZIL — Roraima: Alto Alegre (Ilha de Maracá), 1 female, 26. XI – 2. XII. 1987, O. Mielke & Casagrande leg., DZ 41.790 (DZUP). Pará: (Rio Cuminá), 1 female, 26. III. 1936, d’Almeida leg., OM 11.915 (OM). PERU — Huánuco: Tingo Maria (Cueva de Las Pavas), 1 male, IV. 2016, Ramirez leg., OM 80.822 (OM).	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D50CA46FF7CF941FC98F848.taxon	description	Figs 6, 28, 43, 55, 67, 78 urn: lsid: zoobank. org: act: 0 AAE 3 E 9 D- 3557 - 48 E 6 - AB 04 - 392 D 7 E 4 EB 494	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D50CA46FF7CF941FC98F848.taxon	diagnosis	Diagnosis. Artines panama sp. nov. differs from the other species of the “ aepitus group ” in the following combination of characters: harpe with ventral process wide, slightly shorter than tegumen + uncus and with distal 1 / 3 dilated and rounded; dorsal process of harpe on median portion; sterigma narrower at base; lamella antevaginalis with proximo-ventral folds more or less rectangular; lamella postvaginalis with proximo-lateral projections weakly developed and not bilobed.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D50CA46FF7CF941FC98F848.taxon	description	Description. Male (Fig. 6 A-B). Head: antenna with nudum bearing 12 segments. Forewing length: 13 mm. Dorsal forewing (Fig. 6 A): three apical spots; discal spots as in A. aepitus, but less conspicuous; white spot in CuA 2 – 2 A present. Ventral forewing (Fig. 6 B): as in A. aepitus, except for paler coloration. Ventral hind wing (Fig. 6 B): apical portion of discal cell with few yellow scales; central black spot rectangular and longer than wide. Genitalia (Figs 28 A – L, 43): tegumen more or less rectangular, with proximal margin straight and distal margin convex and angled (Fig. 28 C), in lateral view, shorter than length of gnathos; ventral arm of tegumen longer than dorsal arm of saccus (Fig. 28 A). Saccus shorter than tegumen + uncus, with base approximately as wide as lateral margin (Fig. 28 B). Uncus, in dorsal view, rectangular, about one and a half times longer than wide, shorter than tegumen, without disto-lateral projections, distal margin rounded (Fig. 28 C). Gnathos as in the other species of the group (Figs 28 A, 28 D). Valva about three times longer than wide; sacculus triangular, more than 1 / 2 width of valva; costa narrow and not contiguous with ampulla; ampulla elongated, shorter than costa, distally directed ventrally; ventral process of harpe wide, slightly shorter than tegumen + uncus, distal 1 / 3 dilated and rounded, with apical setae thick and long (setae lost during preparation), in dorsal view narrow and slightly curved; dorsal process of harpe over median portion, triangular in dorsal view, narrower than ventral process (Fig. 28 E – F). Aedeagus as long as valva, moderately curved dorsally; ventral projection distally wide and weakly curved ventrally (Fig. 28 G – J); posterior end of cornutus wide and triangular, with thin and straight spines (Fig. 43). Female (Fig. 6 C – D). Forewing length: 12 – 13 mm. Similar to male, except for the absence of spot in CuA 2 – 2 A on dorsal forewing. Genitalia (Figs 55 A – B, 67): tergum VIII with spiracular opening incomplete. Sterigma rectangular; lamella antevaginalis with proximo-ventral folds more or less rectangular; lamella postvaginalis without lateral fold and with distal projections narrow, separated medially by a relatively deep indentation, proximo-lateral projections weakly developed and not bilobed. Ostium as in A. aepitus; ductus bursae without sclerotized plate. Variation. No relevant variation was observed.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D50CA46FF7CF941FC98F848.taxon	discussion	Comments. This species is well characterized by the male and female genitalia. It is also the only species of the genus that has been recorded from Panama (Fig. 79). For this reason, all literature references to Artines in that country are here interpreted as referring to A. panama sp. nov. (Huntington 1932; Basset et al. 2015). Given the great similarity with A. aepitus, particularly in the female genitalia, Godman (1901) included two specimens from Panama in the description of A. atizies. According to Godman (1901) those specimens were not well preserved. Given that A. aepitus does not occur in Central America, those specimens probably belong to A. panama sp. nov.. Geographic distribution (Fig. 78). The species occurs in forested areas of Panama. PANAMA — Canal Zone: Farfan, Barro Colorado Island (Huntington 1932).	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D50CA46FF7CF941FC98F848.taxon	description	Temporal distribution. This species occurs in January. There is also in the literature record for November (Huntington 1932).	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D50CA46FF7CF941FC98F848.taxon	biology_ecology	Host plant. Unknown.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D50CA46FF7CF941FC98F848.taxon	etymology	Etymology. This species is named in reference to the type locality.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D50CA46FF7CF941FC98F848.taxon	materials_examined	Type material. Holotype male deposited at the OM – DZUP with the following labels: / HOLOTYPUS / 13. I. 1968, Farfan, C [anal] Zone, Panama, Brown leg. / gen. prep. Mielke 1992 / OM 29.165 / Holotypus Artines panama Medeiros & Carneiro det. 2018 /. Allotype female deposited at the OM – DZUP with the following labels: / ALLOTYPUS / 13. I. 1968, Farfan, C [anal] Zone, Panama, Brown leg. / gen. prep. Mielke 1992 / gen. prep. A. Medeiros 2018 / OM 30.179 / Allotypus Artines panama Medeiros & Carneiro det. 2018 /. Paratype: 1 female, same data as holotype, OM 30.083 (OM).	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D56CA41FF7CFF1AFCC6F83A.taxon	description	Figs 7, 29, 56, 68, 78 urn: lsid: zoobank. org: act: 0 B 439081 - EF 4 F- 41 EB-A 728 - 8 B 7781217566	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D56CA41FF7CFF1AFCC6F83A.taxon	diagnosis	Diagnosis. Artines solange sp. nov. differs from the other species of the “ aepitus group ” in the following combination of characters: ventral forewing with distal portion of discal cell densely covered with orange scales; uncus with disto-lateral lobes; ventral process of harpe wide and longer than tegumen + uncus; dorsal process of harpe absent; sterigma more or less rectangular; lamella postvaginalis with lateral sclerotized folds and large distal projections; ostium rounded, surrounded by sclerotized area. Descripiton. Male (Fig. 7 A – B). Head: antenna with nudum bearing 12 segments.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D56CA41FF7CFF1AFCC6F83A.taxon	description	Forewing length: 14 mm. Dorsal forewing (Fig. 7 A): three apical spots; discal spots reduced to few scales; spot in CuA 2 – 2 A absent. Ventral forewing (Fig. 7 B): as in A. aepitus. Ventral hind wing (Fig. 7 B): apical portion of discal cell densely covered by orange to yellow scales; central black spot as wide as long; blue discal spots elongated. Genitalia (Fig. 29 A – L): tegumen more or less rectangular, with proximal margin concave and distal margin convex and strongly angled (Fig. 29 C), in lateral view larger than length of gnathos; ventral arm of tegumen as long as dorsal arm of saccus (Fig. 29 A). Saccus shorter than tegumen + uncus, base larger than lateral margin (Fig. 29 B). Uncus, in dorsal view, quadrate, shorter than tegumen and with two ventrally curved disto-lateral lobes (Fig. 29 C). Gnathos as in the other species of the group (Fig. 29 A, D). Valva approximately four times longer than wide; sacculus elongated, about 1 / 3 of width of valva; costa rectangular and not contiguous with ampulla; ampulla rectangular, longer than costa and directed distally; ventral process of harpe wide, longer than tegumen + uncus, distal 1 / 3 not dilated and approximately rounded, with apical setae thick and long; dorsal process absent (Fig. 29 E – F). Aedeagus lost during preparation. Female (Fig. 7 C – D). Forewing length: 13 mm. Similar to male. Genitalia (Figs 56 A – B, 68): tergum VIII as in A. aepitus. Sterigma more or less rectangular; lamella antevaginalis without proximo-ventral folds; lamella postvaginalis with lateral fold, distal projections wide and rounded, medially separated by shallow and narrow indentation, proximo-lateral projections weakly developed and not bilobed. Ostium rounded, surrounded by sclerotized area; ductus bursae without sclerotized plates. Variation. No relevant variation was observed.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D56CA41FF7CFF1AFCC6F83A.taxon	discussion	Comments. Artines solange sp. nov. is one of the rarest or the least collected species of the “ aepitus group ”. It is also one of the easiest to recognize based on the male and female genitalia. Even though the aedeagus was lost during preparation, before that the general pattern of the “ aepitus group ” species were observed, particularly the narrow, simple and elongate ventral process. The association with the female was based on the color of the ventral hind wing, such as the distal portion of the discal cell densely covered with orange scales and central black spot as wide as long, besides collecting locality. However, in view of the fact that Artines species are similar and different species can potentially occur in the same locality, the association is tentative and for this reason the only female was not included in the type-series. Geographic distribution (Fig. 78). The species is known only from east of Brazilian Amazon Rainforest. BRAZIL — Maranhão: Imperatriz and Sumaúma.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D56CA41FF7CFF1AFCC6F83A.taxon	description	Temporal distribution. This species occurs in July and August.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D56CA41FF7CFF1AFCC6F83A.taxon	biology_ecology	Host plant. Unknown.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D56CA41FF7CFF1AFCC6F83A.taxon	etymology	Etymology. The name of the new species is in honor to Dr. Solange Maria Kerpel, who has been contributing to the study of butterflies in the Brazilian northeast, in addition to her incentive to the first author in getting start the study on the taxonomy of butterflies. The name is a noun in apposition.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D56CA41FF7CFF1AFCC6F83A.taxon	materials_examined	Type material. Holotype male deposited at the DZUP with the followings labels: / HOLOTYPUS / Imperatriz, MA [ranhão, Brazil], 15. VII. 1974, Exc [ursão] Dep [ar] t [ament] o [de] Zool [ogia] leg., / gen. prep. A. Medeiros 2018 / DZ 41.889 / Holotypus Artines solange Medeiros & O. Mielke det. 2018 /. Additional material (only female). BRAZIL — Maranhão: Sumaúma (formerly a district of Montes Altos), 1 female, 10. VIII. 1974, O. Mielke leg., DZ 41.800 (DZUP).	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D57CA40FF7CFF1AFCA9F9B6.taxon	description	Figs 8, 30, 44, 78 urn: lsid: zoobank. org: act: BFB 4 F 773 - D 8 BC- 439 F-BEC 8 - 3663 BD 8 ABBB 6	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D57CA40FF7CFF1AFCA9F9B6.taxon	diagnosis	Diagnosis. Artines delfos sp. nov. differs from the other species of the “ aepitus group ” in the following combination of characters: valva three times longer than wide; harpe with ventral process wide, shorter than tegumen + uncus, distal 1 / 3 angular; dorsal process of harpe over proximal 1 / 2 and distally directed ventrally.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D57CA40FF7CFF1AFCA9F9B6.taxon	description	Description. Male (Fig. 8 A – B). Head: antenna with nudum bearing 12 segments. Forewing length: 14 mm. Dorsal forewing (Fig. 8 A): two apical spots reduced to few scales; discal spot in M 3 – CuA 1 slightly arched; spot in CuA 2 – 2 A absent. Ventral forewing (Fig. 8 B): a row of four blue apical spots directed toward median portion of outer margin. Ventral hind wing (Fig. 8 B): apical portion of discal cell with few yellow scales; central black spot rectangular, longer than wide. Genitalia (Figs 30 A – L, 44): tegumen more or less rectangular, with proximal margin concave and the distal margin not conspicuous (Fig. 30 C), in lateral view shorter than length of gnathos; ventral arm of tegumen longer than dorsal arm of saccus (Fig. 30 A). Saccus shorter than tegumen + uncus, base same size as lateral margin (Fig. 30 B). Uncus, in dorsal view, rectangular, about twice longer than wide, longer than tegumen, without disto-lateral projections, distal margin rounded (Fig. 30 C). Gnathos as in the other species of the group (Fig. 30 A, D). Valva three times longer than wide; sacculus triangular and more than 1 / 2 width of valva; costa narrow and contiguous with ampulla; ampulla irregular and shorter than costa; ventral process of harpe wide, as long as tegumen, distal 1 / 3 angular, with apical setae thick and long; dorsal process of harpe on proximal 1 / 2, narrower than ventral process, with apex over dorsal margin of ventral process (Fig. 30 E – F). Aedeagus shorter than valva, slightly curved dorsally; ventral projection as in A. aepitus (Fig. 30 G – J); posterior end of cornutus triangular, with distal spines thin and curved laterally (Fig. 44). Female. Unknown. Variation. Not observed.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D57CA40FF7CFF1AFCA9F9B6.taxon	discussion	Comments. Artines delfos sp. nov. is apparently rare or rarely collected. The male genitalia are similar to A. rica and A. liege sp. nov., differing in the ventral process of harpe wider and not cylindrical. Geographic distribution (Fig. 78). The species is known only from the type locality. BRAZIL — Amazonas: Maués.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D57CA40FF7CFF1AFCA9F9B6.taxon	description	Temporal distribution. This species occurs in February.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D57CA40FF7CFF1AFCA9F9B6.taxon	biology_ecology	Host plant. Unknown.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D57CA40FF7CFF1AFCA9F9B6.taxon	etymology	Etymology. Delfos is a Greek city that the ancient Greeks believed to be the center of the universe. The name of the species is in reference to its central geographic distribution in relation to the other species of Artines. The name is a noun in apposition.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D57CA40FF7CFF1AFCA9F9B6.taxon	materials_examined	Type material. Holotype male deposited at the OM – DZUP with the following labels: / HOLOTYPUS / 1 – 15. III. 2008, Rio Preto, Maués, Amazonas, [Brazil], [no collector] / gen. prep. A. Medeiros 2018 / OM 70.129 / Holotypus Artines delfos Medeiros & Casagrande det. 2018 /.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D57CA43FF7CF9F0FDCEFBA0.taxon	description	Figs 9, 31, 78 Artines aepitus (misid.); J. de la Maza E. & R. G. de la Maza E., 1985. Rev. Soc. Mex. Lep. 9: 43. — Llorente-Bousquets; Luis- Martínez & Vargas-Fernández, 1990. Publ. esp. Mus. Zool., México, 1: 35. — J. de la Maza E.; White & R. G. de la Maza E., 1991. Rev. Soc. Mex. Lep. 14: 29. — R. G. de la Maza E. & J. de la Maza E., 1993. Marip. Chiapas, p. 206. — Austin et al., 1999. Trop. Lep. 9, Suppl. 2: 13. — J. de la Maza & R. G. de la Maza, 2015, in Carabias; J. de la Maza & Cadena (eds.). Conserv. desarr. sust. Selva Lacandona, p. 648.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D57CA43FF7CF9F0FDCEFBA0.taxon	diagnosis	Diagnosis. Artines maya is very similar to the other species of the “ aepitus group ”, but its distribution is restricted to Central America where it is potentially sympatric with A. rica and A. panama sp. nov., and from which can be easily distinguished by being relatively smaller; discal area of ventral hind wing with the central black spot wider than long; harpe with ventral process triangular and shorter than tegumen and aedeagus longer than valva. These characters are only shared with A. litoralis sp. nov., from which it differs by the tegumen shorter than uncus; ampulla distally curved ventrally and ventral process of harpe shorter than dorsal. Besides that, A. litoralis sp. nov. occurs only on the Brazilian coast, from Paraíba to Espírito Santo. Female. Unknown.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D57CA43FF7CF9F0FDCEFBA0.taxon	description	Variation. Not observed.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D57CA43FF7CF9F0FDCEFBA0.taxon	discussion	Comments. Artines maya is one of the smallest species of the genus. Although it shares great morphological similarity with A. litoralis sp. nov., these two species are widely separated geographically and after relatively intense sampling between their areas of occurrence, no intermediate form was found. Geographic distribution (Fig. 78). The species is known from lowland forest of Central America (Dolibaina et al. 2015). MEXICO — Chiapas: Lacantún. GUATEMALA — Izabal: Morales. BELIZE — Orangewalk.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D57CA43FF7CF9F0FDCEFBA0.taxon	description	Temporal distribution. This species occurs in March, May and September (Dolibaina et al. 2015).	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D57CA43FF7CF9F0FDCEFBA0.taxon	biology_ecology	Host plant. Unknown.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D57CA43FF7CF9F0FDCEFBA0.taxon	etymology	Etymology. According to Dolibaina et al. (2015) this species was named in reference to the Maya people, former inhabitants of the region where the species occurs.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D57CA43FF7CF9F0FDCEFBA0.taxon	materials_examined	Type material. Holotype male deposited at the MGCL with the following labels: / HOLOTYPUS / Belize: Orangewalk dist [rict]., c [irc] a. 5 Km W. Hill Bank, Mar 4 - 1997 Valerie Giles [leg.] / Genitalia Vial # 98 - 72 Andrew D. Warren / Holotypus Artines maya A. Warren & Dolibaina det. 2015 /. Examined material. The holotype and the following two paratypes: MEXICO — Chiapas: Lacantún (Río Lacantún, Chajul), 148 m, 1 male, 27. V. 1981 (MAZA). GUATEMALA — Izabal: Morales (Cayuga), 35 m, 1 male, IX, Schaus & Barnes leg., (USNM).	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D54CA5DFF7CFB0AFB99FF6C.taxon	description	Figs 10, 32, 45, 57, 69, 79 urn: lsid: zoobank. org: act: 29 D 87 F 1 A-B 981 - 411 A-B 007 - 873037944723 Artines atizies (misid.); Cardoso, 1949. Rev. Ent., Rio de Janeiro, 20: 432. Artines aepitus aepitus (misid.); Kesselring & Ebert, [1982]. Rev. Nordestina Biol. 2: 108.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D54CA5DFF7CFB0AFB99FF6C.taxon	diagnosis	Diagnosis. Artines litoralis sp. nov. is externally similar to the other species of the “ aepitus group ”, differing in the following combination of characters: harpe with ventral process triangular and shorter than tegumen; aedeagus longer than valva; proximo-lateral projections of lamella postvaginalis triangular and not bilobed; occurrence restricted to the Brazilian coast, from Paraíba to Espírito Santo.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D54CA5DFF7CFB0AFB99FF6C.taxon	description	Description. Male (10 A – B). Head: antenna with nudum bearing 12 segments. Forewing length: 12 – 15 mm. Dorsal forewing (Fig. 10 A): apical spots generally absent; discal spots reduced or absent; spot in CuA 2 – 2 A absent or represented by a few scales. Ventral forewing (Fig. 10 B): three reduced blue to white apical spots; discal spots as in the other species, but smaller. Ventral hind wing (Fig. 10 B): apical portion of discal cell with few yellow scales; central black spot rectangular, wider than long. Genitalia (Figs 32 A – L, 45): tegumen more or less quadrate, with proximal margin straight and distal margin convex, slightly rounded (Fig. 32 C), in lateral view, shorter than length of gnathos; ventral arm of tegumen longer than dorsal arm of saccus (Fig. 32 A). Saccus as long as tegumen + uncus, base shorter than lateral margin (Fig. 32 B). Uncus, in dorsal view, rectangular, twice longer than wide, as long as tegumen and without disto-lateral projections, distal margin rounded (Fig. 32 C). Gnathos as in the other species of the group (Fig. 32 A, D). Valva twice longer than wide; sacculus triangular reaching 1 / 2 width of valva; costa narrow and contiguous with ampulla; ampulla elongated and shorter than costa; ventral process of harpe short and triangular, shorter than tegumen and slightly longer than dorsal process, with apical setae thin and long; dorsal process of harpe on distal 1 / 2, rounded in dorsal view and wider than ventral process (Fig. 32 E – F). Aedeagus longer than valva, moderately curved dorsally; ventral projection distally wide, in lateral view slightly curved ventrally (Fig. 32 G – J); cornutus similar to A. delfos sp. nov. but narrower and less corrugated (Fig. 45). Female (Fig. 10 C – D). Forewing length: 13 – 14 mm. Similar to male, differing in the more conspicuous marks on dorsal forewing. Genitalia (Figs 57 A – B, 69): tergum VIII with complete spiracular opening. Sterigma triangular; lamella antevaginalis with proximo-ventral folds elongated; lamella postvaginalis without lateral fold, with distal projections wider than in A. aepitus and separated medially by narrow and shallow indentation, proximo-lateral projections triangular and not bilobed. Ostium narrow, surrounded by membranous areas; ductus bursae without sclerotized plates. Variation. The central black spot on ventral hind wing is longer than wide in specimens from Pernambuco and Paraíba, Brazil.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D54CA5DFF7CFB0AFB99FF6C.taxon	discussion	Comments. Artines litoralis sp. nov. is the only species of the “ aepitus group ” which occurs in the low areas of the Brazilian coast, penetrating inland along the Rio Doce Valley, in the states of Espírito Santo and Minas Gerais. The farthest record from the coast is Paraopeba, Minas Gerais. There the species is potentially sympatric with A. liege sp. nov., from which it can be easily distinguished by the male and female genitalia. Geographic distribution. (Fig. 79). The species is restricted to the Atlantic Forest. BRAZIL — Paraíba: João Pessoa. Pernambuco: Camaragibe, Recife, São Lourenço da Mata. Alagoas: Maceió. Espírito Santo: Baixo Guandu, Conceição da Barra, Itaguaçu, Linhares, Santa Teresa (Brown Jr & Freitas 2000).	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D54CA5DFF7CFB0AFB99FF6C.taxon	description	Temporal distribution. This species occurs throughout the year, except in November.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D54CA5DFF7CFB0AFB99FF6C.taxon	biology_ecology	Host plant. Unknown.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D54CA5DFF7CFB0AFB99FF6C.taxon	etymology	Etymology. This species is named in reference of its distribution along the Brazilian coast.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D54CA5DFF7CFB0AFB99FF6C.taxon	materials_examined	Type material. Holotype male deposited at the DZUP with the following labels: / HOLOTYPUS / Linhares, Espírito Santo, Brazil, 17. V. 1972, C. Elias [leg.] / DZ 41.870 / Holotypus Artines litoralis Medeiros & Dolibaina det. 2018 /. Allotype female deposited at the DZUP with the following labels: / ALLOTYPUS / 8 – 9. III. 1994, P [ar] q [ue] Est [adual] Rio Doce, Marliéria, M [inas] G [erais, Brazil], 350 m, Mielke leg. / DZ 41.880 / Allotypus Artines litoralis Medeiros & Dolibaina /. Paratypes: BRAZIL — Paraíba: João Pessoa, 1 female, 24. VII. 1954, Kesselring leg., DZ 41.750, 1 female, 4. VI. 1992, Tangerini leg., DZ 41.809 (DZUP). Pernambuco: Camaragibe, 1 female, 18. VIII. 1957, 1 female, 27. IV. 1958, 1 female, 7. VII. 1958, 1 female, 18. XIII. 1958, 2 females, 7. IX. 1959, 2 males, 2. I. 1960, 1 female, 29. X. 1960, H. Ebert leg., DZ 39.886, DZ 39.903, DZ 39.892, DZ 39.887, DZ 39.890, DZ 39.899, DZ 41.780, DZ 39.884, DZ 39.902 (DZUP); Recife, 1 male, 3. V. 1959, 1 male, 1. IV. 1961, H. Ebert leg., DZ 39.893 (DZUP), OM 10.342 (OM); São Lourenço da Mata, 1 female, 26. VII. 1972, Kesselring leg., DZ 39.896, 1 female, 27. XII. 1961, H. Ebert leg., DZ 39.900 (DZUP). Alagoas: Maceió, 1 female, 11. VIII. 1973, Cardoso leg., DZ 41.770 (DZUP). Espírito Santo: Baixo Guandu, 1 male, 15. IV. 1970, 1 female, 25. IV. 1970, 1 male, 1 female, 4. X. 1971, 1 female, 1 – 10. IV. 1971, C. & C. T. Elias leg., DZ 41.891, DZ 39.885, DZ 41.779, DZ 41.889, DZ 41.839 (DZUP); Conceição da Barra, 1 male, 1 female, 10. IV. 1968, 1 male, 12. IV. 1969, 1 male, 5. V. 1969, 1 male, 14. V. 1969, 1 male, 5. VI. 1969, 1 male, 18. VI. 1969, 1 female, 1. VIII. 1969, 3 males, 1 female, 10. IX. 1969, C. & C. T. Elias leg., DZ 41.902, DZ 41.829, DZ 4.062, DZ 41.801, DZ 41.751, DZ 41.912, DZ 41.761, DZ 39.888, DZ 41.771, DZ 41.781, DZ 41.791, DZ 39.891 (DZUP); Itaguaçu, 1 male, 18. IV. 1970, 1 male, 15. VI. 1970, 1 male, 29. VII. 1970, C. Elias leg., DZ 39.883, DZ 39.880, DZ 41.759 (DZUP); Linhares, 1 male, IV. 1970, 1 male, 11. II. 1972, 1 male, 4. IV. 1972, 1 female, 10. V. 1972, 1 male, 17. V. 1972, 1 male, 22. VI. 1972, 1 male, 20. IX. 1972, 1 male, 22. III. 1973, 1 male, 27. IV. 1973, 1 male, 16 – 22. V. 1973, 1 male, 1 – 9. VI. 1973, 1 male, 1 – 8. IX. 1973, 1 male, 17 – 22. IX. 1973, 1 female, 1 – 8. X. 1973, 2 males, 2. V. 1974, 1 male, 2 – 7. IX. 1974, 1 male, 23 – 30. IX. 1974, C. Elias leg., DZ 41.841, DZ 41.911, DZ 41.851, DZ 41.760, DZ 39.881, DZ 41.821, DZ 41.205, DZ 41.861, DZ 41.811, DZ 41.215, DZ 41.882, DZ 41.769, DZ 41.831, DZ 41.820, DZ 41.901, DZ 41.881, DZ 41.871, DZ 41.810 (DZUP).	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D4ACA5DFF7CFECEFB97F9C5.taxon	description	Figs 11, 33, 46, 58, 70, 78	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D4ACA5DFF7CFECEFB97F9C5.taxon	diagnosis	Diagnosis. Artines rica differs from the other species of the “ aepitus group ” in the following combination of characters: harpe with ventral process narrow and cylindrical, shorter than tegumen + uncus, with setae on apical portion; cornutus with thin and straight spines, directed posteriorly; proximo-ventral folds of lamella antevaginalis similar to A. aepitus, but less developed, distal projections of lamella postvaginalis separated by deeper cleft; distribution restricted to Central America.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D4ACA5DFF7CFECEFB97F9C5.taxon	description	Variation. According to Steinhauser & Austin (1993) there is variation in the presence or absence of discal and apical marks on forewing.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D4ACA5DFF7CFECEFB97F9C5.taxon	discussion	Comments. Steinhauser & Austin (1993) described Artines rica and mentioned some specimens from Rondônia and Distrito Federal, Brazil. They did not designate any of them as paratypes. The identification of the specimen from Distrito Federal is dubious because its genitalia, according to the authors, is damaged. The specimens from these localities studied by us belong to A. liege sp. nov. Therefore, A. rica probably does not occur in South America. Geographic distribution (Fig. 78). The species occurs in forested areas of Central America. COSTA RICA — Cartago: Turrialba. Guanacaste: Guanacaste Conservation Area (Janzen & Hallwachs 2009).	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D4ACA5DFF7CFECEFB97F9C5.taxon	description	Temporal distribution. This species occurs in April, June and July.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D4ACA5DFF7CFECEFB97F9C5.taxon	biology_ecology	Host plant. Olyra caudata and Olyra latifolia (Poaceae) (Janzen & Hallwachs 2009).	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D4ACA5DFF7CFECEFB97F9C5.taxon	etymology	Etymology. According to Steinhauser & Austin (1993) the term " rica " refers to the type locality of the species.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D4ACA5DFF7CFECEFB97F9C5.taxon	materials_examined	Type material. Holotype male deposited at the MGCL (ex AME) with the following labels: / HOLOTYPE Artines rica, S. R. Steinhauser & G. T. Austin / COSTA RICA: CARTAGO, Turriable [sic!] l. vi. 1972, H. L. King / SRS Database, Nº 568 / Allyn Museum Photo / No. 890626 - 19.20 / 890719 - 21, 22 / A. C. Allyn Ace. 1972 - 5 / MGCLFLMNH Specimen nº 47907 /. Examined material. Illustration of the holotype and the following specimens: COSTA RICA — Cartago: Turrialba, 1 female, 5. VII. 1971, 3 males, 1 female, 15 – 30. IV. 1973, Becker leg., DZ 31.561, DZ 4.061, DZ 41.890, DZ 30.956, DZ 41.840 (DZUP), 600 m, 1 male, 29. VI. 1973, Becker leg., DZ 8.870 (DZUP).	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D4ACA5FFF7CF927FDCBF9C4.taxon	description	Figs 12, 34, 47, 59, 71, 79 urn: lsid: zoobank. org: act: 49 C 3626 C-F 581 - 47 A 2 - 94 BD-BA 021 BFFCC 70	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D4ACA5FFF7CF927FDCBF9C4.taxon	description	Artines rica (misid.); Garcia-Salik et al., 2014. Shilap Revta. lepid. 42 (166): 275. Taxonomic history. Artines liege sp. nov. was previously identified as A. aepitus, A. atizies and A. rica by several authors (see catalog).	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D4ACA5FFF7CF927FDCBF9C4.taxon	diagnosis	Diagnosis. Artines liege sp. nov. differs from the species of the “ aepitus group ” in the following combination of characters: harpe with ventral process narrow and cylindrical, shorter than tegumen + uncus and with ventral margin concave; cornutus with thick distal spines, curved laterally; sterigma similar to A. aepitus differing in the proximoventral folds of lamella antevaginalis narrower and elongated; distribution restricted to South America.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D4ACA5FFF7CF927FDCBF9C4.taxon	description	Description. Male (Fig. 12 A – B). Head: antenna with nudum bearing 12 segments. Forewing length: 12 – 14 mm. Dorsal forewing (Fig. 12 A): apical spots generally absent or represented by a few scales; discal spots reduced or absent; spot in CuA 2 – 2 A absent or represented by a few scales. Ventral forewing (Fig. 12 B): as in A. litoralis sp. nov. Ventral hind wing (Fig. 12 B): apex of discal cell with few yellow scales; central black spot very reduced or wider than long. Genitalia (Figs 34 A – B, 47): tegumen more or less quadrate, with proximal margin sinuous and distal margin convex (Fig. 34 C), in lateral view, shorter than length of gnathos; ventral arm of tegumen longer than dorsal arm of saccus (Fig. 34 A). Saccus longer than tegumen + uncus, base shorter than lateral margin (Fig. 34 B). Uncus, in dorsal view, rectangular, twice longer than wide, as long as tegumen and without disto-lateral projections, distal margin rounded (Fig. 34 C). Gnathos as in the other species of the group (Fig. 34 A, D). Valva three times longer than wide; sacculus more than 1 / 2 width of valva; costa rectangular not contiguous with ampulla; ampulla narrow and as long as costa; ventral process of harpe narrow, cylindrical, longer than tegumen + uncus and slightly arched inwards in dorsal view, with apical setae thick and long and ventral margin convex; dorsal process on median portion, rounded in dorsal view and wider than ventral process (Fig. 34 E – F). Aedeagus as long as valva, strongly curved dorsally; ventral projection distally wide and slightly curved ventrally (Fig. 34 G – J); posterior end of cornutus rectangular, with thick distal spines, curved laterally (Fig. 47). Female (Fig. 12 C – D). Forewing length: 12 – 15 mm. Similar to male, differing in the white marks on dorsal forewing more conspicuous (almost all female paratypes with three apical spots on dorsal forewing). Genitalia (Figs 59 A – B, 71): tergum VIII with spiracular opening complete. Sterigma triangular; lamella antevaginalis with proximo-ventral folds narrow and elongated; lamella postvaginalis without lateral fold and with distal projections similar to A. aepitus, proximo-lateral projections weakly developed and weakly bilobed. Ostium narrow, surrounded by membranous areas; ductus bursae without sclerotized plate. Variation. The central black spot on the ventral hind wing is longer than wide in the specimens from Maranhão, Pará and Rondônia and some specimens from Mato Grosso. In the male genitalia, the ventral process of harpe is shorter in specimens from Brasília, more elongated in the specimens from Mato Grosso and slightly wider in the specimens from Rondônia.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D4ACA5FFF7CF927FDCBF9C4.taxon	discussion	Comments. The male genitalia of Artines liege sp. nov. is similar to A. rica, from which it differs in the concave ventral margin of harpe, ventral process of harpe more elongated and dorsal process narrower, leaving more space between them, and cornutus with thick distal spines, laterally curved. These genitalic characters are constant throughout the distribution of A. liege sp. nov., and besides, A. rica is restricted to Central America. Geographic distribution (Fig. 79). This species is widely distributed in the Brazil, Paraguay and Argentina, occurring from eastern Amazon Rainforest throughout the deciduous and semideciduous forests of the western limits of the Atlantic Forest in the Paraguay and Paraná drainage, and in enclaves of humid forests in the Cerrado of the Central Brazil. BRAZIL — Pará: Belém (Mielke 1973), Santo Antônio de Tauá, Paragominas, Dom Eliseu. Maranhão: Feira Nova do Maranhão. Mato Grosso (Brown Jr 1987): Barra do Bugres, Cuiabá, Diamantino, Nova Xavantina. Goiás: Pirenópolis, Santa Rita do Araguaia. Distrito Federal: Brasília, Sobradinho; Minas Gerais: Curvelo. São Paulo: Marília, Pereira Barreto, Rio Claro. Paraná: Diamante do Norte, Foz do Iguaçu, Jussara, Maringá, Planaltina do Paraná, São Pedro do Ivaí. ARGENTINA — Missiones (Núñes-Bustos 2011): General Belgrano, Puerto Iguazú (Núñes-Bustos 2009). PARAGUAY — Alto Parana (Kochalka et al. 1996).	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D4ACA5FFF7CF927FDCBF9C4.taxon	description	Temporal distribution. This species occurs throughout the year, except in November.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D4ACA5FFF7CF927FDCBF9C4.taxon	biology_ecology	Host plant. Unknown.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D4ACA5FFF7CF927FDCBF9C4.taxon	etymology	Etymology. This species is dedicated to Liége Mallmann Granville, by the donation of the Eduardo Emery collection to the Departamento de Zoologia, Setor de Ciências Biológicas, Universidade Federal do Paraná, thus materializing the intentions of her husband and our dear friend. The name is a noun in apposition.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D4ACA5FFF7CF927FDCBF9C4.taxon	materials_examined	Type material. Holotype male deposited at the DZUP with the following labels: / HOLOTYPUS / Brazil, D [istrito] F [ederal], Mata Rib [eirinha], Sobradinho, 3. V. 2001, Eduardo Emery leg. / ex collection Eduardo Emery / gen. prep. A. Medeiros, 2018 / DZ 41.893 / Holotypus Artines liege Medeiros, O. Mielke & Casagrande det. 2018 /. Allotype female deposited at the DZUP with the following labels: / ALLOTYPUS / Brazil, D [istrito] F [ederal, Brazil], Gama Setor Sul, Chácara Floresta de Galeria, 26. VI. 2007 / Ex Coleção Eduardo Emery / DZ 41.850 / Allotypus Artines liege Medeiros, O. Mielke & Casagrande det. 2018 /. Paratypes: BRAZIL — Pará: Santo Antônio de Tauá (Reserva Sonho Azul), 1 male, 13. IV. 2005, P. Jauffret leg., DZ 41.863 (DZUP); Paragominas, 1 female, 22. VII. 1974, Exc. Depto. Zool. leg., DZ 41.899 (DZUP); Dom Eliseu (km 41 da Br 221), 1 female, 29. V. 2005, P. Jauffret leg., DZ 41.909 (DZUP). Maranhão: Feira Nova do Maranhão (26 km L, Faz. Forquilha dos Brejos), 1 male, 28. XI – 4. XII. 2010, O. Mielke & Carneiro leg., DZ 41.843, 1 female, 15 – 25. II. 2012, O. Mielke & C. Mielke leg., DZ 41.859 (DZUP). Mato Grosso: Barra do Bugres, 150 m, 1 female, 25. V. 1974, 1 male, 31. V. 1974, 1 female, 8. VI. 1974, Furtado leg., DZ 30.957, DZ 41.910, DZ 41.315 (DZUP), 1 male, 28. VII. 1974, Furtado leg., OM 30.155 (OM); Diamantino (Fazenda São João), 300 – 400 m, 1 male, 3. VII. 1975, 1 male, 13. III. 1978, 1 female, 7. V. 1978, 1 female, 10. VI. 1983, Furtado leg., DZ 41.255, DZ 8.866, DZ 41.295, DZ 41.792, 1 female, 26. IV. 1978, H. & H. D. Ebert leg., DZ 41.335, 1 female, 16. I. 1978, 1 male, 3 females, 22. I. 1978, O. Mielke & Furtado leg., DZ 41.819, DZ 41.900, DZ 41.752, DZ 41.305, DZ 41.799 (DZUP); Nova Xavantina (Bacaba), 2 females, 17 – 19. VIII. 1997, O. Mielke leg., DZ 41.762, DZ 41.772 (DZUP); São Vicente (São José da Serra), 700 m, 1 female, 24. VI. 1972, 1 female, 15. VII. 1972, O. Mielke & Brown leg., DZ 41.782, DZ 41.325 (DZUP). Goiás: Pirenópolis, 830 m, 1 female, 23. VI. 2011, E. Emery leg., DZ 41.842 (DZUP); Santa Rita do Araguaia, 1 male, 1 female, 20. V. 1969, 1 male, 21. V. 1969, Brown leg., DZ 1.411 (DZUP), OM 66.028, OM 65.993 (OM). Distrito Federal: Brasília 1 female, 30. VI. 1976, Gifford leg., DZ 41.849 (DZUP), (Escola Fazendária), 1 male, 6. VI. 1977, 2 males, 20. V. 1977, Gifford leg., DZ 8.846, DZ 41.245, DZ 41.275 (DZUP), (Km 12 between Buritis – Brasília), 1 male, 30. VI. 1976, Gifford leg., DZ 8.854 (DZUP), (Parque do Gama), 1000 m, 1 female, 17. VI. 1972, O. Mielke & Brown leg., DZ 41.830 (DZUP), 1 female, 7. II. 1978, Brown leg., OM 66.018 (OM). Minas Gerais: Curvelo (cabeceira do Córrego Leitão), 700 m, 1 male, 15. VI. 1972, O. Mielke & Brown leg., DZ 41.255 (DZUP). São Paulo: Pereira Barreto, 300 m, 1 male, 1. V. 1973, H. Ebert leg., DZ 41.883 (DZUP); Marília (Avencas), 600 m, 1 male, 6. I. 1966, H. Ebert leg., DZ 41.235 (DZUP); Rio Claro, 600 m, 1 female, 23. V. 1964, H. Ebert leg., DZ 41.812 (DZUP). Paraná: Diamante do Norte (Estação Ecológica do Caiuá), 300 m, 1 female, 23. IV. 2011, Dolibaina & Salik leg., DZ 41.892 (DZUP), 1 male, 27. I. 2012, Carneiro, Dolibaina & Salik leg., DZ 41.285 (DZUP); Foz do Iguaçu (Parque Nacional do Iguaçu), 2 females, 21 – 24. IV. 1995, O. Mielke & Casagrande leg., DZ 41.879, DZ 41.872 (DZUP); Jussara (Horto CMNP), 500 m, 1 female, 12. XII. 1975, Moure, O. Mielke & Wedderhoff leg., DZ 41.852 (DZUP); Maringá, 1 male, 2 females, 28. I. 1971, Becker leg., DZ 41.789, DZ 41.802, DZ 41.822 (DZUP); Planaltina do Paraná (20 km SW, RPPN Fazenda Duas Barras), 250 m, 1 female, 16. XII. 2009, O. Mielke, Carneiro, Maia & Dolibaina leg., DZ 41.832 (DZUP); São Pedro do Ivaí (RPPN Barbacena), 300 m, 1 male, 1 female, 8. X. 2010, O. Mielke, Dolibaina, Carneiro & M. Maia leg., DZ 41.265, DZ 41.882 (DZUP), 1 female, 2. X. 2012, Exc. LABLEP leg., DZ 41.862 (DZUP). ARGENTINA — Missiones: General Belgrano (Yacutinga Reserve), 2 males, 2 – 5. III. 2007, O. Mielke & Casagrande leg., DZ 8.978, DZ 41.860, 1 female, 17. III. 2003, N. Bustos leg., DZ 41.869 (DZUP).	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D48CA5EFF7CF926FCD2F848.taxon	description	Figs 13, 35, 48, 60, 72, 78 urn: lsid: zoobank. org: act: 4 E 9 AB 23 B-A 26 F- 4961 - A 2 E 6 - 4 B 8790 B 57 A 5 A	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D48CA5EFF7CF926FCD2F848.taxon	diagnosis	Diagnosis. Artines bamba sp. nov. differs from the other species of the “ aepitus group ” in the following combination of characters: harpe with ventral process narrow and cylindrical, strongly arched inwards and distally pointed, with long and thick setae on subapical portion; cornutus with long and robust spines, curved at base and directed anteriorly; proximo-lateral projections of lamella postvaginalis developed; ductus bursae with sclerotized plates.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D48CA5EFF7CF926FCD2F848.taxon	description	Description. Male (Fig. 13 A – B). Head: antenna with nudum bearing 12 segments. Forewing length: 12 – 15 mm. Dorsal forewing (Fig. 13 A): two or three apical spots (absent in most specimens); discal spots absent or reduced to a few scales; spot in CuA 2 – 2 A absent. Ventral forewing (Fig. 13 B): as in A. aepitus. Ventral hind wing (Fig. 13 B): apex of discal cell with few yellow scales; central black spot quadrate; blue discal spots in M 3 – CuA 1 and CuA 1 – CuA 2 quadrate. Genitalia (Figs 35 A – L, 48): tegumen more or less quadrate, with proximal margin straight and distal margin not conspicuous (Fig. 35 C), in lateral view shorter than length of gnathos; ventral arm of tegumen longer than dorsal arm of saccus (Fig. 35 A). Saccus as long as tegumen + uncus, base as long as lateral margin (Fig. 35 B). Uncus, in dorsal view, twice longer than wide, longer than tegumen, without disto-lateral projections, distal margin rounded (Fig. 35 C). Gnathos as in the other species of the group (Fig. 35 A, D). Valva three times longer than wide; sacculus elongate and about 1 / 2 width of valva; costa narrow and contiguous with ampulla; ampulla triangular and shorter than costa; ventral process of harpe narrow, elongate, cylindrical and strongly arched inwards, ventral margin slightly sinuous and distal end pointed, with subapical setae thick and long; dorsal process of harpe on median portion, rounded in dorsal view, wider than ventral process, apex surpassing dorsal margin of valva (Fig. 35 E – F). Aedeagus as long as valva, slightly curved dorsally; ventral projection distally round and strongly curved ventrally (Fig. 35 G – J); posterior end of cornutus wide and triangular, with spines thick, long and curved anteriorly (Fig. 48). Female (Fig. 13 C – D). Forewing length: 13.5 – 14 mm. Similar to male. Genitalia (Figs 60 A – B, 72): tergum VIII with complete spiracular opening. Sterigma triangular; lamella antevaginalis with proximo-ventral folds narrow; lamella postvaginalis without lateral fold, distal projections as in A. aepitus, proximo-lateral projections developed and bilobed. Ostium narrow surrounded by membranous areas; ductus bursae with sclerotized plates near insertion of sterigma. Variation. As in most species of the “ aepitus group ”, the variation is in the shape of the forewing marks. These marks are absent in most individuals analyzed and only the holotype has three apical spots and two male paratypes have conspicuous discal spots. The marks are present in females from Acre, Brazil.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D48CA5EFF7CF926FCD2F848.taxon	discussion	Comments. Species similar to A. rica and A. liege sp. nov. in the genitalia, differing in the cornutus with long, robust spines directed anteriorly and ventral projection of harpe more elongated, distally pointed, strongly arched inwards, with setae on subapical portion. This species is unique among the “ aepitus group ” with the sclerotized plate at base of ductus bursae (Figs 60 and 72). The association of the female in this species was made based on the absence of apical and discal spots on dorsal forewing (see variation above) and distribution. However, in view of the fact that species are similar and different species can potentially occur in the same locality, the association is tentative and for this reason the females were not included in the type-series. Geographic distribution (Fig. 78). The species occurs in the western Amazon. BRAZIL — Amazonas: São Paulo de Olivença. Acre: Senador Guiomard, Porto Acre. PERU — Madre de Díos. San Martin: Juanjui, Moyobamba.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D48CA5EFF7CF926FCD2F848.taxon	description	Temporal distribution. This species occurs in July, September and October.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D48CA5EFF7CF926FCD2F848.taxon	biology_ecology	Host plant. Unknown.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D48CA5EFF7CF926FCD2F848.taxon	etymology	Etymology. The term " bamba " is the termination of the word Moyobamba, the type locality of the species.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D48CA5EFF7CF926FCD2F848.taxon	materials_examined	Type material. Holotype male deposited at the DZUP with the following labels: / HOLOTYPUS / Peru, San Martin, Moyobamba, Calzada. X. 2015, J. Pintado leg. / DZ 20. 673 / Holotypus Artines bamba Medeiros & O. Mielke det. 2018 /. Paratypes: BRAZIL — Amazonas: São Paulo de Olivença, 1 male, no data, no collector, OM 9.115 (OM). PERU — Madre de Díos: (Parque Nacional del Manu, Pakitza), 340 m, 1 male, 2. X. 1991, 1 male, 10. X. 1991, M. Casagrande leg., DZ 39.901, DZ 39.895, 1 male, 20. X. 1991, O Mielke leg., DZ 9.728 (| DZUP), 1 male, 29. IX. 1991, R. Robbins leg., (MUSM). San Martin: Juanjui, 200 m, 1 male, 15. IX. 2002, no collector, OM 60.174 (OM); Moyobamba (Calzada), 1 male, X. 2015, J. Pintado leg., OM 79.553, 850 m, 1 male, X. 2016, J. Pintado leg., OM 83.226, 800 m, 2 males, X. 2016, J. Pintado leg., OM 81.199, OM 83.166, (OM). Additional material (all females). BRAZIL — Amazonas: São Paulo de Olivença, 1 female, no data, no collector, OM 9.116 (OM). Acre: Senador Guiomard (Reserva Catuaba), 1 female, 23 – 27. VII. 2008, O. Mielke & Casagrande leg., DZ 43. 327 (DZUP); Porto Acre (Humaitá), 1 female, 28 – 31. VII. 2008, O. Mielke & Casagrande leg., DZ 43.347 (DZUP). PERU — San Martin: Moyobamba (Calzada), 1 female, X. 2015, J. Pintado leg., OM 79.413, 800 m, 1 female, X. 2016, J. Pintado leg., OM 81.229 (OM).	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D4ECA59FF7CFF19FB1AFD63.taxon	description	Figs 2 – 3, 14 – 18, 20, 21 – 25, 36 – 40, 49 – 53, 61 – 64, 73 – 76, 80 Description. Male and female: dorsal forewing with semi-hyaline and large discal spots; ventral forewing with or without white spot in distal 1 / 2 of CuA 2 – 2 A; ventral hind wing with discal area yellow to orange, with a central black spot and a yellow postdiscal band limited distally by ground stripe, wavy or not. Male: brand present; saccus wide and long, shape variable; uncus long and narrow; arms of gnathos with pointed apexes; valva with distal projection composed of harpe and ampulla, with triangular aspect from 1 / 2 its length; harpe simple and fused with ampulla; aedeagus without disto-dorsal projection; ventral projection of aedeagus wide, distally bifid or trifid, diverging ventrally at angle of approximately 45 degrees with aedeagus; cornutus as a moderately sclerotized plate over most of vesica and with spines of variable thickness and length; fultura inferior arched in lateral view, proximal end simple. Female: externally similar to male; region between seventh sternite and lamella antevaginalis membranous; sterigma partly sclerotized, rectangular and joined with tergum by the lateral plate (Fig. 61 A); lamella postvaginalis with one or two sclerotized plates (Fig. 62 B); lateral fold of lamella postvaginalis membranous (Fig. 61 B); ductus bursae with basal portion sclerotized or membranous. Included species: Artines aquilina (Plötz, 1882), Artines angelica Medeiros sp. nov., Artines focus Evans, 1955, Artines cofus Medeiros & Dolibaina sp. nov. and Artines donia Medeiros & Carneiro sp. nov.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D47CA52FF7CF8EFFAC8FE44.taxon	description	Figs 15, 22, 37, 50, 62, 74, 80 urn: lsid: zoobank. org: act: F 54 ED 9 DC- 3345 - 4396 - 8 A 35 - 76 DAD 7 A 30381	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D47CA52FF7CF8EFFAC8FE44.taxon	diagnosis	Diagnosis. Artines angelica sp. nov. differs from the other species of the “ aquilina group ” in the following combination of characters: ventral hind wing with postdiscal band limited distally by a not wavy dark-brown stripe; uncus with distal 1 / 2 pointed; ampulla protruding dorsally, not exceeding the dorsal margin of costa; ventral projection of the aedeagus with distal portion strongly trifid; cornutus with numerous short and pointed spines from base to distal end of vesica; lamella postvaginalis with proximal sclerotized plate wider than ostium. Descripiton. Male (Fig. 15 A – B). Head: antenna with nudum bearing 11 to 12 segments.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D47CA52FF7CF8EFFAC8FE44.taxon	description	Forewing length: 12 – 14 mm. Dorsal forewing (Fig. 15 A): as in A. aquilina (Fig. 22 A). Ventral forewing (Fig. 15 B): four bluish apical spots, the spots in R 2 – R 3 and R 3 – R 4 elongated; discal spots as on dorsal surface; white, broad and diffuse spot in CuA 2 – 2 A. Ventral hind wing (Fig. 15 B): apical portion of discal cell densely covered by orange to yellow scales; central black spot large, quadrate to rectangular; postdiscal band yellow, limited distally by a not wavy dark brown stripe. Genitalia (Figs 37 A – M, 50): tegumen as long as wide, distally rounded (Fig. 37 C), in lateral view rectangular, dorsal margin slightly convex (Fig. 37 A). Saccus narrower on proximal 1 / 2, anterior portion wider than tegumen, proximal margin straight or slightly concave (Fig. 37 B). Uncus similar to A. aquilina differing in the longer and narrower distal portion (Fig. 37 A, C). Gnathos reaching distal 1 / 3 of uncus, arms parallel and weakly convergent at apex in ventral view (Fig. 37 D), in lateral view moderately curved dorsally toward distal end of uncus (Fig. 37 A). Valva two and a half times longer than wide; sacculus triangular, less than 1 / 2 length of valva, separated from harpe by short distance; costa narrow and triangular, projected inwards, separated from ampulla by distance larger than its length; harpe as long as 1 / 2 valva, uniformly narrow; ampulla as wide as 1 / 4 harpe, dorsally protruding and not exceeding dorsal margin of valva; distal projection of valva pointed, longer and less curved inwards than in A. aquilina (Fig. 37 E – F). Aedeagus uniformly wide; opening of ejaculatory bulb as in A. aquilina; ventral projection diverging ventrally at an angle of approximately 45 degrees, distal apex asymmetrically trifid, rarely quadrifid (Fig. 37 G – J); cornutus as a stripe bearing numerous short spines, from base to apex of vesica (Figs 37 M, 50). Fultura inferior, in lateral view, uniformly narrow (Fig. 37 L). Female (Fig. 15 C – D). Forewing length: 12 – 14 mm. Similar to male. Genitalia (Figs 62 A – B, 74): lamella antevaginalis wide; lamella postvaginalis with two sclerotized plates, one proximal wider than ostium, the other distal with median notches on apical and proximal margins, the latter shaped as an inverted “ U ”. Ostium situated on median portion of sterigma; ductus bursae with 1 / 4 basal sclerotized and wide. Variation. Apical spots of dorsal forewing may be absent and only one female from Bolivia has three conspicuous apical spots. The spot in CuA 2 – 2 A of dorsal forewing may be absent in a few females.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D47CA52FF7CF8EFFAC8FE44.taxon	discussion	Comments. Species similar to A. aquilina in the male and female genitalia (see comments in A. aquilina), however, the color pattern is peculiar by the ventral hind wing without wavy marks, with orange discal area and a large central black spot. Geographic distribution (Fig. 80). The species occurs from southeast to east of Amazon Rainforest, reaching to Atlantic Forest areas of the northeastern Brazil. BRAZIL — Pará: Paragominas. Maranhão: Santa Luzia. Acre: Mâncio Lima, Xapuri. Alagoas: Maceió. Mato Grosso: Sinop. PERU — Madre de Díos: Puerto Maldonado, Parque Nacional del Manu. BOLIVIA — La Paz: Caranavi.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D47CA52FF7CF8EFFAC8FE44.taxon	description	Temporal distribution. This species occurs throughout the year being more abundant between August and January.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D47CA52FF7CF8EFFAC8FE44.taxon	biology_ecology	Host plant. Unknown.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D47CA52FF7CF8EFFAC8FE44.taxon	etymology	Etymology. This species is dedicated to Angelica Faustino Alves, first author’s wife, in gratitude of her continuous support and incentive. The name is a noun in apposition.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D47CA52FF7CF8EFFAC8FE44.taxon	materials_examined	Type material. Holotype male deposited at the DZUP with the following labels: / HOLOTYPUS / Água Azul, PA [rá], Paragominas, [Brazil], 16. VIII. 1974, Mielke leg. / gen. prep. A. Medeiros 2018 / DZ 41. 243 / Holotypus Artines angelica Medeiros det. 2018 /.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D45CA6DFF7CF888FCA7F84C.taxon	description	Figs 16, 23, 38, 51, 63, 75, 80	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D45CA6DFF7CF888FCA7F84C.taxon	description	Taxonomic history. Artines focus was described by Evans (1955) based in five males and five females from Guyana and Roraima, Pará (Santarém) and Pernambuco, Brazil. The other authors mentioned this species in taxonomic comments, faunistic studies and catalogs.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D45CA6DFF7CF888FCA7F84C.taxon	diagnosis	Diagnosis. Artines focus differs from the other species of the “ aquilina group ” in the following combination of characters: ventral forewing without spot in CuA 2 – 2 A; uncus distally wide, with a ventral projection; aedeagus proximal portion narrow; opening of ejaculatory bulb separated from proximal margin of aedeagus by short distance; lamella postvaginalis without proximal plate, distal sclerotized plate without median notch on proximal margin and with distal margin approximately straight.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D45CA6DFF7CF888FCA7F84C.taxon	description	Redescription. Male (Fig. 16 A – B). Head: antenna with nudum bearing 12 to 13 segments. Forewing length: 12 – 15 mm. Dorsal forewing (Fig. 16 A): one to three apical spots, spot in R 3 – R 4, when present, represented by few scales; discal spots as in A. aquilina; a white and yellow spot in CuA 2 – 2 A; brand with lower projection long, upper projection with narrow base and pointed apex (Fig. 23). Ventral forewing (Fig. 16 B): costal area with weak violet aspect; three to four apical spots, spot in R 2 – R 3 not conspicuous and shifted to base; discal spots as on dorsal surface; submarginal band conspicuous and not contiguous in the middle of the cells; without spot in CuA 2 – 2 A. Ventral hind wing (Fig. 16 B): apical portion of discal cell densely covered by orange scales; central black spot quadrate to rectangular; postdiscal band yellow, limited distally by not wavy margin. Genitalia (Figs 38 A – M, 51): tegumen longer than wide, distally trapezoid from 1 / 2 of its extension (Fig. 38 C), in lateral view rectangular, dorsal margin strongly convex (Fig. 38 A). Saccus, in ventral view, narrower on proximal 1 / 2, anterior portion narrower than tegumen, proximal margin straight or convex (Fig. 38 B). Uncus with ventral projection, in dorsal view with base as wide as 1 / 2 width of tegumen, proximal 1 / 2 with lateral margin concave, distal 1 / 2 narrower, tip rounded to slightly bifid (Fig. 38 C), in lateral view, strongly curved ventrally (Fig. 38 A). Gnathos reaching distal 1 / 2 of uncus, arms parallel and slightly divergent at apex in ventral view (Fig. 38 D), tips slightly curved dorsally, directed to distal 1 / 2 of uncus (Fig. 38 A). Valva twice longer than wide; sacculus reduced; costa narrow and triangular, in dorsal view as in A. aquilina, however less projected inwards and separated from ampulla by short distance; harpe longer than 1 / 2 valva, uniformly narrower than in the other species of the group; ampulla as wide as 1 / 2 harpe, dorsally protruding; distal projection of the valva slightly curved inwards and dorsally (Fig. 38 E – F). Aedeagus narrower on proximal portion; opening of ejaculatory bulb ovoid and separated from proximal margin of aedeagus by short distance; ventral projection bifid, narrow at base, diverging at an angle less than 45 degrees and with distal portion asymmetrically bifid (Fig. 38 G – J). Cornutus with spines as setae from base to median portion of vesica (Figs 38 M, 51). Fultura inferior, in lateral view, widened distally (Fig. 38 L). Female (Fig. 16 C – D). Forewing length: 14 mm. Similar to male. Genitalia (Figs 63 A – B, 75): lamella antevaginalis narrow; lamella postvaginalis without proximal plate, distal plate sclerotized, without sclerotized median notch on proximal margin, distal margin approximately straight. Ostium on proximal 1 / 2 of sterigma; ductus bursae narrow. Variation. Some specimens from Reserva Duque, Manaus, Amazonas, Brazil and from French Guiana have vestigial spot in CuA 2 – 2 A on ventral forewing. The male genitalia of the specimens from French Guiana have uncus wider and ventral projection more developed, with distal end dilated (Fig. 38 D).	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D45CA6DFF7CF888FCA7F84C.taxon	discussion	Comments. Artines focus is easily recognized for being the only species of the genus that has uncus with a ventral projection. It is sympatric with A. aepitus in the northern Amazon Rainforest, from which it can be easily differentiated by the broad discal spots on forewing and by the presence of brand in the males. The record of Evans (1955) for Pernambuco, Brazil is probably from a mislabeled specimen. Geographic distribution (Fig. 80). The species occurs in the Guiana Shield. VENEZUELA — Amazonas: Yavita. FRENCH GUIANA — Saint-Laurent-du-Maroni: Régina. Cayenne; Mitaraka (Garrouste 2003, 2007). BRA- ZIL — Roraima: Alto Alegre, Pacaraima. Amazonas: Manaus.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D78CA69FF7CFB8DFB23FAE1.taxon	description	Figs 17, 24, 39, 52, 64, 76, 80 urn: lsid: zoobank. org: act: 1 F 18644 C- 2 C 5 A- 4733 - 8721 - DCD 9 D 5 CF 788 F Artines focus (misid.); O. Mielke; Carneiro & Casagrande, 2010. Biota Neotrop. 10 (4): 296. Taxonomic history. In its only record in the literature this species was misinterpreted as A. focus (Mielke et al. 2010).	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D78CA69FF7CFB8DFB23FAE1.taxon	diagnosis	Diagnosis. Artines cofus sp. nov. differs from the other species of the “ aquilina group ” in the following characters: uncus distally wide and without ventral projection; aedeagus with proximal end narrow; opening of ejaculatory bulb separated from proximal margin of aedeagus by short distance; ventral projection of aedeagus with distal portion bifid; lamella postvaginalis without proximal plate, distal plate with sclerotized median notch on proximal margin and a small median projection on distal margin.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D78CA69FF7CFB8DFB23FAE1.taxon	description	Description. Male (Fig. 17 A – B). Head: antenna with nudum bearing 12 segments. Forewing length: 11.5 – 13 mm. Dorsal forewing (Fig. 17 A): apical spots absent or one or two in R 3 – R 4 and R 4 – R 5; discal spots reduced and spot in CuA 2 – 2 A absent or vestigial; brand shorter, lower projection slightly longer than A. aquilina (Fig. 24). Ventral forewing (Fig. 17 B): three to four reduced apical spots; discal spots as on dorsal surface; submarginal band conspicuous and almost discontinued at the middle of cells; weak spot white and diffuse in CuA 2 – 2 A. Ventral hind wing (Fig. 17 B): apical portion of discal cell densely covered by yellow scales; central black spot large, as wide as long; postdiscal band yellow, limited distally by a not wavy dark brown stripe. Genitalia (Figs 39 K – M, 52): tegumen more or less quadrate, distal margin convex (Fig. 39 C), in lateral view rectangular, dorsal margin slightly convex (Fig. 39 A). Saccus narrow medially, anterior portion slightly dilated and narrower than tegumen, proximal margin rounded (Fig. 39 B). Uncus shorter than tegumen, proximal 1 / 2 with lateral margins convex, distal 1 / 2 narrow, lateral margins concave, distal portion rounded (Fig. 39 C), apex curved ventrally (Fig. 39 A). Gnathos reaching 1 / 2 length of uncus, arms parallel, slightly divergent on apex in ventral view (Fig. 39 D), curved upwards toward distal end of uncus (Fig. 39 A). Valva two and a half times longer than wide; sacculus short and triangular separated from harpe by long distance; costa narrow, triangular, short, projected inwards and separated from ampulla by distance longer than its length; harpe longer than 1 / 2 of valva, uniformly narrow; ampulla 1 / 2 width of harpe, not protruding; distal projection of harpe, in dorsal view, weakly curved inwards and with small distal spines (Fig. 39 E – F). Aedeagus as in A. focus, differing in the opening of the ejaculatory bulb more distant from proximal margin; ventral projection with distal lobes asymmetrical (Fig. 39 G – J). Cornutus similar to A. focus, however, spines less conspicuous and less numerous (Figs 39 M, 52). Fultura inferior, in lateral view, uniformly narrow (Fig. 39 L). Female (Fig. 17 C – D). Forewing length: 12 mm. Similar to male. Genitalia (Figs 64 A – B, 76): lamella antevaginalis narrow; lamella postvaginalis without proximal plate, distal plate with sclerotized median notch on proximal margin, distal margin with small median projection. Ostium on proximal 1 / 2 do sterigma; ductus bursae narrow. Variation. The central black spot on ventral hind wing is sometimes rectangular.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D78CA69FF7CFB8DFB23FAE1.taxon	discussion	Comments. Wing pattern, male and female genitalia as in A. focus, differing in the smaller size, reduced white spots on forewing, absence of ventral projection of uncus and narrower valva. Geographic distribution (Fig. 80). This species occurs in the western Amazon Rainforest. PERU — Madre de Díos: Puerto Maldonado. San Martin: Sacanche. BRAZIL — Acre: Santa Rosa do Purus. Rondônia: Ouro Preto do Oeste.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D78CA69FF7CFB8DFB23FAE1.taxon	description	Temporal distribution. This species occurs between August and October.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D78CA69FF7CFB8DFB23FAE1.taxon	biology_ecology	Host plant. Unknown.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D78CA69FF7CFB8DFB23FAE1.taxon	etymology	Etymology. The name is an anagram of focus, the closest species of A. cofus sp. nov..	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D78CA69FF7CFB8DFB23FAE1.taxon	materials_examined	Type material. Holotype male deposited at the DZUP with the following labels: / HOLOTYPUS / Ouro Preto do Oeste, RO [ndônia, Brazil], 17 – 31. VIII. 1987, C. Elias leg. / Projeto Polo Noroeste / gen. prep. A. Medeiros 2018 / DZ 41.203 / Holotypus Artines cofus Medeiros & Dolibaina det. 2018 /. Allotype female deposited at the DZUP with the following labels: / ALLOTYPUS / Ouro Preto do Oeste, RO [ndônia, Brazil], 17 – 31. VIII. 1987, C. Elias leg. / Projeto Polo Noroeste / gen. prep. A. Medeiros 2018 / DZ 41. 252 / Allotypus Artines cofus Medeiros & Dolibaina det. 2018 /. Paratypes: PERU — Madre de Díos: Puerto Maldonado (Tambopata), 300 m, 1 male, 25. X. 1991, R Robbins leg. (MUSM); San Martin: Sacanche, 1 female, 5. IX. 2012, C. Calderon leg., OM 73.640 (OM). BRAZIL — Acre: Santa Rosa do Purus, 1 male, 6 – 8. VIII. 2008, O. Mielke & Carneiro leg., DZ 41.303 (DZUP).	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D7ECA6AFF7CFA4BFA86FE44.taxon	description	Figs 18, 25, 40, 53, 80 urn: lsid: zoobank. org: act: 5 EA 37 EFE- 671 F- 4 C 96 - B 8 B 5 - 130 B 435 F 3922	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D7ECA6AFF7CFA4BFA86FE44.taxon	diagnosis	Diagnosis. Artines donia sp. nov. differs from the species of the “ aquilina group ” in the following characters: ventral hind wing with central black spot wider than long; uncus distally wide, without ventral projection, apex weakly bifid; valva distally rhomboid; aedeagus uniformly wide; ventral projection of aedeagus with distal portion bifid; cornutus similar to A. focus and A. cofus sp. nov., differing in being more sclerotized and having spines slightly conspicuous; fultura inferior robust and laterally wider than in the other species of the “ aquilina group ”.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D7ECA6AFF7CFA4BFA86FE44.taxon	description	Description. Male (Fig. 18 A – B). Head: antenna with nudum bearing 12 segments. Forewing length: 14 mm. Dorsal forewing (Fig. 18 A): two apical spots in R 4 – R 5 and R 5 – M 1; discal spots as in A. aquilina; white spot in CuA 2 – 2 A very conspicuous, triangular; brand shorter than in other species of the “ aquilina group ”, with lower projection long and upper projection uniformly narrow and with apex rounded. Ventral forewing (Fig. 18 B): apical and discal spots as in A. aquilina; submarginal band as in A. cofus sp. nov.; white spot broad and strongly marked in CuA 2 – 2 A. Ventral hind wing (Fig. 18 B): apical portion of discal cell densely covered by orange scales; central black spot rectangular, wider than long; postdiscal band yellow, limited distally by wavy dark brown stripe. Genitalia (Figs 40 A – L, 53): tegumen more or less oval, longer than wide (Fig. 40 C), in lateral view, rectangular, dorsal margin straight on its entire stretch (Fig. 40 A). Saccus narrowed medially, anterior portion narrower than tegumen, proximal margin rounded (Fig. 40 B). Uncus shorter than tegumen, proximal 1 / 2 with lateral margins slightly concave and distally divergent, distal 1 / 2 narrower with strong lateral concavity, distal end weakly bifid (Fig. 40 C), little curved ventrally (Fig. 40 A). Gnathos in ventral view reaching 1 / 2 of uncus, arms parallel with distal ends straight (Fig. 40 D), curved dorsally toward distal end of uncus (Fig. 40 A). Valva three times longer than wide; costa narrow and triangular, laterally projected inwards, separated from ampulla by distance shorter than its length; harpe longer than 1 / 2 valva, widened from base; ampulla 1 / 2 width of median portion of harpe, dorsally protruding, slightly exceeding dorsal margin of valva; distal projection of valva wide, with thorn-like projection on dorsal-distal portion (Fig. 40 E – F). Aedeagus uniformly wide, weakly curved dorsally; opening of ejaculatory bulb dorsal near proximal margin of aedeagus; ventral projection diverging from aedeagus at an angled of 45 degrees, distal portion asymmetrically bifid (Fig. 40 G – J). Cornutus uniformly sclerotized, bearing setae-like spines at base of vesica (Figs 40 M, 53). Fultura inferior robust, in lateral view wider than in the other species of the group (Fig. 40 L). Female. Unknown. Variation. The specimen from Goiás, Brazil, is paler than the specimen from Rondônia, which may be due to the fact that the wing is worn.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D7ECA6AFF7CFA4BFA86FE44.taxon	discussion	Comments. Collor pattern as in A. focus, A. angelica sp. nov. and A. cofus sp. nov., differing in the central black spot of the ventral hind wing more reduced and wider than long. The male genitalia are similar to A. focus and A. cofus sp. nov., from which it differs in the following characters: uncus slightly bifid, without ventral projection, aedeagus uniformly wide, fultura inferior robust and valva distally not pointed, with projection similar to a distally projected thorn. Geographic distribution (Fig. 80). It is only known from the southern Brazilian Amazon Rainforest. BRA- ZIL — Rondônia: Vilhena. Goiás: Santa Rita do Araguaia. Temporal distribution. This species occurs in May and July. Host plant. Unknown. Etymology. The term “ donia ” is the termination of the word “ Rondônia ”, the type locality of the species. Type material. Holotype male deposited at the DZUP with the following labels: / HOLOTYPUS / Vilhena, RO [ndônia, Brazil], 10. VII. 1972, Mielke leg. / gen. prep. A. Medeiros 2017 / DZ 41.283 / Holotypus Artines donia Medeiros & Carneiro det. 2018 /. Paratype: BRAZIL — Goiás: Santa Rita do Araguaia, 1 male, 21. V. 1969, Brown leg., OM 66.000 (OM).	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D7DCA6AFF7CFDA7FD93FD59.taxon	description	Figs 19, 26, 41, 65, 77, 80 Description. This group is characterized by the diagnosis and description of the only species included. Included species: Artines bipunctata.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D7DCA65FF7CFCA2FC45F9F5.taxon	description	Figs 19, 26, 41, 65, 77, 80	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D7DCA65FF7CFCA2FC45F9F5.taxon	description	Taxonomic history. Artines bipunctata was described by Mielke (1968) based on seven specimens (one male and six females) from Jardim Zoológico, Brasília, Distrito Federal, Brazil and one male from Barbacena, Minas Gerais, Brazil. The other authors mentioned this species in taxonomic comments, faunistic studies and catalogs.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D7DCA65FF7CFCA2FC45F9F5.taxon	diagnosis	Diagnosis. Artines bipunctata differs from other species of the genus in the following combination of characters: dorsal forewing with large and semi-hyaline discal spots; ventral hind wing with discal area pale brown, without central black spot and a violet postdiscal band; male with brand; uncus wide and distally bifid; gnathos arms distally curved inward; valva distally rounded; harpe simple and not fused with ampulla; ventral projection of aedeagus wide and parallel to aedeagus, distal end bilobed, with rounded lobes bearing numerous spines; vesica entirely membranous, with very small and dispersed spines; fultura inferior laterally straight, proximal end bifid; sterigma partially sclerotized; lamella postvaginalis bifid and without lateral folds; region between seventh sternite and lamella antevaginalis completely membranous; ductus bursae with basal portion sclerotized; ostium wide, “ D ” - shaped, surrounded by sclerotized area.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D7DCA65FF7CFCA2FC45F9F5.taxon	description	Description. Male (Fig. 19 A – B). External morphology as described by Mielke (1968). Genitalia (Fig. 41 A – J): tegumen more or less rectangular, wider than long, with short and triangular distal median projection (Fig. 41 C). Saccus triangular, base shorter than lateral margin (Fig. 41 B). Uncus rectangular, one and a half longer than wide, distally bifid, with rounded projections separated medially by shallow concavity (Fig. 41 C). Gnathos arms distally curved inward (Fig. 41 D). Valva twice longer than wide; sacculus triangular, narrow, longer than 1 / 2 of valva; costa rectangular and short, near median portion of valva, distally fused to ampulla; ampulla rectangular, longer than costa; harpe simple and not fused to ampulla, distal margin rounded with arched row of short and triangular spines turned inwards (Fig. 41 E – F). Aedeagus thick, longer than valva; opening of ejaculatory bulb on proximal end of aedeagus; ventral projection of aedeagus wide and parallel to aedeagus, distal end bilobed, with rounded lobes bearing numerous spines (Fig. 41 G – J). Vesica bearing small and short scattered spines (Fig. 39 G). Female (Fig 19 C – D). As described by Mielke (1968). Genitalia (Figs 65 A – B, 77): sterigma rectangular. Tergum VIII with incomplete spiracular opening. Lamella antevaginalis wide and membranous, without proximo-ventral folds. Lamella postvaginalis with two sclerotized plates, one proximal near the ostium, the other distal with two distal lateral projections. Ostium D-shaped, surrounded by sclerotized area. Ductus bursae wide, basal portion sclerotized. Variation. As mentioned by Mielke (1968) some specimens have a second spot / mark in the upper portion of CuA 2 – 2 A of the dorsal forewing, above the larger spot. The spiracular opening of tergum VIII of the female can be complete in some individuals.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D7DCA65FF7CFCA2FC45F9F5.taxon	discussion	Comments. Artines bipunctata shares several morphological characters with the species of the two groups considered before. For example, the presence of a brand on the male forewing and broad white and semi-hyaline spots on the forewing of both sexes are shared with the species of the “ aquilina group ”. On the other hand, characters of the male and female genitalia, as wide uncus, harpe and ampulla not fused, fultura inferior with base bifid, ventral projection of the aedeagus not divergent and sterigma distally bifid, are also present in the species of the “ aepitus group ”. However, the numerous and unique characters mentioned in the diagnosis above justify the inclusion of A. bipunctata in a group apart. Geographic distribution (Fig. 80). This species is only known from islands of humid forests in the Cerrado biome. BRAZIL — Goiás: Formosa, Goianésia, Vianópolis. Distrito Federal: Brasília. Minas Gerais: Barbacena.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D7DCA65FF7CFCA2FC45F9F5.taxon	description	Temporal distribution. This species occurs throughout the year, probably with several generations.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D7DCA65FF7CFCA2FC45F9F5.taxon	biology_ecology	Host plant. Unknown.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D7DCA65FF7CFCA2FC45F9F5.taxon	etymology	Etymology. Although omitted in the original description, the name is a reference of the two white spots on the forewing.	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
CB52563B9D7DCA65FF7CFCA2FC45F9F5.taxon	materials_examined	Type material. Holotype male deposited at the OM – DZUP with the following labels: / HOLOTYPUS / 3. II. 1967, J [ardim] Zoológico, Brasília, D [istrito] F [ederal, Brazil], Tangerini leg. / Artines bipunctata Mielke, 1968, Mielke det. 1968 / [OM] 10.948 /. Allotype female deposited at the OM – DZUP with the following labels: / ALLOTYPUS / 3. II. 1967, J [ardim] Zoológico, Brasília, D [istrito] F [ederal, Brazil], Tangerini leg. / Artines bipunctata Mielke, 1968, Mielke det. 1968 / [OM] 10.949 /. Paratypes: BRAZIL — Distrito Federal: Brasília (Jardim Zoológico de Brasília), 1000 m, 5 females, 3. II. 1967, N. Tangerini leg., OM 10.950, OM 10.951 (OM), DZ 41.234, DZ 41.214 DZ 41.244 (Ex. collection N. Tangerini now deposited at the DZUP). Minas Gerais: Barbacena, 1 male, 27. I. 1967, N. Tangerini leg., OM 10.947 (OM). Examined material. The type material mentioned above and the following specimens: BRAZIL — Goiás: Vianópolis (Ponte Funda), 1 male, 17. I. 1981, N. Tangerini leg., DZ 41.264 (DZUP); Formosa (Itiquira), 1 male, 15. V. 1977, Gifford leg., DZ 41.155 (DZUP); Goianésia, 1 male, IX. 1969, H. Ebert leg., DZ 41.284 (DZUP). Distrito Federal: Brasília, (Córrego Bananal), 1 female, 27. III. 2006, E. O. Emery leg., DZ 41.224 (DZUP), (Jardim Zoológico de Brasília), 1000 m, 1 male, 19. VI. 1972, O. Mielke & Brown leg., DZ 30.961 (DZUP), 1 male, 1 female, 3. II. 1967, N. Tangerini leg., OM 10.948, OM 10.949 (OM), (Parque Nacional de Brasília), 1 female, 31. III. 1977, Gifford leg., DZ 30.960 (DZUP), (Reserva IBGE), 1 male, 5. IV. 2002, 1 female, 20. VII. 2002, 1 male, 1 female, 20. VII. 2005, 1 male, 23. VIII. 2005, E. O. Emery leg., DZ 41.254, DZ 41.334, DZ 41.324, DZ 41.165, DZ 41.274 (DZUP) 1 male, 2 females, 7. IX. 1968, H. Ebert leg., DZ 41.194, DZ 41.184, DZ 41.204 (DZUP). Minas Gerais: Barbacena, 1100 m, 1 male, 16. VIII. 1951, 1 male, 4. IX. 1951, 1 male, 29. V. 1952, 1 male, 24. VII. 1952, 1 male, 16. VIII. 1952, 1 female, 14. XI. 1952, H. Ebert leg., DZ 41.304, DZ 41.145, DZ 41.314, DZ 41.254, DZ 41.294, DZ 41.175 (DZUP), 1 female, 19. V. 1990, N. Alves leg., OM 9.826 (OM).	en	Medeiros, Adalberto Dantas De, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Mielke, Olaf Hermann Hendrik, Casagrande, Mirna Martins (2019): Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species. Zootaxa 4614 (1): 1-49, DOI: 10.11646/zootaxa.4614.1.1
