taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
C97B87ACFFA5FF8EFF77FC645B8DF98B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17321840/files/figure.png	https://doi.org/10.5281/zenodo.17321840	FIGURE 1. Three new species of the Rhyacophila retracta Species Group. 1A, Rhyacophila geranos sp. nov., male; 1B, same, female; 1C, Rhyacophila humerigera sp. nov., male; 1D, same, female; 1E, Rhyacophila koryne sp. nov., male (photographed by K. Inazu); 1F, same, female (same by K. Inazu).	FIGURE 1. Three new species of the Rhyacophila retracta Species Group. 1A, Rhyacophila geranos sp. nov., male; 1B, same, female; 1C, Rhyacophila humerigera sp. nov., male; 1D, same, female; 1E, Rhyacophila koryne sp. nov., male (photographed by K. Inazu); 1F, same, female (same by K. Inazu).	2025-09-10	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
C97B87ACFFA5FF8EFF77FC645B8DF98B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17321842/files/figure.png	https://doi.org/10.5281/zenodo.17321842	FIGURE 2. Rhyacophila geranos sp. nov., male genitalia. 2A, dorsal (left inferior appendage omitted); 2B, ventral (left inferior appendage omitted); 2C, left lateral; 2D, right inferior appendage, mesal; 2E, segment X and phallic apparatus, left lateral; 2F, apical half of segment X, posterior; 2G, anal sclerite and its root, anterior; 2H, phallic apparatus, dorsal. Abbreviations: IX, X = abdominal segments IX, X; ab = apical band; adl = apicodorsal lobe; aed = aedeagus; as = anal sclerite; asia = apical segment of inferior appendage (paired); bsia = basal segment of inferior appendage (paired); dap = dorsal appendage of phallic apparatus; par = paramere (paired); pht = phallotheca; plp = posterolateral process of segment X (paired); tb = tergal band.	FIGURE 2. Rhyacophila geranos sp. nov., male genitalia. 2A, dorsal (left inferior appendage omitted); 2B, ventral (left inferior appendage omitted); 2C, left lateral; 2D, right inferior appendage, mesal; 2E, segment X and phallic apparatus, left lateral; 2F, apical half of segment X, posterior; 2G, anal sclerite and its root, anterior; 2H, phallic apparatus, dorsal. Abbreviations: IX, X = abdominal segments IX, X; ab = apical band; adl = apicodorsal lobe; aed = aedeagus; as = anal sclerite; asia = apical segment of inferior appendage (paired); bsia = basal segment of inferior appendage (paired); dap = dorsal appendage of phallic apparatus; par = paramere (paired); pht = phallotheca; plp = posterolateral process of segment X (paired); tb = tergal band.	2025-09-10	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
C97B87ACFFA5FF8EFF77FC645B8DF98B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17321844/files/figure.png	https://doi.org/10.5281/zenodo.17321844	FIGURE 3. Rhyacophila geranos sp. nov., female genitalia. 3A, segment VIII, dorsal; 3B, same, left lateral; 3C, same, ventral; 3D, vaginal apparatus, left lateral; 3E, same, ventral. Abbreviations: VIII = abdominal segments VIII; ar = apodemal rod of segment VIII (paired); dc = dorsal cleft of segment VIII; ismVIII–X = intersegmental membrane of segment VIII–X; lc = lateral cleft of segment VIII (paired); pp = posterior process; ps = processus spermathecae.	FIGURE 3. Rhyacophila geranos sp. nov., female genitalia. 3A, segment VIII, dorsal; 3B, same, left lateral; 3C, same, ventral; 3D, vaginal apparatus, left lateral; 3E, same, ventral. Abbreviations: VIII = abdominal segments VIII; ar = apodemal rod of segment VIII (paired); dc = dorsal cleft of segment VIII; ismVIII–X = intersegmental membrane of segment VIII–X; lc = lateral cleft of segment VIII (paired); pp = posterior process; ps = processus spermathecae.	2025-09-10	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
C97B87ACFFA5FF8EFF77FC645B8DF98B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17321861/files/figure.png	https://doi.org/10.5281/zenodo.17321861	FIGURE 16. Distributions of seven species of the Rhyacophila retracta Species Group.	FIGURE 16. Distributions of seven species of the Rhyacophila retracta Species Group.	2025-09-10	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
C97B87ACFFA0FF8DFF77F92C5B83F9D3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17321840/files/figure.png	https://doi.org/10.5281/zenodo.17321840	FIGURE 1. Three new species of the Rhyacophila retracta Species Group. 1A, Rhyacophila geranos sp. nov., male; 1B, same, female; 1C, Rhyacophila humerigera sp. nov., male; 1D, same, female; 1E, Rhyacophila koryne sp. nov., male (photographed by K. Inazu); 1F, same, female (same by K. Inazu).	FIGURE 1. Three new species of the Rhyacophila retracta Species Group. 1A, Rhyacophila geranos sp. nov., male; 1B, same, female; 1C, Rhyacophila humerigera sp. nov., male; 1D, same, female; 1E, Rhyacophila koryne sp. nov., male (photographed by K. Inazu); 1F, same, female (same by K. Inazu).	2025-09-10	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
C97B87ACFFA0FF8DFF77F92C5B83F9D3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17321846/files/figure.png	https://doi.org/10.5281/zenodo.17321846	FIGURE 4. Rhyacophila humerigera sp. nov., male genitalia. 4A, dorsal (left inferior appendage omitted); 4Aa, variation of apicodorsal lobe; 4B, ventral (left inferior appendage omitted); 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, segment X, dorsal; 4F, anal sclerite, apical band and tergal band, ventral; 4G, phallic apparatus, dorsal.Abbreviations: IX, X = abdominal segments IX, X; ab = apical band; adl = apicodorsal lobe; aed = aedeagus; as = anal sclerite; asia = apical segment of inferior appendage (paired); bsia = basal segment of inferior appendage (paired); dap = dorsal appendage of phallic apparatus; par = paramere (paired); pht = phallotheca; tb = tergal band.	FIGURE 4. Rhyacophila humerigera sp. nov., male genitalia. 4A, dorsal (left inferior appendage omitted); 4Aa, variation of apicodorsal lobe; 4B, ventral (left inferior appendage omitted); 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, segment X, dorsal; 4F, anal sclerite, apical band and tergal band, ventral; 4G, phallic apparatus, dorsal.Abbreviations: IX, X = abdominal segments IX, X; ab = apical band; adl = apicodorsal lobe; aed = aedeagus; as = anal sclerite; asia = apical segment of inferior appendage (paired); bsia = basal segment of inferior appendage (paired); dap = dorsal appendage of phallic apparatus; par = paramere (paired); pht = phallotheca; tb = tergal band.	2025-09-10	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
C97B87ACFFA0FF8DFF77F92C5B83F9D3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17321848/files/figure.png	https://doi.org/10.5281/zenodo.17321848	FIGURE 5. Rhyacophila humerigera sp. nov., female genitalia. 5A, segment VIII, dorsal; 5B, same, ventral; 5C, same, left lateral; 5D, vaginal apparatus, left lateral; 5E, same, ventral. Abbreviations: VIII = abdominal segments VIII; dc = dorsal cleft of segment VIII; ismVIII–X = intersegmental membrane of segment VIII–X; lc = lateral cleft of segment VIII (paired); pp = posterior process; ps = processus spermathecae.	FIGURE 5. Rhyacophila humerigera sp. nov., female genitalia. 5A, segment VIII, dorsal; 5B, same, ventral; 5C, same, left lateral; 5D, vaginal apparatus, left lateral; 5E, same, ventral. Abbreviations: VIII = abdominal segments VIII; dc = dorsal cleft of segment VIII; ismVIII–X = intersegmental membrane of segment VIII–X; lc = lateral cleft of segment VIII (paired); pp = posterior process; ps = processus spermathecae.	2025-09-10	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
C97B87ACFFA0FF8DFF77F92C5B83F9D3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17321861/files/figure.png	https://doi.org/10.5281/zenodo.17321861	FIGURE 16. Distributions of seven species of the Rhyacophila retracta Species Group.	FIGURE 16. Distributions of seven species of the Rhyacophila retracta Species Group.	2025-09-10	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
C97B87ACFFA0FF8DFF77F92C5B83F9D3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17321859/files/figure.png	https://doi.org/10.5281/zenodo.17321859	FIGURES 12–15. Female genitalia of Rhyacophila spp. 12A–12D, R. retracta Martynov 1914. 13A–13D, R. motakanta Schmid 1970. 14A–14D, R. lambakanta Schmid 1970. 15A–15D, R. orthacantha Emoto 1979. A, segment VIII, dorsal; B, same, lateral; C, vaginal apparatus, ventral; D, same, lateral. Arrows see text.	FIGURES 12–15. Female genitalia of Rhyacophila spp. 12A–12D, R. retracta Martynov 1914. 13A–13D, R. motakanta Schmid 1970. 14A–14D, R. lambakanta Schmid 1970. 15A–15D, R. orthacantha Emoto 1979. A, segment VIII, dorsal; B, same, lateral; C, vaginal apparatus, ventral; D, same, lateral. Arrows see text.	2025-09-10	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
C97B87ACFFA3FF80FF77F9955B83FB83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17321840/files/figure.png	https://doi.org/10.5281/zenodo.17321840	FIGURE 1. Three new species of the Rhyacophila retracta Species Group. 1A, Rhyacophila geranos sp. nov., male; 1B, same, female; 1C, Rhyacophila humerigera sp. nov., male; 1D, same, female; 1E, Rhyacophila koryne sp. nov., male (photographed by K. Inazu); 1F, same, female (same by K. Inazu).	FIGURE 1. Three new species of the Rhyacophila retracta Species Group. 1A, Rhyacophila geranos sp. nov., male; 1B, same, female; 1C, Rhyacophila humerigera sp. nov., male; 1D, same, female; 1E, Rhyacophila koryne sp. nov., male (photographed by K. Inazu); 1F, same, female (same by K. Inazu).	2025-09-10	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
C97B87ACFFA3FF80FF77F9955B83FB83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17321850/files/figure.png	https://doi.org/10.5281/zenodo.17321850	FIGURE 6. Rhyacophila koryne sp. nov., male genitalia. 6A, dorsal (left inferior appendage omitted); 6B, ventral (left inferior appendage omitted); 6C, left lateral; 6D, right inferior appendage, mesal; 6E, segment X and phallic apparatus, left lateral; 6F, segment X, dorsal; 6G, anal sclerite, ventral; 6H, phallic apparatus, dorsal. Abbreviations: IX, X = abdominal segments IX, X; ab = apical band; adl = apicodorsal lobe; aed = aedeagus; as = anal sclerite; asia = apical segment of inferior appendage (paired); bsia = basal segment of inferior appendage (paired); dap = dorsal appendage of phallic apparatus; par = paramere (paired); pht = phallotheca; tb = tergal band.	FIGURE 6. Rhyacophila koryne sp. nov., male genitalia. 6A, dorsal (left inferior appendage omitted); 6B, ventral (left inferior appendage omitted); 6C, left lateral; 6D, right inferior appendage, mesal; 6E, segment X and phallic apparatus, left lateral; 6F, segment X, dorsal; 6G, anal sclerite, ventral; 6H, phallic apparatus, dorsal. Abbreviations: IX, X = abdominal segments IX, X; ab = apical band; adl = apicodorsal lobe; aed = aedeagus; as = anal sclerite; asia = apical segment of inferior appendage (paired); bsia = basal segment of inferior appendage (paired); dap = dorsal appendage of phallic apparatus; par = paramere (paired); pht = phallotheca; tb = tergal band.	2025-09-10	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
C97B87ACFFA3FF80FF77F9955B83FB83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17321853/files/figure.png	https://doi.org/10.5281/zenodo.17321853	FIGURE 7. Rhyacophila koryne sp. nov., female genitalia. 7A, segment VIII, dorsal; 7B, same, ventral; 7C, same, left lateral; 7D, vaginal apparatus, left lateral; 7E, same ventral. Abbreviations: VIII = abdominal segments VIII; dc = dorsal cleft of segment VIII; ismVIII–X = intersegmental membrane of segment VIII–X; lc = lateral cleft of segment VIII (paired); pp = posterior process; ps = processus spermathecae.	FIGURE 7. Rhyacophila koryne sp. nov., female genitalia. 7A, segment VIII, dorsal; 7B, same, ventral; 7C, same, left lateral; 7D, vaginal apparatus, left lateral; 7E, same ventral. Abbreviations: VIII = abdominal segments VIII; dc = dorsal cleft of segment VIII; ismVIII–X = intersegmental membrane of segment VIII–X; lc = lateral cleft of segment VIII (paired); pp = posterior process; ps = processus spermathecae.	2025-09-10	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
C97B87ACFFA3FF80FF77F9955B83FB83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17321861/files/figure.png	https://doi.org/10.5281/zenodo.17321861	FIGURE 16. Distributions of seven species of the Rhyacophila retracta Species Group.	FIGURE 16. Distributions of seven species of the Rhyacophila retracta Species Group.	2025-09-10	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
C97B87ACFFA3FF80FF77F9955B83FB83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17321857/files/figure.png	https://doi.org/10.5281/zenodo.17321857	FIGURES 8–11. Male genitalia of Rhyacophila spp. 8A–8C, R. retracta Martynov 1914. 9A–9C, R. motakanta Schmid 1970. 10A–10C, R. lambakanta Schmid 1970. 11A–11C, R. orthacantha Emoto 1979. A, dorsal; B, lateral; C, segment X and apical part of anal sclerite, dorsal.	FIGURES 8–11. Male genitalia of Rhyacophila spp. 8A–8C, R. retracta Martynov 1914. 9A–9C, R. motakanta Schmid 1970. 10A–10C, R. lambakanta Schmid 1970. 11A–11C, R. orthacantha Emoto 1979. A, dorsal; B, lateral; C, segment X and apical part of anal sclerite, dorsal.	2025-09-10	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
C97B87ACFFAEFF86FF77FB255B9DF808.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17321857/files/figure.png	https://doi.org/10.5281/zenodo.17321857	FIGURES 8–11. Male genitalia of Rhyacophila spp. 8A–8C, R. retracta Martynov 1914. 9A–9C, R. motakanta Schmid 1970. 10A–10C, R. lambakanta Schmid 1970. 11A–11C, R. orthacantha Emoto 1979. A, dorsal; B, lateral; C, segment X and apical part of anal sclerite, dorsal.	FIGURES 8–11. Male genitalia of Rhyacophila spp. 8A–8C, R. retracta Martynov 1914. 9A–9C, R. motakanta Schmid 1970. 10A–10C, R. lambakanta Schmid 1970. 11A–11C, R. orthacantha Emoto 1979. A, dorsal; B, lateral; C, segment X and apical part of anal sclerite, dorsal.	2025-09-10	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
C97B87ACFFAEFF86FF77FB255B9DF808.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17321859/files/figure.png	https://doi.org/10.5281/zenodo.17321859	FIGURES 12–15. Female genitalia of Rhyacophila spp. 12A–12D, R. retracta Martynov 1914. 13A–13D, R. motakanta Schmid 1970. 14A–14D, R. lambakanta Schmid 1970. 15A–15D, R. orthacantha Emoto 1979. A, segment VIII, dorsal; B, same, lateral; C, vaginal apparatus, ventral; D, same, lateral. Arrows see text.	FIGURES 12–15. Female genitalia of Rhyacophila spp. 12A–12D, R. retracta Martynov 1914. 13A–13D, R. motakanta Schmid 1970. 14A–14D, R. lambakanta Schmid 1970. 15A–15D, R. orthacantha Emoto 1979. A, segment VIII, dorsal; B, same, lateral; C, vaginal apparatus, ventral; D, same, lateral. Arrows see text.	2025-09-10	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
C97B87ACFFAEFF86FF77FB255B9DF808.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17321861/files/figure.png	https://doi.org/10.5281/zenodo.17321861	FIGURE 16. Distributions of seven species of the Rhyacophila retracta Species Group.	FIGURE 16. Distributions of seven species of the Rhyacophila retracta Species Group.	2025-09-10	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
C97B87ACFFA9FF87FF77FF7C5BD6FA73.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17321857/files/figure.png	https://doi.org/10.5281/zenodo.17321857	FIGURES 8–11. Male genitalia of Rhyacophila spp. 8A–8C, R. retracta Martynov 1914. 9A–9C, R. motakanta Schmid 1970. 10A–10C, R. lambakanta Schmid 1970. 11A–11C, R. orthacantha Emoto 1979. A, dorsal; B, lateral; C, segment X and apical part of anal sclerite, dorsal.	FIGURES 8–11. Male genitalia of Rhyacophila spp. 8A–8C, R. retracta Martynov 1914. 9A–9C, R. motakanta Schmid 1970. 10A–10C, R. lambakanta Schmid 1970. 11A–11C, R. orthacantha Emoto 1979. A, dorsal; B, lateral; C, segment X and apical part of anal sclerite, dorsal.	2025-09-10	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
C97B87ACFFA9FF87FF77FF7C5BD6FA73.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17321859/files/figure.png	https://doi.org/10.5281/zenodo.17321859	FIGURES 12–15. Female genitalia of Rhyacophila spp. 12A–12D, R. retracta Martynov 1914. 13A–13D, R. motakanta Schmid 1970. 14A–14D, R. lambakanta Schmid 1970. 15A–15D, R. orthacantha Emoto 1979. A, segment VIII, dorsal; B, same, lateral; C, vaginal apparatus, ventral; D, same, lateral. Arrows see text.	FIGURES 12–15. Female genitalia of Rhyacophila spp. 12A–12D, R. retracta Martynov 1914. 13A–13D, R. motakanta Schmid 1970. 14A–14D, R. lambakanta Schmid 1970. 15A–15D, R. orthacantha Emoto 1979. A, segment VIII, dorsal; B, same, lateral; C, vaginal apparatus, ventral; D, same, lateral. Arrows see text.	2025-09-10	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
C97B87ACFFA9FF87FF77FF7C5BD6FA73.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17321861/files/figure.png	https://doi.org/10.5281/zenodo.17321861	FIGURE 16. Distributions of seven species of the Rhyacophila retracta Species Group.	FIGURE 16. Distributions of seven species of the Rhyacophila retracta Species Group.	2025-09-10	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
C97B87ACFFA9FF85FF77FA345895FB83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17321857/files/figure.png	https://doi.org/10.5281/zenodo.17321857	FIGURES 8–11. Male genitalia of Rhyacophila spp. 8A–8C, R. retracta Martynov 1914. 9A–9C, R. motakanta Schmid 1970. 10A–10C, R. lambakanta Schmid 1970. 11A–11C, R. orthacantha Emoto 1979. A, dorsal; B, lateral; C, segment X and apical part of anal sclerite, dorsal.	FIGURES 8–11. Male genitalia of Rhyacophila spp. 8A–8C, R. retracta Martynov 1914. 9A–9C, R. motakanta Schmid 1970. 10A–10C, R. lambakanta Schmid 1970. 11A–11C, R. orthacantha Emoto 1979. A, dorsal; B, lateral; C, segment X and apical part of anal sclerite, dorsal.	2025-09-10	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
C97B87ACFFA9FF85FF77FA345895FB83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17321859/files/figure.png	https://doi.org/10.5281/zenodo.17321859	FIGURES 12–15. Female genitalia of Rhyacophila spp. 12A–12D, R. retracta Martynov 1914. 13A–13D, R. motakanta Schmid 1970. 14A–14D, R. lambakanta Schmid 1970. 15A–15D, R. orthacantha Emoto 1979. A, segment VIII, dorsal; B, same, lateral; C, vaginal apparatus, ventral; D, same, lateral. Arrows see text.	FIGURES 12–15. Female genitalia of Rhyacophila spp. 12A–12D, R. retracta Martynov 1914. 13A–13D, R. motakanta Schmid 1970. 14A–14D, R. lambakanta Schmid 1970. 15A–15D, R. orthacantha Emoto 1979. A, segment VIII, dorsal; B, same, lateral; C, vaginal apparatus, ventral; D, same, lateral. Arrows see text.	2025-09-10	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
C97B87ACFFA9FF85FF77FA345895FB83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17321861/files/figure.png	https://doi.org/10.5281/zenodo.17321861	FIGURE 16. Distributions of seven species of the Rhyacophila retracta Species Group.	FIGURE 16. Distributions of seven species of the Rhyacophila retracta Species Group.	2025-09-10	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
C97B87ACFFA9FF85FF77FA345895FB83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17321848/files/figure.png	https://doi.org/10.5281/zenodo.17321848	FIGURE 5. Rhyacophila humerigera sp. nov., female genitalia. 5A, segment VIII, dorsal; 5B, same, ventral; 5C, same, left lateral; 5D, vaginal apparatus, left lateral; 5E, same, ventral. Abbreviations: VIII = abdominal segments VIII; dc = dorsal cleft of segment VIII; ismVIII–X = intersegmental membrane of segment VIII–X; lc = lateral cleft of segment VIII (paired); pp = posterior process; ps = processus spermathecae.	FIGURE 5. Rhyacophila humerigera sp. nov., female genitalia. 5A, segment VIII, dorsal; 5B, same, ventral; 5C, same, left lateral; 5D, vaginal apparatus, left lateral; 5E, same, ventral. Abbreviations: VIII = abdominal segments VIII; dc = dorsal cleft of segment VIII; ismVIII–X = intersegmental membrane of segment VIII–X; lc = lateral cleft of segment VIII (paired); pp = posterior process; ps = processus spermathecae.	2025-09-10	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
C97B87ACFFABFF85FF77FB255BE7F82C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17321857/files/figure.png	https://doi.org/10.5281/zenodo.17321857	FIGURES 8–11. Male genitalia of Rhyacophila spp. 8A–8C, R. retracta Martynov 1914. 9A–9C, R. motakanta Schmid 1970. 10A–10C, R. lambakanta Schmid 1970. 11A–11C, R. orthacantha Emoto 1979. A, dorsal; B, lateral; C, segment X and apical part of anal sclerite, dorsal.	FIGURES 8–11. Male genitalia of Rhyacophila spp. 8A–8C, R. retracta Martynov 1914. 9A–9C, R. motakanta Schmid 1970. 10A–10C, R. lambakanta Schmid 1970. 11A–11C, R. orthacantha Emoto 1979. A, dorsal; B, lateral; C, segment X and apical part of anal sclerite, dorsal.	2025-09-10	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
C97B87ACFFABFF85FF77FB255BE7F82C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17321859/files/figure.png	https://doi.org/10.5281/zenodo.17321859	FIGURES 12–15. Female genitalia of Rhyacophila spp. 12A–12D, R. retracta Martynov 1914. 13A–13D, R. motakanta Schmid 1970. 14A–14D, R. lambakanta Schmid 1970. 15A–15D, R. orthacantha Emoto 1979. A, segment VIII, dorsal; B, same, lateral; C, vaginal apparatus, ventral; D, same, lateral. Arrows see text.	FIGURES 12–15. Female genitalia of Rhyacophila spp. 12A–12D, R. retracta Martynov 1914. 13A–13D, R. motakanta Schmid 1970. 14A–14D, R. lambakanta Schmid 1970. 15A–15D, R. orthacantha Emoto 1979. A, segment VIII, dorsal; B, same, lateral; C, vaginal apparatus, ventral; D, same, lateral. Arrows see text.	2025-09-10	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
C97B87ACFFABFF85FF77FB255BE7F82C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17321861/files/figure.png	https://doi.org/10.5281/zenodo.17321861	FIGURE 16. Distributions of seven species of the Rhyacophila retracta Species Group.	FIGURE 16. Distributions of seven species of the Rhyacophila retracta Species Group.	2025-09-10	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
