identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
C97B87ACFFA5FF8BFF77FEA15892FCF5.text	C97B87ACFFA5FF8BFF77FEA15892FCF5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyacophila retracta Martynov 1914	<div><p>Rhyacophila retracta Species Group</p><p>Rhyacophila retracta Species Group Ross 1956, 102–103; Schmid 1970, 89–90, 132; Emoto 1979, 556–560; Hattori 2005, 422, 431.</p><p>The males of this species group share the following characters: Segment IX is short ventrally and the long dorsal part bears a large apicodorsal lobe; preanal appendage absent; segment X is oblique downward and bears a longitudinal mesal groove or crack dorsally; a pair of pigmented anal sclerites are fused and form a single board with a distinctive long root; and the dorsal appendage of the phallic apparatus is present with the basal 3/4 being fused with the aedeagus. Males of the R. retracta Species Group are most similar to those of the R. lieftincki Species Group in the shapes of the large, horizontal apicodorsal lobe of segment IX, the slanting segment X, and the subtriangular apical part of the inferior appendage, but can be distinguished from the latter by the presence of the dorsal appendage of the phallic apparatus.</p></div>	https://treatment.plazi.org/id/C97B87ACFFA5FF8BFF77FEA15892FCF5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kawase, Naoki	Kawase, Naoki (2025): Three new species of the Rhyacophila retracta species group (Trichoptera, Rhyacophilidae) from Japan. Zootaxa 5691 (3): 561-578, DOI: 10.11646/zootaxa.5691.3.6, URL: https://doi.org/10.11646/zootaxa.5691.3.6
C97B87ACFFA5FF8EFF77FC645B8DF98B.text	C97B87ACFFA5FF8EFF77FC645B8DF98B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyacophila geranos Kawase 2025	<div><p>Rhyacophila geranos sp. nov.</p><p>(Figs 1A, 1B, 2A–2H, 3A–3E, 16C)</p><p>Rhyacophila sp. 15: Yamamoto et al. 2021, 70, list.</p><p>Diagnosis. Male and female genitalia of this species are very distinctive among the R. retracta Species Group. The male genitalia are distinguishable from those of other species by the following two remarkable characters: The apicodorsal lobe of segment IX bears two pairs of stout and acute projections and segment X curves sharply posteroventrally (Figs 2A, 2C). In females, a pair of lateral clefts of segment VIII are situated rather ventrally and these clefts can be seen in ventral view (Fig. 3C). Moreover, the apical half of the posterior process of the vaginal apparatus is acicular in this species (Figs 3D, 3E).</p><p>Adult (Figs 1A, 1B). Length of each forewing: male 9.0–11.4 mm (mean = 10.4, SD = 0.70, n = 20), female 9.5–13.5 mm (mean =11.9, SD = 0.94, n = 16). Forewings pale yellowish brown in male, yellowish brown in female, with irregular brown reticulation as in Figs 1A, 1B; assembly of rather clear pale patches at middle of hind margin in male and female (Figs 1A, 1B). Abdomen with small midventral process on posterior margin of sternites VI and VII in male, on that of sternite VI in female.</p><p>Male genitalia (Figs 2A–2H). Segment IX (IX) very short ventrally and long dorsally in lateral view (Fig. 2C), rectangular in dorsal view (Fig. 2A). Apicodorsal lobe (adl) large, widely spread and covered basal part of segment X, with two pairs of acute, finger-like projections, middle pair longer than lateral one (Figs 2A, 2C). Segment X (X) long, extending posteroventrad, bend 90 degrees in middle, like crane neck in lateral view (Fig. 2E); with pair of dorsolateral ridges forming longitudinal deep groove, ridges bearing raw of setae (Fig. 2F); apex dark pigmented, with median incision, each posterolateral process (plp) slightly expanded posterolaterad (Fig. 2F). Pair of anal sclerites (as) fused together, bean shaped in ventral view, dark pigmented, with long common root (Figs 2E–2G). Apical band (ab) sclerotized, u-shaped in ventral view, and fused with tergal band basally; tergal band (tb) well sclerotized apically, connecting to dorsum of phallotheca (pht) anteriorly (Fig. 2E). Basal segment of each inferior appendage (bsia) setose, elongate, basal half slightly thicker in lateral view (Figs 2C, 2D). Apical segment of each inferior appendage (asia) subtriangular, posterodorsal margin weakly concaved in lateral view (Figs 2C, 2D), bearing spinules mesally (Fig. 2D). In phallic apparatus, phallotheca cup-shaped in lateral view (Fig. 2E); aedeagus (aed) long, almost cylindrical tapering to apex (Figs 2B, 2E, 2H); dorsal appendage of phallic apparatus (dap) sclerotized, slightly shorter than aedeagus, basal 3/4 fused with aedeagus, apex truncated in dorsal view (Figs 2E, 2H). Pair of parameres (par) long, basal half thick in lateral view (Fig. 2E), apical half thin, directed posterodorsad with acute apex (Figs 2B, 2E).</p><p>Female genitalia (Figs 3A–3E). Segment VIII (VIII) with dorsal cleft (dc) extremely wide and deep, reaching approximately 2/3 of segment in dorsal view (Fig. 3A); with pair of deep lateral clefts (lc) lateroventrally, each extending to middle part of segment (Figs 3B, 3C); pair of apodemal rods (ar) slightly longer than segment VIII in lateral view (Fig. 3B). Intersegmental membrane of segment VIII–X (ismVIII–X) deeply invaginated into segment VIII, slightly sclerotized, anterior part fixed to interior wall of segment VIII (Figs 3A–3C). In vaginal apparatus, posterior process (pp) long, sclerotized, apical half extremely thin (Figs 3D, 3E), basal half thick, subtriangular in lateral view, slightly curved anteroventrad (Fig. 3D); processus spermathecae (ps) small, subrectangular in lateral view, covered by basal part of posterior process (Fig. 3D).</p><p>Immature stage. Unknown.</p><p>Holotype. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=132.91779&amp;materialsCitation.latitude=33.560276" title="Search Plazi for locations around (long 132.91779/lat 33.560276)">Male</a> (in alcohol): <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=132.91779&amp;materialsCitation.latitude=33.560276" title="Search Plazi for locations around (long 132.91779/lat 33.560276)">Namakusa-dani</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=132.91779&amp;materialsCitation.latitude=33.560276" title="Search Plazi for locations around (long 132.91779/lat 33.560276)">Odamiyama</a>, Uchiko-cho, Ehime Prefecture, Shikoku, Japan, 33°33’37” N 132°55’04” E, alt. 1225 m, 11–20.vi.2020, E. Yamamoto (LBM1410012659).</p><p>Paratypes. Shikoku: Ehime : 1 female (pinned), Yonaru-kawa, Yanaigawa, Kuma-kôgen-cho, 25.v.2011, T. Hattori (SPMN); 1 female, same data as the holotype (LBM1410012660); 2 males, 4 females, type locality, 21.vi.1999, E. Yamamoto (SPMN); 6 males, 3 females, type locality, 5.vi.2000, E. Yamamoto (SPMN); 1 female, type locality, 8.viii.2000, E. Yamamoto (SPMN); 2 males, 1 female, type locality, 20.ix.2000, E. Yamamoto (SPMN); 1 female, Honomata, Odamiyama, Uchiko-cho, 1–10.vi.2018, E. Yamamoto (LBM1410012661); 1 male, same locality, 11–20.vi.2018, E. Yamamoto (LBM1410012662); 1 male (pinned, fig. 1A), 1 female (pinned, fig. 1B), Mitaki-keikoku, Kubono, Shirokawa-cho, Seiyo-shi, 22.v.2011, T. Hattori (SPMN) .</p><p>Other specimens examined. Shikoku: Ehime: 2 females, Kanmon, Omogo, Kuma-kôgen-cho, 23–25.v.1999, A. Ohkawa &amp; T. Ito (SPMN) ; 4 males, 6 females, same data as the holotype, 11–20.vi.2020, E. Yamamoto (Nka); 1 male, Hontani, Odamiyama, Uchiko-cho, 5–8.v.2000, E. Yamamoto (SPMN) ; 1 female, same locality, 19.v.2000, E. Yamamoto (SPMN); 1 female, Tarôdani, Odamiyama, Uchiko-cho, 20.vi.2000, E. Yamamoto (SPMN) . Kochi: 1 male, Sakihama-gawa, Muroto-shi, 29.iv.2004, M. Takai (SPMN) ; 2 males, Ioki-gawa, Bettyaku, Aki-shi, 25.iv.2004, K. Nio (SPMN) ; 1 female, Nishikuma-keikoku, Kubo-wakubo, Monobe-cho, Kami-shi, 25.vi.2003, K. Nio (SPMN) ; 1 male, Befu-kyo, Monobe-cho, Kami-shi, 28.v.2004, M. Takai (SPMN) .</p><p>Distribution and biology. Rhyacophila geranos sp. nov. is distributed only in Shikoku, one of Japanese main islands (Fig. 16C). The adults were collected near mountain streams in late April to September by sweeping, Malaise traps or light traps.</p><p>Etymology. The species epithet “ geranos ” (= crane) is a Greek noun derived from the crane neck-like segment X in lateral view, and crane wing-like apicodorsal lobe of segment IX in dorsal view.</p><p>Japanese name. Tsuruo-nagare-tobikera.</p></div>	https://treatment.plazi.org/id/C97B87ACFFA5FF8EFF77FC645B8DF98B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kawase, Naoki	Kawase, Naoki (2025): Three new species of the Rhyacophila retracta species group (Trichoptera, Rhyacophilidae) from Japan. Zootaxa 5691 (3): 561-578, DOI: 10.11646/zootaxa.5691.3.6, URL: https://doi.org/10.11646/zootaxa.5691.3.6
C97B87ACFFA0FF8DFF77F92C5B83F9D3.text	C97B87ACFFA0FF8DFF77F92C5B83F9D3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyacophila humerigera Kawase 2025	<div><p>Rhyacophila humerigera sp. nov.</p><p>(Figs 1C, 1D, 4A–4G, 5A–5E, 16D)</p><p>Rhyacophila sp. 2: Tanaka 2012, 20–22, male; Tanaka 2014, 38, list.</p><p>Rhyacophila sp.: Tanaka 2022, 34, list.</p><p>Rhyacophila sp. aff. motakanta Schmid 1970: Aoya &amp; Tanaka 2023, 26, list.</p><p>Diagnosis. The male genitalia of R. humerigera sp. nov. are somewhat similar to those of R. koryne sp. nov., R. motakanta and R. orthacantha in having the apically bifid apicodorsal lobe of segment IX. The difference is, however, considerable. R. humerigera sp. nov. bears a pair of lateral shoulders in the middle of the apicodorsal lobe (Figs 4A, 4 Aa), whereas other three species bear the simple, short apicodorsal lobe without such a pair of shoulders. Regarding the female genitalia, R. humerigera sp. nov. is similar to R. motakanta Schmid 1970 and R. lambakanta Schmid 1970 in bearing a medium-sized dorsal and a pair of lateral clefts in segment VIII (Figs 5A–5C, 13A–13B, 14A–14B), but this species can be distinguished from the latter by the shape of the posterior processes of the vaginal apparatus: In R. humerigera, the apical part of the posterior process is simply round (Figs 5D, 5E), but in R. motakanta, the apical part of that decreases abruptly in width and height (arrows in figs 13C, 13D); in R. humerigera, the vaginal apparatus bends approximately 45 degrees in lateral view (Fig. 5D), but in R. lambakanta, it bends 90 degrees in the middle in lateral view (arrows in fig. 14D).</p><p>Adult (Figs 1C, 1D). Length of each forewing: male 8.2–10.4 mm (mean = 9.3, SD = 0.77, n = 14), female 10.0–11.3 mm (mean =10.7, SD = 0.50, n = 5). General morphology of R. humerigera sp. nov. similar to those of R. geranos sp. nov., but forewings of this species with many brown reticulations all over (Figs 1C, 1D). In abdomen, small mid ventral process on posterior margin of sternites VI and VII in male, on that of sternite VI in female.</p><p>Male genitalia (Figs 4A–4G). Segment IX (IX) in lateral view, very short ventrally, long dorsally, widely rounded posterolaterally (Fig. 4C); subquadrate in dorsal view, with pair of clefts in both sides of apicodorsal lobe (Fig. 4A). In dorsal view, apicodorsal lobe (adl) long, extending posterad, bifid apically, with pair of pointed shoulders laterally (Fig. 4A) (three specimens in Iwate Prefecture bear pair of rounded shoulders laterally (Fig. 4 Aa)); in lateral view apicodorsal lobe like sharp claws (Fig. 4C). Segment X (X) long, straight, extending posteroventrad in lateral view (Figs 4C, 4D); bilobed apically in dorsal view, each apex round (Fig. 4E). Pair of anal sclerites (as) fused together, wide, round in ventral view, with pair of dark pigmented protuberances apically (Fig. 4F), and with long common root basally.Apical band (ab) sclerotized, u-shaped in ventral view and fused with tergal band basally (Fig. 4F); tergal band (tb) wide, flattened, well sclerotized apically, connecting to dorsum of phallotheca anteriorly (Fig. 4D). Basal segment of each inferior appendage setose, elongate, basal half slightly thicker (Figs 4B, 4C). Apical segment of each inferior appendage rounded, right trapezoid in left lateral view (Fig. 4C), apical half bearing spinules mesally. In phallic apparatus, phallotheca (pht) cup-shaped in lateral view (Fig. 4D); aedeagus (aed) long, almost cylindrical, tapering to apex (Figs 4D, 4G); dorsal appendage of phallic apparatus (dap) sclerotized, shorter than aedeagus, basal 3/4 fused with aedeagus, apical margin slightly rounded, protruded mesally (Figs 4D, 4G). Pair of parameres (par) long cylindrical, each tapering to apex, dilated subapically, bearing many small spines ventrally around mid-length (Figs 4B, 4D).</p><p>Female genitalia (Figs 5A–5E). Segment VIII (VIII) with dorsal cleft (dc) wide and deep in dorsal view (Fig. 5A), with pair of deep lateral clefts (lc) extending to middle part of segment (Fig. 5B), and with posteroventral margin rounded, semicircular (Fig. 5C). Intersegmental membrane of segment VIII–X (ismVIII–X) deeply invaginated into segment VIII, slightly sclerotized, dorsally into anterior margin, ventrally into middle part of segment VIII, anterior part fixed to interior wall of segment VIII (Figs 5A–5C). In vaginal apparatus, posterior process (pp) sclerotized, long bottle shape in ventral view (Fig. 5E), anterior half curved ventrad (Fig. 5D); processus spermathecae (ps) small, long oval in lateral view, covered by basal part of posterior process (Fig. 5D).</p><p>Immature stage. Unknown.</p><p>Holotype. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=140.7061&amp;materialsCitation.latitude=39.93889" title="Search Plazi for locations around (long 140.7061/lat 39.93889)">Male</a> (in alcohol): <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=140.7061&amp;materialsCitation.latitude=39.93889" title="Search Plazi for locations around (long 140.7061/lat 39.93889)">Shibukuro-gawa</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=140.7061&amp;materialsCitation.latitude=39.93889" title="Search Plazi for locations around (long 140.7061/lat 39.93889)">Tazawako-tamagawa</a>, Senboku-shi, Akita Prefecture, Honshu, Japan, 39°56’20” N 140°42’22” E, alt. 485 m, 18–25.vi.2018, K. Aoya (LBM1410012663).</p><p>Paratypes. Honshu: Iwate: 1 male (pinned), 1 female (pinned, fig. 1D), Seiren-no-taki, Uchikawame, Hanamaki-shi, 10.viii.1994, T. Hattori (SPMN) ; 1 male, 1 female, tributary Tobetsu-gawa, Yamagata-cho, Kuji-shi, 12.vi.2004, N. Kuhara (LBM1410012664–1410012665) . Akita: 2 males (pinned, fig. 1C), Toroko, Hachimantai, Kazuno-shi, 4.vii.2019, M. Ookura (SPMN); 1 male, type locality, 28.ix–5.x.2018, K. Aoya (LBM1410012666); 1 male, type locality, 5–13.x.2018, K. Aoya (LBM1410012667) .</p><p>Other specimens examined. Honshu: Aomori: 1 male, 1 female, Akaishi-gawa, Komori-machi, Ajigasawamachi, 6.vi.1982, M. Kobayashi (KPM-NKMK71010) ; 1male, Sasanai-gawa, Iwasaki, Fukaura-machi, 7.vi.1982, M. Kobayashi (KPM-NKMK71080) . Iwate: 4 males, 1 female, Matsukusa, Kuzakai, Miyako-shi, 12.vii.1997, N. Kuhara (KuN-14991) (NKa) ; 1 male, 3 females, same data, N. Kuhara (KuN-14995) (NKa); 6 males, Oyama-gawa, Kadoma, Miyako-shi, 13.vii.1997, N. Kuhara (KuN-14982) (NKu) . Akita: 1 male (pinned), Koirikawa, Hachimori, Happou-cho, 14.vi.2019, M. Ookura (MTa) ; 1 male (pinned), Numanosawa, Tazawako, Senboku-shi, 24.ix.2013, M. Tanaka (MTa) ; 1 male (pinned), Kotaki-zawa, Nakasen, Daisen-shi, 13.vii.2010, M. Tanaka (MTa) .</p><p>Distribution and biology. Rhyacophila humerigera sp. nov. is distributed in northern part of Tohoku District in Honshu (Fig. 16D). Adults were collected near mountain streams in June to October by sweeping, Malaise traps or light traps.</p><p>Etymology. The Latin species epithet “humerigera ” (Latin adjective, with a pair of shoulders) is derived from the shape of the apicodorsal lobe of the segment IX in male genitalia: a pair of shoulders on both sides of the apicodorsal lobe.</p><p>Japanese name. Kadoo-nagare-tobikera.</p></div>	https://treatment.plazi.org/id/C97B87ACFFA0FF8DFF77F92C5B83F9D3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kawase, Naoki	Kawase, Naoki (2025): Three new species of the Rhyacophila retracta species group (Trichoptera, Rhyacophilidae) from Japan. Zootaxa 5691 (3): 561-578, DOI: 10.11646/zootaxa.5691.3.6, URL: https://doi.org/10.11646/zootaxa.5691.3.6
C97B87ACFFA3FF80FF77F9955B83FB83.text	C97B87ACFFA3FF80FF77F9955B83FB83.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyacophila koryne Kawase 2025	<div><p>Rhyacophila koryne sp. nov.</p><p>(Figs 1E, 1F, 6A–6H, 7A–7E, 16D)</p><p>Rhyacophila sp. 3: Inazu 2008, 66, list; Inazu &amp; Nishida 2011, 177, habitus, male, female.</p><p>Rhyacophila sp. A: Nojima 2017, 111, list.</p><p>Diagnosis. The male genitalia of R. koryne sp. nov. are slightly similar to those of R. motakanta and R. orthacantha in having the apically bifid, straight apicodorsal lobe of segment IX, but R. koryne sp. nov. is easily distinguished from the latter by the shape of the apicodorsal lobe which is long, club-shaped in lateral view (Fig. 6C). On the other hand, the shapes of the apicodorsal lobes of the other two species are short, acute apically (not club-shaped) in lateral view (Figs 9B, 11B). Female genitalia of this species are very similar to those of R. orthacantha (Figs</p><p>7A–7E, 15A–15D), and both species cannot be distinguished from each other in the present study. However, the distributions of both species are clearly allopatric (Fig. 16D).</p><p>Adult (Figs 1E, 1F). Length of each forewing: male 8.6–12.3 mm (mean = 10.8, SD = 0.89, n = 21), female 11.8–13.0 mm (mean = 12.5, SD = 0.52, n = 5). Forewing coloration of R. koryne sp. nov. (Figs 1E, 1F) similar to that of R. humerigera sp. nov. Small mesal ventral process on posterior margin of sternites VI and VII in male and sternite VI in female abdomens.</p><p>Male genitalia (Figs 6A–6H). Segment IX in lateral view, very short ventrally, long dorsally and widely rounded posterolaterally (Fig. 6C), subquadrate in dorsal view, with pair of clefts in both sides of apicodorsal lobe (Fig. 6A); apicodorsal lobe long triangular with bifid apex in dorsal view (Fig. 6A), long club shaped in lateral view (Fig. 6C). Segment X long, extending posteroventrad in lateral view (Fig. 6E); bilobed apically, each apex blunt in dorsal view (Fig. 6F). Pair of anal sclerites fused together, wide, rounded in ventral view (Fig. 6G), with pair of dark pigmented protuberances apically, and with long common root basally (Fig. 6G). Apical band sclerotized, u-shaped in ventral view, fused with tergal band basally; tergal band wide, flattened, well sclerotized apically, connecting to dorsum of phallotheca anteriorly (Fig. 6E). Basal segment of each inferior appendage setose, elongate, basal half slightly thicker (Figs 6B–6D). Apical segment of each inferior appendage rounded, right trapezoid in left lateral view (Fig. 6B), apical half bearing spinules mesally (Fig. 6D). In phallic apparatus, phallotheca near cup-shaped in lateral view (Fig. 6E); aedeagus long, almost cylindrical tapering to apex (Figs 6B, 6E, 6H); dorsal appendage of phallic apparatus sclerotized, shorter than aedeagus, basal 3/4 fused with aedeagus, apex slightly protruded in middle (Figs 6E, 6H). Parameres long cylindrical, tapering to apex, dilated subapically, with a few small spines ventromesally around mid-length (Figs 6B, 6E).</p><p>Female genitalia (Figs 7A–7E). Segment VIII with dorsal cleft excavated to apical 1/4, tiny protrusion usually present mesally (Fig. 7A), but occasionally absent, with pair of lateral clefts extending to apical 1/3 (Fig. 7B), and posteroventral margin rounded, semicircular (Fig. 7C). Intersegmental membrane of segment VIII–X (ismVIII–X) deeply invaginated into segment VIII, slightly sclerotized, dorsally into anterior margin, ventrally into middle part of it, anterior part fixed to interior wall of segment VIII (Figs 7A–7C). In vaginal apparatus, posterior process sclerotized, long bottle shape in ventral view (Fig. 7E); anterior half slightly curved anteroventrad in lateral view (Fig. 7D); processus spermathecae small, long rectangular in lateral view, covered by basal part of posterior process (Fig. 7D).</p><p>Immature stage. Unknown.</p><p>Holotype. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=133.32889&amp;materialsCitation.latitude=35.01611" title="Search Plazi for locations around (long 133.32889/lat 35.01611)">Male</a> (in alcohol): <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=133.32889&amp;materialsCitation.latitude=35.01611" title="Search Plazi for locations around (long 133.32889/lat 35.01611)">Shingo-yuno</a>, Niimi-shi, Okayama Prefecture, Honshu, Japan, 35°00’58” N, 133°19’44” E, alt. 550 m, 1.vi.2013, K. Nojima (LBM1410012668).</p><p>Paratypes. Honshu: Hyogo : 1 male, 1 female, Amagoike, Takenouchi, Wadayama-cho, Asago-shi, 25.v.2004, K. Inazu (SPMN) . Okayama: 2 males, 1 female, Uno, Kamo-cho, Tsuyama-shi, 6.v.2013, K. Nojima (LBM1410012669–1410012671); 3 males, 1 female, same data as the holotype (LBM1410012672–1410012675); 6 males, 1 female, Koshiwata, Kagamino-cho, 9.vi.2013, K. Nojima (LBM1410012676–1410012682); 2 males, Ogaya, Nishi-awakura-son, 12.v.2016, K. Nojima (LBM1410012683–1410012684) .</p><p>Other specimens examined. Honshu: Hyogo: 1 male, Sugawara, Kishida, Shin-onsen-cho, 30.v.2004, K. Inazu (NKa) ; 1 male (Fig. 1E), 1 female (Fig. 1F), Mihara, Takeno-cho, Toyooka-shi, 9.vi.2009, K. Inazu (NKa) ; 1 female, Kumanami, Muraoka-ku, Kami-cho, 4.vi.2006, K. Inazu (NKa) ; 1 male, Itabami, Nishigouchi, Chikusa-cho, Shisô-shi, 23.v.2016, K. Inazu (NKa) ; 1 female, Mimuro-no-taki, Kouchi, Chikusa-cho, Shisô-shi, 14.vii.2016, K. Inazu (NKa) . Okayama: 6 males, 3 females, Akawase-keikoku, Kami-saibara, Kagamino-cho, 8.vi.2011, K. Nojima (NKa) . Tottori: 1 female, Kachi-gawa, Wakasa-cho, 12.ix.2002, T . Hattori (SPMN) ; 1 male, Yukawagawa, Yukawa, Nichinan-cho, 11.ix.2008, N. Kuhara (KuN-15026) (NKu) . Shimane: 1 male, Sada-cho, Izumo-shi, 15.ix.1993, H. Nishimoto (SPMN) .</p><p>Distribution and biology. Rhyacophila koryne sp. nov. is distributed in western Honshu (Fig. 16D). Adults were collected near mountainous streams in May to September by sweeping or light traps.</p><p>Etymology. The species epithet “koryne ” (Greek noun, club) is derived from the shape of the apicodorsal lobe of the segment IX of male genitalia in lateral view.</p><p>Japanese name. Maruo-nagare-tobikera.</p></div>	https://treatment.plazi.org/id/C97B87ACFFA3FF80FF77F9955B83FB83	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kawase, Naoki	Kawase, Naoki (2025): Three new species of the Rhyacophila retracta species group (Trichoptera, Rhyacophilidae) from Japan. Zootaxa 5691 (3): 561-578, DOI: 10.11646/zootaxa.5691.3.6, URL: https://doi.org/10.11646/zootaxa.5691.3.6
C97B87ACFFAEFF86FF77FB255B9DF808.text	C97B87ACFFAEFF86FF77FB255B9DF808.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyacophila retracta Martynov 1914	<div><p>Rhyacophila retracta Martynov 1914</p><p>(Figs 8A–8C, 12A–12D, 16A)</p><p>Rhyacophila retracta Martynov 1914, 75–77, male, Russia (Altai); Ross 1956, 76, 103, 121, male; Botosaneanu 1970, 276– 277, 285, 328, female; Lepneva 1964, 250–255, larva; Schmid 1970, 90, 132, pl. 41, male; Tani, 1977, 198, male; Emoto 1979, 559–562, male, female; Tanida 1985, 181, 183, larva; Arefina 1997, 24, 31, male, female; Arefina 2001, 33, 44, female; Hattori 2005, 417, 422, 431, male, female, larva; Nozaki 2016, 311, 313, male; Kang 2020, 218–219, male, larva.</p><p>Rhyacophila uenoi Tsuda 1940, 123–124, male, Hokkaido (Kamikawa). Synonymized by Ross (1956).</p><p>Rhyacophila uchidai Kobayashi 1989, 4–5, male, Korea (Gyeongsangnam-do). Synonymized by Nozaki et al. (2019).</p><p>Diagnosis. Rhyacophila retracta Martynov 1914 is clearly distinguished from other species of the R. retracta Species Group by the following unique genital characters: In males, the short, trilobed apicodorsal lobe of segment IX (Figs 8A, 8B); in females, segment VIII with a deep dorsal cleft and a pair of shallow lateral clefts (Figs 12A, 12B).</p><p>Specimens examined. Hokkaido: 1 female, Horokaishikari-gawa, Kamikawa-cho, 6.vii.1993, M. Sato (SPMN) : 1 female, Yamabe-nishi, Furano-shi, 16.vii.1996, T. Kishimoto (SPMN) ; 1 female, Shikaribetsu, Shikaoicho, 27.viii.1995, unknown collector (SPMN) ; 1 male, Shirogane-no-taki, Ofuyu, Hamamasu-ku, Ishikari-shi, alt. 5 m, 31.vii.2006, T . Ito &amp; T . Hattori (SPMN); 1 male, 4 females, Ponbetsu-gawa, Ikushunbetsu, Mikasa-shi, 22.vi.1997, T . Ito &amp; A. Ohkawa (SPMN); 1 male, 1 female, Takino, Sapporo-shi, 28.vii.1978, T . Ito (SPMN); 1 male, Jozankei, Sapporo-shi, 23.viii.1995, H. Nishimoto (SPMN) ; 2 males, 6 females, Ôtaki, Date-shi, 24.viii.1995, H. Nishimoto (SPMN) ; 32 males, 16 females, Ken-ichi-gawa, Kumaishi-ayukawa-cho, Yakumo-cho, alt. 50 m, 23.vi.1997, T . Ito &amp; A. Ohkawa (SPMN); 3 males, same locality, 22.vii.1997, same collectors (SPMN); 1 male, same locality, 25.viii.1997, same collectors (SPMN); 1 male, Himekawa, Mori-machi, 29.ix.1993, T . Ito (SPMN); 4 males, 9 females, same locality, 24.vi.1997, T. Ito &amp; A. Ohkawa (SPMN); 1 male, Shamisen-daki, Shikabe-cho, 11.ix.2006, T . Ito (SPMN); 1 female, Uzurakawa, Assabu-cho, 30.v.2003, T . Ito &amp; A. Ohkawa (SPMN); 1 female, Kouzurakawa, Assabu-cho, 22.vi.2003, T . Ito &amp; A. Ohkawa (SPMN); 2 females, Ikusagawa, Nanae-cho, 23.vi.2003, T . Ito &amp; A. Ohkawa (SPMN); 1 male, 10 female, same locality, 11.ix.2006, T . Ito (SPMN); 1 female, Idesu-kawa, Siriuchi-cho, 8.vii.1978, T . Ito (SPMN); 1 male, 1 female, Mt. Horoshiri, Toyonuka, Biratori-cho, 25.vii.2017, K. Inazu (NKa). Tsushima Islands: 1 male, Azu-kawa, Izuhara-machi, Tsushima-shi, 3.vi.1984, M. Kobayashi (KPM-NKMK77850) .</p><p>Distribution. Rhyacophila retracta is broadly distributed in Hokkaido, Tsushima Islands, Korea, northeastern parts of China, Sakhalin, Kuril Islands, continental parts of Russian Far East and southern Siberia (Fig.16A: Yang et al. 2005; Ivanov 2011; Taira et al. 2025). Kobayashi (1985) recorded females as this species from Kanagawa, central Honshu, but I identified them (KPM-NKMK76740) deposited in the Kobayashi collection of KPM as an unknown species belonging to the Rhyacophila angulata Species group Ross 1956.</p><p>Japanese name. Ueno-nagare-tobikera.</p></div>	https://treatment.plazi.org/id/C97B87ACFFAEFF86FF77FB255B9DF808	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kawase, Naoki	Kawase, Naoki (2025): Three new species of the Rhyacophila retracta species group (Trichoptera, Rhyacophilidae) from Japan. Zootaxa 5691 (3): 561-578, DOI: 10.11646/zootaxa.5691.3.6, URL: https://doi.org/10.11646/zootaxa.5691.3.6
C97B87ACFFA9FF87FF77FF7C5BD6FA73.text	C97B87ACFFA9FF87FF77FF7C5BD6FA73.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyacophila motakanta Schmid 1970	<div><p>Rhyacophila motakanta Schmid 1970</p><p>(Figs 9A–9C, 13A–13D, 16B)</p><p>Rhyacophila motakanta Schmid 1970, 206, male; Tani 1977, 198, male; Emoto 1979, 560, 564–566, male, female; Hattori 2005, 422, 431, male, female; Inazu &amp; Nishida 2011, 173, male; Tanaka 2012, 20, 22, male; Nozaki 2016, 70, 311, 313, 429, 449, male.</p><p>Diagnosis. The male genitalia of R. motakanta are similar to those of R. orthacantha in having the apically bifid, short apicodorsal lobe, but R. motakanta is easily distinguished from R. orthacantha by bearing the apically truncated anal sclerite without any protuberances apically (Fig. 9C) (whereas R. orthacantha has the anal sclerites with pair of dark pigmented protuberances apically (Fig. 11C)). The female of R. motakanta is easily distinguished from those of other species by the shape of the vaginal apparatus: The apical part of the posterior process narrows abruptly in lateral and ventral views (arrows in figs 13C, 13D).</p><p>Specimens examined. Honshu: Iwate: 3 females, Yakushi-gawa, Taimagura, Etsunagi, Miyako-shi, 10.viii.1994, T. Hattori (SPMN) . Akita: 2 males, 3 females, Asahi-kawa, Nibetsu, Akita-shi, 20.vi.1997, T. Hattori (SPMN) . Yamagata: 4 males, Sagae-shi, 18–22.vii.1994, Murakami (SPMN) ; 3 males, Asahi Mineral Spring, Tateki, Asahi-machi, 2.viii.1983, S. Uchida (KPM-NKMK74290) ; 4 males, 6 females, Higashi-zawa, Hirokawara, Iide-machi, 3.vii.1998, T. Hattori (SPMN) . Tokyo: 2 males, 9 females, Nippara-gawa, Okutama-machi, 5.vi.1991, T. Kagaya &amp; T. Nozaki (TNo) . Kanagawa: 1 male, Karasawa, Nakatsugawa, Kiyokawa-mura, 14.vi.1989, S. Uchida (TNo) ; 1 male, 2 females, Shiraishizawa, Yamakita-machi, 12–13.v.1983, T. Nozaki (TNo) . Nagano: 2 males, 4 females, Narai-gawa, Shiojiri-shi, 12.vi.2012, T. Ito (SPMN) ; 1 female, Kamanashi-gawa, Hakushu-cho, Hokuto-shi, 11.vi.1994, T. Hattori (SPMN) . Gifu: 3 males, Tokuyama-dam, Fujihasi, Ibigawa-cho, 17.vii.1992, F. Hori (SPMN) . Shizuoka: 5 females, Sugehiki-gawa, Nakaizu-cho, Izu-shi, 14.v.1995, T. Hattori (SPMN) . Shiga: 2 males, Takatoki-gawa., Kohara, Yogo-cho, Nagahama-shi, 22.v.2010, H. Morita (NKa) ; 5 males, 47 females, same locaality, 27.v.2008, T. Ushijima (NKa) ; 23 males, 1 female, same locality, 28.v.2008, T. Ushijima (NKa) . Hyogo: 1 male, 1 female, Akioka, Kami-cho, 4.vi.2006, K. Inazu (NKa) ; 1 male, 1 female, Kouro, Chikusa-cho, Shisô-shi, 26.vi.2016, K. Inazu (NKa) . Nara: 1 male, Kawaai, Tenkawa-mura, alt. 600 m, 26.v.1974, T. Hattori (SPMN). Shikoku: Kagawa: 1 female, Kaba-kawa, Shionoe-cho, Takamatsu-shi, 23.v.2000, E. Yamamoto (SPMN) . Ehime: 2 females, Namakusa-dani, Odamiyama, Oda-cho, 29.v.1999, E. Yamamoto (SPMN) . Kochi: 2 males (pinned), Kuroson, Okuyanai, Shimanto-shi, 22.v.2011, T. Hattori (SPMN) ; 11 males, Befu-kyo, Monobe-cho, Kami-shi, 28.v.2004, M. Takai (SPMN) ; 13 males, Shirai-dani, Teragawa, Ino-cho, 1.vii.2005, M. Takai (SPMN) ; 1 male, 2 females, same locality, 20.vi.2006, M. Takai (SPMN) .</p><p>Distribution and biology. Rhyacophila motakanta is widely distributed in Honshu, Shikoku and Kyushu (Fig.16B: Taira et al. 2025). Adults of this species were collected near streams in May to August. Taira (2023) reported that this species had a univoltine life cycle in central Honshu.</p><p>Japanese name. Motakanta-nagare-tobikera.</p></div>	https://treatment.plazi.org/id/C97B87ACFFA9FF87FF77FF7C5BD6FA73	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kawase, Naoki	Kawase, Naoki (2025): Three new species of the Rhyacophila retracta species group (Trichoptera, Rhyacophilidae) from Japan. Zootaxa 5691 (3): 561-578, DOI: 10.11646/zootaxa.5691.3.6, URL: https://doi.org/10.11646/zootaxa.5691.3.6
C97B87ACFFA9FF85FF77FA345895FB83.text	C97B87ACFFA9FF85FF77FA345895FB83.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyacophila lambakanta Schmid 1970	<div><p>Rhyacophila lambakanta Schmid 1970</p><p>(Figs 10A–10C, 14A–14D, 16C)</p><p>Rhyacophila lambakanta Schmid 1970, 209, pl. 41, male, Honshu (Nagano); Tani 1978, 7–8, male; Emoto 1979, 556–560, 562–565, wings, male, female; Hattori 2005, 422, 430, male, female.</p><p>Rhyacophila imagamiensis Kobayashi 1973, 22–23, pl. 3, male. Synonymized by Tani (1978) and Emoto (1979).</p><p>Diagnosis. The male genitalia of R. lambakanta are very distinctive among the R. retracta Species Group in the shape of the apicodorsal lobe of segment IX: Narrow, long, and arcuately curving posteroventrally in apical half (Figs 10A, 10B). The female genitalia of R. lambakanta are similar to those of R. motakanta and R. humerigera sp. nov. in having a medium-sized dorsal cleft and a pair of lateral clefts in segment VIII (Figs 5A–5C, 13A–13B, 14A–14B), but can be distinguished from them by the characters given in the diagnosis of R. humerigera sp. nov.</p><p>Specimens examined. Honshu: Miyagi: 1 male, Natori-gawa, Futakuchi Onsen, Akiu-machi, Sendai-shi, 8.vi.1974, T . Hattori (SPMN) . Yamagata: 1 male, 1 female, Shizu, Nishikawa-machi, 18–19.ix.2004, H. Nishimoto (SPMN) . Tochigi: 1 male, Daiya-gawa, Nikko-shi, 6.x.2002, H. Nishimoto (SPMN) . Tokyo: 6 males, 1 female, Ôkumotori-dani, Nippara-gawa, Okutama-machi, alt. 1300 m, 5.vi.1991, T. Nozaki (Nka) . Kanagawa: 3 males, Shiraishizawa, Yamakita-machi, 12–13.v.1982, T. Nozaki ( Nka). Niigata: 8 males, 5 females, Wasabi-zawa, Ôtokoro, Itoigawa-shi, alt. 1150 m, 14.viii.1998, T. Hattori (SPMN); 1 male, Yahei-gawa, Ôtokoro, Itoigawa-shi, alt. 1600 m, 31.viii.2023, D. Kato (NKa) . Ishikawa: 1 male, Chûgu, Hakusan-shi, 29.ix.2001, H. Nishimoto (SPMN) . Yamanashi: 1 pupa (male), Sakuba-daira-bashi, Ichinose-takahashi, Enzan, Kôshu-shi, alt. 1300 m, T. Hattori (SPMN); 3 males, 2 females, Hirogawara, Ashiyasu-ashikura, Minami-arupusu-shi, 13.viii.1973, T. Hattori (SPMN); 1 male, Kamanashi-gawa, Hakushu-cho, Hokuto-shi, alt. 900 m, 1.vi.1997, T. Hattori (SPMN) . Nagano: 2 males, Kuro-sawa, Yokokawa, Tatsuno-machi, alt. 1050 m, 17.viii.1996, T. Hattori (SPMN); 1 male, Oguro-gawa, Ina-shi, alt. 1400 m, 17.viii.1996, T. Hattori (SPMN); 10 males, 1 female, Shio-kawa, Kashio, Ôshika-mura, alt. 1400 m, T. Hattori (SPMN) . Gifu: 1 male, Namai, Taki-machi, Takayama-shi, 9.vi.2003, H. Nishimoto (SPMN); 5 males, same locality, 24.vi.2004, H. Nishimoto (SPMN); 4 males, 1 female, same locality, 18.vi.2005, H. Nishimoto (SPMN); 1 male (pinned), 2 females (pinned), Iwai-machi, Takayama-shi, 9.vi.2011, T. Hattori (SPMN); 1 female, Otome-keikoku, Kashimo, Nakatsugawa-shi, 8.vi.2003, H. Nishimoto (SPMN); 1 male (pinned), 1 female (pinned), Hora, Miyagawa-cho, Hida-shi, 9.vi.2011, T. Hattori (SPMN); 1 male, Tsurumi, Fujihashi, Ibigawa-cho, 29.v.2002, N. Kawase (NKa) . Shizuoka: 3 males, 5 females, Nishihikage-zawa, Umegashima, alt. 900 m, Shizuoka-shi, 5.vi.1999, T. Hattori (SPMN); 10 males, 2 females, same locality, 28.v.2014, T. Hattori &amp; T. Nozaki (SPMN); 4 males, 6 females, Higashi-zawa-deai, Umegashima, Shizuoka-shi, alt. 900 m, 23.vii.1994, T. Hattori (SPMN); 2 females, Abe-kawa, Nyû-jima, Shizuoka-shi, alt. 375 m, T. Hattori (SPMN) . Shiga: 1 male, Takatoki-gawa, Yogo-cho, Nagahama-shi, 28.v.2008, T. Ushijima (NKa); 1 male, Intani, Kitahira, Ôtsu-shi, 30.iv.2014, S. Takeda (NKa); 1 male, Katsuragawa-sakashita-cho, Ôtsu-shi, 20.v.2013, S. Takeda (NKa) . Kyoto: 1 pupa (male), Kurama, Kyotoshi, 19.v.1974, T. Hattori (SPMN) .</p><p>Distribution and biology. Rhyacophila lambakanta is distributed from northeastern to central Honshu, Japan (Fig. 16C: Taira et al. 2025). Adults of this species were collected near streams in late April to early October. Hattori (2005) indicated that this species was found in rather higher altitude than R. motakanta and R. orthacantha .</p><p>Japanese name. Yumi-nagare-tobikera.</p><p>Remarks. Kobayashi (1984) recorded a male collected from Sado Island (Sadogashima) as R. lambakanta . I examined his specimen (KPM-NKMK72950) and found that the apicodorsal lobe of segment IX of his male is clearly shorter than that of R. lambakanta described above. Further research is needed to confirm the specific identity of this male.</p></div>	https://treatment.plazi.org/id/C97B87ACFFA9FF85FF77FA345895FB83	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kawase, Naoki	Kawase, Naoki (2025): Three new species of the Rhyacophila retracta species group (Trichoptera, Rhyacophilidae) from Japan. Zootaxa 5691 (3): 561-578, DOI: 10.11646/zootaxa.5691.3.6, URL: https://doi.org/10.11646/zootaxa.5691.3.6
C97B87ACFFABFF85FF77FB255BE7F82C.text	C97B87ACFFABFF85FF77FB255BE7F82C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyacophila orthacantha Emoto 1979	<div><p>Rhyacophila orthacantha Emoto 1979</p><p>(Figs 11A–11C, 15A–15D, 16D)</p><p>Rhyacophila orthacantha Emoto 1979, 557–560, 565–568, male, female, Honshu (Aichi); Hattori 2005, 422, 431, male, female.</p><p>Diagnosis. The male genitalia of R. orthacantha are similar to those of R. motakanta in having a short, apically bifid apicodorsal lobe of segment IX, but can be distinguished from the latter by the characters given in the diagnosis of R. motakanta . The female genitalia of this species are indistinguishable from those of R. koryne sp. nov. as indicated in the diagnosis of R. koryne sp. nov.</p><p>Specimens examined. Honshu: Tokyo: 1 male, Ôkumotori-dani, Nippara-gawa, Okutama-machi, alt.1300 m, 5.vi.1991, T. Nozaki (TNo) . Shizuoka: 2 females, Nigori-gawa, Umegashima, Shizuoka-shi, 9.vi.2002, T. Hattori (SPMN) ; 2 males, 1 female, Warashina-gawa, Yunoshima, Shizuoka-shi, alt. 300 m, 7.v.1995, T. Hattori (SPMN) ; 6 males, 2 females, same locality, 11.v.2001, T. Hattori (SPMN) ; 1 female, Ituka-zawa, Tashiro, Ikawa, Shizuoka-shi, alt.750 m, T. Hattori (SPMN) . Mie: 2 males, 2 females, Kawachi-dani, Yamaguchi, Fujiwara-cho, Inabe-shi, alt. 250 m, 4.vi.2006, T. Hattori (SPMN) . Shiga: 1 male, Takatoki-gawa, Yogo-cho, Nagahama-shi, 28.v.2008, T. Ushijima (NKa) . Nara: 7 males, Kawaai, Tenkawa-mura, 26.v.1974, T. Hattori (SPMN) ; 1 male, Dorokawa, Tenkawa-mura, 15.v.2004, H. Nishimoto (SPMN) .</p><p>Distribution and biology. R. orthacantha is distributed in central Honshu (Fig. 16D: Taira et al. 2025). Adults were collected from mountain streams by sweeping or light traps in May to June. Taira (2023) reported that this species had a univoltine life cycle in central Honshu.</p><p>Japanese name. Togari-mijikao-nagare-tobikera.</p></div>	https://treatment.plazi.org/id/C97B87ACFFABFF85FF77FB255BE7F82C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kawase, Naoki	Kawase, Naoki (2025): Three new species of the Rhyacophila retracta species group (Trichoptera, Rhyacophilidae) from Japan. Zootaxa 5691 (3): 561-578, DOI: 10.11646/zootaxa.5691.3.6, URL: https://doi.org/10.11646/zootaxa.5691.3.6
