identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
C97987E73B1EFF93FF607F3EED918CBD.text	C97987E73B1EFF93FF607F3EED918CBD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Luciobarbus Heckel 1843	<div><p>Key to species of Luciobarbus in the African Mediterranean Sea basin</p><p>The key is developed based on materials examined listed at the end of the text.</p><p>1a Barbels thick, rostral-barbel origin close to tip of snout............................................. L. guercifensis</p><p>1b Barbels thin, rostral-barbel origin clearly behind tip of snout.................................................... 2</p><p>2a Last unbranched dorsal-fin ray serrated along 2/3 of its length.................................................. 3</p><p>2b Last unbranched dorsal-fin ray serrated along almost its entire length............................................. 8</p><p>3a Caudal peduncle 1.2–1.3 times longer than deep............................................................. 4</p><p>3b Caudal peduncle 1.3–1.5 times longer than deep............................................................. 6</p><p>4a Predorsal length 46–52% SL; interorbital distance 40–44% HL; 43–47+1–3 lateral line scales...... Luciobarbus setivimensis</p><p>4b Predorsal length 52–59% SL; interorbital distance 36–46% HL; 41–45+1–3 lateral line scales......................... 5</p><p>5a Pectoral-fin length 17–21% SL, 60–79% HL; usually 41+1–2 lateral line scales................ Luciobarbus mascarensis</p><p>5b Pectoral-fin length 21–24% SL, 79–90% HL; 43–45+1–3 lateral line scales................... Luciobarbus lanigarensis</p><p>6a Caudal peduncle depth 13–14% SL; 43–47+1–2 lateral line scales............................ Luciobarbus numidiensis</p><p>6b Caudal peduncle depth 10–13% SL; 41–45+1–2 lateral line scales............................................... 7</p><p>7a Predorsal length 46–52% SL; Interorbital distance 36–39% HL; 41–43+1–2 lateral line scales; 4 unbranched dorsal-fin rays................................................................................... Luciobarbus leptopogon</p><p>7b Predorsal length 52–56% SL; Interorbital distance 33–40% HL; 42–45+1–2 lateral line scales; usually 5, very rarely 4 unbranched dorsal-fin rays............................................................ Luciobarbus yahyaouii</p><p>8a Anal-fin length 19–23% SL.......................................................... Luciobarbus numidiensis</p><p>8b Anal-fin length 16–19% SL............................................................................. 9</p><p>9a Anal-fin base length 8–10% SL; 41–43+1–2 lateral line scales................................ Luciobarbus chelifensis</p><p>9b Anal-fin base length 6–8% SL; 43–47+1–3 lateral line scales................ Luciobarbus callensis / Luciobarbus rifensis</p></div>	https://treatment.plazi.org/id/C97987E73B1EFF93FF607F3EED918CBD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Brahimi, Amina;Libois, Roland;Henrard, Arnaud;Freyhof, Jörg	Brahimi, Amina, Libois, Roland, Henrard, Arnaud, Freyhof, Jörg (2018): Luciobarbus lanigarensis and L. numidiensis, two new species of barbels from the Mediterranean Sea basin in North Africa (Teostei: Cyprinidae). Zootaxa 4433 (3): 542-560, DOI: 10.11646/zootaxa.4433.3.9
C97987E73B1FFF9EFF607A93EA338A1A.text	C97987E73B1FFF9EFF607A93EA338A1A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Luciobarbus lanigarensis Brahimi & Libois & Henrard & Freyhof 2018	<div><p>Luciobarbus lanigarensis, new species</p><p>(Figs. 3–5)</p><p>Holotype. RMCA 2017-022 -P-0024, 134 mm SL; Morocco: Guenfouda prov.: Oued Isly, 34.5118°N, 2.0 5826°W.</p><p>Paratypes. RMCA 2017-022 -P-0019-0042, 21, 71– 150 mm SL; ZFMK ICH-105602-105603, 2, 115– 123 mm SL; same data as holotype .</p><p>Material for molecular analyses. RMCA DNA-2017-022 -P-LO1-LO6; same data as holotype (GenBank accession numbers: Cyt b: MH187170, MH187171, MH187172, MH187173, MH187174; D-loop: MH187193, MH187194, MH187195, MH187196, MH187197).</p><p>Diagnosis. Luciobarbus lanigarensis is distinguished from other Luciobarbus species from the African Mediterranean Sea basin by a combination of characters, none of them unique. Luciobarbus lanigarensis is distributed adjacent to L. mascarensis in the east and L. yahyaouii and L. guercifensis in the west.</p><p>It is distinguished from L. guercifensis by having thin barbels (vs. thick), the rostral-barbel origin placed clearly behind the tip of the snout (vs. rostral-barbel origin at the extreme anterior of the snout) and from L. mascarensis, L. yahyaouii and L. leptopogon, by having a wider interorbital distance (38–47% HL vs. 36–40 in L. mascarensis, 33–40 in L. yahyaouii, 36–39 in L. leptopogon) and a longer pectoral-fin (21–24% SL vs. 16–21 in L. mascarensis, 18–21 in L. yahyaouii). It is further distinguished from L. chelifensis, L. leptopogon and L. mascarensis by having 43–45+1–3 lateral line scales (vs. 41–43+ 1–2 in L. chelifensis and L. leptopogon, usually 41+ 1–2 in L. mascarensis) (Table 8).</p><p>The new species is distinguished from L. callensis, L. chelifensis and L. rifensis by having the last unbranched dorsal-fin ray serrated along 2/3 of its length (vs. last unbranched dorsal-fin ray serrated along almost its entire length) and a longer anal-fin (19–23% SL vs. 17–19 in L. callensis and L. chelifensis, 16–19 in L. rifensis). The new species is distinguished from L. leptopogon and L. setivimensis by having a greater pre-dorsal distance (53– 59% SL vs. 49–52 in L. leptopogon, 47–52 in L. setivimensis).</p><p>Description. See Figures 3–5 for general appearance. Morphometric data are given in Table 2 and meristic data in Tables 7 and 8. Middle sized and stout species, with a moderately long head. Body deepest at dorsal-fin origin. Depth decreasing continuously towards caudal-fin base. Greatest body width between pectoral and dorsal- fin origins. Caudal peduncle compressed, 1.3–1.5 times longer than deep. Section of head roundish, flattened on ventral surface. Snout rounded. Mouth inferior. Dorsal-fin origin situated above pelvic-fin origin. Anal-fin origin at vertical of 50–60% of distance between dorsal and caudal-fin origins. Anal fin reaching to caudal-fin base. Pectoral fin reaching 70–80% of distance between pectoral-fin and pelvic-fin origins. Pelvic fin not reaching vertical of tip of last dorsal-fin ray when folded down. Pelvic fin reaching to about 70–90% of distance between pelvic and analfin origins. Posterior dorsal-fin margin slightly concave. Posterior pectoral- and anal-fin margins convex. Caudal fin forked with rounded lobes of equal length. Largest known specimen 150 mm SL, but expected to grow larger.</p><p>Dorsal fin with 4 unbranched and 8½ branched rays, last unbranched ray ossified and serrated at 2/3 of its length. Anal fin with 3 unbranched and 5½ branched rays. Pectoral-fin with 14–17 (mode 16) and pelvic fin with 8 rays. Lateral line with 43–45 scales on flank and 1–3 scales on caudal-fin base. Between dorsal-fin origin and lateral line 8½–9½ (mode 8½) scale rows, and 5½–6½ (mode 5½) scale rows between pelvic-fin origin and lateral line. Pharyngeal teeth in three rows: 4+3+2. On first gill arch, 3-4 gill rakers on the on the upper and 8 on lower limb.</p><p>Coloration. Brown yellowish-golden in life and preserved individuals without colour pattern. Head golden, cheeks and ventral side of head and body whitish. In life, fins orange.</p><p>Etymology. Luciobarbus lanigarensis is named for Lanigar, the old name of Ouajda province, given by Claudius Ptolemaeus (100-170). An adjective.</p><p>Distribution. Luciobarbus lanigarensis was found in the Isly River in northestern Morocco, which is a headwater stream of the Tafna River. The Tafna River drainage is situated in the border area between Algeria and Morocco, having its estuary in Algeria close to the city of Tlemcen. Therefore, we expect L. lanigarensis to be found in Algeria also. See Figure 2 ( L. lanigarensis are Nr. 28–29) for the distribution of Luciobarbus species from the African Mediterranean Sea basin.</p></div>	https://treatment.plazi.org/id/C97987E73B1FFF9EFF607A93EA338A1A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Brahimi, Amina;Libois, Roland;Henrard, Arnaud;Freyhof, Jörg	Brahimi, Amina, Libois, Roland, Henrard, Arnaud, Freyhof, Jörg (2018): Luciobarbus lanigarensis and L. numidiensis, two new species of barbels from the Mediterranean Sea basin in North Africa (Teostei: Cyprinidae). Zootaxa 4433 (3): 542-560, DOI: 10.11646/zootaxa.4433.3.9
C97987E73B13FF9AFF607C68EBC48EA8.text	C97987E73B13FF9AFF607C68EBC48EA8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Luciobarbus numidiensis Brahimi & Libois & Henrard & Freyhof 2018	<div><p>Luciobarbus numidiensis, new species</p><p>(Figs. 6–8)</p><p>Holotype. RMCA 2017-022 -P-0004, 140 mm SL; Algeria: Mila prov.: Oued El-Kébir at <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=6.2783&amp;materialsCitation.latitude=36.6053" title="Search Plazi for locations around (long 6.2783/lat 36.6053)">Hammam Beni Haroun</a>, 36.6053°N, 6.2783°E.</p><p>Paratypes. RMCA 2017-022 -P-0001-0007, 6, 99– 162 mm SL; same data as holotype .— RMCA 2017-022 -P- 0008-0011, 4, 45– 78 mm SL; Algeria: Jijel prov.: Oued Saleh at <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=5.8894&amp;materialsCitation.latitude=36.4915" title="Search Plazi for locations around (long 5.8894/lat 36.4915)">Borj Bou Akkaz</a>, 36.4915°N, 5.8894°E. — RMCA 2017-022 -P-0012-0018, 5, 73– 150 mm SL; ZFMK ICH-105604-105605, 2, 100– 109 mm SL; Algeria: Mila prov.: Oued El-Kébir at <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=6.2783&amp;materialsCitation.latitude=36.6053" title="Search Plazi for locations around (long 6.2783/lat 36.6053)">Hammam Beni Haroun</a>, 36.6053°N, 6.2783°E.</p><p>Material for molecular analyses. RMCA DNA-2017-022 -P-NH1-NH6, same data as holotype (GenBank accession numbers: Cyt b: MH187177, MH187179, MH187182, MH187183, MH187187; D-loop: MH187200, MH187203, MH187205, MH187206, MH187210) .— RMCA DNA-2017-022 -P-NH16-NH27, Algeria: Mila prov.: Beni-Haroun reservoir at Oued El-Kébir, 36.3319, 6.1611 ( GenBank accession numbers: Cyt b: MH187175, MH187176, MH187178, MH187180, MH187181, MH187184, MH187185, MH187186, MH187188; D-loop: MH187198, MH187199, MH187201, MH187202, MH187204, MH187207, MH187208, MH187209, MH187211) .</p><p>Diagnosis. Luciobarbus numidiensis is distinguished from other Luciobarbus species from the African Mediterranean Sea basin, by a combination of characters, none of them unique. Luciobarbus numidiensis is distributed adjacent to L. callensis in the east and L. setivimensis in the west.</p><p>It is distinguished from L. callensis by having a longer anal-fin (19–23% SL vs. 16–19%), a wider anal-fin base (6.5–10% SL vs. 6.5–8) and a longer distance between the pectoral and pelvic-fin origins (27–30% SL vs. 22– 29). The new species is distinguished from L. setivimensis by the having a longer pectoral-fin (74–94% HL vs. 69– 84), a longer pre-dorsal length (49–55% SL vs. 46–52), a wider caudal peduncle depth (12–14% SL vs. 11–13) and a smaller eye diameter (11–22% HL vs. 11–28%).</p><p>The new species is distinguished from L. guercifensis by having thin barbels (vs. thick), the rostral-barbel origin placed clearly behind the tip of the snout (vs. rostral-barbel origin at the extreme anterior of the snout). It is distinguished from L. chelifensis, L. mascarensis and L. leptopogon by having 43–47+1–2 lateral line scales (vs. 41–43+ 1–2 in L. chelifensis and L. leptopogon, usually 41+ 1–2 in L. mascarensis) (Table 8).</p><p>Luciobarbus numidiesis is further distinguished from L. chelifensis and L. rifensis hy having a longer anal fin (19–22% SL vs. 17–19 in L. chelifensis, 16–19 in L. rifensis). The new species is distinguished from L. leptopogon by having a greater caudal peduncle depth (12–14% SL vs. 10–13) and from L. yahyaouii by having a longer distance between the pelvic and anal-fin origins (23–26% SL vs. 20–24).</p><p>It is also distinguished from L. mascarensis, L. lanigarensis and L. yahyaouii by having a shorter pre-dorsal length (48–55% SL vs. 53–58 in L. mascarensis, 54–59 in L. lanigarensis, 53–57 in L. yahyaouii). The new species is also distinguished from L. lanigarensis by having a smaller pre-pelvic length (49–56% SL vs. 55–60%), a greater post-orbital length (46–52% HL vs. 42–48) and the last unbranched dorsal-fin ray ossified and serrated at 2/3 of its length (vs. last unbranched dorsal-fin ray serrated along almost its entire length).</p><p>Description. See Figures 6–8 for general appearance. Morphometric data are given in Table 3 and meristic data in Table 7 and 8. A large-sized species with a moderately long head. Body deepest at dorsal-fin origin or about midline between nape and dorsal-fin origin. Depth decreasing continuously towards caudal-fin base. Greatest body width in front of dorsal-fin origin. Caudal peduncle compressed, 1.2–1.5 times longer than deep. Section of head roundish, flattened on ventral surface. Snout rounded. Mouth inferior. Dorsal-fin origin situated above pelvic-fin origin. Anal-fin origin slightly behind vertical of middle between dorsal and caudal-fin origins. Anal-fin origin slightly behind vertical of middle of distance between dorsal and caudal-fin origins. Anal fin reaching to caudal-fin base. Pectoral fin reaching 60–80% of distance between pectoral- and pelvic-fin origin. Pelvic fin not reaching vertical of tip of last dorsal-fin ray when folded down. Pelvic fin reaching to 60–90% of distance between pelvic and anal-fin origins. Posterior dorsal-fin margin straight or slightly concave. Posterior pectoral and anal-fin margins convex. Caudal fin forked with rounded lobes of equal length. Largest known specimen 430 mm SL.</p><p>Dorsal fin with 4 or 5 unbranched and 8½ branched rays, last unbranched ray serrated along almost its entire length. Anal fin with 3 unbranched and 5½ branched rays. Pectoral-fin with 13–18 rays (mode 17) and pelvic fin with 8 rays. Lateral line with 43–47 scales on flank and 1–2 scales on caudal-fin base. Between dorsal-fin origin and lateral line 8½–9½ (mode 9½) scale rows and 5½–6½ (mode 5½) scale rows between pelvic-fin origin and lateral line. Pharyngeal teeth in three rows: 4+3+2. On first gill arch, 3-4 gill rakers on upperand 8-10 on lower limb.</p><p>Coloration. In life, whitish-golden on belly, silvery-grey on flank and dark-grey on back. Head golden-grey and cheeks greenish-golden. All fins grey. Pectoral fin with a golden distal edge.</p><p>Etymology. Luciobarbus numidiensis is named for Numidia, an ancient Berber kingdom, located in Algeria. The capital of Numidia was Cirta, the present city of Constantine. The El-Kebir River, the habitat of L. numidiensis, flows through Constantine. An adjective.</p><p>Distribution. Luciobarbus numidiensis was found in the El-Kébir River and in the Beni Haroun reservoir, both belonging to the El-Kébir River drainage in northestern Algeria. See Figure 2 ( L. numidiensis are Nr. 10–12) for the distribution of Luciobarbus species in the African Mediterranean Sea basin.</p><p>Remarks. Luciobarbus numidiensis occurs north of the range of L. biscarensis, which is found in the rivers of the Algerian Sahara. It is distinguished from this species by having 43–47+1–2 lateral line scales (vs. 49–51+1–2), 5½–6½ scale rows between the lateral line and the pelvic-fin origin (vs. 7½–8½), 4 at 5 unbranched dorsal-fin rays (vs. 5), the dorsal-fin origin situated above the pelvic-fin origin (vs. behind), a longer anal fin (19–22% SL vs. 17– 19) and a more wider interorbital distance (37–43% HL vs. 31–39).</p><p>Luciobarbus numidiensis is quite variable in the serration of the last unbranched dorsal-fin ray. Specimens collected from the Beni-Haroun reservoir have the last unbranched dorsal-fin ray serrated along 2/3 of its length with sparsely set denticulations while fish collected from the El-Kébir River have the last unbranched dorsal-fin ray serrated along almost its entire length with densely set denticulations.</p></div>	https://treatment.plazi.org/id/C97987E73B13FF9AFF607C68EBC48EA8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Brahimi, Amina;Libois, Roland;Henrard, Arnaud;Freyhof, Jörg	Brahimi, Amina, Libois, Roland, Henrard, Arnaud, Freyhof, Jörg (2018): Luciobarbus lanigarensis and L. numidiensis, two new species of barbels from the Mediterranean Sea basin in North Africa (Teostei: Cyprinidae). Zootaxa 4433 (3): 542-560, DOI: 10.11646/zootaxa.4433.3.9
