taxonID	type	description	language	source
C84487BFC739782B4F2AFC09FDCE2185.taxon	type_taxon	Type species Nassa paupera Gould, 1850 by original designation (Iredale 1936: 322).	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC739782B4F2AFC09FDCE2185.taxon	diagnosis	Diagnosis Morphological features of the protoconch are of great importance when recognizing species of Reticunassa. Several species have a multispiral protoconch, indicating a planktotrophic / planktonic larval development, resulting in a larger range of distribution. Others have a paucispiral protoconch, indicating a short, non-planktotrophic / direct larval development. All species of Reticunassa possess an unkeeled protoconch (Fig. 2 A – B), as do other Nassariinae genera in the Atlantic Ocean (Galindo et al. 2016). However, Reticunassa lives in sympatry with the Indo-Pacific genus Nassarius s. str., characterized by a keeled protoconch. Protoconchs of Reticunassa can be rather broad and flattened; a few are pointed. They might possess microscopic sculpture, such as spiral ridges, axial striae and minute pustules (Fig. 2 A – B), but frequently a much higher resolution is needed to see details of this sculpture. Due to the obvious importance of protoconch characters, the identification of specimens in this genus lacking a protoconch becomes more difficult. Species of Reticunassa have strongly or weakly developed continuous spiral cords, from 10 to 15 on the body whorl (Fig. 2 D – F), crossing over the axial ribs. The space between these spiral cords, the intercordal surface, can be smooth, axially striated or can display a number of (occasionally extremely) fine spiral striae (Fig. 2 F). This makes the sculpture of the intercordal surface important for identification purposes within Reticunassa. Fig. 1 A – B shows that differences on these intercordal surfaces are useful synapomorphies to separate SC 1 from SC 2. Species within Reticunassa have an axially-ribbed teleoconch (Fig. 2 C). These axial ribs may become weaker or absent on the body whorl towards the base and / or the outer lip. Ribs can be round or flat. The number of ribs differs among species. Most species develop a strong varix near the outer lip. The suture is usually deeply impressed. The aperture is oval to sub-circular, the inside of the outer lip with a number of denticles or lirae (Fig. 2 G – H). A columellar callus is present and varies in shape and sculpture. It can be limited to the columella or partly extend over the body whorl. In some species, the color is variable; generally, it is cream-yellowish to orange, or white to dark brown. Banding with darker colors may also be present. Sometimes only some of the spiral cords are darker than the remaining part of the shell. Reticunassa species are small, usually varying between 5 and 12 mm, but one is up to 16.5 mm (R. goliath sp. nov.). Most species live in littoral to sublittoral ecosystems, only empty shells having been found in waters deeper than 260 m. Their occurrence is in the Indo-Pacific Ocean, from the Miocene to the present.	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC739782B4F2AFC09FDCE2185.taxon	discussion	Remarks The genus name Reticunassa was introduced to group together “ species around pauperus Gould ” by Iredale (1936: 322). Cernohorsky (1984: 175) regarded Reticunassa as a synonym of the subgenus “ Hima Leach in Gray, 1852 ”, within the genus Nassarius Duméril, 1805. However, Hima is not an available name, but an incorrect subsequent spelling of Hinia Leach in Gray, 1847 (Gray 1847 a: 269), since “ Hima ” included the same three species as in the original description of Hinia (during a period in which original descriptions only consisted of introducing new names). Nonetheless, “ Hima ” and not Hinia was listed by Adams & Adams (1853). Cossmann (1901) considered both Hinia and “ Hima ” as valid names, with “ Hima ” treated sensu Adams & Adams (1853), not Leach in Gray, 1852, and with “ type species ” Nassa incrassata “ Müller ”. Woodring (1928) noted the misspelling and this was reiterated by Lee (2013). Despite this, the error in the use of “ Hima ” has persisted. On the other hand, Hinia Leach in Gray, 1847 is a junior objective synonym of Tritia Risso, 1826 as they have the same type species, Buccinum reticulatum Linnaeus, 1758, designated for both genera by Gray (1847 b: 139). Molecular results of Galindo et al. (2016) indicated that species considered part of the Nassarius pauperus complex, including “ Nassarius pauper ”, form a clade distant from Buccinum reticulatum (accepted as Tritia reticulata), the type species of Tritia (= Hinia), also shown in Fig. 1 A. The only available genus group name for this pauperus clade is Reticunassa Iredale, 1936, which is herein reinstated with full generic status. Other species might be confused as members of the genus Reticunassa because of their morphological similarities. As it is impossible to treat all the small and ridged species, we focus on the following characters: (1) Reticunassa is an Indo-Pacific group of molluscs, not occurring in Atlantic waters (Galindo et al. 2016); (2) species of Reticunassa lack keeled protoconchs (all species with this keel belong to the genus Nassarius); (3) fine spiral striae or axial striae on the intercordal surface (Fig. 2 D – F) may be an indication of at least kinship with Reticunassa; (4) all species of Reticunassa have a distinct, elongate and slender shape (Figs 3, 8). These criteria exclude from Reticunassa any Mediterranean species, such as Tritia incrassata (Strøm, 1768) (previously Nassarius incrassatus), West African species, such as Tritia senegalensis (Von Maltzan, 1884) (previously Nassarius senegalensis) and Caribbean species, such as Phrontis candidissima (C. B. Adams, 1845) (previously Nassarius candidissimus) (Galindo et al. 2016). Most Indo-Pacific species, such as Nassarius vanuatuensis Kool & Galindo, 2014, can be distinguished from Reticunassa by the protoconch features. Additionally, Kool & Galindo (2014) demonstrated a 19 % genetic distance between Nassarius vanuatuensis and Reticunassa paupera. Naytia priscardi (Bozzetti, 2006) from Madagascar is a member of the sister group of Reticunassa and exhibits great morphological similarity to some species of this genus. Molecular synapomorphies indicate that N. priscardi belongs to Naytia.	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC73D78334F38FE96FEF52589.taxon	description	Figs 2 A, C, F – L, 3 A – C, 4; Tables 1 – 2	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC73D78334F38FE96FEF52589.taxon	materials_examined	Type material Lectotype (ICZN Art. 74.5; Johnson 1964: 124) PACIFIC OCEAN: length 9.1 mm, width 4.8 mm (USNM 5726, Fig. 2 I – J, http: // n 2 t. net / ark: / 65665 / 3 f 1 f 5 ebf 5 - d 0 d 6 - 4 c 9 a-bddd- 3677 d 3 efb 7 e 2). Paralectotype (ICZN Art. 74.5; Johnson 1964: 124) PACIFIC OCEAN: 1 spm, type locality, length 7.2 mm, width 3.5 mm (MCZ 216602, ex USNM, Fig. 2 K – L). Neotype Both the lectotype and the paralectotype have been affected by Byne’s disease (Kool & Dekker 2006). However, remnants of microscopic spiral sculpture on the intercordal surface of both specimens are vaguely visible. Gould (1850) did not mention this feature in his original description. The protoconch of the lectotype is badly damaged and that of the paralectotype is completely absent. The protoconch, figured by Cernohorsky (1984: 5, fig. 12) cannot be the protoconch of either of the types, but most likely belongs to Reticunassa tringa, which he considered a synonym of Nassarius pauperus. The poor condition of both existing types calls for the designation of a neotype for this species. Among the specimens in the MNHN collection, there is one sequenced individual that is very similar to the original types of the species. A proposal to designate a neotype is in the process of being submitted to the International Commission on Zoological Nomenclature (ICZN). If accepted, the neotype will be MNHN IM- 2007 - 31782, length 6.9 mm, width 3.6 mm (Fig. 3 A – C; http: // coldb. mnhn. fr / catalognumber / mnhn / im / 2007 - 31782). Type locality Pacific Ocean. If the designation of a neotype is accepted by the ICZN, the locality of MNHN IM- 2007 - 31782 becomes the new type locality: SANTO 2006 Expedition, stn DB 12, Vanuatu, South of Aore Island, 15 ° 13.6 ′ S, 167 ° 10.1 ′ E, 10 – 18 m. Other material examined INDONESIA: Bali, Benoa Port, in sand dumps, leg. T. de Boer (HK 995.06, 1 spm); Bali, Nusa Dua, Benoa Beach, leg. J. Post (HK 995.09, 1 spm); Flores Sea, Pulau Sabalana, Lamuruang, 10 – 20 m (HD 17295, 1 spm); Arafura Sea, near Pulau Kuringarankeke, W of Makassar, southern Sulawesi, 5 ° 5 ′ 34 ″ S, 119 ° 17 ′ 09 ″ E, leg. H. Morrison (GH, 1 spm); West-Papua, Manokwari, leg. D. Smits (HK 995.03, 5 spm; HD 38513, 6 spm); SNELLIUS 1929, Kera, near Timor (RMNH, 1 spm); Java, Bay of Jakarta, Pulau Dapur, leg. J. D. F. Hardenberg, 1938 (ZMA. Moll. 096216, 1 spm); Java, Bay of Jakarta, Pulau Ubi, leg. J. D. F. Hardenberg, 1938 (ZMA. Moll. 096218, 4 spm); Java, Bay of Jakarta, Pulau Kelor, leg. J. D. F. Hardenberg, 1938 (ZMA. Moll. 096219, 5 spm); Java, Bay of Jakarta, Pulau Untung Djava, leg. J. D. F. Hardenberg, 1938 (ZMA. Moll. 096220, 3 spm); Java, Bay of Jakarta, Pulau Dapur, leg. J. D. F. Hardenberg, 1938 (ZMA. Moll. 096221, 9 spm); Java, Bay of Jakarta, Pulau Damar, leg. J. D. F. Hardenberg, 1938 (ZMA. Moll. 096222, 2 spm); Java, Bay of Jakarta, Pulau Sakit, leg. J. D. F. Hardenberg, 1938 (ZMA. Moll. 096223, 3 spm); Java, Bay of Jakarta, Pulau Njamoek, leg. J. D. F. Hardenberg, 1938 (ZMA. Moll. 096224, 10 spm); Java, Bay of Jakarta, Pulau Noesi, leg. J. D. F. Hardenberg, 1938 (ZMA. Moll. 096225, 15 spm); Java, Bay of Jakarta, Pulau Ubi, leg. J. D. F. Hardenberg, 1938 (ZMA. Moll. 096226, 9 spm); Java, Bay of Jakarta, Pulau Ramboet, leg. J. D. F. Hardenberg, 1938 (ZMA. Moll. 096227, 1 spm); KEPULAUAN SERIBU 2005, Java, Hantu Besar, 5 ° 31.8 ′ S, 106 ° 32.3 ′ E, 5 – 26 m (ZMA. Moll. 175389, 1 spm); KEPULAUAN SERIBU 2005, stn SER 05 / 33, Java, Pari East, 5 ° 51.3 ′ S, 106 ° 38.3 ′ E, 5 – 25 m (ZMA, 1 spm); KEPULAUAN SERIBU 2005, stn SER 05 / 43, Java, Payung Besar, 5 ° 40.3 ′ S, 106 ° 33.8 ′ E, 5 – 25 m (ZMA. Moll. 154531, 2 spm). PHILIPPINES: all PANGLAO 2004: stn M 10, Panglao I., Bingag / Tabalong, 9 ° 37.8 ′ N, 123 ° 48.4 ′ E, 0 – 3 m (1 spm); stn M 26, Bohol I., Taloto, 9 ° 40.9 ′ N, 123 ° 51.1 ′ E, 0 – 2 m (1 spm); stn S 25, Bohol I., Ubajan, 9 ° 41.5 ′ N, 123 ° 51.0 ′ E, 21 m (1 spm); stn B 14, Panglao I., Sungcolan Bay, 9 ° 38.5 ′ N, 123 ° 49.2 ′ E, 2 – 4 m (3 spm). AUSTRALIA: Queensland, Fitzroy I. (HK 995.08, 2 spm; HD 17298, 1 spm; PS, 1 spm); Queensland, Innisfail, leg. Pini (ZMA. Moll. 096207, 1 spm). GUAM: Neye I. Channel, leg. S. P. Kool (HK 995.04, 2 spm); Apra Harbor, by diving to 10 m, leg. H. Morrison (HK 995.05, 1 spm); Glass Breakwall, snorkeling, 1 – 2 m (HD 12297, 2 spm); Glass Breakwall, 6 m, in rubble (GH, 1 spm). CAROLINE ISLANDS: Truk I., Moen I. (HK 995.10, 2 spm). NEW CALEDONIA: CHALCAL 1: stn D 17, Plateau Chesterfield-Bellona, 19 ° 11 ′ S, 158 ° 55.8 ′ E, 44 m (1 spm); stn D 57, Plateau Chesterfield-Bellona, 21 ° 29.5 ′ S, 159 ° 16.4 ′ E, 62 m (1 spm). – LAGON: Ile Quen, Baie Prony, stn 64, 22 ° 28 ′ S, 166 ° 25 ′ E, 15 m (2 spm; NBC, 1 spm; HK 995.01, 1 spm); Ile Quen, Baie Prony, stn 78, 22 ° 28 ′ S, 166 ° 31 ′ E, 35 m (1 spm); Ile Quen, Baie Prony, stn 240, 22 ° 23 ′ S, 166 ° 32 ′ E, 42 m (1 spm); Nouméa, stn 290, 22 ° 16 ′ S, 166 ° 32 ′ E, 11 m (1 spm); Yaté, stn 620, 22 ° 02 ′ S, 166 ° 56 ′ E, 50 – 52 m (1 spm); Yaté, stn 625, 21 ° 59 ′ S, 166 ° 54 ′ E, 34 – 40 m (1 spm); Poindimié, stn 789, 21 ° 03 ′ S, 164 ° 34 ′ E, 29 m (1 spm); Poum, stn 1039, Balabio Section, 20 ° 01 ′ S, 164 ° 02 ′ E, 20 – 21 m (1 spm; HK 995.02, 1 spm); Belep, stn 1105, 19 ° 40 ′ S, 163 ° 57 ′ E, 25 m (1 spm); Belep, stn 1129, 19 ° 29 ′ S, 163 ° 49 ′ E, 40 m (1 spm); Belep, stn 1168, 19 ° 16 ′ S, 163 ° 09 ′ E, 50 m (2 spm); Nouméa, Pointe Magnin, stn 1355, 22 ° 18.9 ′ S, 166 ° 26.6 ′ E, 7 – 10 m (3 spm). – MONTROUZIER 1993: Touho, Lagon du grand Récif Mengalia, stn 1253, 20 ° 46.6 ′ S, 165 ° 14.1 ′ E, 2 – 3 m (1 spm); Touho, Lagon du grand Récif Mengalia, stn 1266, 20 ° 39.6 ′ S, 165 ° 14.7 ′ E, 10 – 15 m (1 spm); Touho, Lagon du grand Récif Mengalia, stn 1269, 20 ° 35.1 ′ S, 165 ° 08.1 ′ E, 15 – 20 m (2 spm); Koumac, stn 1298, 20 ° 35.2 ′ S, 164 ° 16.6 ′ E, 2 – 4 m (1 spm); Koumac, stn 1299, 20 ° 34 ′ S, 164 ° 13.0 ′ E, 12 – 14 m (11 spm; NBC, 1 spm; HD 24147, 1 spm); Koumac, stn 1308, 24 ° 40 ′ S, 164 ° 15.2 ′ E, 15 – 20 m (3 spm; ZMA, 1 spm); Koumac, stn 1312, 20 ° 40 ′ S, 164 ° 14.9 ′ E, 26 – 40 m (6 spm; HK 995.11, 1 spm); Koumac, stn 1314, 20 ° 39.8 ′ S, 164 ° 15.3 ′ E, 30 – 63 m (7 spm); Koumac, stn 1315, 20 ° 40.7 ′ S, 164 ° 14.7 ′ E, 66 – 87 m (3 spm); Koumac, stn 1318, 20 ° 41.4 ′ S, 164 ° 14.8 ′ E, 20 – 30 m (2 spm); Koumac, stn 1319, 20 ° 44.7 ′ S, 164 ° 15.5 ′ E, 15 – 20 m (2 spm); Koumac, stn 1322, 20 ° 45.2 ′ S, 164 ° 15.2 ′ E, 53 – 71 m (1 spm); Koumac, stn 1323, 20 ° 40.9 ′ S, 164 ° 14.8 ′ E, 82 – 120 m (1 spm). – CORAIL 2: stn DW 32, Plateau de Chesterfield, 19 ° 25 ′ S, 158 ° 49 ′ E, 55 m (1 spm); stn DW 34, Plateau de Chesterfield, 19 ° 18 ′ S, 158 ° 47 ′ E, 47 m (2 spm); stn DW 38, Plateau de Chesterfield, 19 ° 22 ′ S, 158 ° 43 ′ E, 61 m (1 spm); stn DW 56, Plateau de Chesterfield, 19 ° 18 ′ S, 158 ° 47 ′ E, 66 m (1 spm); stn DW 59, Plateau de Chesterfield, 19 ° 19 ′ S, 158 ° 57 ′ E, 50 m (1 spm); stn DW 65, Plateau de Chesterfield, 19 ° 15 ′ S, 158 ° 41 ′ E, 62 m (1 spm); stn DW 71, Plateau de Chesterfield, 19 ° 15 ′ S, 158 ° 24 ′ E, 55 m (HD 22467, 1 spm); stn DW 72, Plateau de Chesterfield, 19 ° 15 ′ S, 158 ° 21 ′ E, 32 m (1 spm); stn DW 156, Plateau de Chesterfield, 19 ° 49 ′ S, 158 ° 21 ′ E, 42 m (1 spm). – PALEO-SURPRISE, stn DW 1382, 18 ° 26.5 ′ S, 163 ° 05.9 ′ E, 41 – 42 m (1 spm). LOYALTY ISLANDS: both LIFOU 2000: Lifou, Santal Bay, stn 1423, 20 ° 54.0 ′ S, 167 ° 07.1 ′ E, 12 m (2 spm); Lifou, Santal Bay, stn 1444, 20 ° 55.0 ′ S, 167 ° 05.2 ′ E, 9 – 20 m (1 spm). VANUATU: all SANTO 2006: stn FB 56, NE of Tangisi I., 15 ° 35.2 ′ S, 167 ° 02.1 ′ E, 3 – 18 m (3 spm); stn DB 08, E of Aore I., 15 ° 34.6 ′ S, 167 ° 13.8 ′ E, 12 m (1 spm); stn DB 33, E of Aore I., 15 ° 34.7 ′ S, 167 ° 13.8 ′ E, 14 – 25 m (1 spm); stn ZB 36, E of Aore I., 15 ° 34.3 ′ S, 167 ° 12.4 ′ E, intertidal (1 spm); stn ZS 22, W of Aore I., 15 ° 31.1 ′ S, 167 ° 09.6 ′ E, 27 – 29 m (15 spm); stn ZB 16, Aore I., 15 ° 32.4 ′ S, 167 ° 12.1 ′ E, 5 m (2 spm); stn DB 12, S of Aore I., 15 ° 36.6 ′ S, 167 ° 10.1 ′ E, on sand (9 spm); stn LD 35, Aore I., 15 ° 32.7 / 32.8 ′ S, 167 ° 11.5 / 11.6 ′ E, 3 – 8 m (1 spm); stn DB 48, W of Malo I., 16 ° 38.7 ′ S, 167 ° 5.2 ′ E, 10 – 17 m (1 spm); stn DB 83, E of Malo I., 15 ° 43.4 ′ S, 167 ° 15.0 ′ E, on sand (2 spm); stn DS 49, Malo I., 15 ° 38.7 ′ S, 167 ° 05.2 ′ E, 10 – 17 m (4 spm); stn ZM 15, NW of Malo I., 15 ° 38.1 ′ S, 167 ° 05.9 ′ E, intertidal (1 spm); stn DB 75, SE of Matewulu, 15 ° 22.9 ′ S, 167 ° 11.9 ′ E, 20 m (4 spm); Segond Channel, stn DB 14, 15 ° 39.9 ′ S, 167 ° 11.0 ′ E, 10 – 14 m (3 spm); Segond Channel, stn DS 04, 15 ° 31.4 ′ S, 167 ° 14.1 ′ E, 25 m (33 spm; HD 38512, 2 spm; HK 995.12, 3 spm); Segond Channel, stn DS 06, 15 ° 30.9 ′ S, 167 ° 11.1 ′ E, 8 – 15 m (1 spm); Segond Channel, stn DS 91, 15 ° 33.7 ′ S, 167 ° 08.4 ′ E, 7 m (20 spm); Segond Channel, stn FS 54, 15 ° 31.4 ′ S, 167 ° 14.1 ′ E, 20 – 31 m (25 spm; HD 35303, 2 spm); Segond Channel, stn FS 77, 15 ° 33.1 ′ S, 167 ° 09.6 ′ E, 29 m (17 spm; HK 995.14, 2 spm); Segond Channel, stn FS 79, 15 ° 33.1 ′ S, 167 ° 09.6 ′ E, 2 m (3 spm); Segond Channel, stn FS 96, 15 ° 33.1 ′ S, 167 ° 09.6 ′ E, 35 m (6 spm); Segond Channel, stn FB 80, 15 ° 33.1 ′ S, 167 ° 09.6 ′ E, 2 m (1 spm); Segond Channel, stn LS 17, 15 ° 31.1 ′ S, 167 ° 10.5 ′ E, 7 m (2 spm); Segond Channel, stn ZS 19, 15 ° 31.4 ′ S, 167 ° 14.1 ′ E, 15 – 30 m (20 spm); Segond Channel, stn NB 12, 15 ° 33.1 ′ S, 167 ° 09.6 ′ E, 20 m (1 spm); Segond Channel, stn NS 37, 15 ° 31.4 ′ S, 167 ° 09.8 ′ E, 2 – 3 m (4 spm); Palikulo Bay, stn DB 53, 15 ° 28.8 ′ S, 167 ° 15.2 ′ E, 5 m (2 spm); Palikulo Bay, stn DB 65, 15 ° 25.8 ′ S, 167 ° 13.0 ′ E, 13 m (2 spm); Palikulo Bay, stn DB 77, 15 ° 27.9 ′ S, 167 ° 14.7 ′ E, 42 – 45 m (2 spm); Palikulo Bay, stn DS 43, 15 ° 27.9 ′ S, 167 ° 14.3 ′ E, 22 m (5 spm); Palikulo Bay, stn DS 54, 15 ° 28.8 ′ S, 167 ° 15.2 ′ E, 5 m (6 spm); Palikulo Bay, stn DS 59, Aésé I., 15 ° 24.6 ′ S, 167 ° 14.3 ′ E, 6 – 43 m (1 spm); Palikulo Bay, stn DB 16, Tutuba I., 15 ° 35.5 ′ S, 167 ° 15.8 ′ E, 32 – 40 m (4 spm); Palikulo Bay, stn DS 103, West Tutuba I., 15 ° 34.0 ′ S, 167 ° 16.0 ′ E, 70 – 80 m (4 spm); Palikulo Bay, stn LD 36, West Taluba I., 15 ° 39.5 ′ S, 167 ° 16.8 ′ E, 8 – 10 m (1 spm); Palikulo Bay, stn DS 105, NW of Tutuba I., 15 ° 33.0 ′ S, 167 ° 16.7 ′ E, 92 m (3 spm); Palikulo Bay, stn DS 110, Bruat Channel, 15 ° 36.8 ′ S, 167 ° 08.8 ′ E, 35 m (1 spm); Palikulo Bay, stn LD 12, Bruat Channel, 15 ° 36.6 ′ S, 167 ° 11.3 ′ E, 2 – 4 m (1 spm); Palikulo Bay, stn ZB 06, SW Urélapa I., 15 ° 36.8 ′ S, 167 ° 01.3 ′ E (1 spm); SE corner of SANTO 2006, shallow water (10 spm; NBC, 1 spm; HD 38511, 1 spm; HK 995.13, 2 spm). FIJI: MUSORSTOM 10, stn DW 1384, 18 ° 18.5 ′ S, 178 ° 06 ′ E, 260 – 305 m (1 spm). TONGA: BORDAU 2, stn DW 1581, 18 ° 41 ′ S, 174 ° 02 ′ E, 75 – 85 m (1 spm).	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC73D78334F38FE96FEF52589.taxon	description	Description PROTOCONCH. Approximately 2.25 shiny whorls (Fig. 2 A). SEM photography shows fine axial striae and minute pustules on last whorl. SHELL. Teleoconch consists of approximately 5 convex whorls. Suture deep. Penultimate whorl with 12 – 14 and body whorl with 14 – 16 pronounced and rounded axial ribs. Outer lip with a strong varix. SPIRAL CORDS. Penultimate whorl with 5 – 6 and body whorl with 10 – 11 continuous, narrow, string-like spiral cords, some more pronounced on axial ribs than in between them. INTERCORDAL SCULPTURE. Between spiral cords, spiral striae faintly visible on well-preserved specimens (Fig. 2 F). APERTURE. Aperture oval-rounded, outer lip with 5 – 7 fairly prominent, lirate denticles, peripheral denticle most prominent. Columella (Fig. 2 G – H) with 6 – 9 fine denticles, columellar callus sharply bordered and limited to columella. Parietal denticle strong, anal canal wide. OPERCULUM. Whitish, serrated. SIPHONAL CANAL. Fasciole strong, siphonal area with 5 – 7 weak cords, last one most prominent. COLOR. Creamy-white to yellowish brown, occasionally with 1 – 3 darker spiral bands. ADULT SIZE. 6.8 – 10.3 mm, usually 7.5 – 8.5 mm.	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC73D78334F38FE96FEF52589.taxon	discussion	Remarks Several species, previously synonymized with R. paupera by Cernohorsky (1984), have been taken out of synonymy in the past few years. A series of published reviews (Kool & Dekker 2006, 2007), including the present article, demonstrate that the Nassarius pauperus complex is composed of several valid species. Therefore, the synonymy given for Reticunassa paupera s. str. has become very short and caution is advised with citations of this species name in the literature. The combination of the multispiral protoconch, the many pronounced, round ribs, the deep suture, the broad varix, the very fine spirals, the sharply-bordered columella and the columellar denticles not lirae characterize this species, one of the smaller within the genus. The differences between R. paupera and the other species of Reticunassa, described below, will be discussed in the remarks section in each case. In the taxonomic history of Nassa paupera, several declensions on the specific epithet have been used. In Latin, there are two different adjectives with the same meaning: the classical pauper (3 rd declension, nominative unchanged) and the post-classical (medieval, generally clerical) pauperus (1 st declension, paupera feminine, pauperum neuter), both meaning “ poor. ” The second adjective is obviously a derivation of the first. The fact that Gould (1850) used paupera in his original description indicates that he was referring to the second meaning. Combined with Reticunassa, the new combination is R. paupera (Gould, 1850).	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC73D78334F38FE96FEF52589.taxon	biology_ecology	Habitat Specimens were collected from the intertidal to a depth of 82 m, mostly from 0 to 25 m; one empty shell was obtained at 260 m (Fiji Islands). The proposed neotype was collected on sand with dead corals. Specimens from Guam (HD 12797) were found in rubble.	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC73D78334F38FE96FEF52589.taxon	distribution	Distribution Widely distributed in the western Pacific Ocean, from Indonesia and the Philippines to the Tonga Islands (Fig. 4).	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC72378354F46FC22FD4320EA.taxon	description	Figs 2 E, 3 D – I, 5; Tables 1 – 2	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC72378354F46FC22FD4320EA.taxon	materials_examined	Type material NEW CALEDONIA: 6 syntypes of Nassa tringa (MHNBx 2014. TY. 89.0, 2 spm; MNHN IM- 2000 - 28389, 4 spm, http: // coldb. mnhn. fr / catalognumber / mnhn / im / 2000 - 28389). We designate MHNBx 2004. TY. 89.1 (from the former MHNBx 2004. TY. 89.0) as the lectotype (Fig. 3 F – H), length 9.0 mm, width 5.0 mm. NEW CALEDONIA: holotype of N. mamillata (NHMUK 1923.2.20.8). Other material examined NEW CALEDONIA: MONTROUZIER 1993: Touho, stn 1237, 20 ° 46.9 ′ S, 165 ° 13.8 ′ E, 0 – 1 m (15 spm; HD, 2 spm; HK 994.02, 3 spm); Touho, stn 1240, 20 ° 46.5 ′ S, 165 ° 14.0 ′ E, 0 – 2 m (19 spm; HK 994.01, 3 spm; HD 24146, 1 spm); Touho, stn 1241, 20 ° 48.0 ′ S, 165 ° 15.7 ′ E, 0 – 2 m (2 spm; NBC, 1 spm); Touho, stn 1242, 20 ° 46.2 ′ S, 165 ° 14.5 ′ E (29 spm); Touho, stn 1244, 20 ° 49.3 ′ S, 165 ° 15.2 ′ E (1 spm); Touho, stn 1245, 20 ° 45.2 ′ S, 165 ° 16.3 ′ E (3 spm); Touho, stn 1246, 20 ° 42.8 ′ S, 165 ° 08.7 ′ E (4 spm); Touho, stn 1253, 20 ° 46.6 ′ S, 165 ° 14.1 ′ E, 2 – 3 m (15 spm); Touho, stn 1259, 20 ° 44.6 ′ S, 165 ° 13.7 ′ E, 15 – 35 m (2 spm); Touho, stn 1261, 20 ° 46 ′ – 20 ° 47 ′ S, 165 ° 15 ′ – 165 ° 16.5 ′ E, 45 – 56 m (2 spm); Koumac, stn 1277, 20 ° 34 ′ S, 164 ° 16 ′ E, 0 – 2 m (4 spm); Koumac, stn 1278, 20 ° 34 ′ S, 164 ° 16 ′ E, 0 – 2 m (2 spm); Koumac, stn 1279, 20 ° 35 ′ S, 164 ° 15.5 ′ E (3 spm); Koumac, stn 1282, 20 ° 33.5 ′ S, 164 ° 13 ′ E (10 spm); Koumac, stn 1283, 20 ° 33.5 ′ S, 164 ° 12.2 ′ E (1 spm); Koumac, stn 1284, 20 ° 33.7 ′ S, 164 ° 19 ′ E (19 spm); Koumac, stn 1286, 20 ° 38 ′ – 20 ° 39 ′ S, 164 ° 16 ′ – 164 ° 17 ′ E (2 spm); Koumac, stn 1288, 20 ° 40 ′ S, 164 ° 19 ′ E (1 spm); Koumac, stn 1289, 20 ° 29.2 ′ S, 164 ° 10.2 ′ E (6 spm); Koumac, stn 1290, 20 ° 29.2 ′ S, 164 ° 10.2 ′ E (1 spm); Koumac, stn 1291, 20 ° 22.4 ′ S, 164 ° 06.8 ′ E (1 spm); Koumac, stn 1292, 20 ° 22.4 ′ S, 164 ° 06.8 ′ E (4 spm); Koumac, stn 1298, 20 ° 35.2 ′ S, 164 ° 16.6 ′ E, 2 – 4 m (1 spm). – LAGON: stn 77, Ile Ouen-Baie du Prony, 22 ° 26 ′ S, 166 ° 32 ′ E, 22 m (1 spm); stn 384 bis, Grand Récif Sud, 22 ° 34 ′ S, 167 ° 11 ′ E, 72 m (1 spm); stn 441, Atoll de Huon, 18 ° 04 ′ S, 162 ° 56 ′ E, 37 m (1 spm). – New Caledonia, leg. Rossiter, ex coll. Dautzenberg (ZMA. Moll. 098635, 5 spm); beach S of Noumea, leg. Mrs Wijsman-Best, 1968 (ZMA Moll. 096210, 2 spm); Ile des Pins (PS, 2 spm); Nouméa, leg. A. P. Greeven-Lels, 1965 (HD 21379, 9 spm). VANUATU: SANTO 2006: stn AT 52, Segond Channel, 15 ° 31.5 ′ S, 167 ° 12.7 ′ E, 52 – 62 m (MNHN IM- 2007 - 31753, 1 lv, l 9.5 mm, w 4.6 mm, Fig. 3 D – E, I); stn LM 23, Segond Channel, 15 ° 31.5 ′ S, 167 ° 09.6 ′ E, intertidal (1 spm); stn VM 53, Segond Channel, 15 ° 31.0 ′ S, 167 ° 11.9 ′ E, intertidal (6 spm); stn VM 69, Tutuba I., 15 ° 33.4 ′ S, 167 ° 16.7 ′ E, intertidal (MNHN IM- 2007 - 31758, 1 lv); stn FB 92, Tutuba I., 15 ° 33.6 ′ S, 167 ° 16.6 ′ E, 2 – 4 m (1 spm); stn VM 32, W of Aésé I., 15 ° 26.6 ′ S, 167 ° 15.2 ′ E, intertidal (2 spm). – Efate, snorkeling, 2 – 3 m, leg. T. McCleery, 2002 (HD 17091, 4 spm; HK 994.20, 3 spm). FIJI: Yasawa I., leg. S. P. Kool (coll. S. P. Kool, 3 spm; HK 994.26, 1 spm).	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC72378354F46FC22FD4320EA.taxon	description	Description PROTOCONCH. Whitish, paucispiral, consisting of 1.5 – 1.75 whorls (Fig. 3 I). SEM photograph shows very fine growth lines, and in addition minute pustules on part of last whorl. SHELL. Elongate-ovate, teleococh with 5 – 6 whorls, suture impressed. On penultimate whorl 13 – 17 and on body whorl 12 – 17 pronounced, round axial ribs, on ventral side of body whorl often decreasing in height towards base. SPIRAL CORDS. Continuous, moderately broad, rounded primary cords, approximately 4 – 5 on penultimate and approximately 10 – 12 on body whorl. INTERCORDAL SCULPTURE. Fine, evenly-spaced, continuous striae (Fig. 2 E), 7 – 9 on body whorl, sometimes visible between, as well as on spiral cords. APERTURE. Roundish; outer lip thick, plicate inside, with strong varix; peripheral denticle and more pronounced pleat near siphonal canal. Columella callused, callus sharply bordered, posteriorly slightly extending over body whorl. Columellar lip lirate throughout. Parietal denticle moderately strong. OPERCULUM. Yellowish, serrated. SIPHONAL CANAL. Narrow, with strong fasciole. Siphonal area with weak cords and in some specimens faint axial striae between cords. COLOR. Off white, yellowish to orange or brown. Banding very variable: some specimens with chestnut to dark brown, or nearly black bands; on penultimate whorl a sub- and / or a supra-sutural band; on body whorl occasionally one band directly below periphery and one at base. Occasionally, some or most spiral cords darker, especially between axial ribs. Aperture yellowish, showing outside banding, if present. ADULT SIZE. 6.7 – 12.2 mm, usually 8 – 9.5 mm.	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC72378354F46FC22FD4320EA.taxon	discussion	Remarks Characteristics of this species are a paucispiral protoconch, low spiral cords, in combination with numerous, evenly-spaced intercordal spiral striae (Fig. 2 E), axial ribs decreasing in height anteriorly on the ventral side of the body whorl, and the presence of lirae on the columellar callus. In wellpreserved specimens, spiral striae are also visible on top of the spiral cords. The intercordal spiral striae were not mentioned in the original description by Souverbie (in Souverbie & Montrouzier, 1864). However, Preston (1907) mentioned this sculpture in the description of Nassa mamillata. We agree with Cernohorsky (1984), that the latter is a synonym of Reticunassa tringa. R. tringa can easily be distinguished from R. paupera by its paucispiral protoconch, probably figured by Cernohorsky (1984: 5, fig. 12). Cernohorsky did not discuss protoconch morphology or other conchological features shared by species within the Nassarius pauperus complex, resulting in numerous synonyms of R. paupera, followed by many authors. R. tringa closely resembles R. visayaensis sp. nov. and R. poppeorum sp. nov. For the differences between R. tringa and these species, see respective Remarks sections.	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC72378354F46FC22FD4320EA.taxon	biology_ecology	Habitat Intertidal to 72 m, more frequently within the first 10 m of depth.	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC72378354F46FC22FD4320EA.taxon	distribution	Distribution New Caledonia, Vanuatu and Fiji (Fig. 5).	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC72578364F24F8BFFD1D2062.taxon	description	urn: lsid: zoobank. org: act: 742 DB 3 E 4 - 4 D 0 F- 40 DD- 985 C- 948 F 113 C 4 C 65 Fig. 3 J – M, 6; Tables 1 – 2	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC72578364F24F8BFFD1D2062.taxon	etymology	Etymology The name refers to the Visayas, a group of central Philippine Islands where this species is common.	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC72578364F24F8BFFD1D2062.taxon	materials_examined	Type material Holotype PHILIPPINES: lv, Panglao Island, 9 ° 35.7 ′ N, 123 ° 44.7 ′ E, 0 – 3 m, seagrass and hard bottom, length 9.5 mm, width 4.6 mm, sequenced (MNHN IM- 2007 - 31912). Paratypes PHILIPPINES: Panglao Island, Momo Beach, 9 ° 36.1 ′ N, 123 ° 45.2 ′ E, 0 – 3 m (MNHN IM- 2000 - 28395, 9.2 mm; MNHN IM- 2000 28405; MNHN IM- 2000 - 28404, 3 spm); Panglao I. area (HD 24336, 60 spm; HK 184.13, 2 spm). Other material examined PHILIPPINES: PANGLAO 2004: Panglao I., stn B 8, Napaling, 9 ° 37.1 ′ N, 123 ° 46.1 ′ E, 3 m (2 spm); Panglao I., stn M 1, Alona Beach, 9 ° 32.9 ′ N, 123 ° 50.5 ′ E, 5 m (2 spm); Panglao I., stn M 5, Doljo Point, 9 ° 35.5 ′ N, 123 ° 43.3 ′ / 123 ° 44.3 ′ E, 0 – 2 m (1 spm); Panglao I., stn M 7, Momo Beach, 9 ° 36.1 ′ N, 123 ° 45.2 ′ E, 0 – 3 m (2 spm); Panglao I., stn M 9, near Doljo Point, 9 ° 35.1 ′ N, 123 ° 43.6 ′ E, 0.5 m (1 spm); Panglao I., stn M 10, Bingag / Tabalong, 9 ° 37.8 ′ N, 123 ° 48.4 ′ E, 0 – 3 m (1 spm); Panglao I., stn M 22, Napaling, 9 ° 37.2 ′ N, 123 ° 46.4 ′ E, 0 – 3 m (1 spm); Panglao I., stn M 40, Looc, 9 ° 35.7 ′ N, 123 ° 44.7 ′ E, 0 – 3 m (1 spm); Panglao I., stn R 19, Napaling, 9 ° 37.1 ′ N, 123 ° 46.1 ′ E, 2 – 54 m (2 spm); Panglao I., stn S 24, Momo Beach, 9 ° 36.1 ′ N, 123 ° 45.0 ′ E, 2 – 4 m (1 spm); Pamilacan I., stn S 42, 9 ° 30.1 ′ N, 123 ° 55.5 ′ E, 15 – 20 m (1 spm). – Cebu, leg. R. Martin (ZMA. Moll. 175495, 10 spm); Cebu (HK 184.09, 1 spm; ZMA Moll. 080981, 1 spm); Cebu, Liloan, 50 – 150 m, 2013 (HD 35803, 1 spm); Mactan I., Maribago, Buyon, in sandy tide pools, leg. O. K. McCausland (HK 184.07, 1 spm); Punta Engaño, in tangle nets, 150 m (HK 184.05, 3 spm); Punta Engaño, in coral rubble, 20 m (HK 184.14, 3 spm); Punta Engaño (HD 11610, 8 spm); Punta Engaño, 80 – 100 m (HD 17300, 3 spm); Mactan I., Punta Engaño, 25 – 50 m (HD 33879, 1 spm); Punta Engaño, leg. S. P. Kool, 2005 (HK 184.11, 1 spm); Punta Engaño, 1 – 2 m, live (HK 184.12, 2 spm); Punta Engaño, 25 – 30 m, 2009 (AMD, 2 spm); Punta Engaño, 100 – 150 m, 2009 (AMD, 2 spm); Balicasag I. (HD 15930, 2 spm); Olango I., night dive, 20 – 25 m (HD 17292, 3 spm); Olango I., 2014 (HD 35872, 4 spm); Olango I., 20 – 25 m, 2009 (AMD, 2 spm); Cuyo I., 15 – 20 m (HD 35488); Calituban I., 10 m (HK 184.01, 6 spm); SW side of Catanduanes, San Rafael, leg. S. P. Kool, 2005 (HK 184.10, 1 spm); Aliguay I., tangle nets, 150 – 180 m (HK 184.08, 2 spm); Palawan, 10 – 25 m, 2009 (AMD, 2 spm). INDONESIA: SNELLIUS 1929: Ternate I., 1 – 2 Apr. 1930 (RMNH, 1 spm); Tidore I., 24 – 29 Nov. 1929 (RMNH, 1 spm). – RUMPHIUS 1990: Ambon, stn 05 Leitimur, Ambon Bay, outer bay, Tg. Benteng (RMNH, 2 spm); Ambon, stn 17, SE side of Pombo I. (RMNH, 2 spm); Ambon, stn 20 Hitu, N coast, Hitulama (RMNH, 1 spm); Ambon, stn 21 Hitu, N coast, Mamala (RMNH, 2 spm); Ambon, stn 23 Hitu, Kaitetu (near Hila), 22 – 23 Nov. 1990 (RMNH, 1 spm); Ambon, stn 26 Hitu, 4 km W of Kaitetu, 23 Nov. 1990 (RMNH, 3 spm); Ambon, stn 27 Leitimur, S coast, Hutumuri (RMNH, 1 spm); Ambon, stn 34 Hitu, Ruhmatiga 3 – 5 Dec. 1990 (RMNH, 4 spm; HK 184.03, 1 spm). – LAGON, stn Seith, Karubar, Amboine, low tide (1 spm); South Moluccas, leg. Rijkschroeff (ZMA. Moll. 096203, ex coll. Butot 12403, 7 spm); South Moluccas, leg. Rijkschroeff (ZMA. Moll. 096204, ex coll. Butot 12404, 5 spm); Ceram, N coast, Seleman Bay, leg. H. Strack (HK 184.04, 1 spm); Sulawesi, Lintido, leg. Semmelink (ZMA. Moll. 096205, ex coll. Schepman, 1 spm); Bali, Sanur, leg. K. van Duin, 1989 (ZMA. Moll. 099329, 1 spm); Bali, Kaliyasem, Lovina area, 8 ° 09.7 ′ S, 115 ° 01.7 ′ E, 1 – 2 m (HK 184.16, 3 spm); Flores, Labuan Bajo, Binongko Beach, leg. J. N. J. Post (HK 184.15, 1 spm); West Papua, Manokwari, near Uriami River, leg. D. Smits 1958 / 1961 (HK 184. 02, 11 spm; HD 38514, 11 spm). PAPUA NEW GUINEA: PAPUA NIUGINI: stn PM 22, Sek I., 05 ° 04.7 ′ S, 145 ° 48.9 ′ E, 0 – 1 m (MNHN IM- 2013 - 13184; MNHN IM- 2013 - 13192, 2 lv); stn PM 19, Islet SE of Megas Islet, 05 ° 05.4 ′ S, 145 ° 48.6 ′ E, 0 – 1 m (2 spm); stn PB 23, Lauhamug I., outer slope, 04 ° 59.5 ′ S, 145 ° 47.7 ′ E, 13 m (1 spm); stn DP 31, Alexishafen, 05 ° 05.3 ′ S, 145 ° 48.1 ′ E, 1 – 6 m (4 spm); stn PM 38, Biliau I., 05 ° 11.8 ′ S, 145 ° 48.2 ′ E (1 spm); stn PD 78, Tabad I., 05 ° 08.2 ′ S, 145 ° 48.7 ′ E, 5 m (1 spm); stn PR 202, S of Tab I., 05 ° 10.3 ′ S, 145 ° 50.3 ′ E, 2 – 4 m (2 spm).	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC72578364F24F8BFFD1D2062.taxon	description	Description Holotype PROTOCONCH. Smooth, multispiral, whitish-yellowish, consisting of 2.5 whorls. Beset with rows of minute pustules (Fig. 3 M). SHELL. Elongate-ovate, 5.5 postnuclear whorls, suture impressed. Penultimate whorl with 15 nearly equally pronounced, round ribs; body whorl with 14 ribs, decreasing in height or disappearing on ventral side. Varix broad and strong. SPIRAL CORDS. Continuous, flat and narrow, 7 cords on penultimate, and 11 on body whorl, peripheral one somewhat darker. INTERCORDAL SCULPTURE. Approximately 6 very fine, evenly spaced striae between spiral cords. APERTURE. Oval, inside outer lip with 9 lirate denticles, peripheral denticle slightly more pronounced. Parietal denticle moderate, anal canal wide. Columellar callus wide, anteriorly somewhat elevated, posteriorly partly extending over whorl; well delineated border. Fine lirae over entire surface. OPERCULUM. Yellowish, serrated. SIPHONAL CANAL. Fasciole strong, siphonal area with 1 strong and 5 weak cords. COLOR. Yellowish, most spiral cords reddish between ribs. ADULT SIZE. 8.5 – 12.7 mm, usually 8.5 – 10.5 mm.	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC72578364F24F8BFFD1D2062.taxon	discussion	Remarks The intraspecific variability is considerable. The number of ribs and spiral cords may vary, and the color is extremely variable, white to yellow and orange to brown, unicolor or with narrow or broad yellow, brown, dark brown, or grey bands on all whorls or only on the body whorl. Reticunassa visayaensis sp. nov. has a broader, less pointed protoconch than R. paupera (Gould, 1850). R. visayaensis sp. nov. also differs from R. paupera by its larger size, its more bulbous shape and its lower ribs, especially on the body whorl, and by usually displaying dark bands. R. visayaensis sp. nov. is very similar to R. tringa. When the protoconch is missing, a positive identification is almost impossible. The protoconch of R. tringa is paucispiral with 1.5 – 1.75 whorls, whereas R. visayaensis sp. nov. has a protoconch of 2.25 – 2.5 whorls. The protoconch of R. tringa is nipple-shaped, hence the name “ mamillata ” (Preston 1907); the protoconch of R. visayaensis sp. nov. is dome-shaped. Preston’s description is accompanied by a drawing of the paucispiral protoconch. R. visayaensis sp. nov. is the Reticunassa species most commonly offered in the shell trade from the central Philippines, as well as R. crenulicostata (Shuto, 1969). The latter is smaller (5 – 7 mm) and has a large multispiral protoconch of 3.5 whorls (Cernohorsky 1984: pl. 38, figs 1 – 2; Martin 2008: pl. 354, figs 4 – 5). These features are the most distinguishing differences between R. crenulicostata and R. visayaensis sp. nov. Reticunassa tringa has a paucispiral protoconch. Identification based on geographical distribution alone (Fig. 6) may be possible in specimens lacking a protoconch or welldefined teleoconch sculpture.	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC72578364F24F8BFFD1D2062.taxon	biology_ecology	Habitat Intertidal to 150 m, commonly from 0 to 20 m.	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC72578364F24F8BFFD1D2062.taxon	distribution	Distribution Philippines, Indonesia and New Guinea (Fig. 6).	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC726783A4F2BF907FC1225FF.taxon	description	urn: lsid: zoobank. org: act: 21521 D 97 - 7 E 7 D- 4627 - A 806 - 43 AF 0 E 2 AE 7 EA Fig. 3 N – Q, 7; Tables 1 – 2	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC726783A4F2BF907FC1225FF.taxon	etymology	Etymology The name is given to honor Guido T. and Philippe Poppe for their many valuable contributions to the knowledge of and myriad publications on molluscs, including four volumes on Philippine marine molluscs and the magazine “ Visaya. ”	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC726783A4F2BF907FC1225FF.taxon	materials_examined	Type material Holotype VANUATU: SANTO 2006, stn FB 61, Aore Island, Port Benier, 15 ° 34.4 ′ S, 167 ° 12.6 ′ E, 2 – 3 m, in fine coral sand (MNHN IM- 2007 - 31779, lv, length 9.0 mm, width 4.5 mm). Paratypes VANUATU: SANTO 2006, stn DR 03, Segond Channel, 15 ° 31.0 ′ S, 167 ° 15.8 ′ E, 2 – 30 m, reef slope, fine rubble with Halimeda (MNHN IM- 2007 - 32742, 1 lv); Éfate, snorkeling, 2 – 3 m, leg. T. McCleery, 2002 (HD 35802, 1 spm); Tutuba I., 15 ° 32.3 ′ S, 167 ° 16.9 ′ E, 13 m (HK 137.23, 1 spm). PHILIPPINES: PANGLAO 2004, stn S 2, Bohol I., Baclayon, 9 ° 37.4 ′ N, 123 ° 54.5 ′ E, 4 – 5 m (MNHN IM- 2007 - 31938, MNHN IM- 2007 - 31937, 2 lv; HK 137.21, 2 spm); stn S 15, Cortes Takot, 9 ° 41.3 ′ N, 123 ° 49.5 ′ E, 4 – 6 m (MNHN IM- 2000 - 22721, 1 lv; HK 137.01, 3 spm; HD 24148, 2 spm). Other material examined MALAYSIA: Sarawak, South China Sea, Muri I., off NW coast, 01 ° 54 ′ 13 ″ N, 108 ° 38 ′ 49.4 ″ E, 10 – 30 m, leg. H. Morrison (GH, 1 spm); Sarawak, Miri, outer reef off Tanjung Lobang (JGMR, 1 spm); Sabah, Kota Kinabalu, Tanjung Aru (JGMR, 1 spm). PHILIPPINES: PANGLAO 2004, stn S 2, Bohol I., Baclayon, 9 ° 37.4 ′ N, 123 ° 54.5 ′ E, 4 – 5 m (41 spm); stn S 15, Bohol I., Cortes Takot, 9 ° 41.3 ′ N, 123 ° 49.5 ′ E, 4 – 6 m (15 spm; ZMA. Moll. 4.09.449, 1 spm); stn S 1, Panglao I., Biking, 9 ° 35.6 ′ N, 123 ° 50.5 ′ E, 5 m (2 spm); stn B 5, Panglao I., Biking, 9 ° 35.2 ′ N, 123 ° 50.4 ′ E, 4 m (2 spm); stn B 8, Panglao I., Napaling, 9 ° 37.1 ′ N, 123 ° 46.1 ′ E, 3 m (3 spm); stn S 5, Panglao I., Napaling, 9 ° 37.1 ′ N, 123 ° 46.1 ′ E, 2 – 4 m (10 spm); stn S 7, Panglao I., Sungcolan Bay, 9 ° 38.5 ′ N, 123 ° 49.9 ′ E, 1 – 4 m (2 spm); stn S 17, Panglao I., San Isidro, 9 ° 34.6 ′ N, 123 ° 49.9 ′ E, 6 m (1 spm); stn S 24, Panglao I., Momo Beach, 9 ° 36.1 ′ N, 123 ° 45.0 ′ E, 2 – 4 m (1 spm); stn S 32, Panglao I., Looc, 9 ° 35.8 ′ N, 123 ° 44.6 ′ E, 2 – 3 m (12 spm); Pamilacan I., stn B 11, 9 ° 29.4 ′ N, 123 ° 56.0 ′ E, 2 – 4 m (1 spm); Pamilacan I., stn B 24, 929.4 ′ N, 123 ° 56.0 ′ E, 38 m (2 spm); Pamilacan I., stn S 10, 9 ° 29.4 ′ N, 123 ° 56.0 ′ E, 6 – 14 m (2 spm); Pamilacan I., stn S 14, 9 ° 29.3 ′ N, 123 ° 55.1 ′ E, 5 – 12 m (1 spm). – Cebu, leg. R. Martin (ZMA. Moll. 175495, 2 spm); Cebu, by lumen lumen, 50 – 150 m (HK 137.19, 2 spm); Panglao I. area (HD 14809, 15 spm; HK 137.14, 1 spm); Siquijor I., tangle nets, 100 m (HD 17294, 1 spm); Balicasag I. (MUZEE, 2 spm); Mactan I., 10 – 30 m, 2012 (HD 28994, 7 spm); Mactan I., 20 m, 2014 (HD 35816, 1 spm); Olango I., 2014 (HD 35873, 1 spm); Camotes I., Santiago, 15 m, 2007 (HD 21366, 1 spm); Palawan I., 10 – 15 m, 2009 (HD 33880, 2 spm). INDONESIA: Kalimantan, Berau Is, 02 ° 12 ′ 16.3 ″ N, 118 ° 35 ′ 18.9 ″ E, 10 – 25 m (ZMA. Moll. 175383, 1 spm); RUMPHIUS 1990, Ambon, stn 39 Hitu, W coast, S side of Larike (RMNH, 1 spm); Nusa Tenggara, Alor Strait, SE bay of Pantar I., 9 – 25 m, leg. H. Morrison (GH, 1 spm); Banda Sea, Wakatobi Group, E side of Moromaho I., 06 ° 08 ′ 59 ″ S, 124 ° 35 ′ 11 ″ E, leg. H. Morrison (GH, 6 spm); Flores Sea, S of Buton, 7 – 40 m, leg. H. Morrison (HK 137.08, 2 spm); N coast of Nusa Penida, leg. H. Morrison (GH, 1 spm); NW side of Sulawesi, Makassar Strait, Tambu Bay, sand and rubble slope, 10 – 40 m, among gorgonian corals, 00 ° 01 ′ 07 ″ S, 119 ° 43 ′ 77 ″ E (GH, 6 spm); Karang Kapota Atoll, SE of Buton I., 5 ° 27 ′ 73 ″ S, 123 ° 24 ′ 10 ″ E, on ledges of wall, 10 – 40 m, leg. H. Morrison (GH, 1 spm); Berau I., W side of Maratua I., 02 ° 12 ′ 16.3 ″ N, 118 ° 35 ′ 18.9 ″ E, 10 – 25 m (ZMA. Moll. 175383, 1 spm); near Kalimantan, Pejantan I., 0 ° 30 ′ 48.8 ″ N, 107 ° 13 ′ 24 ″ E, 10 – 20 m, leg. H. Morrison (GH, 1 spm); sand, near wreck of Japanese fighter, 0 ° 45 ′ 16 ″ N, 105 ° 36 ′ 11 ″ E, 45 m, leg. H. Morrison (GH, 2 spm); near wreck, Acasta Rock, 01 ° 39 ′ 12 ″ N, 106 ° 18 ′ 26 ″ E, 9 – 50 m (GH, 1 spm); sand, near wreck, 0 ° 45 ′ 15 ″ N, 105 ° 36 ′ 11 ″ E, leg. H. Morrison (GH, 2 spm); Badas I. Group, 0 ° 33 ′ 8.9 ″ N, 106 ° 58 ′ 47.4 ″ E, 8 – 30 m, leg. H. Morrison (GH, 1 spm); 01 ° 38 ′ 7 ″ N, 106 ° 21 ′ 23 ″ E, on reef, 10 – 25 m, leg. H. Morrison (GH, 1 spm); Sebangmawang I., S of Bunguran I., 03 ° 33.92 ′ N, 108 ° 02.08 ′ E, 10 – 15 m, leg. H. Morrison (GH, 4 spm); Sumatra, Kelulekabung I. (HK 137.07, 1 spm); West Papua, Manokwari, mouth of Uriami River, leg. D. Smits (HK 137.06, 2 spm); Bay of Hollandia, leg. Mrs H. Grootens-Boerefijn (HK 137.11, 6 spm); off NW coast, Waigeo I., Sorong, leg. H. Morrison (HK 137.04, 2 spm); off Sorong, Kri I., 10 – 15 m, leg. H. Morrison (HK 137.05, 2 spm). PAPUA NEW GUINEA: PAPUA NIUGINI: stn PB 18, Sek I., outer slope, 05 ° 06.3 ′ S, 145 ° 49.1 ′ E, 26 m (1 spm); stn PB 28, E of Kranket I., 05 ° 11.9 ′ S, 145 ° 49.6 ′ E, 10 m (2 spm); stn PB 45, Sinub I., 05 ° 07.9 ′ S, 145 ° 48.9 ′ E, 8 m (4 spm); stn PB 47, N of Kranket I., 05 ° 11.3 ′ S, 145 ° 49.6 ′ E, 5 m (MNHN IM- 2013 - 17574, 1 lv); stn PB 47, N of Kranket I., 05 ° 11.3 ′ S, 145 ° 49.6 ′ E, 5 m (3 spm); stn PB 48, N of Riwo, 05 ° 08.7 ′ S, 145 ° 48.2 ′ E, 2 m (MNHN IM- 2013 - 17567, 1 lv); stn PB 51, N of Bilbil I., 05 ° 17.7 ′ S, 145 ° 46.9 ′ E, 5 m (3 spm); stn PD 31, Alexishafen, 05 ° 05.3 ′ S, 145 ° 48.1 ′ E, 1 – 6 m (1 spm); stn PD 32, N of Sek I., Ottilien Passage, 05 ° 04.4 ′ S, 145 ° 48.7 ′ E, 1 – 8 m (1 spm); stn PD 36, Rempi Area, W of Barag I., 05 ° 01.2 ′ S, 145 ° 47.9 ′ E, 5 – 10 m (1 spm); stn PM 19, Alexishafen, islet SE of Megas Islet, 05 ° 05.4 ′ S, 145 ° 48.6 ′ E, 0 – 1 m (1 spm); stn PM 22, Sek I., 05 ° 04.7 ′ S, 145 ° 48.9 ′ E, 0 – 1 m (MNHN IM- 2013 - 13185, 1 lv); stn PM 25, Rempi Area, Barag I., 05 ° 01.1 ′ S, 145 ° 47.9 ′ E (1 spm); stn PR 202, S of Tab I., 05 ° 10.3 ′ S, 145 ° 50.3 ′ E, 2 – 4 m (MNHN IM- 2013 - 17555, IM- 2013 - 17556, IM- 2013 - 17557, IM- 2013 - 17558, IM- 2013 - 17559, IM- 2013 - 17560, 6 lv); stn PS 04, Tab I., 05 ° 10.0 ′ S, 145 ° 50.1 ′ E, 12 m (1 spm); stn PS 27, Bilbil I., 05 ° 17.9 ′ S, 145 ° 46.7 ′ E, 14 m (1 spm); stn PS 41, S of Urembo I., outer slope, 05 ° 15.9 ′ S, 145 ° 47.1 ′ E, 10 m (HK 137.20, 3 spm); stn PS 45, W Yabob I., 05 ° 15.4 ′ S, 145 ° 47.0 ′ E, 6 m (MNHN IM- 2013 - 17562, 1 lv); stn PS 47, N of Sek I., inner slope, 05 ° 04.7 ′ S, 145 ° 48.9 ′ E, 8 m (1 spm). – Bismarck Archipelago, New Ireland, Kavieng, Baudisson Bay, 15 – 35 m, leg. H. Morrison (HK 137.03, 2 spm). SOLOMON ISLANDS: SALOMON 2, stn DW 2234, Choiseul I., 6 ° 51 ′ S, 156 ° 24 ′ E, 182 – 277 m (1 spm); SALOMONBOA 3, stn DW 2827, N of San Cristobal, 10 ° 26 ′ S, 161 ° 51 ′ E, 134 – 272 m (2 spm). FIJI: Suva Point, leg. S. P. Kool (HK 137.15, 1 spm); Yasawa Group, Wayasewa I., 17 ° 21 ′ S, 177 ° 08 ′ E, intertidal (HK 137.17, 1 spm). SAMOA: Savaii I., Asau Bay (PS, 4 spm; HK 137.10, 2 spm).	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC726783A4F2BF907FC1225FF.taxon	description	Description Holotype PROTOCONCH. Multispiral consisting of 2.5 whitish whorls, body whorl with rows of microscopic pustules, otherwise smooth (Fig. 3 Q). SHELL. Teleoconch consisting of 5.5 whorls; penultimate whorl with 15 and body whorl with 12 rounded ribs of equal size; broad varix. SPIRAL CORDS. Body whorl with approximately 10 continuous, weak spiral cords. INTERCORDAL SCULPTURE. Very fine, evenly spaced spiral striae, occasionally not only between, but also on spiral cords. APERTURE. Six low denticles on outer lip, that at periphery more pronounced. Columellar callus elevated anteriorly, backside of elevation with microscopic growth lines. Callus partly extending posteriorly over body whorl onto anal canal. Columellar lip lirate onto moderately strong parietal denticle; anal canal deep. OPERCULUM. Cream colored, serrated. SIPHONAL CANAL. Narrow. A strong fasciole; siphonal area with several fine, one dominating, cords. COLOR. Whitish to yellowish; some spiral cords light brown between ribs. Lower half of dorsal side of body whorl frequently with broad, orange-brown band (Fig. 3 O). ADULT SIZE. 6.9 – 12.0 mm.	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC726783A4F2BF907FC1225FF.taxon	discussion	Remarks The color varies from white to yellow and orange; the spiral cords occasionally brown, sometimes only locally between the axial ribs. Some specimens have 2 brown bands on the body whorl, occasionally only visible on the varix. Some specimens, including the holotype, display a broad orange-brown band on the lower part of the body whorl. R. poppeorum sp. nov. is often recognizable by a broad, dark orange-brown band on the last half or the last third of the ultimate whorl, and is characterized by its more slender shape compared to other species in this genus. Reticunassa poppeorum sp. nov. shows resemblance to R. tringa and R. visayaensis sp. nov. The greatest difference among the three species is the morphology of the protoconch. R. tringa has a paucispiral protoconch with 1.5 – 1.75 whorls, R. visayaensis sp. nov. has a multispiral protoconch of 2.25 – 2.5 whorls, whereas the protoconch of R. poppeorum sp. nov. has 2.5 – 2.75 whorls. The axial ribs of R. poppeorum sp. nov. are more pronounced and are present on the entire body whorl, whereas they decrease in height on the ventral side of the body whorl in R. tringa and R. visayaensis sp. nov. The most noticeable difference between R. paupera on the one hand and R. tringa, R. visayaensis sp. nov. and R. poppeorum sp. nov. on the other, is the presence of lirae in the latter three species over the entire surface of the extended columella. Besides this sculptural feature, the shape of the columellar callus differentiates R. paupera from the other three species; the callus of R. paupera is practically limited to the columella, whereas the callus of the other three partly extends over the body whorl. There are no morphological differences between R. neoproducta Kool & Dekker, 2007 and the closely related R. poppeorum sp. nov. (Fig. 3 N – Q). However, we decided to considerer these two lineages as different species based on the reciprocal monophyly indicated by the taxonomic trees (COI and 28 S separately). Furthermore, R. neoproducta is found in the western Indian Ocean and R. poppeorum occurs in the western Pacific Ocean. To date, no specimens of these two species have been found east of Sri Lanka or west of Sumatra, respectively. It seems very unlikely that these two allopatric lineages still share any gene flux.	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC726783A4F2BF907FC1225FF.taxon	biology_ecology	Habitat In sand and coral rubble, 0 – 40 m, mainly between 1 to 20 m. Some freshly dead specimens with operculum were collected on a hard bottom with small pockets of sediment and in coral sand at a depth of 4 – 6 m. Some specimens were collected in mangrove areas and in seagrass.	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC726783A4F2BF907FC1225FF.taxon	distribution	Distribution Malaysia, Indonesia, the Philippines, Fiji, and Western Samoa (Fig. 7).	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC72A783F4F2EFB88FD0627E0.taxon	description	urn: lsid: zoobank. org: act: 781 F 25 AD-B 216 - 47 A 1 - 8084 - FC 46 ED 4 C 8029 Fig. 8 A – D, 9; Tables 1 – 2	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC72A783F4F2EFB88FD0627E0.taxon	etymology	Etymology The species is dedicated to Mrs Anna Bolte, the second author’s partner and travel companion for more than 20 years.	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC72A783F4F2EFB88FD0627E0.taxon	materials_examined	Type material Holotype MADAGASCAR: ATIMO VATAE, stn BM 06, South Madagascar, Ambatobe, Bavarama, 25 ° 27.9 ′ S, 44 ° 57.6 ′ E, 0 – 1 m, in rolling stones (MNHN IM- 2009 - 12843, lv, sequenced, length 11.5 mm, width 5.3 mm). Paratypes MADAGASCAR: same collection data as holotype (MNHN IM- 2009 - 12812, MNHN IM- 2009 - 12849, MNHN IM- 2009 - 12862, MNHN IM- 2009 - 12868, 4 lv; HK 302.01, 3 spm; HD 37306, 2 spm); S coast, Lavanono (HK 302.05, 5 spm; HK 302.06, 2 spm). Other material examined MADAGASCAR: ATIMO VATAE, South Madagascar: stn BB 03, Lavanono, 25 ° 26.4 ′ S, 44 ° 56.1 ′ E, 14 – 18 m (MNHN IM- 2009 - 12865, 1 lv); stn BM 02, Ambatobe, Soamanitse, 25 ° 27.4 ′ S, 44 ° 57.4 ′ E, 0 – 1 m (MNHN IM- 2009 - 12772, 1 lv); stn BM 06, Ambatobe, Bavarama, 25 ° 27.9 ′ S, 44 ° 57.6 ′ E, 0 – 1 m (MNHN IM- 2009 - 12778, MNHN IM- 2009 - 12783, MNHN IM- 2009 - 12825, MNHN IM- 2009 - 12841, MNHN IM- 2009 - 12844, 5 lv); stn BM 09, Saririaka, 25 ° 29.0 ′ S, 44 ° 59.6 ′ E, 0 – 1 m (MNHN IM- 2009 - 12228, 1 lv); stn BM 10, Andramara, 25 ° 28.8 ′ S, 44 ° 58.3 ′ E, 0 – 1 m (MNHN IM- 2009 - 12873, 1 lv); stn BS 14, Cap Sainte Marie, 25 ° 36.0 ′ S, 45 ° 08.7 ′ E, 16 m (MNHN IM- 2009 - 12329, 1 lv); stn CP 3547, Cap Andavaka, 25 ° 18.0 ′ S, 46 ° 40.3 ′ E, 69 – 70 m (MNHN IM- 2009 - 12893, 1 lv); stn TM 01, Monseigneur Bay, 25 ° 02.1 - 2 ′ S, 46 ° 59.9 ′ E, 0 – 1 m (MNHN IM- 2009 - 12793, 1 lv); stn TM 05, Lokaro I., 24 ° 56.5 ′ S, 47 ° 07.1 ′ E, 0 – 1 m (MNHN IM- 2009 - 12306, MNHN IM- 2009 - 12888, MNHN IM- 2009 - 12889, MNHN IM- 2009 - 12897, MNHN IM- 2009 - 12899, MNHN IM- 2009 - 12905, MNHN IM- 2009 - 13087, 7 lv); stn TM 08, NE of Evatra lighthouse, 24 ° 58.7 ′ S, 47 ° 05.9 ′ E, 0 – 1 m (MNHN IM- 2009 - 12894, 1 lv); stn TS 09, Lokaro Bay, Fort Dauphin, 24 ° 57.0 ′ S, 47 ° 06.4 ′ E, 5 – 6 m (MNHN IM- 2009 - 12834, MNHN IM- 2009 - 12881, 2 lv); stn TS 10, Lokaro Bay, Fort Dauphin, 24 ° 56.8 ′ S, 47 ° 06.6 ′ E, 7 m (HK 302.02, 2 spm); stn TS 21, Phare Flacourt, 25 ° 01.5 ′ S, 47 ° 00.0 ′ E, 2 – 4 m (HK 302.03, 2 spm); stn TV 07, Phare Flacourt, Amparihy, 25 ° 01.3 ′ S, 47 ° 00.2 ′ E, 12 – 16 m (MNHN IM- 2009 - 12885, MNHN IM- 2009 - 12898, 2 lv). – South Madagascar (HK 302.04, 1 spm); Lavanono (HD 21156, 2 spm); Lavanono (HD 36646, 21 spm); Soamanitse (HD 20287, 3 spm).	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC72A783F4F2EFB88FD0627E0.taxon	description	Description Holotype PROTOCONCH. Paucispiral consisting of 1.5 – 1.75 broad, flattened, axially-striated whorls. Separation between protoconch and teleoconch indistinct (Fig. 8 D). SHELL. Elongate-ovate, consisting of 5.5 convex whorls, suture deep; axial ribs poorly developed, decreasing in height on body whorl; 16 ribs on penultimate whorl and 14 on body whorl; broad varix. SPIRAL CORDS. Fine, weak compared to other species in genus, more pronounced where they cross axial ribs. Two anterior spiral cords more developed, resulting in two deep spiral grooves. INTERCORDAL SCULPTURE. Very fine spiral striae between spiral cords. APERTURE. Oval, inside outer lip 8 lirate denticles; columellar callus plicate throughout, somewhat elevated anteriorly, posteriorly slightly extended over body whorl; parietal denticle weak, anal canal wide; siphonal area with 4 – 5 low cords. OPERCULUM. Yellowish, serrated. SIPHONAL CANAL. Deep and narrow. Fasciole deep. COLOR. Yellow-orange, narrow brown band above suture and broad brown band below periphery. Most spiral cords darker. ADULT SIZE. 7.5 – 12.6 mm, usually 9 – 11 mm.	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC72A783F4F2EFB88FD0627E0.taxon	discussion	Remarks Besides the variability in color, from off-white to yellowish-orange, the difference in size of adult specimens is remarkable. Some specimens have somewhat more developed spiral cords. Comparison with some other Reticunassa species with a paucispiral protoconch: R. tringa is a smaller, more bulbous species with a pointed protoconch; R. goliath sp. nov. also has fine spiral cords as found in R. annabolteae sp. nov., but differs in its pointed protoconch; R. intrudens sp. nov. has a protoconch intruded by the teleoonch spiral cords; R. zanzibarensis (Kool & Dekker, 2007) has a smooth protoconch contrasting the axially-striated protoconch in R. annabolteae sp. nov.	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC72A783F4F2EFB88FD0627E0.taxon	biology_ecology	Habitat Intertidal to 70 m, frequently between 0 and 10 m. Some specimens were found living on rocky platforms with algae, associated to eunicid tubes, coarse sand, and rolling stones.	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC72A783F4F2EFB88FD0627E0.taxon	distribution	Distribution Only known from southern Madagascar (Fig 9).	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC72F78004F0EFD85FE6C2223.taxon	description	urn: lsid: zoobank. org: act: 5 C 34 E 90 F- 3 AD 2 - 42 ED-B 7 A 6 - 582525 AD 11 DB Fig. 8 E – H, 10; Table 2	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC72F78004F0EFD85FE6C2223.taxon	etymology	Etymology This species is named after the biblical Goliath, the giant who fought against David. This name is chosen because Reticunassa goliath sp. nov. is to date the largest known species in the genus Reticunassa.	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC72F78004F0EFD85FE6C2223.taxon	materials_examined	Type material Holotype NEW CALEDONIA: Nouméa, in sandy areas within coral reef, snorkeling at night, leg. D. Massemin (MNHN IM- 2000 - 22724, ex coll. H. Kool, length 15.4 mm, width 7.9 mm). Paratypes NEW CALEDONIA: PLOUVEAL, stn 1221, Lagon d’Ouvéa, 20 ° 29 ′ S, 166 ° 31 ′ E, 10 m (MNHN IM- 2000 - 22725, MNHN IM- 2000 - 22723, MNHN IM- 2000 - 28394, 3 spm; ZMA. Moll. 4.09.050, 1 spm; GH, 1 spm; HD 24143, 2 spm; HK 997.01, 3 spm; HUJ, 1 spm); PLOUVEAL, stn 1227, 12 ° 37 ′ S, 166 ° 25 ′ E, 12 m (HD 35304, 2 spm; HK 997.05, 2 spm); Nouméa, sand between coral reefs, snorkeled at night (HK 997.07, 1 spm); Nouméa, Nouville Beach, brown sand, 10 m (HK 997.08, 1 spm); New Caledonia, 5 – 10 m (HD 26939, 1 spm). Other material examined INDONESIA: West Papua, Manokwari, leg. D. Smits (HK 997.03, 2 spm). AUSTRALIA: North Queensland, Bowen, Grey’s Bay, ex coll. Hessel (ZMA, 4 spm; HK 997.06, 1 spm). VANUATA: SANTO 2006, stn FB 52, Malokilikili, 15 ° 42.7 ′ S, 167 ° 15.1 ′ E, 7 m (1 spm). NEW CALEDONIA: LAGON: stn 161, Ile Ouen-Baie du Prony, 22 ° 34 ′ S, 166 ° 38 ′ E, 20 m (1 spm); stn 943, Koumac, 20 ° 37 ′ S, 164 ° 11 ′ E, 15 m (1 spm). – PLOUVEAL, Lagon d’Ouvéa: stn 1218, 20 ° 36 ′ S, 166 ° 30 ′ E, 13 m (8 spm); stn 1219, 20 ° 30 ′ S, 166 ° 28 ′ E, 15 m (26 spm); stn 1220, 20 ° 29 ′ S, 166 ° 29 ′ E, 14 m (4 spm); stn 1221, 20 ° 29 ′ S, 166 ° 31 ′ E, 10 m (69 spm); stn 1222, 20 ° 28 ′ S, 166 ° 30 ′ E, 15 m (13 spm); stn 1223, 20 ° 28 ′ S, 166 ° 28 ′ E, 19 m (1 spm); stn 1224, 20 ° 32 ′ S, 166 ° 28 ′ E, 18 m (25 spm); stn 1225, 20 ° 36 ′ S, 166 ° 28 ′ E, 18 m (6 spm); stn 1226, 20 ° 32 ′ S, 166 ° 24 ′ E, 21 m (5 spm); stn 1227, 12 ° 37 ′ S, 166 ° 25 ′ E, 12 m (29 spm); stn 1228, 20 ° 36 ′ S, 166 ° 24 ′ E, 18 m (8 spm); stn 1229, 20 ° 37 ′ S, 166 ° 23 ′ E, 16 m (19 spm); stn 1230, 20 ° 35 ′ S, 166 ° 23 ′ E, 18 m (22 spm); stn 1231, 20 ° 31 ′ S, 166 ° 23 ′ E, 23 m, 6 spm); stn 1232, 20 ° 32 ′ S, 166 ° 24 ′ E, 31 m (7 spm); stn 1233, 20 ° 29.1 ′ S, 166 ° 29.0 ′ E, 15 m (1 spm). – MONTROUZIER 1993: Koumac, stn 1271, 20 ° 52.7 ′ S, 165 ° 19.5 ′ E, 5 – 25 m (3 spm); Koumac, stn 1286, Plateau Karembé, 20 ° 38 ′ – 20 ° 39 ′ S, 164 ° 16 ′ – 164 ° 17 ′ E (3 spm); Koumac, stn 1287, Récif de l’Infernet, 20 ° 37 ′ S, 164 ° 14 ′ E (2 spm); Koumac, stn 1301, Récif de l’Infernet, 20 ° 37.1 ′ – 20 ° 37.5 ′ S, 164 ° 14.7 ′ – 164 ° 15 ′ E, 1 – 5 m (3 spm); Koumac, stn 1303, Lagon, parages du Plateau Karembé, 20 ° 37.7 ′ – 20 ° 38.8 ′ S, 164 ° 15.9 ′ – 164 ° 17.1 ′ E, 0 – 8 m (2 spm); Koumac, stn 1304, Chenal de l’Infernet, 20 ° 38.6 ′ S, 164 ° 13.2 ′ E, 12 – 15 m (2 spm); Koumac, stn 1306, Chenal de l’Infernet, 20 ° 39.1 ′ S, 164 ° 12.4 ′ E, 11 – 13 m (1 spm); Koumac, stn 1312, 20 ° 40 ′ S, 164 ° 14.9 ′ E, 26 – 40 m (1 spm); Touho, stn 1245, Grand Récif Mengalia, 20 ° 45.2 ′ S, 165 ° 16.3 ′ E (4 spm); Touho, stn 1273, Touho region, outer reef, 20 ° 50.4 ′ S, 165 ° 22.8 ′ E, 20 m (2 spm). – PALEO-SURPRISE 1999, stn CP 1388, 18 ° 23.8 ′ S, 163 ° 06.9 ′ E, 40 m (2 spm). LOYALTY ISLANDS: LIFOU 2000, stn 1419, Lifou, Santal Bay, Bay of Gaatcha, 20 ° 55.6 ′ S, 167 ° 04.5 ′ E, 5 m (8 spm; HK 997.02, 1 spm); LIFOU 2000, stn 1426, Lifou, Santal Bay, 20 ° 45.9 ′ S, 167 ° 06.2 ′ E, 4 – 7 m (2 spm); Lifou, leg. J. R. le B. Tomlin (ZMA. Moll. 096228, ex coll. Schepman, 6 spm); Lifou, leg. R. P. Goubin (ZMA. Moll. 089599, ex coll. Dautzenberg, 3 spm); Lifou (HUJ 9055, ex coll. Dautzenberg, ex coll. Coen 7553, 3 spm).	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC72F78004F0EFD85FE6C2223.taxon	description	Description Holotype PROTOCONCH. Paucispiral, pointed, 1.75 milky whorls, beset with rows of microscopic pustules. Protoconch missing in holotype, seen in paratype (Fig. 8 H). SHELL. Rather heavy, conical, acuminate, teleoconch consisting of 7 whorls, suture impressed. On penultimate 13 and on body whorl 12 well pronounced, equidistant axial ribs and strong varix. Ribs equally strong on dorsal and ventral side of body whorl. SPIRAL CORDS. Continuous, moderately broad on top of axial ribs, but between ribs narrower and weak, approximately 6 cords on penultimate and 10 on body whorl. INTERCORDAL SCULPTURE. Numerous very fine, evenly spaced spiral striae, occasionally also on top of spiral cords. APERTURE. Round, proportionally small, approximately 1 / 5 of shell length. Outer lip with 7 unequal denticles. Columella callused, somewhat elevated anteriorly and with evenly spaced lirae throughout. Callus sharply bordered, bending over part of fasciole and somewhat extending over body whorl posteriorly. Inside outer lip with 8 lirae and pronounced tooth at siphonal canal. Elevated part of callus with fine growth lines on left (out) side. Parietal denticle strong, anal canal deep. OPERCULUM. Yellowish, serrated. SIPHONAL CANAL. Narrow, fasciole strong. Siphonal area with one strong and some weak cords. COLOR. Off white to yellow, narrow band just above suture and broader band below periphery; columella and outer lip white, aperture yellowish. ADULT SIZE. 8.2 – 16.5 mm, usually 13 – 15 mm.	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC72F78004F0EFD85FE6C2223.taxon	discussion	Remarks Color and banding pattern variable; some specimens have light to dark brown bands and others lack banding. In banded specimens, one band occurs on the body whorl, with an additional band on the sutural area. In some specimens, the spiral cords may be colored orange / brown, but only between the axial ribs. Aperture white, inside faintly showing the outside bands if present. Reticunassa visayaensis sp. nov. and R. poppeorum sp. nov. have a multispiral protoconch, whereas the protoconch of R. goliath sp. nov. is paucispiral. The strong columellar callus, together with the thickening of the inside of the outer lip and the strong varix, gives the aperture a round, “ open mouth ” appearance, whereas the other species in this group have a more oval-shaped aperture. R. goliath sp. nov. and R. tringa both have a paucispiral protoconch, but the former has a considerably heavier and larger shell. It is similar to R. tringa, R. visayaensis sp. nov. and R. poppeorum sp. nov. in having weak spiral cords. R. goliath sp. nov. differs from R. paupera in weight, in its larger size, in the number of whorls, and in the size of the aperture relative to the total shell length.	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC72F78004F0EFD85FE6C2223.taxon	biology_ecology	Habitat Sublittoral to 0 – 40 m, frequently between 10 and 20 m.	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC72F78004F0EFD85FE6C2223.taxon	distribution	Distribution Western Pacific; Indonesia (Papua), Australia (Queensland), Vanuatu, New Caledonia, and Loyalty Islands (Fig. 10).	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC71078024F1EFBCFFD292224.taxon	description	urn: lsid: zoobank. org: act: C 79 E 7864 - DE 80 - 4 A 17 - 94 BC- 54932 F 8532 CA Fig. 8 I – L, 11; Table 2	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC71078024F1EFBCFFD292224.taxon	etymology	Etymology The name of this species refers to the spiral cords that seemingly intrude and continue over part of the body whorl of the protoconch.	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC71078024F1EFBCFFD292224.taxon	materials_examined	Type material Holotype NEW CALEDONIA: MONTROUZIER 1993, stn 1260, Touho, Banc de Touho, 20 ° 44 ′ S, 165 ° 14 ′ E, 49 – 59 m, Sep. 1993 (MNHN IM- 2000 - 22726, length 7.9 mm, width 4.3 mm). Paratypes NEW CALEDONIA: same collection data as holotype (MNHN IM- 2000 - 22722, MNHN IM- 2000 - 28406, MNHN IM- 2000 - 28407, 3 spm; ZMA. Moll. 4.09.051, 1 spm; HD 35302, 3 spm; HK 196.01, 4 spm). Other material examined NEW CALEDONIA: LAGON: stn 146, Ile Ouen-Baie du Prony, 22 ° 24 ′ S, 166 ° 55 ′ E, 40 – 52 m (1 spm); stn 357, Grand Récif Sud, 22 ° 30 ′ S, 167 ° 07 ′ E, 77 m (1 spm); stn 603, Yaté, 22 ° 16 ′ S, 167 ° 05 ′ E, 78 – 80 m (1 spm). – MUSORSTOM 4, stn DW 203, 22 ° 36 ′ S, 167 ° 05 ′ E, 105 – 110 m (1 spm); SMIB 5, stn DW 81, S New Caledonia, 22 ° 38 ′ S, 167 ° 35 ′ E, 110 m (5 spm); BATHUS 1, stn DW 678, W coast, 20 ° 49 ′ S, 165 ° 19 ′ E, 94 – 100 m (7 spm). – MONTROUZIER 1993, Touho, stn 1237, Touho Bay, 20 ° 46.9 ′ S, 165 ° 13.8 ′ E, 0 – 1 m (5 spm); Touho, stn 1249, Passe de Touho, 20 ° 49 ′ S, 165 ° 19 ′ E, 80 – 140 m (3 spm); Touho, stn 1250, 20 ° 46.7 ′ S, 165 ° 13.7 ′ E, 3 – 6 m (7 spm); Touho, stn 1251, 20 ° 46.0 ′ – 20 ° 46.5 ′ S, 165 ° 13.0 ′ E, 15 m (4 spm); Touho, stn 1253, 20 ° 46.6 ′ S, 165 ° 14.1 ′ E, 2 – 3 m (1 spm); Touho, stn 1257, lagoon in front of Vieux-Touho, 20 ° 44 ′ S, 165 ° 11 ′ E, 25 – 28 m (3 spm); Touho, stn 1260, 20 ° 44 ′ S, 165 ° 14 ′ E, 49 – 59 m (70 spm); Touho, stn 1261, 20 ° 46 ′ – 20 ° 47 ′ S, 165 ° 15 ′ – 165 ° 16.5 ′ E, 45 – 56 m (33 spm); Touho, stn 1268, 20 ° 45.2 ′ S, 165 ° 08.0 ′ E, 9 – 11 m (1 spm); Touho, stn 1275, 20 ° 49 ′ S, 165 ° 17 ′ E, 50 – 62 m (1 spm).	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC71078024F1EFBCFFD292224.taxon	description	Description Holotype PROTOCONCH. Bulbous, flattened, paucispiral, 1.75 whorls, yellowish, axially striate; separation between protoconch and teleoconch indistinct, last half whorl appears to have been “ intruded ” by the 5 spiral cords on first whorl of teleoconch. SHELL. Teleoconch consisting of 5 strongly impressed whorls. Penultimate whorl and body whorl with 13 pronounced strong, round axial ribs, varix strong. SPIRAL CORDS. Penultimate whorl with 7 and body whorl with 10 narrow, fine, equally wide spiral cords. INTERCORDAL SCULPTURE. Minute spiral striae, occasionally also located on spiral cords. APERTURE. Outer lip with 5 short and low lirate denticles. Columella wrinkled, columellar callus margined, elevated anteriorly and somewhat curved towards fasciole, posteriorly slightly extending onto body whorl. Parietal denticle prominent, anal canal rather wide. OPERCULUM. Unknown. SIPHONAL CANAL. Siphonal area with 1 stronger and 4 weak cords. Fasciole strong. COLOR. Creamy-yellowish, some spiral cords darker. Whorls with light subsutural band, body whorl with 3 bands, darkest on the thick varix and at base of shell. ADULT SIZE. 5.4 – 9.7 mm, usually 7.5 – 8.5 mm.	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC71078024F1EFBCFFD292224.taxon	discussion	Remarks The most characteristic feature of this species is the broad, flattened, paucispiral and axially striated protoconch with, on the last part of it, 1 – 5 spiral cords that intrude from the teleoconch (Fig. 8 L). This character was also mentioned by Cernohorsky (1984) in his description of Nassarius (Hima) pauperus (Gould, 1850) and was treated as intraspecific variation. These features, the broad, flat paucispiral protoconch and the extremely fine intercordal spiral striae of Reticunassa intrudens sp. nov., make it easy to distinguish it from R. paupera and the other species described in this paper. Moreover, R. intrudens sp. nov. lacks the minute pustules present in some other species described herein and most of the others also have a wider distribution; R. intrudens sp. nov. is only known from New Caledonia. It is one of the smallest species within this group.	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC71078024F1EFBCFFD292224.taxon	biology_ecology	Habitat 0 – 110 m in muddy sand, with the largest concentration of specimens collected at 45 – 60 m.	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC71078024F1EFBCFFD292224.taxon	distribution	Distribution Only known from New Caledonia (Fig. 11).	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC71278044F37FBC1FD0C25E8.taxon	description	urn: lsid: zoobank. org: act: E 0731396 - 9 A 77 - 4 C 1 C- 99 F 7 - 24 B 72 E 5 B 55 DD Fig. 8 M – P, 12; Table 2	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC71278044F37FBC1FD0C25E8.taxon	etymology	Etymology This species is named after Thailand, the type locality.	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC71278044F37FBC1FD0C25E8.taxon	materials_examined	Type material Holotype THAILAND: SE coast, Gulf of Thailand, Pattani Province, Ko Lopi, 06 ° 40.313 ′ N, 101 ° 43.852 ′ E, scuba diving, 3 – 15 m, leg. R. G. Moolenbeek and H. Dekker, 2003 (ZMA. Moll. 4.09.047, length 7.35 mm, width 3.8 mm). Paratypes THAILAND: same collection data as for holotype (ZMA. Moll. 4.09.048, 222 spm; 5 spm; MNHN IM- 2000 - 32481, 10 spm; GH, 2 spm; HD 24145, 6 spm; HK 139.01, 6 spm); Gulf of Thailand, Rayong Province, Talengam, near Suan Son, E of Ban Phe beach, leg. O. K. McCausland (HK 139.03, 1 spm). Other material examined THAILAND: Gulf of Thailand: type locality (4 spm); Ban Pak Nam Sakom, local fishery, 06 ° 57 ′ N, 100 ° 49.36 ′ E (ZMA. Moll. 4.09.062, 3 spm; HD 1789, 6 spm); Ban Pak Nam Sakom, near Leela Resort, 06 ° 57.4 ′ N, 100 ° 51.6 ′ E, muddy pools (ZMA, 3 spm; HD 1190, 6 spm); Ban Talokapo, 06 ° 54.0 ′ N, 101 ° 22.4 ′ E, local fishery (HD 2146, 3 spm); Trat Province, Ko Chang, beach (HD 2793, 1 spm); Ko Samui, NE side of Chaweng Beach, 09 ° 32.6 ′ N, 100 ° 05.7 ′ E, in seagrass (HD 9572, 5 spm); Ko Samui, Ban Bangkao, 09 ° 25. 2 ′ N, 99 ° 59.7 ′ E, sand flat (HD 9574, 1 spm); Ko Kham, 06 ° 58.4 ′ N, 100 ° 51.6 ′ E, scuba diving, 3 – 5 m (ZMA. Moll. 093866, 14 spm); Pattani, in shell grit (ZMA. Moll. 098947, 2 spm); off Pattaya, Ko Lan, scuba diving, reef, 3 – 6 m, leg. R. G. Moolenbeek (ZMA. Moll. 4.09.063, 12 spm). SINGAPORE: Bedok River area of East Coast Park, beached, leg. Chan Sow Yan (HK 139.04, 6 spm). MALAYSIA: E coast, South China Sea, 02 ° 38 ′ 11 ″ N, 105 ° 13 ′ 30 ″ E, 30 – 35 m, leg. H. Morrison (GH, 2 spm; HK 139.02, 3 spm); South China Sea, on sand near wreck, 0 ° 45 ′ 15 ″ N, 105 ° 36 ′ 11 ″ E, leg. H. Morrison (GH, 1 spm). INDONESIA: Laboratorium Onderzoek der Zee, 1938, stn Bat 38 / 07 a, Bay of Jakarta (ZMA. Moll. 096217, 2 spm). – KEPULAUAN SERIBU 2005, stn Ser 05 / 03, Java, Bay of Jakarta, Damar Kecil, 5 ° 59 ′ 00 ″ S, 106 ° 50 ′ 43 ″ E, scuba diving, 5 – 15 m (ZMA. Moll. 175169, 7 spm); stn ser 05 / 07, Java, Bidadari, scuba diving, 6 ° 01 ′ 55 ″ S, 106 ° 44 ′ 47 ″ E (ZMA. Moll. 175470, 12 spm); stn ser 05 / 08, Java, Ayer Besar, 6 ° 00 ′ 06 ″ S, 106 ° 46 ′ 48 ″ E (ZMA. Moll. 154506, 1 spm); stn ser 05 / 09, Java, Ubi Besar, scuba diving, 5 ° 59 ′ 55 ″ S, 106 ° 44 ′ 23 ″ E (ZMA. Moll. 175172, 8 spm); stn ser 05 / 10, Java, Kelor, scuba diving, 5 ° 51 ′ 00 ″ S, 106 ° 35 ′ 05 ″ E, 5 – 35 m (ZMA. Moll. 175173, 3 spm). – Java, W coast, Tjilintjing (ZMA. Moll. 096215, ex coll. Butot 12405, 1 spm); Flores, E coast, near Larantuka, between Weri and Waibalun, leg. Fr. M. Vianney (ZMA. Moll. 096211, 1 spm); Kalimantan, South China Sea, Pejantan I., 0 ° 30 ′ 48.8 ″ N, 107 ° 13 ′ 24 ″ E, 10 – 20 m, leg. H. Morrison (GH, 5 spm); NETHERLANDS BERAU 2003, stn ber 03 / 38, Berau Is, Maratua, 10 – 25 m (ZMA. Moll. 175049, 1 spm). PHILIPPINES: Palawan, Balabac I. (RMNH, 5 spm; 5 spm; HD 24144, 20 spm; GH, 6 spm; HK 139.05, 140 spm; coll. L. van Gemert, Zeist, the Netherlands, 20 spm; AMD,> 300 spm); Olango I., Ziczac Point, 10 – 20 m, 2010 (HD 33413, 1 spm); Mindanao, Surigao del Norte, Basul I., 80 – 90 m, 2013 (HD 33414, 1 spm).	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC71278044F37FBC1FD0C25E8.taxon	description	Description Holotype PROTOCONCH. Bulbous, flattened, paucispiral, 1.3 whorls, whitish, part of body whorl with growth lines. SHELL. Teleoconch consisting of 4.5 impressed whorls. Axial ribs pronounced, rounded, 14 on penultimate and 13 on body whorl. Varix strong and broad. SPIRAL CORDS. Continuous, narrow, thread-like spiral cords, increasing in height where they cross over axial ribs; 6 on penultimate and approximately 9 on body whorl. Spiral cord on periphery, the two above it, and the three at the base of body whorl more distinct; intercordal distances unequal. INTERCORDAL SCULPTURE. Fine, distinct spiral striae, number depending on distance between spiral cords. APERTURE. Oval, outer lip with 8 unequal denticles, one near periphery strongest. Columellar callus well bordered, slightly extending over body whorl; entire lip with short lirae. Parietal denticle strong. OPERCULUM. Yellowish, finely serrated. SIPHONAL CANAL. Narrow; fasciole strong. Siphonal area with approximately 4 cords; parietal denticle strong, anal canal wide. COLOR. Yellowish-orange, penultimate whorl with two, body whorl with three orange-brown bands. ADULT SIZE. 5.5 – 9.1 mm, usually 7 – 8 mm. One of the smaller species of the group.	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC71278044F37FBC1FD0C25E8.taxon	discussion	Remarks The color is variable. Well-preserved specimens are cream or orange to light brown, either unicolor or with several darker-colored spiral cords. Occasionally remnants of bands are only present on the varix and on the fasciole. The main difference between Reticunassa intrudens sp. nov and R. thailandensis sp. nov. is to be found in the protoconch morphology: the former has a protoconch with axial striae, in the latter it is partly smooth. Both are paucispiral and bulbous, and lack the microscopic pustules found in the other species described in this paper. R. paupera (Gould, 1850), R. visayaensis sp. nov. and R. poppeorum sp. nov. have a multispiral protoconch. R. thailandensis sp. nov. has more well developed spiral cords than the other species described here. Furthermore, in some specimens the 2 – 3 most anterior spiral cords are more strongly developed. In general, the distribution of the species, as shown on Fig. 12, might help with the identification.	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC71278044F37FBC1FD0C25E8.taxon	biology_ecology	Habitat This species lives in sand and between dead corals, in depths of 1 – 90 m, usually found between 15 and 20 m.	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
C84487BFC71278044F37FBC1FD0C25E8.taxon	distribution	Distribution Thailand, Singapore, Malaysia, Indonesia, and the Philippines, where it is common at Balabac I., but apparently rare in the central Visayas (Fig. 12).	en	Galindo, Lee Ann, Kool, Hugo H., Dekker, Henk (2017): Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species. European Journal of Taxonomy 275: 1-43, DOI: 10.5852/ejt.2017.275
