taxonID	type	description	language	source
CF1687E1FFF6FB4F269A2D99FE08D3D7.taxon	description	Leeonychiurus sp. 1: Lee et al., 2014, Fig. 10.	en	Park, Kyung-Hwa (2017): Two new species of the genus Leeonychiurus Sun & Arbea, 2014 (Collembola: Onychiuridae) from Korea. Zootaxa 4329 (3): 292-300, DOI: 10.11646/zootaxa.4329.3.7
CF1687E1FFF6FB4F269A2D99FE08D3D7.taxon	diagnosis	Diagnosis. Dorsal pseudocellar formula 32 / 022 / 33243 (Fig. 1 A), ventral 11 / 000 / 0100; subcoxa 1 of legs I – III with one pseudocellus each. Postantennal organ with 13 – 15 compound vesicles. Abd. I – III terga with 3 + 3 chaetae along axial line, and without unpaired axial chaetae. Th. I – III sterna with 0 + 0, 1 + 1, 1 + 1 chaetae, respectively. Upper anal valve without chaetae a 1.	en	Park, Kyung-Hwa (2017): Two new species of the genus Leeonychiurus Sun & Arbea, 2014 (Collembola: Onychiuridae) from Korea. Zootaxa 4329 (3): 292-300, DOI: 10.11646/zootaxa.4329.3.7
CF1687E1FFF6FB4F269A2D99FE08D3D7.taxon	materials_examined	Type material. Holotype: male on slide, Korea, Jeoksangsan mountain (N 127 ° 41´46 ´´ E 35 ° 58´01 ´´), Jeoksang-myeon, Muju-gun, Jeollabuk-do, 8 – IV – 2016 in litter of a coniferous forest dominated by pine trees, collected by K. H. Park and I. A. Lee. Paratypes: 3 females and 3 males on slides, same data as holotype. 3 females and 2 males on slides, 22 – III – 2013, same locality as holotypes. 4 females on slides, Jogyeongdan (N 127 ° 08´01 ´´ E 35 ° 51´02 ´´), Deokjin-dong, Jeonju-si, Jeollabuk-do, 9 – VII – 2013, litter of mixed forest, collected by I. A. Lee and H. S. Kim.	en	Park, Kyung-Hwa (2017): Two new species of the genus Leeonychiurus Sun & Arbea, 2014 (Collembola: Onychiuridae) from Korea. Zootaxa 4329 (3): 292-300, DOI: 10.11646/zootaxa.4329.3.7
CF1687E1FFF6FB4F269A2D99FE08D3D7.taxon	description	Description. Body length (excluding antennae) of adults: males 0.85 – 0.96 mm, females 1.18 – 1.35 mm. Color white in alcohol. Cuticle granulation more or less uniform. Dorsal pseudocellar formula 32 / 022 / 33243 (Fig. 1 A), ventral 11 / 000 / 0100; subcoxa 1 of legs I – III with one pseudocellus each. Psx not visible. Psp formula: 00 / 011 / 11110 dorsally and 00 / 111 / 01 m 0 1 m 0 0 ventrally (Fig. 5 A). Head: Antennal bases well marked (Fig. 1 A). Antennae slightly shorter than head; ratio antennae / head = 0.8 – 0.9. Ant. IV subapical organite with globular apex; invaginated apical bulb absent; microsensillum in lateroexternal position approximately one-third length from the base; 7 – 8 sensilla, clearly distinguished from ordinary chaetae, present (Fig. 1 B). Ant. III sensory organ composed of five papillae, five guard chaetae, 2 rods and 2 granulated sensory clubs (Fig. 2 A); lateral ms just behind sensory organ. Ant. II with 16 chaetae. Ant. I with 9 chaetae. Postantennal organ with 13 – 15 compound vesicles arranged in two rows along axis of organ (Figs 2 C, 2 D). Head with unpaired dorsal chaeta d 0. 4 + 4 p-chaetae between two inner posterior pso on head, p 1 anterior to others. Mandible with strong molar plate and four apical teeth, maxilla bearing 3 teeth and 6 lamellae. Maxillary palp simple with 1 basal chaeta and 2 sublobal hairs. Labral formula as 4 / 342. Labial palp of type AC (Fig. 2 B: after Fjellberg, 1999), labial papillae A, B, C, D and E with 1,4,0,3,3 guard chaetae respectively (Fig. 2 B). Labium with 6 proximal, 4 basomedian and 6 basolateral chaetae. 4 + 4 postlabial chaetae present along ventral groove. Body chaetotaxy: s-chaeta distinct, their formulas as 2 / 012 / 222120 dorsally and 11 / 000 / 000100 ventrally. Tiny and blunt ms, present on Th. II and III. Th. I tergum with 6 – 8 + 6 – 8 chaetae. Th. II – Abd. III with 3 (4), 3 (4), 3,3,3 chaetae respectively on both side of axial line and without unpaired axial chaetae. Abd. IV tergum with axial chaeta p 0; Abd. V tergum with axial chaeta m 0; Abd. VI tergum with 2 unpaired axial chaetae: a 0, p 0 (Fig. 1 A). Th. I, II and III sterna with 0 + 0, 1 + 1 and 1 + 1 chaetae between legs, respectively. Subcoxae 1 of legs I – III with 3, 3, 3 chaetae respectively (Fig. 1 A), subcoxa 2 with 1, 4, 4 chaetae. Appendages: Tibiotarsi I, II and III with 22 (11, 8, 3), 21 (11, 8, 2) and 21 (11, 8, 2) chaetae respectively (Fig. 1 C). Unguis without teeth. Empodial appendage with basal lamella, appendage length shorter than inner edge of unguis (Fig. 1 C). Ventral tube with 6 + 6 distal chaetae, without anterior or basal chaetae. Furca reduced to a finely granulated area, with four small dental chaetae in one row posterior to furcal rudiment; three manubrial rows of chaetae present (Fig. 1 D). Anal valves chaetotaxy: each lateral valve with chaetae a 0, 2 a 1, b 0, 2 b 1 and 2 b 2; upper valve with chaetae a 0, b 0, 2 b 1, 2 b 2, c 0, 2 c 1, and 2 c 2 (Fig. 1 E). Anal spines set on distinct papillae, as long as inner edge of hind unguis.	en	Park, Kyung-Hwa (2017): Two new species of the genus Leeonychiurus Sun & Arbea, 2014 (Collembola: Onychiuridae) from Korea. Zootaxa 4329 (3): 292-300, DOI: 10.11646/zootaxa.4329.3.7
CF1687E1FFF6FB4F269A2D99FE08D3D7.taxon	biology_ecology	Ecology. Found in leaf litter of coniferous and mixed forest.	en	Park, Kyung-Hwa (2017): Two new species of the genus Leeonychiurus Sun & Arbea, 2014 (Collembola: Onychiuridae) from Korea. Zootaxa 4329 (3): 292-300, DOI: 10.11646/zootaxa.4329.3.7
CF1687E1FFF6FB4F269A2D99FE08D3D7.taxon	etymology	Etymology. The name “ koreanus ” derives from the Korean Peninsula, where the new species was collected. Korean name of the species: Hankook-lee-eori-tok-to-gi.	en	Park, Kyung-Hwa (2017): Two new species of the genus Leeonychiurus Sun & Arbea, 2014 (Collembola: Onychiuridae) from Korea. Zootaxa 4329 (3): 292-300, DOI: 10.11646/zootaxa.4329.3.7
CF1687E1FFF6FB4F269A2D99FE08D3D7.taxon	discussion	Discussion. The new species, L. koreanus sp. nov., can be easily distinguished from all other known species of the genus Leeonychiurus by the number of dorsal and ventral pseudocelli (Table 1 and Discussion in Leeonychiurus analis sp. nov.), first of all by the presence of two (rather than three) distinct pseudocelli on each side of Abd. III tergum.	en	Park, Kyung-Hwa (2017): Two new species of the genus Leeonychiurus Sun & Arbea, 2014 (Collembola: Onychiuridae) from Korea. Zootaxa 4329 (3): 292-300, DOI: 10.11646/zootaxa.4329.3.7
CF1687E1FFF4FB4B269A2C33FBD8D6CD.taxon	diagnosis	Diagnosis. Dorsal pseudocellar formula 32 / 12 (3) 2 / 33343, ventral 11 / 000 / 0100; subcoxa 1 of legs I – III with one pseudocellus each. Postantennal organ with 18 – 22 compound vesicles. Abd. I – III terga with 3 + 3 chaetae along axial line. Th. I – III sterna with 0 + 0, 1 + 1, 1 + 1 chaetae respectively. Chaeta c 0 on upper anal valve absent, but chaetae a 1 present.	en	Park, Kyung-Hwa (2017): Two new species of the genus Leeonychiurus Sun & Arbea, 2014 (Collembola: Onychiuridae) from Korea. Zootaxa 4329 (3): 292-300, DOI: 10.11646/zootaxa.4329.3.7
CF1687E1FFF4FB4B269A2C33FBD8D6CD.taxon	materials_examined	Type material. Holotype: female on slide, Korea, Sinwol-ri, Dongsang-myeon, Wanju-gun, Jeollabuk-do, (N 127 ° 17´23 ´´ E 35 ° 56´40 ´´), 27 – III – 2015 in roots of grass, collected by K. H. Park. Paratypes: 4 females and 1 male on slides, same data as holotype.	en	Park, Kyung-Hwa (2017): Two new species of the genus Leeonychiurus Sun & Arbea, 2014 (Collembola: Onychiuridae) from Korea. Zootaxa 4329 (3): 292-300, DOI: 10.11646/zootaxa.4329.3.7
CF1687E1FFF4FB4B269A2C33FBD8D6CD.taxon	description	Description. Body length (excluding antennae) of adults: male 1.65 mm, females 1.80 – 2.12 mm. Color white in alcohol. Cuticle granulation more or less uniform. Dorsal pseudocellar formula 32 / 12 (3) 2 / 33343 (Fig. 3 A), ventral 11 / 000 / 0100; subcoxa 1 of legs I – III with one pseudocellus each. Parapseudocellar formula ventrally 0 / 000 / 101001 m (Fig. 5 B), dorsal psx absent. Subcoxa 1 of legs I – III with 1 psx each. Psp formulas: 00 / 011 / 11110 dorsally and 00 / 111 / 0001 m 0 0 ventrally (Fig. 5 B). Head: Antennal bases well marked (Fig. 3 A). Antennae slightly shorter than head; ratio antenna / head = 0.83 – 0.88. Ant. IV subapical organite with globular apex; invaginated apical bulb absent; microsensillum in lateroexternal position approximately one-third length from the base; sensilla not well distinguish from ordinary chaetae (Fig. 3 D). Ant. III sensory organ composed of five papillae, five guard chaetae, 2 rods and 2 granulated sensory clubs (Fig. 3 C); lateral ms just behind sensory organ. Ant. II with 16 chaetae. Ant. I with 9 chaetae. Postantennal organ with 18 – 22 compound vesicles arranged in two rows along axis of organ (Fig. 4 A). Head with unpaired dorsal chaeta d 0. 4 + 4 p-chaetae between two inner posterior pso on head, p 1 anterior to others. Mandible with strong molar plate and four apical teeth, maxilla bearing 3 teeth and 6 lamellae. Maxillary palp simple with 1 basal chaeta and 2 sublobal hairs. Labral formula as 4 / 342 (Fig. 4 C). Labial palp of type AC (Fig. 4 B: After Fjellberg, 1999), labial papillae A, B, C, D and E with 1, 4, 0, 3, 3 guard chaetae respectively (Fig. 4 B). Labium with 6 proximal, 4 basomedian and 6 basolateral chaetae. 4 + 4 postlabial chaetae present along ventral groove (Fig. 3 B). Body chaetotaxy: s-chaeta formula as 2 / 012 / 222120 dorsally. Tiny and blunt ms, present on Th. II and III. Th. I tergum with 8 + 8 chaetae on each side. Th. II – Abd. III with 3, 3, 3, 3, 3 chaetae respectively on both sides of axial line and without unpaired axial chaetae, sometimes with asymmetrical chaetae. Abd. IV tergum with unpaired axial chaeta p 0, sometimes with m 0 chaetae; Abd. V tergum with axial chaeta m 0; Abd. VI tergum with 2 unpaired axial chaetae: a 0, p 0 (Fig. 3 A). Th. I, II and III sterna with 0 + 0, 1 + 1 and 1 + 1 chaetae between legs, respectively. Subcoxae 1 of legs I – III with 4, 5, 5 chaetae respectively, subcoxa 2 with 1, 4, 4 chaetae. Appendages: Tibiotarsi I, II and III with 22 (11, 8, 3), 21 (11, 8, 2) and 21 (11, 8, 2) chaetae, respectively. Claw without teeth. Empodial appendage with basal lamella, appendage length shorter than inner edge of unguis. Ventral tube with 8 – 9 + 8 – 9 distal chaetae, without anterior and basal chaetae. Furca reduced to a finely granulated area, with four small dental chaetae in one row posterior to furcal rudiment; three manubrial rows of chaetae present (Fig. 4 D). Anal valves chaetotaxy: each lateral valve with chaetae a 0, 2 a 1, b 0 and 2 b 1; upper valve with chaetae a 0, 2 a 1, b 0, 2 b 1, 2 b 2, 2 c 1 and 2 c 2 (Fig. 4 E). Anal spines set on distinct papillae, as long as inner edge of hind unguis.	en	Park, Kyung-Hwa (2017): Two new species of the genus Leeonychiurus Sun & Arbea, 2014 (Collembola: Onychiuridae) from Korea. Zootaxa 4329 (3): 292-300, DOI: 10.11646/zootaxa.4329.3.7
CF1687E1FFF4FB4B269A2C33FBD8D6CD.taxon	biology_ecology	Ecology. Found in roots of grass.	en	Park, Kyung-Hwa (2017): Two new species of the genus Leeonychiurus Sun & Arbea, 2014 (Collembola: Onychiuridae) from Korea. Zootaxa 4329 (3): 292-300, DOI: 10.11646/zootaxa.4329.3.7
CF1687E1FFF4FB4B269A2C33FBD8D6CD.taxon	etymology	Etymology. The name of the species refers to the anal lobe that has specific chaetotaxy. Korean name of the species: Dongsang-lee-eori-tok-to-gi. TABLE]. Major characters of all known species in the genus Leeonychiurus. North Distribution Korea Korea China China China Korea (cave) America References: 1) Sun & Zhang 2 O 12; 2) Sun & Arbea 2 O 14; 3) Lee & Park 1986; 4) Bernarđ 2 O 15	en	Park, Kyung-Hwa (2017): Two new species of the genus Leeonychiurus Sun & Arbea, 2014 (Collembola: Onychiuridae) from Korea. Zootaxa 4329 (3): 292-300, DOI: 10.11646/zootaxa.4329.3.7
CF1687E1FFF4FB4B269A2C33FBD8D6CD.taxon	discussion	Discussion. This new species is most similar to L. koreanus sp. nov. having identical number of ventral pseudocelli (11 / 000 / 0100) and the same set of unpaired axial chaetae on Abd. IV – VI. Nevertheless it can easily be distinguished due to the presence of dorsal pso on Th. I (absent in L. koreanus sp. nov.) and three dorsal pso on Abd. III (only two pso in L. koreanus sp. nov.), chaetotaxy of upper anal valve appears to be also rather specific (Table. 1). The new species is also close to L. gulinensis (Sun & Zhang, 2012) and L. antennalis (Sun & Zhang, 2012) as sharing same dorsal pseudocellar formula, but it can be distinguished from two latter species by the ventral pseudocellar formula on abdomen (0100 in L. analis sp. nov. vs. 0 101 in L. gulinensis & L. antennalis) and the absence of chaetae c 0 on upper anal valve (present in L. gulinensis & L. antennalis).	en	Park, Kyung-Hwa (2017): Two new species of the genus Leeonychiurus Sun & Arbea, 2014 (Collembola: Onychiuridae) from Korea. Zootaxa 4329 (3): 292-300, DOI: 10.11646/zootaxa.4329.3.7
