taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
3DE8B6EDF3D6561A840315D7FEE3D654.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461985	­10.3897/vz.75.e162650.figure6	Figure 6. Holotype of Ptyctolaemus gularis (ZMB 5004). A Lateral view; B dorsal view; C ventral view; D lateral view of head; E dorsal view of head; F ventral view of head. Scale bar = 10 mm.	Figure 6. Holotype of Ptyctolaemus gularis (ZMB 5004). A Lateral view; B dorsal view; C ventral view; D lateral view of head; E dorsal view of head; F ventral view of head. Scale bar = 10 mm.	2025-11-11	Balan, Amirtha;Das, Abhijit;Boruah, Bitupan;Tillack, Frank;Lalronunga, Samuel;Deepak, Veerappan		Zenodo	biologists	Balan, Amirtha;Das, Abhijit;Boruah, Bitupan;Tillack, Frank;Lalronunga, Samuel;Deepak, Veerappan			
3DE8B6EDF3D6561A840315D7FEE3D654.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461986	­10.3897/vz.75.e162650.figure7	Figure 7. Gular fold arrangement in males of the three species of Ptyctolaemus found in India. Ptyctolaemus gularis A WII - ADR 3477, B WII - ADR 3180, C WII - ADR 3482, D WII - ADR 1436; P. siangensis sp. nov., E WII - ADR 1210, F ZSI - R- 29479, G WII - ADR 1176, H WII - ADR 2542 and P. namdaphaensis sp. nov., I WII - ADR 3283, J WII - ADR 3178, K. WII - ADR 3061, L ZSI - R- 29481. Scale bar = 10 mm.	Figure 7. Gular fold arrangement in males of the three species of Ptyctolaemus found in India. Ptyctolaemus gularis A WII - ADR 3477, B WII - ADR 3180, C WII - ADR 3482, D WII - ADR 1436; P. siangensis sp. nov., E WII - ADR 1210, F ZSI - R- 29479, G WII - ADR 1176, H WII - ADR 2542 and P. namdaphaensis sp. nov., I WII - ADR 3283, J WII - ADR 3178, K. WII - ADR 3061, L ZSI - R- 29481. Scale bar = 10 mm.	2025-11-11	Balan, Amirtha;Das, Abhijit;Boruah, Bitupan;Tillack, Frank;Lalronunga, Samuel;Deepak, Veerappan		Zenodo	biologists	Balan, Amirtha;Das, Abhijit;Boruah, Bitupan;Tillack, Frank;Lalronunga, Samuel;Deepak, Veerappan			
3DE8B6EDF3D6561A840315D7FEE3D654.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461987	­10.3897/vz.75.e162650.figure8	Figure 8. Variation in the live gular colour pattern in P. gularis. A Uncollected female from Namdapha Tiger Reserve, Arunachal Pradesh; B a female (WII - ADR 3292) from Namdapha, Tiger Reserve, Arunachal Pradesh; C a female (WII - ADR 3473) from Kamlang Tiger Reserve, Arunachal Pradesh; D a female (WII - ADR 1152) from Dampa Tiger Reserve, Mizoram; E a male (WII - ADR 1436) from Daribokgre, Garo Hills, Meghalaya; F a male (WII - ADR 3180) from Shillong, Khasi Hills, Meghalaya.	Figure 8. Variation in the live gular colour pattern in P. gularis. A Uncollected female from Namdapha Tiger Reserve, Arunachal Pradesh; B a female (WII - ADR 3292) from Namdapha, Tiger Reserve, Arunachal Pradesh; C a female (WII - ADR 3473) from Kamlang Tiger Reserve, Arunachal Pradesh; D a female (WII - ADR 1152) from Dampa Tiger Reserve, Mizoram; E a male (WII - ADR 1436) from Daribokgre, Garo Hills, Meghalaya; F a male (WII - ADR 3180) from Shillong, Khasi Hills, Meghalaya.	2025-11-11	Balan, Amirtha;Das, Abhijit;Boruah, Bitupan;Tillack, Frank;Lalronunga, Samuel;Deepak, Veerappan		Zenodo	biologists	Balan, Amirtha;Das, Abhijit;Boruah, Bitupan;Tillack, Frank;Lalronunga, Samuel;Deepak, Veerappan			
3DE8B6EDF3D6561A840315D7FEE3D654.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461989	­10.3897/vz.75.e162650.figure10	Figure 10. Ptyctolaemus gularis in life. A WII - ADR 3180 from Shillong, Meghalaya; B WII - ADR 3473 from Kamlang Tiger Reserve, Arunachal Pradesh; C WII - ADR 1436 from Daribokgre, Meghalaya; D WII - ADR 1406 from Namdapha Tiger Reserve, Arunachal Pradesh; E WII - ADR 1152 from Dampa Tiger Reserve, Mizoram; F ZSI - R- 29477 from Narpuh Wildlife Sanctuary, Meghalaya.	Figure 10. Ptyctolaemus gularis in life. A WII - ADR 3180 from Shillong, Meghalaya; B WII - ADR 3473 from Kamlang Tiger Reserve, Arunachal Pradesh; C WII - ADR 1436 from Daribokgre, Meghalaya; D WII - ADR 1406 from Namdapha Tiger Reserve, Arunachal Pradesh; E WII - ADR 1152 from Dampa Tiger Reserve, Mizoram; F ZSI - R- 29477 from Narpuh Wildlife Sanctuary, Meghalaya.	2025-11-11	Balan, Amirtha;Das, Abhijit;Boruah, Bitupan;Tillack, Frank;Lalronunga, Samuel;Deepak, Veerappan		Zenodo	biologists	Balan, Amirtha;Das, Abhijit;Boruah, Bitupan;Tillack, Frank;Lalronunga, Samuel;Deepak, Veerappan			
3DE8B6EDF3D6561A840315D7FEE3D654.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461990	­10.3897/vz.75.e162650.figure11	Figure 11. Habitat of Ptyctolaemus species in northeast India. A, B Habitat of P. gularis in Shillong, Khasi Hills, Meghalaya; C habitat of P. siangensis sp. nov. at Potin, Arunachal Pradesh; D habitat of P. namdaphaensis sp. nov. at Kamala valley, Namdapha Tiger Reserve, Arunachal Pradesh.	Figure 11. Habitat of Ptyctolaemus species in northeast India. A, B Habitat of P. gularis in Shillong, Khasi Hills, Meghalaya; C habitat of P. siangensis sp. nov. at Potin, Arunachal Pradesh; D habitat of P. namdaphaensis sp. nov. at Kamala valley, Namdapha Tiger Reserve, Arunachal Pradesh.	2025-11-11	Balan, Amirtha;Das, Abhijit;Boruah, Bitupan;Tillack, Frank;Lalronunga, Samuel;Deepak, Veerappan		Zenodo	biologists	Balan, Amirtha;Das, Abhijit;Boruah, Bitupan;Tillack, Frank;Lalronunga, Samuel;Deepak, Veerappan			
3DE8B6EDF3D6561A840315D7FEE3D654.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461980	­10.3897/vz.75.e162650.figure1	Figure 1. Sampling localities of the current study and type localities of the Ptyctolaemus species. See Table 1 for sampling locality (1–31) details. Type localities are indicated by stars and other localities are indicated with a circle. Green circles: Ptyctolaemus gularis; purple star and circle: Ptyctolaemus siangensis sp. nov.; blue star and circle: Ptyctolaemus namdaphaensis sp. nov.; yellow star: Ptyctolaemus collicristatus; pink star: Ptyctolaemus chindwinensis.	Figure 1. Sampling localities of the current study and type localities of the Ptyctolaemus species. See Table 1 for sampling locality (1–31) details. Type localities are indicated by stars and other localities are indicated with a circle. Green circles: Ptyctolaemus gularis; purple star and circle: Ptyctolaemus siangensis sp. nov.; blue star and circle: Ptyctolaemus namdaphaensis sp. nov.; yellow star: Ptyctolaemus collicristatus; pink star: Ptyctolaemus chindwinensis.	2025-11-11	Balan, Amirtha;Das, Abhijit;Boruah, Bitupan;Tillack, Frank;Lalronunga, Samuel;Deepak, Veerappan		Zenodo	biologists	Balan, Amirtha;Das, Abhijit;Boruah, Bitupan;Tillack, Frank;Lalronunga, Samuel;Deepak, Veerappan			
1D82C091228E55DE822E62DEF5A1020F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461993	­10.3897/vz.75.e162650.figure14	Figure 14. Holotype of P. namdaphaensis sp. nov. (WII - ADR 3178). A Dorsal, B ventral and C lateral views of the body and D, E lateral, dorsal and ventral views of the head. Scale bar = 10 mm.	Figure 14. Holotype of P. namdaphaensis sp. nov. (WII - ADR 3178). A Dorsal, B ventral and C lateral views of the body and D, E lateral, dorsal and ventral views of the head. Scale bar = 10 mm.	2025-11-11	Balan, Amirtha;Das, Abhijit;Boruah, Bitupan;Tillack, Frank;Lalronunga, Samuel;Deepak, Veerappan		Zenodo	biologists	Balan, Amirtha;Das, Abhijit;Boruah, Bitupan;Tillack, Frank;Lalronunga, Samuel;Deepak, Veerappan			
1D82C091228E55DE822E62DEF5A1020F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461994	­10.3897/vz.75.e162650.figure15	Figure 15. Gular colouration of P. namdaphaensis sp. nov. A Male (WII - ADR 1409) from Namdapha; B male (WII - ADR 1427) from Namdapha; C male (WII - ADR 3061) from Namdapha; D holotype, male (WII - ADR 3178) from Gandhigram; E female (WII - ADR 1411) from Namdapha; F female (WII - ADR 1412) from Namdapha.	Figure 15. Gular colouration of P. namdaphaensis sp. nov. A Male (WII - ADR 1409) from Namdapha; B male (WII - ADR 1427) from Namdapha; C male (WII - ADR 3061) from Namdapha; D holotype, male (WII - ADR 3178) from Gandhigram; E female (WII - ADR 1411) from Namdapha; F female (WII - ADR 1412) from Namdapha.	2025-11-11	Balan, Amirtha;Das, Abhijit;Boruah, Bitupan;Tillack, Frank;Lalronunga, Samuel;Deepak, Veerappan		Zenodo	biologists	Balan, Amirtha;Das, Abhijit;Boruah, Bitupan;Tillack, Frank;Lalronunga, Samuel;Deepak, Veerappan			
1D82C091228E55DE822E62DEF5A1020F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461995	­10.3897/vz.75.e162650.figure16	Figure 16. Ptyctolaemus namdaphaensis sp. nov. in life condition. A, B Holotype (WII - ADR 3178); C, D male (WII - ADR 1409) and E – G female (WII - ADR 1410) from Nampdapha Tiger Reserve.	Figure 16. Ptyctolaemus namdaphaensis sp. nov. in life condition. A, B Holotype (WII - ADR 3178); C, D male (WII - ADR 1409) and E – G female (WII - ADR 1410) from Nampdapha Tiger Reserve.	2025-11-11	Balan, Amirtha;Das, Abhijit;Boruah, Bitupan;Tillack, Frank;Lalronunga, Samuel;Deepak, Veerappan		Zenodo	biologists	Balan, Amirtha;Das, Abhijit;Boruah, Bitupan;Tillack, Frank;Lalronunga, Samuel;Deepak, Veerappan			
1D82C091228E55DE822E62DEF5A1020F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461986	­10.3897/vz.75.e162650.figure7	Figure 7. Gular fold arrangement in males of the three species of Ptyctolaemus found in India. Ptyctolaemus gularis A WII - ADR 3477, B WII - ADR 3180, C WII - ADR 3482, D WII - ADR 1436; P. siangensis sp. nov., E WII - ADR 1210, F ZSI - R- 29479, G WII - ADR 1176, H WII - ADR 2542 and P. namdaphaensis sp. nov., I WII - ADR 3283, J WII - ADR 3178, K. WII - ADR 3061, L ZSI - R- 29481. Scale bar = 10 mm.	Figure 7. Gular fold arrangement in males of the three species of Ptyctolaemus found in India. Ptyctolaemus gularis A WII - ADR 3477, B WII - ADR 3180, C WII - ADR 3482, D WII - ADR 1436; P. siangensis sp. nov., E WII - ADR 1210, F ZSI - R- 29479, G WII - ADR 1176, H WII - ADR 2542 and P. namdaphaensis sp. nov., I WII - ADR 3283, J WII - ADR 3178, K. WII - ADR 3061, L ZSI - R- 29481. Scale bar = 10 mm.	2025-11-11	Balan, Amirtha;Das, Abhijit;Boruah, Bitupan;Tillack, Frank;Lalronunga, Samuel;Deepak, Veerappan		Zenodo	biologists	Balan, Amirtha;Das, Abhijit;Boruah, Bitupan;Tillack, Frank;Lalronunga, Samuel;Deepak, Veerappan			
1D82C091228E55DE822E62DEF5A1020F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461990	­10.3897/vz.75.e162650.figure11	Figure 11. Habitat of Ptyctolaemus species in northeast India. A, B Habitat of P. gularis in Shillong, Khasi Hills, Meghalaya; C habitat of P. siangensis sp. nov. at Potin, Arunachal Pradesh; D habitat of P. namdaphaensis sp. nov. at Kamala valley, Namdapha Tiger Reserve, Arunachal Pradesh.	Figure 11. Habitat of Ptyctolaemus species in northeast India. A, B Habitat of P. gularis in Shillong, Khasi Hills, Meghalaya; C habitat of P. siangensis sp. nov. at Potin, Arunachal Pradesh; D habitat of P. namdaphaensis sp. nov. at Kamala valley, Namdapha Tiger Reserve, Arunachal Pradesh.	2025-11-11	Balan, Amirtha;Das, Abhijit;Boruah, Bitupan;Tillack, Frank;Lalronunga, Samuel;Deepak, Veerappan		Zenodo	biologists	Balan, Amirtha;Das, Abhijit;Boruah, Bitupan;Tillack, Frank;Lalronunga, Samuel;Deepak, Veerappan			
BC376BD0C3D656DFB39F51F2E49D1064.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461991	­10.3897/vz.75.e162650.figure12	Figure 12. Holotype of P. siangensis sp. nov. (WII - ADR 1176) in preserved condition. A Dorsal, B ventral and C lateral views of the body and D – E lateral, dorsal and ventral views of the head. Scale bar = 10 mm.	Figure 12. Holotype of P. siangensis sp. nov. (WII - ADR 1176) in preserved condition. A Dorsal, B ventral and C lateral views of the body and D – E lateral, dorsal and ventral views of the head. Scale bar = 10 mm.	2025-11-11	Balan, Amirtha;Das, Abhijit;Boruah, Bitupan;Tillack, Frank;Lalronunga, Samuel;Deepak, Veerappan		Zenodo	biologists	Balan, Amirtha;Das, Abhijit;Boruah, Bitupan;Tillack, Frank;Lalronunga, Samuel;Deepak, Veerappan			
BC376BD0C3D656DFB39F51F2E49D1064.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461992	­10.3897/vz.75.e162650.figure13	Figure 13. Ptyctolaemus siangensis sp. nov. showing variation in gular and / or body colouration; A, B Male (ZSI - R- 29479) from Adi hills, Arunachal Pradesh; C, D female (WII - ADR 465) from Potin, Arunachal Pradesh; E, F an uncollected male from Nameri National Park, Assam.	Figure 13. Ptyctolaemus siangensis sp. nov. showing variation in gular and / or body colouration; A, B Male (ZSI - R- 29479) from Adi hills, Arunachal Pradesh; C, D female (WII - ADR 465) from Potin, Arunachal Pradesh; E, F an uncollected male from Nameri National Park, Assam.	2025-11-11	Balan, Amirtha;Das, Abhijit;Boruah, Bitupan;Tillack, Frank;Lalronunga, Samuel;Deepak, Veerappan		Zenodo	biologists	Balan, Amirtha;Das, Abhijit;Boruah, Bitupan;Tillack, Frank;Lalronunga, Samuel;Deepak, Veerappan			
BC376BD0C3D656DFB39F51F2E49D1064.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461986	­10.3897/vz.75.e162650.figure7	Figure 7. Gular fold arrangement in males of the three species of Ptyctolaemus found in India. Ptyctolaemus gularis A WII - ADR 3477, B WII - ADR 3180, C WII - ADR 3482, D WII - ADR 1436; P. siangensis sp. nov., E WII - ADR 1210, F ZSI - R- 29479, G WII - ADR 1176, H WII - ADR 2542 and P. namdaphaensis sp. nov., I WII - ADR 3283, J WII - ADR 3178, K. WII - ADR 3061, L ZSI - R- 29481. Scale bar = 10 mm.	Figure 7. Gular fold arrangement in males of the three species of Ptyctolaemus found in India. Ptyctolaemus gularis A WII - ADR 3477, B WII - ADR 3180, C WII - ADR 3482, D WII - ADR 1436; P. siangensis sp. nov., E WII - ADR 1210, F ZSI - R- 29479, G WII - ADR 1176, H WII - ADR 2542 and P. namdaphaensis sp. nov., I WII - ADR 3283, J WII - ADR 3178, K. WII - ADR 3061, L ZSI - R- 29481. Scale bar = 10 mm.	2025-11-11	Balan, Amirtha;Das, Abhijit;Boruah, Bitupan;Tillack, Frank;Lalronunga, Samuel;Deepak, Veerappan		Zenodo	biologists	Balan, Amirtha;Das, Abhijit;Boruah, Bitupan;Tillack, Frank;Lalronunga, Samuel;Deepak, Veerappan			
BC376BD0C3D656DFB39F51F2E49D1064.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1461990	­10.3897/vz.75.e162650.figure11	Figure 11. Habitat of Ptyctolaemus species in northeast India. A, B Habitat of P. gularis in Shillong, Khasi Hills, Meghalaya; C habitat of P. siangensis sp. nov. at Potin, Arunachal Pradesh; D habitat of P. namdaphaensis sp. nov. at Kamala valley, Namdapha Tiger Reserve, Arunachal Pradesh.	Figure 11. Habitat of Ptyctolaemus species in northeast India. A, B Habitat of P. gularis in Shillong, Khasi Hills, Meghalaya; C habitat of P. siangensis sp. nov. at Potin, Arunachal Pradesh; D habitat of P. namdaphaensis sp. nov. at Kamala valley, Namdapha Tiger Reserve, Arunachal Pradesh.	2025-11-11	Balan, Amirtha;Das, Abhijit;Boruah, Bitupan;Tillack, Frank;Lalronunga, Samuel;Deepak, Veerappan		Zenodo	biologists	Balan, Amirtha;Das, Abhijit;Boruah, Bitupan;Tillack, Frank;Lalronunga, Samuel;Deepak, Veerappan			
