taxonID	type	description	language	source
CE0987BF145F263EFCAAFDBA7231EB2F.taxon	type_taxon	Type Species. Ectospinula graduata Opitz, new species.	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF145F263EFCAAFDBA7231EB2F.taxon	diagnosis	Diagnosis. The gradual increase in the width of the distal antennomeres of the funicle (Fig. 3) will distinguish the members of this genus from others in Korynetinae.	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF145F263EFCAAFDBA7231EB2F.taxon	description	Description. Size: Length 8.0 mm; width 3.5 mm. Form: Oblong, hind body suboval, robust, deep body, about 2.5 times longer than broad. Vestiture: Disc of cranium and pronotum vested with long, dark setae, elytral disc vested with 22) P. astrictus. epipleural fold wide, ventrally positioned and expanded in proximal half, ends just before elytral apex, outer margin of elytral apex crenulated, anterior margin carinate; metathoracic wing as in Fig. 1, wedge cell closed; tibial spur formula 2 - 2 - 2, tarsal pulvillar formula 3 - 3 - 3, unguis with well-developed denticle. Abdomen: With 6 visible sternites, sternites 3 and 4 without setal patches.	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF145F263EFCAAFDBA7231EB2F.taxon	distribution	Distribution. This monotypic genus is known only from Zambia.	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF145F263EFCAAFDBA7231EB2F.taxon	etymology	Etymology. The generic name Ectospinula is a compound name derived from the Greek ekto (= without) and the Latin spinula (= dim). I refer to the slightly developed spine near the posterior angles of the pronotum. Gender: feminine. 1 ° setae and shorter, profusely distributed 2 ° setae. Head: Cranium quadrate, frons very wide, indented with large, setiferous punctation, latter nearly contiguous; gula small, rectangular, sutures vertical, gular process short, narrow, and forked; labrum short, medial incision deep, curvate concave; mandibular body short, stout; maxillary terminal palpomere subsecuriform; labial terminal palpomere digitiform (Fig. 5); eyes moderately sized, coarsely faceted, ocular notch large; antenna capitate (Fig. 3), capitulum lax, scape large, longer than combined length of pedicel and antennomeres 1 – 3, basal 3 funicular antennomeres oblong, distal 3 funicular antennomeres progressively increasing in diameter, capitular antennomeres not compacted, antennomeres 9 and 10 transverse, antennomere 11 subovoid. Thorax: Pronotum subovoid, convex, lateral margins minutely crenulated, with a small spine near pronotal hind angle, coarsely sculptured with round, setiferous punctation, prebasal fissure deep, prointercoxal process very expanded distally; pronotal projections long, but do not contact prointercoxal process; elytron sculptured with large, spheroid, asetiferous punctation, latter subseriate, 1 ° setae always adjacent to asetiferous punctation, 2 ° setae present and somewhat arranged serially, Ectospinula graduata Opitz, new species (Figs. 1, 3, 5, 19, 36, 37)	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF145F263EFCAAFDBA7231EB2F.taxon	materials_examined	Holotype. ♀. Mongu, Barotseld., 1952, J. M. Niemeyer (Zambia, Mongu, Barotseland) (TMSA).	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF145F263EFCAAFDBA7231EB2F.taxon	diagnosis	Diagnosis. The shape of the funicular antennomeres (Fig. 3) will distinguish the members of this species from any other in Korynetinae.	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF145F263EFCAAFDBA7231EB2F.taxon	description	Description. Size: Length 8.0 mm; width 3.5 mm. Form: As in Fig. 19. Color: Cranium and pronotum black with a shiny blue luster, antennae and legs black, elytra black with a cupreous luster, abdomen testaceous. Head: Antennal capitulum lax (Fig. 3), antennomeres 9 and 10 transverse and cupuliform, 11 subovoid; last labial palpomere digitiform (Fig. 5); eyes much narrower than frons (EW / FW 18 / 62). Thorax: Pronotum transverse (PW / PL 145 / 120), lateral margins serrulated; elytral punctation large to subseriate, interstitial spaces smooth and shiny; elytral apex subacuminate. Abdomen: Setal patches on 3 rd and 4 th sternites; female pygidium scutiform.	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF145F263EFCAAFDBA7231EB2F.taxon	distribution	Distribution. This species is known only from Zambia (Fig. 37).	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF145F263EFCAAFDBA7231EB2F.taxon	etymology	Etymology. The specific epithet graduata is a Latin adjective that stems from gradus (= step). The name describes the gradual increase in width of the distal funicular antennomeres.	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF1453263CFF6AFE277596EA9C.taxon	materials_examined	Holotype. ♂. Kenya, Eastern, E 729, 202 km E of Thika, 20.11.07, Snížek (ISNB). Paratypes. 105 specimens. KENYA: Garissa env., 30 xi 1999, M. Snížek (ISBN, 17; WOPC 16); Kenya Eastern E 729, Sosoma, 202 km E of Thika, 20.11.07, Snížek (ISNB, 14; WOPC, 13); idem, E of Thika, SW Kangonde, 28.12.2007, Lgt. Snížek (ISNB, 6; WOPC, 5); Coast Garissa, N of Bura, 5.12.2007, Lgt. Snížek (ISNB, 2; WOPC, 1); S. E. Tana riv. Prov. 14 km N of Garsen, 9. IV. 2004, Snížek leg. (ISNB, 1; RGCG, 1); SE Voi, S Foot Sagala, 12.12.2007, Mts., Lgt. Snížek (ISNB, 1); SE Taveta env., 16. IV. 2004, M. Snížek lgt (WOPC, 2; RGCG, 3); Machakos, 11.23, Dr. van Someron (BMNH. 1); Taita. Distr., surroundings of Voi, at light, 30. V. - 2. VI. 1994, legit. L. Bartolozzi, B. Cecchi, A. Sfozi (WOPC, 2). BURUNDI: 10 km W Bujumbura, 9.2.92, fc. Arndt (RGCG, 1). SOMALIA: El Wach Samarole ¾. XI. 1986, legit S. B. S. (MRAC, 4; WOPC, 1); El Buur Sar, 2. XI. 1986, S. B. S (MRAC, 2; WOPC, 2). ETHIOPIA: Harar Prov.: Errer, 12 m, 21 / 22. VIII. 1971 R. O. S. Clark (MRAC, 3; WOPC, 2); Shoa Prov., Awash Park, VII / VIII. 71, G. de Rougemont (MRAC, 2); Bubassa, VI- 1911 (WOPC, 1); Abyssinie, Raffray, 1882 (MNHN, 1). BURKINA FASO: J. Vulilet, V. 1909 (MNHN, 1).	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF1453263CFF6AFE277596EA9C.taxon	diagnosis	Diagnosis. The members of this species are superficially similar to those of P. livens, but P. adustus specimens differ by their dark brown coloration. 32) P. livens; 33) P. mulleri; 34) P. rudis; 35) P. testaceiventris.	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF1453263CFF6AFE277596EA9C.taxon	description	Description. Size: Length 6.0 mm; width 2.5 mm. Form: As in Fig. 20. Color: Cranium, prothorax, elytra, pterothorax dark brown, legs, antennae, abdomen testaceous. Head: Antennal capitulum lax, antennomere 9 cupuliform, antennomere 10 triangular; eyes much narrower than frons (EW / FW 27 / 55). Thorax: Pronotum transverse (PW / PL 130 / 95), spine on pronotal posterior angle well-developed; elytral disc sculptured with 12 striae of large punctation that extends to elytral apex, elytral interstitial spaces smooth and shiny. Abdomen: Setal patches on 3 rd and 4 th sternites; male pygidium scutiform; aedeagus as in Fig. 28. Natural History. Specimens were collected throughout the year, some at light and some at 12 m elevation.	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF1453263CFF6AFE277596EA9C.taxon	distribution	Distribution. This species is known from Burkina Faso, Burundi, Ethiopia, Kenya, and Somalia (Fig. 37).	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF1453263CFF6AFE277596EA9C.taxon	etymology	Etymology. The specific epithet adustus (= brown) is a Latin adjective. The name describes the coloration of these beetles.	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF1452263BFCA0FAB375C7EF23.taxon	materials_examined	Holotype. ♂. Mozambique, Sofala P, 30 km S Caia, 76 m, 18.02 S- 35.12 E. A second label reads: 21.12.2005, E-Y: 3709, light trap, leg. Gussmann, Müller (TMSA).	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF1452263BFCA0FAB375C7EF23.taxon	diagnosis	Diagnosis. The members of this species resemble those of P. mulleri Opitz, new species. But in specimens of P. arsus specimens, the elytral punctures are larger and the dorsum shows less of a metallic sheen.	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF1452263BFCA0FAB375C7EF23.taxon	description	Description. Size: Length 6.5 mm; width 2.8 mm. Form: As in Fig. 21. Color: Cranium, pronotum, and elytra red-brown, antennae, pterothorax, abdomen, and legs testaceous, elytra with a slight metallic sheen. Head: Antennal capitulum lax, antennomere 9 cupuliform, antennomere 10 triangular; eyes much narrower than frons (EW / FW 25 / 47). Thorax: Pronotum transverse (PW / PL 130 / 115), no spine on pronotal posterior angle; elytral disc sculptured with 12 striae of punctation that extends to elytral apex, elytral interstitial spaces smooth and shiny. Abdomen: Setal patches on 4 th sternite; male pygidium scutiform; aedeagus as in Fig. 29. Natural History. The holotype was collected in December, at 76 m elevation.	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF1452263BFCA0FAB375C7EF23.taxon	distribution	Distribution. This species is known only from Mozambique (Fig. 37).	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF1452263BFCA0FAB375C7EF23.taxon	etymology	Etymology. The specific epithet arsus (= shine) is a Latin adjective. The name describes the shine that emits from the elytral disc.	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF1455263AFD7CFE1C7490ED0D.taxon	materials_examined	Holotype. ♂. Kenya, Kibwezi, Tsavo Park, 1 - XI- 1981, 1300 M, leg: Rautenstrauch (FSCA).	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF1455263AFD7CFE1C7490ED0D.taxon	diagnosis	Diagnosis. The slender shape of the last labial palpomere (Fig. 5) will distinguish the members of this species from congeners.	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF1455263AFD7CFE1C7490ED0D.taxon	description	Description. Size: Length 7.5 mm; width 4.0 mm. Form: As in Fig. 22. Color: Cranium and pronotum dark castaneous, antennae, pterothorax, abdomen, and legs testaceous, elytra admixture of castaneous and black coloration, most elytral punctation black. Head: Antennal capitulum lax, antennomere 9 cupuliform, antennomere 10 triangular, antennomere 11 ovoid; eyes much narrower than frons (EW / FW 40 / 62). Thorax: Pronotum transverse (PW / PL 170 / 135); elytral disc sculptured with swellings and large, randomly distributed punctation (as in Fig. 15), elytral interstitial spaces variable in size, smooth and shiny; 1 ° setae very long, 2 ° setae often aggregated into small tufts. Abdomen: Setal patches on 3 rd and 4 th sternites; male pygidium scutiform; aedeagus as in Fig. 30. Natural History. The holotype was collected in November, at 1300 m elevation.	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF1455263AFD7CFE1C7490ED0D.taxon	distribution	Distribution. This species is known only from Kenya (Fig. 38).	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF1455263AFD7CFE1C7490ED0D.taxon	etymology	Etymology. The specific epithet astrictus (= narrow) is a Latin adjective. The name describes the slender form of the terminal labial palpomere.	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF1454263AFF72FC0172B4EBDA.taxon	materials_examined	Holotype. ♂. S. Afr,: Mpumalanga, Grobiersdal, 100 m, 2518 S- 2921 E. A second label reads: 10.3.2005; E-Y 3657, health trap, leg. J. Harrison (TMSA). Paratypes. 20 specimens. SOUTH AFRICA: Mpumalanga, Grobiersdal, Blaauwbank, 25 ° 15 ′ S- 29 ° 35 ′ E, 19 - 20.1.2004; E-Y: 3602, UV light, leg. TMSA staff (TMSA, 1); Transvaal, Pienaars Riv., 8 km S, 25 ° 17 ′ S- 28 ° 17 ′ E, 28.12.1994. E Y: 3084, beating, C. L. Bellamy (WOPC, 1); Kruger Nat. Pk, Skukuza, 3 km NW, 24 ° 58 ′ S- 31 ° 34 ′ E, 6.3.1996; E Y: 3213, pyrethrum fogging, leg. Endrödy-Younga (TMSA, 2; WOPC, 1); idem, 12 km S 25 ° 04 ′ S- 31 ° 37 ′ E, 6.3.1996; E Y: 3216, general beating, leg. Endrödy-Younga (TMSA, 1); Skukuza, 13 km SE, 25 ° 04 ′ S- 31 ° 37 ′ E, 7.3.1996; E Y: 3218, pyrethrum fogging, on Acacia nigrescens leg. Endrödy-Younga (WOPC, 1); Skukuza, fishingtr, 24 ° 59 ′ S- 31 ° 38 ′ E, 9.3.1996; E Y: 3223, pyrethrum fogging, on Diospyros misipiliformus, leg. Endrödy-Younga (WOPC, 1); Skukuza, Numbi Gate, 25 ° 06 ′ S- 31 ° 30 ′ E, 11.3.1996; E Y: 3231, pyrethrum fogging, on Acacia nigrescens leg. Endrödy-Younga (WOPC, 1); Skukuza, 5 km NE, 24 ° 59 ′ S- 31 ° 36 ′ E, 9.12.1997; E Y: 3301, misc. beating, leg. C. L. Bellamy (TMSA, 1). NAMIBIA: Khomas Region, road C 28, 75 km W Windhoek, 22 ° 45 ′ S 14 ° 23 ′ E, 1780 m, 9.11.2012, leg. M. Hornburg (RGCG, 1); Khomas Region, road C 28, near Bosua Pass, 22 ° 39 ′ S 16 ° 00 ′ E, 1300 m, 10.11.2012, M. Hornburg (RGCG, 2; WOPC, 1); E Otavi, 14. I. 1996, leg. K + F. Adlbauer (RGCG, 1). KENYA: Voi, Sagala Reg., 12.1991, leg. K. Werner (RGCG, 1); Meru Distr., Materi (Mitunguu), 800 m, 5 - 13.10.1988, leg. Mourglia (RGCG, 1); Tsavo NP, Kibwezi, 1300 m, 12.1989, leg. Werner (WOPC, 1). ETHIOPIA: Agere Maryam to Shakisso, 13.5.2002, leg. Werner & Sudre (RGCG, 1). TANZANIA: Ardai, 22 - VI- 1961 (BMNH, 1).	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF1454263AFF72FC0172B4EBDA.taxon	diagnosis	Diagnosis. The males can be distinguished from superficially similar specimens of P. rudis by characteristics of the male genitalia. In P. brevipenis, the aedeagus is shorter than the abdomen, whereas in P. rudis it is longer than the abdomen. The females of this species can only be credibly identified when they are associated with males from the same locality.	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF1454263AFF72FC0172B4EBDA.taxon	description	Description. Size: Length 8.0 mm; width 3.5 mm. Form: As in Fig. 23. Color: Cranium and pronotum dark castaneous, antennae, pterothorax, abdomen, and legs testaceous, elytra admixture of castaneous and black coloration. Head: Antennal capitulum lax, antennomeres 9 and 10 triangular, antennomere 11 ovoid; eyes much narrower than frons (EW / FW 30 / 80). Thorax: Pronotum transverse (PW / PL 170 / 140); elytral disc sculptured with swellings and large, randomly distributed punctation, elytral interstitial spaces variable in size, smooth and shiny; 1 ° setae very long, 2 ° setae often aggregated into small tufts. Abdomen: Setal patches on 3 rd and 4 th sternites; male pygidium scutiform; aedeagus as in Fig. 31. Natural History. Specimens were collected throughout the year at 800 – 1,780 m elevation. Collecting techniques involve UV light, beating, and pyrethrum fogging on A. nigrescens and D. mespiliformi.	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF1454263AFF72FC0172B4EBDA.taxon	distribution	Distribution. This species is known from Ethiopia, Kenya, Namibia, South Africa, and Tanzania (Fig. 38).	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF1454263AFF72FC0172B4EBDA.taxon	etymology	Etymology. The specific epithet brevipenis is a compound Latin name that stems from brevis (= short) and penis (= male copulatory organ). The name describes the short aedeagus of these beetles.	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF1450263DFC94FAE37716EFF3.taxon	type_taxon	Type Species. Prosymnus cribripennis Laporte, 1836: 52. By monotypy.	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF1450263DFC94FAE37716EFF3.taxon	diagnosis	Diagnosis. Beetles of this genus resemble those of the scarab genus Trox. They are deep-bodied beetles whose pronotum and elytral disc are sculptured with large, deep punctation. These characteristics will distinguish Prosymnus beetles from others in Korynetinae.	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF1450263DFC94FAE37716EFF3.taxon	description	Description. This genus was recently described and illustrated by Opitz (2011).	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF1450263DFC94FAE37716EFF3.taxon	distribution	Distribution. Throughout central and southern Africa.	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF14572627FD42FE877740EC52.taxon	materials_examined	Holotype. ♂. S. Afr /: N. Province, Amatola, Scott farm, 22. 56 S 29.23 E. Another label reads: 26.11.1998, E. Y. 3316, at light, leg. R. Müller (TMSA). Paratypes. 2 specimens. South Africa: N. Province, Amatola, Scott farm, 22 ° 56 ′ S- 29 ° 23 ′ E, 26.1.1998, at light, leg. R. Müller (WOPC); E. Transvaal, Penge, 13 - 17.11.1972, A. Strydom (TMSA).	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF14572627FD42FE877740EC52.taxon	diagnosis	Diagnosis. The members of this species resemble those of C. arsus. However, in P. mulleri the elytral punctation is smaller, and the dorsum shows a more prominent metallic sheen.	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF14572627FD42FE877740EC52.taxon	description	Description. Size: Length 7.0 mm; width 2.8 mm. Form: As in Fig. 25. Color: Cranium, pronotum, and elytra violaceous, antennae, pterothorax, abdomen, and legs testaceous, elytra with a prominent metallic sheen. Head: Antennal capitulum lax, antennomere 9 cupuliform, antennomere 10 triangular, antennomere 11 oval; eyes much narrower than frons (EW / FW 30 / 45). Thorax: Pronotum transverse (PW / PL 135 / 111), spine on pronotal posterior angle absent; elytral disc sculptured with 12 striae of large punctation that extends to elytral apex, elytral interstitial spaces smooth and shiny. Abdomen: Setal patches on 4 th sternite (Fig. 8); male pygidium scutiform; aedeagus as in Fig. 33. Variation. Size: Length 6.0 – 7.0 mm; width 2.3 – 2.8 mm. Except for size, the beetles before me are quite homogeneous. Natural History. The available specimens were collected in November or January, two at light.	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF14572627FD42FE877740EC52.taxon	distribution	Distribution. This species is known only from South Africa (Fig. 39).	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF14572627FD42FE877740EC52.taxon	etymology	Etymology. From the surname Müller, in honor of the collector of the holotype, Ruth Müller. Prosymnus rudis (Boheman, 1851) (Figs. 4, 6, 7, 9 – 18, 26, 34, 36, 40)	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF14572627FD42FE877740EC52.taxon	diagnosis	Diagnosis. The males can be distinguished from superficially similar specimens of P. brevipenis by characteristics of the male genitalia. In P. rudis, the aedeagus is longer than the abdomen, whereas in P. brevipenis it is shorter than the abdomen. The females of this species can only be credibly identified when they are associated with males from the same locality.	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF14572627FD42FE877740EC52.taxon	description	Redescription. Size: Length 8.0 mm; width 3.5 mm. Form: As in Fig. 26. Color: Cranium and pronotum dark castaneous, antenna, pterothorax, abdomen, and legs testaceous, elytra admixture of castaneous and black coloration. Head: Antennal capitulum lax (Fig. 4), antennomeres 9 and 10 triangular, antennomere 11 ovoid; eyes (Fig. 9) much narrower than frons (EW / FW 27 / 80); ventrally as in Figs. 10 – 11. Thorax: Pronotum transverse (Figs. 12 – 14) (PW / PL 170 / 135); elytral disc sculptured with swellings and large, randomly distributed punctation (Fig. 15 – 18), elytral interstitial spaces variable in size, smooth and shiny; 1 ° setae very long, 2 ° setae often aggregated into small tufts. Abdomen: Setal patches on 3 rd and 4 th sternites; male pygidium scutiform; aedeagus as in Fig 34. Natural History. Specimens have been collected throughout the year, at altitudes ranging from 18 to 1,500 m. Among the collecting techniques used were beating sheet, light trap, Lindgren funnel, and mercury vapor light. Some specimens were collected on the flowers of the angiosperm Protea. R. Murphy collected one specimen at 550 m, in woods laden with trees of the angiosperm Brachystegia.	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
CE0987BF14572627FD42FE877740EC52.taxon	distribution	Distribution. In addition to the type, I examined 125 specimens: SOUTH AFRICA: Victoria Falls, 6 - 13. XI. 1938, D. L. Uyttenboogaart; Waterberg, Geelhoutbush farm, 24 ° 22 ′ S 27 ′ 33 ″ E, 19.12.1995, misc. beating camp, leg. C. L. Bellamy; Northwestern Prov., Soutpan, Tswaing, 25 ° 24 ′ S 28 ° 06 ′ E, 12.12.1996, beating, Dombrowsky & Krüger; idem, Sitkaatsneck, 25 ° 40 ′ S 27 ° 55 ′ E, 30.11.1995, beating, leg. C. L. Bellamy; idem, Kommanddoneck, 25 ° 45 ′ S 27 ° 47 ′ E, 1.12.1995, beating, leg. C. L. Bellamy; idem, Warmbaths, Mabalingwe Res., 20.3.1996, Light trap, leg. B. Dombrowsky; idem, 20 km W Thabazimbi, 24 ° 35 ′ S 27 ° 14 ′ E, 12. XII- 5. II. 2000, Lindgren Funnel, T. K. Phillips; Transvaal, Waterval farm, 25 ° 35 ′ S 28 ° 16 ′ E, 8.1.1992, at light, leg. R. Dixon; idem, Pienaars Riv., 8 km S, 25 ° 17 ′ S 28 ° 17 ′ E, 28.12.1994, beating, leg. C. L. Bellamy; idem, d’ Nyala Natur Res. IV- 2 - 1992, UV light, C. W. & L. B. O’ Brien; idem, Saartjiesnek, 30 km W Pretoria, I. 1989, leg. K. Werner; idem, Rustenburg, 26 - 30. XI. 1948, leg. A. L. Capener; idem, Middlefontein near Nylstroom, 15 - 17.12.1953, leg. A. L. Capener; idem, Nmabolela estate, 22 ° 40 ′ S 28 ° 15 ′ E 6.3.1973, mercury vapor light, leg. Endrödy- Younga; idem, 30 km NE Thabazimbi, 24, 25. X. 1984, H. & A. Howden; idem, Ben Alberta Nat. Res., Thabazimbi, 24 ° 37 ′ S 27 ° 23 ′ E, 24 - 28. XI. 1980, S. J. van Tonder; idem, Potgietersrus, 1 - 3. XII. 1992, Ex. Protea flowers, J. Klimaszewski; Gauteng, Hennops River District, Portion 53 of Vlakaas, 24. XI. 2008, leg., Martin Paulsen; Makueni, Ukamba, 11.1947, von Somoren. SENEGAL: M’ Bambey, 8 - VIII- 1939, M. Risbec; Casamanche, 6 - VI- 1981, leg. B. Sigwalt; Thies, 1882; Ziguinchor, Boukitimbo, 12 ° 26 ′ 41 ″ N 16 ° 35 ′ 53 ″ E, 5. VIII. 2007, light trap, 18 m, P. Moretto & F. Génier. BURKINA FASO: Comoé, Koflandé (village), 10 ° 14 ′ 42 ″ N 4 ° 27 ′ 50 ″ E, 4. VII. 2006, 290 m, savanna wood, light trap, F. & S. Génier; Nahouri, Forêt de Nazinga, 11 ° 07 ′ 52 ″ N 1 ° 34 ′ 38 ″ E, 24. VII. 2006, 270 m, savanna wood, light trap, F. & S. Génier; Loroum, Toulfé, 13 ° 53 ′ 43 ″ N 1 ° 52 ′ 25 ″ E, 16. VII. 2006, 300 m, light trap, F. & S. Génier; Bobo Dioulasso, XII- 1974. GAMBIA: Abuko Nat. Res. 13.10.1999, leg. W. Schacht; Janjangbureh, 12 - 13.7.2000, R. Grimm. DEMOCRATIC REPUBLIC OF THE CONGO: Lualaba, Zilo, 15 - XI- 1960, Dr. V. Allard; idem, Ruwe, I- 1960, at light, Dr. V. Allard; Elisabethville, 8 - XII- 1930, H. J. Bredo; Baudouinville, I- 1933, L. Burgeon; Kapanga, XII- 1932, F. G. Overlaet. KENYA: Eastern, E 729 Sosoma, 202 km E of Thika, 20.11.07, Snižek; idem, E of Thika, Mwingi, Nguini, 29. XII. 2007, Legt. Snižek. ETHIOPIA: Baffray, 1882. SUDAN: Koulikoro, 1913, Paul Linarix. CHAD: N’ Gouri, distr. De Kanem, P. Renaud. TANZANIA: Longido, Masai Distr. 17 / 20 - IV- 1957, 1500 m, P. Basilewsky & N. Leleup; Ngarenanyuki. BENIN: Dahomey. NIGER: Maradi, 11 - VII- 1980, Paul Bouchard. RWANDA: P. N. Akagera, 6. XII. 1985, R. Jocqué. ZIMBABWE: Sebakwe; Mazoe, Dec. 1905, G. A. K. Marshall; Salisbury, Dec. 1897, G. A. K. Marshall; Masvingo Pro. Lake Mutirikwe, 20 ° 13 ′ S- 31 ° 00 ′ E, 27.11.2006, light trap, leg Gussmann & Müller; Chivhu, The Range env., 30. XI. 1998, leg. M. Snižek; Matopos, 8 - XII- 1990, R. Miller & L. Strange. CÔTE D ’ IVORE: Elfenbeinküste, Comoé National Park, V. 1993, lg. Rödel. MOZAMBIQUE: Nyassa. ZAMBIA: Kashitu, N of Broken Hill, 26. IV. 1915, H. C. Dollman; Lochinvar, 6 / 26. V. 1964, F. Van Noten; Kashitu, 26 - IV- 1915, Mopan. MALAWI: Salima Dist., Senga Bay Forest Res., 13 - 16 - II- 2009, 550 m, in Brachystegia woods, R. Murphy. NIGERIA: Lama Burra G. R. 11 ° 04 ′ 57 ″ N 8 ° 37 ′ 26 ″ E, 21 - 25 - VII- 2006, 830 m, Leonard & Vingerhoedt (Fig. 40). Specimens are deposited in BMNH, CMNC, FSCA, ISNB, MNHN, MRAC, NBCN, RGCG, TMSA, UCDC, and WOPC.	en	Opitz, Weston (2016): Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte. The Coleopterists Bulletin 70 (3): 429-446, DOI: 10.1649/0010-065X-70.3.429, URL: http://dx.doi.org/10.1649/0010-065x-70.3.429
