identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
12A30B7D8C72F78846DDBF255657332D.text	12A30B7D8C72F78846DDBF255657332D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oahutanais Morales-Núñez & Larsen & Cooke 2016	<div><p>Oahutanais gen. n.</p><p>Generic diagnosis.</p><p>Female: Small, 0.8-0.9 mm, slender body, parallel-sided. Carapace extending laterally to cover (dorsally) the cheliped attachments. Carapace not connected with the cheliped sclerites Pereonites wider than long. Antennule with four articles and minute terminal segment, shorter than carapace, article-2 with dorsal symmetric projection overlapping basal part of article-3, terminal segment minute and covered by article-4 (only visible with scanning electron microscope image). Antenna with six articles. Labium without distolateral spines. Maxillule with seven distal spiniform setae (two bifid); maxillipedal palp article-2 with geniculate, finely-pectinate spiniform seta on distal inner margin (serrations visible at magnification 100 ×). Cheliped attached via sclerite just anterior to the posterior margin of the cephalothorax, very close to the midventral line. Pereopods 1 to 6 attached ventrally. Pereopods 1-3 relatively slender; ischial seta shorter than merus. Pereopods 4-6 not stouter than pereopods 1-3; ischial setae shorter than merus. Pleopods absent in females. Uropods longer than pleotelson; basal article shorter than pleotelson, without distal apophyses; exopod uni-articulated, slightly longer than endopod article-1.</p><p>Male unknown.</p><p>Type species.</p><p>Oahutanais makalii sp. n.</p><p>Etymology.</p><p>Named after “O’ahu Island", where the material used in this study was collected, plus suffix - tanais.</p><p>Gender.</p><p>Masculine.</p><p>Distribution.</p><p>Hawaii Islands at depths ranging from 19 to 102 meters.</p><p>Remarks.</p><p>The family Colletteidae has been considered as polyphyletic (Larsen and Wilson 2002; Błażewicz-Paszkowycz and Poore 2008; Błażewicz-Paszkowycz et al. 2013) and it is currently composed of 15 genera (WoRMS); unfortunately, most of the genera included within this diverse family need revision (e.g. Leptognathiella Hansen, 1913 and Filitanais Kudinova-Pasternak, 1973). The females of Oahutanais gen. n. can be identified by having a small body (less than 1 mm in length of reproductively active specimens), cheliped attached just anterior to the posterior margin of the cephalothorax, very close to the midventral line, not in contact with carapace lateral margin, and pereopods 1 to 6 attached ventrally. Oahutanais gen. n. appears to be most closely related to the genera, Leptognathiella Bird and Holdich, 1984 from the Atlantic or Gulf of México (Larsen 2005; Larsen et al. 2006), Leptognathiopsis Holdich &amp; Bird, 1986 from the North Atlantic (Holdich and Bird 1986), and several species of Leptognathia G.O. Sars, 1882 sensu stricto (see Larsen and Shimomura 2007: 12) in having pereonites wider than long, pointed molars, females without pleopods (only in some species of Leptognathiella and Leptognathia), and uropod structure. However, the Oahutanais can be distinguished from the Atlantic species of Leptognathiopsis and Leptognathiella by having 1) the maxilliped palp article-2 with geniculate, finely pectinate spiniform seta on sub-distal margin, 2) pereopods 1-3 with basis slender, and 3) pereopod ischial setae shorter than merus.</p><p>The new genus also shows similarities with the monotypic genus Nippognathiopsis Błażewicz-Paszkowycz, Bamber &amp; Jóźwiak, 2013, recently described from 517-1356 m in waters off Japan. However, Oahutanais can be differentiated by 1) its pereonites are wider than long, 2) the antennal article-1 is longer than the distal three articles and minute terminal segment combined (shorter in Nippognathiopsis), 3) the maxilliped endites have a medial small process (two oval tubercles in Nippognathiopsis), 4) the maxilliped palp article-2 has a geniculate, finely pectinate spiniform seta on the sub-distal margin, and 5) pereopods 1-6 are slender. Although the presence of the geniculate, finely pectinate spiniform seta in the maxilliped palp article-2 separates Oahutanais from the other genera within the family Colletteidae, it is possible that this spiniform seta has been overlooked in the original description of Nippognathiopsis, as well as in other colletteids due to their small overall size. Unfortunately, no information is available on the form of the cheliped attachment in Nippognathiopsis .</p><p>Bird and Larsen (2009) mentioned that this character, the cheliped-cephalothorax attachment position, is an important feature that has been overlooked or has not been recorded or illustrated by many authors, even today. So far within the family Colletteidae, only Błażewicz-Paszkowycz and Bamber (2012) and this study have included detailed information and illustrations showing the real point of insertion of the cheliped and how far it is located from pereonite-1. Thus, Oahutanais can be separated from Bascestus Błażewicz-Paszkowycz &amp; Bamber, 2012 by having 1) the cheliped attached just anterior to the posterior margin of the cephalothorax (more anterior in Bascestus), 2) females without pleopods (pleopods present in Bascestus), and 3) exopod uni-articulated (bi-articulated in Bascestus).</p><p>The ventral cheliped attachment without contact with the carapace lateral margin (Fig. 8B-C) is an interesting and rare character among tanaidomorphans, and it has so far only been recorded from another colletteid, Isopodidus Larsen &amp; Heard, 2002, but this highly modified genus differs in many other aspects from Oahutanais .</p></div>	https://treatment.plazi.org/id/12A30B7D8C72F78846DDBF255657332D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Morales-Nunez, Andres G.;Larsen, Kim;Cooke, William J.	Morales-Nunez, Andres G., Larsen, Kim, Cooke, William J. (2016): Oahutanais makalii, a new genus and species of colletteid tanaidacean (Crustacea, Peracarida) from shelf-waters off Hawaii, with a taxonomic key. Zoosystematics and Evolution 92 (1): 1-12, DOI: http://dx.doi.org/10.3897/zse.92.5581, URL: http://dx.doi.org/10.3897/zse.92.5581
A657F81AB46BE6C8ACF0D185F5EF53BE.text	A657F81AB46BE6C8ACF0D185F5EF53BE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oahutanais makalii Morales-Núñez & Larsen & Cooke 2016	<div><p>Oahutanais makalii sp. n. Figures 2, 3, 4, 5, 6, 7, 8</p><p>Type material.</p><p>Holotype Adult ♀, length 0.9 mm, (USMN 1283305), BPOBS Station (Stn) HB4-R1 (21° 16 ’47.7’’ N - 158° 01 ’38.1’’ W), depth 62 m, substrata: "predominantly fine and medium sand", coll. by City and County of Honolulu Oceanographic Team, March 2013.</p><p>Paratypes. Four ♀♀ (USMN 1283306); four ♀♀ (GCRL 06534); and six ♀♀ (BPBM 2015.097; four on SEM stubs and two in alcohol), SIOBS Stn D3A (21° 16 ’55.3’’ N - 157° 53 ’49.9’’ W), depth 49 m, substrata "primarily coarse sediment including shell fragments", coll. by City and County of Honolulu Oceanographic Team, October 2014. Additional specimens from the type locality are in the collection of the authors.</p><p>Other material examined.</p><p>Thirteen ♀♀, one ovigerous ♀, two ♀♀ with remains of marsupium, SIOBS Stn D 3A (21° 16 ’55.3’ N - 157° 53 ’49.9’ W), depth 49 m, substrata "primarily coarse sediment including shell fragments", coll. by City and County of Honolulu Oceanographic Team, October 2014 ; 27 specimens (spec), MOBS Stn D (21° 25' 32.3"N - 157° 42' 53.6"W), depth 30 m, October 2013; two spec, SIOBS Stn C1A (21° 17' 38.3"N - 157° 55' 28.3"W), depth 19.2 m, October 2014; one spec, SIOBS Stn C5A (21° 16' 53.9"N - 157° 51' 25.4"W), depth 20.1 m, October 2014; two spec, SIOBS Stn D1 (21° 17' 23.2"N - 157° 55' 29.8"W), depth 49.1 m, October 2014; three spec, SIOBS Stn D2 (21° 16' 55.2"N - 157° 54' 36.3"W), depth 56.4 m, October 2014; 40 spec, SIOBS Stn D3A (21° 16' 55.9"N - 157° 53' 48.8"W), depth 50 m, October 2014; 16 spec, SIOBS Stn D5 (21° 16' 36.8"N - 157° 51' 33.9"W) depth 52.1 m, October 2014; eight spec, SIOBS - Stn D6 (21° 16' 02.5"N - 157° 50' 46.4"W) depth 50.0 m, October 2014; two spec, SIOBS Stn E1 (21° 17' 09.5"N - 157° 55' 32.2"W), depth 102.4 m, October 2014; 49 spec, SIOBS Stn E3 (21° 16' 42.9"N - 157° 53' 49.5"W), depth 84.4 m, October 2014; three spec, SIOBS Stn E5 (21° 16' 22.5"N - 157° 51' 40.3"W) 101.5 m, October 2014; four spec, SIOBS Stn E6 (21° 15' 51.1"N - 157° 50' 57.2"W) depth 102.4 m, October 2014; 7 spec, WOBS Stn Z (21° 25' 38.8"N - 158° 11' 48.1"W) depth 29.3 m, October 2014.</p><p>Diagnosis.</p><p>As the generic diagnosis above.</p><p>Etymology.</p><p>The species name, makali’i, is Hawaiian for tiny or minute and reflects both where the material used in this study was collected and its small size relative to other tanaidaceans within this benthic community.</p><p>Type locality.</p><p>Off Barbers Point Oahu, BPOBS study area (21° 16 ’47.7’’ N - 158° 01 ’38.1’’ W), Hawaii, May 2013.</p><p>Description.</p><p>Based in holotype female, length 0.9 mm (USMN 1283305).).</p><p>Body (Fig. 2) length about 0.9 mm, about 8.6 times width.</p><p>Cephalothorax (Figs 2, 6A-B) about 15% of TL, slightly longer than first two pereonites combined, about 1.5 times longer than wide, oval shape (Fig. 6B); with distolateral seta. Eye-lobes absent.</p><p>Pereon (Fig. 2): about 60% of TL; pereonites 1-3 and 6 sub-rectangular, wider than long; pereonites 4-5 quadrate.</p><p>Pleon (Figs 2, 7A): about 20% of TL; combined length of pleonites 1 to 5 slightly shorter to that of pereonites 5 and 6 combined; all pleonites subequal, much wider than long; pereonite-1 with two simple setae distally.</p><p>Pleotelson (Figs 2, 5B) about 5% of TL, same length of pleonites 5 and 6 combined; sub-rectangular, with two broom setae and two simple setae, apex blunt; pleonite-5 with two simple setae distally.</p><p>Antennule (Figs 3A, 6A-B): slightly longer than 2/3 length of cephalothorax. Article-1 about 3.1 times long than wide, longer than distal three articles, with three setulose setae and one simple seta along lateral margin. Article-2 about 1.1 times longer than wide, with distodorsal simple seta; with two setulose setae and one long (longer than articles 2 to 4 combined) simple seta on distoventral margin. Article-3 about 1.5 times wider than long, with two simple setae dorsally and one simple seta lateral. Article-4 about 1.4 times longer than wide, with four simple setae of different length. Terminal segment minute and covered by article-4, with one seta and one aesthetasc (only visible with SEM images) (Fig. 6B).</p><p>Antenna (Figs 3B, 6B): article-1 short, asetose. Article-2 about 1.2 times wider than long, with distolateral short seta. Article-3 about 1.3 times wider than long, with distodorsal long simple seta. Article-4 about 3.0 times longer than wide, longer than articles 2 and 3 combined, with one setulose seta and two simple setae on distoventral setae margin. Article-5 about 2.0 times longer than wide, with one simple seta on distolateral margin. Article-6 minute, with five simple setae of unequal length.</p><p>Mouthparts: Labrum (Figs 3C, 7A): hood-shaped and finely setose. Mandibles (Fig. 3D-E): left mandible, incisor with two to three uneven denticles; lacinia mobilis narrow, apparently smooth (Fig. 3D). Right mandible incisor with broad and crenulate upper margin (Fig. 3E). Molar process pointed, with small distoventral spines (Fig. 3D-E). Labium (Figs 3F, 6B, 7A): bilobed with distolateral processes. Maxillule (Figs 3G, 6B, 7A): endite with seven distal spiniform setae (two bifid), two sub-distal simple setae, and cluster of setules on distal margin; palp bearing two long terminal setae of unequal length. Maxilla: not recovered.</p><p>Maxilliped (Figs 3H, 7A): basis fused, apparently asetose. Endites unfused, with one simple seta and medial small process, outer margin with small spine. Palp: Article-1 asetose. Article-2 with setulose seta on outer margin, inner sub-distal margin with two simple setae and geniculate, finely-pectinate spiniform seta (serrations visible at magnification 100x). Article-3 with two setulose setae on inner margin. Article-4 with subdistal setulose setae and cluster of setules on outer margin (Fig. 7A), inner and distal margin with five setulose setae.</p><p>Epignath: not recovered.</p><p>Cheliped: (Figs 4A-B, 6A-C, 8): cheliped attached just anterior to the posterior margin of cephalothorax and very close to the midventral line, via sclerite (Figs 6B-C, 8). Basis about 1.9 times longer than wide, with subdistal short seta. Merus triangular, with simple seta on ventral margin. Carpus about 1.8 times longer than wide, anterior margin with distodorsal seta; ventral margin with two simple setae of different length. Propodus about 1.4 times longer than wide, with small simple seta near insertion of dactylus. Fixed finger with crenulated ventral margin, with two ventral setae and three simple setae on outer incisive margin, with two to three sharp denticles on inner margin. Inner surface (Fig. 4B) with three short simple setae (one distinctly longest) at articulation with dactylus. Dactylus with long simple proximal seta on inner side.</p><p>Pereopod-1 (Figs 5A, 8): attached ventrally, coxa with simple seta on anterodistal margin. Basis about 4.0 times longer than wide, asetose. Ischium wider than long, with simple seta shorter than merus. Merus about 1.5 times longer than wide, with two distoventral pectinate setae (Fig. 5H) (one short and one robust, just longer than carpus). Carpus about 1.7 times longer than wide; two pectinate distodorsal setae and one robust pectinate distoventral seta (Fig. 5D). Propodus about 3.1 times longer than wide; distodorsal margin with spine-like apophysis; distoventral margin with one pectinate subdistal seta. Dactylus elongate, together with unguis longer than propodus, dactylus shorter than unguis.</p><p>Pereopod-2 (Fig. 5B): similar to pereopod-1, except basis and propodus longer. Propodus with ventrodistal pectinate small seta. Dactylus and unguis shorter than propodus.</p><p>Pereopod- 3 (Fig. 5C): similar in form to pereopod-2, except shorter than other five pereopods.</p><p>Pereopod-4 (Figs 5D, 8): attached ventrally, basis about 4.0 times longer than wide, with setulose seta on proximal dorsal margin. Ischium wider than long with two simple setae of unequal length on posterior margin, long seta shorter than merus. Merus about 2.0 times longer than wide, with two distoventral pectinate setae. Carpus about 2.0 times longer than wide, with four pectinate setae of unequal length. Propodus about 3.5 times longer than wide; distodorsal margin with long pectinate setae, reaching beyond the dactylus; distoventral margin with two pectinate setae. Dactylus and unguis longer than propodus, dactylus shorter than unguis.</p><p>Pereopod-5 (Fig. 5E): Similar to pereopod-4, except carpus and dactylus longer. Basis with setulose seta on mid-ventral margin.</p><p>Pereopod- 6 (Fig. 5F): similar to pereopod-5, except basis, carpus, and propodus slightly longer; ischium, merus, and dactylus shorter.</p><p>Pleopods: Absent. (Fig. 7B)</p><p>Uropod (Figs 5G, 7B): biramous, twice as long as pleotelson, but half as long as entire pleon. Basal article shorter than pleotelson, without distal apophyses. Exopod uni-articulate, slightly longer than endopod article-1, with simple seta on mid-lateral? margin, and two simple distal setae (one longer). Endopod biarticulate, article-1 with three setae (two setulose and one simple) on subdistal inner margin; article-2 with subdistal simple lateral seta, with one setulose and five (four long and one short) simple setae distally.</p><p>Male. Unknown.</p><p>Ovigerous female. As above. When embryos were present, six (smaller embryos) was the most observed; typically only three or fewer were present.</p><p>Remarks.</p><p>Among the family Colletteidae, Oahutanais makalii sp. n. shows some similarities with Cetiopyge mira Larsen and Heard, 2002, Isopodidus janum Larsen and Heard, 2002, and Collettea minima Hansen, 1913 (see Larsen 2000) in having the unusual presence of bifurcate spiniform terminal setae on the maxillule endite; however, the presence of this kind of setae has also been reported in some species of other families such as the Tanaellidae Larsen &amp; Wilson, 2002 ( Arthrura andriashevi Kudinova-Pasternak, 1966), Cryptocopidae Sieg, 1977 ( Curtichelia expressa Kudinova-Pasternak, 1987), and Paratanaoidea incertae sedis ( Parafilitanais mexicanus Larsen, 2002). Larsen and Heard (2002) suggested that this setal character could have a wider occurrence in the deep-sea species, since it has only been reported in specimens collected in deep waters. Our results indicated that the presence of these unusual setae is not restricted to deep-sea Tanaidacea, since Oahutanais makalii was found in shallow waters (&lt;105 m).</p></div>	https://treatment.plazi.org/id/A657F81AB46BE6C8ACF0D185F5EF53BE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Morales-Nunez, Andres G.;Larsen, Kim;Cooke, William J.	Morales-Nunez, Andres G., Larsen, Kim, Cooke, William J. (2016): Oahutanais makalii, a new genus and species of colletteid tanaidacean (Crustacea, Peracarida) from shelf-waters off Hawaii, with a taxonomic key. Zoosystematics and Evolution 92 (1): 1-12, DOI: http://dx.doi.org/10.3897/zse.92.5581, URL: http://dx.doi.org/10.3897/zse.92.5581
