taxonID	type	description	language	source
0FFBEDDECE8E5002BE42573A9CA558E8.taxon	diagnosis	Diagnosis. Larva. Following combination of characters: A) dorsal surface of labrum with submarginal arc of 13 - 15 long, clavate setae; B) labial palp segment II with a broad, thumb-like distomedial protuberance, segment III slightly pentagonal; C) left mandible without setae at apex of mola; D) fore femur rather broad, length 3.4 x maximum width, dorsal margin with 8 - 11 curved, spine-like setae; E) paraproct distally expanded, with 34 - 39 marginal, stout spines.	en	Kaltenbach, Thomas, Gattolliat, Jean-Luc (2020): Labiobaetis Novikova & Kluge in Borneo (Ephemeroptera, Baetidae). ZooKeys 914: 43-79, DOI: http://dx.doi.org/10.3897/zookeys.914.47067, URL: http://dx.doi.org/10.3897/zookeys.914.47067
0FFBEDDECE8E5002BE42573A9CA558E8.taxon	description	Description. Larva (Figs 1, 2, 10 a). Body length 3.5 - 4.3 mm; antennae and cerci broken. Colouration. Head, thorax and abdomen dorsally brown; head and thorax with bright median, dorsal suture. Head, thorax and abdomen ventrally light brown; femur ecru, with brown dorsal margin and brown ventrodistomedial spot, tibia and tarsus brown, caudal filaments ecru. Antenna (Fig. 2 g) with scape and pedicel subcylindrical, with poorly developed distolateral process at scape. Labrum (Fig. 1 a). Rectangular, length 0.6 x maximum width. Distal margin with medial emargination and a small process. Dorsally with medium, fine, simple setae scattered over surface; submarginal arc of setae composed of 13 - 15 long, clavate setae. Ventrally with marginal row of setae composed of anterolateral long, feathered setae and medial long, bifid setae; ventral surface with four short, spine-like setae near lateral and anterolateral margin. Right mandible (Fig. 1 b, c). Incisors fused. Outer and inner sets of denticles with 4 + 3 denticles and one minute intermediate denticle. Inner margin of innermost denticle with a row of thin setae. Prostheca robust, apically and distolaterally denticulate. Margin between prostheca and mola straight, with minute denticles. Tuft of setae at apex of mola present. Left mandible (Fig. 1 d, e). Incisors fused. Outer and inner sets of denticles with 4 + 3 denticles and one minute intermediate denticle. Prostheca robust, apically with small denticles and comb-shaped structure. Margin between prostheca and mola straight, with minute denticles towards subtriangular process. Subtriangular process long and slender, above level of area between prostheca and mola. Denticles of mola apically constricted. Tuft of setae at apex of mola absent. Both mandibles with lateral margins almost straight. Basal half with fine, simple setae scattered over dorsal surface. Hypopharynx (Fig. 1 f). Lingua approx. as long as superlingua. Lingua longer than broad; medial tuft of stout setae well developed; distal half laterally expanded. Superlingua straight; lateral margin rounded; fine, long, simple setae along distal margin. Maxilla (Fig. 1 g). Galea-lacinia with two simple, robust apical setae under crown. Inner dorsal row of setae with three denti-setae, distal denti-seta tooth-like, middle and proximal denti-setae slender, bifid and pectinate. Medially with one bipectinate, spine-like seta and 3 - 4 medium, simple setae. Maxillary palp 1.5 x as long as length of galea-lacinia; 2 - segmented; palp segment II 1.5 x length of segment I; setae on maxillary palp fine and simple, scattered over surface of segments I and II; apex of last segment rounded, with excavation at inner distolateral margin. Labium (Fig. 1 h). Glossa basally broad, narrowing toward apex; shorter than paraglossa; inner margin with five spine-like setae increasing in length distally; apex with two long and one medium, robust, pectinate setae; outer margin with five long, spine-like setae; ventral surface with fine, simple, scattered setae. Paraglossa sub-rectangular, curved inward; apex rounded; with three rows of long, robust, distally pectinate setae in apical area and two or three medium, simple setae in anteromedial area; dorsally with a row of three long, spine-like setae near inner margin. Labial palp with segment I 0.8 x length of segments II and III combined. Segment I with fine, simple setae along margins. Segment II with broad, thumb-like distomedial protuberance; distomedial protuberance 0.6 x width of base of segment III; inner and outer margins with short, fine, simple setae; dorsally with one or two long, spine-like seta near outer margin. Segment III slightly pentagonal; apex rounded; length 1.1 x width; ventrally covered with short, spine-like, simple setae and short, fine, simple setae. Hindwing pads absent. Foreleg (Fig. 2 a, b, c). Ratio of foreleg segments 1.3: 1.0: 0.6: 0.2. Femur. Length ca. 3 x maximum width. Dorsal margin with a row of 8 - 11 curved, spine-like setae; length of setae 0.29 x maximum width of femur. Apex rounded; with one pair of curved, spine-like setae, one or a few short stout setae and some fine, simple setae. Stout, lanceolate setae scattered along the ventral margin; femoral patch absent. Tibia. Dorsal margin with a row of stout, apically rounded setae, apically one longer, apically rounded seta. Ventral margin with a row of curved, spine-like setae, on apex a few stout, spine-like, partly bipectinate setae and a tuft of fine, simple setae. Anterior surface scattered with stout, lanceolate setae. Patellotibial suture present on basal 1 / 3 area. Tarsus. Dorsal margin almost bare. Ventral margin with a row of curved, spine-like setae. Tarsal claw with one row of 9 - 11 denticles; distally pointed; with two stripes; subapical setae absent. Terga (Fig. 2 d). Surface with rows of U-shaped scale bases and scattered fine, simple, setae. Posterior margin of tergum IV with triangular spines, wider than long. Gills (Fig. 2 e). Present on segments II - VII. Margin with small denticles intercalating fine simple setae. Tracheae partly extending from main trunk towards outer and inner margins. Gill IV as long as length of segments V and 1 / 3 VI combined. Gill VII as long as length of segments VIII and 1 / 4 IX combined. Paraproct (Fig. 2 f). Distally expanded, with 34 - 39 stout marginal spines. Surface scattered with U-shaped scale bases, fine simple setae and micropores. Cercotractor with small marginal spines.	en	Kaltenbach, Thomas, Gattolliat, Jean-Luc (2020): Labiobaetis Novikova & Kluge in Borneo (Ephemeroptera, Baetidae). ZooKeys 914: 43-79, DOI: http://dx.doi.org/10.3897/zookeys.914.47067, URL: http://dx.doi.org/10.3897/zookeys.914.47067
0FFBEDDECE8E5002BE42573A9CA558E8.taxon	etymology	Etymology. Dedicated to Dr. Kate Baker (University of Exeter, UK), who collected the specimens in Brunei.	en	Kaltenbach, Thomas, Gattolliat, Jean-Luc (2020): Labiobaetis Novikova & Kluge in Borneo (Ephemeroptera, Baetidae). ZooKeys 914: 43-79, DOI: http://dx.doi.org/10.3897/zookeys.914.47067, URL: http://dx.doi.org/10.3897/zookeys.914.47067
0FFBEDDECE8E5002BE42573A9CA558E8.taxon	distribution	Distribution. Brunei (Fig. 15 c).	en	Kaltenbach, Thomas, Gattolliat, Jean-Luc (2020): Labiobaetis Novikova & Kluge in Borneo (Ephemeroptera, Baetidae). ZooKeys 914: 43-79, DOI: http://dx.doi.org/10.3897/zookeys.914.47067, URL: http://dx.doi.org/10.3897/zookeys.914.47067
5DF31DA7EE105AEE9D71458B922A3965.taxon	diagnosis	Diagnosis. Larva. Following combination of characters: A) dorsal surface of labrum with submarginal arc of 9 - 10 feathered setae (Mueller-Liebenau 1984 b: fig. 2 a); B) labial palp segment II with a large, lobed distomedial protuberance, segment III oblong, apically slightly pointed; C) fore femur rather slender, length 3.6 x maximum width, dorsal margin with a row of 11 - 13 curved, spine-like setae (Mueller-Liebenau 1984 b: fig. 2 i); D) seven pairs of gills; E) hindwing pads present, small; F) distolateral process at scape well developed (Mueller-Liebenau 1984 b: fig. 2 f).	en	Kaltenbach, Thomas, Gattolliat, Jean-Luc (2020): Labiobaetis Novikova & Kluge in Borneo (Ephemeroptera, Baetidae). ZooKeys 914: 43-79, DOI: http://dx.doi.org/10.3897/zookeys.914.47067, URL: http://dx.doi.org/10.3897/zookeys.914.47067
5DF31DA7EE105AEE9D71458B922A3965.taxon	description	Description. Male imago (Fig. 12 b, 13 b). Body length 4.6 mm, forewing length 4.5 mm. Colouration. Head beige. Turbinate eyes dark orange brown, shaft slightly lighter. Thorax beige, pronotum dark olive brown, mesonotum olive. Wings hyaline, venation hyaline. Abdomen: terga olive, sterna transparent. Forewing (Fig. 12 b). Pterostigma with seven cross-veins, only two proximal ones reaching subcostal vein; double intercalary veins shorter than distance between corresponding main veins at wing margin. Genitalia (Fig. 13 b). Basal segment of gonostylus (unistyliger) with inner margin apically slightly expanded; segments I and II almost completely fused; constriction at base of segment II; segment III quadrangular. Styliger plate between unistyligers trapezoidal, distal margin slightly concave.	en	Kaltenbach, Thomas, Gattolliat, Jean-Luc (2020): Labiobaetis Novikova & Kluge in Borneo (Ephemeroptera, Baetidae). ZooKeys 914: 43-79, DOI: http://dx.doi.org/10.3897/zookeys.914.47067, URL: http://dx.doi.org/10.3897/zookeys.914.47067
5DF31DA7EE105AEE9D71458B922A3965.taxon	distribution	Distribution. Indonesia: Kalimantan, Malaysia: Sabah, Brunei (Fig. 15 b).	en	Kaltenbach, Thomas, Gattolliat, Jean-Luc (2020): Labiobaetis Novikova & Kluge in Borneo (Ephemeroptera, Baetidae). ZooKeys 914: 43-79, DOI: http://dx.doi.org/10.3897/zookeys.914.47067, URL: http://dx.doi.org/10.3897/zookeys.914.47067
5DF31DA7EE105AEE9D71458B922A3965.taxon	materials_examined	Examined material. 11 larvae (2 on slides, GBIFCH 00592240, GBIFCH 00658085, 9 in alcohol, GBIFCH 00515368, GBIFCH 00515370), Brunei, Temburong District, Ulu Temburong National Park, Belalong River (near field station), 04 ° 32.82 ' N, 115 ° 09.50 ' E, 100 m, V. 2014, K. Baker leg.; 1 larva (in alcohol, GBIFCH 00515369), Brunei, Temburong District, Ulu Temburong National Park, Sungai Seluju (small tributary to Temburong River, near station), 04 ° 33.83 ' N, 115 ° 08.92 ' E, 90 m, V. 2014, K. Baker leg.; 3 larvae (2 on slides, GBIFCH 00658081, GBIFCH 00592244, 1 in alcohol, GBIFCH 00515372), Malaysia, Sabah, Tawau River, primary forest, 04 ° 24.08 ' N, 117 ° 53.35 ' E, 280 m, 12. III. 2008, Mendoza leg.; 1 male imago (in alcohol and wing on slide, GBIFCH 00672289, GBIFCH 00606854), Malaysia, Sabah, Tawau River, primary forest, 04 ° 24.08 ' N, 117 ° 53.35 ' E, 280 m, 12. III. 2008, Mendoza leg.; 9 larvae (1 on slide, GBIFCH 00465236, 8 in alcohol, GBIFCH 00515394, GBIFCH 00515309, GBIFCH 00515296, GBIFCH 00515376, GBIFCH 00515299), Indonesia, East Kalimantan, Bas. Malinau, River Rian, loc. Langap South (1997 - bloc 6), trib. Belakau, 03 ° 04.07 ' N, 116 ° 30.43 ' E, 07. VII. 2000, P. Derleth leg.; 14 larvae (in alcohol, GBIFCH 00515392, GBIFCH 00515393, GBIFCH 515315, GBIFCH 515312, GBIFCH 515306), Indonesia, East Kalimantan, Bas. Malinau, River Seturan, loc. Seturan (2001 - bloc 57), trib. Bengahau, 02 ° 59.37 ' N, 116 ° 30.77 ' E, 08. VIII. 2000, P. Derleth leg.; 6 larvae (in alcohol, GBIFCH 00515317, GBIFCH 00515321, GBIFCH 00515383), Indonesia, East Kalimantan, Bas. Malinau, River Seturan, loc. Seturan (1999 - block 39 - 40), trib. Temalat (Sungai Guang), 03 ° 00.17 ' N, 116 ° 32.40 ' E, 01. VII. 2000, P. Derleth leg.; 1 larva (in alcohol, GBIFCH 00515313), Indonesia, East Kalimantan, Bas. Malinau, River Rian, loc. Langap South (1995), trib. Ngayo, 03 ° 01.80 ' N, 116 ° 29.80 ' E, 08. VII. 2000, P. Derleth leg.; 2 larvae (in alcohol, GBIFCH 00515382, GBIFCH 00515310), Indonesia, East Kalimantan, Bas. Malinau, River Rian, Langap South (1995), trib. Ngayo, 03 ° 04.93 ' N, 116 ° 30.97 ' E, 13. VII. 2000, P. Derleth leg.; 3 larvae (in alcohol, GBIFCH 00515395, GBIFCH 00515297, GBIFCH 00515378), Indonesia, East Kalimantan, Bas. Malinau, River Seturan, loc. Seturan (2000 - block 43), trib. Temalat (Sungai Guang), 02 ° 59.48 ' N, 116 ° 33.48 ' E, 16. VIII. 2000, P. Derleth and R. Schlaepfer leg.; 3 larvae (1 on slide, GBIFCH 00465237, 2 in alcohol, GBIFCH 00515307, GBIFCH 00515375), Indonesia, East Kalimantan, Bas. Malinau, River Rian, loc. Seturan (1998 - block 32 - 33), tributary, 03 ° 00.95 ' N, 116 ° 32.27 ' E, 30. III. 2001, P. Derleth leg.; 1 larva (in alcohol, GBIFCH 00515316), Indonesia, East Kalimantan, Bas. Malinau, River Seturan, loc. Seturan (1999 - block 27), tributary, 03 ° 00.95 ' N, 116 ° 30.52 ' E, 10. VII. 2000, P. Derleth leg.; 4 larvae (in alcohol, GBIFCH 00515322, GBIFCH 00515377, GBIFCH 515379), Indonesia, East Kalimantan, Bas. Malinau, River Rian, loc. Langap South (1999 - block 24), tributary, 03 ° 01.67 ' N, 116 ° 31.08 ' E, 11. VII. 2000, P. Derleth leg.; 3 larvae (in alcohol, GBIFCH 00515380, GBIFCH 00515381), Indonesia, East Kalimantan, Bas. Malinau, River Seturan, loc. Seturan, main river, 03 ° 00.08 ' N, 116 ° 30.80 ' E, 28. III. 2001, P. Derleth and B. Feldmeyer leg.; 5 larvae (in alcohol, GBIFCH 00515374), Indonesia, East Kalimantan, Bas. Malinau, River Seturan, loc. Seturan, tributary, 02 ° 59.82 ' N, 116 ° 31.37 ' E, 27. IV. 2001, P. Derleth and M. Sartori leg. All material deposited in MZL.	en	Kaltenbach, Thomas, Gattolliat, Jean-Luc (2020): Labiobaetis Novikova & Kluge in Borneo (Ephemeroptera, Baetidae). ZooKeys 914: 43-79, DOI: http://dx.doi.org/10.3897/zookeys.914.47067, URL: http://dx.doi.org/10.3897/zookeys.914.47067
DB83AAA0FC2E594BA767F8339E6624A0.taxon	diagnosis	Diagnosis. Larva. Following combination of characters: A) dorsal surface of labrum with submarginal arc of 10 - 12 long, feathered setae; B) labial palp segment II with a large, lobed distomedial protuberance, segment III slightly pentagonal; C) fore femur rather broad, length ca. 4 x maximum width, dorsal margin with a row of 12 - 14 curved, spine-like setae; D) hindwing pads well developed; E) paraproct distally not expanded, with 30 - 37 marginal, stout spines.	en	Kaltenbach, Thomas, Gattolliat, Jean-Luc (2020): Labiobaetis Novikova & Kluge in Borneo (Ephemeroptera, Baetidae). ZooKeys 914: 43-79, DOI: http://dx.doi.org/10.3897/zookeys.914.47067, URL: http://dx.doi.org/10.3897/zookeys.914.47067
DB83AAA0FC2E594BA767F8339E6624A0.taxon	description	Description. Larva (Figs 5, 6, 11 a). Body length 5.2 mm; antenna: approximately 2.5 x as long as head length; cerci broken. Colouration. Head, thorax and abdomen dorsally brown; head and thorax with bright median, dorsal suture, abdominal segment X light brown. Head, thorax and abdomen ventrally light brown, legs light brown with a brown spot medially and apically on femur, caudal filaments light brown. Antenna (Fig. 6 g) with scape and pedicel subcylindrical, with well-developed distolateral process at scape. Labrum (Fig. 5 a). Rectangular, length 0.7 x maximum width. Distal margin with medial emargination and a small process. Dorsally with medium to long, fine, simple setae scattered over surface; submarginal arc of setae composed of 10 - 12 long, feathered setae. Ventrally with marginal row of setae composed of anterolateral long, feathered setae and medial long, bifid setae; ventral surface with five short, spine-like setae near lateral and anterolateral margin. Right mandible (Fig. 5 b, c). Incisors fused. Outer and inner sets of denticles with 4 + 3 denticles and one minute intermediate denticle. Inner margin of innermost denticle with a row of thin setae. Prostheca robust, apically denticulate. Margin between prostheca and mola slightly convex, with minute denticles. Tuft of setae at apex of mola present. Left mandible (Fig. 5 d, e). Incisors fused. Outer and inner sets of denticles with 4 + 3 denticles and one minute intermediate denticle. Prostheca robust, apically with small denticles and comb-shaped structure. Margin between prostheca and mola straight, with minute denticles towards subtriangular process. Subtriangular process long and slender, above level of area between prostheca and mola. Denticles of mola apically constricted. Tuft of setae at apex of mola present. Both mandibles with lateral margins almost straight. Basal half with fine, simple setae scattered over dorsal surface. Hypopharynx (Fig. 5 f). Lingua approx. as long as superlingua. Lingua longer than broad; medial tuft of stout setae poorly developed; distal half laterally expanded. Superlingua rounded; lateral margin rounded; fine, long, simple setae along distal margin. Maxilla (Fig. 5 g). Galea-lacinia with two simple, robust apical seta under crown. Inner dorsal row of setae with three denti-setae, distal denti-seta tooth-like, middle and proximal denti-setae slender, bifid and pectinate. Medially with one bipectinate, spine-like seta and four medium, simple setae. Maxillary palp 1.2 x as long as length of galea-lacinia; 2 - segmented; palp segment II 1.6 x length of segment I; setae on maxillary palp fine and simple, scattered over surface of segments I and II; apex of last segment rounded, with excavation at inner distolateral margin. Labium (Fig. 5 h, i). Glossa basally broad, narrowing toward apex; shorter than paraglossa; inner margin with seven or eight spine-like setae increasing in length distally; apex with two long and one medium, robust, pectinate setae; outer margin with five or six long, spine-like setae; ventral surface with short, fine, simple, scattered setae. Paraglossa sub-rectangular, curved inward; apex rounded; with three rows of long, robust, distally pectinate setae in apical area and two medium, simple setae in anteromedial area; dorsally with a row of three long, spine-like setae near inner margin. Labial palp with segment I 0.9 x length of segments II and III combined. Segment I ventrally with short, fine, simple setae. Segment II with large, lobed distomedial protuberance; distomedial protuberance 0.7 x width of base of segment III; inner and outer margin with short, fine, simple setae; dorsally with two medium, spine-like, simple setae near outer margin. Segment III slightly pentagonal; apex truncate; length 1.1 x width; ventrally covered with short, spine-like, simple setae and short, fine, simple setae. Hindwing pads (Fig. 6 h) well developed. Foreleg (Fig. 6 a, b). Ratio of foreleg segments 1.1: 1.0: 0.4: 0.2. Femur. Length ca. 4 x maximum width. Dorsal margin with a row of 12 - 14 curved, spine-like setae; length of setae 0.26 x maximum width of femur. Apex rounded, with one pair of curved, spine-like setae and some short, stout setae. Many short, stout, lanceolate setae scattered along the ventral margin; femoral patch absent. Tibia. Dorsal margin with a row of short, stout setae, on apex one longer seta, and a row of short, stout setae close to dorsal margin. Ventral margin with a row of curved, spine-like setae, on apex two spine-like seta and a tuft of long, fine, simple setae. Anterior surface scattered with stout, lanceolate setae. Patellotibial suture present on basal 1 / 3 area. Tarsus. Dorsal margin with a row of short, stout setae. Ventral margin with a row of curved, spine-like setae. Tarsal claw with one row of 9 - 13 denticles; distally pointed; with four stripes; subapical setae absent. Terga (Fig. 6 c, d). Surface with irregular rows of U-shaped scale bases and scattered fine, simple setae. Posterior margin of tergum IV with rounded or triangular spines, wider than long. Gills (Fig. 6 e). Present on segments I - VII. Margin with small denticles intercalating fine simple setae. Tracheae extending from main trunk to inner and outer margins. Gill I as long as length of 1 / 2 segment II. Gill IV as long as length of segments V and 1 / 3 VI combined. Gill VII as long as length of segments VIII and 1 / 3 IX combined. Paraproct (Fig. 6 f). Distally not expanded with 30 - 37 stout marginal spines. Surface scattered with U-shaped scale bases, fine, simple setae and micropores. Cercotractor with medium marginal spines.	en	Kaltenbach, Thomas, Gattolliat, Jean-Luc (2020): Labiobaetis Novikova & Kluge in Borneo (Ephemeroptera, Baetidae). ZooKeys 914: 43-79, DOI: http://dx.doi.org/10.3897/zookeys.914.47067, URL: http://dx.doi.org/10.3897/zookeys.914.47067
DB83AAA0FC2E594BA767F8339E6624A0.taxon	etymology	Etymology. Dedicated to the indigenous Dayak people of Borneo.	en	Kaltenbach, Thomas, Gattolliat, Jean-Luc (2020): Labiobaetis Novikova & Kluge in Borneo (Ephemeroptera, Baetidae). ZooKeys 914: 43-79, DOI: http://dx.doi.org/10.3897/zookeys.914.47067, URL: http://dx.doi.org/10.3897/zookeys.914.47067
DB83AAA0FC2E594BA767F8339E6624A0.taxon	distribution	Distribution. Indonesia: Kalimantan (Fig. 15 c).	en	Kaltenbach, Thomas, Gattolliat, Jean-Luc (2020): Labiobaetis Novikova & Kluge in Borneo (Ephemeroptera, Baetidae). ZooKeys 914: 43-79, DOI: http://dx.doi.org/10.3897/zookeys.914.47067, URL: http://dx.doi.org/10.3897/zookeys.914.47067
CD22467EC6F05D53BDCDEA5B6307E20B.taxon	diagnosis	Diagnosis. Larva. Following combination of characters: A) dorsal surface of labrum with submarginal arc of 1 + 8 - 10 simple setae, the first three after central seta longer than others and decreasing in length; B) labial palp segment II with a large, lobed distomedial protuberance, segment III conical, apically slightly truncate; C) fore femur rather broad, length 3.4 x maximum width, dorsal margin with a row of ca. 10 curved, spine-like setae; D) six pairs of gills; E) hindwing pads present, minute; F) scape with well-developed distolateral process (Mueller-Liebenau 1984 a: fig. 10 f); G) paraproct distally not expanded, with ca. 12 stout marginal spines (Mueller-Liebenau 1984 a: fig. 10 l).	en	Kaltenbach, Thomas, Gattolliat, Jean-Luc (2020): Labiobaetis Novikova & Kluge in Borneo (Ephemeroptera, Baetidae). ZooKeys 914: 43-79, DOI: http://dx.doi.org/10.3897/zookeys.914.47067, URL: http://dx.doi.org/10.3897/zookeys.914.47067
CD22467EC6F05D53BDCDEA5B6307E20B.taxon	distribution	Distribution. Malaysia: Selangor, Sabah; Vietnam; Brunei (Fig. 15 a).	en	Kaltenbach, Thomas, Gattolliat, Jean-Luc (2020): Labiobaetis Novikova & Kluge in Borneo (Ephemeroptera, Baetidae). ZooKeys 914: 43-79, DOI: http://dx.doi.org/10.3897/zookeys.914.47067, URL: http://dx.doi.org/10.3897/zookeys.914.47067
CD22467EC6F05D53BDCDEA5B6307E20B.taxon	materials_examined	Examined material. Paratype. 1 larva (on slide, no. 41), W. Malaysia, Trib. of Gombak River, 16 1 / 2 miles N of Kuala Lumpur, 14. XI. [19] 68, Coll. Bishop. Other material. 1 larva (on slide, GBIFCH 00658106), Brunei, Temburong District, Ulu Temburong National Park, Belalong River (near field station), 04 ° 32.82 ' N, 115 ° 09.50 ' E, 100 m, V. 2014, K. Baker leg.; 1 larva (on slide, GBIFCH 00592243), Brunei, Temburong District, Ulu Temborong National Park, Belalong River tributary, 04 ° 32.63 ' N, 115 ° 08.85 ' E, 170 m, V. 2014, K. Baker leg.; 5 larvae (2 on slides, GBIFCH 00592241, GBIFCH 00658112, 3 in alcohol, GBIFCH 00515325), Malaysia, Sabah, Tawau River, primary forest, 04 ° 24.23 ' N, 117 ° 53.58 ' E, 280 m, 12. III. 2008, Mendoza leg. All material deposited in MZL, except paratype in Zoologische Staatssammlung Muenchen (ZSM).	en	Kaltenbach, Thomas, Gattolliat, Jean-Luc (2020): Labiobaetis Novikova & Kluge in Borneo (Ephemeroptera, Baetidae). ZooKeys 914: 43-79, DOI: http://dx.doi.org/10.3897/zookeys.914.47067, URL: http://dx.doi.org/10.3897/zookeys.914.47067
21BE64F66B0B55AABF3A4CB998A2938E.taxon	diagnosis	Diagnosis. Larva. Following combination of characters: A) dorsal surface of labrum with submarginal arc of 18 - 22 clavate setae; B) labial palp segment II with a broad, thumb-like distomedial protuberance, segment III oblong; C) left mandible without setae at apex of mola; D) fore femur rather broad, length 3.4 x maximum width, dorsal margin with a row of 15 - 19 curved, spine-like setae; E) paraproct distally expanded, with 27 - 33 marginal, stout spines, some of them with split tips.	en	Kaltenbach, Thomas, Gattolliat, Jean-Luc (2020): Labiobaetis Novikova & Kluge in Borneo (Ephemeroptera, Baetidae). ZooKeys 914: 43-79, DOI: http://dx.doi.org/10.3897/zookeys.914.47067, URL: http://dx.doi.org/10.3897/zookeys.914.47067
21BE64F66B0B55AABF3A4CB998A2938E.taxon	description	Description. Larva (Figs 3, 4, 10 b). Body length 3.8 - 6 mm. Cerci: approx. as long as body length. Terminal filament: approx. as long as 1 / 2 length of cerci. Antenna: approximately 3 x as long as head length. Colouration. Head, thorax, and abdomen dorsally brown; head and thorax with bright, median, dorsal suture. Head, thorax, and abdomen ventrally light brown, legs light brown, caudal filaments light brown. Antenna (Fig. 4 i) with scape and pedicel subcylindrical, without distolateral process at scape. Labrum (Fig. 3 a). Rectangular, length 0.6 x maximum width. Distal margin with medial emargination and a small process. Dorsally with medium, fine, simple setae scattered over surface; submarginal arc of setae composed of 18 - 22 long, clavate setae. Ventrally with marginal row of setae composed of lateral and anterolateral long, feathered setae and medial long, bifid setae; ventral surface with five short, spine-like setae near lateral and anterolateral margin. Right mandible (Fig. 3 b, c). Incisors fused. Outer and inner sets of denticles with 4 + 3 denticles and one minute intermediate denticle. Inner margin of innermost denticle with a row of thin setae. Prostheca robust, apically denticulate. Margin between prostheca and mola slightly convex, with a few minute setae. Tuft of setae at apex of mola present. Left mandible (Fig. 3 d, e). Incisors fused. Outer and inner sets of denticles with 4 + 3 denticles and one minute intermediate denticle. Prostheca robust, apically with small denticles and comb-shaped structure. Margin between prostheca and mola straight, with minute denticles towards subtriangular process. Subtriangular process long and slender, above level of area between prostheca and mola. Denticles of mola apically constricted. Tuft of setae at apex of mola absent. Both mandibles with lateral margins almost straight. Basal half with fine, simple setae scattered over dorsal surface. Hypopharynx (Fig. 3 f). Lingua approx. as long as superlingua. Lingua approx. as broad as long; medial tuft of stout setae well developed; distal half not expanded. Superlingua rounded; lateral margin rounded; fine, long, simple setae along distal margin. Maxilla (Fig. 3 g, h). Galea-lacinia with two simple, robust apical setae under crown. Inner dorsal row of setae with three denti-setae, distal denti-seta tooth-like, middle and proximal denti-setae slender, bifid and pectinate. Medially with one bipectinate, spine-like seta and three medium, simple setae. Maxillary palp 1.4 x as long as length of galea-lacinia; 2 - segmented. Palp segment II 1.4 x length of segment I. Setae on maxillary palp fine and simple, scattered over surface of segments I and II. Apex of last segment rounded, with strong excavation at inner distolateral margin. Labium (Fig. 3 i). Glossa basally broad, narrowing toward apex; shorter than paraglossa; inner margin with five spine-like setae increasing in length distally; apex with two long and one medium, robust, pectinate setae; outer margin with four long, spine-like setae; ventral surface with short, fine, simple and short, spine-like setae. Paraglossa sub-rectangular, curved inward; apex rounded; with three rows of long, robust, distally pectinate setae in apical area and three medium, simple setae in anteromedial area; dorsally with a row of three long, spine-like setae near inner margin. Labial palp with segment I 0,7 x length of segments II and III combined. Segment I ventrally with short, fine, simple setae. Segment II with broad, thumb-like distomedial protuberance; distomedial protuberance 1.0 x width of base of segment III; inner and outer margin with short, fine, simple setae; dorsally with two long, spine-like, simple setae near outer margin. Segment III oblong; apex rounded; length 1.4 x width; ventrally covered with short to medium, spine-like, simple setae and short, fine, simple setae. Hindwing pads absent. Foreleg (Fig. 4 a-d). Ratio of foreleg segments 1.1: 1.0: 0.4: 0.1. Femur. Length ca. 3 x maximum width. Dorsal margin with a row of 15 - 19 curved, spine-like, apically rounded setae and many long, fine, simple setae and partly a few stout setae near margin; length of setae 0.28 x maximum width of femur. Apex rounded; with one pair of curved, spine-like setae and some short, stout setae. Many stout, lanceolate setae scattered along ventral margin; femoral patch poorly developed. Tibia. Dorsal margin with a row of stout, lanceolate, apically rounded setae and fine, simple setae; on apex one larger, lanceolate, apically rounded seta. Ventral margin with a row of curved, spine-like setae, on apex one bipectinate, spine-like seta and a tuft of long, fine, simple setae. Anterior surface scattered with stout, lanceolate setae. Patellotibial suture present on basal 1 / 3 area. Tarsus. Dorsal margin with a row of small, stout setae and fine, simple setae. Ventral margin with a row of curved, spine-like setae. Tarsal claw with one row of 9 - 11 denticles; distally pointed; with three stripes; subapical setae absent. Terga (Fig. 4 e, f). Surface with rows of U-shaped scale bases. Posterior margin of tergum IV with triangular or rounded spines, wider than long. Gills (Fig. 4 g). Present on segments II - VII. Margin with small denticles intercalating both short and medium, fine, simple setae. Tracheae extending from main trunk to inner and outer margins. Gill IV as long as length of segments V and 1 / 3 VI combined. Gill VII as long as length of segment VIII. Paraproct (Fig. 4 h). Distally expanded, with 27 - 33 stout marginal spines, some of them with split tips. Surface scattered with U-shaped scale bases, fine, simple setae and micropores. Cercotractor with small marginal spines.	en	Kaltenbach, Thomas, Gattolliat, Jean-Luc (2020): Labiobaetis Novikova & Kluge in Borneo (Ephemeroptera, Baetidae). ZooKeys 914: 43-79, DOI: http://dx.doi.org/10.3897/zookeys.914.47067, URL: http://dx.doi.org/10.3897/zookeys.914.47067
21BE64F66B0B55AABF3A4CB998A2938E.taxon	description	Description. Male imago (Fig. 12 a, 13 a, c). Body length 3.8 mm, forewing length 4.4 mm. Colouration. Head light beige. Turbinate eyes orange, shaft proximally lighter. Thorax light beige with lateral brown markings (Fig. 13 c). Legs light brown. Wings hyaline, venation hyaline. Abdomen dorsally whitish with lateral orange brown markings (Fig. 13 c), segment VII dorsally orange brown. Forewing (Fig. 12 a). Pterostigma with three cross-veins, distal one bifurcated and reaching subcostal vein, in the middle a short one not reaching subcostal vein and the proximal one reaching subcostal vein; double intercalary veins generally shorter than distance between corresponding main veins at wing margin. Hindwing absent. Genitalia (Fig. 13 a). Basal segment of gonostylus (unistyliger) with inner margin apically only slightly expanded; segments I and II almost completely fused; constriction at base of segment II; segment III ovoid. Styliger plate between unistyligers poorly developed, distal margin straight.	en	Kaltenbach, Thomas, Gattolliat, Jean-Luc (2020): Labiobaetis Novikova & Kluge in Borneo (Ephemeroptera, Baetidae). ZooKeys 914: 43-79, DOI: http://dx.doi.org/10.3897/zookeys.914.47067, URL: http://dx.doi.org/10.3897/zookeys.914.47067
21BE64F66B0B55AABF3A4CB998A2938E.taxon	etymology	Etymology. Dedicated to the indigenous Penan people of Borneo.	en	Kaltenbach, Thomas, Gattolliat, Jean-Luc (2020): Labiobaetis Novikova & Kluge in Borneo (Ephemeroptera, Baetidae). ZooKeys 914: 43-79, DOI: http://dx.doi.org/10.3897/zookeys.914.47067, URL: http://dx.doi.org/10.3897/zookeys.914.47067
21BE64F66B0B55AABF3A4CB998A2938E.taxon	distribution	Distribution. Indonesia: Kalimantan, Brunei, Malaysia: Sabah (Fig. 15 d).	en	Kaltenbach, Thomas, Gattolliat, Jean-Luc (2020): Labiobaetis Novikova & Kluge in Borneo (Ephemeroptera, Baetidae). ZooKeys 914: 43-79, DOI: http://dx.doi.org/10.3897/zookeys.914.47067, URL: http://dx.doi.org/10.3897/zookeys.914.47067
