identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
CD4D87D9FFAF6529FF33BEB46135F8C9.text	CD4D87D9FFAF6529FF33BEB46135F8C9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Campsurus cristales Molineri & Granados-Martinez 2019	<div><p>Campsurus cristales sp. nov.</p><p>(Figs. 1 D–G, 4A–E)</p><p>http://zoobank.org/NomenclaturalActs/ 418316B8-092E-4C03-960C-DF24C22F8EC2</p><p>Diagnosis. Campsurus cristales sp. nov. is described from male and female adults. It can be distinguished from other species of Campsurus by: 1) small to medium size (length of male FW 7.0–8.0 mm, female FW 10.7 mm); 2) posterior margin of male abdominal sternum IX tri–lobed (Figs. 1D); 3) pedestal almost touching each other medially at base, strongly elongated, parastylus broadly rounded apically, relatively wide and somewhat projected distally (Fig. 1D, 4D), inner corner rounded and slightly projected (Fig. 1 D–4D); 4) penes with a strongly expanded dorsal area (ea in Fig. 4E), main lobe sclerotized, long and relatively acute apically, secondary membranous lobe shorter, rounded apically and cylindrical (Figs. 1 D–E, 4D–E).</p><p>Type material. Holotype ♂ imago from COLOMBIA, Meta, La Macarena, Pailones, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.78306&amp;materialsCitation.latitude=2.2677777" title="Search Plazi for locations around (long -73.78306/lat 2.2677777)">Caño Cristales</a>, N 2° 16' 4" / W 73° 46' 59", 257 m, 26.x.2016, light trap 18–20 hs, C Granados col. (MUSENUV) . Paratypes: 16 ♂ imagos (1 on slide IBN803 CM) and 1 ♀ imago (slide IBN804 CM), same data as holotype (8 ♂ and 1♀ in IBN, 4♂ in MUSENUV, 4♂ in CEBUC) .</p><p>Male imago. Length (mm): body, 7.2–7.7; fore wing, 7.0–8.0; hind wing, 3.5–4.0; fore leg, 3.5–4.2; cerci, 20.0–21.0. General coloration yellowish white, widely shaded with gray (Fig. 4A). Head completely shaded with black dorsally, and on antenna (except for hyaline flagellum), ventrally with gray marks along carinae. Thorax (Fig. 4A, 4C). Pronotum translucent, extensively shaded with black, darker on anterior portion, medial line, and margins; prosternum yellowish white. Mesonotum yellowish, shaded with gray medially on parapsidal sutures, and on a triangle between PSP; pleura and sterna yellowish white, shaded with slightly gray. Metanotum yellowish white, shaded with gray medially. Meso furcasternal plates as in Fig. 4C. Legs. Fore leg completely shaded with gray; middle and hind legs yellowish translucent, shaded completely with gray. Wings. Membrane hyaline, except C and Sc areas shaded with brownish gray, veins hyaline, except C, Sc, and R 1 brownish on basal 2/3. Hind wing hyaline, except base of veins yellowish. Abdomen (Fig. 4A) whitish translucent, extensively shaded with gray on terga, except on small paler dots. Abdominal sterna translucent white, shaded with gray on medial line of sterna VII–IX. Genitalia (Figs. 1 D–E, 4D–E)): yellowish white; sternum IX with three slightly marked lobes on hind margin (arrows in Figs. 1D), medial line and hind margin black; pedestal of each side touching basally, with apically rounded parastylus (mo in Fig. 4D), round inner corner small, slightly projected (mi in Fig. 4D); forceps short and translucent, shaded with gray basally; base of main lobe of penes sclerotized, very expanded dorsally (ea in Figs. 4 D–E), apical portion narrowing abruptly and finger like, main lobe sclerotized and acute apically, secondary lobe membranous and rounded apically, slender and cylindrical. Caudal filament hyaline translucent.</p><p>Female adult. Length (mm): body, 8.0; fore wing, 10.7; hind wing, 4.5; fore leg, 1.2; cerci broken. Similar to male imago, except pigmentation more heavily marked (Fig. 4B). Thorax. Pronotum shaded with gray, except pale transverse line between anterior and posterior portions; thin black medial line surrounded by a pale area along anterior portion of pronotum. Mesonotum (Fig. 4B) additionally shaded gray on anterior triangular mark (after mesonotal protuberance). Legs shaded completely with gray. Wing membranes yellowish translucent. Abdomen shaded extensively with black on dorsum (Fig. 4B); sternum VIII with small anteromedian subovate socket (Figs. 1 F–G). Caudal filaments translucent whitish.</p><p>Egg. Length, 275–300 µm; width, 225–250 µm. Oval, cup–shaped, yellowish with whitish polar cap.</p><p>Etymology. The name alludes to the type locality (Caño Cristales).</p><p>Distribution (Fig. 5). Colombia (Meta).</p><p>Discussion. Campsurus cristales sp. nov. is very similar to C. janae Molineri &amp; Salles, and will key to that species in Molineri &amp; Salles (2017). Nevertheless, the pedestals are broader and parastylus more projected distally in the new species. Also, the penes are even more expanded in a dorsal ovate flat structure (ea in Figs. 4 D–E), which is more circular in C. janae . Additionally, the main lobe of penes in the new species is longer and apically acute (it is shorter and rounded in C. janae). Finally, the tri–lobed hind margin of the sternum IX is unique for the new species.</p></div>	https://treatment.plazi.org/id/CD4D87D9FFAF6529FF33BEB46135F8C9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Molineri, Carlos;Granados-Martinez, Cristian E.	Molineri, Carlos, Granados-Martinez, Cristian E. (2019): Two new species of Campsurus Eaton (Ephemeroptera: Polymitarcyidae) from Colombia. Zootaxa 4543 (1), DOI: 10.11646/zootaxa.4543.1.5
CD4D87D9FFAB652FFF33BCF6658BF873.text	CD4D87D9FFAB652FFF33BCF6658BF873.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Campsurus vichada Molineri & Granados-Martinez 2019	<div><p>Campsurus vichada sp. nov.</p><p>(Figs. 1 A–C, Figs. 3 A–D)</p><p>http://zoobank.org/NomenclaturalActs/ C6E4D995-4CC1-4B77-91FC-725E7BE84B0F</p><p>Diagnosis. Campsurus vichada sp. nov., here described from male and female imagos, can be separated from other species of the genus by the characters listed for the albifilum species group (Molineri &amp; Salles 2013) and from the species of this group, by: 1) FW 10.0–11.0 mm (male), 13.5–16.0 (female); 2) dorsum of head shaded black widely, some specimens with Y–shaped pale area between ocelli and extending through occiput (Figs. 3 A–B); 3) abdominal terga shaded slightly darker posteriorly, with a pale median band and other markings as in Figs. 3 A–B; 4) pedestals relatively short, subquadrate, main lobes of penes very long and slender, secondary lobe reduced and relatively short (Figs. 3 E–D).</p><p>Type material. Holotype ♂ imago from COLOMBIA, Vichada, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-68.49305&amp;materialsCitation.latitude=5.5772223" title="Search Plazi for locations around (long -68.49305/lat 5.5772223)">rio Tomo</a>, N 5° 34' 38" / W 68° 29' 35"), 119 m, 04.iv.2017, light trap 18–20 hs, C Granados col. (MUSENUV) . Paratypes: 52 ♂ (slides IBN800 CM, IBN801 CM) and 16 ♀ (slide IBN802 CM), same data as holotype (10 ♂ and 4 ♀ in MUSENUV, 10 ♂ and 4♀ in CEBUC, 32 ♂ and 8 ♀ in IBN) .</p><p>Male imago. Length (mm): body, 10.5–11.0; FW, 10.0–11.0; HW, 4.5–4.7; forelegs, 4.9–5.1; cerci, 27.5–28.5. General coloration yellowish white, shaded with grey dorsally. Head (Fig. 3A) shaded dorsally with grey, some males with lighter coloration on medial line and around median ocellus; vertex with grey shading; some specimens with Y–shaped pale area between ocelli and extending through occiput. Antennae whitish shaded with grey, stronger at margins of scape and pedicel. Thorax (Fig. 3A): pronotum hyaline translucent shaded very slightly with grey on anteromedian zone; mesonotum shaded gray on carinae and on posterior scutal protuberances (PSP); metanotum yellowish, shaded slightly with grey medially. Forelegs whitish shaded completely with purplish grey, lighter on tarsi; legs II–III yellowish translucent with apical black dash (Fig. 3D). Wings: veins whitish translucent, shaded with purplish grey at the base of longitudinal veins, shading more extensive on veins (and membrane around them) Sc and R1. Abdomen (Fig. 3A): terga shaded with grey along a median band; tergum I lightest, IX darkest; tergum I with a median mark (not broken by a pale area medially; terga II–VI shaded with gray except on medial pale band and pale subcircular mark at each side of medial band (Fig. 3A); terga VII–VIII similar but more widely pigmented with gray, and with two pairs of pale circular marks; tergum IX widely pigmented, darker medially, a pair of pale circular marks on anteromedian margin. Abdominal sterna pale not shaded. Genitalia (Figs. 1A, 3 E–D): yellowish white; pedestals relatively short, with outer corner indented so apical half is directed slightly towards the outer side; main lobes of penes very long and slender, secondary lobe relatively short. Cerci whitish translucent.</p><p>Female imago. Length (mm): body, 13.2–19.3; FW, 13.5– 16.0; HW, 6.1–7.7; forelegs broken; cerci, 5.5–6.0. General coloration yellowish (Fig. 3B). Head with dorsal grey shading except on median zone of vertex (Fig. 3B). Thorax: pronotum shaded grey widely, except some pale areas as in Fig. 3B. Mesonotum as male. Foreleg broken off in all specimens, similar to middle and hind legs in shape. Middle and hind legs with apical black dash as in male (Fig. 3C). Abdomen similar to male except broader pale medial band on terga II–VII (Fig. 3B). Sternum VIII with a pair of relatively large anteromedian sockets (Fig. 1 B–C), sockets partially fused forming a single structure. Caudal filaments whitish translucent.</p><p>Eggs. Length, 245–275 µm; width, 200–210 µm. Hemispheric, yellowish, with one whitish polar cap formed by a compound thread (ca. 5.0 mm long) formed by numerous thin filaments tightly braided.</p><p>Etymology. Vichada, from the type locality.</p><p>Distribution (Fig. 5). Colombia (Vichada).</p><p>Discussion. Campsurus vichada sp. nov. shows characters in common with the two clades in the albifilum group found by Molineri &amp; Salles (2013). The large size of the body and the slightly curved pedestals are shared with C. albifilum and C. fuliginatus, but the extremely long penes are more similar to C. gracilipenis and C. homaulos .</p><p>We added the new species to the matrix used in Molineri &amp; Salles (2013), and the searches resulted in the same topology shown by these authors, with the new species resulting sister to the pair C. homaulus + C. gracilipenis (Fig. 2), with good support. The three species share two synapomorphies: 1) a change in continuous character number 1 – Pedestal length/width ratio (from 1.12–1.40 to 1.05), thus the general shape of the pedestal change from subrectangular (plesiomorphy) to subquadrate; and 2) a change in color pattern of abdominal terga (character 7, discrete) from the state 0 (with][–shaped marks) to state 1 (marks closed, forming rectangle or circle).</p><p>The key published in Molineri &amp; Salles (2013) should be modified as follows to include the new species here described. Please note that couplet 5 was also modified to include new data regarding C. gracilipenis and C. homaulus:</p><p>1. Abdominal color pattern pale on terga I–III and very dark on terga IV–IX; secondary lobe of penes short, knoblike (see other characters in Molineri &amp; Salles 2006)........................................................... C. fuliginatus</p><p>- Abdominal color pattern uniform on terga except darker on posterior terga IX or VIII–IX; secondary lobe of penes distinct (Figs. 1A, 3 E–D); FW length variable; pedestals variable..................................................... 2</p><p>2. Pedestal subrectangular, length/width ratio = 1.12–1.40; abdominal terga with submedian][–shaped gray mark............ 3</p><p>- Pedestal subquadrate (Figs. 1A, 3D), length/width ratio = 1.05 or less; abdominal terga more widely pigmented with gray, except small rectangular to ovate pale mark at each side (Fig. 3A)................................................ 4</p><p>3. Pedestals very long (length/width ratio&gt; 2.1) and with the apical inner margin strongly projected posteriorly...... C. yavari</p><p>- Pedestals shorter (length/width ratio&gt; 1.7), inner apical margin broad, not so strongly projected.............. C. albifilum</p><p>4. Body and wings large to medium–sized (FW length&gt; 9.5 mm); secondary lobe of penes reduced, short (Figs. 1A, 3 D–E).................................................................................. Campsurus vichada sp. nov.</p><p>- Body and wings medium to small in size (FW length &lt;9.0 mm); secondary lobe of penes relatively long................. 5</p><p>5. Penes long and slender (length/width mostly&gt; 0.85); basal inner membrane of penes very short in relation to total length of penean lobe (ratio &lt;0.28).................................................................... C. gracilipenis</p><p>- Penes shorter (length/width mostly &lt;0.80); basal inner membrane of penes not so short in relation to total length of penean lobe (ratio&gt;0.32)............................................................................. C. homaulos</p></div>	https://treatment.plazi.org/id/CD4D87D9FFAB652FFF33BCF6658BF873	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Molineri, Carlos;Granados-Martinez, Cristian E.	Molineri, Carlos, Granados-Martinez, Cristian E. (2019): Two new species of Campsurus Eaton (Ephemeroptera: Polymitarcyidae) from Colombia. Zootaxa 4543 (1), DOI: 10.11646/zootaxa.4543.1.5
