taxonID	type	description	language	source
CC4187C9716B3B3DFF4D159B24F21D95.taxon	materials_examined	Material Examined. HOLOTYPE: Australia, Western Australia, Point Peron, 25 ° 51 ' S, 113 ° 34 ' E, 1946, (AM W. 6820) fixed in formalin. Non-types: Australia, Queensland, Lizard Island, off Casuarina Beach, 14 ° 40 ' 46 " S, 145 ° 26 ' 49 " E, Aug 17, 2013, AM W. 44353 (1 incomplete specimen), AM W. 44391 (1 complete and 2 incomplete specimens), fixed in formalin; Aug 20, 2013, AM W. 44633 (1 complete and 2 incomplete specimens), fixed in formalin; Aug 21, 2013, AM W. 44920 (3 complete and 1 incomplete specimens), AM W. 44911 (1 complete and 1 incomplete specimens), fixed in formalin; north west of Watson's Bay, 14 ° 39 ' 41 " S, 145 ° 22 ' 27 " E, Aug 23, 2013, AM W. 44977 (3 incomplete specimens), AM W. 44983 (3 incomplete specimens) fixed in formalin and AM W. 44982 (1 incomplete specimen) fixed in ethanol.	en	Zanol, Joana, Da Silva, Thauane Dos S. C., Hutchings, Pat (2017): One new species and two redescriptions of Marphysa (Eunicidae, Annelida) species of the Aenea-group from Australia. Zootaxa 4268 (3): 411-426, DOI: 10.11646/zootaxa.4268.3.6
CC4187C9716B3B3DFF4D159B24F21D95.taxon	description	Measurements. Holotype incomplete with 163 chaetigers, 53 mm in total length, 5 mm in length through chaetiger 10 and 3 mm wide at this chaetiger with parapodia. Non-types, six complete specimens ranging from 96 – 150 chaetigers, 23 – 45 mm in total length, 2 – 4.4 mm in length through chaetiger 10 and 1.36 – 1.92 mm wide at this chaetiger with parapodia; 14 incomplete specimens ranging from 26 – 148 chaetigers, 10 – 45 mm in total length, 2.25 – 5.4 mm in length through chaetiger 10 and 0.68 – 3 mm wide at this chaetiger with parapodia. Description. Live specimens translucent with reddish hues due to blood (Fig. 1 A). Fixed specimens beige to brown in color (Fig. 1 B – E). Body long and slender, widest at anterior region, evenly tapering towards posterior end; cross section dorsally flattened and ventrally slightly rounded at anterior and median regions, flattened to round at posterior region of body. Chaetigers less than 10 times wider than long at widest region of body. Prostomium approximately as long as, half as deep as and narrower to as wide as peristomium; dorsally flat with anterior end higher (Fig. 1 B, C, D), anteriorly rounded to tapering; not bilobed; median sulcus absent (Fig. 1 C) or inconspicuous (Fig. 1 B) at anterior end, absent at dorsal side and conspicuously present at ventral side (Fig. 1 E). Prostomial appendages short, reaching around one third length of prostomium. Median and lateral antennae in straight line, palps inserted slightly more anteriorly (Fig. 1 B, C). Median antenna longest prostomial appendage; isolated by gap from lateral antennae and palps. Antennal styles and palpostyles tapering to fusiform and wrinkled under light microscope. Antennophores and palpophores ring shaped. One pair of eyes present, conspicuous in shape of dark line between lateral antennae antennophore and palpophore (Fig. 1 D). Separation between peristomial rings distinct all around. Second peristomial ring about as long as 1 / 5 – 1 / 3 of complete peristomium. Peristomial ventro-lateral lips distinct laterally as elevated surface (Fig. 1 D). Peristomial anterior margin of dorsal and ventral sides in shallow arc, lateral margins longer; shortest on dorsal side (Fig. 1 E). Posterior end of muscularized pharynx at chaetiger 3 – 5. Mandibles with small delicate cutting plates, diamond to oval shape, some with medio-anterior teeth (Fig. 1 F, G). MxI around twice as long as carriers and 6 – 8 times longer than locking system (Fig. 1 H, I). MxIII at least in part located ventral to MxII,; attachment lamella oval to rectangular present only at anterior end of median margin of MxIII (Fig. 1 I). Left MxIV wider than long, semicircular; attachment lamella shorter than plate, almost as wide as plate, along most of anterior edge, missing on lateral edge of plate (Fig. 1 I, J). Right MxIV attachment lamella starting between lateralmost teeth 1 – 2, along around ¾ of plate, longest at lateral side and evenly shorter towards opposite side (Fig. 1 I, K). Maxillary formula: I = 1 + 1, II = 5 (4) + 6 (5), III = 6 (5, 7) + 0, IV = 5 (4, 6) + 8 (7 – 10), V = 1 + 1, VI absent. Pre-neurochaetal lobe shorter than neurochaetal lobe along whole body. Post-neurochaetal lobe longer than neurochaetal lobe in anterior chaetigers decreasing in length along body, becoming as long as or shorter than neurochaetal lobe in median and posterior chaetigers; in anterior chaetigers wide, distally truncate with longer dorsal edge (Fig. 2 A, B). Neurochaetal lobe rounded in anterior (Fig. 2 A), tapering in median and posterior chaetigers (Fig. 2 C). Anterior notopodial cirri evenly tapering; from around start of branchiae to end of body some abruptly tapering. Abruptly tapering notopodial cirri vary from simple (Fig. 2 A – C), semi-bifurcated (Fig. 2 D) to bifurcated (Fig. 2 E – G), not consistently present in all chaetigers. Notopodial cirri longer than neurochaetal lobe in anterior chaetigers (Fig. 2 A, B), decreasing in length towards posterior region, about as long as neurochaetal lobe along most of body (Fig. 2 C). Anteriormost ventral cirri digitiform to tapering; basally inflated from chaetiger 4 - 6 until end of body; inflated base round shaped (Fig. 2 A) with rounded tip conspicuous from chaetiger 8 to end of body; inflated base gradually decreasing in size from around chaetigers 21 to 50; rounded tip gradually increasing in length along body. Ventral cirri as long as or longer than neurochaetal lobe at anterior region (Fig. 2 A), slightly shorter to as long as neurochaetal lobe at median and posterior region (Fig. 2 C), sometimes longer than neurochaetal lobe in posteriormost chaetigers. Branchiae palmate present along most of body, with one filament from chaetiger 16 – 24, reaching maximum of 2 – 4 filaments at chaetiger 21 – 62; where best developed, branchial filaments around 2 – 8 and 4 – 6 times longer, respectively, than branchial stems and notopodial cirri; length of filaments start to decrease around chaetiger 61 – 82. Branchial stems completely attached to body wall (Fig. 2 B, E, F). Notopodial aciculae yellow, present in notopodial cirri along whole body (Fig. 2 F). Neurochaetae in two distinct bundles; supra-acicular with limbate chaetae and pectinate chaetae, subacicular with bidentate compound falcigers chaetae and subacicular hooks. Neuroaciculae tip blunt to tapering, brown to yellow until first quarter of body, thereafter only yellow; distributed in oblique row, anteriormost neuroaciculae also dorsalmost in parapodium. Number of neuroaciculae decreasing towards posterior end, 2 – 4 in anteriormost chaetigers, two in first quarter of body and thereafter only one; distributed from median to dorsal region of neurochaetal lobe in most anterior parapodia (P 1) and midline thereafter. Number of limbate chaetae and compound falcigers chaetae also varying along body. Limbate chaetae longer than all other chaetae, serrated, narrowing towards posterior end (Fig. 2 H); around 8 – 18 limbate in anterior region, 3 – 7 in median and 6 – 11 in posterior. Compound falciger chaetae around 14 – 26 in anteriormost chaetigers (P 1), 9 – 14 in first quarter, 3 – 6 in the remaining part of body. Shaft and appendage of compound falciger chaetae marginally serrated; appendage bidentate with guards symmetrically blunt without mucros; both teeth directed laterally; distal tooth tapering and curved; proximal tooth perpendicular to length of chaetae, as long as or longer than distal tooth (Fig. 2 J). One – three thin pectinate chaetae with around 10 – 22 teeth present at anteroventral edge of supra-acicular bundle until second quarter of body (P 3); inner teeth long, equal in length, evenly tapering; outer teeth not equal in length and longer than inner teeth, longest about twice as long as inner teeth (Fig. 2 K, L). Curved thick pectinate chaetae present in median (P 3 – P 4) to posterior regions with around 10 – 14 coarse teeth, all with about same length at distal end but with increasingly deeper proximal ends from margins to center, teeth with deepest proximal end off center (Fig. 2 M); 1 - 3 thick pectinate per parapodium present until end of body. Pseudocompound chaetae and compound spinigers chaetae absent. Subacicular hooks first present from chaetiger 20 – 32, present in all chaetigers thereafter, always single; guards not always present (Fig. 2 N – R); hook light yellow to brown, bidentate, narrower or as wide as aciculae; distal tooth rounded to tapering, shorter than proximal tooth, erect (Fig. 2 N – Q) to curved (Fig. 2 R); proximal tooth about twice as long as distal tooth, perpendicular to length of hook or directed distally, rounded or tapering (Fig. 2 N – R). Pygidium longer on ventral side, with two pairs of pygidial cirri attached to ventral margin; pygidial cirri tapering with irregular cylindrical articulations, dorsal 2 – 4 times longer than ventral cirri. Variation. In some specimens jaws are semi-everted which make prostomium seem shorter and less deep than in those with jaws withdrawn. Eyes are absent in the holotype, this could be due to preservation. The most conspicuously variable feature among examined specimens is the shape of notopodial cirri. They can be simple, semi-bifurcated or bifurcated. All examined specimens have simple notopodial cirri in anteriormost chaetigers. The two largest specimens examined (AM W. 6820 and AM W. 44977) have bifurcated notopodial cirri along most of the examined fragments. The remaining specimens have semi-bifurcate cirri, which may be irregularly distributed along the body. Almost all specimens have abruptly tapering notopodial cirri at least in some chaetigers. Number of limbate chaetae decreases towards median region but in some cases increases in posteriormost chaetigers. Thin pectinate chaetae have more teeth in the longest specimen examined (AM W. 44977) than in the other specimens. Laterally directed proximal teeth of subacicular hooks are usually present in posteriormost chaetigers. Subacicular hooks with smaller teeth may be worn down or characteristic of larger specimens (e. g., AM W. 6820 and AM W. 44977).	en	Zanol, Joana, Da Silva, Thauane Dos S. C., Hutchings, Pat (2017): One new species and two redescriptions of Marphysa (Eunicidae, Annelida) species of the Aenea-group from Australia. Zootaxa 4268 (3): 411-426, DOI: 10.11646/zootaxa.4268.3.6
CC4187C9716B3B3DFF4D159B24F21D95.taxon	discussion	Remarks. Our description complements the original one by Kott (1951) in describing features not mentioned and including comments about variations occurring along the body and intraspecific variation as exhibited by the additional material examined. The main additional features observed in our study are the shape of notopodial cirri, pectinate chaetae and subacicular hooks .. Notopodial cirri may not be completely bifurcated, the semi-bifurcated shape is the most common among the studied specimens. In the original description, pectinate chaetae were described only as pectinate of general type, whereas we refer to it as a thin pectinate. We found two types of pectinate chaetae, thin and thick. Thick pectinate chaetae are restricted to the posterior region, which is absent in the incomplete holotype.	en	Zanol, Joana, Da Silva, Thauane Dos S. C., Hutchings, Pat (2017): One new species and two redescriptions of Marphysa (Eunicidae, Annelida) species of the Aenea-group from Australia. Zootaxa 4268 (3): 411-426, DOI: 10.11646/zootaxa.4268.3.6
CC4187C9716B3B3DFF4D159B24F21D95.taxon	distribution	Distribution. Australia: Western Australia, Queensland (Lizard Island). Arabian Gulf: Saudi Arabia coast (Joydas et al. 2012). South Vietnam: Bay of Nha Trang (Gallardo 1968). Indian Ocean: Mozambique (Day 1967).	en	Zanol, Joana, Da Silva, Thauane Dos S. C., Hutchings, Pat (2017): One new species and two redescriptions of Marphysa (Eunicidae, Annelida) species of the Aenea-group from Australia. Zootaxa 4268 (3): 411-426, DOI: 10.11646/zootaxa.4268.3.6
CC4187C9716F3B32FF4D15FD239118FE.taxon	materials_examined	Material examined. HOLOTYPE: Australia, New South Wales, Pittwater, Careel Bay, 33 ° 36 ' 54 " S, 151 ° 19 ' 24 " E, 17 Feb 2011 (AM W. 37249). PARATYPES: same locality as for holotype, AM W. 37089, AM W. 37088, AM W. 37246, AM W. 37250 (4 specimens).	en	Zanol, Joana, Da Silva, Thauane Dos S. C., Hutchings, Pat (2017): One new species and two redescriptions of Marphysa (Eunicidae, Annelida) species of the Aenea-group from Australia. Zootaxa 4268 (3): 411-426, DOI: 10.11646/zootaxa.4268.3.6
CC4187C9716F3B32FF4D15FD239118FE.taxon	description	Measurements. Holotype, complete female with regenerating pygidium, 164 mm in total length, 379 chaetigers, in four fragments, 132 anteriormost chaetigers, 53 median and 174 posteriormost chaetigers fixed in 8 % formalin, 20 median chaetigers fixed in 95 % ethanol, 6.5 mm length through chaetiger 10 and about 3.5 mm with parapodia (2 mm without parapodia) in width at chaetiger 10 (maximal width). One paratype complete specimen, with 100 mm in total length, 239 chaetigers, 5.2 mm in length through chaetiger 10 and 2 mm with parapodia (1 mm without parapodia) in width at chaetiger 10. Three paratypes incomplete specimens or regenerating posterior end ranging from 50 – 55 mm in total length, 85 – 249 chaetigers, 6.5 – 7 mm in length through chaetiger 10, 2.5 – 3 mm with parapodia (1.6 mm without parapodia) in width at chaetiger 10. Description. (Of holotype, values in parentheses for paratypes). Live specimens whitish translucent with pink to red hues due to red blood. Fixed specimens white, lacking color pattern. Body long, slender, abruptly tapering towards anterior end from around chaetiger 5, evenly tapering towards posterior end; cross section dorsoventrally flattened at anterior region and rounded at median and posterior regions. Chaetigers around 3 – 5 times wider than long at widest region of body. Prostomium as long and wide as peristomium, two-thirds (half) as deep as peristomium; dorsally flat with anterior end higher (completely flat), anteriorly triangular (round); not bilobed; median sulcus absent at anterior and dorsal sides, conspicuously present at ventral side (Fig. 3 A – D). Median and lateral antennae in straight line, palps inserted slightly more anteriorly (Fig. 3 A, B). Median antenna isolated by gap from lateral antennae and palps; longest prostomial appendage, as long as prostomium, folding back to posterior margin of second peristomium ring (mid second peristomial ring). Lateral antennae around two thirds length of prostomium, folding back to mid of second peristomium ring (posterior margin of first to anterior margin of second peristomium ring). Palps shortest, around as long as half of prostomium, folding back to mid of first peristomial ring. Antennal styles and palpostyle tapering (palpostyle digitiform), all irregularly articulated and sensory papillae distributed in rings, last visible only under SEM, under stereomicroscope prostomial appendages appearing wrinkled (Fig. 3 C, E). Antennophores and palpophores ring shaped. One pair of inconspicuous eyes present, in shape of dark line at base of palpophores (lateral antennae antennophores) (Fig. 3 D). Separation between peristomial rings distinct all around. Second peristomial ring about as long as 1 / 3 – 1 / 2 of complete peristomium. Peristomial ventro-lateral lips distinct laterally just as elevated surface. Peristomial anterior margin of dorsal and ventral sides in shallow arc, lateral margins longer; shortest on dorsal side (Fig. 3 D). Posterior end of muscularized pharynx at chaetiger 4. Mandibles missing calcareous cutting plates. MxI twice as long as carriers and six times longer than locking system. MxIII at least in part located ventral to MxII; attachment lamella upside down drop-shaped at anterior end of median margin of MxIII (Fig. 3 F). Left MxIV plate semi-circular; attachment lamella shorter than plate, almost as wide as plate, along most of anterior edge, missing on lateral edge of plate (Fig. 3 F). Right MxIV attachment lamella starting between lateralmost teeth 2 – 3, along around 2 / 3 of plate, longest at lateral side and evenly shorter towards opposite side (Fig. 3 F). Maxillary formula: I = 1 + 1, II = 5 (6) + 6, III = 6 (5) + 0, IV = 6 (7) + 9 (8), V = 1 + 1, VI absent. Pre-neurochaetal lobe shorter than neurochaetal lobe along whole body. Post-neurochaetal lobe longer than neurochaetal lobe in anterior chaetigers, decreasing in length along body, becoming as long as or shorter than neurochaetal lobe in median and posterior chaetigers. Anterior post-neurochaetal lobe wide, distally truncate with longer dorsal edge (Fig. 3 G, H). Neurochaetal lobe round in anteriormost region (P 1) (Fig. 3 I), tapering from parapodia P 2 – P 6 (Fig. 3 H, J). Anterior notopodial cirri fusiform (tapering), longer than neurochaetal lobe (Fig. 3 I); median and posterior cirri slender and slightly shorter than anterior ones, but still longer than neurochaetal lobes (Fig. 3 J) (shorter than neurochaetal lobe in branchiated region). Lateral sense organs slightly elevated in P 1, as conspicuously ciliated bump in remaining of body (Fig. 3 H). Ventral cirri tapering with round wide tips at chaetigers 1 to 4, around 2 / 3 as long as notopodial cirri (Fig. 3 I); basally inflated from chaetiger 5, inflated base of round shape with round tip (Fig. 3 J), gradually decreasing from chaetiger 61 (35, 43) to 152 (85); round (triangular) tapering with distinct tip from chaetiger 153 (86). Branchiae palmate (Fig. 3 J, K), with 2 (1) filaments from chaetiger 28 (21 – 26), reaching maximum of 6 (4, 5) filaments at chaetiger 129 (71 – 106; first 1 / 3 of body), terminating at last chaetiger (21 chaetigers before pygidium). Best developed branchial filaments around 4 times longer than notopodial cirri and 8 times longer than branchial stems, length of filaments decreasing at median region of body. Branchial stems distally loose, basally attached to notopodial cirri and body (Fig. 3 K). Notopodial aciculae yellow, present in notopodial cirri along whole body. Neurochaetae in two distinct bundles; supra-acicular with limbate chaetae and pectinate chaetae, subacicular with bidentate compound falciger chaetae and subacicular hooks. Neuroaciculae blunt to tapering, brown in anterior chaetigers, becoming yellow at posterior most chaetigers, dorsal to midline in P 1 and in midline thereafter; distributed in oblique row, anteriormost neuroaciculae also dorsalmost in parapodium (Fig. 3 L). Number of neuroaciculae, limbate chaetae and compound falciger chaetae decreasing towards posterior end. Three neuroaciculae present in P 1 and P 2, in posterior chaetiger mostly one present. Around 15 limbate chaetae at anteriormost region (P 1), 10 in P 2 and more posterior chaetigers with as few as 6. Limbate chaetae longer than all other chaetae, strongly serrated, serration increases towards flat distal end (spike like serration surround middle of its length). Compound falciger chaetae numerous in P 1, around 40 (30) present, 15 (10) by P 2 and 4 – 6 in more posterior parapodia; shaft and appendage of compound falciger chaetae strongly serrated (shaft serration spike like toward proximal end of chaetae); appendage bidentate with guards symmetrically blunt, marginally serrated, without mucros; both teeth directed laterally, distal tooth curved, proximal tooth straight, perpendicular to length of chaetae, about twice as long as distal tooth (Fig. 3 M – O). One – two curved thin pectinate chaetae present at anterior edge of supra-acicular bundle from P 1 – P 3; around 11 – 16 teeth present (Fig. 3 H, Q, R); inner teeth equal in length; outer teeth not equal in length and longer than inner teeth. Curved thick pectinate chaetae in median to posterior regions with around 12 – 14 coarse teeth with about same length at distal end but with increasingly deeper proximal ends from margins to center, teeth with deepest proximal end off center (Fig. 3 M, S); 2 – 3 present in P 4, 2 – 4 in P 5 and 1 – 2 in P 6. Pseudocompound chaetae and compound spiniger chaetae absent. Subacicular hook first present from chaetiger 37 (27 – 36), present in all chaetigers thereafter, always single; yellow, clearly bidentate, thinner than aciculae; distal tooth erect tapering directed distally and much smaller than proximal tooth; proximal tooth perpendicular to length of hook, directed laterally to distally, tip tapering to round; guards truncate present on both sides of teeth (Fig. 3 M, Q, T). Pygidium longer on ventral side, with two pairs of pygidial cirri attached to ventral margin; pygidial cirri triangular, about as long as pygidium, dorsal cirri as long as ventral (tapering, longer than pygidium, dorsal 3 times longer than ventral cirri in paratype AM W. 37089). Variation. Calcareous cutting plates of mandibles are probably dissolved due to formalin fixation. Thin pectinate is inconspicuous in P 1 under light microscopy. It was only clearly observed in the holotype, largest specimen. Spike-like serration present in some chaetae may be artifact of preparation. The appearance of pygidial cirri in the holotype is probably due to recent regeneration.	en	Zanol, Joana, Da Silva, Thauane Dos S. C., Hutchings, Pat (2017): One new species and two redescriptions of Marphysa (Eunicidae, Annelida) species of the Aenea-group from Australia. Zootaxa 4268 (3): 411-426, DOI: 10.11646/zootaxa.4268.3.6
CC4187C9716F3B32FF4D15FD239118FE.taxon	discussion	Remarks. Among the species of the Aenea-group, M. mortenseni, M. bifurcata, M. sessilobranchiata and Marphysa galluccii Orensanz, 1990 are the most similar to this new species in having a tapering non-bilobed prostomium. Marphysa mortenseni, M. bifurcata and M. galluccii are clearly distinct from M. pseudosessiloa n. sp. in having prostomial appendages shorter than half the length of the prostomium; the first two species may also have bifurcated notopodial cirri, which is simple in the current species. Marphysa galluccii also differs from the current species in lacking thin pectinate chaetae, having only single filament branchiae and unidentate subacicular hooks. General features are similar among the specimens examined here, paratypes and non-types of M. sessilobranchiata. The main divergences among them are the shape of the left MxIV plate, left and right MxIV attachment lamellae, teeth of subacicular hooks and position of branchial stem. In the current species branchiae are attached to a loose stem connected only basally to notopodial cirri and body wall, while in M. sessilobranchiata, as well as M. bifurcata, branchial stems are completely attached to the body wall, giving the appearance that branchial filaments are individually connected to the body wall as previously described. Molecular identification clearly distinguishes this species from other species of the genus (K 2 P = 18 – 25 %) and from M. bifurcata (K 2 P = 17 %), the only species of the Aenea-group included in the analyses (Fig. 4).	en	Zanol, Joana, Da Silva, Thauane Dos S. C., Hutchings, Pat (2017): One new species and two redescriptions of Marphysa (Eunicidae, Annelida) species of the Aenea-group from Australia. Zootaxa 4268 (3): 411-426, DOI: 10.11646/zootaxa.4268.3.6
CC4187C9716F3B32FF4D15FD239118FE.taxon	etymology	Etymology. It is very similar to M. sessilobranchiata in general morphology but it lacks the sessile branchiae and hence the specific name of M. pseudosessiloa.	en	Zanol, Joana, Da Silva, Thauane Dos S. C., Hutchings, Pat (2017): One new species and two redescriptions of Marphysa (Eunicidae, Annelida) species of the Aenea-group from Australia. Zootaxa 4268 (3): 411-426, DOI: 10.11646/zootaxa.4268.3.6
CC4187C9716F3B32FF4D15FD239118FE.taxon	materials_examined	Type locality. Australia, New South Wales, Pittwater, Careel Bay. Habitat. Intertidal mud in seagrass beds of Zostera capricorni.	en	Zanol, Joana, Da Silva, Thauane Dos S. C., Hutchings, Pat (2017): One new species and two redescriptions of Marphysa (Eunicidae, Annelida) species of the Aenea-group from Australia. Zootaxa 4268 (3): 411-426, DOI: 10.11646/zootaxa.4268.3.6
CC4187C9716F3B32FF4D15FD239118FE.taxon	distribution	Distribution. Only known from the type locality.	en	Zanol, Joana, Da Silva, Thauane Dos S. C., Hutchings, Pat (2017): One new species and two redescriptions of Marphysa (Eunicidae, Annelida) species of the Aenea-group from Australia. Zootaxa 4268 (3): 411-426, DOI: 10.11646/zootaxa.4268.3.6
CC4187C971633B37FF4D17BD22CE1A53.taxon	materials_examined	Material examined. PARATYPES: Australia, Western Australia, Princess Royal Harbor, Little Grove, 35 ° 04 ' S, 117 ° 52 ' E, 22 Nov 1975, AM W. 198420 (2 specimens). NON-TYPE: Western Australia, Albany, Princess Royal Harbor, Quaranup Jetty, 35 ° 03 ' S, 117 ° 55 ' E, Jan 1988, AM W. 20292 (1 specimen).	en	Zanol, Joana, Da Silva, Thauane Dos S. C., Hutchings, Pat (2017): One new species and two redescriptions of Marphysa (Eunicidae, Annelida) species of the Aenea-group from Australia. Zootaxa 4268 (3): 411-426, DOI: 10.11646/zootaxa.4268.3.6
CC4187C971633B37FF4D17BD22CE1A53.taxon	description	Measurements. Paratypes incomplete with 48 and 58 mm of total length, 147 and 157 chaetigers, 5 and 5.25 mm long to chaetiger 10, 3 mm with parapodia (1.6 mm without parapodia) and 3.5 mm with parapodia (1.76 mm without parapodia) wide at this chaetiger. Non-type, incomplete with 38 mm of total length, 123 chaetigers, 5.3 mm long to chaetiger 10 and 3 mm with parapodia (1.96 mm without parapodia) wide at this chaetiger. Description. Fixed specimens brown, lacking any color pattern. Body long, slender, median region widest; cross-section dorsally or dorsoventrally flattened at anterior region and rounded at remaining of body. Chaetigers around 5 times wider than long at widest region of body. Prostomium as long as or longer, narrower or as wide as and less than half as deep as peristomium (Fig. 5 A, B). Prostomium dorsally completely flat or with anterior end higher; anteriorly rounded to triangular; not bilobed; median sulcus absent at anterior and dorsal sides, conspicuously present at ventral side. Median and lateral antennae in straight line, palps slightly more anteriorly (Fig. 5 A). Median antenna isolated by gap from lateral antennae and palps; longest prostomial appendage, almost as long as to little longer than prostomium, folding back to first chaetiger. Lateral antennae around two thirds length of prostomium, folding back to mid or posterior margin of second peristomium ring. Palps shortest, little longer than half of prostomium, fold back to mid or posterior edge of first peristomial ring. Antennal styles tapering, palpostyle digitiform, all irregularly articulated, appearing wrinkled. Antennophores and palpophores ring shaped. Eyes absent. Separation between peristomial rings distinct all around. Second peristomial ring about as long as 1 / 3 – 1 / 2 of complete peristomium (Fig. 5 A, B). Peristomial ventro-lateral lips distinct laterally as elevated surface or by groove. Peristomial ventro-anterior margin straight or shallow arc, ventral side longer than dorsal. Posterior end of muscularized pharynx at chaetiger 4 – 5. Mandibles with small delicate calcareous cutting plates more than twice as long as sclerotized matrix; almost diamond shape with curved anterior inner edge, two anterior teeth. MxI about twice as long as carriers (Fig. 5 C) and six times longer than locking system. MxIII at least in part located ventral to MxII; attachment lamella oval at anterior end of median margin of MxIII (Fig. 5 D). Left MxIV plate semi-ellipse; attachment lamella shorter than plate, as wide as plate, along complete anterior and lateral edges of plate (Fig. 5 D). Right MxIV attachment lamella starting between lateralmost teeth 1 – 2, along around ¾ of plate, longest at lateral side (Fig. 5 E). Maxillary formula: I = 1 + 1, II = 5 + 6, III = 6 + 0, IV = 4 + 9, V = 1 + 1, VI absent. Pre-neurochaetal lobe shorter than neurochaetal lobe along whole body. Post-neurochaetal lobe longer than neurochaetal lobe in anterior chaetigers, decreasing in length along body, becoming as long as to shorter than neurochaetal lobe in median and posterior chaetigers. Anterior post-neurochaetal lobe wide, distally truncate with longer dorsal edge (Fig. 5 A, F). Neurochaetal lobe round at anterior region (Fig. 5 G, I), tapering in remaining of body (Fig. 5 J). Anterior notopodial cirri tapering, longer than neurochaetal lobe (Fig. 5 G, I); median and posterior cirri slender and shorter than neurochaetal lobes (Fig. 5 H). Lateral sense organ slightly elevated to conspicuously ciliated bump. Ventral cirri thumb shaped to tapering with round wide tips at chaetigers 1 to 4, around 2 / 3 to as long as notopodial cirri (Fig. 5 F); basally inflated from chaetiger 5, inflated base of round to fusiform shape with round tip (Fig. 5 G, I), gradually decreasing in size at median region of body. Branchiae palmate, with 1 – 2 filaments from chaetiger 25, reaching maximum of 4 filaments at chaetiger 70 – 99. Best developed branchial filaments around 3 – 5 times longer than notopodial cirri and 3 times longer than branchial stems. Branchial stems completely attached to body wall (Fig. 5 H, J). Notopodial aciculae yellow, present in notopodial cirri along whole body. Neurochaetae distributed in two distinct bundles, supra-acicular with limbate chaetae and pectinate chaetae, subacicular with bidentate compound falciger chaetae and subacicular hooks. Neuroaciculae blunt, brown; three at anteriormost parapodia, distributed in oblique row, anteriormost neuroaciculae also dorsalmost in parapodium (Fig. 5 G, K). Towards posterior end, number of neuroaciculae decreasing, mostly one present, as well as of limbate chaetae and compound falciger chaetae. Limbate chaetae longer than all other chaetae, serrated, spike-like serrations may surround middle of its length and tips. Shaft and appendage of compound falciger chaetae serrated; appendage bidentate with guards symmetrically blunt, without mucros; both teeth directed laterally; distal tooth curved; proximal tooth straight, perpendicular to length of chaetae, as long as to longer than distal tooth (Fig. 5 L – N). One – three curved thin pectinate chaetae present at anterior edge of supra-acicular bundle at anterior and median regions, around 9 – 13 long teeth present; inner teeth equal in length; both outer teeth longer than inner teeth, equal or unequal in length (Fig. 5 O). One – two slightly curved thick pectinate chaetae in postero-median region with around 11 – 13 coarse teeth, one outer tooth longer, inner teeth about same length at distal end but with increasingly deeper proximal ends from margins to center, teeth with deepest proximal end off center (Fig. 5 P). Pseudocompound chaetae and compound spiniger chaetae absent. Subacicular hook first present from chaetiger 35 – 38, present in all chaetigers thereafter, always single; yellow, unidentate to bidentate, thinner than aciculae. Bidentate subacicular hooks with small rounded to tapering teeth, both teeth erect (Fig. 5 Q, R). Pygidium missing. Variation. In all examined specimens, jaws are semi-everted, which usually make prostomium seem less deep than when jaws are fully withdrawn. Maxillae are described based on non-type specimen. Both paratypes have symmetric maxillae, the left side is copy of the right side (MxI 1 + 1, II = 7 + 7, III absent, IV = 10 + 10, V = 1 + 1, VI absent), which may be a malformation. Mandible calcareous cutting plates are missing, probably dissolved due to formalin fixation. Spike-like serration present in some chaetae may be artifact of preparation. Unidentate subacicular hooks are, probably, result of wear.	en	Zanol, Joana, Da Silva, Thauane Dos S. C., Hutchings, Pat (2017): One new species and two redescriptions of Marphysa (Eunicidae, Annelida) species of the Aenea-group from Australia. Zootaxa 4268 (3): 411-426, DOI: 10.11646/zootaxa.4268.3.6
CC4187C971633B37FF4D17BD22CE1A53.taxon	discussion	Remarks. General features are similar to the original description (Hartmann-Schröder 1984). The main divergences are the shape of the branchiae, number of teeth on MxIV and of pectinate chaetae. Branchiae are palmate with stem attached to the body wall. Branchial filaments are connected to the stems and not individually to the body wall as originally described. In the original description, number of teeth on left and right MxIV appears to be reversed. As in many Marphysa species, M. sessilobranchiata has thin pectinate at the anterior region and thick pectinate at median and posterior regions. In the original description only the thin pectinate were described. Attempts to recollect from the type locality for molecular studies were unsuccessful. Habitat. Fine sand close to Avicenia mangroves, covered with shell pieces or in seagrass bed of Posidonia sinuosa, intertidal to 3 m deep.	en	Zanol, Joana, Da Silva, Thauane Dos S. C., Hutchings, Pat (2017): One new species and two redescriptions of Marphysa (Eunicidae, Annelida) species of the Aenea-group from Australia. Zootaxa 4268 (3): 411-426, DOI: 10.11646/zootaxa.4268.3.6
CC4187C971633B37FF4D17BD22CE1A53.taxon	distribution	Distribution. Southern Western Australia: Princess Royal Harbor.	en	Zanol, Joana, Da Silva, Thauane Dos S. C., Hutchings, Pat (2017): One new species and two redescriptions of Marphysa (Eunicidae, Annelida) species of the Aenea-group from Australia. Zootaxa 4268 (3): 411-426, DOI: 10.11646/zootaxa.4268.3.6
