identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
CC3D87EEFFEEFFE8FED6FDA0FDDD2268.text	CC3D87EEFFEEFFE8FED6FDA0FDDD2268.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypostomus denticulatus Zawadzki & Weber & Pavanelli 2008	<div><p>Hypostomus denticulatus, new species</p><p>Fig. 1</p><p>Hypostomus sp. 7 .- Zawadzki, 2001: 21 [photo; rio Corumbá, affluent of rio Paranaíba; comparison to other species of Hypostomus; allozymes].- Pavanelli et al., 2007: 61 [rio Corumbá basin, affluent of rio Paranaíba].</p><p>Holotype. MZUSP 98770, 161.9 mm SL, Brazil, Goiás state, Município de Caldas Novas, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.497223&amp;materialsCitation.latitude=-17.573334" title="Search Plazi for locations around (long -48.497223/lat -17.573334)">rio do Peixe</a>, tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.497223&amp;materialsCitation.latitude=-17.573334" title="Search Plazi for locations around (long -48.497223/lat -17.573334)">rio Corumbá</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.497223&amp;materialsCitation.latitude=-17.573334" title="Search Plazi for locations around (long -48.497223/lat -17.573334)">rio Paranaíba</a> drainage, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.497223&amp;materialsCitation.latitude=-17.573334" title="Search Plazi for locations around (long -48.497223/lat -17.573334)">Pires do Rio</a>, 17°34’24”S, 48°29’50”W, 21 Sep 1996, Nupélia.</p><p>Paratypes. All from Brazil, Goiás State, upper <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Paraná basin</a>: ANSP 187499, 1, 165.7 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio do Peixe</a>, tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Corumbá</a>, Município de Caldas Novas, Pires do Rio, 17°36’06”S, 48°28’01”W, 8 Oct 1999, Nupélia; MHNG 2709.052, 1, 172.9 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Pirapitinga</a>, tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Corumbá</a>, Município de Caldas Novas, Pires do Rio, 17°43’37”S, 48°32’54”W, 6 Nov 1999, Nupélia; NUP 480, 3, 107.3- 147.1 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Corumbá</a> (Corumbá Reservoir), tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Paranaíba</a>, Município de Caldas Novas, Ipameri, 17°42’43”S, 48°29’52”W, 12 Nov 1996 to 11 May 1999, Nupélia; NUP 4233, 1, 165.8 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio do Peixe</a>, tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Corumbá</a>, Município de Caldas Novas, Pires do Rio, 17°34’24”S, 48°29’50”W, 5 Oct 1999, Nupélia; NUP 4306, 2, 144.3- 158.5 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Corumbá</a>, tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Paranaíba</a>, Município de Pires do Rio, Ipameri, 17°29’02”S, 48°22’13”W, 22 May 1996, Nupélia; NUP 4308, 2, 166.7- 186.4 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Corumbá</a> (downstream the Corumbá Reservoir), tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Paranaíba</a>, Município de Caldas Novas, Corumbaíba, 18°00’33”S, 48°32’19”W, 7 Feb 1999, Nupélia; NUP 5574, 2, 117.7- 136.9 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Corumbá</a> (Corumbá Reservoir), tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Paranaíba</a>, Município de Caldas Novas, Corumbaíba, 17°43’37”S, 48°32’54”W, 10 Nov 1996, Nupélia; NUP 5617, 2, 93.1-99.8 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio do Peixe</a>, tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Corumbá</a>, Município de Pires do Rio, Ipameri, 17°36’S, 48°24’”W, 28 Oct 1997, Nupélia; NUP 5619, 1, 134.5 mm SL, Corumbá Reservoir, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Corumbá</a>, tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Paranaíba</a>, Município de Caldas Novas, Corumbaíba, 17°43’37”S, 48°32’54”W, 15 Dec 1996, Nupélia; NUP 5620, 2, 182.6- 191.3 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Corumbá</a> (Corumbá Reservoir), tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Paranaíba</a>, Município de Caldas Novas, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">Ipameri</a>, 17°42’43”S, 48°29’52”W, Aug 1997, Nupélia; NUP 5635, 1, 156.6 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio do Peixe</a>, tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Corumbá</a>, Município de Pires do Rio, Ipameri, 17°36’06”S, 48°28’01”W, 20 May 1996, Nupélia; NUP 5637, 1, 120.1 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Corumbá</a>, tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Paranaíba</a>, Município de Pires do Rio, Ipameri, 17°29’02”S, 48°22’13”W, 6 Dec 1998, Nupélia. Non-measured paratypes: Goiás State: NUP 479, 6, 107.9- 132.4 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio do Peixe</a>, tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Corumbá</a>, Município de Pires do Rio, Ipameri, 17°36’S, 48°24’W, 12 Nov 1996, Nupélia; NUP 5571, 5, 90.0- 133.2 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio do Peixe</a>, tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Corumbá</a>, Município de Pires do Rio, Ipameri, 17°36’S, 48°24’W, 10 Nov 1996, Nupélia; NUP 5572, 1, 90.1 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio do Peixe</a>, tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Corumbá</a>, Município de Pires do Rio, Ipameri, 17°36’S, 48°24’W, 17 Oct 1996, Nupélia; NUP 5573, 1, 111.7 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio do Peixe</a>, tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Corumbá</a>, Município de Pires do Rio, Ipameri, 17°36’S, 48°24’W, 30 Jun 2001, Nupélia; NUP 5621, 1, 109.1 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Corumbá</a> (Corumbá Reservoir), tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Paranaíba</a>, Município de Caldas Novas, Ipameri, 17°43’37”S, 48°32’54”W, 21 Oct 1996, Nupélia; NUP 5622, 2, 65.0- 145.9 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Corumbá</a> (Corumbá Reservoir), tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Paranaíba</a>, Município de Caldas Novas, Ipameri, 17°43’37”S, 48°32’54”W, 11 Dec 1996, Nupélia. Minas Gerais State: MZUSP 36788, 2, 136.3- 154.4 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Paranaíba</a> (São Simão Reservoir), tributary to upper <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Paraná</a>, Município de Cachoeira Dourada, Oct 1985, A.C. Beaumord. Indefinite State: MNRJ 17624, 1, 147.0 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Preto</a>, corredeira, right margin of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Paranaíba</a>, city and State not specified on label, 3 Aug 1997, O. Sagim Jr .</p><p>Diagnosis. Hypostomus denticulatus is distinguished from all congeners, except H. multidens and H. ternetzi, by having more than 115 teeth (vs. less than 109) per ramus on dentary and premaxilla. Hypostomus denticulatus is distinguished from similar-looking H. ternetzi by having 116-214 teeth per jaw ramus (mean 167, n = 20) vs. 39-127 teeth per ramus (mean 81.4, n = 20); beige to light-brown body color vs. dark brown; larger orbital diameter, 18.5 to 21.2% of HL (n = 20) vs. 14.5 to 16.1% (n = 20); abdomen naked vs. abdomen partially plated; and teeth with two symmetrical cusps (Fig. 2a) vs. two asymmetrical cusps (Fig. 2b). Hypostomus denticulatus is distinguished from H. multidens by having dark (vs. pale) spots over body and fins (Fig. 3).</p><p>Description. Counts and measurements in Table 1. Body relatively short with rough plates. Cleithral width greater than head depth. Dorsal profile rising somewhat irregularly at an angle slightly less than 45 o from snout tip to dorsal-fin origin, and decreasing gently from that point to end of caudal peduncle. Head broad and moderately depressed, covered dorsally with dermal ossifications, except for small amorphous naked area on snout tip. Anterior profile of snout rounded in dorsal view. Median longitudinal bulge associated with mesethmoid usually conspicuous from snout tip to transverse line between nares. Eye large (18.5 to 21.2% of HL), dorsolaterally placed. Interorbital space moderately concave in frontal view. A pair of weak ridges on dorsal surface of head, each beginning lateral to naris, passing through upper margin of orbit and terminating on posterior portion of pteroticsupracleithrum. Supraoccipital with weak median ridge and slight, broadly triangular posterior process bordered by single predorsal plate, partially divided in some specimens. Predorsal region with weak medial keel; its uppermost portion flat.</p><p>Lips well developed and transversely ellipsoidal. Lower lip reaching or almost reaching transverse line between gill openings, its inner surface covered with numerous small papillae. Maxillary barbel short with free tip. Mouth large. Teeth long, very slender, bicuspid, and curved inward distally; mesial cusp similar in length and form to lateral cusp (Fig. 2a). Dentary teeth usually not overlapping medially. Dentary ramus flat, straight, and approximately forming a straight angle (usually ranging from 160° to 180°).</p><p>Five rows of slightly spinulose plates on side of body. Dorsal series of plates starting at vertical through dorsal-fin origin. A weak longitudinal keel begins on first plate of dorsal series and continues until dorsal procurrent caudal-fin plates. Mid-dorsal series of plates without keel or with a very weak keel. Median series bearing lateral line. Mid-ventral series forming longitudinal ridge along first five or six plates. Plates in ventral series right-angled transversely on caudal peduncle, forming a strong keel along lower portion of peduncle. Caudal peduncle roughly triangular in cross section.</p><p>Upper and lower lips without scutelets. Ventral surfaces of head and abdomen completely naked in specimens up to 131.2 mm SL; larger specimens with small platelets distributed just anterior to gill opening, in narrow transverse band between gill openings, laterally on abdomen between pectoral and pelvic fins (contacting ventral plates), and anterior to urogenital opening. Preanal plate partially exposed in specimens up to 129.8 mm SL, usually completely covered with skin in larger specimens.</p><p>Dorsal-fin origin situated two lateral-plate rows anterior to vertical through pelvic-fin origin. Dorsal fin large with flexible spine and seven branched rays; two posteriormost rays reaching or almost reaching adipose-fin origin. Adipose-fin spine straight and well developed. Pectoral fin with moderately inward-curved spine and six branched rays; distal margin straight. Pectoral-fin spine covered with poorly-developed odontodes; odontodes more developed distally in some larger specimens. Pectoral fin inserted on same horizontal plane as pelvic fin such that pectoral spine, when adpressed, lies on top of and reaches to approximately middle of pelvicfin spine. Pelvic fin with flexible and slightly inward-curved spine and five branched rays; distal border straight. Pelvicfin spine flattened ventrally; when adpressed spine tip reaches to about middle of anal-fin spine. Anal fin with flexible spine and four branched rays; spine tip reaching fifth plate posterior to its origin. Caudal fin slightly emarginate, with two longer outer spines and 14 inner branched rays; ventral lobe longer than dorsal lobe.</p><p>Color in alcohol. Background color of dorsal and lateral surfaces of head and body beige to light-brown. Head, dorsum, flanks and fins covered by many large dark spots. Dark spots generally merge into short and narrow vermiculations on anterior portion of head, becoming isolated and roundish on pterotic-supracleithrum and more elongate posteriorly. Spots most conspicuous on head and usually fade gradually on trunk towards caudal peduncle. Background color of ventral surfaces of head and body lighter than dorsal portions, except ventral portion of snout uniformly dark brown. Upper and lower lips and ventral surface of head without spots. Abdomen usually without spots or with very faded irregularly spaced dark spots in a few specimens. Ventral surface of caudal peduncle without spots.</p><p>Dorsal fin with roundish dark spots only on its base; spots merging distally into bands on interradial membranes paralleling rays. Adipose fin and caudal fin brown. Pectoral fin and pelvic fin with dark spots on spine, rays and interradial membranes. Anal fin with diffuse dark markings.</p><p>Distribution and habitat. Hypostomus denticulatus is known from the rio Corumbá in the vicinity of the Corumbá Reservoir, upper rio Paraná basin, Brazil (Fig. 4). This species was mostly collected in the reservoir of the rio Corumbá just upstream from the dam, and its tributary the rio Areia. Water was turbid, substrates rocky with sand, and riparian vegetation variable and remnant. This species was not found associated with woody debris, and occurs syntopically with H. ancistroides, H. iheringii, H. margaritifer, H. regani, Hypostomus sp., and the second new species described herein.</p><p>Etymology. From the Latin denticulus, meaning small teeth, plus - atus, meaning provided with.</p></div>	https://treatment.plazi.org/id/CC3D87EEFFEEFFE8FED6FDA0FDDD2268	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Zawadzki, Cláudio H.;Weber, Claude;Pavanelli, Carla S.	Zawadzki, Cláudio H., Weber, Claude, Pavanelli, Carla S. (2008): Two new species of Hypostomus Lacépède (Teleostei: Loricariidae) from the upper rio Paraná basin, Central Brazil. Neotropical Ichthyology 6 (3): 403-412, DOI: 10.1590/S1679-62252008000300013, URL: http://www.scielo.br/scielo.php?script=sci_arttext&pid=S1679-62252008000300013&lng=en&tlng=en
CC3D87EEFFEAFFEAFEC9FD7AFAE72548.text	CC3D87EEFFEAFFEAFEC9FD7AFAE72548.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypostomus heraldoi Zawadzki & Weber & Pavanelli 2008	<div><p>Hypostomus heraldoi, new species</p><p>Fig. 5</p><p>Hypostomus sp. 1.- Zawadzki, 2001: 61 [photo; rio Corumbá, affluent of rio Paranaíba; comparison to other species of Hypostomus; allozymes].- Pavanelli et al., 2007: 61 [rio Corumbá basin, affluent of rio Paranaíba].</p><p>Holotype. MZUSP 98771, 217.9 mm SL, Brazil, Goiás State, Município de Caldas Novas, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Pirapitinga</a>, tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Corumbá</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.548332&amp;materialsCitation.latitude=-17.726946" title="Search Plazi for locations around (long -48.548332/lat -17.726946)">rio Paranaíba</a> drainage, 17°43’37”S, 48°32’54”W, 10 Dec 1999, Nupélia.</p><p>Paratypes. All from Brazil, upper <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Paraná basin</a>: Goiás State: ANSP 187500, 1, 152.6 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Corumbá</a> (Corumbá Reservoir), tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Paranaíba</a>, Município de Caldas Novas, Corumbaíba, 17°58’25”S, 48°31’04”W, 17 Oct 1996, Nupélia; MHNG 2709.051, 1, 140.9 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Corumbá</a> (Corumbá Reservoir), tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Paranaíba</a> basin, Município de Caldas Novas, Corumbaíba, 17°50’51”S, 48°31’54”W, 11 Nov 1996, Nupélia; NUP 2280, 3, 178.4- 194.6 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Pirapitinga</a>, tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Corumbá</a>, Município de Caldas Novas, Pires do Rio, 17°43’37”S, 48°32’54”W, 9 Abr 1999, Nupélia; NUP 2281, 1, 200.8 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Corumbá</a> (Corumbá Reservoir), tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Paranaíba</a>, Município de Caldas Novas, Ipameri, 17°43’37”S, 48°32’54”W, 9 Apr 1999, Nupélia; NUP 4042, 1, 141.8 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio do Peixe</a>, tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Corumbá</a>, Município de Caldas Novas, Pires do Rio, 17°34’24”S, 48°29’50”W, 20 May 1996, Nupélia; NUP 4048, 2, 157.2- 189.3 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Pirapitinga</a>, tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Corumbá</a>, Município de Caldas Novas, Pires do Rio, 17°43’37”S, 48°32’54”W, 9 Dec 1999, Nupélia; NUP 4051, 1, 138.3 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Corumbá</a> (Corumbá Reservoir), tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Paranaíba</a>, Município de Caldas Novas, Ipameri, 17°42’43”S, 48°29’52”W, 25 Dec 1997, Nupélia; NUP 5623, 5, 138.2- 183.1 mm SL (3, 174.1- 176.6 mm SL), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Pirapitinga</a>, tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Corumbá</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Paranaíba</a> drainage, upper <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Paraná basin</a>, Município de Caldas Novas, 17°43’37”S, 48°32’54”W, 12 Jan 2000, Nupélia; NUP 5624, 5, 133.2- 207.6 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Pirapitinga</a>, tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Corumbá</a>, Município de Caldas Novas, Pires do Rio, 17°43’37”S, 48°32’54”W, 25 Jan 1998, Nupélia; NUP 5625, 4, 138.4- 163.6 mm SL (1, 160.1 mm SL), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Pirapitinga</a>, tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Corumbá</a>, Município de Caldas Novas, Pires do Rio, 17°43’37”S, 48°32’54”W, 24 Oct 1997, Nupélia. Non-measured paratypes: Goiás State: NUP 2282, 6, 70.6-192.8 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Corumbá</a>, tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Paranaíba</a>, Município de Caldas Novas, Pires do Rio, 17°43’37”S, 48°32’54”W, 9 Apr 1999, Nupélia; NUP 5648, 3, 162.4- 189.4 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Corumbá</a>, tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Paranaíba</a>, Município de Caldas Novas, Pires do Rio, 17°43’37”S, 48°32’54”W, 20 Sep 1996, Nupélia; NUP 5650, 1, 146.4 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Corumbá</a>, tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Paranaíba</a>, Município de Caldas Novas, Pires do Rio, 17°43’37”S, 48°32’54”W, 21 Jun 1997, Nupélia; NUP 5651, 1, 112.3 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Corumbá</a>, tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Paranaíba</a>, Município de Caldas Novas, Pires do Rio, 17°43’37”S, 48°32’54”W, 11 Nov 1996. NUP 6049, 5, 36.5-104.0 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">córrego Gameleira</a>, tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Corumbá</a>, Município de Corumbaíba, 17°59’05”S, 48°29’46”W, 17 Jul 2008, C.H. Zawadzki, B.F. Morales &amp; V.S. Ferreira; NUP 6050, 1, 75.0 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">córrego Libório</a>, tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Corumbá</a>, Município de Corumbaíba, 17°55’48”S, 48°28’57”W, 17 Jul 2008, C.H. Zawadzki, B.F. Morales &amp; V.S. Ferreira. Minas Gerais State: MZUSP 73222, 2, 79.0-84.0 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Piedade</a> (Fazenda Piedade), tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Grande</a>, Município de Centralina, 18°38’11”S, 49°05’28”W, 2 Aug 2001, P. Gerhard &amp; F.C.T. Lima; MZUSP 73229, 2, 226.1- 236.1 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Piedade</a> (Fazenda Piedade), tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Grande</a>, Município de Centralina, 18°38’11”S, 49°05’28”W, 3 Aug 2001, P. Gerhard &amp; F.C.T. Lima; NUP 5696, 1, 115.5 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Muzambinho</a>, tributary to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Grande</a>, Município de Poços de Caldas, 21°44’50”S, 46°28’07”W, 24 Nov 2007, L.H.G. Pereira; NUP 5697, 1, 198.8 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Grande</a> (Funil Reservoir), tributary to upper <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.016666&amp;materialsCitation.latitude=-21.133333" title="Search Plazi for locations around (long -45.016666/lat -21.133333)">rio Paraná</a>, Município de Perdões, 21°08’S, 45°01’W, 26 Sep 2002, UFLA collectors .</p><p>Diagnosis. Hypostomus heraldoi is distinguished from all congeners, except H. albopunctatus, by having the pectoralfin spine length equal to or smaller than the pelvic-fin spine. From H. albopunctatus, it is distinguished by having roundish dark (vs. pale) spots on body and fins, and generally with more premaxillary teeth, 39-59 per ramus (vs. 32-42; see Table 2).</p><p>Description. Counts and measurements in Table 1. Body relatively short with rough plates. Cleithral width greater than head depth. Dorsal profile rising convexly at an angle of approximately 30° from snout tip to dorsal-fin origin, then decreasing linearly to end of caudal peduncle. Head broad and depressed, dorsally covered with dermal ossifications, except for small naked area on snout tip. Anterior profile of snout rounded in dorsal view. Median longitudinal bulge associated with mesethmoid usually conspicuous from snout tip to transverse line between nares. Eye small (12.0 to 13.7% of HL), dorsolaterally placed. Interorbital space straight or slightly convex in frontal view. A pair of weak ridges on dorsal surface of head, each one beginning lateral to naris, passing through upper margin of orbit and finishing on central portion of pterotic-supracleithrum. Supraoccipital bone without median ridge, and with short and rounded posterior process bordered by partially divided predorsal plate.</p><p>Lips small and transversely ellipsoid. Lower lip not reaching transverse line between gill openings, its inner surface covered with numerous small papillae. Maxillary barbel very short, largely coalesced with lower lip but with free tip. Mouth moderate. Teeth medium sized, robust, bicuspid, and curved inward distally; inner cusp with elongated crown, about twice length of lateral cusp. Dentary teeth usually not overlapping medially. Dentary ramus forms angle of approximately 140°.</p><p>Five rows of slightly spinulose plates on side of body. Predorsal region and all plates in lateral series usually without conspicuous keels. Dorsal series of plates starting at vertical through dorsal-fin origin. Mid-dorsal series interrupted by first plate of dorsal series. Median series bearing lateral line; interrupted by ventrally oriented expansions of plates in mid-dorsal series in region just below dorsal-fin base. Midventral series without keel or with very weak keel restricted to first three plates. Plates in ventral series weakly angled transversely on caudal peduncle; angle more pronounced posteriorly. Caudal peduncle roughly triangular in cross section.</p><p>Ventral surface of head mostly covered with minute platelets except beneath lower lip and along outer edge of upper lip. Abdomen covered with minute platelets, even in smaller specimens, except for small naked areas near pectoral and pelvic-fin insertions and surrounding urogenital opening in some specimens. Preanal plate partially exposed.</p><p>Dorsal-fin origin situated two lateral plates anterior to vertical through pelvic-fin origin. Dorsal fin small with flexible spine and seven branched rays; two posteriormost rays touching second plate anterior to azygous pre-adipose plate. Adipose-fin spine well developed in most specimens, straight or slightly curved inward. Pectoral fin with slightly inwardcurved spine and six branched rays; posterior margin straight. Pectoral-fin spine covered with poorly-developed odontodes; odontodes more developed distally in larger specimens. Pectoral fin inserted on same horizontal plane as pelvic fin such that pectoral spine, when adpressed, lies on top of and reaches between basal one-fourth to one-third of pelvic-fin spine. Pelvic fin with flexible and slightly inward-curved spine and five branched rays; distal margin straight. Pelvic-fin spine slightly flattened ventrally; when adpressed, tip reaches to distal one-third of anal-fin spine. Anal fin with flexible spine and four branched rays; spine tip reachs to sixth plate posterior to its origin. Caudal fin slightly forked, with two outer spines and 14 inner branched rays; ventral lobe longer than dorsal lobe.</p><p>Color in alcohol. Background color of dorsal and lateral surfaces of head and trunk brown and covered by darker spots. Spots closely to moderately spaced, and slightly faded in some specimens. Dark spots smaller, more compact and numerous on head, becoming slightly larger, and more dispersed posteriorly. Background color of ventral surfaces of head and body lighter than superior portions. Upper lip uniformly dark except for lighter border. Spots on ventral surface of head and body present but vary in conspicuousness among specimens. Ventral spots roundish and isolated in some specimens; spots merged into vermiculations in others. Caudal peduncle region with very faded dark spots ventrally.</p><p>Dorsal, pectoral, pelvic and caudal fins with roundish dark spots on spines and rays; spots weakly aligned to form five to seven transverse bands. Dorsal fin sometimes with dark spots merging to form irregular bands on interradial membranes paralleling rays. Adipose fin spotless or with irregular dark spots. Anal fin with dark spots generally merged into two or three transverse bands.</p><p>Distribution and habitat. Hypostomus heraldoi is known from the rio Corumbá and rio Grande drainages, upper rio Paraná basin (Fig. 4). The Corumbá Reservoir and its tributary rio Pirapitinga are the main sites where H. heraldoi was collected. The water was turbid, substrates rocky with sand, and riparian vegetation variable and remnant. This species was not found associated with woody debris, and, like H. denticulatus, occurs syntopically with H. ancistroides, H. iheringii, H. margaritifer, H. regani and Hypostomus sp.</p><p>Etymology. Hypostomus heraldoi is named in honor of Heraldo Antônio Britski, MZUSP (Museu de Zoologia da Universidade de São Paulo), for his extensive contributions to our knowledge of neotropical fish diversity.</p></div>	https://treatment.plazi.org/id/CC3D87EEFFEAFFEAFEC9FD7AFAE72548	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Zawadzki, Cláudio H.;Weber, Claude;Pavanelli, Carla S.	Zawadzki, Cláudio H., Weber, Claude, Pavanelli, Carla S. (2008): Two new species of Hypostomus Lacépède (Teleostei: Loricariidae) from the upper rio Paraná basin, Central Brazil. Neotropical Ichthyology 6 (3): 403-412, DOI: 10.1590/S1679-62252008000300013, URL: http://www.scielo.br/scielo.php?script=sci_arttext&pid=S1679-62252008000300013&lng=en&tlng=en
