identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
BF7F5914F919CFBE3752226F5E743EB8.text	BF7F5914F919CFBE3752226F5E743EB8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eublaberus distanti (Kirby 1903) Kirby 1903	<div><p>Taxon classification Animalia Blattodea Blaberidae</p><p>Eublaberus distanti (Kirby, 1903)</p><p>Materials.</p><p>Adult ♂.</p><p>Voucher number: DEKBO0843.</p><p>Collection locale. Karanambu Ranch, Rupununi, Guyana.</p><p>GPS: 3°45'2.2"N, 59°18'31.2"W.</p><p>Date: 7 - June - 2013.</p><p>Collectors. Dominic A. Evangelista, Oswin Ambrose, Susan George, and Megan M. Wilson.</p><p>Collection/ecological information.</p><p>This specimen was collected in the bathroom of one of the cabins at the camp of Karanambu Ranch.</p><p>Known geographic distribution.</p><p>Guatemala, Costa Rica, Panama, Colombia, Trinidad and Tobago, French Guiana, Suriname, Guyana and Brazil</p></div>	https://treatment.plazi.org/id/BF7F5914F919CFBE3752226F5E743EB8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Evangelista, Dominic A.;Chan, Kimberly;Kaplan, Kayla L.;Wilson, Megan M.;Ware, Jessica L.	Evangelista, Dominic A., Chan, Kimberly, Kaplan, Kayla L., Wilson, Megan M., Ware, Jessica L. (2015): The Blattodeas. s. (Insecta, Dictyoptera) of the Guiana Shield. ZooKeys 475: 37-87, DOI: http://dx.doi.org/10.3897/zookeys.475.7877, URL: http://dx.doi.org/10.3897/zookeys.475.7877
107EC6F18790057E9EE0C2DBFF5B0135.text	107EC6F18790057E9EE0C2DBFF5B0135.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neorhicnoda maronensis (Hebard 1921) Hebard 1921	<div><p>Taxon classification Animalia Blattodea Blaberidae</p><p>Neorhicnoda maronensis (Hebard, 1921)</p><p>Materials.</p><p>Adult ♂ Figure 1.</p><p>Voucher number: DECBA0615.</p><p>GenBank accession number: KF155090.</p><p>Collection locale. CEIBA Biological Station, Madewini, Guyana.</p><p>GPS: 6°29'N, 58°13'W.</p><p>Date: 02 - January - 2012.</p><p>Collectors. Dominic A. Evangelista, Ian Biazzo, Joseph A. Evangelista, Paul Frandsen, William R. Kuhn, and Jessica L. Ware.</p><p>Collection/ecological information.</p><p>This specimen was caught in a pitfall trap baited with beer in an uplands secondary forest.</p><p>Morphological identification.</p><p>This specimen agrees with the description of the male genitalia in Grandcolas (1992b).</p><p>Known geographic distribution.</p><p>Guyana (new record), Suriname, and French Guiana.</p></div>	https://treatment.plazi.org/id/107EC6F18790057E9EE0C2DBFF5B0135	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Evangelista, Dominic A.;Chan, Kimberly;Kaplan, Kayla L.;Wilson, Megan M.;Ware, Jessica L.	Evangelista, Dominic A., Chan, Kimberly, Kaplan, Kayla L., Wilson, Megan M., Ware, Jessica L. (2015): The Blattodeas. s. (Insecta, Dictyoptera) of the Guiana Shield. ZooKeys 475: 37-87, DOI: http://dx.doi.org/10.3897/zookeys.475.7877, URL: http://dx.doi.org/10.3897/zookeys.475.7877
D29F1557D625A3B82218338D6291065C.text	D29F1557D625A3B82218338D6291065C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Colapteroblatta surinama (Saussure 1868) Saussure 1868	<div><p>Taxon classification Animalia Blattodea Blaberidae</p><p>Colapteroblatta surinama (Saussure, 1868)</p><p>Materials.</p><p>Adult ♂ Figure 2E.</p><p>Voucher number: DECBA0703.</p><p>GenBank accession number: KF155029.</p><p>Collection locale. CEIBA Biological Station, Madewini, Guyana.</p><p>GPS: 6°29'N, 58°13'W.</p><p>Date: 05 - August - 2011.</p><p>Collectors. Dominic A. Evangelista, Ian Biazzo, Manpreet K. Kohli, Melissa Sanchez-Herrera, Nicole Sroczinski, and Jessica L. Ware.</p><p>Collection/ecological information.</p><p>This specimen was collected in an uplands secondary forest from within a rotting vine.</p><p>Morphological identification.</p><p>This specimen was identified using Roth and Gutierrez (1998).</p><p>Adult ♀ Figure 2D.</p><p>Voucher number: DECBA1810.</p><p>GenBank accession number: KF155126.</p><p>Collection locale. CEIBA Biological Station, Madewini, Guyana.</p><p>GPS: 6°29'N, 58°13'W.</p><p>Date: 20 - August - 2011.</p><p>Collectors. Dominic Evangelista and William R. Kuhn.</p><p>Collection/ecological information.</p><p>This specimen was collected in an uplands secondary forest from within an arboreal bromeliad.</p><p>Morphological identification.</p><p>This specimen was identified using Roth and Gutierrez (1998).</p><p>Juvenile Figure 2 A–C .</p><p>Voucher number: DECBA1811.</p><p>GenBank accession number: KF155112.</p><p>Collection locale. CEIBA Biological Station, Madewini, Guyana.</p><p>GPS: 6°29'N, 58°13'W.</p><p>Date: 17 - August - 2013.</p><p>Collectors. Dominic Evangelista and William R. Kuhn.</p><p>Collection/ecological information.</p><p>This specimen was collected on vegetation in an uplands secondary forest.</p><p>Morphological identification.</p><p>This specimen was associated to its adult morph using barcodes in Evangelista et al. (2014). The overall coloration of the juvenile specimens of this species is more similar to that of Colapteroblatta darlingtoni Roth &amp; Gutiérrez, 1998 and Colapteroblatta rehni Roth &amp; Gutiérrez, 1998 than to that of the adults of its own species (see Figure 2).</p><p>Genetic information and evolutionary placement. All three specimens have nearly identical cytochrome oxidase I (COI) haplotypes but their position could not be determined relative to other cockroach species with the data evaluated by Evangelista et al. (2014).</p><p>Known geographic distribution.</p><p>Guyana, Suriname.</p></div>	https://treatment.plazi.org/id/D29F1557D625A3B82218338D6291065C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Evangelista, Dominic A.;Chan, Kimberly;Kaplan, Kayla L.;Wilson, Megan M.;Ware, Jessica L.	Evangelista, Dominic A., Chan, Kimberly, Kaplan, Kayla L., Wilson, Megan M., Ware, Jessica L. (2015): The Blattodeas. s. (Insecta, Dictyoptera) of the Guiana Shield. ZooKeys 475: 37-87, DOI: http://dx.doi.org/10.3897/zookeys.475.7877, URL: http://dx.doi.org/10.3897/zookeys.475.7877
38447DFDB65DA661E898C1F58D0C5B64.text	38447DFDB65DA661E898C1F58D0C5B64.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epilampra opaca Walker 1868	<div><p>Taxon classification Animalia Blattodea Blaberidae</p><p>Epilampra opaca Walker, 1868</p><p>Materials.</p><p>Adult ♂ Figure 3B.</p><p>Voucher number: DECBA1845.</p><p>GenBank accession number: KF155125.</p><p>Collection locale. CEIBA Biological Station, Madewini, Guyana.</p><p>GPS: 6°29'N, 58°13'W.</p><p>Date: 18 - August - 2012.</p><p>Collectors. Dominic A. Evangelista and William R. Kuhn.</p><p>Adult ♀</p><p>Voucher number: DECBA1847.</p><p>GenBank accession number: KF155124.</p><p>Collection locale. CEIBA Biological Station, Madewini, Guyana.</p><p>GPS: 6°29"N, 58°13"W.</p><p>Date: 5 - August - 2011.</p><p>Collectors. Dominic A. Evangelista, Ian Biazzo, Manpreet K. Kohli, Melissa Sanchez-Herrera, Nicole Sroczinski and Jessica L. Ware.</p><p>Collection/ecological information.</p><p>The adult male (DECBA1845) was collected at a light trap. Adult female (DECBA1847) was collected by hand in the leaf litter by a small pond. Most late instar individuals of this species were also collected at the edge of this pond and some were collected in pitfall traps baited with beer. Early instar individuals of this species were collected from within bromeliads.</p><p>Genetic information.</p><p>The two adult specimens reported here, as well as three juvenile individuals (Voucher and accession numbers: DEDSM0141 - KF155097, DECBA1706 - KF155089, DECBA0205 - KF155088) have identical COI barcodes and are sister to each other on the tree. However, other individuals (similar to Epilampra opaca) included in the analysis (Voucher and accession numbers: DECBA0214 - KF155018, DECBA0216 - KF155017, DECBA0606 - KF155013, DECBA1101 - KF155016, DECBA0605 - KF155012, DECBA0608 - KF155015) are more genetically diverse and are only supported as monophyletic by 63% bootstrap support.</p><p>Morphological identification.</p><p>There is a great deal of intraspecific variation in the morphology of this species. Early instar nymphs are difficult to associate to later instar nymphs, all of which are entirely unrecognizable from the adults (Figure 3 A–C). Furthermore, there is variation within instars, where some later instar nymphs will appear to have a medially divided subgenital plate and others do not. This trait was not found to correlate with genetic differences (Evangelista et al. 2014).</p><p>The external morphology of this species provides little assistance in its identification, as most descriptions of it emphasize coloration that is both subtle and variable. However, the allometry of our specimens (Table 2) agree with those of Bruijning (1959). A definitive identification was made by comparison of genital morphology using Roth (1970b), particularly in the shape of the prepuce.</p><p>Known geographic distribution.</p><p>Venezuela (unverified), Guyana, Suriname, French Guiana and Brazil</p><p>History and synonymy. Walker (1868) first described both Epilampra opaca Walker, 1868 and Epilampra substrigata Walker, 1868. Hebard (1926) noted that Epilampra opaca Walker, 1868 has a highly variable morphology and may be synonymous with a few other Epilampra (e.g. Epilampra conferta Walker, 1868 syn. stigmosa Giglio-Tos, 1898, Epilampra maculicollis (Serville, 1838)). This variability is evident in the work published by Roth (1970b), which shows a great deal of variation in the genital morphology, in particular for L2d. Although it is not clear if anyone before Roth (1970b) examined the genitalia of these two species, both Shelford (1910) and Princis (1963) considered them to be synonyms. Roth’s (1970b) photos show that, although each species is intraspecifically variable, both are distinct and separable by the shape of L2d and the prepuce. Roth himself acknowledged this and considered the species as being separate. Although we have not examined any Epilampra substrigata Walker, 1868, we agree with Roth’s interpretation of the morphology and follow from his precedence in considering these separate (see Roth 1970b for the opinions of Princis and Gurney on the status of these two species).</p></div>	https://treatment.plazi.org/id/38447DFDB65DA661E898C1F58D0C5B64	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Evangelista, Dominic A.;Chan, Kimberly;Kaplan, Kayla L.;Wilson, Megan M.;Ware, Jessica L.	Evangelista, Dominic A., Chan, Kimberly, Kaplan, Kayla L., Wilson, Megan M., Ware, Jessica L. (2015): The Blattodeas. s. (Insecta, Dictyoptera) of the Guiana Shield. ZooKeys 475: 37-87, DOI: http://dx.doi.org/10.3897/zookeys.475.7877, URL: http://dx.doi.org/10.3897/zookeys.475.7877
09C6F4763C53F1734B7F607A9F54CD6B.text	09C6F4763C53F1734B7F607A9F54CD6B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epilampra sodalis Walker 1868	<div><p>Taxon classification Animalia Blattodea Blaberidae</p><p>Epilampra sodalis Walker, 1868</p><p>Materials.</p><p>Adult ♂ Figure 4A.</p><p>Voucher number: DECBA0401.</p><p>GenBank accession number: KF155063.</p><p>Collection locale. CEIBA Biological Station, Madewini, Guyana.</p><p>GPS: 6°29'N, 58°13'W.</p><p>Date: 31 - July to 6 - August - 2011.</p><p>Collectors. Dominic A. Evangelista, Ian Biazzo, Manpreet K. Kohli, Melissa Sanchez-Herrera, Nicole Sroczinski, and Jessica L. Ware.</p><p>Collection/ecological information.</p><p>This specimen was collected at a light trap.</p><p>Morphological identification.</p><p>This specimen agrees with the description the synonym Epilampra cinnamomea (Hebard 1926).</p><p>Juvenile</p><p>Voucher number: DECBA1702.</p><p>GenBank accession number: KF155068.</p><p>Collection locale. CEIBA Biological Station, Madewini, Guyana.</p><p>GPS: 6°29'N, 58°13'W.</p><p>Date: 27 - December - 2011.</p><p>Collectors. Dominic A. Evangelista, Ian Biazzo, Joseph A. Evangelista, Paul Frandsen, William R. Kuhn and Jessica L. Ware.</p><p>Juvenile</p><p>Voucher number: DECBA1701.</p><p>GenBank accession number: KF155069.</p><p>Collection locale. CEIBA Biological Station, Madewini, Guyana.</p><p>GPS: 6°29'N, 58°13'W.</p><p>Date: 10 - January - 2012.</p><p>Collectors. Dominic A. Evangelista, Ian Biazzo, Joseph A. Evangelista, Paul Frandsen, William R. Kuhn and Jessica L. Ware.</p><p>Collection/ecological information.</p><p>Both of these juvenile specimens were collected at the edge of a small pond.</p><p>Genetic information and evolutionary placement. These three specimens (previous reported as " Blaberidae sp. 04") were placed in the same clade with 90% bootstrap support.</p><p>Known geographic distribution.</p><p>Venezuela, Guyana (new record), Suriname, French Guiana and Brazil</p></div>	https://treatment.plazi.org/id/09C6F4763C53F1734B7F607A9F54CD6B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Evangelista, Dominic A.;Chan, Kimberly;Kaplan, Kayla L.;Wilson, Megan M.;Ware, Jessica L.	Evangelista, Dominic A., Chan, Kimberly, Kaplan, Kayla L., Wilson, Megan M., Ware, Jessica L. (2015): The Blattodeas. s. (Insecta, Dictyoptera) of the Guiana Shield. ZooKeys 475: 37-87, DOI: http://dx.doi.org/10.3897/zookeys.475.7877, URL: http://dx.doi.org/10.3897/zookeys.475.7877
A41F64DED03669F361577A4153DACE2F.text	A41F64DED03669F361577A4153DACE2F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thanatophyllum akinetum Grandcolas 1991	<div><p>Taxon classification Animalia Blattodea Blaberidae</p><p>Thanatophyllum akinetum Grandcolas, 1991</p><p>Materials.</p><p>Adult ♂ Figure 5.</p><p>Voucher number: DECBA0611.</p><p>GenBank accession number: KF155066.</p><p>Collection locale. CEIBA Biological Station, Madewini, Guyana.</p><p>GPS: 6°29'N, 58°13'W.</p><p>Date: 28 - December - 2011.</p><p>Collectors. Dominic A. Evangelista, Ian Biazzo, Joseph A. Evangelista, Paul Frandsen, William R. Kuhn and Jessica L. Ware.</p><p>Collection/ecological information.</p><p>This specimen was collected by hand on vegetation in an uplands secondary forest.</p><p>Morphological identification.</p><p>This specimen agrees with the description of the head and male genitalia of Grandcolas (1990).</p><p>Known geographic distribution.</p><p>Guyana (new record) and French Guiana.</p></div>	https://treatment.plazi.org/id/A41F64DED03669F361577A4153DACE2F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Evangelista, Dominic A.;Chan, Kimberly;Kaplan, Kayla L.;Wilson, Megan M.;Ware, Jessica L.	Evangelista, Dominic A., Chan, Kimberly, Kaplan, Kayla L., Wilson, Megan M., Ware, Jessica L. (2015): The Blattodeas. s. (Insecta, Dictyoptera) of the Guiana Shield. ZooKeys 475: 37-87, DOI: http://dx.doi.org/10.3897/zookeys.475.7877, URL: http://dx.doi.org/10.3897/zookeys.475.7877
6BE9CB5A9C946BF4A895692964D17EB6.text	6BE9CB5A9C946BF4A895692964D17EB6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anaplecta parviceps (Walker 1868) Walker 1868	<div><p>Taxon classification Animalia Blattodea Ectobiidae</p><p>Anaplecta parviceps (Walker, 1868)</p><p>Materials.</p><p>Adult ♂ Figure 6.</p><p>Voucher number: DECBA1843.</p><p>GenBank accession number: KF155137.</p><p>Collection locale. CEIBA Biological Station, Madewini, Guyana.</p><p>GPS: 6°29'N, 58°13'W.</p><p>Date: 16 - August - 2012.</p><p>Collectors . Dominic A. Evangelista and William R. Kuhn.</p><p>Collection/ecological information.</p><p>This specimen and another adult male (Voucher number: DECBA1841) were collected at a light trap near the camp of CEIBA Biological Station on the date noted above. A juvenile of this species was also collected at the same locale, found crawling through a benab between 21 and 24 of August 2012 (Voucher number: DECBA1842).</p><p>Morphological identification.</p><p>The specimen agrees with the description of the synonym Anaplecta insignis of Hebard (1926). Other specimens were identified by comparison with specimen DECBA1843.</p><p>Genetic information and evolutionary placement. The COI barcodes of this specimen (previously reported as " Blattodea sp. 18") falls sister to another specimen identified as Anaplecta sp. (previously reported as " Ectobiidae sp. 04"; Voucher number: DEDSM0111; GenBank accession number: KF155041) but with 25% bootstrap support. This other species is not reported in this paper due to an uncertainty in specific identification.</p><p>Known geographic distribution.</p><p>Guyana, Suriname, French Guiana, Brazil (Rio de Janeiro), Brazil ( Pará), and Brazil ( Amapá).</p></div>	https://treatment.plazi.org/id/6BE9CB5A9C946BF4A895692964D17EB6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Evangelista, Dominic A.;Chan, Kimberly;Kaplan, Kayla L.;Wilson, Megan M.;Ware, Jessica L.	Evangelista, Dominic A., Chan, Kimberly, Kaplan, Kayla L., Wilson, Megan M., Ware, Jessica L. (2015): The Blattodeas. s. (Insecta, Dictyoptera) of the Guiana Shield. ZooKeys 475: 37-87, DOI: http://dx.doi.org/10.3897/zookeys.475.7877, URL: http://dx.doi.org/10.3897/zookeys.475.7877
16A283D1EAD00D121B2AFB19D1B42A69.text	16A283D1EAD00D121B2AFB19D1B42A69.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anisopygia decora Hebard 1926	<div><p>Taxon classification Animalia Blattodea Ectobiidae</p><p>Anisopygia decora Hebard, 1926</p><p>Materials.</p><p>Adult ♀ Figure 7.</p><p>Voucher number: DEKBO0504.</p><p>Collection locale. Capuchin Trail, Karanambo Ranch, Rupununi, Guyana.</p><p>GPS: 3°44'43.70"N, 59°18'51.88"W.</p><p>Date: 10 - June - 2013.</p><p>Collectors. Dominic A. Evangelista, Oswin Ambrose, Susan George, and Megan M. Wilson.</p><p>Collection/ecological information.</p><p>This specimen was collected by hand in an undisturbed forested area. This is the first record of this specimen from Guyana.</p><p>Morphological identification.</p><p>This specimen was identified by comparison with Hebard’s description (1926).</p><p>Known geographic distribution.</p><p>Guyana (new record) and French Guiana.</p></div>	https://treatment.plazi.org/id/16A283D1EAD00D121B2AFB19D1B42A69	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Evangelista, Dominic A.;Chan, Kimberly;Kaplan, Kayla L.;Wilson, Megan M.;Ware, Jessica L.	Evangelista, Dominic A., Chan, Kimberly, Kaplan, Kayla L., Wilson, Megan M., Ware, Jessica L. (2015): The Blattodeas. s. (Insecta, Dictyoptera) of the Guiana Shield. ZooKeys 475: 37-87, DOI: http://dx.doi.org/10.3897/zookeys.475.7877, URL: http://dx.doi.org/10.3897/zookeys.475.7877
50AFE6E5CB1C6C45F763B02274361024.text	50AFE6E5CB1C6C45F763B02274361024.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ischnoptera atrata Hebard 1916	<div><p>Taxon classification Animalia Blattodea Ectobiidae</p><p>Ischnoptera atrata Hebard, 1916</p><p>Materials.</p><p>Adult ♂.</p><p>Voucher number: DECBA2153.</p><p>Collection locale. CEIBA Biological Station, Madewini, Guyana.</p><p>GPS: 6°29'N, 58°13'W.</p><p>Collection date: December - 2011.</p><p>Collectors. Dominic Evangelista, Ian Biazzo, Joseph A. Evangelista, Paul Frandsen, William R. Kuhn, and Jessica L. Ware.</p><p>Collection/ecological information.</p><p>This specimen was collected in a pitfall trap baited with beer in an uplands secondary forest area.</p><p>Adult ♂ Figure 8.</p><p>Voucher number: DEKBO0594.</p><p>Collection locale. Karanamabu Ranch, Rupununi, Guyana.</p><p>GPS: 3°45'0.1"N, 59°18'53.7"W.</p><p>Collection date: 10 - June - 2013.</p><p>Collectors. Dominic A. Evangelista, Oswin Ambrose, Susan George, and Megan M. Wilson.</p><p>Collection/ecological information.</p><p>This specimen was collected in a pitfall trap baited with beer in a forest proximal to the Rupununi River.</p><p>Morphological identification.</p><p>Both specimens mostly agree with the description and figures of Hebard (1916). However, there are slight differences in the supra-anal plate when compared to Hebard’s illustration. The white region on the SA plate of our specimen is slightly larger than in Hebard’s illustration. It is possible that this is a different species than that described by Hebard, but this cannot be fully determined without a full phylogenetic treatment of sexual morphology and genetic information of individuals from both Trinidad and Guyana.</p><p>Known geographic distribution.</p><p>Trinidad and Tobago, and Guyana</p></div>	https://treatment.plazi.org/id/50AFE6E5CB1C6C45F763B02274361024	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Evangelista, Dominic A.;Chan, Kimberly;Kaplan, Kayla L.;Wilson, Megan M.;Ware, Jessica L.	Evangelista, Dominic A., Chan, Kimberly, Kaplan, Kayla L., Wilson, Megan M., Ware, Jessica L. (2015): The Blattodeas. s. (Insecta, Dictyoptera) of the Guiana Shield. ZooKeys 475: 37-87, DOI: http://dx.doi.org/10.3897/zookeys.475.7877, URL: http://dx.doi.org/10.3897/zookeys.475.7877
94A8328DADB3CDFDB48A6D1A5FD08C88.text	94A8328DADB3CDFDB48A6D1A5FD08C88.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Xestoblatta berenbaumae Evangelista, Kaplan, & Ware	<div><p>Taxon classification Animalia Blattodea Ectobiidae</p><p>Xestoblatta berenbaumae Evangelista, Kaplan, &amp; Ware sp. n.</p><p>Authors of description.</p><p>Evangelista, Kaplan, &amp; Ware.</p><p>Holotype.</p><p>Adult ♂ Figure 9 B–E, G.</p><p>Voucher number: DECBA2109.</p><p>Type locality. CEIBA Biological Station, Madewini, Guyana.</p><p>GPS: 6°29'N, 58°13'W.</p><p>Collection date: 17 to 18 - August - 2012.</p><p>Collectors. Dominic A. Evangelista and William R. Kuhn.</p><p>Type information.</p><p>The holotype specimen is stored in ethanol with genitalia in a separate ethanol vial and is deposited at the Center for Biodiversity at the University of Guyana.</p><p>Collection/ecological information.</p><p>This specimen was collected in a pitfall trap baited with beer and fruit in an uplands secondary forest in CEIBA Biological Station.</p><p>Morphological identification.</p><p>This specimen was identified as Xestoblatta Hebard, 1916 by the position of the hooked phallomere (left), the presence of the external modification of the tergum as part of the dorsal tergal gland (Figure 9A), incomplete rami on the ulnar vein of the hind wing (Figure 9I) and the spination (type A) on the ventro-anterior margin of the fore-femur.</p><p>Holotype morphological description.</p><p>Head uniformly colored a deep mahogany. Clypeus pale buffy. Ocellar spots easily distinguishable, smaller than antennal pits and white. Head otherwise without distinguishing features. Ocellar spots slightly closer together than eyes. Facial grooves on lateral most edge. See Figure 9H for a representative photo of the head.</p><p>Pronotum a uniformly reddish mahogany color (Figure 10A). Medial expansion on posterior margin of pronotum is barely noticeable. Ventral margin of pronotum not lined with hairs. Anterior margin of pronotum significantly conformed around the head. Leg coloration deep orange amber. Coxae with some diffuse black regions. Ventro-anterior margin of fore-femur with 14 (left) or 13 (right) spines decreasing in size from basal to apical, one slightly larger pre-apical spine and one large apical spine (16 total left, 15 total right). Ventro-posterior margin of forelimbs with 4 large spines and 1 apical spine. Ventro-anterior margin of middle leg has seven large spines and one apical spine. Middle leg also with one large genicular spine. Hind leg ventro-anterior margin has six spines, one apical spine, and one genicular spine. Pulvilli present on all tarsomeres. Arolia present but not surpassing the tips of the pretarsal claws. Claws symmetrical and unspecialized.</p><p>Ulnar vein with three incomplete rami and three complete rami (Figure 9I). Tegmina reddish mahogany with small patch of white under the base of the subcostal vein.</p><p>Supra-anal plate subtriangular with a blunt tip from dorsal view. Left paraproct modified into a tri-dentate spine (Figure 9F; bi-dentate in some other specimens). Sub-genital plate has both styli highly modified (Figure 9F, G). The right stylus is projecting dorso-medially from posterior margin, curving back posteriorly and terminating in a shape reminiscent of a bifurcated serpentine tongue. Left stylus projecting dorsally, shorter than right stylus and tipped with a small, translucent, irregularly shaped ball (Figure 9F, G).</p><p>Left phallomere (Figure 9B, C) hooked in apical third. (Hooked phallomere is about 1.5 mm long). Medial phallomere (Figure 9D) approximately three times the length of the left phallomere, roughly uniform width, and a slight slender curve in the posterior end. R2c (Figure 9E) divided into two sclerites that form dual concave cups that meet dorsally.</p><p>Dorsal modification of terga as part of the dorsal tergal gland. Modification represented by a small patch of hairs with a concave semi-circular modification of the margin of the segment anterior to the gland. See Figure 9A for an illustration of a representative dorsum.</p><p>Medium sized hairs (~ 2 mm) covering entire body roughly uniformly, yet sparsely.</p><p>Other adult male paratypes.</p><p>Voucher numbers: DECBA1967, DECBA0801, DECBA1958, DECBA2182, DECBA2092, DECBA2039</p><p>Collection/ecological information.</p><p>All additional male individuals reported here were collected in leaf litter pitfall traps baited with beer at various locations (dryer secondary uplands forest and wet primary lowlands forest) in CEIBA biological station.</p><p>Adult female paratype morphological description.</p><p>Voucher number: DECBA2074.</p><p>Head slightly darker in color than male with a more reflective surface. Other features of head similar to male.</p><p>Description of legs similar or identical to that of male with the following spination on the ventro-anterior margin of fore-femur: 13 (left) and 12 (right) spines decreasing in size from basal to apical, two larger preapical spines and one large apical spine (16 total left and 15 total right). Ventro-posterior margin of fore-femur four large spines and one apical spine. Ventro-anterior margin of mid-leg with seven large spines, one apical spine, and one genicular spine. Ventro-anterior margin of hind-leg with five large spines, one apical spine, and one genicular spine.</p><p>Tegmina and wings reduced and not reaching end of abdomen. Three incomplete and three complete rami on ulnar vein. Ulnar vein very faint in the reduced wings of the female (Figure 10B; Table 3).</p><p>Pronotum matches description of the male.</p><p>Subgenital plate slightly more abbreviated than in male. Paraprocts simple and unspecialized. Sub-genital plate simple and symmetrical.</p><p>Other adult female paratypes.</p><p>Voucher numbers: DECBA1787, DECBA1791, DECBA1792, and DECBA1793</p><p>Collection/ecological information.</p><p>All additional female individuals reported here were collected in leaf litter pitfall traps baited with beer in an uplands secondary forest at CEIBA biological station.</p><p>Summary of female morphology.</p><p>All individuals match the description of the above female and have the following spination on the vento-anterior margin of the fore-limb: 13 spines decreasing in size from basal to apical, one or two slightly larger preapical spines and one large apical spine making a total of 15 or 16 spines.</p><p>Juvenile paratypes.</p><p>Voucher numbers: DECBA1788, DECBA1789, DECBA1790, DECBA1796.</p><p>Collection/ecological information.</p><p>All additional juvenile individuals reported here were collected in leaf litter pitfall traps baited with beer in an uplands secondary forest at CEIBA biological station.</p><p>Summary of juvenile morphology.</p><p>Juveniles are apterous and largely match the morphology of adults except for in the following. Simple styli present on the subgenital plate in some individuals but are short and abbreviated. Spines on ventro-anterior margin of forelimb are as follows: 12 to 14 spines decreasing in size basally to apically, one or two slightly larger preapical spines and one large apical spine making a sum total of 15 or 16 total spines.</p><p>Molecular data and evolutionary placement.</p><p>Vouchers numbers and GenBank accession numbers: DECBA1791 - KF155114, DECBA1789 - KF155105, DECBA0801 - CBA0801, DECBA1827 - KF155103, DECBA1826 - KF155107, DECBA1814 - KF155115. The clade containing the above haplotypes (formerly reported as " Blattodea sp.1") is supported by 96% bootstrap support and the haplotypes are nearly identical.</p><p>Diagnostic features of Xestoblatta berenbaumae .</p><p>The morphology of modified styles on the subgenital plate is the most useful trait for discerning this species with other Xestoblatta Hebard, 1916. The simple dorsal tergal gland, shape of the paraprocts (left modified into a tri-dentate or bi-dentate spine), and morphology of the internal genital sclerites of the male are also useful in identifying this species. Unfortunately the adult females and juveniles are largely lacking obvious identifying characteristics and there may be errors made in associating juveniles to the adults without the use of genetic information.</p><p>Etymology.</p><p>We give this species the specific epithet “berenbaumae” in honor of the esteemed entomologist, Dr. May Berenbaum, who has made huge contributions to entomology through scientific products, service and public outreach.</p><p>Known geographic distribution.</p><p>Guyana</p></div>	https://treatment.plazi.org/id/94A8328DADB3CDFDB48A6D1A5FD08C88	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Evangelista, Dominic A.;Chan, Kimberly;Kaplan, Kayla L.;Wilson, Megan M.;Ware, Jessica L.	Evangelista, Dominic A., Chan, Kimberly, Kaplan, Kayla L., Wilson, Megan M., Ware, Jessica L. (2015): The Blattodeas. s. (Insecta, Dictyoptera) of the Guiana Shield. ZooKeys 475: 37-87, DOI: http://dx.doi.org/10.3897/zookeys.475.7877, URL: http://dx.doi.org/10.3897/zookeys.475.7877
5E85DDB177A7AA8CE81030B284321FE6.text	5E85DDB177A7AA8CE81030B284321FE6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Xestoblatta agautierae Grandcolas 1992	<div><p>Taxon classification Animalia Blattodea Ectobiidae</p><p>Xestoblatta agautierae Grandcolas, 1992</p><p>Materials.</p><p>Adult ♂.</p><p>Voucher number: DEKBO0827.</p><p>Collection locale. Wilson’s pond trail (Honey pond trail), Karanambu Ranch, Rupununi, Guyana.</p><p>GPS: 3°44'42.36"N, 59°19'15.21"W.</p><p>Collection date: 10 - June - 2013.</p><p>Collectors. Dominic A. Evangelista, Oswin Ambrose, Susan George, and Megan M. Wilson.</p><p>Adult ♀.</p><p>Voucher number: DEKBO0826.</p><p>Collection locale. Forest Island “Darwin”, Karanambu Ranch, Rupununi, Guyana.</p><p>GPS: 3°47'47.62"N, 59°22'6.77"W.</p><p>Collection date: 14 - June - 2013.</p><p>Collectors. Dominic A. Evangelista, Oswin Ambrose, Susan George, and Megan M. Wilson.</p><p>Collection/ecological information.</p><p>Both specimens above were collected in pitfall traps baited with beer in the forests of the Rupununi savannah.</p><p>Morphological identification.</p><p>The left genital phallomere, right genital phallomere, absence of a dorsal tergal gland and body coloration match closely with the species description (Grandcolas 1992a). The styli differ slightly to the illustrations in the original description in that the left stylus of our specimen is shorter and originates more medially. The female was associated to the male by comparison of gross morphology and body coloration. See Figure 11 for photos of adult male and adult female.</p><p>Collection/ecological information for other specimens not reported here.</p><p>We collected many individuals of this species from most forested areas surrounding Karanambu Ranch. We collected only one individual of this species in a similar trap at the edge of a forest, near open savannah. We found this species and Xestoblatta berenbaumae sp. n. to be extremely abundant in their respective localities (&gt;100 individuals of each collected). However, both are previously unreported for Guyana. We believe this can be attributed to the fact that we used beer and fermenting fruit to bait out pitfall traps. As Gurney (1939) reports, Xestoblatta Hebard, 1916 were rare in collections until the contributions of an entomologist trapping fruit flies in Panama. We can speculate that these fruit flies were also collected with some sort of aromatic bait (as this is common for fruit fly trapping) that attracted the Xestoblatta Hebard, 1916 as by-catch.</p><p>Known geographic distribution.</p><p>Guyana (new record) and French Guiana.</p></div>	https://treatment.plazi.org/id/5E85DDB177A7AA8CE81030B284321FE6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Evangelista, Dominic A.;Chan, Kimberly;Kaplan, Kayla L.;Wilson, Megan M.;Ware, Jessica L.	Evangelista, Dominic A., Chan, Kimberly, Kaplan, Kayla L., Wilson, Megan M., Ware, Jessica L. (2015): The Blattodeas. s. (Insecta, Dictyoptera) of the Guiana Shield. ZooKeys 475: 37-87, DOI: http://dx.doi.org/10.3897/zookeys.475.7877, URL: http://dx.doi.org/10.3897/zookeys.475.7877
5E95AA97A505D9C30D3BEBC2737223EF.text	5E95AA97A505D9C30D3BEBC2737223EF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nyctibora dichropoda Hebard 1926	<div><p>Taxon classification Animalia Blattodea Ectobiidae</p><p>Nyctibora dichropoda Hebard, 1926</p><p>Materials.</p><p>Adult ♂ Figure 12.</p><p>Voucher number: DECBA0302.</p><p>GenBank accession number: KF155061.</p><p>Collection locale. CEIBA Biological Station, Madewini, Guyana.</p><p>GPS: 6°29'N, 58°13'W.</p><p>Collection date: 29 - July - 2011.</p><p>Collectors. Dominic A. Evangelista, Ian Biazzo, Manpreet K. Kohli, Melissa Sanchez-Herrera, Nicole Sroczinski and Jessica L. Ware.</p><p>Collection/ecological information.</p><p>This specimen was collected in the leaf litter.</p><p>Morphological identification.</p><p>This specimen matches the illustration and description by Hebard (1926) in the "striking pale" coloration on the surfaces of the tibiae, the definitive character for this species. However, the male we have is much larger than that which he described. It is matching in all other ways.</p><p>Molecular identification.</p><p>The COI barcodes of this specimen are close to an adult female (Voucher number: DECBA0235; GenBank accession number: KF155062) and juvenile specimen (Voucher number: DECBA0104; GenBank accession number: KF155024) of Nyctibora . Based on both genetic distance and morphological dissimilarity, these individuals are likely members of a separate species. We do not report them further here.</p><p>Known geographic distribution.</p><p>Guyana (new record), Suriname and French Guiana.</p></div>	https://treatment.plazi.org/id/5E95AA97A505D9C30D3BEBC2737223EF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Evangelista, Dominic A.;Chan, Kimberly;Kaplan, Kayla L.;Wilson, Megan M.;Ware, Jessica L.	Evangelista, Dominic A., Chan, Kimberly, Kaplan, Kayla L., Wilson, Megan M., Ware, Jessica L. (2015): The Blattodeas. s. (Insecta, Dictyoptera) of the Guiana Shield. ZooKeys 475: 37-87, DOI: http://dx.doi.org/10.3897/zookeys.475.7877, URL: http://dx.doi.org/10.3897/zookeys.475.7877
84AF0166B4510840A5B3D7BAA6BCB071.text	84AF0166B4510840A5B3D7BAA6BCB071.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chorisoneura inversa Hebard 1926	<div><p>Taxon classification Animalia Blattodea Chorisoneura</p><p>Chorisoneura inversa Hebard, 1926</p><p>Materials.</p><p>Adult ♂ Figure 13.</p><p>Voucher number: DECBA1782.</p><p>GenBank accession number: KF155130.</p><p>Collection locale. CEIBA Biological Station, Madewini, Guyana.</p><p>GPS: 6°29'N, 58°13'W.</p><p>Date: 7 to 11 - August - 2013.</p><p>Collectors. Dominic A. Evangelista, Ian Biazzo, Manpreet K. Kohli, Melissa Sanchez-Herrera, Nicole Sroczinski and, Jessica L. Ware.</p><p>Morphological identification.</p><p>This individual was recognizable when comparing to the description of Hebard (1926) and the presence of the anteriorly pointing “V” shape on pronotum.</p><p>Genetic information and evolutionary placement.</p><p>As discussed below, this specimen was placed near Calhypnorna Saussure &amp; Zehntner, 1893 with 75% bootstrap support.</p><p>Known geographic distribution.</p><p>Guyana, Suriname, French Guiana and Brazil.</p></div>	https://treatment.plazi.org/id/84AF0166B4510840A5B3D7BAA6BCB071	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Evangelista, Dominic A.;Chan, Kimberly;Kaplan, Kayla L.;Wilson, Megan M.;Ware, Jessica L.	Evangelista, Dominic A., Chan, Kimberly, Kaplan, Kayla L., Wilson, Megan M., Ware, Jessica L. (2015): The Blattodeas. s. (Insecta, Dictyoptera) of the Guiana Shield. ZooKeys 475: 37-87, DOI: http://dx.doi.org/10.3897/zookeys.475.7877, URL: http://dx.doi.org/10.3897/zookeys.475.7877
560593B5B7E86F79035EF8290B6799C8.text	560593B5B7E86F79035EF8290B6799C8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dendroblatta callizona Rehn 1928	<div><p>Taxon classification Animalia Blattodea Ectobiidae</p><p>Dendroblatta callizona Rehn, 1928</p><p>Materials.</p><p>Adult ♀ Figure 14.</p><p>Voucher number: DECBA0805.</p><p>Collection locale. CEIBA Biological Station, Madewini, Guyana.</p><p>GPS: 6°29'57.75"N, 58°13'7.28"W.</p><p>Date: 14 - August - 2011.</p><p>Collectors. Dominic A. Evangelista, Ian Biazzo, Manpreet K. Kohli, Melissa Sanchez-Herrera, Nicole Sroczinski, and Jessica L. Ware.</p><p>Juvenile</p><p>Voucher number. DECBA0901.</p><p>GenBank accession number: KF155067.</p><p>Collection locale. CEIBA Biological Station, Madewini, Guyana.</p><p>GPS: 6°29'57.75"N, 58°13'7.28"W.</p><p>Date: 13 - August - 2011.</p><p>Collectors. Dominic A. Evangelista, Ian Biazzo, Manpreet K. Kohli, Melissa Sanchez-Herrera, Nicole Sroczinski, and Jessica L. Ware.</p><p>Collection/ecological information.</p><p>Both of these specimens were collected in a cup baited with beer placed in the canopy. The cup was tied to the trunk of a tree 13.8 meters above the ground. The tree chosen was close to a swampy primary forest area and on the edge of grassy hillside (most likely a plot that had been burned in the past). There were traps placed in the same tree at other heights but both individuals of this species were caught in this particular trap.</p><p>Morphological identification.</p><p>Our female specimen of Dendroblatta callizona Rehn, 1928 is within the variation described by Rehn (1928). The juvenile specimen was identified by comparison with the adult and using genetic data as well.</p><p>Genetic information.</p><p>In the tree of Evangelista et al. (2014) this species is placed near two individuals reported as " Ectobiidae sp. 10". The morphology of these specimens is consistent with Dendroblatta cnephaia Hebard, 1926, although we do not report them here because of a lack of adults to confirm identification.</p><p>Known geographic distribution.</p><p>Trinidad and Tobago, Guyana, and Suriname.</p></div>	https://treatment.plazi.org/id/560593B5B7E86F79035EF8290B6799C8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Evangelista, Dominic A.;Chan, Kimberly;Kaplan, Kayla L.;Wilson, Megan M.;Ware, Jessica L.	Evangelista, Dominic A., Chan, Kimberly, Kaplan, Kayla L., Wilson, Megan M., Ware, Jessica L. (2015): The Blattodeas. s. (Insecta, Dictyoptera) of the Guiana Shield. ZooKeys 475: 37-87, DOI: http://dx.doi.org/10.3897/zookeys.475.7877, URL: http://dx.doi.org/10.3897/zookeys.475.7877
040FD72A86AC5E23B20192FB3F8EC0FE.text	040FD72A86AC5E23B20192FB3F8EC0FE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Calhypnorna sp. A	<div><p>Taxon classification Animalia Blattodea Ectobiidae</p><p>Calhypnorna sp. A</p><p>Authors of the description.</p><p>Evangelista, Wilson, &amp; Ware.</p><p>Materials.</p><p>Juvenile Figure 15.</p><p>Voucher number: DECBA1802.</p><p>GenBank accession number: KF155118.</p><p>Collection locale. CEIBA Biological Station, Madewini, Guyana.</p><p>GPS: 6°29'N, 58°13'W.</p><p>Collection date: 15 - August - 2012.</p><p>Collectors. Dominic A. Evangelista and William R. Kuhn.</p><p>Specimen information.</p><p>This specimen is stored in ethanol and is deposited in the Center for Biodiversity at the University of Guyana.</p><p>Identification and differential diagnosis.</p><p>We identified this specimen as Calhypnorna based on the following comparisons. Our specimen is not lacking an interocular carina as in Hypnornoides (Rehn 1917). Our specimen also has a definitively truncate posterior margin of the pronotum (Figure 15B), which differentiates it from Euhypnorna (Hebard 1921). Our specimen is lacking the hairs covering most of the body as in Hypnorna (1893) and most closely matches the illustration of Calhypnorna by Saussure &amp; Zehntner (1893).</p><p>Description .</p><p>The specimen is a juvenile that is likely in its penultimate instar. Overall, the body shape is elongated for a typical cockroach, and even for a typical Pseudophyllodromiinae . A large portion of the head is visible from a dorsal perspective, and reaches anteriorly past the pronotum significantly. The black coloration on the pronotum is the same width as the width of the head where it meets with the pronotal margin (Figure 15B).</p><p>Antennae are hirsute to nearly plumose. The antennae are slightly clubbed basally with the widest point occurring at first segment of the flagellum. There are two major color regions of the antennae: a dark basal region and a light distal region. The dark basal region begins as slightly lighter than the remainder but becomes a dark black color by the end of the dark region. The 25th segment of the antennae is the final dark segment. The 26th antennal segment begins the light region of the antennae. The 26th or 27th and subsequent segments are nearly white, becoming more brownish orange after the 7th white segment (33 total). The total number of antennal segments on the specimen is 38 (left) and 44 (right).</p><p>The head is very large in relation to the remainder of the body, triangular, and wider than typical for a Pseudophyllodromiinae (Figure 15A). Inter-ocular space is sharply angled creating a carina that begins where the compound eye meets the antennae. The antennal pits are closer together than the eyes. Eyes are prominent and appear to bulge the head laterally. Facial grooves spanning from the posterior portion of the eye towards the mouthparts are prominent. Coloration on head is brown-orange overall with a slightly lighter, less brown, patch above and below the carina. Ocellar spots are either absent or not readily visible.</p><p>The pronotum is colored with a dark black region taking up the major two fifths of the medial area. The black area is opaque and reaches forward to the anterior margin but just stops short of completion in the posterior eighth of the segment. The black region is nearly rectangular, slightly rounded anteriorly and widened posteriorly (Figure 15B). Bordering the black region laterally and posteriorly are translucent regions colored brown-orange similar to the remainder of the body.</p><p>Meta- and meso-thoracic segments are both strongly lobed, presumably due to the developing wings within. Color is orange-brown overall with small amounts of black on the tips of the posterior pair of wing pads. Legs are light in color with a slight orange tinge overall. Dark regions are present on the medial side of the base of the fore-coxae.</p><p>The ventro-anterior margin of the fore-femur have five (right) or eight (left) large piliform spines basally followed by 27 (right) and 20 (left) shorter piliform spines, which are then each followed by one larger piliform spine and finally one large distal spine that is not piliform. Arolia are large and extend beyond the tips of the pretarsal claws on all legs. Claws are symmetrical and unspecialized.</p><p>Both the venter and dorsum of the abdomen is the same orange-brown color as the remainder of the body, but with a slightly redder tinge. Soft black color borders the abdomen laterally and posteriorly.</p><p>The dorsal abdomen is mostly glabrous. Hairs that are present are most dense laterally and on segments five and six. Ventral abdomen is glabrous as well, with fewer hairs than on the dorsal side and no regions with any dense pubescence. Supra-anal plate is unspecialized and broadly subtrapezoidal or triangular. Subgenital plate is broadly subtrapezoidal with the posterior margin being broader than that of the subgenital plate. The posterior margin of the subgenital plate is not perfectly uniform and conforms around two large styli. Styli are equal in length to the entire subgenital plate. Their width is equal to half of the length of the visible portion of the styli.</p><p>Genetic information and evolutionary placement.</p><p>Evangelista et al. (2014) recovers this sequence as being most closely related to a species reported as " Ectobiidae sp. 6" with 75% bootstrap support. This species is identified above as Chorisonuera inversa Hebard, 1926. Hebard hypothesized that these are closely related genera (Hebard 1921a) and we can now say that genetic data supports this hypothesis. We cannot definitively say, however, that they are sister taxa because of incomplete phylogenetic sampling in this tree. Thus, we follow Hebard (1921a) and not Beccaloni (Beccaloni 2007) and consider this to be in the Psuedophylodromiinae .</p><p>Known geographical distribution of Calhypnorna.</p><p>Guyana (new record), Para Brazil, Bolivia and Panama.</p><p>Collection/ecological information.</p><p>This specimen was found crawling through a benab. The only individual of this species observed in the field was the one collected and described here. Given that our overall collecting effort was significant (&gt;1000 individuals of Blattodea s.s.) and we only found a single individual of Calhypnorna sp. A, we consider this species to be quite rare.</p><p>Previous work (Shelford 1912) has cited species of this genus as being beetle mimics. However, we observed no beetle model in the field that this species may have been mimicking. We did notice a similarity in body coloration of a wasp and Hemipteran sympatric with this conspicuously colored Blattodea (Figure 16).</p><p>Notes on historical records of this genus.</p><p>The genus Calhypnorna Saussure &amp; Zehntner, 1893 was originally established as a subgenus of Hypnorna Stål, 1860. It was then given generic status by Kirby (1904). The genera Calhypnorna, Hypnorna, Hypnornoides Rehn, 1917 and Euhypnorna Hebard, 1921 are thought to be closely related (Hebard 1921). These are known from a number of regions (Para and Rio de Janiero Brazil, Bolivia and Panama) but there are no records from the Guiana Shield. Therefore, a new record of this species from the coastal rainforests of Guyana is geographically disjointed from all other records of these taxa. On this basis alone, we might distinguish this specimen as a new species. However, since our lone specimen is a juvenile, we have limited morphological basis for differentiating this from known taxa. We refrain from establishing this as new species until adult specimens can be found but we still give a synopsis of the biological traits of this specimen. This new record extends the potential range of Calhypnorna Saussure &amp; Zehntner, 1893 and it has now been recorded from Para Brazil (south of Amazon), Bolivia, Panama, and Guyana (new record).</p></div>	https://treatment.plazi.org/id/040FD72A86AC5E23B20192FB3F8EC0FE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Evangelista, Dominic A.;Chan, Kimberly;Kaplan, Kayla L.;Wilson, Megan M.;Ware, Jessica L.	Evangelista, Dominic A., Chan, Kimberly, Kaplan, Kayla L., Wilson, Megan M., Ware, Jessica L. (2015): The Blattodeas. s. (Insecta, Dictyoptera) of the Guiana Shield. ZooKeys 475: 37-87, DOI: http://dx.doi.org/10.3897/zookeys.475.7877, URL: http://dx.doi.org/10.3897/zookeys.475.7877
