taxonID	type	description	language	source
CE5F2D45B1E3525FBE311C7E06221AF7.taxon	description	Figs 5, 6	en	Huang, Yong, Li, Hongyu, Wang, Yilin, Li, Maojin, Hou, Mian, Cai, Bo (2023): Taxonomic review of the Calotes versicolor complex (Agamidae, Sauria, Squamata) in China, with description of a new species and subspecies. ZooKeys 1187: 63-89, DOI: http://dx.doi.org/10.3897/zookeys.1187.110704, URL: http://dx.doi.org/10.3897/zookeys.1187.110704
CE5F2D45B1E3525FBE311C7E06221AF7.taxon	description	Description of holotype. Adult male, medium-sized body, SVL 96.40 mm, trunk length 49.82 mm, head length 24.29 mm, head depth 13.18 mm, head width 15.42 mm, interorbital width 9.47 mm, snout width 5.87 mm, eye-ear length 5.97 mm, eye diameter 8.56 mm, naris-eye length 4.58 mm, jaw width 16.14 mm, snout-eye length 8.82 mm, snout-forelimb length 32.65 mm, tail length 256.15 mm, finger IV length 14.10 mm, toe IV length 19.04 mm, upper arm length 15.80 mm, lower arm length 14.34 mm, upper leg length 21.94 mm, crus length 16.87 mm, forelimb length 46.03 mm, hind limb length 73.17 mm, when hind limbs adpressed forward to reach between eyes and tympanum. Supralabial scale count 10: 11, infralabial scale count 10: 11, nasal-supralabial scale rows 1: 1, suborbital scale rows 3: 3, gular scale count 30, ventral scale count 48, finger IV subdigital lamellae count 20: 21, toe IV subdigital lamellae count 25: 25, post-tympanic scale count 3: 3, post-occipital scale count 3: 2, vertebral scales 42, mid-body scale rows 42, dorsal eyelid scales 13: 13, scales between anterior chin-shield 1, scales between nasal shield and orbit 5: 5, state of scales on side of neck and adjacent shoulder area pointing obliquely upward, keels on these scales are weakly to strongly developed, nuchal and dorsal crest scales short, nuchal crest scales significantly larger than dorsal crest scales, dorsal crest shortens progressively before mid-body, gular fold and fold in front of the shoulder are absent, moderate to large scales in front of forelimb insertion, postorbital spine absent, scales on side of trunk point obliquely upward. Under stress, the color is khaki (240,230,140) or dark khaki (189,183,107) with pale gray (105,105,105) markings, three black (0,0,0) transverse stripes on top of head, nine blank radial stripes around eyes, throat coloration burly wood (222,184,135) with blank throat stripes, inner-lip coloration is white-smoke (245,245,245), oral cavity coloration is pale flesh (239, 205, 197), tongue coloration is orange (255,165,0), ventral body coloration tan (210,180,140) with dark stripes, presence of dark line on vent midline from throat to pelvis, seven dark khaki bands on dorsum of trunk between axilla and inguen, 7 - 8 dark khaki fore- and hindlimb cross-bands, striping ventrally on trunk continuous striping, 22 tail cross-bands (the front 9 bands are black and the rear 13 bands are brown [165, 42, 42]).	en	Huang, Yong, Li, Hongyu, Wang, Yilin, Li, Maojin, Hou, Mian, Cai, Bo (2023): Taxonomic review of the Calotes versicolor complex (Agamidae, Sauria, Squamata) in China, with description of a new species and subspecies. ZooKeys 1187: 63-89, DOI: http://dx.doi.org/10.3897/zookeys.1187.110704, URL: http://dx.doi.org/10.3897/zookeys.1187.110704
CE5F2D45B1E3525FBE311C7E06221AF7.taxon	etymology	Etymology. The species name wangi is named after Prof. Yuezhao Wang, a former director of the Amphibian and Reptile Research Laboratory (CIB, CAS) and Museum of Herpetology (CIB, CAS) for his research on Chinese herpetology and his contributions in leading the Amphibian and Reptile Research Laboratory through many difficulties. We suggest the English common name Wang's garden lizard and the Chinese name 中国树蜥 (zhōng guo shu xī).	en	Huang, Yong, Li, Hongyu, Wang, Yilin, Li, Maojin, Hou, Mian, Cai, Bo (2023): Taxonomic review of the Calotes versicolor complex (Agamidae, Sauria, Squamata) in China, with description of a new species and subspecies. ZooKeys 1187: 63-89, DOI: http://dx.doi.org/10.3897/zookeys.1187.110704, URL: http://dx.doi.org/10.3897/zookeys.1187.110704
6CF7CF9AEE175B6DB43C90E42C4B44EE.taxon	description	Fig. 4	en	Huang, Yong, Li, Hongyu, Wang, Yilin, Li, Maojin, Hou, Mian, Cai, Bo (2023): Taxonomic review of the Calotes versicolor complex (Agamidae, Sauria, Squamata) in China, with description of a new species and subspecies. ZooKeys 1187: 63-89, DOI: http://dx.doi.org/10.3897/zookeys.1187.110704, URL: http://dx.doi.org/10.3897/zookeys.1187.110704
6CF7CF9AEE175B6DB43C90E42C4B44EE.taxon	materials_examined	Other material examined. Hainan Province, China: 1. Qiongzhong, Wanling Town (2 ♂, 1 ♀), Hongmao Town (1 ♂, 1 ♀), Zayun Town (1 ♂), Hongmao Town (3 ♂, 1 ♀), Limushan Town (1 ♀). 2. Wuzhishan, Hongshan Town (3 ♂, 2 ♀), Fanyang Town (5 ♀). 3. Dingan, Donghong (1 ♀). 4. Ledong, Yongming (1 ♀). 5. Tunchang, Dalupo (1 ♀).	en	Huang, Yong, Li, Hongyu, Wang, Yilin, Li, Maojin, Hou, Mian, Cai, Bo (2023): Taxonomic review of the Calotes versicolor complex (Agamidae, Sauria, Squamata) in China, with description of a new species and subspecies. ZooKeys 1187: 63-89, DOI: http://dx.doi.org/10.3897/zookeys.1187.110704, URL: http://dx.doi.org/10.3897/zookeys.1187.110704
6CF7CF9AEE175B6DB43C90E42C4B44EE.taxon	description	Description of holotype. Adult male, medium-sized body, SVL 88.34 mm. Trunk length 43.82 mm, head length 24.47 mm, head depth 14.41 mm, head width 16.08 mm, interorbital width 8.51 mm, snout width 5.29 mm, eye-ear length 6.30 mm, eye diameter 7.66 mm, naris-eye length 4.49 mm, jaw width 16.06 mm, snout-eye length 7.85 mm, snout-forelimb length 30.74 mm, tail length 283.89 mm, toe IV length 21.57 mm, finger IV length 13.41 mm, upper arm length 16.64 mm, lower arm length 13.59 mm, upper leg length 23.01 mm, Crus length 20.26 mm, forelimb length 44.82 mm, hindlimb length 71.77 mm, when hind limbs are adpressed forward to reach between eyes and tympanum. Supralabial scale count 10: 10, infralabial scale count 11: 11, nasal-supralabial scale rows 1: 1, suborbital scale rows 3: 3, gular scale count 21, ventral scale count 46, finger IV subdigital lamellae count 22: 22, toe IV subdigital lamellae count 27: 27, post-tympanic scale count 1: 1, post-occipital scale count 2: 2, vertebral scales 38, mid-body scale rows 38, dorsal eyelid scales 13: 13, no scales between anterior chin-shield, scales between nasal shield and orbit 5: 5, state of scales on side of neck and adjacent shoulder area pointing obliquely upward, keels on these scales are weakly to strongly developed, nuchal and dorsal crest scales short, nuchal crest scales significantly larger than dorsal crest scales, dorsal crest shortening progressively before mid-body, gular fold and fold in front of the shoulder absent, no patch of granular scales in front of forelimb insertion, pre-axillary area with moderate to large scales, postorbital spine absent, scales on side of trunk pointing obliquely upward. In ethanol, color is dark khaki or khaki with black or pale gray (105, 105, 105) markings. Tan and black mixed marking on top of head, nine brown (165, 42, 42) radial stripes around eyes, throat coloration burly wood with blank throat stripes, inner lip, oral cavity and tongue coloration is smoky white, ventral body is tan with dark stripes, presence of blank line on vent midline from throat to pelvis, six dark-khaki bands on dorsum of trunk between axilla and inguen, Seven to eight dark-khaki cross-bands on the fore- and hindlimbs, continuous ventral striping on trunk, and 22 cross-bands on tail (anterior nine bands are black, posterior 13 are brown).	en	Huang, Yong, Li, Hongyu, Wang, Yilin, Li, Maojin, Hou, Mian, Cai, Bo (2023): Taxonomic review of the Calotes versicolor complex (Agamidae, Sauria, Squamata) in China, with description of a new species and subspecies. ZooKeys 1187: 63-89, DOI: http://dx.doi.org/10.3897/zookeys.1187.110704, URL: http://dx.doi.org/10.3897/zookeys.1187.110704
6CF7CF9AEE175B6DB43C90E42C4B44EE.taxon	etymology	Etymology. The specific epithet of hainanensis refers to Hainan Island where the new subspecies was discovered. We suggest the English common name Hainan garden lizard and the Chinese name 中国树蜥雷公马亚种 (zhōng guo shu xī lei gōng mǎ ya zhǒng), which comes from a colloquial name for Calotes wangi hainanensis in Hainan Province, China, meaning Thor's mount that can predict the weather.	en	Huang, Yong, Li, Hongyu, Wang, Yilin, Li, Maojin, Hou, Mian, Cai, Bo (2023): Taxonomic review of the Calotes versicolor complex (Agamidae, Sauria, Squamata) in China, with description of a new species and subspecies. ZooKeys 1187: 63-89, DOI: http://dx.doi.org/10.3897/zookeys.1187.110704, URL: http://dx.doi.org/10.3897/zookeys.1187.110704
6CF7CF9AEE175B6DB43C90E42C4B44EE.taxon	diagnosis	Diagnosis. Calotes wangi sp. nov. can be separated from all other species of Calotes by having the following characters: 1. Medium-sized adult male, SVL <90 mm (66.86 - 109.1 mm, average 85.64 mm). 2. Smaller HeadW in males, 11.64 - 19.72 (average 14.84). 3. Larger HindLimbL / SVL 0.64 - 0.89 (average 0.75). 4. Larger 4 ToeLng / SVL 0.08 - 0.3 (average 0.21). 5. Larger 4 FingLng / SVL 0.11 - 0.21 (average 0.14). 6. Eyelid 10 - 14 (average 12.4). 7. NarEyeS 4 - 5 (average 4.9). 8. Scales on side of trunk point obliquely upward. 9. Patch of granular scales in front of forelimb insertion absent. 10. Scales on side of neck and adjacent shoulder area point obliquely upward; keels on neck scales are weakly to strongly developed. 11. Paired dark patches are approximately triangular, extending from the front of the shoulder to the jaw angles. 12. Coloration of tongue is orange. 13. Nuchal and dorsal crest scales short; dorsal crest shortening progressively before mid-body; nuchal crest distinctly larger than dorsal crest. 14. Supratympanic spines are short. 15. Fourth toe with claw can stretch between the eyes and tympanum when hind limbs are adpressed forward. 16. Fold in front of the shoulder is absent. 17. During breeding season, males are uniformly dark orange to orange on the front half (except for the fingers) with black patches on both sides of neck and darker radial stripes around eyes; Hind body and toes a uniform khaki or dark khaki.	en	Huang, Yong, Li, Hongyu, Wang, Yilin, Li, Maojin, Hou, Mian, Cai, Bo (2023): Taxonomic review of the Calotes versicolor complex (Agamidae, Sauria, Squamata) in China, with description of a new species and subspecies. ZooKeys 1187: 63-89, DOI: http://dx.doi.org/10.3897/zookeys.1187.110704, URL: http://dx.doi.org/10.3897/zookeys.1187.110704
6CF7CF9AEE175B6DB43C90E42C4B44EE.taxon	distribution	Distribution and natural history. Calotes wangi sp. nov. is a transboundary species, ranging from southern China (Fujian Province, Guangdong Province, Hainan Province and Guangxi Zhuang Autono-mous Region, southern Hunan Province, Macao, and Hong Kong) to northern Vietnam (Lang Son). The eastern Yunnan region in China is also likely to harbor this species (Yang and Rao 2008; Shuo Liu, pers. comm. 17 April 2023). The known altitude range of C. w. wangi ssp. nov. is 15 ~ 2000 m and C. w. hainanensis is 47 ~ 1867 m. It is found in subtropical evergreen broad-leaved forests and tropical monsoon forests in southern China and northern Vietnam, mostly in mountainous areas, hills and plains on forest edges, arable land, shrub lands, and even urban green belts. It is active at the edge of the forest, and when it is in danger, it rushes into bushes or climbs tree trunks to hide. It is active from April to October every year, while in the tropics it is active from March to November or even longer. The authors investigated in Hainan and found that C. wangi sp. nov. was active from 7: 30 ~ 11: 30 hrs in summer and 15: 00 ~ 19: 00 hrs in the afternoon. Investigations in Guangxi found that the lizards lie on sloping shrub branches at night, sleeping close to the branches. Both Calotes wangi sp. nov. and C. irawadi are oviparous. It has been reported that females of C. wangi sp. nov. contained 5 - 12 (17 in one specimen in Mt. Diaoluo, Hainan) mature eggs from late April to September, while females of C. irawadi contained 12 mature eggs in June (Zhao et al. 1999). After our dissections, 13 specimens of C. wangi sp. nov. from Hainan obtained in August had 5 - 8 eggs (average 6), and five C. irawadi from Lushui, Yunnan obtained in late May were pregnant with 5 - 15 eggs (average 11). Calotes wangi sp. nov. eats a variety of insects, spiders, and other arthropods (Zhao et al. 1999).	en	Huang, Yong, Li, Hongyu, Wang, Yilin, Li, Maojin, Hou, Mian, Cai, Bo (2023): Taxonomic review of the Calotes versicolor complex (Agamidae, Sauria, Squamata) in China, with description of a new species and subspecies. ZooKeys 1187: 63-89, DOI: http://dx.doi.org/10.3897/zookeys.1187.110704, URL: http://dx.doi.org/10.3897/zookeys.1187.110704
6CF7CF9AEE175B6DB43C90E42C4B44EE.taxon	materials_examined	Specimens examined. Calotes irawadi (n = 18). Myanmar: Kawnglanghpu (CIB 097515). China: Yunnan Province, Lushui City (Shangjiang CIB 001824 - 26), Baoshan City (Shangjiang CIB 001819 - 23, CIB 001827 - 28), Dehong Dai and Jingpo Autonomous Prefecture (Hongbenghe CIB 116106 and CIB 116212, Tongbiguan KIZ 059191, KIZ NB 20180905, KIZ HBH 20200913 and KIZ HBH 20200914), Xishuangbanna Dai Autonomous Prefecture (CIB 001610).	en	Huang, Yong, Li, Hongyu, Wang, Yilin, Li, Maojin, Hou, Mian, Cai, Bo (2023): Taxonomic review of the Calotes versicolor complex (Agamidae, Sauria, Squamata) in China, with description of a new species and subspecies. ZooKeys 1187: 63-89, DOI: http://dx.doi.org/10.3897/zookeys.1187.110704, URL: http://dx.doi.org/10.3897/zookeys.1187.110704
