taxonID	type	description	language	source
D076D419B504FFC1FF23F2C12649F9B7.taxon	discussion	Large damselflies partly metallic red, purple or green that inhabit flowing water. Adults perch with the wings together over the abdomen. The nymphs are long-legged, antennal segment 1 is as long as the rest of the antenna, and lateral caudal gills are long and triangular in cross section. Traditionally, all species in Ecuador were placed in two genera (Hetaerina and Mnesarete); Garrison (1990) added Ormenophlebia. A recent molecular and morphological study indicates that all the species that have been placed in these genera are congeneric (Seth Bybee, pers. comm.); however, taxonomic problems with the higher classification of these taxa remain, and Rosser Garrison (pers. comm.) maintains that Ormenophlebia is a valid genus.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B504FFCCFF23F04D27D4FE6F.taxon	discussion	Most species of Hetaerina have a red patch in the base of the wings and some have a small red or brown spot at the tips of the wings (Fig. 9). Nine of the 37 known species occur in Ecuador. They inhabit small to medium-sized streams, usually in partly to heavily forested areas; nymphs of five species have been described. The most recent revision was by Garrison (1990). Hetaerina caja (Drury, 1773). Publ. Recs. ECU: Calvert (1901), Tsuda (1986, 1991), Garrison (1990), Paulson (1983) (H 08), (P 11), (H 14); CR: * Campos (1922); EO: Campos (1927); GY: Ris (1918), Campos (1922); LR: Campos (1922); OR: Daigle (1996), von Ellenrieder & Garrison (2011); PA: * McLachlan (1881 a); SD: Daigle et al. (1997). Also CEN, COL, GUF, PER, VEN, TTO: (H 08), (P 11); plus SUR: (P 16). Addl. Data. CN, La Troncal: [SWD]; EO, Arenillas: [FSCA, JTJ, KJT]; Piñas: [ACRP]; Santa Rosa: [JTJ, KJT]; GY, Balao: [FSCA, JTJ, KJT]; Naranjal: [FSCA]; LJ, Chaquarpamba: [FSCA, KJT]; Macará: [ANSP, FSCA]; LR, Babahoyo: [RWG]; Buena Fé: [DRP, FSCA, RWG, USNM]; Montalvo: [JTJ, KJT]; Ventanas: [JTJ, KJT]; MN, Chone: [FSCA, KJT, USNM]; El Carmen: [JTJ]; Junin: [JTJ, RWG]; Pedernales: [JTJ, KJT]; NA, Tena: [FSCA]; OR, Francisco de Orellana: [FSCA, RWG]; PA, Pastaza: [KJT, TWD, USNM]; SD: [FSCA, UTIC, JJD, JTJ, KJT, RWG, SWD]; SU, Cascales: [SWD]; Shushufindi: [DRP, FSCA]. Hetaerina capitalis Selys, 1873. Publ. Recs. ECU: * Garrison (1990), Tsuda (1991), (H 08), (P 11), (H 14). Also CEN, COL, VEN: (H 08), (P 11). Addl. Data. MS, Limón Indanza: 1 ♂, stream, Hwy. 45, 25.3 km N of Indanza (- 2.9217, - 78.4031, 1110 m), 7 xi 1997, leg. & det. KJT [KJT]; NA, El Chaco: 1 ♂, stream 3.5 km N of El Chaco, Hwy. 28 (- 0.3164, - 77.7919, 1425 m), 7 vi 1995, leg. & det. KJT [KJT]; PA, Pastaza: 1 ♂, Puyo, Río Pastaza (- 1.4667, - 77.9833, 980 m), 26 xi 1936, leg. WCM, det. RWG- 1990, from USMN [RWG]; Mera: 1 ♂, Abitagua, Río Pastaza watershed (- 1.45, - 78.15, 1200 m), iv 1937, leg. WCM, det. RWG- 1990 [UMMZ]; TU, Baños: 1 ♂, Río Topo, Río Pastaza watershed (- 1.10, - 78.65, 2800 m), xii 1948, leg. WCM, det. RWG, from UMMZ [RWG]; SD: 1 ♂, Tinalandia, 12 km E of Santo Domingo de los Colorados, streams S of river (- 0.3069, - 79.0492, 610 m), 5 x 1988, leg. & det. SWD [SWD]; 1 ♀, same data but 6 x 1988 [SWD]; 1 ♂, About 93 Km SW Quito, Chiriboga Road, 7 ix 1990, leg. & det. OSF [USNM]. Hetaerina charca Calvert, 1890. Publ. Recs. ECU: Tsuda (1991), (H 08), (P 11), (H 14); MS: * Garrison (1990). Also BOL, PER: (P 11); plus ARG: (RWG, pers. comm., 2019). Addl. Data. CB, Alausí: 1 ♂, 1 ♀, Huigra, upslope (- 2.28, - 78.98, 2134 m), 7 iii 1931, leg. W. Coxey, det. MJW [FSCA]; Camada: 2 ♂♂, Dos Puentes, km 99 along Quito-Guayaquil Railway, Río Chanchan just E of Ventura (- 2.25, - 79.10, 518 m), 10 I 1929, leg. W. Coxey, det. MJW [FSCA]; GY, Naranjal: 3 ♂♂, Bucay (- 2.2, - 79.18, 233 m), 13 ii 1911, leg. S. Rhoads [FSCA, ANSP]; LR, Quevedo: 1 ♂, stream 16 km E of Hwy. E 25 [from Formisa] (- 0.7256, - 79.3484, 200 m), 24 ii 2010, leg. & det. KJT [FSCA]; MS, Santiago: 1 ♂, Patuca, 5.2 km S, 23 x 1992, leg. M. Kippenhan, det. SWD [SWD]; OR, Loreto: 1 ♂, small stream near Reserva Río Bigal, about 30 km N of Loreto (- 0.5642, - 77.4114, 900 m), 17 v 2009, leg. & det. KJT [KJT]; PC, San Miguel Bancos: 1 ♀, Mindo, 30 iv 1990, leg. E. Smith, det. WFM [SWD]. Hetaerina duplex Selys, 1869. Publ. Recs. ECU: Tsuda (1986 —? as H. longipes), Hagen in Selys (1853 —? as H. carnifex), Paulson (1983 —? as H. carnifax), Tsuda (1991), Selys (1869), (H 08 — also as H. aurora), (P 11), (H 14); EO: Garrison (1990); LR: Campos (1922 — as mistaken identity H. hebe): LJ: Tennessen & Johnson (2008); PC: Selys (1873 a), Hagen (1875 —? as H. carnifex); ZC: * Garrison (1990). Also COL: (H 08), (P 11); plus PER: (H 08), (P 16). Addl. Data. AZ, Santa Isabel: [FSCA, JTJ, KJT]; BO, Caluma: [FSCA, JTJ, KJT]; San Miguel: [ANSP, DRP, FSCA, KJT]; CN, Ca ñ ar: [FSCA, JTJ, JJD, KJT, SWD]; CB, Cumandá: [FSCA]; Pallatanga: [ANSP, FSCA]; CT, Pangua: [FSCA, JTJ, KJT]; Sigchos: [FSCA]; EO, Pi ñ as: [FSCA, JTJ, RWG]; Portovello: [FAL, RWG]; GY, Guayaquil: [FSCA]; Daule: [FSCA]; Naranjal: [FSCA]; IM: Cotacachi [FSCA, KJT, USFQ]; LJ, Catamayo: [FSCA, JTJ]; Loja: [FSCA, FAL, UTIC, JTJ, KJT, RWG]; NA, Archidona: [RWG, SWD]; PC, Mejía: [DRP, RWG]; Pedro Vicente Maldonado: [FSCA]; Quito: [DRP, USNM]; San Miguel Bancos: [FSCA, KJT, RWG, USFQ, USNM]; SD: [DRP, FSCA, JJD, JTJ, KJT, RWG, USNM]; ZC, Zamora: [UTIC]. Note: Specimens in various collections may be labeled H. aurora (see Appendix A). Hetaerina flavipennis Garrison, 1990. Publ. Recs. ECU: Tsuda (1991), (H 08), (P 11), (H 14); MS: * Garrison (1990), Tennessen (2006); ZC: * Garrison (1990 — Type Locality: “ Zumba, ” Holotype [UMMZ]), Bridges (1993), Garrison et al. (2003). Addl. Data. MS, Huamboya: 2 ♂, Macas, about 56 km N on Hwy. 45, 1.4 km S of Río Pasanac, small stream with shallow mucky bottom (- 1.9503, - 77.8628, 823 m), 16 ix 2005 leg. & det. WFM [FSCA]; Morona: 1 ♂, 1 ♀, Mangosisa, Río Upano (- 2.5244, - 77.8947, 708 m), date?, leg. L Gomez Alonzo, det. RWG, from CHK [UMMZ]; NA, Archidona: 1 ♂, 3 ♀♀, km 45 of Hollin-Loreto Road (- 0.7228, - 77.5653, 1130 m), 18 xii 1989, leg. H. Real, det. RWG [RWG]; Tena: 2 ♀♀, Río Jatun Yacu, Ilupulin Creek (- 1.0167, - 77.8333, 700 m), i 1935, leg. WCM, det. RWG, from UMMZ [RWG]; 1 ♂, same data but 15 iv 1935 [RWG]; ZC, Zamora: Holotype ♂, [Zumbi] “ Zumba, ” Río Zamora, 31 x 1941, leg. D. Laddey, det. RWG, from CHK [UMMZ]. Notes: So far reported only from Ecuador. The specimens WFM collected were in the deep shade along with Heteragrion spp. See page 14 for correction of error regarding “ Zumba. ” Hetaerina fuscoguttata Selys, 1878. Publ. Recs. ECU: * Garrison (1990), Tsuda (1991), (H 08), (P 11), (H 14); ES: Daigle et al. (1997); SD: Daigle et al. (1997). Also CEN, COL: (H 08), (P 11); plus VEN: (H 08). Addl. Data. CB, Cumandá: [RWG]; EO, Piñas: [DRP]; ES, San Lorenzo: [FAL, FSCA, KJT, LSAM, QZAC]; LR, Buena Fé: [DRP]; MN, Pedernales: [ACRP, FSCA, KJT]; MS, Gualaquiza: 1 ♂, Bomboiza, 17 km S Guala- quiza 20 ix 1990 leg. OSF, det. SWD [USNM]; PC, San Miguel Bancos: [DRP, KJT, USFQ]; SD: [FSCA, JJD, KJT, RWG]. Note: The Gualaquiza specimen represents the only record from the east slope. Hetaerina miniata Selys, 1879. Publ. Recs. ECU: von Ellenrieder (2007), (H 08), (P 11), (H 14); SD: * Garrison (1990 — as “ PC ”). Also CEN, COL, VEN: (H 08), (P 11). Addl. Data. LR, Buena Fé: 2 ♂♂, 1 ♀, Santo Domingo, 47 km S, stream at Río Palenque (200 m), 18 vii 1977, leg. & det. DRP [DRP]; 2 ♂♂, same data but 9 x 1988, leg. & det. SWD [SWD]; 1 ♂, [Consumulo] S Río Palenque (- 0.58, - 79.35, 175 m), 17 vi 1974, leg. J. Longino, det. MJW [FSCA]; Quevedo: 1 ♂, stream 16 km E of Hwy. E 25, from Formisa (- 0.7256, - 79.3483, 200 m), 23 ii 2010, leg. & det. KJT [FSCA]; 2 ♂♂, same data but leg. & det. JTJ [JTJ]; MN, Junin: 1 ♀, [RWG]. Palmar (- 0.9333, - 80.2333, 200 m), 10 iv 1941, leg. D. Laddey, det. RWG, from UMMZ [RWG]. Hetaerina occisa Hagen in Selys, 1853. Publ. Recs. ECU: * Garrison (1990), (H 08), (P 11), (H 14). Also CEN, COL, PER, TTO, VEN: (P 11); plus BRA: (H 08). Addl. Data. MS: Gualaquiza: [FSCA, USNM]; Huamboya: [SWD]; Limón Indanza: [FSCA, KJT]; Morona: [FSCA, RWG]; Santiago: [SWD]; NA, Archidona: [FSCA, KJT, SWD, USNM]; Carlos Julio Arosemena Tola: [FAL]; Tena: [KJT, USNM]; OR, Loreto: [FSCA, KJT, SWD, TWD, USNM]; PA, Mera: [FSCA, JTJ]; Pastaza: [FSCA, JTJ, KJT]; SU, Cascales: [SWD]; ZC, El Pangui: [KJT, USNM]; Yantzaza: [FSCA, KJT]; Zamora: [JTJ, USNM]. Hetaerina sanguinea Selys, 1853. Publ. Recs. ECU: * Garrison (1990), (L 05), (H 08), (P 11), (H 14); NA: Daigle (1996); OR: von Ellenrieder & Garrison (2011). Also ARG, BOL, BRA, COL, PER, VEN: (L 05), (H 08), (P 11). Addl. Data. MS, Gualaquiza: [ANSP, FSCA, USNM]; Huamboya: [KJT, SWD]; NA, Carlos Julio Arosemena Tola: [FSCA]; Tena: [DRP, FSCA, UTIC, JJD, KJT, RWG, TWD, USNM]; OR, Francisco de Orellana: [FSCA, KJT, RWG, SWD]; Loreto: [JTJ, USNM]; PA, Mera: [FSCA, JTJ]; Pastaza: [FSCA, JTJ, KJT, USNM]; Santa Clara: [FSCA]; SU, Cascales: [KJT]; Lago Agrio: [UTIC, KJT, RWG, USNM]; Shushufindi: [DRP, FSCA, KJT, USNM]; ZC, El Pangui: [FSCA, KJT]; Yantzaza: [FSCA, KJT, USNM]; Zamora: [JTJ, KJT, USNM]. Note: A specimen in the QZAC collection, labeled H. sanguinea, from Esmeraldas, San Lorenzo, IORI # 16475, is a mislabeled H. fuscoguttata Selys, 1878.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B509FFCDFF23F435206DF9EF.taxon	discussion	The six species recorded in Ecuador are metallic purple, usually with dark wing tips (Fig. 10). The nymphs, which resemble those of Hetaerina, are found in small forested streams, sometimes partly open; most of the species are unknown in the nymph stage. The genus was revised by Garrison (2006). Mnesarete devillei (Selys, 1880). Publ. Recs. ECU: Kirby (1890 — as Lais devillei), Campos (1922 — as L. devillei), Fraser (1946 a), Davies & Tobin (1984), Tsuda (1986, 1991), Paulson (1983), Steinmann (1997 a), (H 08), (P 11), (H 14); NA: * Selys (1880 — description), Tennessen (1996 b), Daigle (1996), Garrison (2006); PA: McLachlan (1881 a — as L. devillei). Also PER: (P 11); plus BOL: (P 16). Addl. Data. NA, Archidona: [RWG]; Tena: [FSCA, JJD, KJT, LSAM, QZAC, RWG, TWD, USNM]; OR, Loreto: [JTL, USNM]; PA, Arajuno: [RWG]; Pastaza: [RWG]. Mnesarete drepane Garrison, 2006. Publ. Recs. ECU: (H 08), (P 11), (H 14); OR: * Garrison (2006), von Ellenrieder & Garrison (2011); SU: * Garrison (2006 — Holotype: Shushufindi, Limoncocha on Río Napo (- 0.40, - 76.6167, 269 m), 3 xi 1980, leg. MJW [FSCA]; Allotype: same data but 8 xi 1980 (FSCA)]. Also COL: (H 08, P & P); plus PER: Garrison (2006), (P 11). Addl. Data. OR, Francisco de Orellana: [ACRP, BMNH, FSCA, JJD, KJT, NVE, RWG, USFQ, USNM]; Loreto: [JTJ, KJT]; SU, Lago Agrio: [USNM]; Shushufindi: [AEB, DRP, FAL, FSCA, KJT, RWG, USNM]. Mnesarete ephippium Garrison, 2006. Publ. Recs. ECU: (H 08), (P 11), (H 14); SU: * Garrison (2006, SU, Putumayo: 5 ♂♂, 5 ♀♀, Río Cuyabeno, 60 km E of Lago Agrío (- 0.0167, - 76.3167), 16 ii 1998, leg. G. Kareofelas, det. RWG (ACR, RWG]; ZC, Nangaritza: * Garrison (2006 — Holotype: “ Mayaicu ” as “ Morona-Santiago Province: Mayalico ” on Río Naganitza ” (- 3.0592, - 77.9956, 490 m) [corrected as Mayaicu, Río Nangaritza (- 4.0290, - 78.6340, 900 m), 2 xii 1941, leg. D. Laddey, det. RWG [UMMZ]; Allotype: same data but UMMZ). Also COL: (H 08); plus PER: Garrison (2006), (P 11). Addl. Data. NA, Tena: 1 ♂, about 5 km S of Jatun Sacha Road (- 1.0964, - 77.6418), 31 x 2013, leg. & det. KJT [FSCA]; SU, La Joya de los Sachas: 1 ♀, La Joya de los Sachas, E at stream (- 0.2066, - 76.6916, 284 m), 29 x 2013, leg. & det. C. Kerst [FSCA]. Mnesarete fulgida (Selys, 1879). Publ. Recs. ECU: Selys (1879), Kirby (1890), Calvert (1909), Campos (1922), Schmidt (1942), Fraser (1946 a — as Lais hauxwelli). Kimmins (1969 b — designated Lectotype in BMNH), Bridges (1993), Davies & Tobin (1984 — as synonym of M. hauxwelli), (H 14); NA: * Selys (1879 — Type Locality “ Río Napo ”, as Lais fulgida); Selys (1880); OR: Garrison (2006), von Ellenrieder & Garrison (2011). Also, COL, PER Garrison (2006), (H 08). Addl. Data. OR, Francisco de Orellana: (ACR, FSCA, KJT, LSAM, QZAC, RWG, USNM]; SU, Shushufindi: 1 ♂, stream about 3 km SW of Shushufindi (- 0.2136, - 76.6786), 12 v 2009, leg. & det. KJT [KJT]. Note: Garrison (2006) discussed its confused taxonomic history and states that it is a valid species distinct from M. hauxwelli. The type could have been from either Napo or Orellana Province. Mnesarete hauxwelli (Selys, 1869). Publ. Recs. ECU: Campos (1922 — “ por Dr. Martin ” as Lais hauxwelli), Schmidt (1942), Fraser (1946), Tsuda (1986, 1991), Paulson (1983), Steinmann (1997 a), (H 08), (P 11), H 14); ES: * Selys (1880); MS: Garrison (2006); NA: Garrison (2006), Daigle (1996); PA: Garrison (2006), Tennessen (2006); SU: Garrison (2006); TU: Garrison (2006); ZC: Garrison (2006). Also, PER: Garrison (2006), (H 08), (P 11); plus BRA, COL (P 11). Addl. Data. MS, Morona: [FAL, RWG]; Gualaquiza: [USNM]; NA, Archidona: [FSCA, JJD, KJT, RWG]; Tena: [RWG]; OR, Francisco de Orellana: [FSCA, USNM]; Loreto: [FSCA, FAL, KJT, RWG]; PA, Mera: [FSCA, KJT, USNM]; Pastaza: [FAL, FSCA, RWG, SWD]; Santa Clara: [RWG]; SU, Lago Agrio: [USNM]; Shushufindi: [RWG]; TU, Baños: [RWG]; ZC, Zamora: [RWG]. Note: Heckman (2008) commented that some recent illustrations and descriptions of this species refer to M. fulgida. Two specimens in the QZAC collected by WFM labeled “ M. acerifolia ms. ” should be H. hauxwelli. Mnesarete metallica (Selys, 1869). Publ. Recs. ECU: Fraser (1946 — as Mnesarete hincksi, description), Tsuda (1986, 1991), Paulson (1983), Steinmann (1997 a — as M. hincksi), (H 08), (P 11), (H 14); NA: Tennessen (1996 b), Daigle (1996), Garrison (2006); OR: von Ellenrieder & Garrison (2011); PA: * McLachlan (1881 a), Garrison (2006); SU: Garrison (2006). Also COL, VEN: Garrison (2006), (H 08, P 11). Addl. Data. NA, Tena: [DRP, FAL, FSCA, KJT, MNHM, RWG, TWD, UMMZ); OR, Francisco de Orellana: [FSCA, RWG]; Loreto: [FSCA, JJD, JTJ, KJT, RWG]; PA, Pastaza: [FSCA, KJT, QZAC, RWG, TWD RWG]; SU, Lago Agrio: [USNM]; Shushufindi: [DRP, FAL, RWG]. Note: Montgomery (1969) and De Marmels (1989) both stated that M. hincksi is the same as M. metallica.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B508FFCDFF23F0B42699F8F3.taxon	discussion	Only one of the four species in this genus has been recorded in Ecuador. The adults are more elongate than the other calopterygid genera and are largely metallic green (Fig. 11); females often have a reddish tinge in the wings. The nymphs inhabit small, high elevation forested streams often below waterfalls. The genus was revised by Garrison (2006), who also described the nymph of the one species in Ecuador.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B50FFFCAFF23F3A824AAF8AB.taxon	discussion	Coenagrionidae is the largest family of damselflies in the world with more than 130 genera and over 1200 species. These totals are based on recent molecular phylogenetic studies indicating that the Pseudostigmatidae and New World Protoneuridae belong in the Coenagrionidae (Dijkstra et al. 2014). As such, the family in the New World contains 71 genera and around 600 species. Most of the genera in the New World are endemic (> 90 %) to the Americas. These damselflies are extremely diverse in regard to size (total length ranging from 19 to 155 mm), color and color pattern, and they inhabit almost every type of water body, including lakes, ponds, ditches, streams, seepage areas and phytotelmata (water-containing tree holes and bromeliads). Much taxonomic and phylogenetic study remains to be done in this group.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B50FFFCAFF23F2172680FADA.taxon	description	Addl. Data. NA, Quijos: [DRP, FSCA, FAL, JJD, JTJ, KJT, RWG, TWD, USNM]; PA, Mera: [KJT, RWG]; Pastaza: [RWG]; PC, Quito: [RWG]; ZC, Zamora: [JTJ, RWG].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B50FFFF6FF23F1772132F82B.taxon	discussion	A large genus of slender, usually small damselflies, mostly black but usually with blue, green or orange stripes on the thorax and tip of the abdomen (Fig. 12). The appendages of males slant posteroventrally and the apical rim of abdominal segment 10 is often elevated; females usually have paired mesepisternal fossae. Habitat includes vegetated ponds, lake margins, and slow areas of streams. For decades, species identification relied on Leonard (1977), but numerous errors and synonymies have been found in that publication. For recent taxonomic changes and additions, see Lozano (2007, 2013), von Ellenrieder & Lozano (2008) and Machado (2012 a). The genus is in need of revision. Acanthagrion ascendens Calvert, 1909. Publ. Recs. ECU: Paulson (1983 “? ”), Tsuda (1986, 1991), (H 08), Lozano (2007), (H 14 — as A. cf. ascendens), (P 16); GY: * Campos (1922 — as A. gracile var ascendens, redetermined as A. floridense by KJT), Leonard (1977); OR: Francisco de Orellana: Tennessen (2004 — “ tentatively identified ”). PA: Gloger (1967). Also BRA, COL, GUF, GUY, SUR, TTO, VEN: (L 06), (P 11); plus BOL, PER (H 08), (P 16); and PRY, (H 08). Addl. Data. OR, Francisco de Orellana, 1 ♂, pond near Río Savaleto, km marker 31, P. N. Y., 17 vii 1996, leg. & det. KJT [FSCA]; adult, T. B. S., Río Tiputini, 55 km SE of Pompeya, 94 km ESE of Coca (- 0.6370, - 76.1500, 230 m), no date provided, leg. & det. WAH & DLW. Note: The ♂ examined has color variation — S 9 is orange, as discussed by Tennessen (2004). Note: A ♂ and ♀ from F. Campos taken at Tena were identified as A. ascendens. We checked the ♂ which is in FSCA and it is A. floridense. The ♀ was sent back to ANSP and may also be A. floridense. Acanthagrion apicale Selys, 1976. Publ. Recs. ECU: (L 06), (Lozano 2007), (H 08), (P 11), (H 14). NA: Machado (2012 a — as A. chicomendesi paratypes); OR: * Tennessen (1996 b), Muzón & Lozano (2005), von Ellenrieder & Garrison, (2011). Also BOL, BRA, COL, GUF, PER, VEN: (L 06), (H 08), (P 11); plus GUY: (H 08); and SUR: (P 16). Addl. Data. MS, Gualaquiza: 3 ♂♂, 3 ♀♀, Gualaquiza (- 3.40, - 78.58, 800 m), vi – v 1935, leg. F. Campos, det. MJW [FSCA]; OR, Francisco de Orellana: 4 ♂♂, 4 ♀♀, [La Nueva Esperanza] [across Río Napo from] near Primavera “ Limoncocha ” Río Napo & Lake Taracoa (- 0.45, - 76.77, 300 m), 27 vii 1977, leg. DRP & S. Hills, det. DRP [DRP]; 3 ♂♂, same data but 4 xi 1980, leg. & det. MJW [FSCA]; 1 ♂, same data but swamp N of Lake, 26 viii 1980, leg. & det. SWD [SWD]; 1 ♂, [Limoncocha] [about 40 km east] or 100 km E of Coca [on Río Napo] Lago Garzacocha, swamp, La Selva Jungle Lodge (- 0.4819, - 76.3733, 235 m), 12 x 1988, leg. & det. SWD [SWD]; 1 ♂, [Pompeya Sur] S on [Pompeya Sur-Iro road] at km marker 8, swamp & stream (- 0.4694, - 76.5869, 253 m), 12 vi 1995, leg. & det. WFM [QZAC]; 7 ♂♂, [Pompeya Sur] S on [Pompeya Sur-Iro road] at km marker 31, borrow pit near Río Sabaleto (- 0.60, - 76.47, 229 m), 17 vii 1996, leg. & det. WFM [FSCA, LSAM, QZAC, USNM]; 1 ♂, 1 ♀, same data but leg. & det. KJT [KJT]; 1 ♂, swamp pond, km marker 38 on Yasuni Road 12 vi 1995, leg. & det. KJT [KJT]; 5 ♂♂, open & forest swamps, main road, P. N. Y., 15 xi 1997, leg. & det. KJT [KJT]; 1 ♂, just S of Bogi 2 Este River, S of km 42 on Yasuni Road, swamp (- 0.38, - 76.30), 11 vi 1995, leg. & det. SWD [SWD]; 3 ♂♂, P. N. Y., 2.1 km E of Yasuni Biological Research sta., pond and swamp (- 0.6741, - 76.3685, 244 m), 15 xi 1997, leg. & det. WFM [FSCA, QZAC, USNM]; 1 ♂, stream, road to Waorani Reserve, 5.6 km S of Yasuni Sci. Res. Sta., 10 vi 1995, leg. & det. KJT [KJT]; 1 ♂, same data but leg. & det. WFM [FSCA]; 1 ♀, P. N. Y., 6.8 km E of Yasuni Biological Research Sta., small forested stream (- 0.6545, - 76.3417, 238 m), 15 xi 1997, leg. & det. WFM [QZAC]; 1 ♂, P. N. Y., stream (- 0.6542, - 76.3542, 200 m), 15 xi 1997, leg. & det. TWD [SWD]; 1 ♂, P. N. Y., swamp (- 0.6508, - 76.3558, 200 m), 16 xi 1997, leg. & det. TWD [SWD]; 1 ♂, P. N. Y., stream (- 0.7100, - 76.4750, 200 m), 16 xi 1997, leg. & det. TWD [SWD]; 10 ♂♂, 1 ♀, T. B. S., USFQ, black water stream crossing, Guacamayo trail at marker 2050 (- 0.6356, - 76.1492, 220 m), 15 – 21 i 2009, leg. & det. RWG & NVE [MLPA, NVE, RWG, USFQ]; 1 ♀, T. B. S., USFQ, swamp at marker 650 of Parahuaco trail (- 0.6397, - 76.1453, 229 m), 21 i 2009, leg. & det. RWG & NVE [NVE]; 1 ♂, T. B. S., USFQ, stream crossing Matapalo trail at marker 450, Ti 15 (- 0.6369, - 76.1458, 245 m), 15 i 2009, leg. & det. RWG & NVE [RWG]; SU, Shushufindi: 1 ♂, Limoncocha, 25 viii 1980, leg. & det. SWD [SWD]; 1 ♀, Limoncocha, Río Napo (- 0.40, 76.62, 300 m), 21 ix 1974, leg. B. Drummond III, det. SWD [FSCA]; 1 ♀, Limoncocha, swamp, 26 vii 1977, leg. DRP & S. Hills, det. DRP [DRP]; 1 ♂, trails, small streams 3 km W of Shushufindi, 2 + km N off Hwy. 26 at km marker 17 (- 0.1664, - 76.6917, 290 m), 13 x 2009, leg. & det. JTJ [JTJ]; 1 ♂, same data but leg. & det. KJT [KJT]. Note: Machado (2012 a) split A. apicale into several species. After discussion with and evidence provided by Rosser Garrison (pers. comm., 2017), we recognize A. apicale as the species in Ecuador and A. chicomendesi as a possible junior synonym of A. apicale. Acanthagrion cuyabae Calvert, 1909. Publ. Recs. ECU: (L 06 — also as A. lancea), Lozano (2007 — also as A. lancea), (H 08 — also as A. lancea), Rojas & Sanchez (2009 — as A. viridescens), (P 11 — also as A. lancea) (P 11 — as A. viridescens), (H 14 — as both A. lancea and A. viridescens). OR: * Tennessen (1996 b — as A. lancea), von Ellenrie- der & Garrison (2011 — as A. lancea). Also BOL, BRA: (L 06), (H 08), (P 11); plus COL — as A. viridescens, PER (P 16). Addl. Data. NA, El Chaco: 1 ♂, Chaco, 53 km NE on road to Lago Agrio, ponds and stream, 18 viii 1980, leg. & det. SWD as A. lancea [SWD]; OR, Francisco de Orellana: 1 ♂, [La Nueva Esperanza] [across Río Napo from] near Primavera, Laguna Taracoa (- 0.45, - 76.77, 300 m), 27 vii 1977, leg. DRP & S. Hills, det. DRP as A. viridescens [DRP]; 2 ♂♂, same data but 4 xi 1980, leg. MJW & D. Robinson, det. MJW as A lancea [FSCA]; 4 ♂, same data but 26 viii 1980, leg. & det. SWD as A. lancea [SWD]; 3 ♂♂, P. N. Y., 6.3 km S of Yasuni Road on Waorani Reserve Road, marshy pond (- 0.7120, - 76.4745, 260 m), 10 vi 1995, leg. & det. WFM as A. lancea [FSCA, LSAM, QZAC, SWD]; 4 ♂♂, same data but leg. & det. KJT as A. viridescens [KJT]; 2 ♂♂, swamp pond at km 8, S of Río Napo on Yasuni Road (- 0.6333, - 76.5000), 12 vi 1995 leg. & det. SWD as A. lancea [RWG]; 3 ♂♂, same data but leg. & det. KJT as A. viridescens [KJT]; 1 cop. pair, P. N. Y., 29 km N of Waorani Road on Yasuni Road roadside ditch and pond (- 0.4917, - 76.5584, 183 m), 11 vi 1995, leg. & det. KJT as A. viridescens [KJT]; 1 ♂, 1 ♀, same data but leg. & det. WFM as A. lancea [FSCA]; 2 ♂♂, [Pompeya Sur], S on [Pompeya Sur-Iro road] at km marker 31, borrow pit near Río Sabaleto (- 0.60, - 76.47, 229 m), 12 vi 1995, leg. & det. WFM as A. lancea [FSCA]; 1 cop. pair, same data but 17 vii 1996, leg. & det. KJT as A. viridescens [KJT]; 3 ♂♂, same data but leg. & det. SWD as A. lancea [SWD]; 2 cop. pairs, 2 ♂♂, same data but leg. & det. WFM as A. lancea [FSCA, QZAC, USNM]; 1 ♂, same data but 17 xi 1997 [FSCA]; 1 ♂, 1 ♀, same data but leg. & det. JJD as A. viridescens [DRP, JJD]; 3 ♂♂, 1 ♀, same data but leg. & det. TWD as A. viridescens [SWD]; 2 cop. pairs, 2 ♂♂, Puerto Francisco de Orellana, 12.3 km W on road to Loreto from Coca-Sachas Road, pond (- 0.4750, - 77.0747, 290 m), 13 vi 1995, leg. & det. WFM as A. lancea [FSCA, USNM]; 1 ♂, same data but leg. & det. SWD as A. lancea [UTIC]; 2 ♂♂, 1 ♀, Francisco de Orellana, 1.7 km W on Loreto Road from Coca-Sachas Road, stream, pools & flowing shallow marsh (- 0.4367, - 77.0017, 258 m), 13 vi 1995, leg. & det. WFM as A. lancea [FSCA, QZAC, USNM]; 1 ♂, same data but 18 xi 1997 [FSCA]; 4 ♂♂, Puerto Francisco de Orellana 0.6 km W on road to Loreto from Río Payamino, marsh (- 0.4347, - 77.0011, 200 m), 20 vii 1996, leg. & det. WFM as A. lancea [FSCA, QZAC, USNM]; 1 ♂, same data but 18 xi 1997 [FSCA]; 2 ♂♂, same data but leg. & det. TWD as A. viridescens [FSCA, SWD]; SU, Shushufindi: 6 ♂♂, Limoncocha (- 0.40, - 76.62, 300 m), 27 xii 1971, leg. D. Pearson, det. DRP as A. viridescens [DRP]; 1 ♀, same data but 6 i 1972 [DRP]; 2 ♂♂, 11 – 16 viii 1972 [DRP]; 6 ♂♂, same data but 3 – 16 vi 1977, leg. P. Spangler & D. Givens, det.? as A. lancea [USNM]; 3 ♂♂, same data but 23 viii 1980, leg. & det. SWD as A. lancea [SWD]; 2 ♂♂, same data but 25 viii 1980 [SWD]; 1 ♂, same data but 14 v 1975, leg. T. Emmel, det. MJW as A. lancea [FSCA]; 1 ♀, same data but 23 viii 1980, leg. SWD, det. KJT as A. viridescns [KJT]; 16 cop. pairs, 62 ♂♂, 4 ♀♀, same data but 1 – 27 xi 1980, leg. MJW & M. Robertson, det. MJW as A. lancea [FSCA]; 2 cop. pair, 52 ♂♂, 10 ♀♀, same data but Limoncocha, lakeshore, 23 – 26 vii 1977, leg. & det. DRP as A. viridescens [DRP]; 3 ♂♂, [Limoncocha] [about 40 km E] 100 km E of Coca [on Río Napo], Lago Garzacocha, La Selva Jungle Lodge (- 0.4819, - 76.3733, 235 m), 12 x 1988, leg. & det. SWD as A. lancea [FSCA, SWD]; 1 ♂, same data but 23 xi 1997, leg. & det. TWD as A. viridescens [SWD]; 1 ♂, La Selva, Heron Lake, 4 v 1990, leg. E Smith, det. WFM [SWD]. Note: Leonard (1977) considered A. cuyabae and its two subspecies as synonyms of A. lancea Selys, 1876. Federico Lozano (pers. comm., 2019) found that A. cuyabae is a valid species and that A. viridescens is a synonym of A. cuyabae. Most published records and many specimens in various collections from Ecuador are misidentified as either A. viridescens or A. lancea; probably all such records should be referred to A. cuyabae. Acanthagrion floridense Fraser, 1946. Publ. Recs. ECU: Paulson (1983), Tsuda (1986, 1991), (L 06), Lozano (2007 — also as A. gracile), (H 08), Rojas & Sanchez (2009), (P 11), (H 14). NA: Gloger (1967 — as A. gracile); Leonard (1977 — as A. peruvianum); OR: Tennessen (1996 b), von Ellenrieder & Garrison (2011 — as A. peruvianum); PA: * Leonard (1977); ZC: Tennessen (2000), Tennessen & Johnson (2008). Also ARG, PER: (L 06), (H 08), (P 11); plus COL: (P 11, P & P); and BRA: (H 08); and URY: (L 06), (P 11). Addl. Data. LJ, Loja: [UTIC]; MS, Santiago: [FSCA, TWD, USNM]; NA, Carlos Julio Arosemena Tola: [USNM; El Chaco: [SWD]; Tena: [DRP, FSCA, UTIC, JTJ, KJT, RWG, SWD, TWD, USNM]; OR, Francisco de Orellana: [DRP, FSCA, UTIC, KJT, RWG, SWD, TWD, USNM]; Loreto: [FSCA, KJT, SWD, SWD]; PA, Mera: [USNM]; Pastaza: [USNM]; Santa Clara: [USNM]; SU, Cascales: [SWD, USNM]; Lago Agrio: [FSCA, KJT, SWD, USNM]; Shushufindi: [FAL, FSCA, KJT, SWD, USNM]; TU, Baños: [DRP, RWG]; ZC, El Pangui: [FSCA]; Yantzaza: [FSCA, USNM]; Zamora: [FSCA, UTIC, JTJ, KJT, USNM]. Note: All the published records and a majority of the specimens in various museums under “ Addl. Data ” above are labeled A. peruvianum which has only recently been determined as a junior synonym of A. floridense (Lozano et al. 2017). A ♂ and ♀ from F. Campos collected at Tena were labeled as A. ascendens; we checked the ♂ which is in FSCA and it is A. floridense. The ♀ was sent back to ANSP and may be labeled A. ascendens but it is probably A. floridense also. Acanthagrion longispinosum Leonard, 1977. Publ Recs. BOL, BRA (H 08). Addl. Data. OR, Francisco de Orellana: 1 ♀, T. B. S. Río Tiputini, 55 km southeast of Pompeya, 94 km ESE of Coca (- 0.6370, - 76.1500), 21 i 2012, leg. DLW, John Ascher, WAH, Fred Morrison, det. RWG, data provided by RWG. Acanthagrion obsoletum (Förster, 1914). Publ. Recs. ECU: Paulson (1983), Tsuda (1986, 1991), Lozano (2007 — also as A. hartei), (P 11 — also as A. hartei), (H 14 — also as A. hartei); MS: * Muzón & Lozano (2005 — as A. hartei, Holotype & Allotype: Bomboiza, 17 km S of Gualaquiza (- 3.4333, - 78.5167, 880 m), 10 ix 1990, leg. OSF [USNM]); NA: * Förster (1914 — type locality: “ Napo ”), Bridges (1993), Gloger (1967 — as A. leonora); Leonard (1977), Muzón & Lozano (2005), Machado (2012 a); OR: Tennessen (1996 b), von Ellenrieder & Garrison (2011); PA: * Förster (1914 — ALLOTYPE), Leonard (1977); SU: * Muzón & Lozano (2005 — as A. hartei); ZC: Tennessen (2000), Tennessen & Johnson (2008 — as A. hartei). Also BOL, COL, PER: (H 08), (P 11). Addl. Data. MS, Huamboya: [FSCA, KJT, USNM]; Morona: [JJD]; Santiago: [FSCA, SWD]; NA, Carlos Julio Arsemena Tola: [FSCA, USNM]; El Chaco: [SWD]; Tena: [DRP, FSCA, UTIC, KJT, RWG, SWD, TWD, USNM]; OR, Francisco de Orellana: [DRP, FAL, FSCA, KJT, RWG, SWD, TWD, USNM]; SU, Lago Agrio: [FSCA, KJT, RWG, SWD]; Shushufindi: [DRP, FSCA, KJT, SWD]; PA, Pastaza: [FSCA, JTJ, KJT]; Santa Clara: [FSCA, UTIC, RWG, USNM]; ZC, El Pangui: [FSCA]; Yantzaza: [FSCA, USNM]; Zamora: [FSCA, UTIC, USNM]. Note: According to Garrison (pers. comm. 2017), A. hartei Muzón & Lozano is a junior synonym of A. obsoletum. Acanthagrion phallicorne Leonard, 1977. Publ. Recs. ECU: * (L 06), Lozano (2007), (P 11), (H 14). Also BRA, PER: (L 06), (H 08), (P 11). Addl. Data. NA, Tena: 1 ♂, Río Chinchipino, oxbow pond, 27 km E of Puerto Napo (- 1.0728, - 77.5864, 385 m), 13 xi 1997, leg. & det. KJT [KJT]; OR, Francisco de Orellana: 1 ♂, [Pompeya Sur] S on [Pompeya Sur-Iro road] at km marker 31, borrow pit near Río Sabaleto [road to Yasuni Research station] (- 0.60, - 76.47, 230 m), 12 vi 1995, leg. & det. WFM [FSCA]. Acanthagrion trilobatum Lenoard, 1977. Publ. Recs. ECU: Lozano (2007), (P 11), (H 14). SD: * Daigle et al. (1997); ES: * Daigle et al. (1997); IM: * Daigle et al. (1997); LJ: Tennessen & Johnson (2008). Also CEN, COL, VEN: (H 08), (P 11). Addl. Data. BO, Caluma: [FSCA, KJT]; CT, Pangua: [FSCA, JTJ, KJT]; EO, Arenillas: [FSCA, KJT]; Santa Rosa: [FSCA, JTJ, KJT]; ES, San Lorenzo: [FAL, FSCA, KJT, USNM]; GY, Balao: [FSCA, JTJ, KJT]; Naranjal: [FSCA]; LJ, Loja: [FSCA, KJT]; LR, Buena Fé: [DRP, FAL, SWD, USNM]; MN, El Carmen: [FSCA, JTJ, KJT]; Flavio Alfaro: [JTJ]; PC, Quito: [FSCA, KJT, USFQ]; San Miguel Bancos: [FSCA, KJT, RWG, USFQ]; SD: [FSCA, JJD, JTJ, KJT, SWD, USNM]. Acanthagrion yungarum Ris, 1918. Publ. Recs. ECU: Lozano (2007), (P 11), (H 14); NA: Tennessen (1996 b), * Daigle (1996); ZC: Tennessen & Johnson (2008). Also PER: (H 08), (P 11); plus COL: (P 16). Addl. Data. MS, Gualaquiza: 1 ♂, 1 ♀, Gualaquiza (- 3.40, 78.58, 800 m), iv – v 1935, leg. F. Campos, det. KJT 2011 — labeled A. gracile by P. Calvert, [ANSP, FSCA]; Morona: 1 cop. pair, 1 ♂, 1 ♀, Río Copueno, E side of Macas (- 2.3019, - 78.1133), 17 ix 2005, leg. & det. KJT [KJT]; 2 ♂♂, same data but leg. & det. JJD [JJD]; NA, Archidona: 1 ♂, [Mondayacu], 28 km N of Tena on road between Tena & Baeza (- 0.80, - 77.77, 884 m), 27 x 1980, leg. & det. MJW [FSCA]; 2 cop. pairs, 9 ♂♂, 1 ♀, [Osayacu] 33 km N of Tena on road between Tena & Baeza (0.78, - 77.78, 975 m), 27 x 1980, leg. & det. MJW [FSCA]; 1 ♂, 1 cop. pair, [Narupa] E on road to Loreto, 43.5 km W of Loreto, seep near large rocky river [Río Hollin] (- 0.7022, - 77.7189, 1036 m), 14 vi 1995, leg. & det. WFM [FSCA, USNM]; 2 ♂♂, same data but 15 vii 1996 [QZAC, USNM]; 2 ♂♂, same data but leg. & det. SWD [SWD]; 2 ♂♂, [Narupa] E on road to Loreto, 43.5 km W of Loreto, seep near large rocky river [Río Hollin] (- 0.7022, - 77.7189, 1036 m), 14 xi 1997, leg. & det. WFM [FSCA]; 1 ♂, pool, Loreto Road, 44 km E of Baeza / Tena Road, 15 vii 1996, leg. & det. KJT [KJT]; 1 ♂, km 45 on Hollen-Loreto Road (- 0.7228, - 77.5653, 1130 m), 22 xii 1987, leg. M. & J. Wasbauer, det. RWG [RWG]; 2 ♂♂, [Narupa] E off road to Loreto, 1 km NW Bridge, Río Hollin (- 0.6875, - 77.7333, 914 m), 4 xi 2001, leg. & det. F. Sibley [FSCA]; El Chaco: 12 ♂♂, 1 ♀, El Chaco 28.3 km N on Hwy. 45 near Salado, adjacent to Río Quijos, marsh and borrow pit (- 0.1831, - 77.6789, 1237 m), 7 vi 1995, leg. & det. WFM [FSCA, QZAC, USNM]; 3 ♂♂, same data but leg. & det. SWD [RWG, SWD]; 5 ♂♂, 1 ♀, same data but leg. & det. KJT [KJT]; OR, Loreto: 1 ♂, Loreto, 30.7 km W on road to Narupa, trickle (- 0.7381, - 77.5119, 920 m), 14 vi 1995, leg. & det. WFM [FSCA]; 2 ♂♂, pond & small stream, 34 km W of Loreto (835 m), 14 vi 1995, leg. & det. SWD [FAL]; 1 cop. pair, 5 ♂♂, same data but leg. & det. WFM [FSCA, LSAM, QZAC, USNM]; 2 ♂♂, same data but leg. & det. KJT [KJT]; 1 cop. pair, 2 ♂♂, Loreto, 38.9 km W on road to Narupa, pond (- 0.7411, - 77.5569, 988 m), 14 vi 1995, leg. & det. WFM [FSCA, USNM]; 1 ♂, same data but leg. & det. KJT [KJT]; 3 ♂♂, same data but leg. & det. SWD [SWD]; 1 ♂, Loreto, 35.2 W on road to Narupa, pond (- 0.7414, - 77.5481, 927 m), 19 xi 1997, leg. & det. WFM [FSCA]; 1 ♂, same data but leg. & det. KJT [KJT] 1 ♂, same data but leg. & det. TWD [FSCA]; 6 ♂♂, 2 ♀♀, pond along Loreto-Archidona road (- 0.7417, - 77.5517, 900 m), 19 xi 1997, leg. & det. TWD [SWD]; PA, Mera: 1 ♂, Shell (- 1.52, - 78.07, 914 m), 7 xi 2001, leg. & det. F. Sibley [FSCA]; Pastaza: 1 ♂, 1 ♀, pond near Comunidad Chuwitayo, S of Puyo, 9 xi 1997, leg. & det. KJT [KJT]; 1 ♂, 1 ♀, pond 3 km N of Comunidad Chu- witayo, S of Puyo, 9 xi 1997, leg. & det. KJT [KJT]; 1 ♂, pond, 8.7 km S of Veracruz, Hwy. 45, 11 xi 1997, leg. & det. KJT [KJT]; 1 ♂, pond, about 4 km SE off Hwy. 45, N of Puyo (- 1.3422, - 77.8681, 945 m), 8 x 2009, leg. & det. KJT [KJT]; 2 ♂♂, same data but leg. & det. JTJ [JTJ]; 2 ♂♂, Puyo, 16 km N on Tena Road (Hwy. 45), near Teniente Hugo Ortiz, small stream [listed at 2000 feet], (- 1.3475, - 77.9183, 1036 m), 26 x 1980, leg. & det. MJW [FSCA]; 2 ♂♂, Cuasha, about 1.5 km N on Hwy. 45 [91.5 km N of Sucua], hard bottomed pond (- 1.85, - 77.82, 884 m), 9 xi 1997, leg. & det. WFM [FSCA, QZAC]; 2 tan. pairs, 3 ♂♂, Cuasha approx. 4 km N on Hwy. 45 [94 km N of Sucua], cut grass pond (- 1.83, - 77.82, 884 m), 9 xi 1997, leg. & det. WFM [FSCA, QZAC, USNM]; 2 ♂♂, pond on road between Macas and Puyo (- 1.8417, - 77.8283, 900 m), 9 xi 1997, leg. & det. TWD [SWD]; 1 ♂, 1 ♀, 12 km E of Puyo, 2 ii 1976, leg. P. Spangler et al., det.? [USNM]; Santa Clara: 1 ♂, 1 ♀, 39 km NE of Puyo, 5 xi 1977, leg. P. Spangler & D. Givens, det.? [USNM]; SU, Gonzalo Pizarro: 1 ♂, 65 km W of Lago Agrio, roadside pond, 18 v 1975, leg. A Gurney, det.? [USNM]; ZC, Zamora: 1 ♂, roadside ditch, 2.1 km ENE of Zamora (915 m), 5 xi 1997, leg. & det. KJT [KJT]; 1 ♂, pool near Río Sabanillo, 10 km W of Zamora (1220 m), 5 xi 1997, leg. & det. KJT [KJT]; 1 ♂, tilapia ponds and small streams, Hosteria El Arenal, about 2 km S of Cumbaratza (- 4.0017, - 78.8753, 950 m), 2 iv 2008, leg. & det. KJT [KJT]; 1 ♂, same data but leg. & det. JTJ [JTJ].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B532FFF7FF23F6882035FA30.taxon	discussion	The small damselflies in this genus are rarely seen. Adults have the basal half of the wings dark (Fig. 13). See Williamson & Williamson (1924) for identification. Habitat is small streams in Amazonian forest. Acanthallagma caeruleum Williamson & Williamson, 1924. Publ. Recs. ECU: (L 06), (H 08), (P 11), (H 14); NA: * Tennessen (1996 b). Also BRA: (L 06), (H 08), (P 11); plus PER: (P 16). Addl. Data. OR, Francisco de Orellana: 1 ♀, Puerto Francisco de Orellana, 1.7 km W on Loreto Road from Coca-Sachas Road, stream, pools & flowing shallow marsh (- 0.4367, 77.0017, 258 m), 13 vi 1995, leg. & det. WFM [FSCA]; 1 ♂, same data but leg. & det. KJT [KJT]; 1 ♀, La Nueva Esperanza, across Río Napo from near Primavera “ Limoncocha ”, Río Napo & Lake Taracoa (- 0.45, - 76.77, 300 m), 4 xi 1980, leg. & det. MJW [FSCA]; 1 ♂, 2 ♀♀, Primavera, Laguna Taracoa, 27 vii 1977, leg. & det. DRP [DRP]; SU, Lago Agrio: 2 ♂♂, 3 ♀♀, stream 24 km W of Lago Agrio, brushy ponds on stream (- 0.12, - 77.071, 335 m), 20 viii 1980, leg. & det. SWD [RWG]; Shushufindi: 1 ♂, Shushufindi, 2 km S of Yamaramsuku School on road to Limoncocha, stream & marsh (- 0.2147, - 76.6494, 255 m), 19 vii 1996, leg. & det. WFM [QZAC]; 1 ♂, same data but leg. & det. KJT [KJT]; Río Sin Nombre, 5 km SE of Shushufindi on road to Limoncocha (- 0.23, - 76.65, 265 m), 18 i 2011, leg. DLW, det. WAH (WAH).	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B532FFF5FF23F3CD2639FD4F.taxon	discussion	Small to medium-sized brown or black damselflies with pale markings usually blue (Fig. 14). Adults perch mostly in the shade along ponds or backwaters of streams. For species identification, see Dunkle (1991 a) and Tennessen (2009). Aeolagrion axine Dunkle, 1991. Publ. Recs. ECU: Tsuda (1991), (H 08), (P 11), (H 14); OR: * Dunkle (1991 a — Holotype: “ Napo, Primavera ” [Francisco de Orellana], [La Nueva Esperanza] S of [across Río Napo from] Primavera, Laguna Taracoa (- 00.45, - 76.77, 244 m), [FSCA]), Bridges (1993), Tennessen (1996 b), Daigle (1996), Tennessen (2009). Also PER (P 16). Addl. Data. NA, Carlos Julio Arosemena Tola: 1 ♂, Río Ila, Pupuyacu (- 1.2000, - 77.8333, 500 m), xi 1934, leg. D. Laddey, det. RWG, from UMMZ [RWG]; Tena: 1 ♂, J. S. B. S., forest near stream (450 m), 30 xi 1994, leg. A. Jaramillo A, det. DRP [DRP]; 1 paratype ♂, pond near J. S. B. S., 23 km E of Puerto Napo (- 1.0667, - 77.6167, 400 m), 18 x 1988, leg. & det. SWD [SWD]; 2 ♂♂, 3.2 km E of Tena Road on road to J. S. B. S. Road, sloughs along Napo River, 23 vii 1996, leg. & det. SWD [SWD]; 1 ♂, Puerto Napo, 5.3 km E of Hwy. 45 on J. S. B. S. Road, Río Sinde (- 1.0500, - 77.7481, 450 m), 16 x 1988, leg. & det. SWD [RWG]; 1 ♂, same data but 15 vii 1995 [SWD]; 13 ♂♂, same data but 13 vii 1996, leg. & det. WFM [FSCA, LSAM, QZAC, USNM]; 4 ♂♂, 1 ♀, same data but leg. & det. JJD [JJD]; 5 ♂♂, same data but leg. & det. KJT [FAL, KJT]; 1 ♂, same data but leg. & det. SWD [RWG]; 1 cop. pair, 2 ♂♂, same data but 23 vii 1996, leg. & det. KJT [KJT]; 6 ♂♂, same data but 12 xi 1997 [KJT, RWG]; 7 ♂♂, same data but leg. & det. WFM [FSCA, QZAC, USNM]; 2 ♂♂, same data but leg. & det. TWD [SWD]; 1 ♂, same data but leg. & det. JJD [DRP]; 1 ♂, Río Chinchipino, oxbow pond, 27 km E of Puerto Napo (- 1.0728, - 77.5864, 385 m), 13 xi 1997, leg. & det. KJT [KJT]; OR, Francisco de Orellana: 3 ♂♂, 1 ♀, [La Nueva Esperanza] S of [across Río Napo from] Primavera Laguna Taracoa, (- 0.45, - 76.77, 244 m), 27 vii 1977, leg. & det. DRP [DRP]; 3 ♂♂, same data but forest trail, leg. DRP & S. Hills, det. DRP [DRP]; 1 paratype ♂, same data but 26 viii 1980, leg. & det. SWD [SWD]; 7 ♂♂, 1 ♀, same data but 4 xi 1980, leg. & det. MJW [FSCA]; 1 paratype ♂, same data but leg. MJW & D. Robinson, det. SWD [RWG]; SU, Shushufindi: 2 ♂♂, Limoncocha, forest stream, 24 vii 1977, leg. DRP, det. SWD [DRP]; 1 ♂, Limoncocha on Río Napo (- 0.4000, - 76.6167, 269 m), 7 xi 1977, leg. KWK, det. SWD [KWK]; 1 ♀ (labeled “ Allotype ” but not the one used by Tennessen, 2009), [Limoncocha] [about 40 km E] or 100 km E of Coca [on Río Napo], Lago Garzacocha, La Selva Jungle Lodge (- 0.4819, - 76.3733, 235 m), 12 x 1988, leg. & det. SWD [SWD]. Aeolagrion dorsale (Burmeister, 1839). Publ. Recs. ECU: * (L 06), (H 08), (P 11), (H 14), Bota-Sierra et al. (2015); SU: Tennessen (2009). Also BRA, PER, SUR, VEN, TTO: (L 06), (H 08), (P 11); plus BOL: (L 06), (H 08), Tennessen (2009); and GUF (P 16); and COL: Palacino et al. (2017). Addl. Data. OR, La Joya de los Sachas: 4 ♂♂, Pompeya, 0.5 km W of Puerto Pompeya ferry landing, ditches (- 0.4444, - 76.6263, 238 m), 19 vii 1996, leg. & det. WFM [LSAM, QZAC, USNM]; 7 ♂♂, same data but leg. & det. KJT (KJT, FSCA]; 4 ♂♂, same data but leg. & det. JJD [FSCA, JJD]; 1 ♂, same data but leg. & det. SWD [SWD]; SU, Lago Agrio: 5 ♂♂, swamp, 17 km S of Lago Agrio, 19 viii 1980, leg. & det. SWD [KJT, SWD]; Shushufindi: 1 ♂, Limoncocha, 23 viii 1980, leg. & det. SWD; 2 ♂♂, Limoncocha, lakeshore and nearby areas, 23 viii 1978, leg. KWK, det. SWD [SWD]. Aeolagrion inca (Selys, 1876). Publ. Recs. ECU: * Tsuda (1991 — as A. foliaceum), Daigle (1996), (L 06), (H 08), (P 11), (H 14). Also BOL, BRA, PER: (L 06), (H 08), (P 11); plus COL: (P 16). Addl. Data. SU, Shushufindi: 8 ♂♂, 3 ♀♀, Limoncocha, lakeshore, 23 vii 1977, leg. & det. DRP [DRP]; 1 ♂, same data but 24 vii 1977, 4 ♂♂, same data but 26 vii 1977.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B530FFF5FF23F52A26D7F9F4.taxon	discussion	Medium-sized, green and black damselflies with a pale preapical abdominal tip (Fig. 15) that inhabit marsh and seepage areas thick with aquatic plants; the thorax and anterior abdominal segments of immature individuals are mostly orange. For species identification see De Marmels & Garrison (2005); Tennessen (2012 b) described the nymph of A. inornatum. Anisagrion inornatum (Selys, 1876). Publ. Recs. ECU: (H 14); LJ: * De Marmels & Garrison (2005), Tennessen & Johnson (2008), Tennessen (2012 b — nymph description). Also VEN: (H 08), (P 11); plus COL: Rojas-Riano (2011 b), (P 16). Addl. Data. AZ, Santa Isabel: 1 ♂, 1 ♀, stream 21 km E of Hwy. 25, road from Balao jct. (- 2.9400, - 79.5653, 730 m), 11 iv 2008, leg. & det. KJT [FSCA]; 2 ♂♂, 2 ♀♀, same data but leg. & det. JTJ [JTJ]; EO, Piñas: 1 ♂, 15 km E Piñas (- 3.6667, - 79.65, 876 m), 4 vii 1989, leg. R. Miller & L. Stange, det. RWG [RWG]; LJ, Catamayo: 1 ♂, 2 ♀♀, trickle, 23 km from Hwy. 50 on road to Zaruma (- 3.8917, - 79.5511, 1095 m), 7 iv 2008, leg. & det. KJT [FSCA]; 1 ♂, 1 ♀, same data but leg. & det. JTJ [JTJ]; 1 ♂, stream about 18 km from Hwy. 50 on road to Zaruma (- 3.9092, - 79.5912, 1160 m), 7 iv 2008, leg. & det. JTJ [JTJ]; 1 ♂, small seep between Catamayo and Zaruma (- 3.9472, - 79.3958, 1276 m), 6 iv 2008, leg. & det. KJT [KJT]; Loja: 1 ♂, 3 ♀♀, Río Chamba at Vilcabamba (- 4.2625, - 79.2192), 16 – 17 viii 2006, leg. & det. JCA [UTIC]; 7 ♂♂, 9 ♀♀, seep, trib. Río Chamba, Vilcabamba (- 4.2636, - 79.2172, 1555 m), 30 iii 2008, leg. & det. KJT [FSCA]; 1 ♂, 1 ♀, same data but 31 iii 2008 [FSCA]; 6 ♂♂, 3 ♀♀, same data but 30 iii 2008, leg. & det. JTJ [JTJ]; 2 ♂♂, Vilcabamba, Natural Reserve Rumi-Wilco, floodplain trails along Río Chamba (- 4.2558, - 79.2197, 1530 m), 31 iii 2008, leg. & det. JTJ [JTJ]; 1 ♂, Malacatos, 15 mi S, 3 mi W of Loja- (- 4.2333, - 79.2833, 1500 m), 18 viii 1941, leg. D. Laddey, det. RWG, from UMMZ [RWG]; 1 ♀, same data but 23 viii 1941 [RWG]; 5 ♂♂, 4 ♀♀, same data but 24 viii 1941 [RWG].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B530FFFAFF23F08A21B3F9EF.taxon	discussion	Argia is the largest genus of damselflies in the New World, with over 138 valid species names and at least seven species yet to described (Garrison & von Ellenrieder 2018). Adults range from small to large and are variable in color pattern, the thoracic dorsum being black with blue or purple stripes (rarely orange) to entirely metallic red or copper, the thoracic sides usually blue, and the abdomen from mostly blue with black markings to mostly black with small blue markings (Figs. 16 a & 16 b), rarely largely orange. Species identification relies on Calvert (1902), and Garrison & von Ellenrieder (2015, 2017, 2018). Thirty-two species have been recorded in Ecuador, of which three are considered endemic. Habitat of most species in Ecuador is small streams to mid-sized rivers, with a few found in pooled areas of slow streams, seepage streams or sheet flow over boulders. Argia acridens Garrison & von Ellenrieder, 2018. Publ. Recs. ES: * Garrison & von Ellenrieder (2018); MN: * Garrison & von Ellenrieder (2018 — Holotype: Flavio-Alfaro Canton: 79 km west of Santo Domingo de los Colorados (- 0.3333, - 79.6666, 260 m), 7 v 1975, leg. Paul J. Spangler et al. [USNM]); SD: * Garrison & von Ellenrieder (2018). Note: Endemic to Ecuador’s Pacific slope. Argia adamsi Calvert, 1902. Publ. Recs. ECU: (P 11), (H 14); LR: Garrison & von Ellenrieder (2018); MN: Garrison & von Ellenrieder (2018); MS: Garrison & von Ellenrieder (2018); NA: * Tennessen (1996 b), Daigle (1996), Garrison & von Ellenrieder (2018); OR: Garrison & von Ellenrieder (2018); PA: Garrison & von Ellenrieder (2018); PC: Garrison & von Ellenrieder (2018); SD: Garrison & von Ellenrieder (2018); TU: Garrison & von Ellenrieder (2018). Also CEN, PER, VEN: (H 08), (P 11); Plus COL: (H 08), P & P (2011), (P 16), and BOL, BRA: Garrison & von Ellenrieder (2018). Addl. Data. ES, San Lorenzo: [FSCA. JJD, KJT]; LR, Quevedo: [FSCA, JTJ, KJT]; MS, Gualaquiza: [USNM]; Huambaya: [KJT, RWG]; NA, Tena: [FAL, FSCA, UTIC, JJD, KJT, RWG, TWD, USNM]; OR, Francisco de Orellana: [FSCA, JJD, KJT, RWG, SWD]; Loreto [FSCA, JTJ, KJT, RWG, USNM]; PA, Pastaza: [FSCA, JJD, JTJ, KJT, RWG, USNM]; SD: [RWG]; SU, Lago Agrio: [SWD]; Shushufindi: [UTIC]. Note: In east slope form the light coloration on the thorax is blue, and in some west slope forms it is lighter blue or pale yellowish, especially from northwest Ecuador. According to Garrison & von Ellenrieder (2018), L. K. Gloyd considered A. adamsi from Ecuador west of the Andes to represent a new species to which she had applied the manuscript name A. lorena. However, Garrison & von Ellenrieder (2018) found no significant differences in male caudal appendages nor female mesostigmal plates. Argia collata Selys, 1865. Publ. Recs. ECU: * Tennessen (1996 b — as A. thespis), (L 06 — as A. thespis), (H 08), (H 14), Garrison & von Ellenrieder (2018); NA: * Garrison & von Ellenrieder (2015); OR: * Garrison & von Ellenrieder (2015); SU: * Garrison and von Ellenrieder (2015). Also BRA: (L 06 — as A. thespis), Garrison & von Ellenrieder (2015), (H 08), (P 11 — as A. thespis), GUF: (H 08); plus COL, PER, VEN: Garrison & von Ellenrieder (2015); and BOL (P 16). Addl. Data. EO, Santa Rosa: 1 ♂, stream 12 km E of Arenillas, Hwy. 25 (- 3.5244, - 79.9831, 30 m), 29 iii 2008, leg. & det. KJT — as A. thespis [FSCA]; GY, Balao: 1 ♂, stream, banana plantation, 2 km E of Hwy. 25 & Balao Jct. (- 2.9100, - 79.6919, 70 m), 28 iii 2008, leg. & det. KJT — as A. thespis, [FSCA]; NA, Tena: 4 ♂♂, 1 ♀, Río Chinchipino, 27.7 km E of Puerto Napo (- 1.0728, - 77.5864, 385 m), 13 xi 1997, leg. & det. J. Daigle — as A. thespis [JJD]; OR, Francisco de Orellana: 1 ♂, stream 2.3 km E of Yasuni Sci. Res. Sta., P. N. Y., 9 vi 1995, leg. & det. KJT — as A. thespis [KJT]; 1 ♂, same data but leg. & det. WFM — as A. thespis [FSCA]; 1 ♂, seepage 2.1 km E of first road past research station, Yasuni Preserve, 15 xi 1997, leg. & det. JJD — as A. thespis [JJD]; 1 ♀, stream Waorani Road, 2.0 km past cutoff, Yasuni Preserve, 16 xi 1997, leg. & det. JJD — as A. thespis [JJD]; SU, Cascales: 1 ♀, stream 50 km W of Lago Agrío (395 m), 8 vi 1995, leg. & det. KJT — as A. thespis [KJT]; Shushufindi: 1 ♂, stream about 3 km NW of Shushufindi (- 0.1664, - 76.6167, 280 m), 14 v 2009, leg. & det. KJT — as A. thespis [KJT]. Note: According to Garrison (pers. comm. 2011), “ A. collata was thought to be a synonym of A. thespis, but it has been elevated back to distinct species. Specimens labeled and / or published as A. thespis from Ecuador should be this species. ” Argia cuspidata Garrison & von Ellenrieder, 2018. Publ. Recs. ES: * Garrison & von Ellenrieder (2018 — ♀ para- type, from tandem pair), San Lorenzo: seeps 1.1 km NW of Lita at bridge, 5: 00 pm [0.8850, - 78.4755, 470 m], 2 ii 1997, leg JJD, [JJD]); SD: * Garrison & von Ellenrieder (2018 — Holotype: 19 km E of Santo Domingo de los Colorados [- 0.3137, - 79.0289, 740 m], 7 v 1975, leg. A. Langley & J. Cohen [USNM]). Note: Endemic to limited area of Pacific slope. Argia difficilis Selys, 1965. Publ. Recs. ECU: (H 08); (P 11); MS: Garrison & von Ellenrieder (2007); NA: Garrison & von Ellenrieder (2007); OR: Garrison & von Ellenrieder (2007); PA: Garrison & von Ellenrieder (2007); SU: Garrison & von Ellenrieder (2007). Also BRA, CEN, COL, PER, VEN, T & T: (H 08). Addl. Data. MS, Gualaquiza: [RWG, UMMZ, USNM]; NA, Tena: [DRP, FSCA, RWG, UMMZ]; OR, Francisco de Orellana: [FSCA, KJT, NVE, SWD, TWD, USFQ, USNM]; Loreto: [KJT, RWG, UMMZ]; PA, Mera: [FSCA]; Pastaza: [RWG]; SU, Shushufindi: [DRP, FSCA, RWG, SWD]. Argia dives Förster, 1914. Publ. Recs. ECU: (L 06), (H 08), (P 11), (H 14); MS: * Tennessen (1998), Garrison & von Ellenrieder (2018); NA: Garrison & von Ellenrieder (2018); OR: Garrison & von Ellenrieder (2018); PA: Tennessen (2006), Garrison & von Ellenrieder (2018); SU: (Garrison & von Ellenrieder, 2018); TU: Garrison & von Ellenrieder (2018). Also PER: (L 06), (P 11); plus BRA: (L 06); and COL: P & P (2011), (P 16). Addl. Data. MS, Huamboya: [JJD]; Morona: [RWG]; Gualaquiza: [FSCA]; NA, Archidona: [RWG]; Carlos Julio Arsemena Tola: [RWG]; Tena: [FSCA, KJT, RWG, USNM]; OR, La Joya de los Sachas: [FSCA]; Loreto: [DRP, FSCA, JJD, JTJ, KJT, RWG, TWD, USNM]; PA, Mera: [FSCA, JJD, KJT, RWG, USNM]; Pastaza: [FAL, FSCA, JJD, KJT, RWG, SWD, SWD]; SU, Shushufindi: [FSCA, RWG, SWD]; ZC, El Pangui: [FSCA, JJD, KJT]. Argia euphorbia Fraser, 1946. Publ. Recs. ECU: (P 16), Garrison & von Ellenrieder (2018); * OR: von Ellenrieder & Garrison (2011), Garrison & von Ellenrieder (2015). Addl. Data. OR, Francisco de Orellana: 1 ♂, T. B. S. (- 0.6333, - 76.1500, 220 m), date?, leg. K. Jackson, D. Inward, det. RWG- 2003 [BMNH] Also PER: (H 08), (P 11); plus ARG, BRA: (H 08); plus COL, VEN: (P 16). Note: The Argentina record is an error (Garrison & von Ellenrieder 2015). Argia fraudatricula Förster, 1914. Publ. Recs. ECU: (* H 08), (P 11), (H 14), Garrison & von Ellenrieder (2018). Also PER: (H 08), (P 11). Addl. Data. NA, Tena: 1 ♂, Jatun Yacu, near mouth of Ilapulin, 8 iii 1937, leg. WCM, det. JJD [FSCA]; same but 4 ♂♂, det. L. Gloyd, from RWG [RWG, UMMZ]; 3 ♂♂, Jatun Yacu, Río Napo watershed (- 1.0167, - 77.8332, 700 m), 4 iii 1937, leg. WCM, det. L. Gloyd, from UMMZ [RWG, UMMZ]; same but 3 ♂♂, 2 ♀♀, 21 iii 1937, [RWG, UMMZ]; PA: 4 ♂♂, Partidero River trail [El Partidero, Indian village on east bank of Río Anzu, jct. of trails from Mera and Puyo northward to Napo], Río Napo watershed (- 1.0167, - 78.000, 1000 m), 16 ix 1935, leg. WCM, det. L. Gloyd, from RWG [RWG, UMMZ]; ZC, Nangaritza: 7 ♂♂, 2 ♀♀, Río Nangaritza (- 4.55, - 78.8167, 1400 m), 2 xii 1941, leg. D. Laddey, det. L. Gloyd [RWG, UMNZ]; same but 1 ♂, 1 ♀, Mayaicu, Río Nangaritza (- 3.9894, - 78.6328, 1136 m) [RWG]; same but 2 v 1941 [RWG]. Note: There are no recent records of this species despite numerous field surveys within that part of Ecuador where it was recorded more than 75 years prior to this report. Argia fulgida Navás, 1934. Publ. Recs. BO: * Garrison & von Ellenrieder (2017, 2018); EO: * Garrison & von Ellenrieder (2017, 2018); ES: * Garrison & von Ellenrieder (2017, 2018); LR: * Garrison & von Ellenrieder (2017, 2018); MN: * Garrison & von Ellenrieder (2017, 2018); PC: Garrison & von Ellenrieder (2018); SD: * Garrison & von Ellenrieder (2017, 2018);. Also CEN, COL: Garrison & von Ellenrieder (2017). Addl. Data. BO, Caluma: 4 ♂♂, rocky stream, 5 km E of Caluma, Guaranda Road (- 1.5942, - 79.2308, 480 m), 5 iii 2010, leg. JJD, det. RWG [JJD, RWG]; 4 ♂♂, same data but KJT [FSCA, KJT]; CN, Cañar: 2 ♂♂, Cochancay 20.2 km E of Prov. Border on Hwy. 70 rocky stream (- 2.4686, - 79.2853, 195 m), 3 xi 1997, leg, det. JJD [FSCA]; EO, Portovelo: 1 ♂, Portovelo (- 3.7167, - 79.6167, 650 m), 1 vii 1941, leg. D. Laddey, det. RWG [RWG]; ES, San Lorenzo: 1 ♂, rocky stream 36.5 km NW of Lita, near Urbina, 3 ii 1997, leg. & det. JJD; 1 tan. pair, 1 ♂, [San Antonio], nearby, 36.5 km WNW of Lita on [old] San Lorenzo Hwy, pools / seeps / stream (0.95, - 78.68, 200 m), 3 ii 1997, leg. & det. RWG [FSCA]; LR, Babahoyo 2 ♂♂, 2 ♀♀, Playas de Juan Montalvo, near Balzapamba (- 1.6833, - 79.2833, 15 m), 28 ii 1938, leg. WCM, det. RWG, from UMMZ [RWG]; 1 ♂, 1 ♀, same data but 8 iv 1938 [RWG]; 2 ♂, same data but 15 iv 1938 [RWG]; 2 ♂♂, same data but 17 iv 1938 [RWG]; 1 ♂, same data but 31 vii 1941 [RWG]; PC, Quito: 6 ♂♂, upper Mashpi Road, Rio Huaycuyacu, above waterfall (0.1952, - 78.8657, 850 m), 2 ix 2018, leg. & det. WFM [FSCA, USFQ]; 2 ♂♂, 1 tan. pair, same data but leg & det. KJT [KJT]; SD: 3 ♂♂, 1 ♀, Río Pove, Santo Domingo de los Colorados (- 0.25, - 79.15, 600 m), 27 ix 1940, leg. D. Laddey, det. RWG, from UMMZ [RWG]; 1 ♂, 1 ♀, same data but 28 ix 1940 [RWG]; 1 ♂, Río Mulaute, near Santo Domingo de los Colorados; virgin forest, 3 iii 1973, leg. M. Deyrup, det. RWG, from UMMZ [RWG]; 1 ♂, small stream, Hwy. 30, near km marker 71, near La Unión del Toachi (- 0.3139, - 78.9436, 900 m), 25 ii 2010, leg. & det. JTJ [JTJ]. Note: All records are from Pacific slope. Argia fumigata Hagen in Selys, 1865. Publ. Recs. ECU: (P 16), Garrison & von Ellenrieder (2018); * OR: Garrison & von Ellenrieder (2015). Also: BRA, GUF, GUY, SUR, VEN: (P 16). Addl. Data. OR: Francisco de Orellana: 1 ♀, Reserva Étnica Waorani, 1 km S Okone Gare Camp, canopy fogging project, Lot # 1546, TR 3 # 6, terra firme forest (- 0.6569, - 76.4528, 216 m), 21 vi 1996, leg. T. L. Erwin et al., det. RWG [USNM]. Note: This record is the westernmost for this predominately Guiana shield species. Argia gerhardi Calvert, 1909. Publ. Recs. ECU: (H 08), (P 11), (H 14). Also BOL, COL, PER: (H 08), (P 11). Addl. Data. Provided by R. Garrison (pers. comm., 25 vi 2018): LJ, Loja: 1 ♂, 1 ♀, Estación Cientifica San Francisco, Río San Francisco, small creek, oviposition in soil (- 3.9744, - 79.0778, 1870 m), 28 x 2014, leg. C. Willigalla, det. RWG [C. Willigalla]; NA, Archidona: 4 ♂♂, 1 ♀, seepages near river, 43.5 km W Loreto (- 0.7212, - 77.6410, 880 m), 14 vi 1995, leg & det. SWD [RWG]; 1 ♂, seep 44 km east of Narupa jct., Loreto Road, 14 xi 1997, leg. KJT, det. RWG [RWG]; PA, Mera: 1 ♀, 20 km on Puyo to Baños Hwy. (- 1.4543, - 78.1221, 1130 m), 19 x 1997, leg. M. Kippenrand, det. RWG [RWG]. SU, Gonzalo Pizarro: 1 ♂, 1 ♀, San Rafael Falls, Río Quijos, 52 km northeast of Baeza (- 0.1020, - 77.5890, 1290 m), 22 i 2009; leg DLW, WAH, Michael Thomas, Fred Morrison & Ronald Vargas, det. RWG [WAH]; TU, Baños: 1 ♀, Abitagua, Río Pastaza watershed, Hda. Guillermina (- 1.4431, - 78.1665, 1200 m), 7 xii 1939, leg. WCM, det. L. Gloyd [RWG]; same but, 1 ♂, 9 xii 1939 [RWG]; ZC, Zamora: 1 ♂, seepage near Río Sabanillo, W of Zamora (- 4.0453, - 79.0147, 1225 m), 1 iv 2008, leg. & det. JTJ [JTJ]; 1 ♂, 1 ♀, tributary of Bombuscara River on road towards Zamora (- 4.0428, - 79.0142, 1280 m), 18 viii 2006, leg. JCA, det. RWG [UTIC]. Notes: According to Rosser Garrison (pers. comm. 25 - vi- 2018), “ The identification within the Argia gerhardi / kokama / nigrior complex is difficult. I have seen (and illustrated) types of all three and have thus far done a lot of work on this group .... Putting names on various populations is not always easy as I have had to compare various characters from Ecuadorean and Peruvian populations with the types (Bolivia, Peru) and the characters I have thus far seen to be of diagnostic value do not seem to be concordant. In particular, males of Argia kokama approach those of A. nigrior closely and the females of A. gerhardi and A. nigrior are difficult to separate. All of this is going to require more work ..... the records I have for Argia gerhardi from Ecuador are listed with the proviso that not all of the records may pertain to that species. ” All records are from relatively high elevation, from 880 – 1870 m. Argia hamulata Fraser, 1946. Publ. Recs. ECU: * Garrison & von Ellenrieder (2018), Also BOL (RWG, pers. comm.), PER: (H 08). So far known from only one record: SD: 1 tan. pair, stream, Tinalandia, E of Santo Domingo, Hwy. 30 (- 0.3000, - 79.0667, 800 m), 31 i 1997, leg. KJT, det. RWG, data provided by RWG [KJT]. Argia huanacina Förster, 1914. Publ. Recs. ZC: stream, north side of Guaguayme Bajo, 4 iv 2008, leg. & det. KJT [FSCA] * Tennessen & Johnson (2008), Garrison & von Ellenrieder (2018). Also BOL, PER (H 08) (P 11), Garrison & von Ellenrieder (2018). Notes: From Rosser Garrison (pers. comm., 2018): “ This species was treated and illustrated in Garrison & von Ellenrieder (2015). von Ellenrieder & Garrison (2007) followed by Heckman (2008) illustrated the appendages of a male in the RBINS with the manuscript name “ lateralis ” by Selys as A. huanacina. This is incorrect. Subsequent examination of these specimens showed the manuscript name lateralis Selys to represent A. infrequentula as can be seen by comparing Figs. 6 l-n in von Ellenrieder & Garrison (2007) and Fig. 3.1. 427 in Heckman (2008) to Figs. 10 l- 2 and 10 p- 2 in Garrison & von Ellenrieder (2015). ” This southern Ecuador record seems to be the northernmost record for this species. Argia inculta Hagen in Selys, 1865. Publ. Recs. ECU: (* H 08), (P 11), (H 14), Garrison & von Ellenrieder (2018). Also PER: (H 08), (P 11). Addl. Data. ZC: 2 ♂♂, 1 ♀, Pacocha (- 1.0577, - 80.8472, 120 m), 27 iv 1946, leg. L. Gomez Alonzo, det. L. Gloyd [RWG, UMMZ]; EO, Arenillas: 1 tan. pair, 2 ♂, clay-bottom stream, 14 km S of Arenillas, road to Alamar (- 3.6656, - 80.0869, 95 m), 10 iv 2008, leg. & det. KJT [FSCA]; 3 ♂♂, same data but leg. & det. JTJ [JTJ]; 1 tan pair, farm stream, 11 km S of Arenillas, road to Alamar (- 3.6436, - 80.0989), 10 iv 2008, leg. & det. KJT [FSCA]; Piñas: 1 ♀, Piñas (- 3.6667, - 79.65, 1000 m), 23 vii 1941, leg. D. Laddey, det. L. Gloyd, from CHK / UMMZ [RWG]; Portoviejo: 9 ♂♂, 2 ♀♀, Cerro de Hojas (- 1.0432, - 80.4311, 200 m), 19 iv 1946, leg. L. Gomez Alonzo, det. MJW [CSCA, RWG, UMMZ], same but, 21 iv 1946, 10 ♂♂, 3 ♀♀, [CSCA, RWG, UMMZ]; same but, 16 ♂♂, 2 ♀♀, [CSXA, RWG, UMMZ]; 1 ♀, concrete ditch, Hwy. E 40, E of Portoviejo (- 1.0672, - 80.1350, 420 m), 4 iii 2010, leg. & det. KJT [FSCA]; Santa Rosa: 1 ♂, stream, 12 km E of Arenillas, Hwy. 25 (- 3.5244, - 79.9831, 30 m), 29 iii 2008, leg. & det. KJT [FSCA]; 2 ♂♂, same data but leg. & det. JTJ [JTJ]; 2 ♂♂, same data but 9 iv 2008, leg. & det. KJT [FSCA]; LJ, Catamayo: 1 ♂, 1 ♀, small stream and tilapia pond on old road from Malacatos to Catamayo, (- 4.0312, - 79.3558, 1290 m), 18 i 2014, leg. F. Morrison, R. Vargas, & E. Wyman, det. RWG [RWG]; MN, Chone: 2 ♂♂, 7.2 km NNE of Chone, Boca de Mosquito, between Río Grande and Río Mosquito (- 0.6833, - 80.1000, 110 m), 24 x 1978, leg. J. Anderson, det. RWG, from USNM [JJD, RWG]; 1 ♂, small stream / trib. 3 km N of Canuto (- 0.7678, - 80.1231, 60 m), 3 iii 10, leg. & det. JTJ [JTJ]; 9 ♂♂, same data but leg. & det. JJD [JJD]; Flavio Alfaro: 1 ♂, borrow pit / small stream about 4 km N of Flavio Alfaro (- 0.3750, - 79.8875, 295 m), 3 iii 2010, leg. & det. JTJ [JTJ]; Jipijapa: 1 ♂, [Playa] San Jose, 4 miles inland (- 1.38, - 80.75, 60 m), 12 iii 1970, leg.? det. KJT [FSCA]; 2 ♂♂, Puerto Lopez, 20 km to Salinas, 5 iv 1998, leg. E. Tapia, det. JJD [JJD]; Junin: 2 ♂♂, 1 ♀, stream just S of Junin (- 0.9317, - 80.2256), 3 iii 2010, leg. & det. KJT [FSCA]; Manta: 1 ♂, Santa Rosa (- 1.0576, - 80.8472, 20 m), 11 v 1946, leg. L. Gomez Alonzo, det. L. Gloyd [RWG]; same but, 6 ♂♂, 2 ♀♀, 12 v 1946 [CSCA, RWG, UMMZ]; same but, 6 ♂♂, 1 ♀, 14 v 1946 [CSCA, RWG, UMMZ]; Pedernales: 2 ♂, small stream about 2 km E of Pedernales (0.0836, - 80.0139, 90 m), 28 ii 2010, leg. & det. KJT [FSCA]; 1 ♂, same data but leg. & det. JTJ [JTJ]. Note: Collection data indicate that A. inculta is a low elevation west slope species. Argia indicatrix Calvert, 1902. Publ. Recs. ECU: (L 06), (H 08), (P 11), (H 14), Garrison & von Ellenrieder (2018); CN: Garrison & von Ellenrieder (2015); EO: Garrison & von Ellenrieder (2015); * ES: Daigle et al. (1997), Garrison & von Ellenrieder (2015); LR: Garrison & von Ellenrieder (2015); NA: Garrison & von Ellenrieder (2015); OR: von Ellenrieder & Garrison (2011), Garrison & von Ellenrieder (2015); PC: Garrison & von Ellenrieder (2015); * SD: Daigle et al. (1997) Garrison & von Ellenrieder (2015); SU: Garrison & von Ellenrieder (2015). Also CEN, COL, PER, VEN: (L 06), (H 08), (P 11) plus BRA: (L 06), (H 08) and BOL: (P 16). Addl. Data: BO, San Miguel: [RWG]; CN, La Troncal: [RWG, SWD]; EO, Santa Rosa: [FSCA, JTJ, KJT]; ES, San Lorenzo: [FAL, FSCA, JA, JJD, KJT, RWG, SWD]; LR, Babahoyo: [RWG]; Buena Fé: [DRP, SWD]; MN, Flavio Alfaro: [FSCA, KJT]; OR, Francisco de Orellana: [JJD, KJT, RWG, SWD]; Loreto: [JTJ, KJT]; PC, San Miguel Bancos: [RWG]; SD: [DRP, FAL, FSCA, JJD, KJT, RWG, SWD]; SU, Lago Agrío: [RWG]. Argia infrequentula Fraser, 1946. Publ. Recs. ECU: (H 08), (P 11), (H 14), Garrison & von Ellenrieder (2018); NA: * Daigle (1996). Also COL: (H 08, P 11, P & P); and PER: (P 16). Addl. Data. MS, Morona: [UMMZ]; NA, Tena: [BMNH, CSCA, FSCA, JJD, KJT, RWG, UMMZ]; OR, Francisco de Orellana: [FSCA, KJT, TWD, USNM]; Loreto: [JJD, KJT, USNM]; PA, Mera: [JJD, KJT, RWG, UMMZ]; Pastaza: [BMNH, CSCA, RWG, UMMZ]; Santa Clara: [JJD, RWG]; SU, Cascales: [FSCA, USNM]; Lago Agrio: [RWG]; Sushufindi: [FAL, FSCA, JJD, JTJ, KJT, RWG]; TU, Baños: [UMMZ]; ZC, Zamora: [RWG]. Note: Some of these older records need to be rechecked as they may be Argia selysi Garrison & von Ellenrieder (2018). Argia jocosa Hagen in Selys, 1865. Publ Recs. S U: 1 ♂, Bellavista River, 32 km SE of Lago Agrio, 11 i 2014, leg. D. Wagner, W. Haber, F. Morrison, det. RWG [RWG] Garrison & von Ellenrieder (2018). Also COL, VEN: (Garrison & von Ellenrieder 2018). Argia kokama Calvert, 1909. Publ. Recs. ECU: Garrison & von Ellenrieder (2018); MS: * Tennessen (2006); OR: von Ellenrieder & Garrison (2011, 2017); SU: Garrison & von Ellenrieder (2017); Also PER: (H 08), (P 11); plus COL: P & P (2011), (P 16). Addl. Data. MS, Huamboya: 6 ♂♂, 1 tan. pair, Macas, 58 km N on Hwy. 45 (70 km N of Sucua), roadside ditch and tribs. near Río Pasanac (- 1.9447, - 77.8536, 762 m), 16 ix 2005, leg. & det. WFM [FSCA, QZAC, USNM]; 3 ♂♂, 1 ♀, same but: 10 xi 197, leg. & det. JJD — as “ A. makoka ” [JJD]; 1 ♂, same data but, leg. TWD, det. JJD — as “ A. makoka ” [JJD]; Morona; 8 ♂♂, Macas 15.7 km W on Macas-Guamote Road small stream (- 2.2689, - 78.1936, 1463 m), 17 ix 2005, leg. & det. WFM [FSCA, QZAC, USNM]; 8 ♂♂, Macas 22.9 km W on Guamote Road, 3.3 km W on Río Albanico rocky stream and flowing roadside ditches (- 2.2342, - 78.218, 1690 m), 18 ix 2005, 8 ♂♂, leg. & det. WFM [FSCA, QZAC, USNM]; OR, Francisco de Orellana, 1 ♀, T. B. S., USFQ, open grassy area behind din- ing room, surrounded by bushes and trees and some seepage areas (- 0.6383, - 76.1492, 220 m), 14 i 2009, leg. & det. RWG & NVE [RWG]; 2 tan. pairs, same data but 15 i 2009; 1 tan. pair, 1 ♂, same data but, 16 i 2009, 1 ♂, same data but 16 i 2009, 1 ♂, 1 ♀, same data but 18 i 2009; 1 ♂, T. B. S., USFQ, black water stream crossing Guacamayo trail at marker 2050, 16 i 2009, 1 ♂, same data but T. B. S., USFQ, Mauritia palm swamp off Guacamayo trail at marker 2100, 20 i 2009, 2 ♂♂, 1 ♀, same data but T. B. S., USFQ, Guacamayo trail between markers 450 – 2100, 22 i 2009, leg. & det. RWG & NVE [USFQ]; 1 ♂, 1 ♀, General Plaza Gutierrez: [about 7 km N] & 22.5 km N of Indanza on Hwy. 45, roadside ditch, 10 i 2009, leg. & det. RWG & NVE [RWG]; PA, Santa Clara: El Partidero, Río Napo watershed (- 1.2833, - 77.9000, 800 m), 4 ix 1935, leg. WCM, det. RWG, from C. H. Kennedy [UMMZ]; Mera; 5 ♂♂, Mera between Mera and Shell, Los Copales trails and forest N of Río Metolo (- 1.4886, - 78.0708, 1021 m), 20 ix 2005, leg. det. WFM [FSCA, QZAC, USNM]. Argia limitata Navás, 1924. Publ. Recs. - NA: * Garrison & von Ellenrieder (2018); OR: * Garrison & von Ellenrie- der (2018); PA: * Garrison & von Ellenrieder (2018); SU: * Garrison & von Ellenrieder (2018); ZC: * Garrison & von Ellenrieder (2018). Also PER: (H 08), (P 11), (H 14). Addl. Data: NA, El Chaco: [FSCA]; Tena: [JJD, KJT, RWG, SWD, SWD]; OR, Loreto: [JTJ, KJT]; PA, Mera: [JJD]; Pastaza: [FAL, JJD, KJT, RWG, SWD]; SU, Cascales: [SWD]; Lago Agrio: [RWG]. Note: Often referred to as “ near ” A. risi, then in the mid 1990 ’ s it was referred to as “ A. dunklei, ” a manuscript name since it was thought to be a new species. Many specimens in various museums around the world are labeled “ A. dunklei ms. ” Another manuscript name that was proposed more recently was “ A. upano; ” Rosser Garrison (pers. comm., 2017) has determined that most of these refer to A. limitata, a very common, mid-elevation east slope species. A few specimens labeled “ A. upano ms. ” turned out to be A. philipi Garrison & von Ellenrieder, 2018. Argia loutoni Garrison & von Ellenrieder, 2015. Publ. Recs. ECU: (P 16), Garrison & von Ellenrieder (2018); SU, Shushufindi: * Garrison & von Ellenrieder (2015). Also BRA, COL, PER, VEN: Garrison & von Ellenrieder (2015). Addl. Data. SU, Shushufindi: 1 ♀, small stream about 3 km W of Shushufindi (- 0.1647, - 76.6878, 286 m), 28 x 2013, leg. KJT, det. RWG [KJT]. Argia mauffrayi Garrison & Ramón Cabrera, 2019. Publ. Recs. ECU: PC, San Miguel de Los Bancos Cantón: Garrison & Ramón Cabrera (2019 — Holotype ♂, Recinto Milpe, Milpe Bird Sanctuary, Mirador Uno Trail (0.0333, - 78.8661, 1101 m), 4 ix 2018, leg. WFM [USFQ]. ES, Quinindé: Paratype ♂, Reserva Canande (0.5203, - 79.0866, 470 m), 15 v 2012, leg. Jessica Ware & Melissa Sanchez [RWG]). Note: Known only from these two localities at present. Argia medullaris Hagen in Selys, 1865. Publ. Recs. ECU: (H 08), (P 11), (H 14); IM: Garrison & von Ellenrieder (2018); LJ: Garrison & von Ellenrieder (2018); MN: Garrison & von Ellenrieder (2018); MS: Garrison & von Ellenrieder (2018); NA: * Tennessen (1996 b), Daigle (1996), Garrison & von Ellenrieder (2018); PA: Garrison & von Ellenrieder (2018); PC: Garrison & von Ellenrieder (2018); SD: Garrison & von Ellenrieder (2018); TU: Garrison & von Ellenrieder (2018). Also CEN, COL, VEN: (H 08), (P 11), Garrison & von Ellenrieder (2018). Addl. Data. CT, Pangua: [FSCA, KJT]; Sigchos: [FSCA]; IM, Cotacachi: [FSCA, KJT, USFQ]; San Miguel Urcuq: [JTJ]; LJ, Catamayo: [JTJ]; MS, Morona: [FSCA, JJD, RWG]; NA, Archidona: [FSCA, JTJ, KJT, SWD]; Quijos: [FAL, FSCA, JJD, KJT, RWG, TWD, USNM]; PC, Mejía: [FSCA, JJD, USNM]; San Miguel Bancos: [FSCA, USNM]; SD: [FAL, KJT]; SU, Gonzalo Pizarro: [FSCA]; TU, Baños: [RWG]; ZC, Zamora: [JTJ]. Argia nigrior Calvert, 1909. Publ. Recs. ECU: (H 08), (P 11), (H 14); Garrison & von Ellenrieder (2018); ZC: * Tennessen & Johnson (2008). Also PER: (H 08), (P 11). Note: Rosser Garrison (pers. comm., 2017) wrote “ A pair labeled Argia sp. “ D ”: ECUADOR: Zamora-Chinchipe Province, 5.6 km east of Zamora, 3.7 km south of Timbara [“ small river with boulders, tributary of Río Zamora, end of gravel road, trickle on roadside and along trail bordering river, secondary forest and pasture ”], 4.0658, - 78.9044; 4 iv 2008; collected by Dave Wagner, William A. Haber, Michael Thomas, Fred Morrison & Ronald Vargas matches well the type of A. nigrior from Bolivia and have so labeled it this species. ” Argia oculata Hagen in Selys, 1865. Publ. Recs. ECU: * Calvert (1902), Calvert (1902 b), Ris (1918), Soukop (1954), Paulson (1983), Tsuda (1991), Steinmann (1997 a), (L 06), (H 08), (P 11), (H 14); CR: Campos (1922); CB: Calvert (1907), Campos (1922); EO: Campos (1927); LR: Campos (1922); AZ, BO, CT, ES, LJ, LR, PC, SD: Garrison & von Ellenrieder (2015), MS: Garrison & von Ellenrieder (2007); NA: Daigle (1996), Garrison & von Ellenrieder (2007); OR: Tennessen (1996), Garrison & von Ellenrieder (2007, 2011); PA: Garrison & von Ellenrieder (2007); SU: Garrison & von Ellenrieder (2007). Also BRA, COL, PER — as A. difficilis, TTO as A. difficilis, VEN: (L 06), (H 08), (P 11); plus CEN, PER, TTO: (H 08); and BOL, GUF, GUY, SUR: (P 16). Addl. Data. AZ, Santa Isabel: [FSCA, KJT, RWG, SWD]; BO, Caluma: [FSCA, KJT, JTJ]; CB, Camada: [RWG]; CN, Cañar: [FSCA]; La Troncal: [JJD, KJT, SWD]; San Miguel: [FSCA, KJT]; CT, Pangua: [FSCA, JTJ, KJT]; Sigchos: [FSCA]; EO, Arenillas: [FSCA]; Piñas: [FSCA, JTJ, KJT]; Santa Rosa: [JTJ, KJT]; ES, San Lorenzo: [FSCA, JJD, KJT, RWG, USNM]; GY, Naranjal: [FSCA, JTJ, KJT]; IM, Ibarra: [FSCA]; LJ, Catamayo: [FSCA, JTJ, KJT]; Loja: [FSCA, JTJ, KJT, UTIC]; LR, Buena Fé: [SWD]; Quevedo: [JTJ, KJT]; Montalvo: [KJT]; MN, Flavio Alfaro: [KJT]; Pedernales: [JTJ, KJT]; PC, Mejía: [FSCA]; San Miguel Bancos: [FSCA, KJT, RWG, USFQ]; SD: [FSCA, JJD, JTJ, KJT, RWG, SWD, USNM]. Note: Many specimens of A. oculata in various collections are labeled “ Argia cf. oculata, ” “ A. oculata?, ” “ A. difficilis, ” “ A. forficula ” and “ A. elipitca ”. Garrison & von Ellenrieder (2015) defined A. oculata and discussed its varability. It is very common in Ecuador on the Pacific slope. Argia philipi Garrison & von Ellenrieder, 2018. Publ. Recs. MS: * Garrison & von Ellenrieder (2018); NA: * Garrison & von Ellenrieder (2018); OR: * Garrison & von Ellenrieder (2018); SU: * Garrison & von Ellenrieder (2018). Also BOL, COL, PER: Garrison & von Ellenrieder (2018). Addl. Data. OR, Loreto: 2 ♂♂, 1 cop. pair, forest stream about 12 km NE of Loreto (- 0.6217, - 77.2600, 365 m), 12 x 2009, leg. KJT [KJT]; same data but 1 ♂, small streams and forest trails 12 km E of Loreto, 2 + km S of Hwy. 45, leg. JTJ, det. RWG [JTJ]; PA, Mera: 1 ♀, near Shell, trails, forest, and Río Metolo, at Los Copales (- 1.4919, - 78.0725, 1070 m), 20 ix 2005, leg. KJT, det. RWG [KJT]; SU, Shushufindi: 1 ♂, stream about 3 km NW of Shushu- findi (- 0.1664, - 76.6917, 290 m), 13 x 2009, leg. KJT, det. RWG [KJT]. Argia popoluca Calvert, 1902. Publ. Recs. ECU: (H 14), * Garrison & von Ellenrieder (2018). Also COL (as A. variata): (H 08), (P 11), (P & P). Addl. Data. BO, Caluma: 1 ♂, tributary of rocky stream 5 km E of Caluma, Gauranda Road (- 1.5942 - 79.2308, 480 m), 5 Mar 2010, leg. JJD, det. RWG [JJD]; ES, Quinindé: 1 ♂, Río Canande Reserve, 35 km NE of Quininde, tributary of Río Hoja Blanca, small gravel stream in forest (0.5264, - 79.2122, 345 m), 20 x 2012, leg. WAH, det. RWG, data provided by RWG (2018) [WAH]; PC, Quito: 2 ♂♂, 2 ♀♀, Río Tulambi, Forestry Station Maquipucuna (about 5 km E of Nanegalito) (- 0.1169, - 78.6297, 1344 m), 5 ix 1990, leg. OSF, det. RWG [RWG, USNM]; 1 ♂, 2 km NE of Mindo, Septimo Paraiso Lodge, small stream in forest (0.0308, - 78.7633, 1555 m), leg. WAH, det. RWG, data provided by RWG (2018) [WAH]. Argia pulla Hagen in Selys, 1865. Publ. Recs. ECU: (L 06), von Ellenrieder & Muzón (2007), (H 08), (P 11), (H 14), Garrison & von Ellenrieder (2018); NA: * Daigle (1996). Also BRA, CEN, COL, PER, TTO, VEN: (L 06), (H 08), (P 11); plus ARG: (H 08); and SUR: (P 16). Addl. Data. ES, Esmeraldas: [DRP]; MN, Chone: [JJD, JTJ]; Flavio Alfaro: [JTJ]; Junin: [JTJ, KJT]; MS, Santiago: [SWD]; NA, El Chaco: [SWD]; Tena: [FSCA, JTJ, KJT, SWD, TWD, USNM]; OR, Francisco de Orellana: [FSCA, JJD, KJT, RWG, TWD, SWD, USNM]; Loreto: [JTJ]; SD: [FSCA, JJD, KJT, USNM]; SU, Lago Agrio: [SWD, USNM]; Shushufindi: [FSCA, KJT, SWD, USNM]. Argia schneideri Garrison & von Ellenrieder, 2017. Publ. Recs. MS: * Garrison & von Ellenrieder (2017); NA: * Garrison & von Ellenrieder (2017 — Holotype: Las Palmas, on Río Anzu in Río Napo watershed, 11 xii 1936, leg. WCM, [UMMZ]; PA: * Garrison & von Ellenrieder (2017); TU: * Garrison & von Ellenrieder (2017). Also COL: Bota-Sierra et al. (2018). Addl. Data. OR, Loreto: 1 cop. pair, forest stream about 12 km NE of Loreto (- 0.6217, - 77.2600, 365 m), 12 x 2009, leg. & det. KJT [KJT]; 1 ♂, same data but 15 x 2009, leg. & det. JTJ [JTJ]; 1 ♂, small streams and forest trails 12 km E of Loreto, 2 + km S of Hwy. 45 (- 0.6217, - 77.2600, 365 m), 15 x 2009, leg. & det. JTJ [JTJ]; SU, Shushufindi: 1 ♂, trails, small streams 3 km W of Shushufindi, 2 + km N off Hwy. 26 at km marker 17 (- 0.1664, - 76.6917, 290 m), 13 x 2009, leg. & det. JTJ [JTJ]. Note: The manuscript name “ camura ” was originally applied to this species. Argia selsyi Garrison & von Ellenrieder, 2018. Publ. Recs. MS: * Garrison & von Ellenrieder (2018); NA: * Garrison & von Ellenrieder (2018 — Holotype: [Tena]: Jatun Yacu, Napo River watershed [- 1.0166, - 77.8333, 700 m], 18 iv 1935, leg. WCM [UMMZ]); OR: * Garrison & von Ellenrieder (2018); PA: * Garrison & von Ellenrieder (2018); TU: * Garrison & von Ellenrieder (2018). Note: Endemic but common when found. Argia talamanca? Calvert, 1907. Publ. Recs. ECU: * Tennessen (1998), (P 11), (H 14). Also CEN, COL, VEN: (H 08), (P 11). Addl. Data. BO, San Miguel: 1 ♂, Balzapamba (- 1.77, - 79.18 W, 732 m), ix 1935, leg. R. Campos, det. L. Gloyd [FSCA]; CN, La Troncal: 2 ♂♂, small stream and waterfall, 25 km E of Cochancay, Hwy. 70, (- 2.4692, - 79.1697, 1540 m), 3 xi 1997, leg. & det. JJD [FSCA]; 4 ♂♂, same data but leg. & det. KJT [FSCA]; 5 ♂♂, 2 ♀♀, same data but leg. & det. TWD [RWG, TWD, FSCA]; 3 ♂♂, same data but leg. & det. WFM [FSCA, LSAM, USNM]; CT, Sigchos: 3 ♂♂, San Francisco de las Pampas Otonga Reserve (- 0.42, - 79.0, 1700 m), 14 – 16 ii 1997, leg. & det. Elicio Tapia, [FSCA, QZAC]; ES, San Lorenzo: 1 ♂, 5 km NW of Lita, seeps along RR tracks, 5 ii 1997, leg. & det. JJD [FSCA]; MN, Jipijapa: 3 ♂♂, 3 ♀♀, Puerto Lopez “ 20 km to Salinas ” (- 1.57, - 80.82, 61 m), 5 iv 97, leg. & det. Elicio Tapia [FSCA, QZAC]; PC, Mejía: 1 ♂, Manuel Cornejo 2 km S of Tandapi, waterfalls (- 0.43, - 78.78, 1737 m), 21 x 1980, leg. & det. MJW [FSCA]; 1 ♂, Manual Cornejo E near Tandapi, “ east of Santo Domingo de los Colorados ” (- 0.4167, - 78.7833, 1585 m), 21 x 1980, leg. & det. MJW [FSCA]; 7 ♂♂, same data but 24 x 1980. [FSCA]; TU, Baños: 1 ♂, La Ventana, upper Río Pastaza watershed, ix 1935, leg. & det. F Campos R, [FSCA]. Note: South American specimens may represent a new undescribed species, since the true A. talamanca may occur only in Central America (R. Garrison, pers. comm., 2018). Very similar to A variegata. Argia talamanca appears to be a west slope species whereas A. variegata is mainly an east slope species. Argia tennesseni Garrison & von Ellenrieder, 2018. Publ. Recs. NA: * Garrison & von Ellenrieder (2018); OR: * Garrison & von Ellenrieder (2018 — Holotype: [Loreto]: stream 8.5 km E of Loreto [- 0.6183, - 77.2950, 400 m], 14 xi 1997, leg. KJT [FSCA]; ♀ Paratype: stream on road to Waorani Reserve, 6.2 km south of main Yasuni Road [- 0.7120, - 76.4744, 230 m], 10 vi 1995, leg. KJT [FSCA]; SU: * Garrison & von Ellenrieder (2018). Also BRA, PER: Garrison & von Ellenrieder (2018). Addl. Data. OR, Francisco de Orellana: 3 ♂♂, 1 ♀, stream, road to Waorani Reserve, 6.2 km S of Yasuni Sci. Res. Sta., 10 vi 1995, leg. KJT [KJT, RWG]; 1 ♀, P. N. Y., on road to Waorani Reserve, 1.8 km W of Waorani Re- serve Road, on Markus Oil Co. Road PC 29 - 30, small stream (- 0.7000, - 76.4667, 200 m), leg. WFM, det. RWG & NVE [RWG]; 1 ♂, 2 ♀♀, T. B. S. (- 0.6333, - 76.1500, 220 m), 20 ix 2000, leg. Kelly Jackson & D. Inward, det. RWG [BMNH]; 1 ♀, T. B. S., USFQ, stream crossing Matapalo trail at marker 450 (- 0.6369, - 76.1458, 245 m), 14 i 2009, leg. & det. RWG & NVE [RWG]; 1 ♂, 4 ♀♀, T. B. S., USFQ, stream crossing Chorongo trail at marker 225 (- 0.6350, - 76.1503, 220 m), 16 i 2009, leg. & det. RWG & NVE [RWG]; 1 ♂, 4 ♀♀, same data but 18 i 2009 [RWG]; 2 ♀♀, T. B. S., USFQ, black water stream crossing Guacamayo trail at marker 2050 (- 0.6383, - 76.1492, 220 m), 17 i 2009, leg. & det. RWG & NVE [RWG]; 2 ♀♀, T. B. S., USFQ, Guacamayo trail between markers 450 – 2100 (- 0.6350, - 76.1533, 237 m), 22 i 2009, leg. & det. RWG & NVE [RWG]; Loreto: 3 ♂♂, small streams and forest trails 12 km E of Loreto, 2 + km S of Hwy. 45 (- 0.6271, - 77.2600, 365 m), 12 x 2009, leg. JTJ [JTJ]; 1 ♂, 2 ♀♀, 1 reared, same data but leg. KJT [KJT]. Argia translata Hagen in Selys, 1865. Publ. Recs. ECU: (H 08), (P 11). CR, GY, LR: * Campos (1922), EO: Campos (1927). Also ARG, CEN, COL, GUF, PER, TTO, VEN: (H 08), (P 11); plus BOL, GUY, SUR: (P 16). Addl. Data. EO, Piñas: [FSCA, JTJ]; Santa Rosa: [KJT]; GU, Balao: [FSCA, KJT]; Naranjal: [FSCA]; LR, Buena Fé: [DRP, FSCA, RWG, SWD]; Quevedo: [FSCA, KJT]; Vantanas: [FSCA, KJT]; MN, Chone: [USNM]; Manta: [RWG, USNM]; NA, Tena: [FSCA, JJD, RWG, SWD, TWD, USNM]; SD: [RWG]; SU, Gonzalo Pizarro: [USNM]. Argia variegata Förster, 1914. Publ. Recs. ECU: * Förster (1914 - Description, no specific type locality), Davies & Tobin (1984), Tsuda (1986, 1991), Bridges (1993), Steinmann (1997 a), Garrison et al. (2003), (H 08), (P 11), (H 14); MS: Tennessen (2006); NA: Daigle (1996), Tennessen (1998); ZC: Tennessen & Johnson (2008). Also COL, PER: (H 08), (P 11). Addl. Data. MS, Huamboya: [FAL, FSCA, JJD, KJT, TWD, USNM]; Morona: [FSCA]; NA, Archidona: [FAL, FSCA, JJD, JTJ, KJT, SWD, TWD, USNM]; Carlos Julio Arsemena Tola: [RWG]; El Chaco: [SWD]; Quijos: [JJD]; Tena: [SWD]; LJ, Loja [JTA]; OR, Loreto: [DRP, FSCA, JJD, KJT, RWG, TWD, USNM]; PA, Mera: [FSCA, JJD, KJT, RWG, UMMZ, USNM]; PC, San Miguel de Los Bancos: [FSCA, USFQ]; SU, Gonzalo Pizarro: [FSCA]; TU, Baños: [CSCA, FSCA, RWG, UMMZ, USNM]; ZC, Zamora: [FSCA, JTJ, KJT, USNM]. Note: This species is closely related to A. talamanca and may be found to be a junior synonym of it; the issue is currently under review by Rosser Garrison. Argia unresolved: There are numerous specimens and records for Argia which we have not listed here. Many belong to the Argia gerhardi-kokama-nigrior group that still needs to be resolved. Museum specimens are often labeled by alphabetical designation (e. g. “ Argia sp. g ”). Garrison & von Ellenrieder have been working on this complex for some time and hopefully someday will provide a solution for these unresolved Argia.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B53FFFFBFF23F0B42482F9EC.taxon	discussion	Relatively large coenagrionids with a long abdomen, mostly black with slight metallic sheen and pale blue or yellow markings (Fig. 17). Two species of this genus occur in Ecuador; see De Marmels & Garrison (2005) for species identification. Females lay eggs in tank bromeliads in forest. Nymphs have been discovered recently, with descriptions by De Marmels in Garrison (2005) and Torreias et al. (2008). Bromeliagrion fernandezianum (Racenis, 1958). Publ. Recs. PA: * Tennessen (2006). Also VEN: De Marmels & Garrison (2005), (H 08), (P 11). Addl. Data. OR, Francisco de Orellana: 1 ♂, Reserva Etnica Waorani, 1 km S Okone Gare Camp, canopy fogging project, Lot # 971, TR 1 # 2, terra firme forest (- 0.6569, - 76.4528, 216 m), 9 ii 1995, leg. T. L. Erwin, det. RWG [USNM]; 1 ♂, T. B. S. (- 0.6514, - 76.1466, 236 m), 6 xii 2012, leg. & det. ACRP [ACRP]; 1 ♂, same data but, camiño Harpía (- 0.6258, - 76.1463, 257 m), 12 vi 2013; PA, Mera; 2 ♀♀, trails and forest, Los Copales, between Mera and Shell (- 1.4919, - 78.0725, 1070 m), 20 ix 2005, leg. & det. KJT [KJT]; 2 ♂♂, same data but 19 ix 2005, leg. & det. JJD (JJD). Bromeliagrion rehni Garrison in De Marmels & Garrison, 2005. Publ. Recs. ECU: (L 06), Torreias et al. (2008), (P 11), (H 14); OR: De Marmels & Garrison (2005 — Holotype & Allotype: Scientific Research Station, P. N. Y. (- 0.6714, - 76.4006, 220 m), 07 – 21 xi 2001, leg. A. Rehn); SU: De Marmels & Garrison (2005), von Ellenrieder & Garrison (2011). Also BRA: (H 08), (P 11). Addl. Data. MS, Huamboya: 1 ♀, waterfall / seeps 80 km N of Sucua, Hwy. 45, 10 xi 1997, leg. & det. JJD [JJD]; NA, Tena: 1 ♀, J. S. B. S., near Tena, wet rocky seep (- 1.0500, - 77.7683, 450 m), 11 xi 1994, leg. A. Jaramillo, det. DRP [DRP]; OR, Francisco de Orellana: 2 ♂♂, Reserva Etnica Waorani, 1 km S Onkone Gare Camp (- 0.6528 - 76.4333, 220 m), 7 x 1994, leg. T. Erwin, det.? [USNM]; 3 ♂♂, 2 ♀♀, Scientific Research Station, P. N. Y. (- 0.6714, - 76.4006, 220 m), 7 – 21 xi 2001, leg. A. Rehn, det. RWG [ACR, UMMZ]; 1 ♀, T. B. S., TLE Plot T- 4, Sta 8, Lot # 2237 fogging canopy (- 0.6319, - 76.1442, 220 – 250 m), 1 x 2000, leg. T. Erwin, det. RWG [USNM]; 1 ♂, T. B. S., at base of canopy tower at marker 2600 of Guacamayo trail (- 0.6364, - 76.1642, 219 m), 17 i 2009, leg. & det. NVE 2009 [RWG]; 1 ♂, T. B. S., sendero Harpía (- 0.6336, - 76.1471, 239 m), 6 xii 2012, leg. & det. ACRP [ACRP]; SU, Putumayo: 2 ♂♂, Río Cuyabeno, 60 km E Lago Agrío (0.0167, - 76.3167), 16 – 26 ii 1998, leg. G. Kareofelas, det. RWG (ACR, RWG].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B53EFFF8FF23F0B12119FF47.taxon	discussion	Small damselflies, the one species recorded in Ecuador marked with blue and black on head and thorax, abdomen mostly black with narrow green rings and yellow on sides, but young adults are orange-ocher with the abdomen greenish yellow (Fig. 18). See Tennessen (2015) for identification. Habitat includes small marshy streams and shallow pools and ditches near rainforest. Nymph is unknown. Calvertagrion mauffrayi Tennessen, 2015. Publ. Recs. OR: * Tennessen (2015 — Holotype ♂: P. N. Y., ditch near Río Savaleto, Yasuni Road km marker 31 (- 0.4700, - 76.5850), 12 vi 1995, leg. KJT (FSCA); Allotype ♀: same data but, 7 vi 1996, leg. KJT [FSCA]); NA: Tennessen (2015); SU: Tennessen (2015). Also COL: Bota-Sierra et al. (2018). Note: From Tennessen (2015), “ Calvertagrion mauffrayi was collected in a variety of shallow ponds and ditch- es filled with water in the upper Amazon basin of Ecuador, perching on and / or flying low amidst grasses and other partly shady vegetation. They did not occur in nearby deeply shaded rainforest. ”	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B53DFFF9FF23F2292113FEA3.taxon	discussion	A small genus with only three known species. Adults are medium-sized damselflies, reddish-brown to orange with light blue markings (Fig. 19); for species identification see Machado (2009 a). Nymphs are unknown; adults have been found in lowland Amazonian forest near streams. * Denticulobasis dunklei Machado, 2009. Publ. Recs. PER: (Machado 2009 a). Addl. Data. SU, Shushufindi: Limoncocha (- 0.4000, - 76.6167, 269 m), 15 xi 1980, leg. MJW, det. JJD [FSCA]. * Denticulobasis garrisoni Machado, 2009. Publ. Recs. BRA: Machado (2009 a); plus COL: (P 16). Addl. Data. NA, Tena: 1 ♂, J. S. B. S., Bosque hacia o Napo (- 1.0652, - 77.6196, 404 m), 7 xi 2014, leg. & det. ACRP [ACRP]; OR, Orellana: 1 ♂, T. B. S. (- 0.6379, - 76.1511, 243 m), 13 xii 2012, leg. & det. ACRP [RWG]; 1 ♂, same data but camino Chichico (- 0.6415, - 76.1449, 226 m), 6 xii 2013 [ACRP].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B53CFFF9FF23F3172400FAD6.taxon	materials_examined	Addl. Data. SU, Lago Agrio: 1 ♂, 18 km E of Lago Agrio, 30 viii 1975, leg. A. Langley & J. Cohen, det. RWG [USNM].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B53CFFF9FF23F387270BF9DA.taxon	materials_examined	Publ. Recs. ECU: (L 06), (H 08), (P 11), (H 14); GY: * Campos (1922 — as E. coecum var. novae-hispaniae). Also ARG, BRA, CEN, COL, PER, TTO, VEN: (L 06), (H 08), (P 11). Addl. Data. EO, Santa Rosa: [FSCA, JTJ, KJT]; GY, Balao: [FSCA, KJT]; Naranjal: [FSCA, RWG]; MN, El Carmen: [JTJ]; Pedernales: [FSCA, KJT]; MS, Morona: [KJT]; NA, Tena: [FSCA, UTIC, KJT, SWD, TWD, USNM]; OR, Francisco de Orellana: [FSCA, SWD]; SD: [FSCA]; SU, Gonzalo Pizarro: [RWG]; Lago Agrio: [SWD]; ZC, Yantzaza: [FSCA].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B53CFFE7FF23F0A7215FFEA3.taxon	discussion	A relatively large genus of small damselflies, adults of which are mostly black with green and / or blue stripes or spots (Fig. 21), and in one species thorax and abdomen with yellow markings. Habitats include ponds, marshes, lake margins and slow streams and ditches with aquatic vegetation. Four species are known in Ecuador and an undescribed species has been collected. No comprehensive study on the taxonomy of the South American species is available, however the key in Westfall & May (2006) contains most of the species found in Ecuador. Ischnura capreolus (Hagen, 1861). Publ. Recs. ECU: Paulson (1983), Tsuda (1986), (L 06), (H 08), (P 11), (H 14); CR: Campos (1922), GY: * Ris (1918 — as Ceratura capreolus), Campos (1922); EO: Campos (1927); LR: Campos (1922). Also ARG, BRA, CEN, COL, GUF, GUY, PRY, PER, SUR, VEN, TTO: (L 06), (H 08), (P 11). Addl. Data. EO, Arenillas: [JTJ, KJT]; Santa Rosa: [JTJ, KJT]; ES, Esmeraldas: [DRP, FSCA, KJT, QZAC, USNM]; San Lorenzo: [FSCA]; GY, Balao: [JTJ, KJT]; Daule: [RWG, USNM]; Duran: [FSCA]; Guayaquil: [ANSP, DRP, FSCA]; Naranjal: [FSCA]; LR, Babahoyo: [RWG, USNM]; Quevedo: [FSCA, USNM]; Ventanas: [KJT]; MN, Bolivar: [USNM]; Chone: [USNM]; El Carmen: [JTJ, KJT]; Flavio Alfaro: [KJT]; Sucre: [RWG, USNM]; OR, Francisco de Orellana: [FSCA, KJT, SWD]; Loreto: [RWG]; SD: [DRP, FSCA, JJD, KJT, LSAM, QZAC, SWD, USNM]; SU, Shushufindi: [FSCA]. Note: The Pacific slope specimens differ somewhat in the shape of the terminal appendages and could represent an undescribed species (R. Garrison, pers. comm., 2017). Ischnura fluviatilis Selys, 1876. Publ. Recs. ECU: (H 14); IM: * Daigle et al. (1997); LJ: Tennessen & Johnson (2008). Also ARG, BOL, BRA, CHL GUF, PRY, PER, VEN, URY: (H 08), (P 14); plus SUR: (P 16). Addl. Data. EO: Arenillas [JTJ, KJT]; Machala: [KJT]; Santa Rosa [JTJ, KJT]; GY, Durar: [JTJ]; Naranjal [FSCA]; Simon Bolivar: [KJT]; IM, Ibarra: [KJT]; LJ, Loja: [JTJ, KJT]; Macará: [ANSP, FSCA]; LR, Quevedo: [FSCA]; Ventanas: [KJT]; MN, Sucre: [RWG]; ZC, Zamora: [JTJ]. Ischnura hastata (Say, 1840). Agrion hastatum Say, 1840; Anomalagrion hastatum (Say, 1840). Synonyms: Agrion anomala Rambur, 1842; A. venerinotata Halderman, 1844. Publ. Recs. ECU: Tsuda (1986, 1991) GA: * Currie (1901), Calvert (1906), Campos (1922), Schmidt (1938), Gloger (1964), Turner (1967), Belle (1991 a — all as A. hastatum), (H 08), (P 11). Isabela: Peck (1992), San Cristibal: Peck (1992), Santa Cruz: Peck (1992). Also CEN, VEN: (H 08); plus COL, GUF, SUR: (P 16). Addl. Data. GA, Isabela: [USNM]; San Cristobal: [ACRP, USNM]; Santa Cruz: [USNM]. Note: Peck (1992) found this species “ in littoral saline meadows to pampa zones around permanent wet spots and pools. ” No records from the mainland yet. Ischnura ramburii (Selys, 1850). Publ. Recs. ECU: (L 06), (P 11), (H 14 — as “ near ramburii ”); IM: * Daigle et al. (1997). Also BRA, COL, GUF, SUR, TTO, VEN: (L 06), (H 08), (P 11); plus CEN: (H 08), (P 11); plus CHL, PER, PRY: (L 06), (H 08). Addl. Data. GY, Daule: adults, 5.5 km S of Nobol, 12 i 1978, leg. J. Anderson, det.? [USNM]; IM, Ibarra: 1 cop. pair, 3 ♂♂, 1 ♀, pool 1.5 km S of Salinas (0.4853, - 78.1333), 2 ii 1997, leg. & det. KJT [KJT]; 2 ♂♂, 2 ♀♀, Sa- linas, 0.5 km E, pools / small irrigation stream (- 0.55, - 78.12, 1676 m), 2 ii 1997, leg. & det. WFM [FSCA, QZAC]; 1 cop. pair, 4 ♂♂, 45.8 km E of Lita, rocky stream (- 0.78, - 78.32, 853 m), 6 ii 1997, leg. & det. WFM [FSCA, LSAM, QZAC, USNM]; LJ, Loja: 1 ♂, Río Chamba at Vilcabamba (- 4.2625 - 79.2192), 16 – 17 viii 2006, leg. & det. JCA [UTIC]; LR, Babahoyo: adults, 6 km N of Babahoyo, 21 vi 1975, leg. & det.? [USNM]; MN, Bolivar: adults, 4 km W Calceta, lake margin, 20 x 1978, leg. J. Anderson & det.? [USNM]; Sucre: adults, 35 km SE of Bahia de Caraquez, oxbow pond, 5 x 1975, leg. A. Langley, det.? [USNM]; SU, Lago Agrio: adults, 18 km E of Lago Agrio, 20 viii 1975, leg. A. Langley & J. Cohen, det.? [USNM]. Note: There may be two separate species that are currently referred to I. ramburii.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B522FFE4FF23F76021A8FA7F.taxon	description	Small to medium slender damsefllies, only two of the nine species being found in Ecuador; undetermined specimens also exist in collections that could represent an undescribed species. The thorax usually has green stripes and the slender, elongate abdomen is often orange or yellowish (Fig. 22). See Garrison & von Ellenrieder (2010) and Johnson (2016) for identification. The Ecuador species inhabit open to partly shaded shallow marshes and the edges of small slow streams. Leptobasis linda Johnson, 2016. Publ. Recs. MN: Johnson (2016 — Holotype: ♂, from a small forested stream, N of Chindul, (0.2389, - 79.8742, about 30 m), 1 iii 2010, leg. JTJ [FSCA]); LR: Johnson (2016 — Allotype ♀: from a small stream, 1.8 km N of Hwy. 491, approx. 25 km E of Babahoyo, (- 1.8111, - 79.3189, about 60 m), 7 iii 2010, leg. JTJ [FSCA]); EO: Johnson (2016). Addl. Data. GY, Naranjal: 1 ♀, “ Finca de la Gringa, ” 15 km E of Tres Cerritos, near Cerro Cimalón (- 2.4456, - 79.5556, 25 m), 12 iv 2008, leg. & det. JTJ [JTJ]. Note: Kennedy (1919) recognized that “ the species (of Leptobasis labeled vacillans) in the Cornell University collection from Ecuador appears to be entirely new and have long forcipate inferiors. ” He showed figures of the Cornell specimen genital ligula. Leptobasis vacillans Selys, 1877. Publ. Recs. ECU: Paulson (1983), Tsuda (1986 — as L. v. ecuadorica), Tsuda (1991), (H 08), (P 11), (H 14); EO: Campos (1927); GY: * Calvert (1909), Campos (1922), Garrison & von Ellenrieder (2010); LR: * Calvert (1909 — as “ L. vacillans race ecuadorica ”), Campos (1922); SD: Daigle et al. (1997), Garrison & von Ellenrieder (2010 — as Pinchincha). Also BRA, CEN, GUF, SUR, VEN: (H 08), (P 11); plus COL: (P 11), (P & P). Addl. Data. GY, Guayaquil: 7 ♂♂, 2 ♀♀, Quayaquil (- 2.7333, - 79.75, 17 m), date?, leg. F. Campos, det. P. Calvert, data provided by RWG, [ANSP, USNM]; Naranjal: 2 ♀♀, [San Rafael], listed as “ San Raphael ” (- 2.18, - 79.23, 200 m), date?, leg. F. Campos, det.? [FSCA]; LR, Quevedo: 5 ♂♂, 1 ♀, Quevedo (- 1.0333, - 79.4500, 11 m), date?, leg. F. Campos, det. P. Calvert 1909, data provided by RWG [ANSP]; MN, El Carmen: borrow pit / small slow stream, NW of El Carmen (- 0.1375, - 79.5728, 185 m), 28 ii 2010, leg. & det. JTJ [JTJ]; SD: 2 ♂♂, 2 ♀♀, seepage fen at Hotel Colorado, Hwy. 30 about 16 km E of Santo Domingo (- 0.31, - 79.03, 762 m), 31 i 1997, leg. & det. KJT [KJT]; 2 ♂♂, same data but leg. & det. JJD [FAL, JJD]; 1 ♂, 1 ♀, same data but 30 i 1997, leg. JJD, det. RWG [RWG]; 2 ♀, same data but leg. & det. WFM [FSCA, QZAC]; 2 ♂♂, 3 ♀♀, 29 km W Santo Domingo de los Colorados, 5 vii 1975, leg. A. Langley & P. Spangler, det. RWG [USNM]. Leptobasis unresolved: GY, Duran: Leptobasis nov. sp. “ as por Calvert ” (* Campos 1922); LR, Babahoyo: 6 ♂♂, 5 ♀♀, 10 km N of Babahoyo, banana plantation, (- 1.8167, - 79.5167, 56 m), 22 vi 1975, leg. J. Cohen, A. Langley, P. Monnig, det. RWG & NVE 2009 [USNM]; Buena Fé: 37 km N of Quevedo, 7 iv 1975, leg. J. Cohen & A. Langley, det.? [USNM]; Quevedo: 1 ♀, Pichilingue (- 1.1000 - 79.4833, 40 m), 2 ii 1955, leg. E. I. Schlinger & E. S. Ross, det. RWG 2009 CAS [DRP]. Note: These are all probably Leptobasis linda, pending re-examination of the specimens.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B521FFE5FF23F0042173FA00.taxon	discussion	Two of the fourteen species are recorded for Ecuador, and there is possibly a third yet unknown species. Adults are small to medium in size and usually are black with blue markings (Fig. 23), although young individuals are black and red. For identification, see von Ellenrieder & Garrison (2008 a). They breed in small pools of open marshy seepage at high elevations. Mesamphiagrion dunklei von Ellenrieder & Garrison, 2008. Publ. Recs. ECU: Dunkle (1981 — as Cyanallagma laterale), Daigle (1996 — as C. laterale) (P 11), Bota-Sierra & Wolf (2013), (H 14); NA: * von Ellenrieder & Garrison (2008 a — Holotype & Allotype: [El Chaco]; 13 km E of Coyuja on road to Chaco, marsh [coordinates for Coyuja] (- 0.4, - 78.0, 2934 m), 17 viii 1980, leg. SWD [FSCA]), Daigle (1996 — as C. laterale). Addl. Data. NA, El Chaco: Paratypes: 26 ♂♂, 9 ♀♀, 13 km E of Coyuja on road to Chaco, marsh, (coordinates for Cuyuja: - 0.4, - 78.0, 2934 m), 17 viii 1980, leg. SWD, det. NVE & RWG [DRP, FSCA, KJT, NVE, RWG, SWD, USNM]; Archidona: 1 paratype ♂, 1 ♂, 1 ♀, seep, Baeza / Tena Road, 26.7 km N of Narupa jct. (- 0.6033, - 77.8433, 2240 m), 19 xi 1997, leg. KJT, det. NVE & RWG [FSCA, KJT]; 1 paratype ♂, seep and marsh along Archidona- Baeza road (- 0.6033, - 77.8433, 2100 m), 19 xi 1997, leg. TWD, det. NVE & RWG [SWD]; Quijos: 1 paratype ♂, [Bermejo], 10.6 km S of Baeza on Hwy. 45 near Bermejo, seepage marsh (- 0.5083, - 77.8789, 1800 m), 24 vii 1996, leg. WFM, det. NVE & RWG [FSCA]; 1 ♀, [Cosanga], along Baeza-Archidona Road just S of Cosanga, seep and marsh (- 0.6006, - 77.8350, 2100 m), 19 xi 1997, leg. TWD, det. NVE & RWG [SWD]; 3 ♂♂, 1 ♀, marsh / seep, about 2 km N of Cosanga (- 0.5631, - 77.8756, 1925 m), 16 x 2009, leg. & det. KJT [KJT]; 2 ♂♂, dame but, leg. & det. JTJ [JTJ]; 1 ♂, 1 ♀, Papallacta, 4.6 km E on Hwy. E 20, seepage hillside with pools (- 0.3739, - 78.0779, 2597 m), leg. & det. CDK [FSCA]; 3 ♂♂, 1 ♀, same data but leg. & det. KJT [FSCA]. Mesamphiagrion ecuatoriale von Ellenrieder & Garrison, 2008. Publ. Recs. ECU: (P 11), Bota & Wolf (2013), (H 14); NA: * von Ellenrieder & Garrison (2008 a — Holotype & Allotype: [Archidona], seep and marsh along Archidona-Baeza road (- 0.6033, - 77.8433, 2100 m), 19 xi 1997, leg. TWD [FSCA]); 3 ♂♂ and 1 ♀ paratypes, same data but [RWG, SWD]; 2 ♂♂, 2 ♀♀, same data but leg. KJT [KJT]). Note: Endemic. Mesamphiagrion unresolved: NA, Quijos: Yanayacu Biological Station, 3.5 km SW of Cosanga, marshy seeps and ditches in pasture (- 0.5800, - 77.8650, 2045 m), date not provided; data provided by WAH & DLW. WAH (pers. comm. 2016) listed it as “ cf. laterale, ID not confirmed, but not dunklei or ecuatoriale. ”	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B520FFE3FF23F01D24AEFC1E.taxon	discussion	This genus was recently revised by von Ellenrieder (2013 b) who added 18 new species. Eight of the 31 species are known to occur in Ecuador, and it is likely that others will be discovered as they are secretive forest dwellers quite easily overlooked. The exact habitat is unknown as the nymph of none of the species in this large genus has been found. Adults are quite distinctive, having an orange thorax with a dark (often metallic green) mid-dorsal stripe and mostly black abdomen (Fig. 24); they perch on twigs or leaf tips in deep shade and usually fly only upon disturbance. Metaleptobasis gibbosa Tennessen, 2012. Publ. Recs. ECU: (H 14); PA, Mera: * Tennessen (2012 a — Holotype ♂: Mera, between Mera and Shell, Los Copales, trails and forest wetlands (- 1.4919, - 78.0725, 1070 m), 22 ix 2005, leg. & det. KJT [FSCA]; 1 paratype ♂, same data but leg. JJD [FSCA]; 1 paratype ♂, same data but leg. WFM [FSCA]; Allotype ♀: same data but 20 ix 2005 [FSCA]). Addl. Data. PA, Mera: 1 ♂, same data as holotype except leg. WFM [LSAM]. Note: Apparently endemic. Type specimens were found near a small, partially shaded seepage pool at the edge of the forest at Los Copales, with very little water. Adults perched low on stems along a path, but when disturbed flew high toward the overhanging bushes (Tennessen 2012 a). Upon return to Los Copales in 2012, it was discovered that the type locality had been cut over and houses built in the area. Metaleptobasis knopfi Tennessen, 2012. Publ. Recs. ECU: (H 14); NA, El Chaco: * Tennessen (2012 a — as “ Sucumbíos ”, Holotype and Allotype plus 7 paratype ♂♂, Baeza-Lago Agrío road, 52 km NE Chaco, swamp, forest and stream (- 0.0011, - 77.4019, 685 m), 18 viii 1980, leg. KWK (FSCA); 3 paratype males, same data but leg. SWD [FSCA]; Tena: 1 paratype male, pond along Río Sinde, about 5 km E of Puerto Napo bridge (- 1.0517, - 77.7544, 450 m), 12 xi 1997, leg. KJT [FSCA], von Ellenrieder (2013 b); S U, Shushufindi: 1 paratype female, forest S of Limoncocha (- 0.4203, - 76.6108, 250 m), 11 v 2009, leg. KJT [FSCA]; NVE (2013). Also from COL: (P 16). Metaleptobasis lillianae Daigle, 2003. Publ. Recs. ECU: (H 14); SU, Shushifindi: Limoncocha, left or S fork of nature trail in forest (- 0.24, - 76.36, 243 m), 25 viii 1980, leg. SWD, det. NVE [RWG]. Also BOL, BRA, PER: (von Ellenrieder 2013). Metaleptobasis mauffrayi Daigle, 2000. Publ. Recs. ECU: (H 08), (P 11), (H 14). OR: * Daigle (2000 — Holotype: as “ Napo ” P. N. Y., ditch on road PC 29 - 30 about 3.2 km from main road junction, 18 vi 1996, leg. JJD [FSCA]; Allotype: as “ NA ”, forest pool on main road, 3.6 km N of Yasuni Biological Research Station, 16 xi 1997, leg. KJT [FSCA]); NVE (2013); SU: * Daigle (2000 — as “ Napo ”), von Ellenrieder (2013 b). Also PER: Daigle (2000), (H 08), (P 11) plus COL: von Ellenrieder (2013 b). Addl. Data. MS, Morona: [RWG]; NA, Tena: [ACRP, FSCA, KJT, RWG,); OR, Francisco de Orellana: (ACR, ACRP, FSCA, JJD, KJT, NVE, RWG, SWD, TWD, USNM]; Loreto: [KJT, FSCA]; La Joya de los Sachas: [KJT]; SU, Shushufindi: [FSCA, KJT, RWG, SWD, SWD]. Note: Seems to be the most abundant and widespread species of Metaleptobasis in Ecuador. Metaleptobasis minteri Daigle, 2003. Publ. Recs. ECU: (H 08), (P 11), (H 14); MS: NVE (2013); OR: * Daigle (2003 — Holotype & Allotype: as “ NA ”, Limoncocha [across Río Napo from], [La Nueva Esperanza], Primavera, Río Napo, at Lake Taracoa (- 0.45, - 76.77, 243 m), 26 viii 1978, leg. KWK [FSCA], von Ellenrieder (2013 b); SU: von Ellenrieder (2013 b). Also BRA, PER: von Ellenrieder (2013 b); plus COL: Palacino et al. (2017). Addl. Data. MS, Morona: 2 ♂♂, Cusuimi, Río Cusuimi, 150 km SE of Puyo (- 2.7225, - 77.6753, 320 m), 15 vi 1971, leg. B. Malkin & B. Yuma, det. NVE, data provided by RWG [NMNHL]; OR, Francisco de Orellana: 1 cop. pair, 1 paratype ♂, Limoncocha [across Río Napo from], [La Nueva Esperanza], primavera, Río Napo, at Lake Taracoa (- 0.45, - 76.77, 243 m), 26 viii 1978, leg. KWK, det. JJD [FAL, FSCA]; 4 ♂♂, 2 ♀♀, same data but 27 vii 1977, leg. DRP & S. Hills, det. NVE [DRP]; 1 tan. pair, 2 ♂♂, 2 ♀♀, same data but leg. & det. SWD, det. WFM [SWD]; 1 ♂, 1 ♀, [Pompeya Sur], S on [Pompeya Sur-Iro road] at km marker 31, borrow pit near Río Sabaleto (- 0.6569, - 76.4528, 216 m), 12 vi 1995, leg. SWD, det. WFM [SWD]; La Joya de los Sachas: 1 ♀, ditch, Pompaya, 0.5 km N of Río Napo ferry, 19 vii 1996, leg. & det. KJT [FSCA]; SU, Lago Agrio: 1 ♂, Lago Agrio, 7 km W on Hwy. 45, swamp & stream, 8 vi 1995, leg. SWD, det. WFM [SWD]; Shushufindi: 1 ♂, Limoncocha, 23 viii 1980, leg. SWD, det. WFM [SWD]; 3 ♂♂, 1 ♀, Limoncocha, lakeshore and nearby areas, (- 0.4000, - 76.6167, 269 m), 23 viii 1980, leg. KWK, det. JJD [FSCA]; 1 ♂, [Limoncocha] [about 40 km E] or 100 km E of Coca [on Río Napo], Lago Garzacocha, La Selva Jungle Lodge (- 0.4819, - 76.3733, 235 m), 12 x 1988, leg. SWD, det. WFM [SWD]; 1 ♀, forest S of Limoncocha (- 0.4203, - 76.6108, 250 m), 11 v 2009, leg. & det. KJT [FSCA]. Metaleptobasis orthogonia von Ellenrieder, 2013. Publ. Recs. ECU: (H 14); OR: von Ellenrieder (2013 b) — paratype ♀, Río Shiripuno, Shiripuno Lodge, 26 km SE of Shiripuno bridge (- 1.1047, - 76.7317, 235 m), 23 – 26 i 2009, leg. D Wagner, det. NVE (UCMS); SU: 1 paratype ♀, Río Cuyabeno, 60 km E Lago Agrío (- 0.0317, - 76.3208, 241 m), 16 – 26 ii 1998, leg. G. Kareofelas, det. NVE [RWG]). Also PER: von Ellenrieder (2013 b). Addl. Data. OR: Estacion Cientifica Yasuni, Río Tiputini, 39 km SE of Pompeya, 61 km ESE of Coca (- 0.674, - 76.397, 210 m), no date provided, data provided by WAH & DLW. Metaleptobasis prostrata von Ellenrieder, 2013. Publ. Recs. ECU: (H 14); * SU: von Ellenrieder (2013 b) — paratype teneral ♂, swamp-stream, 7 km W of Lago Agrío (0.0833, - 76.9833, 280 m), 8 vi 1995, leg. SWD. det. NVE [RWG]. Also PER: von Ellenrieder (2013 b). Metaleptobasis spatulata von Ellenrieder, 2013. * MS, OR, SU all by von Ellenrieder (2013 b). Also BRA, PER: von Ellenrieder (2013 b).	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B526FFE3FF23F5F821E1F846.taxon	discussion	Of the two species in this genus, one occurs in Ecuador. The bright yellow face is distinctive; the thorax is blue on the sides with black stripes and the abdomen is mostly black with the tip blue (Fig. 25). Habitat consists of small, clear, open lakes and ponds with floating vegetation. See Williamson & Williamson (1930 b) and González-Soriano et al. (2004) for identification. * Neoerythromma cultellatum (Hagen in Selys, 1876). Publ. Recs. CEN, VEN: (H 08), (P 11), (H 14). plus COL: (P & P). Addl. Data. MN, Bolivar: 2 ♂♂, 1 ♀, 4 km W Calceta, lake margin, 20 x 1978, leg. J. Anderson, det.? [USNM]; El Carmen: 2 ♂♂, borrow pit lake, 18 km NW of El Carmen (- 0.1350, - 79.5797, 185 m), 2 iii 2010, leg. & det. JJD [JJD]; 4 ♂♂, 1 ♀, same data but leg. & det. JTJ [JTJ]; 7 ♂♂, 4 cop. pairs, same data but leg. & det. KJT [FSCA, KJT]; Sucre: 1 ♀, 39 km E Bahia de Caraquez, 9 v 1975, leg. P. Spangler, det.? [USNM].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B525FFE0FF23F688211CF8BF.taxon	discussion	Two of the five known species occur in Ecuador. Adults are mainly blue and black, of medium size with an elongate abdomen (Fig. 26); young adults are reddish brown. The latest taxonomic treatment was by von Ellenrieder & Garri- son (2008 a). Little is known about the habitat, but at least one species was found to breed in water-filled bromeliads, at relatively high elevations. Oreiallagma oreas (Ris, 1918). Publ. Recs. ECU: * (H 14). Also COL (H 08), (P 16). Addl. Data. PC, San Miguel Bancos: Reserva Gralarias, 7 km NE of Mindo, open area under powerline (- 0.011, - 78.731, 2040 m), no date provided, data provided by WAH & DLW; 2 ♂♂, 1 ♀, Mindo (1524 m), 15 v 2011, data provided by ACRP [ACRP, RWG]. Oreiallagma prothoracium (Kimmins, 1945). Publ. Recs. ECU: Kimmins (1970), Paulson (1983), Davies & Tobin (1984), Tsuda (1986, 1991), all as Leptagrion prothoracium, Steinmann (1997 a — as Teleagrion prothoracicum) (P 11), (H 14); IM: * Kimmins (1945 — as T. prothoracicum, Holotype & Paratype: “ Intag ”, [Apuela, NE on Apuela- Cuicocha Road, along Río Intag and tributaries between 1800 – 2800 m (0.358817, - 78.450904) at jct of Río Intag and tributary], examined by RWG [BMNH]), Bridges (1993), von Ellenrieder & Garrison (2008 a — assigned to Oreiallagma). Addl. Data. A specimen which was examined by both RWG and KJT could be this species: NA, Quijos: 1 ♀, stream 5 km S of Baeza (- 0.4767, - 77.8700, 1900 m), 20 xi 1997, leg. KJT [KJT]	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B525FFE1FF23F15B2765FB0B.taxon	discussion	Only one species of this relatively large genus of 25 species occurs in Ecuador. Adults are small and are usually mostly red, rarely with blue on head and tip of abdomen (Fig. 27). See Mauffray (1999) and von Ellenrieder & Lozano (2008) for identification and diagnoses. Habitat in Ecuador is mainly small seeps and slow stream edges at high elevations. Oxyagrion tennesseni Mauffray, 1999. Publ. Recs. ECU: Paulson (2004 a), von Ellenrieder & Garrison (2006), (P 11), (H 14); NA: * Mauffray (1999 — Holotype & Allotype, Baeza, 10.6 km S on Hwy. 45, near Bermejo, seepage marsh, 24 vii 1996, leg. WFM [FSCA], Muzón et al. (2007). Addl. Data. MS, Morona: [FSCA, KJT]; NA, Archidona: [FAL, FSCA, KJT, SWD]; El Chaco: [DRP, FAL, FSCA, JJD, KJT, RWG, SWD, SWD]; Quijos: [FSCA, KJT, QZAC, RWG, SWD, TWD, USNM]; Tena: [UTIC, USNM]; TU: Banos [FSCA]. Note: Before being described, this species was thought to be a form of Oxyagrion bruchi Navás, 1924. So far found only in a limited area on the east slope of the Andes in Ecuador. Specific locality information on many of these records can be found in Mauffray (1999).	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B524FFEFFF23F3182453FEA3.taxon	discussion	One of the two known species of this genus is found in Ecuador. Adults are relatively small but stocky and are patterned with red (Fig. 28). See Kennedy (1939 b) and Bota-Sierra (2014 a) for identification and diagnoses. Habitat includes lakes and ponds in the La Sierra region. Oxyallagma dissidens (Selys, 1876). Publ. Recs. ECU: Soukop (1954 — as Protallagma runtuni), Paulson (1983 — as P. dissidens), Tsuda (1986 — as P. dissidens), Tsuda (1991), Steinmann (1997 a), (H 08), (P 11), (H 14); CB: * Kirby (1890), Campos (1922); CT: * Kirby (1890), Campos (1922); IM: Daigle et al. (1997), Hunger & Schiel (2012), LJ: Tennessen & Johnson (2008); PC: * Selys (1876 — type Series “ Quito ” (- 0.2167, - 78.5, 2763 m), leg. M. Emile de Ville, data from RWG [IRSNB]), Kirby (1890), Campos (1922), Kennedy (1919); TU: * Kirby (1890), Campos (1922), Kennedy (1939 b — as P. runtuni, Holotype: Runtuni, Baños, 4 vii 1935, leg. WCM [UMMZ]; Allotype: Río Pastaza, Baños (1820 m), 30 iii 1936, leg. WCM, [UMMZ]), Bridges (1993), Garrison et al. (2003 — as P. runtuni but mentions considering it a synonym), Kennedy (1946 b) sunk the species in favor of dissidens (Bota-Sierra 2014 a). Also PER: (H 08), (P 11); plus COL: Bota-Sierra (2014 a). Addl. Data. AZ, Gualaceo: [FAL, FSCA, SWD]; Nabón: [FSCA, JJD, KJT, RWG, USNM]; CB, Alausí: [FSCA]; Colta: [FSCA, RWG]; Pallatanga: [ANSP, FSCA]; CR, Espejo: [ACRP]; CT, Latacunga: [FSCA]; IM, Cotacachi: [FSCA, SWD, USFQ, USNM]; Otavalo: [FSCA, JJD, KJT, RWG, USFQ, USNM]; LJ, Catamayo: [FSCA]; Loja: [FSCA, UTIC, JJD, JTJ, KJT]; Saraguro: [FSCA, USNM]; PC, Cayambe: [DRP, RWG]; Quito: [FSCA, UTIC, RWG]; TU, Baños: [FSCA, RWG, USNM]. Note: widespread at higher elevations of the Andes, especially the central valley. Phoenicagrion von Ellenrieder, 2008 One of the three described species known in this genus occurs in Ecuador and an apparently undescribed species has recently been collected in the country. Adults are mid-sized with a reddish-orange thorax devoid of black stripes, and the sides of the thorax have oblique blue stripes resulting in a unique color pattern (Fig. 29). They occur near lowland forested pools. For identification use von Ellenrieder (2008). Phoenicagrion paulsoni von Ellenrieder, 2008. Publ. Recs. ECU: (H 14); OR: von Ellenrieder & Garrison (2011). Also PER: (P 11). Addl. Data: OR, Francisco de Orellana: 1 ♂, T. B. S., USFQ, Numa trail at Tiputini inlet (- 0.6397, - 76.1483, 220 m), 20 i 2009, leg. & det. RWG & NVE [RWG]. Phoenicagrion unresolved: OR, Francisco de Orellana: Río Shiripuno, Shiripuno Lodge, 26 km SE of Shiripuno bridge (- 1.1047, - 76.7317, 235 m), data provided by WAH & DLW; 1 ♂, T. B. S., camino Matapalo (- 0.6417, - 76.1451, 192 m), 12 iii 2014, leg. & det. ACRP [ACRP]; 1 ♂, same data but quebrada en Chichico (- 0.6420, - 76.1451, 261 m).	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B52AFFEBFF23F760243CFEFF.taxon	discussion	A large genus with 57 species, 12 of which occur in Ecuador. Most species are small with various red or blue and black color patterns, some species with green stripes (Figs. 30 a & b). The latest revision with illustrated keys is by Garrison (2009); Machado (2010) provided an update. Most species inhabit weed-choked ponds, marshes and swamps with floating aquatic plants, some in deeply shaded forest. Telebasis bickorum Daigle, 2002. Publ. Recs. ECU: Paulson (2011); SU: * Garrison (2009). Also BOL: Daigle (2002), (H 08), Garrison (2009), (P 11), (H 14). Addl. Data: SU, Shushufindi: 2 ♂, Limoncocha, palm swamp (- 0.4000, - 76.6167, 269 m), 25 vii 1977, leg. DRP, det. RWG [DRP, RWG]. Telebasis brevis Bick & Bick, 1995. Publ. Recs. ECU: (H 08), Paulson (2009); EO: Tennessen & Johnson (2008), Garrison (2009), (H 14); GY: Garrison (2009); LR: Garrison (2009); MN: * Bick & Bick (1995); SD: * Bick & Bick (1995 — as “ Pichincha ” Holotype & Allotype: Tinalandia, 12 km E of Santo Domingo de los Colorados, 06 x 1988, leg. SWD, [FSCA]), Daigle et al. (1997), Garrison (2009). Addl. Data. EO, Arenillas: [KJT]; Santa Rosa [JTJ, KJT]; GY, Daule: [USNM]; Duran: [FSCA]; Guayaquil: [DRP, FSCA, RWG]; Naranjal: [KJT]; LR, Babahoyo: [FSCA, RWG, USNM]; Buena Fé: [DRP, FAL, SWD, USNM]; Urdaneta: [JTJ, KJT]; Ventanas: [KJT]; MN, Bolivar: [USNM]; El Carmen: [JTJ, KJT]; Pedernales: [FAL, FSCA, RWG, SWD]; Sucre: [USNM]; SD: [DRP, FAL, FSCA, JJD, JTJ, KJT, RWG, SWD, USNM]. Note: “ Telebasis nov. sp. ” from Guayaquil, Naranjito, Venecia, San Rafael, Chimbo, Bucay, Babahoyo and Quevedo as reported by Campos (1922) are probably all T. brevis; we have examined the Babahoyo specimens and they are T. brevis. Endemic to west slope. Telebasis carota Kennedy, 1936. Publ. Recs. ECU: Davies & Tobin (1984), Tsuda (1986, 1991), Bick & Bick (1995), Steinmann (1997 a), (H 08), (P 11), Pinto & Carvalho (2012), (H 14); NA: * Kennedy (1936), Garrison (2009); MS: Garrison (2009), Tennessen (2006); PA: * Kennedy (1936 — Holotype ♂: “ Oriente: near Partidero, Río Anzu ”, [El Partidero, a village on the Río Anzu, two and one-half days’ travel up from the town of Napo], 800 m, 11 x 1935, leg. WCM, data from RWG (2009) [UMMZ]), Bridges (1993), Garrison et al. (2003), Tennessen (1998), Garrison (2009); TU: Garrison (2009). Also PER: (H 08), Garrison (2009), (P 11); plus COL: Bota-Sierra et al. (2018). Addl. Data. MS, Huamboya: [FAL, FSCA, JJD, KJT, USNM]; Santiago: [FSCA, SWD]; Taisha: [DRP, RWG]; NA, Archidona: [RWG]; Carlos Julio Arsemena Tola: [RWG]; Tena: [FSCA, RWG, USNM]; OR, Loreto: [SWD, SWD]; PA, Mera: [KJT]; Pastaza: [DRP, FSCA, KJT, RWG]; Santa Clara: [RWG]; SU, Shushufindi: [FSCA, KJT]; TU, Baños: [FSCA, RWG]. Telebasis dunklei Bick & Bick, 1995. Publ. Recs. ECU: (H 08), Garrison (2009), (P 11), (H 14), Bota-Sierra et al. (2015); SU: * Donnelly (1998). Also PER: (H 08), (P 11); plus COL: (P 16). Addl. Data. SU, Shushufindi: 6 ♂♂, La Selva Lodge; L. Mandicocha and outlet stream (- 0.4983, - 76.3733, 200 m), 25 xi 1997, leg. TWD, det. KJT [KJT, FSCA, RWG, SWD]; 1 ♂, La Selva, Hyacinth Lake, 2 v 1990, leg. E. Smith, det. WFM [SWD]. Telebasis flammeola Kennedy, 1936. Publ. Recs. ECU: Paulson (1983), Davies & Tobin (1984), Tsuda (1986, 1991), Bick & Bick (1995), Steinmann (1997 a), (H 08), (P 11), (H 14); NA: Daigle (1996), Garrison (2009); PA: * Kennedy (1936 — Holotype ♂: “ Río Yanamanaca, Oriente ” Yanamanaca [next Indian village up Río Anzu from Yanamanaca is El Partidero; coordinates for El Partidero] (- 1.28, - 77.90, 800 m), 29 ix 1935, leg. WCM, data provided by RWG [UMMZ]), Bridges (1993), Garrison et al. (2003), Garrison (2009). Also COL: Bota-Sierra et al. (2018). Addl. Data. NA, Tena: 2 ♂♂, 1 ♀, Río Sinde, pond, 5.3 km E of Puerto Napo (- 1.0517, - 77.7544, 450 m), 22 vii 1996. leg. & det. JJD [JJD, KJT]; 1 ♂, same data but 23 vii 1996, leg. & det. JJD [KJT]; 2 ♂♂, same data but leg. & det. KJT [KJT]; 1 cop. pair, 1 ♂, same data but 12 xi 1997 [KJT, RWG]; 3 ♂♂, same data but 16 x 1988, leg. & det. SWD [SWD]; 2 ♂♂, pool near Río Napo, 3.2 km E of Puerto Napo (- 1.0456, - 77.7467, 420 m), 23 vii 1996, leg. & det. KJT [KJT]; 2 ♂♂, same data but leg. & det. WFM [LSAM, QZAC]; 3 ♂♂, same data but leg. & det. SWD [SWD]; 2 ♂♂, Puerto Napo, 19.1 km E on J. S. B. S. Road, seep, 23 vii 1996, leg. & det. JJD [DRP, FAL]; PA, Santa Clara: 2 ♂♂, paratypes, Yanamanaca [next Indian village up Río Anzu from Yanamanaca is El Partidero; coordi- nates for El Partidero] (- 01.28, - 77.90, 800 m), 29 ix 1935, leg. WCM, det. CHK, from UMMZ [RWG]; 1 ♂, Santa Clara W – trib. of Río Anzu Río Yanamanaca (- 1.28, - 77.90, 800 m), 20 viii 1935, leg. WCM, det. CHK [FSCA]. Note: Until recently, this species had been reported only along a 50 km stretch of the upper Río Napo and lower Río Anzu, at ponds and marshes near the road between Puerto Napo and J. S. B. S. Lodge which runs along the south bank of the Río Napo. Telebasis garleppi Ris, 1918. Publ. Recs. ECU: (H 08), (P 11); ES: * Daigle et al. (1997), Garrison (2009), (H 14). Also CEN, COL: (H 08), Garrison (2009), (P 11). Addl. Data. ES, San Lorenzo: 3 ♂♂, [San Antonio], nearby, 36.5 km WNW of Lita on [old] San Lorenzo Hwy., pools / seeps / stream (0.95, - 78.6667, 200 m), 3 ii 1997, leg. JJD, det. RWG [JJD]; 1 ♂, same data but leg. & det. WFM [FSCA, JJD]. Telebasis griffinii (Martin, 1896). Publ. Recs. ECU: (Paulson, 2009), (H 14); NA: Daigle (1996), Garrison (2009); OR: * Bick & Bick (1995 — as “ NA ”. Holotype & Allotype of T. limoncocha: [Francisco de Orellana], [La Nueva Esperanza] [across Río Napo from] near Primavera “ Limoncocha ” Río Napo & lake Taracoa (- 0.45, - 76.77, 300 m), 4 xi 1980, leg. MJW, D. Robinson & H. Smith [FSCA]); Tennessen (1998 — as T. limoncocha), Garrison (2009), von Ellenrieder & Garrison (2011); SU: Tennessen (1996 b), (L 06), Garrison (2009). Also ARG, BRA, COL, PER, TTO, VEN: (L 06), Garrison (2009), (P 11); plus BOL, CEN: (H 08), Garrison (2009), (P 11). Addl. Data. NA, El Chaco: 5 ♂♂, swamp, 52 km NE Chaco on road to Lago Agrio, 18 viii 1980, leg. & det. SWD [SWD]; Tena: 4 ♂♂, 23 km E of Puerto Napo, J. S. B. S. (400 m), 18 x 1988, leg. SWD, det. Bick & Bick [SWD]; 4 ♂♂, sloughs along Napo River, 3.2 km E of Puerto Napo on J. S. B. S. Road, 23 vii 1996, leg. & det. SWD [SWD]; 1 ♂, 1 ♀, Puerto Napo, 5.3 km E on J. S. B. S. Road, Río Sinde, pond upstream (- 1.0517, - 77.7544, 450 m), 13 vii 1996, leg. & det. JJD [DRP]; 2 ♂♂, 1 ♀, same data but leg. KJT, det. RWG [FAL, KJT]; 4 ♂♂, 1 ♀, same data but leg. & det. SWD [SWD]; 2 ♂♂, same data but 22 vii 1996, leg. & det. JJD [DRP]; 1 ♂, same data but 12 xi 1997, leg. & det. KJT [KJT]; 2 ♂♂, same data but 23 vii 1996 [KJT]; 2 ♂♂, same data but leg. & det. SWD [RWG]; 1 ♂, Río Chinchipino, oxbow pond, 27 km E of Puerto Napo (- 1.0728, - 77.5864, 385 m), 13 xi 1997, leg. & det. KJT [KJT]; OR, Francisco de Orellana: 5 ♂♂, 1 ♀, Laguna Taracoa, Primavera (- 0.45, - 76.77, 300 m), 27 vii 1977, leg. DRP & S. Hills, det. DRP [DRP]; 5 ♂♂, same data but 26 viii 1980, leg. & det. SWD [SWD]; 4 ♂♂, 1 cop. pair, trib. of Río Payamino, road to Loreto, 1.7 km W of jct. W Coca Road (- 0.4367, - 77.0017, 258 m), 13 vi 1995, leg. & det. KJT [KJT]; 8 ♂♂, same data but leg. & det. SWD [FAL, RWG, SWD]; 2 ♂♂, P. N. Y., open swamp along main road, 15 xi 1997, leg. & det. KJT [KJT]; 1 ♂, pond, Loreto Road, 8.2 km W of Río Payamino ferry, 18 xi 1997, leg. & det. KJT [KJT]; PA, Pastaza: 3 ♂♂, west branch of Ishpinga-Jacu (- 2.2167, - 77.0666, 500 m), 19 ix 1942, leg. WCM, det. RWG, from UMMZ [FSCA, RWG]; SU, Lago Agrio: 2 ♂♂, swamp, 17 km S of Lago Agrio, 19 viii 1980, leg. & det. SWD [SWD]; Shushufindi: 1 ♂, Limoncocha, palm swamp (- 0.4000, - 76.6167, 269 m), 25 vii 1977, leg. DRP, det. RWG [DRP]; 5 ♂♂, 1 ♀, same data but 26 vii 1977, leg. DRP & S. Hills [DRP]; 1 ♂, same data but lake shore around lake, 24 viii 1978, leg. KWK, det. RWG [RWG]; 1 ♂, same data but swamp canal, 25 viii 1980, leg. SWD, det. RWG [RWG]; 2 paratype tan. pairs, same data but 23 viii 1980, det. Bick & Bick [SWD]; 1 ♂, 1 ♀, cop. pair, [Limoncocha] [about 40 km E] or 100 km E of Coca [on Río Napo], Lago Garzacocha, La Selva Jungle Lodge (- 0.4819, - 76.3733, 235 m), 12 x 1988, leg. & det. SWD [RWG, SWD]; 3 ♂♂, 1 ♀, La Selva, 100 km E Coca, Manicocha, shady forest rivulet (- 0.4983, - 76.3733, 262 m), 12 x 1988, leg. SWD, det. R, Garrison [RWG]; 1 ♂, pond, Limoncocha Road, 15.7 km S of Shushufindi Road, 19 vii 1996, leg. & det. KJT [KJT]. Note: Many specimens, especially in the FSCA, are labeled T. limoncocha Bick & Bick, 1995. Telebasis inalata (Calvert, 1961). Publ. Recs. ECU: (H 08), (P 11), (H 14), Bota-Sierra et al. (2015); GY: Garrison (2009); MN: * Bick & Bick (1995), Garrison (2009); LR: Garrison (2009); SU: Garrison (2009). Also BOL, PER: (H 08), (L 06), Garrison (2009), (P 11); plus ARG: (L 06), Garrison (2009), (P 11) and COL (P 16). Addl. Data. EO, Machala: [KJT]; Santa Rosa: [KJT]; GY, Duran: [FSCA, JTJ, KJT]; Naranjal: [DRP, FSCA, RWG]; LR, Babahoyo: [FSCA, RWG]; MN, Bolivar: [RWG, USNM]; El Carmen: [JTJ, KJT]; Pedernales: [KJT]; Tosagua: [RWG]; SU, Lago Agrio: [USNM]; Shushufindi: [DRP, FSCA, RWG, SWD, USNM]. Telebasis livida Kennedy, 1936. Publ. Recs. ECU: Paulson (1983), Davies & Tobin (1984), Tsuda (1986, 1991), Bick & Bick (1995), Steinmann (1997 a), (H 08), (P 11), (H 14). NA: * Kennedy (1936 — Holotype: [Carlos Julio Arosemena Tola], “ between Río Ila and Río Pupuyaca ”, Río Ila / Pupuyacu (- 0.2000, - 77.8333, 800 m), xi 1934, leg. WCM, data provided by RWG [UMMZ]), Bridges (1993), Tennessen (1996 b), Daigle (1996), Garrison et al. (2003), Garrison (2009). Also PER: Garrison (2009), (P 11); plus BOL: (H 08), (P 11). Addl. Data. NA, Tena: 12 ♂♂, J. S. B. S., 23 km E Puerto Napo, tropical wet forest, ponds nearby (- 1.0500, - 77.7833, 450 m), 18 x 1988, leg. & det. SWD [FAL, FSCA, KJT, RWG, SWD]; 2 cop. pairs, 2 ♂♂, Puerto Napo, 5.3 km E at on Jatun Sacha Road, Río Sinde swamp and pond approx. ½ km upstream (- 1.0517, - 77.7544, 450 m), 13 vii 1996, leg. & det. KJT [KJT]; 3 cop. pairs, 1 ♂, same data but 12 xi 1997 [KJT, RWG]; 1 ♂, same data but leg. & det. WFM [FSCA, QZAC, USNM]; 1 ♂, 1 ♀, same data but 2 vii 1996, leg. & det. JJD [DRP, JJD]; OR, Francisco de Orellana: 1 cop. pair, Puerto Francisco de Orellana, 1.7 km W on Loreto Road from Coca-Sachas Road, stream, pools and flowing shallow marsh (- 0.4367, - 77.0017, 258 m), 13 vi 1995, leg. & det. WFM [FSCA]; 1 cop. pair, Puerto Francisco de Orellana, 12.3 km W on road to Loreto from Coca-Sachas Road, pond (- 0.4750, - 77.0747, 290 m), 13 vi 1995, leg. & det. WFM [FSCA]; 1 tan. pair, same data but leg. & det. SWD [SWD]. Telebasis obsoleta (Selys, 1876). Publ. Recs. ECU: (L 06), (H 08), (P 11), (H 14), Bota-Sierra et al. (2015); OR: * Tennessen (1996 b — as Helveciagrion chirihuanum from NA), Daigle (1996 — as H. chirihuanum); SU: Donnelly (1998), Garrison (2009). Also BOL, BRA, PER: (L 06), (H 08), (P 11); plus ARG, PRY: Garrison (2009), (P 11); and COL (P 16). Addl. Data. OR, Francisco de Orellana: 2 ♂♂, Puerto Francisco de Orellana 1.7 km W on Loreto Road from Coca-Sachas Road, stream, pools & flowing shallow marsh (- 0.4367, - 77.0017, 258 m), 13 vi 1995, leg. & det. WFM [FSCA]; 1 ♂, same data but leg. & det. KJT [KJT]; 1 tan. pair, 4 ♂♂, same data but leg. & det. SWD [JJD, SWD]; SU, Lago Agrio: 1 ♂, swamp, 17 km S of Lago Agrio, 19 viii 1980, leg. & det. SWD [SWD]; Shushufindi: 7 ♂♂, Shushufindi, 15.7 km S on Limoncocha Road, marshy stream and pond (- 0.2219, - 76.6478, 253 m), 19 vii 1996, leg. & det. WFM [FSCA, LSAM, QZAC, RWG, USNM]; 4 ♂♂, same data but leg. & det. SWD [SWD]; 4 ♂♂, Limon- cocha (- 0.4000, - 76.6167, 269 m), 3 – 16 vi 1977, leg. P. Spangler & D. Givens, det. RWG [RWG, USNM]; 4 ♂♂, 1 ♀, same data but lake edge, 27 xii 1971, leg. D. Pearson, det. DRP [DRP, FSCA]; 4 ♂♂, same data but 23 vii 1977 [DRP]; 3 ♂♂, 1 ♀, same data but 26 vii 1977; 1 ♂, same data but 29 viii 1978, leg. KWK, det. RWG [RWG]; 1 ♂, same data but Río Napo, 14 xi 1980, leg. & det. MJW [FSCA]; 7 ♂♂, same data but 25 xi 1980 [FSCA]; 2 ♂♂, 1 ♀, same data but 25 viii 1980, leg. SWD, det. L. Gloyd [SWD]; 6 ♂♂, Shushufindi, pond, Limoncocha Road, 2 km S of Yamaramsuku School, 19 vii 1996, leg. & det. KJT [KJT]; 4 ♂♂, N of La Selva Lodge, Lago Mandicocha & outlet stream “ coca ” - about 100 km E on Río Napo (- 0.4983, - 76.3733, 200 m), 25 xi 1997, leg. & det. TWD [FSCA, SWD]; 1 ♂, La Selva, Hyacinth Lake, 13 vi 1992, leg. E. Smith, det. WFM [SWD]. Note: Many specimens in various collections were initially identified as and may be still labeled Helveciagrion chirihuanum (Calvert, 1909). Telebasis rubricauda Bick & Bick, 1995. Publ. Recs. ECU: * (L 06), (H 08), (P 11), (H 14); OR: * Tennessen (1996 b — as NA), Garrison (2009 — as NA), Tennessen (1998), von Ellenrieder & Garrison (2011); SU: Donnelly (1998). Also BOL, BRA, PER, VEN: (L 06), (H 08), (G 09), (P 11); plus COL (P 16). Addl. Data. OR, Francisco de Orellana: 5 tan. pairs, 3 ♂♂, 1 ♀, sandy stream & marshes, 1.7 km W jct Coca Road on Loreto Road (- 0.4367, - 77.0017, 258 m), 13 vi 1995, leg. & det. SWD [FAL, JJD, SWD, USNM]; 1 tan. pair, 6 ♂♂, 1 ♀, same data but 1 ♂, leg. & det. KJT [FAL, KJT, RWG]; 4 cop. pairs, 4 ♂♂, 2 ♀♀, same data but leg. & det. WFM [FSCA, QZAC, USNM]; 2 ♂♂, same data but 18 xi 1997 [FSCA, QZAC]; 1 ♀, same data but leg. & det. KJT [KJT]; 6 ♂♂, same data but leg. & det. TWD; 2 ♂♂, Puerto Francisco de Orellana 8.2 km W on road to Loreto from Coca-Sachas Road, pond (- 0.4578, - 77.0381, 288 m), 18 xi 1997, leg. & det. WFM [QZAC, USNM]; 1 ♂, 1 ♀, same data but leg. JJD, det. RWG [JJD]; 3 ♂♂, Puerto Francisco de Orellana 11.8 km W on Loreto Road, large pond (- 0.4742, - 77.0583, 285 m), 18 xi 1997, leg. & det. WFM [FSCA, QZAC, USNM]; 2 ♂♂, same data but leg. & det. TWD [FSCA]; 1 ♂, Puerto Francisco de Orellana 12.3 km W on road to Loreto from Coca-Sachas Road, pond (- 0.4750, - 77.0747, 290 m), 13 vi 1995, leg. & det. WFM [FSCA]; 1 ♂, same data but leg. & det. KJT [KJT]; 1 ♂, same data but leg. & det. SWD [SWD]; 1 ♂, ♀, pond on Coca-Loreto road (- 0.4750, - 77.0747, 290 m), 18 xi 1997, leg. & det. TWD [SWD]; 1 ♂, P. N. Y., pond at Río Tiputini (- 0.6101, - 76.4676, 200 m), 17 xi 1997, leg. & det. TWD [FSCA]; Loreto: 3 ♂♂, 2 ♀♀, Loreto, E on road to Coca (- 0.5125, - 77.1511, 400 m), 18 xi 1997, leg. & det. TWD [FSCA, SWD]; SU, Shushufindi: La Selva Lodge; Lake Mandicocha and outlet stream (- 0.4983, - 76.3733, 200 m), 25 xi 1997, leg. & det. TWD [SWD]. Telebasis versicolor Fraser, 1946. Publ. Recs. ECU: Machado (2010), (P 11), (H 14); OR: * Garrison (2009 — as NA), von Ellenrieder & Garrison (2011). Also COL: (H 08), Garrison (2009), (P 11), (P & P); plus PER (P 16). Addl. Data. OR, Francisco de Orellana: 3 ♂, Reserva Etnica Waorani, 1 km S Onkone Gare Camp, Transect Ent. Fogged Lot # 927 (- 0.6528, - 76.4333, 220 m), 7 x 1994, leg. T. Erwin, det. RWG. [RWG, USNM]; Río Shiripuno, Shiripuno Lodge, 26 km SE of Shiripuno bridge (- 1.1047, - 76.7317, 235 m), data provided by WAH & DLW; Loreto: 2 ♂♂, forest pool about 12 km NE of Loreto, (0.6215, - 77.2601), 3 xi 2013, leg. & det. KJT [FSCA]; SU, Shushufindi: small ponds and tributaries of Río Shushufindi, 2 km W of Shushufindi on oil pipeline road (- 0.1810, - 76.6920, 265 m), data provided by WAH & DLW; 11 km NNE of Tarapoa, large non-flowing laguna below high bridge N of Cuyabeno village and deep muddy pool bordered by forest (- 0.0270, - 76.3179, 230 m), data provided by WAH & DLW.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B52EFFEBFF23F784244FF98E.taxon	discussion	This Neotropical genus of 12 known species is represented by one species in Ecuador of uncertain identity. The genus was described and treated in Machado (2009 a). Tuberculobasis sp. LR, Buena Fé: 1 ♀, Río Palenque Biological Sta., 47 km S of Santo Domingo de los Colorados, (- 0.5500, - 79.3667, 151 m), 8 x 1988, leg. SWD, det. RWG [?]; 1 ♂, 1 ♀, same data but 9 x 1988, data provided by RWG. Adults have a green-striped thorax and orange-tipped abdomen (Fig. 31). Note: Although Rosser Garrison examined these specimens, they could not be located at the time of publication to be re-studied.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B52DFFE9FF23F6882780FE13.taxon	discussion	Shade-dwelling, dark damselflies that tend to remain perched (Fig. 32); for identification, see the papers by Machado (2001, 2004, 2009 b). Habitats are seepage areas and slow, muddy, small streams in forest. Two of the three known species occur in Ecuador; the nymphs of this genus are unknown. Amazoneura ephippigera (Selys, 1886). Publ. Recs. ECU: Machado (2000), (L 05), Pessacq (2008), (H 08), (P 11), (H 14); OR: Tennessen (1996 b), Daigle (1996), von Ellenrieder & Garrison (2011). Also BRA, PER: (L 05), (H 08), (P 11). Addl. Data. SU, Shushufindi: 4 ♂♂, 1 ♀, Limoncocha, Río Napo (- 0.4000, - 76.6176, 269 m), 1 xi 1980, leg. MJW & D. Robinson, det. MJW [FSCA]; 7 ♂♂, same data but 5 xi 1980 [FSCA]; 7 ♂♂, same data but 18 xi 1980 [FSCA]; 1 ♂, same data but 24 xi 1980 [FSCA]; 2 ♂♂, same data but 25 xi 1980 [FSCA]; OR, Francisco de Orellana, 1 ♀, Reserva Etnica Waorani, 500 m N Onkone Gare Camp (- 0.6333 - 76.6000, 220 m), 10 i 1994, leg. T. Er- win, det.? [USNM]; 2 ♂♂, P. N. Y., 14.5 km N of Waorani Road on Yasuni Road, flowing Heliconia swamp (- 0.65, - 76.46, 200 m), 11 vi 1995, leg. & det. WFM [FSCA]; 4 ♂♂, P. N. Y., tiny stream (- 0.6508, - 763558, 200 m), 15 xi 1997, leg. & det. TWD [SWD]. Amazoneura westfalli (Machado, 2001). Publ. Recs. ECU: (H 08), (P 11), (H 14); OR: von Ellenrieder & Garrison (2011); SU: * Machado (2001 — Holotype: “ Napo: Limoncocha on Río Napo, 300 m, 15 xi 1980, leg. MJW [FSCA]; Allotype: same data but edge of ravine on hunting trail, 23 – 28 viii 1980, leg. MJW & SWD [FSCA]). Also PER: (P 11). Addl. Data. NA, Tena: 2 ♂♂, Puerto Napo, 23 km E, J. S. B. S. (400 m), 17 x 1988, leg. & det. SWD [SWD]; 1 ♀, Río Chinchipino, 27 km E of Puerto Napo (- 1.0728, - 77.5864, 385 m), 22 vii 1996, leg. & det. KJT [KJT]; OR, Francisco de Orellana: 1 ♂, small stream, 3.6 km from Yasuni Sci. Res. Sta., 18 vii 1996, leg. & det. KJT [FSCA]; 2 ♂♂, same data but leg. & det. JJD [JJD]; 3 ♂♂, forest seep, main road, P. N. Y., 15 xi 1997, leg. & det. KJT [KJT]; 2 ♂♂, seepage stream 2.1 km on Road E of station, 15 xi 1997, leg. & det. JJD [JJD]; 30 ♂♂, 6 ♀♀, T. B. S., USFQ, [various locations] (- 0.6180, - 76.1600, 200 – 245 m), 14 – 22 xi 2009, leg. & det. RWG & NVE [NMNHL, NVE, RWG, USFQ]; 2 ♂♂, Río Shiripuno, Shiripuno Lodge, 26 km SE of Shiripuno bridge (- 1.1047, - 76.7317, 235 m), 23 i 2009, leg. DLW & WAH, det. RWG [RWG]; Loreto: 1 ♂, Concepcion, Río Napo (- 1.6667, - 77.4167, 400 m), 28 xii 1939, leg. WCM, det. RWG, from UMMZ [RWG]; 1 ♂, forest stream, about 12 km NE of Loreto, 12 x 2009, leg. & det. KJT [FSCA]; 1 ♂, small streams and forest trails, 12 km E of Loreto, 2 + km S of Hwy. 45 (- 0.6217, - 77.2600, 365 m), 12 x 2009, leg. & det. JTJ [JTJ]; PA, Pastaza: 1 ♀, Canelos [a mission and jungle town on Río Bobonaza at headquarters of canoe navigation in Río Pastaza watershed] (- 1.5833, - 77.8000, 600 m), 16 xii 1938, leg. WCM, det. RWG, from UMMZ [RWG]; SU, Shushufindi: 1 ♀, Limoncocha, forest trail (- 0.4000, - 76.6167, 269 m), 24 i 1972, leg. D. Pearson, det. DRP [DRP]; 1 ♀, same data but 13 iii 1972; 1 cop. pair, 7 ♂♂, 1 ♀, same data but forest stream, 24 – 25 vii 1977, leg. & det. DRP [DRP, RWG]; 1 ♂, same data but 1 ix 1978, leg. KWK, det. RWG [RWG]; 3 ♂♂, 2 ♀♀, same data but, Río Napo, 15 – 24 xi 1980, leg. MJW, det. KJT [FSCA]; 1 ♂, 3 ♀♀, same data but edge of ravine on hunting trail, 28 viii 1980, leg. & det. SWD [FSCA]; 1 ♂, Limoncocha Road, 15.7 km S of Shushufindi Road, 19 vii 1996, leg. & det. KJT [FSCA]. Note: The distinctness of A. westfalli from A. ephippigera is questionable, as the male cerci are nearly identical; more study should be devoted to these taxa.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B52CFF97FF23F411268AFAC7.taxon	discussion	Dark, very small damselflies that easily avoid being seen (Fig. 33). See von Ellenrieder & Garrison (2008 b) for a key to the three species recorded in Ecuador. Eggs are laid in rootlets in slow or moderately flowing portions of small to mid-sized forested streams (pers. obs., KJT). Drepanoneura loutoni von Ellenrieder & Garrison, 2008. Publ. Recs. ECU: (H 14); MS: * von Ellenrieder & Garrison (2008 b). Also PER: Paulson (2011); plus BOL (P 16). Addl. Data. ZC, Nangaritza: 1 paratype ♂, “ Mayaicu ” [mistakenly published as “ Morona-Santiago Prov- ince: Mayalico ” on Río Naganitza ” (- 3.0592, - 77.9956, 490 m) [corrected as Mayaicu, Río Nangaritza, (- 4.0290, - 78.6340, 900 m)], D. Laddey, det. NVE & RWG [UMMZ]. Note: Should be found elsewhere on the eastern slope of the Andes in southern Ecuador since it has been found at a number of locations in Peru. Drepanoneura muzoni von Ellenrieder & Garrison, 2008. Publ. Recs. ECU: (P 11), (H 14), Bota-Sierra et al. (2015); NA: * von Ellenrieder & Garrison (2008 b); OR: * von Ellenrieder & Garrison (2008 b — Holotype: Francisco de Orellana; P. N. Y., small stream down Waorani Road (- 0.6333, - 76.5000, 180 m), 18 vii 1996, leg. KJT, det. NVE & RWG [FSCA]), von Ellenrieder & Garrison (2011); SU: * von Ellenrieder & Garrison (2008 b). Also PER: (P 11); plus COL (P 16). Addl. Data. NA, Tena: [FSCA, JJD, KJT, RWG, SWD]; OR, Francisco de Orellana: [FSCA, JJD, KJT, SWD, RWG]; Loreto: [JTJ]; PA, Santa Clara: [RWG]; SU, Shushufindi: [DRP, FSCA, JTJ, RWG]. Note: Specific location data can be found in von Ellenrieder & Garrison (2008 b). Drepanoneura tennesseni von Ellenrieder & Garrison, 2008. Publ. Recs. ECU: (P 11), (H 14); NA: * von Ellenrie- der & Garrison (2008 b — Holotype: Río Chinchipino, E of J. S. B. S., 27.7 km E of Puerto Napo (- 1.0728, - 77.5864, 385 m), 13 vii 1996, leg. KJT [FSCA]); OR: * von Ellenrieder & Garrison (2008 b); PA: * von Ellenrieder & Garrison (2008 b); SU: * von Ellenrieder & Garrison (2008 b). Addl. Data. NA: Carlos Julio Arsemena Tola: [RWG]; Tena: [FSCA, JJD, KJT, RWG]; OR, Loreto: [KJT, RWG]; PA, Pastaza: [FSCA, KJT]; SU, Lago Agrio: [RWG, USNM]; Shushufindi: [RWG]. Note: Endemic. Detailed locality data for most of the above records are in von Ellenrieder & Garrison (2008). Epipleoneura Williamson 1915 Small dark damselflies closely resembling Drepanoneura (Fig. 34). For a synopsis of the known species of Epi- pleoneura consult Pessacq (2014). Other references include Selys (1886), Williamson (1915), De Marmels (1989), Racenis (1955) and Garrison et al. (2010). Habitats are shaded streams, similar to those where Drepanoneura is found and oxbow lakes; the two genera are sometimes found on the same stream. * Epipleoneura humeralis (Selys, 1886). Publ. Recs. BRA: (H 08). Addl. Data. OR, Francisco de Orellana: 5 tan. pairs, 3 ♂♂, 1 ♀, T. B. S. (- 0.6353, - 76.1653, 200 m), leg. ACR, det. RWG [ACR, RWG]; SU, Shushufindi: 3 ♂♂, La Selva, jungle lodge, Río Napo, 70 km east of Coca; Laguna Garzacocha (- 0.4978, - 76.3741, 262 m), 12 x 1988, leg. SWD, det. JJD [JJD]; 1 ♂, same but, 22 x 2009; leg. WAH & Fred Morrison, det. RWG [RWG]. Note: this specimen was listed in (H 14) as “ sp. A ” cf. pallida. Epipleoneura lamina Williamson, 1915. Publ. Recs. ECU: (H 14); OR: * von Ellenrieder & Garrison (2011). Also GUY, PER, VEN: (H 08), (P 11); plus BRA, SUR: (P 16). Addl. Data. OR, Francisco de Orellana: 1 ♂, T. B. S., USFQ, Lago trail crossing river (- 0.6381, - 76.1558, 219 m), 21 i 2009, leg. & det. RWG & NVE [RWG]; SU, Shushufindi: 1 ♂, 1 ♀, La Selva Lodge, Río Napo, 70 km E of Coca, oxbow / laguna (- 0.4978, - 76.3741, 220 m), date not provided, data provided by WAH & DLW; SU, Putumayo: 1 ♂, Cuyabeno Reserve, Río Canabuena, 13 km east of Cuyabeno village, Río Canabuena, seasonal outflow of Laguna Canabuena to Río Cuyabeno (- 0.0404, - 76.1970), 7 vii 2015, leg, DLW, WAH, Fred Morrison, Michael Veit, Daniel Padilla & Valerie Giles, det. RWG, data provided by RWG; 1 ♀, same but at vegetated inlet edge (- 0.0424, - 76.1966). Epipleoneura venezuelensis Racenis, 1955. Publ. Recs. (H 14). Also BRA, VEN: (H 08), (P 16); plus COL: (Palacino 2017). Addl. Data. SU, La Joya de los Sachas: 1 ♀, La Joya de los Sachas, E at small stream (- 0.2838, - 76.7817), 29 x 2013, leg. & det. KJT [FSCA]; Shushufindi: 1 cop. pair, 1 ♀, Shushufindi, SW at small stream (- 0.2067, - 76.6919), 29 x 20132, leg. & det. KJT [FSCA]; 1 ♂, 1 ♀, Río Eno tributary, small stream in pasture on road from Lago Agrío to La Joya, (- 0.0509, - 76.8064, 300 m), date not provided, data provided by WAH & DLW; 1 ♂, 1 ♀, small stream in pasture, 2 m wide, deep and meandering, 9 km W Shushufindi on highway from Shushufindi to La Joya (- 0.1860, - 76.7305, 270 m), date not provided, data provided by WAH & DLW; 1 ♂, 1 ♀, slow, deep meandering stream in forest and pasture, 5 km S of Shushufindi on road to Limoncocha (- 0.2393, - 76.6477, 260 m), date not provided, data provided by WAH & DLW. Note: Rosser Garrison (pers. comm. 2019) suspects that more than one species may be involved under the name E. venezuelensis. Epipleoneura unresolved: a. Epipleoneura n. sp. NA, Tena: ♂, road south to Pulini from main road to airport and La Punta, 21 km southeast of Tena, 1 km south of road junction, large pond (- 1.0743, - 77.6521); 22 i 2011; leg. DLW, WAH, Fred Morrison & Ronald Vargus. det. RWG [RWG]; OR: Francisco de Orellana: 8 ♂♂, 8.2 km W of Río Payamino ferry (- 0.4578, - 77.0381, 288 m), 18 xi 97, leg. KJT [FSCA] 5 ♂♂, same data but leg. & det. JJD [JJD]; 1 ♂, pond, Loreto Road, 11.8 km W of Río Payamino ferry (- 0.4742, - 77.0058, 285 m), 18 ix 1997, leg. KJT [FSCA]. b. Epipleoneura haroldoi? Santos 1964 [USNM # 391333). PA, Santa Clara: Santa Clara, 4 viii 1976, leg. P. Turner, det.? [USNM]. Note: This specimen was not checked and the determination is doubtful. c. Epipleoneura cf. solitaria De Marmels, 1989. OR, Francisco de Orellana: 1 ♂, 1 ♀, Río Shiripuno, Shi- ripuno Lodge, 26 km SE of Shiripuno bridge, shaded bottomland forest away from water, oxbow forest (- 1.1047, - 76.7317, 235 m), 23 i 2009, leg. DLW & WAH, det. RWG [RWG].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B552FF95FF23F3DD21F4FBC3.taxon	discussion	Neoneura damselflies are slightly stockier than most others in the subfamily. Four species have been recorded for Ecuador; adults are often predominantly red and black (Fig. 35 a), although one species is green and yellow with black stripes (Fig. 35 b). Another species that is mostly blue and red has not been found in Ecuador but most likely occurs there. The genus was revised by Garrison (1999), although one species was described since (see Wasscher & Van’T Bosch 2013). Most species inhabit slow to moderate streams and rivers in forest, hovering close to the water surface; they can be extremely wary of movement. Neoneura confundens Wasscher & Van’t Bosch, 2013. Publ Recs. ECU: * Wasscher & Van’t Bosch (2013), (P 16). Also ARG, BOL, BRA, COL, GUF, GUY, PER, SUR, TTO, VEN: Wasscher & Van’t Bosch (2013), (P 16). Addl. Data. EO, Santa Rosa: 2 ♂♂, stream, 12 km E of Arenillas, Hwy. 25 (- 3.5244, - 79.9831, 30 m), 29 iii 2008, leg. & det. KJT — as N. bilinearis [FSCA]; LR, Montalvo: 3 ♂♂, small stream 1.8 km N of Hwy. 491, 25 km E of Babahoyo (- 1.8111, - 79.3189), 7 iii 2010, leg. & det. JJD [JJD]; 2 ♂♂, same data but leg. & det. KJT — as N. bilinearis [FSCA]; 2 ♂♂, same data but leg. & det. JTJ [JTJ]. Note: All prior records for N. bilinearis from Ecuador should be this species. The listing by Wasscher & Van’t Bosch (2013) “ E of Babahoyo, pers. comm. JTJ ” is based on the locality data herein. The Ecuador record is the only Pacific slope record based on the locality map of Wasscher and Van’t Bosch (2013). This widespread species should be found on the east slope also. Neoneura denticulata Williamson, 1917. Publ. Recs. ECU: Garrison (1999), (L 05) (H 08), (P 11), (H 14); OR: * Daigle (1996), von Ellenrieder & Garrison (2011). Also BRA, PER, VEN: (H 08), (P 11); plus SUR: (P 16). Addl. Data. OR, Francisco de Orellana: 1 ♂, Río Savaleto 2, Yasuni road at km 31.0, Yasuni Preserve (- 0.6569, - 76.4528, 216 m), 17 vii 1996, leg. & det. JJD [JJD]; 1 ♂, road S from Pompeya, Río Bogi 1, Río Tiputini tributary, 32 km SE of Pompeya, 5 km W of Yasuni Station, (- 0.6758, - 76.4394, 230 m), date not provided, data provided by WAH & DLW. Neoneura rubriventris Selys, 1860. Publ. Recs. ECU: Garrison (1999), (H 08), (P 11), (H 14); * Tennessen (1996 b); OR: Dunkle (1981), SU: Donnelly (1998). Also BRA GUY, PER, VEN: (L 05); (P 11); plus BOL: (H 08); and COL: Palacino et al. (2017). Addl. Data. OR, Francisco de Orellana: 25 ♂♂, 1 ♀, [La Nueva Esperanza] [across Río Napo from] near Pri- mavera “ Limoncocha ” Río Napo & lake Taracoa (- 0.45, - 76.77, 300 m), 27 vii 1977, leg. DRP & S. Hills, det. DRP [DRP, FSCA, RWG]; 2 ♂♂, same data but 26 viii 1980, leg. & det. SWD [KJT, SWD]; 1 ♂, same data but 4 xi 80, leg. MJW & D. Robinson, det. MWJ [FSCA]; 3 ♂♂, trib. of Río Payamino, road to Loreto, 1.7 km W of jct. with Coca Road (- 0.4367, - 77.0017, 258 m), 13 vi 1995, leg. & det. KJT [KJT, FSCA]; 1 ♀, stream 6.2 km down Waorani Road, P. N. Y., 16 xi 1997, leg. & det. KJT [FSCA]; SU, Shushufindi: 4 ♂♂, [Limoncocha] E, N of La Selva Lodge, Lago Mandicocha & outlet stream “ coca ” (about 100 km E on Río Napo) (- 0.4983, - 76.3733, 200 m), 25 xi 1997, leg. & det. TWD [FSCA, SWD]; 2 ♂♂, La Selva, Hyacinth Lake, 2 v 1990, leg. E. Smith, det. WFM [SWD]. Neoneura rufithorax Selys, 1886. (H 14 as “ cf. ”) Publ. Recs. BRA, PER: (H 08), (P 16); plus COL: (P 16); and (VEN): (DeMarmels 2016). Addl. Data. OR, Francisco de Orellana: 1 ♂, Estacion Cientifica Yasuni, Río Tiputini, 39 km SE of Pompeya, 61 km ESE of Coca, (- 0.6758, - 76.4394, 230 m), date not provided, data provided by WAH & DLW.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B550FF90FF23F2C124DCFCAB.taxon	discussion	Very slender, beautifully colored damselflies (Figs. 36 a & 36 b), with at least six species in Ecuador. The synopsis by von Ellenrieder & Garrison (2017) is the main source for identification of Ecuadorian species in addition to the papers by Cowley (1941), Gloyd (1939) and Kennedy (1939 c). Habitat includes small, sluggish forested streams, where adults perch mostly in shade. Protoneura amatoria Calvert, 1907. Publ. Recs. ECU: Lencioni (2005), (H 08), (P 11), (H 14); GY: * Campos (1922), von Ellrieder & Garrison (2017); EO: Campos (1927), von Ellenrieder & Gasrrison (2017); LR: von Ellrieder & Garrison (2017); MN: von Ellrieder & Garrison (2017). Also PER, VEN, TTO: (L 05), Pessacq (2008), (H 08), (P 11); plus CEN: (H 08), (P 11); and COL: (P & P); and BRA: (P 16). Addl. Data. GY, Naranjal: 5 ♂♂, San Rafael, [as “ San Raphael ”] (- 2.18, - 79.23, 200 m), iv-v (no year pro- vided), leg. F. Campos R, det. GHB [FSCA]; LR, Buena Fé: 2 ♂♂, 2 ♀♀, forest slough at Río Palenque Biological Sta., 47 km S Santo Domingo de los Colorados (- 0.5500, - 79.3667, 200 m), 18 vii 1977, leg. & det. DRP [DRP]; Montalvo: 3 ♂♂, 3 ♀♀, small stream 1.8 km N of Hwy. 491, 25 km E of Babahoyo (- 1.8111, - 79.3189, 60 m), 7 iii 2010, leg. & det. JJD [JJD, FSCA]; 2 ♂♂, same data but leg. & det. KJT [FSCA]; 1 ♂, 1 ♀, same data but leg. & det. JTJ [JTJ]; Quevedo: 1 ♀, stream 16 km E of Hwy. E 25 [from Formisa] (- 0.7256, - 79.3483, 200 m), 24 iii 2010, leg. & det. JTJ [JTJ]; MN, Flavio Alfaro: 1 pair, 2 ♂♂, small forest stream 0.5 km N of Chindul, Hwy. E 15 (0.2389, - 79.8742, 30 m), 1 iii 2010, leg. & det. KJT [FSCA]; 1 cop. pair, 1 ♂, same data but leg. & det. JTJ [JTJ]; Pedernales: 1 ♂, Río Nalpe, about 3 km E of El Achote (0.0547, - 79.9739, 130 m), 28 ii 2010, leg. & det. JTJ [JTJ]. Protoneura klugi Cowley, 1941. Publ. Recs. ECU: (H 08), (P 11), (H 14); OR: von Ellenrieder & Garrison (2011, 2017); SU: von Ellenrieder & Garrison (2017). Also PER: (H 08), (P 11). Addl. Data. OR, Francisco de Orellana: 1 ♂, P. N. Y, swamp (- 0.6508, - 76.3558, 200 m), 16 xi 1997, leg. & det. TWD [SWD]; 38 ♂♂, 3 ♀♀, T. B. S., USFQ, [various locations within station] (- 0.6370, - 76.1500, 220 – 245 m), 11 – 18 i 2009, leg. & det. RWG & NVE [DRP, RWG, NMNHL, NVE, USFQ]; 1 ♂, Río Shiripuno, Shiripuno Lodge, 26 km SE of Shiripuno bridge; oxbow forest swamp (- 1.1047 - 76.7317, 235 m), 23 – 26 i 2009, leg. D. Wagner & W. Haber, det. RWG [RWG]. Protoneura macintyrei Kennedy, 1939. Publ. Recs. ECU: Paulson (1983), Davies & Tobin (1984), Tsuda (1986, 1991), Steinmann (1997 a), (H 08), (P 11), (H 14); ES: Daigle et al. (1997), von Ellenrieder & Garrison (2017); GY: von Ellenrieder & Garrison (2017); LR: * Kennedy (1939 c — Holotype & Allotype: “ Prov. Los Ríos, 30 m, 3 ♀♀ ” [UMMZ]), Bridges (1993), Garrison et al. (2003), von Ellenrieder & Garrison (2017); SD: von Ellenrieder & Garrison (2017). Also COL Palacino et al. (2017). Addl. Data. ES, San Lorenzo: 2 ♂♂, [San Antonio] near Urbina, 36.5 km WNW of Lita on [old] San Lorenzo Hwy. pools / seeps / stream (0.95, - 78.68, 200 m), 3 ii 1997, leg. & det. WFM [FSCA]; 3 ♂♂, same data but leg. & det. KJT [JJD, KJT]; LR, Buena Fé: 1 ♂, 1 ♀, [just S of Patricia Piliar], Río Palenque Biological Sta., 47 km S Santo Domingo de los Colorados (- 0.5500, - 79.3667, 200 m), 18 vii 1977, leg. & det. DRP [DRP]; 4 ♂♂, same data but 9 x 1988, leg. & det. SWD [RWG, SWD]; Quevedo: 3 ♂♂, stream 16 km E of Hwy. E 25 [from Formisa] (- 0.72565, - 79.3483, 200 m), 23 ii 2010, leg. & det. KJT [KJT, FSCA]; 1 cop. pair, 3 ♂♂, leg. & det. JTJ [JTJ]; 1 ♂, 24 ii 2010, leg. & det. KJT [FSCA]. Note: IUCN Red List status is NT, Near Threatened (IUCN 2018). Protoneura paucinervis Selys, 1886. Publ. Recs. ECU: * (L 05), (H 08), (P 11), (H 14); SU: von Ellenrieder & Garrison (2017). Also PER, BRA: Lencioni (2005), (H 08), (P 11); plus COL Bota-Sierra et al. (2015), (P 16); and VEN: De Marmels (2016); and BOL, GUY: von Ellenrieder & Garrison (2017. Addl. Data. SU, Shushufindi: 6 ♂♂, 3 ♀♀, Limoncocha, lake edge, 5 – 20 vii 1971, leg. D. Pearson, det. DRP [DRP]; 1 ♂, 1 ♀, 26 vii 1977, leg. & det. DRP [DRP]; 1 ♂, 1 ♀, same data but 22 viii 1980, leg. & det. SWD [SWD]; 14 cop. pairs, 1 ♂, 1 ♀, Limoncocha on Río Napo (300 m), 12 – 25 xi 1980, leg. & det. MJW [FSCA]. Protoneura scintilla Gloyd, 1939. Publ. Recs. ECU: (L 05), (H 08), (P 11), (H 14), Bota-Sierra et al. (2015); NA: * Tennessen (1998); SU: von Ellenrieder & Garrison (2017). Also BRA: (L 05), (H 08), (P 11); plus COL, PER: (P 16). Addl. Data. NA, Tena: 1 cop. pair, 1 ♂, 1 ♀, Río Chinchipino, 27 km E of Puerto Napo (- 1.0728, - 77.5864, 385 m), 22 vii 1996, leg. & det. KJT [FSCA]; 2 cop. pairs, same data but 13 xi 1997, leg. & det. KJT [FSCA]; OR, Loreto: 1 ♂, small streams and forest trails 12 km E of Loreto, 2 + km S of Hwy. 45 (- 0.6217, - 77.2600, 365 m), 15 x 2009, leg. & det. JTJ [JTJ]; SU, Shushufindi: 2 ♂♂, stream about 3 km NW of Shushufindi, 2 + km N off Hwy. 26 at km marker 17 (- 0.1664, - 76.6917, 290 m), 13 x 2009, leg. & det. KJT [FSCA]; 2 ♂♂, same data but leg. & det. JTJ [JTJ]; 1 ♂, Shushufindi, about 3 km W, small stream, (- 0.1646, - 76.6878), leg. & det. KJT [FSCA]. Protoneura woytkowskii Gloyd, 1939. Publ. Recs. ECU: Cowley (1941 — as “ subspecies a ”), Soukop (1954), Paulson (1983), Tsuda (1991), Daigle (2005), Pessacq (2008), (H 08), (P 11), (H 14); NA: * Gloyd (1939 — Allotype: “ Oriente Prov. Río Jatun Yacu, 2200 feet, i 1935, leg. WCM, from CHK [UMMZ]); OR: Daigle (1996), von Ellenrieder & Garrison (2011, 2017); PA * Gloyd (1939), Tennessen (2006), von Ellenrieder & Garrison (2017); SU: von Ellenrieder & Garrison (2017). Also PER, BOL: (H 08), (P 11); plus BRA: (P 16); plus COL: Bota-Sierra et al. (2018). Addl. Data. NA, Tena: 1 ♂, Jatun Yacu, Río Napo watershed (- 1.0167, - 77.8333, 700 m), 1 – 31 i 1935, leg. WCM, det. L. Gloyd, from UMMZ [RWG]; Río Chinchipino, oxbow pool, 27 km E of Puerto Napo (- 1.0728, - 77.5864, 385 m), 13 xi 1997, leg. & det. KJT [FSCA]); 3 ♂♂, same data but leg. & det. JJD [FSCA]; OR, Francisco de Orellana: 2 ♂♂, stream, road to Waorani Reserve, 5.6 km S of Yasuni Sci. Res. Sta., 10 vi 1995, leg. & det. KJT [KJT]; 1 ♂, stream, road to Waorani Reserve, 6.2 km S of Yasuni Sci. Res. Sta., 10 v 1995, leg. & det. KJT [KJT]; 2 cop. pairs, 12 ♂♂, P. N. Y., 29 km N of Waorani Road on Yasuni Road, stream (- 0.4917, - 76.5584, 183 m), 11 vi 1995, leg. & det. WFM [FSCA, QZAC, USNM]; 2 ♂♂, same data but leg. & det. KJT [KJT]; 4 ♂♂, same data but leg. & det. SWD [JJD, SWD]; 1 ♀, P. N. Y., 14.5 km N of Waorani Road on Yasuni Road, flowing Heliconia swamp (- 0.65, - 76.46, 200 m), 11 vi 1995, leg. & det. WFM [FSCA]; 1 ♂, [Pompeya Sur] S on [Pompeya Sur-Iro road] at km marker 38, small stream (- 0.6419, - 76.4606, 260 m), 12 vi 1995, leg. & det. WFM [FSCA]; 5 ♂♂, same data but leg. & det. KJT [KJT]; 1 ♂, same data but leg. & det. SWD [SWD]; 3 ♂♂, P. N. Y., Yasuni Biological Research Sta. along trails and roads within 5 km of Res. Sta. (- 0.6719, - 76.3992, 245 m), 18 vi 1996, leg. & det. WFM [FSCA, QZAC, USNM]; 1 tan. pair, 3 ♂♂, P. N. Y., 1.8 km W of Waorani Reserve Road on Markus Oil Co Road PC 29 - 30, small stream (- 0.6861, - 76.4726, 200 m), 16 xi 1997, leg. & det. WFM [FSCA, LSAM, QZAC]; 1 ♂, same data but leg. & det. KJT [KJT]; 5 ♂♂, 1 ♀, stream, PC 29 - 30 Road, 3.2 km from jct. with park road., P. N. Y., 18 vii 1996, leg. & det. KJT [KJT]; 1 ♂, same data but 10 vi 1995, leg. & det. SWD [SWD]; 2 tan. pairs, 10 ♂♂, T. B. S., USFQ, black water stream crossing Guacamayo trail at marker 2050 (- 0.6383, - 76.1492, 220 m), 16 – 17 i 2009, leg. & det. RWG & NVE [NMNHL, NVE, RWG, USFQ]; 2 tan. pairs, 21 ♂♂, 5 ♀♀, T. B. S., USFQ, stream crossing Numa trail at marker 100, muddy bottom, shady (- 0.6383, - 76.0103, 220 m), 10 i 2009 [NMNHL, NVE, RWG, USFQ]; 4 ♂♂, T. B. S., USFQ, stream crossing, Chorongo trail (- 0.6378, - 76.1486, 224 m), 12 i 2009, leg. & det. RWG & NVE [NVE, RWG, USFQ]; 1 tan. pair, 2 ♂♂, 1 ♀, T. B. S., USFQ, stream crossing Chorongo trail at marker 225 (- 0.6350, - 76.1503, 220 m), 16 i 2009, leg. & det. RWG & NVE [NVE, RWG, USFQ]; 2 ♂♂, T. B. S., USFQ, swamp at marker 650 of Parahuaco trail (- 0.6397, - 76.1453, 229 m), 2 i 2009, leg. & det, RWG & NVE [RWG]; T. B. S., USFQ, stream crossing Matapalo trail at marker 450 (- 0.6369, - 76.1458, 245 m), 14 i 2009, leg. & det. RWG & NVE [NVE, RWG, USFQ]; 1 ♂, T. B. S., USFQ, stream crossing Matapalo trail (- 0.6294, - 76.1544, 246 m), 20 i 2009, leg. & det. RWG & NVE [USFQ]; 2 ♂♂, Río Shiripuno, Shiripuno Lodge, 26 km SE of Shiripuno bridge; forest swamp with Calathea, (- 1.1047, - 76.7317, 235 m), 23 – 26 i 2009, leg. D. Wagner, det. RWG [RWG]; Loreto: 1 ♂, 1 ♀, Cotopino (- 1.00, - 78.70, 400 m), 1 – 28 ii 1950, leg. WCM, det. L. Gloyd, from UMMZ [RWG]; 3 ♂♂, forest stream about 12 km NE of Loreto (- 0.6217, - 77.2600, 365 m), 12 x 2009, leg. & det. KJT [KJT]; PA, Santa Clara: 1 ♂, Río Arajuno, Dayuma Camp (- 1.1000, - 77.5833, 434 m), 22 – 25 ix 1990, leg. J. Belle, det. RWG [RWG]; Mera: 5 ♂♂, 1 ♀, Río Pastaza, near Puyo (- 1.4667, - 77.9833, 1000 m), 1 – 31 xii 1948, leg. WCM, det. L. Gloyd, from UMMZ [RWG]; 2 ♂♂, small streams / trails, Los Copales, between Mera and Shell (- 1.4919, - 78.0725, 1070 m), 20 ix 2005, leg. & det. KJT [KJT]; 1 ♂, same data but 22 ix 2005, leg. & det. WFM [FSCA; 1 cop. pair, same data but 7 x 2009, leg. & det. JTJ [JTJ]; SU, Shushufindi: 1 ♂, Limoncocha, small forest stream, 24 viii 1980, leg. & det. SWD [SWD]; 2 ♂♂, trails, small streams 3 km W of Shushufindi, 2 + km N off Hwy. 26 at km marker 17 (- 0.1664, - 76.6917, 290 m), 14 x 2009, leg. & det. JTJ [JTJ].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B555FF91FF23F57924FAFD3B.taxon	discussion	Very slender, small damselflies, the three species in Ecuador being mostly dark with red markings (Figs. 37 a & 37 b). For identification, see Garrison (2004) and Tennessen (2016). Nymphs unknown; habitat appears to be small seepage streams in forest. Psaironeura angeloi Tennessen, 2016. Publ. Recs. ES, San Lorenzo: * Tennessen (2016 — Holotype & Allotype: small stream 5.6 km NW of Lita [on San Lorenzo Hwy, near Finca Consuegros] (0.893, - 78.510, 1967 m), 4 ii 1997, leg. KJT [FSCA]). Addl. Data. ES, San Lorenzo: 5 ♂♂, 1 ♀, Lita, 1 km NW along RR tracks, seeps, (- 0.8792, - 78.4589, 535 m), 5 ii 1997, leg. JJD, det. KJT [JJD, FSCA, RWG]. LR, Quevedo: 1 ♂, stream 16 km E of Hwy. E 25 (from Formisa) (- 0.7356, - 79.3483, 200 m), 23 ii 2010, leg. JTJ, det. KJT [JTJ]. MN, Flavio Alfaro: 3 ♂♂, Chindul, 0.5 km N, small forested stream (0.2389, - 79.8742, 30 m), 1 iii 2010, leg. JTJ, det. KJT [JTJ]. 2 ♂♂, same data but leg. JJD [JJD, FSCA]. Note: All previous references to Psaironeura remissa Calvert from Ecuador should refer to this species. It appears to be a west slope species. Also CEN: KJT (2016); plus COL: (Palacino 2017). Psaironeura bifurcata (Sjöstedt, 1918). Publ. Recs. ECU: (H 14); NA: * Garrison (2004); PA: * Garrison (2004), Tennessen (2006). Also BRA: (L 05), (H 08), (P 11); Plus VEN: (H 08), (P 16); plus PER (P 16); plus COL: Bota-Sierra et al. (2018). Addl. Data. OR, Loreto, 1 ♂, small stream near Reserva Río Bigal, about 30 km N of Loreto (- 0.5642, - 77.4114, 900 m), 16 v 2009, leg. & det. KJT [FSCA]; PA, Mera, 2 ♂♂, seepage streams, Los Copales Cabanas between Mera and Shell, W of Puyo, Hwy. 50 (- 1.4886, - 78.0708, 1021 m), 19 ix 2005, leg. & det. JJD [JJD]; 8 ♂♂, same data but 20 v 2005, leg. & det. WFM [FSCA, LSAM, QZAC, USNM]; 2 ♂♂, same data but 22 v 2005 [FSCA]; 2 ♂♂, 1 ♀, same data but leg. & det. KJT [KJT]; 1 ♂, same data but 4 x 2009 [FSCA]; 1 ♂, same data but 5 x 2009, leg. & det. JTJ [JTJ]; SU, Shushufindi: 1 ♂, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 8 xi 1974, leg. B. Drummond III, det. JJD [FSCA]: Note: Garrison (2004) lists additional specific locality data. Psaironeura tenuissima (Selys, 1886). Publ. Recs. ECU: Tsuda (1986, 1991), (L 05), (H 08), (P 11), (H 14), Bota- Sierra et al. (2015); NA: * Santos (1968 — as P. cerasina), Garrison (2004); OR: von Ellenrieder & Garrison (2011); PA: * Santos (1968 — as P. cerasina), Garrison (2004). Also BRA, GUY, PER: (L 05), (H 08), (P 11); plus GUF: (P 11), COL, SUR: (P 16). Addl. Data. NA, Tena: 1 ♂, J. S. B. S., small stream (- 1.0500, - 77.7833, 450 m), 16 vi 1994, leg. A. Jaramillo A, det. DRP [DRP]; 1 ♂, same data but 17 x 1988, leg. & det. SWD [SWD]; OR, Francisco de Orellana: 2 ♂♂, stream near Yasuni Research Station, Yasuni Preserve, 18 vii 1996, leg. & det. KJT [JJD]; 1 ♂, same data but leg. SWD [FSCA]; 3 ♂♂, stream, 3.6 km from Yasuni Sci. Res. Sta., 18 vii 1996, leg. & det. KJT [FAL, KJT]; 2 ♂♂, stream, PC 29 - 30 Road, 3.2 km from jct. with Yasuni Road, 18 vii 1996, leg. & det. KJT [KJT]; 1 ♂, forest seep, main road., P. N. Y., 15 xi 1997, leg. & det. KJT [KJT]; 2 ♂♂, 1 ♀, P. N. Y., swamp (- 0.6508, - 76.3558, 200 m), 16 xi 1997, leg. & det. TWD [SWD]; 1 cop. pair, 11 ♂♂, 1 ♀, T. B. S., USFQ, [various locations] (- 0.6383, - 76.1492, 220 - 240 m), 1 – 2 i 2009, leg. & det. RWG & NVE [NVE, RWG, USFQ]; Loreto: 1 ♂, stream about 12 km NE of Loreto (- 0.6217, - 77.2600, 365 m), 15 x 2009, leg. & det. KJT [FSCA]; 1 ♂, same data but leg. & det. JTJ [JTJ]; 1 ♂, Río Pacai-Yacu “ Paciyacu ”, “ Concepcion-Río Napo ” (- 1.6667, - 77.4167, 400 m), 28 xii 1939, leg. WCM, det. RWG [UMMZ]; 2 ♂♂, Cotopino [? Río Cotopino, between Napo and Francisco de Orellana] (- 1.00, - 78.70, 400 m), ii 1950, leg. WCM, det. RWG [RWG, UMMZ]; SU, Shushufindi: 1 ♂, Limoncocha, Río Napo (300 m), leg. B. Drummond III, det. WFM, from KWK [FSCA]; 1 ♂, streams in hill forest S of Napo River from La Selva, 13 x 1988, leg. & det. SWD [SWD].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B55BFF9FFF23F33A2619FE13.taxon	discussion	A monotypic genus of large damselflies (Fig. 38), the nymph of which is unknown. Assumed to breed in phytotelmata as do all others of the subfamily for which the nymphs are known. Anomisma abnorme McLachlan, 1877. Publ. Recs. ECU: * Kirby (1890 — as Microstigma terminatum), Campos (1922 — as M. terminatum), Schmidt (1942 — as A. terminatum), Paulson (1983), Tsuda (1986, 1991), Steinmann (1997 a), (H 08), (P 11), (H 14); OR: von Ellenrieder & Garrison (2011); PA: Selys (1886 — “ pebas-Río Napo ”). Also COL, PER, VEN: (H 08), (P 11). Addl. Data. NA, Carlos Julio Arsemena Tola: 1 ♂, Río Anzu (- 1.2000, - 77.8333, 700 m), 12 viii 1934, leg. WCM, det. RWG [UMMZ]; 1 ♀, Río Chucapi (- 0.95, - 77.75, 599 m), 26 ix 1934, leg. WCM, det. RWG [UMMZ]; Tena: 1 ♂, 1 ♀, Río Napo (- 1.0333, - 77.5000, 429 m), date?, leg. & det. McLachlan- 1877 [BMNH]; 1 ♀, Río Ja- tun Yacu (- 1.0167, - 77.8333, 700 m), iv 1937, leg. WCM, det. CHK [UMMZ]; OR, Francisco de Orellana: 1 ♀, Reserva Etnica Waorani, 1 km S of Okone Gare Camp (- 0.6571, - 76.4530, 216 m), 16 i 1994, leg. T. Erwin, det.? [USNM]; 1 ♀, T. B. S., USFQ, along Guacamayo trail at marker 800 (- 0.6314, - 76.1608, 241 m), 16 i 2009, leg. & det. RWG & NVE [NVE]; 1 ♀, same data but open grassy area behind dining room, surrounded by bushes and trees and some seepage areas (- 0.6383, - 76.1492, 220 m), 16 i 2009, leg. & det. RWG & NVE 2009 [RWG]; 1 ♂, 1 ♀, same data but Matapalo trail at marker 1200 (- 0.6403, - 78.1444, 230 m), 20 i 2009, leg. & det. R. W. Garrison & NVE [NVE]; Loreto: 1 ♂, forest about 12 km NE of Loreto (- 0.6217, - 77.2600, 365 m) 15 x 2009, leg. & det. KJT [FSCA]; PA, Pastaza: 1 ♂, Río Bobonaza (- 1.5833, - 77.8000, 600 m), date?, leg. M. Buckley, det. McLachlan, data provided by RWG [BMNH]; 1 ♂, Ish Pinga-Yacu, Río Napo (- 2.2167, - 77.0666, 500 m), 26 ix 1934 leg. WCM, det. MJW [FSCA]; SU, Shushufindi: 1 ♂, Limoncocha, forest trail (- 0.4000, - 76.6167, 269 m), 1 iii 1972, leg. D. Pearson, det. DRP [DRP-FSCA]; 1 ♂, same data but 13 iii 1972 [DRP-FSCA]; 1 ♀, same data but 22 vii 1977 [DRP-FSCA]; 1 ♀, same data but Río Napo, 5 ix 1972, leg. T. Emmel, det. MJW [FSCA]; 1 ♂, same data but 13 v 75 [FSCA]; 1 ♂, same data but 14 v 1975 [FSCA]; 1 ♀, same data but 14 xi 73, leg. B. Drummond III, det. MJW [FSCA]; 1 ♂, same data but 4 iii 1974, [FSCA]; same data but 12 ix 1974 [FSCA]; 1 ♀, Providencia trail to Lago Anago Río Napo (- 0.47, - 76.47, 300 m), 9 v 1975, leg. B. Drummond III, det. MJW [FSCA]; TU, Baños: 1 ♂, 1 ♀, Baños “ supposedly ” (- 1.40, - 78.42, 1829 m), vi 1985, by Peter Rosier where it was purchased [FSCA]; this record needs verification by further collecting in the Baños area.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B55AFF9DFF23F1BC2742F8CF.taxon	discussion	Largest genus in the subfamily with ten species, six of which are recorded for Ecuador. These are very long damselflies with narrow wings and yellow or white wing tips (Fig. 39). Numerous subspecies have been named; the group is in need of revision. References include Selys (1886), Calvert (1901, 1948), and Hedström & Sahlén (2001). A recent revision by Machado & Lacerda (2017) removed some of the species from Mecistogaster and placed them in Kennedy’s Platystigma, a genus that was previously synonymized. We have not followed this change in this paper. Mecistogaster buckleyi McLachlan, 1881. Publ. Recs. ECU: Kirby (1890), Campos (1922), Paulson (1983), Davies & Tobin (1984), Tsuda (1986, 1991), Steinmann (1997 a — also as M. garleppi), (H 08), (P 11), (H 14); PA: * McLachlan (1881 a — Type: “ Río Bobonaza ” [- 1.5833, - 77.8, 600 m]), Selys (1886), Kimmins (1970 — Lectotype [BMNH]), Bridges (1993), (L 05). Also PER: (L 05), (H 08), (P 11); plus BRA: (H 08). Mecistogaster jocaste Hagen, 1869. Publ. Recs. ECU: Kirby (1890), Campos (1922 — “ indicated by Calvert ”), Paulson (1983), Tsuda (1986, 1991), (H 08), (P 11), (H 14); PA: * McLachlan (1881 a), Selys (1886). Also BRA, COL, PER, VEN: (H 08), (P 11). Addl. Data. LR, Quevedo: 1 ♀, Pichilingue (- 1.1000, - 79.4833, 84 m), 20 iv 1970, leg. C. Williner, det. RWG (Senckenberg); NA, Archidona: 1 ♀, Cotundo, about 15 km N. Tena, (800 m), 17 xii 1996, leg. Nicolay, det.? [USNM]; Tena: 1 ♀, Río Sinde, 5 km E of Puerto Napo (- 1.0500, - 77.7481, 450 m), 13 vii 1996 leg. & det. KJT [KJT-FSCA]; 2 ♀♀, Río Chinchipino, 27 km E of Puerto Napo (- 1.0728, - 77.5864, 385 m), 22 vii 1996, leg. & det. KJT [KJT-FSCA]; SU, Lago Agrio: 1 ♂, Lago Agrío (0.1000, - 76.8667, 396 m), 24 v 1976, leg. C. Williner, det. RWG (Senckenberg); 1 ♂, Lago Agrio, 16 iv 1976, leg. Langley, det.? [USNM]; Shushufindi: 1 ♂, Limonco- cha, lake edge, 16 i 1972, leg. D. Pearson, det. DRP [DRP-FSCA]; 1 ♀, Limoncocha, 8 vi 1977, leg. Pinto, det.? [USNM]. Mecistogaster linearis (Fabricius, 1777). Publ. Recs. ECU: Campos (1922 — “ por Dr. Calvert ”), Geijskes (1971), Paulson (1983), Tsuda (1986, 1991), (L 05), (H 08), (P 11), (H 14); OR: von Ellenrieder & Garrison (2011); PA: * McLachlan (1881 a), Selys (1886); SU: Donnelly (1998). Also BOL, BRA, COL, GUF, GUY, PER, SUR, VEN: (L 05), (H 08), (P 11); plus CEN: (P 11). Addl. Data. NA: 1 ♂, 1 ♀, (500 m), 11 vi 1988, leg. R. Robbin, det.? [USNM]; Archidona: 1 ♂, Archidona (- 0.9167, - 77.8000, 604 m), 4 xii 1976, leg. Herman G. Real, det. RWG [RWG]; OR, Francisco de Orellana: 1 ♂, forest, Río Savaleto, P. N. Y., 17 vi 1996, leg. & det. KJT [FSCA]; 1 ♀, T. B. S., TLE Plot T- 2, Sta. 9, Lot # 2818, fogging canopy (- 0.6319, - 76.1442, 220 – 250 m), 24 vii 2002, leg. T. Erwin, et. al, det. WFM [USNM]; 1 ♀, T. B. S., USFQ, Parahuaco trail between markers 300 – 1300 (- 0.6397, - 76.1453, 229 m), 21 i 2009, leg. & det. RWG & NVE [USFQ]; SU Lago Agrio: 1 ♀, Santa Cecilia, Río Francisco de Orellana, 25 – 31 iii 1969, leg. P. Spangler, det.? [USNM]; Shushufindi: 1 ♂, 1 ♀, Limoncocha, forest trail (- 0.4000, - 76.6167, 269 m), 5 v 1971, leg. D. Pearson, det. DRP [DRP-FSCA]; 1 ♀, same data but 24 i 1972, [DRP-FSCA]; 1 ♂, same data but Río Napo, 9 ix 1958, leg. T. Emmel, det. MJW [FSCA]; 1 ♀, same data but 9 ix 1958, [FSCA]; 2 ♂♂, 5 xi 1980, leg. & det. MJW [FSCA]; 1 ♂, 1 ♀, same data but 7 xi 1980, [FSCA]; 1 ♀, 28 i 1974, leg. B. Drummond III, det. MJW [FSCA]; 1 ♂, same data but 3 – 16 vi 1977, leg. P. Spangler, det.? [USNM]; 1 ♂, La Selva Lodge; forest (- 0.4983, - 76.3733, 200 m), 22 xi 1997, leg. & det. TWD [SWD-FSCA]; TU, Baños: 1 ♂, Baños, (- 1.38, - 78.42, 1800 m), 1938, leg. J. Schunke, det. MJW [FSCA]. Mecistogaster lucretia (Drury, 1773). Publ. Recs. ECU: Paulson (1983), Tsuda (1991), (L 05), (H 08), (P 11), (H 14); OR: von Ellenrieder & Garrison (2011); PA: * McLachlan (1881 a). Also BRA, GUF, GUY, PER, SUR, VEN: (L 05), (H 08), (P 11); plus ARG, COL: (P 16). Addl. Data. OR, Francisco de Orellana: 1 ♀, T. B. S., USFQ, stream crossing Chorongo trail at marker 225 (- 0.6350, - 76.1503, 221 m), 18 i 2009, leg. & det. RWG & NVE [NVE]; 1 ♂, same data but along Guacamayo trail between markers 2250 – 2500 (- 0.6181, - 76.1578, 245 m), 20 i 2009, leg. & det. RWG & NVE [NVE]; Loreto: 1 ♂, small streams and forest trails 12 km E of Loreto, 2 + km S of Hwy. 45 (- 0.6217, - 77.2600, 365 m), 12 x 2009, leg. & det. JTJ [JTJ]; SU, Shushufindi: 1 ♂, Limoncocha, forest trail, 14 i 1972, leg. D. Pearson, det. DRP [DRP- FSCA]. * Mecistogaster modesta Selys, 1860. Publ. Recs. ECU: (H 14). Also CEN, COL, VEN: (H 08), (P 11). Addl. Data. CB, Cumandá: 1 ♀, Naranjapata, (- 2.27, - 79.08, 550 m), leg. F. Campos, det. P. Calvert, checked by GHB, checked by WFM, from ANSP [FSCA]. Mecistogaster ornata Rambur, 1842. Publ. Recs. ECU: (L 05), (H 08), (P 11), (H 14); EO: Campos (1927); LR: * Campos (1922). Also ARG, BRA, COL, GUF, PER, SUR, TTO, VEN: (L 05), (H 08), (P 11); Plus CEN: (H 08), (P 11); and COL (P 16). Addl. Data. GY Naranjito: 1 ♀, Naranjito (- 2.17, - 79.47, 30 m), leg. F. Campos, det. GHB [FSCA]; LR, Quevedo: 1 ♂, Quevedo, Río Palenque (- 1.03, - 79.43, 100 m), date?, leg. F. Campos, det. GHB [FSCA]; Babahoyo: 1 ♀, Babahoyo (- 1.80, - 79.53, 10 m), v, leg. F. Campos, det. GHB [FSCA]; MN, Pedernales: 1 ♂, Bosque Seco Lalo Loor, arroyo casi seco (- 0.0837, - 80.1472, 88 m), 30 xi 2012, leg. & det. ACRP [ACRP]; OR, Loreto: 1 ♂, near stream 8.5 km E of Loreto (- 0.6183, - 77.2950, 400 m), 14 xi 1997, leg. & det. KJT [FSCA]; SU, Shushufindi: 1 ♀, Limoncocha, forest trail (- 0.4000, - 76.6167, 269 m), 27 ix 1971, leg. D. Pearson, det. DRP [DRP-FSCA]; 1 ♀, same data but 6 i 1972 to 24 ii 1972 [DRP].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B558FF9BFF23F1B02085FE4B.taxon	discussion	The largest damselfly in the world; wings wide, tips subapically black with blue reflections plus a variable white mark (Fig. 40). The genus is monotypic, although size and coloration is variable among populations. This large spe- cies breeds in water-filled tree holes, including stumps. Megaloprepus caerulatus (Drury, 1782). Publ. Recs. ECU: * Selys (1876, 1886), Calvert (1906), Paulson (1983), Tsuda (1986, 1991), (P 11), (H 14). EO: Campos (1927); GY: * Campos (1922). Also BOL, CEN, COL, GUY, PER, VEN: (H 08), (P 11). Addl. Data. ES, San Lorenzo: 1 ♂, Playa de Oro, primary forest (220 m) 30 vii 1998, leg. T. Enriquez, det. DRP [DRP-FSCA]; GY, Naranjito: 1 ♀, Naranjito (- 2.17, - 79.47, 30 m), iv, leg. F. Campos, det.? [FSCA]; LR, Buena Fé: 1 ♀, Río Palenque Biological Sta., 47 km S Santo Domingo de los Colorados (- 0.5500, - 79.3667, 151 m), 28 ii 1977, leg. H. Real, det. SWD & RWG [RWG]; 1 ♀, [Consumulo] S Río Palenque (- 0.58, - 79.35, 175 m), 13 vi 1974, leg. J. Longino, det. MJW [FSCA]; Quevedo: 1 ♀, Quevedo, Río Palenque (- 1.03, - 79.43, 100 m), 23 v 1976, leg. T. Emmel, det. GHB [FSCA]; NA, Tena: 1 ♂, 1 ♀, Río Verde Yacu (- 0.7167, - 77.9667, 695 m), 4 iv 1984, leg. H. Real, det. RWG [RWG]; PC, Quito: sight record, 1 ♂, upper Mashpi road, Río Huaycuyacu above waterfall (0.1952, - 78.8657, 850 m), 2 ix 2018, det. KJT; SD: 1 ♀, Santo Domingo de los Colorados, 12 km E on Hwy. 30 near Hotel Tinalandia (- 0.30, - 79.67, 671 m), 13 v 85, leg. J. Nation, det. MJW [FSCA]; ZA: Zamora: Estacion Scientifica San Franzisco, Copalinga blue trail, stream, 1000 – 1900 m, no date provided, leg. & det. C. Willigalla.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B55EFF98FF23F42E200BFF6B.taxon	discussion	Of the three species in this genus, only one has been found in Ecuador. The wings are wide with brown and yellow tips (Fig. 41); breeding habitat is water-filled tree holes. Last revised by Schmidt (1958), this genus is in need of further revision (Garrison et al. 2010). Microstigma rotundatum Selys, 1860. Publ. Recs. ECU: * Kirby (1890), Campos (1922), Paulson (1983), Davies & Tobin (1984), Tsuda (1991), Steinmann (1997 a), (L 05), (H 08), (P 11), (H 14); NA: Schmidt (1958); OR: von Ellenrieder & Garrison (2011), Ingley et al. (2012); PA: * McLachlan (1881 a), Schmidt (1958); SU: Dunkle (1981), Donnelly (1998), Ingley et al. (2012); TU: Schmidt (1958). Also BOL, BRA COL, PER, VEN: (L 05), (H 08), (P 11). Addl. Data. NA, Archidona: 1 ♂, Archidona, about 8 km N of Tena (700 m), date?, leg. Nicolay, det.?, [USNM]; 1 ♀, Archidona, (- 0.9167, - 77.8000, 604 m), 3 x 1976, leg. H. Real, det. RWG 1977 [RWG]; 1 ♀, same data but 26 x 76 [RWG]; 2 ♀♀, same data but 2 xii 1976 [RWG]; Tena: 1 ♂, J. S. B. S., near Tena, primary forest (450 m), 18 vi 1994, leg. A. Jaramillo, det. DRP [DRP-FSCA]; 1 ♂, 1 ♀, Río Chinchipino (- 1.0728, - 77.5864, 385 m), 13 xi 1997, leg. & det. TWD [SWD-FSCA]; OR, Francisco de Orellana: 1 ♀, Primavera, forest trail, 27 vi 1977, leg. & det. DRP [DRP-FSCA]; 1 ♂, [La Nueva Esperanza] S of [across Río Napo from] Primavera Laguna Taracoa (- 0.45, - 76.77, 300 m), 26 vi 1980, leg. I. Finkelstein, det. MJW [FSCA]; 1 ♂, streams in hill forest S of Napo River from La Selva, 13 x 1988, leg. & det. SWD [SWD]; 1 ♂, P. N. Y., 18 x 2001, leg. D. Bogar, det. WFM [FSCA]; 1 ♀, P. N. Y., Yasuni Biological Research Sta. along trails and roads within 5 km of Res. Sta. (- 0.6719, - 76.3992, 244 m), 28 viii 2000, leg. T. Emmel, det. JJD [FSCA]; 1 ♂, T. B. S., USFQ, stream crossing Matapalo trail at marker 450 (- 0.6369, - 76.1458, 245 m), 14 i 2009, leg. & det. RWG & NVE [NVE]; 1 ♀, same data but stream crossing Chorongo trail at marker 225 (- 0.6350, - 76.1503, 220 m), 16 i 2009, leg. & det. RWG & NVE [NVE]; 1 ♀, same data but along Guacamayo trail at marker 1700 [Ti 23] (- 0.6350, - 76.1533, 237 m), 17 i 2009, leg. & det. RWG & NVE [RWG]; 1 ♂, 1 ♀, same data but Parahuaco trail between markers 300 – 1300 (- 0.6397, - 76.1453, 229 m), 21 i 2009, leg. & det. RWG & NVE [RWG, USFQ]; 1 ♀, same data but at base of canopy tower at marker 2600 of Gua- camayo trail (- 0.6364, - 76.1642, 219 m), 17 i 2009, leg. & det. RWG & NVE [USFQ]; 1 ♀, same data but clearing in forest along Harpia trail (- 0.6347, - 76.1511, 244 m), 11 i 2009, leg. & det. RWG & NVE [USFQ]; 1 ♂, same data but along Guacamayo trail at marker 800 (- 0.6314, - 76.1608, 241 m), 16 i 2009, leg. & det. RWG & NVE [RWG]; Loreto: 1 ♂, trickle, 30.7 km W of Loreto on road to Narupa, 14 vi 1995, leg. & det. SWD [SWD]; 1 ♂, Loreto, ne- arby streams (0.7164, - 77.6640, 376 m), 26 x 2013, leg. & det. Cary Kerst [Cary Kerst]; PA, Pastaza: 2 ♂♂, 2 ♀♀, [Sarayacu], date?, leg.?, det.? [USNM]; Santa Clara: 2 ♂♂, Río Arajuno, Camp Dayuma, 390 – 420 m (- 1.0931, - 77.5853), 19 – 22 iv 1990, leg. S. Weller, P. Batra, M Ryan, det.? [USNM]; SU, Gonzalo Pizarro: 1 ♀, 65 km W of Lago Agrio, roadside pond, 18 v 1975, leg. Langley, det.? [USNM]; Shushufindi: 1 ♂, Limoncocha (- 0.4000, - 76.6167, 269 m), 1972, leg. P. Kazan, det. SWD [SWD]; 1 ♂, 1 ♀, same data but 23 – 28 viii 1980, leg. & det. SWD [SWD]; 2 ♂♂, Limoncocha, forest trail, 19 vi 1971, leg. D. Pearson, det. DRP [DRP-FSCA]; 1 ♀, same data but 5 ix 1971 [DRP-FSCA]; 1 ♂, same data but 4 i 1972 [DRP-FSCA]; 1 ♂, same data but 10 i 1972 [DRP-FSCA]; 1 ♀, same data but forest stream, 22 ii 1972 [DRP-FSCA]; 1 ♂, same data but Río Napo, 9 i 1974, leg. B. Drummond III, det. MJW [FSCA]: 1 ♀, same data but 5 x 1974 [FSCA]; 1 cop. pair, same data but 19 vii 1974; 1 ♂, same data but 8 xi 1974 [FSCA]; 1 ♀, same data but 13 v 1975; 1 ♂, same data but 19 iii 1974, leg. T. Emmel, det. MJW [FSCA]; 1 ♀, same data but 16 v 1975 [FSCA]; 2 ♂♂, same data but 5 xi 1980, leg. & det. MJW [FSCA]; 2 ♂♂, forest near Río Napo, S of Limoncocha (- 0.4203, - 76.6108, 250 m), 11 v 2009, leg. & det. KJT [FSCA]; 1 ♀, forest about 3 km SW of Shushufindi (- 0.4203, - 76.6108, 250 m), 12 v 2009, leg. & det. S. Bybee & KJT [KJT-FSCA]; 1 ♂, 2 ♀♀, La Selva Lodge, forest (- 0.4983, - 76.3733, 200 m), 24 xi 1997, leg. & det. TWD [SWD-FSCA]; TU, Baños: 1 ♂, [Baños] Río Pastaza [presumably] (- 1.38, - 78.42, 1800 m), 26 xi 1936, leg. WCM, det. MJW [FSCA].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B55DFF98FF23F0DC246AF8CA.taxon	discussion	The Dicteriadidae contains only two monotypic genera. They are large to medium-sized damselflies that perch with the wings held out horizontally; the legs are very long and slender without conspicuous setae. Only one genus occurs in Ecuador.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B55DFF86FF23F1902112FF6B.taxon	discussion	A monotypic genus of large, blue and green damseflies that from a distance resemble slender gomphids (Fig. 42). For identification see Dunkle (1991 b). Adults perch on large leaves in sunny spots above flowing water. The wings flash multi-colored irridescence during flight. The nymphs live in root masses along the banks of small forested streams. Heliocharis amazona Selys, 1853. Publ. Recs. ECU: Silsby (2001), (L 05), (H 08), (P 11), (H 14); NA: * Dunkle (1991 b), Daigle (1996); OR: Tennessen (1998), von Ellenrieder & Garrison (2011). Also ARG, BOL, BRA, COL, GUF, GUY, PER, PRY, SUR, VEN: (H 08), (P 11). Addl. Data. NA, Tena: 1 ♂, 1 ♀, [Puerto Napo], Río Napo [presumably Puerto Napo] (- 1.04, - 77.78, 436 m), iii 1937, leg. WCM, det. SWD [FSCA]; 2 ♂♂, [Puerto Napo], [upstream near jct with Río Mulatos], Jatun Yacu Río Napo watershed (- 1.05, - 77.98, 700 m), xii 1934, leg. WCM, det. SWD [FSCA]; 1 ♂, 1 ♀, same data but 18 iv 1935 [FSCA]; 1 ♂, same data but 12 ii 1937 [FSCA]; 1 ♀, same data but iii 1937 [FSCA]; 1 ♀, [Puerto Napo], upstream of off Río Jatun Yacu Río Lloquillin — as “ Llupulin Creek ” (- 1.0935, - 77.9305, 541 m), i 1935, leg. WCM, det. SWD [FSCA]; 1 ♂, Puerto Napo 11.2 km E of Hwy. 45 on J. S. B. S. Road, Río Tiyuyacu (- 1.0516, - 77.7017, 400 m), 23 vii 1996, leg. & det. WFM [FSCA]; 1 ♂, 1 ♀, stream 14.6 km E of Puerto Napo, 23 vii 1996, leg. & det. JJD [JJD]; 7 ♂♂, Río Chinchipino, 27 km E of Puerto Napo (- 1.0728, - 77.5864, 385 m), 22 – 23 vii 1996, leg. & det. KJT [KJT]; 1 ♂, same data but 13 xi 1997 [KJT]; 4 ♂♂, same data but leg. & det. TWD [SWD]; OR, Francisco de Orellana: 1 ♂, slow stream, 6.4 km down Waorani Road, P. N. Y., 16 xi 1997, leg. & det. KJT; Loreto: 1 ♂, forest stream about 12 km NE of Loreto (- 0.6217, - 77.2600, 365 m), 12 x 2009, leg. & det. KJT [KJT]; PA, Aranjuno: 2 ♂♂, 1 ♀, Aranjuno upstream — SSW Río Aranjuno headwaters (- 1.3000, - 77.8010, 1000 m), 26 – 27 iv 1941, leg. WCM, det. SWD [FSCA]; 1 ♂, same data but 30 x 1941 [FSCA]; Pastaza: 2 ♂♂, 1 ♀, Puyo, Río Churi Yacu, downstream near mouth at Río Puyo (- 1.47, - 78.00, 900 m), 22 – 27 iii 1941, leg. WCM, det. SWD [FSCA]; SU, Lago Agrio: adults, Santa Cecilia, Río Francisco de Orellana, 25 – 31 iii 1969, leg. P. Spangler, det.? [USNM]; 1 ♂, [El Ono, about 8 km N], 26 km S of Lago Agrio, stream, 19 viii 1980, leg. KWK, det. SWD [KWK]; Shushufindi: 1 ♂, Limoncocha, forest stream, 16 ii 1972; leg. D. Pearson, det. DRP [DRP]; 4 ♂♂, same data but 6 – 17 iii 1972 [DRP, FSCA]; TU, Baños: 1 ♀, Río Pastaza watershed, 20 xi 1936, leg. WCM, det. SWD [FSCA].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B543FF86FF23F70E2185FE23.taxon	discussion	Two genera of Heteragrionidae occur in Ecuador. They are large to medium-sized damselflies that perch with the wings held out horizontally. All are forest stream species, from low to medium elevations.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B543FF85FF23F7F6208DFB57.taxon	discussion	Medium to large damselflies; the species in Ecuador are usually brown and black with yellow to yellowish orange markings (Fig. 43). Adults perch at an angle on stems and twigs mostly in shade or small sunny spots. Eight species have been recorded in Ecuador, but there is potential for undescribed species to be discovered as the genus is in need of revision. For identification see Williamson (1919), Lencioni (2005) and Vilela et al. (2019). Heckman (2008) is useful for narrowing down some of the choices but we advise caution in using this reference for this genus. Heteragrion aequatoriale Selys, 1886. Publ. Recs. ECU: Kirby (1890), Williamson (1919), Campos (1922), Schmidt (1942), Paulson (1983), Tsuda (1986, 1991), Steinmann (1997 a), (H 08), (P 11), (H 14); MS: Tennessen (2006); PA: * Selys (1886 — description). Also PER: (H 08), (P 16) plus COL: (P & P), (P 16). Addl. Data. CT, Sigchos: 1 ♀, San Francisco de las Pampas, Río Esmeraldes (- 0.43, - 79.0, 1219 m), 4 iv 1997, leg. E. Tapia, det. JJD [FSCA]. MS, Gualaquiza: 1 ♀, Gualaquiza (- 3.40, - 78.58, 800 m), v 1935, leg. F. Campos, det. MJW [ANSP]; Morona: 9 ♂♂, 1 ♀, forested stream 0.3 km past [N] Río Albanico, 19.8 km W of Macas on Guamote Road (- 2.2494, - 78.2022, 1540 m), 17 ix 2005, leg. & det. JJD [JJD, RWG]; 1 ♂, same data but leg. & det. KJT [RWG]: 1 ♂, same data but 18 ix 2005, leg. & det. WFM [FSCA]; 8 ♂♂, 1 ♀, small stream, 20.1 km W of Macas, road to Guamote (- 2.2489, - 78.2031), 17 ix 2005, leg. & det. KJT [KJT]; NA, Archidona: 1 ♂, 2 ♀, stream 47.4 km W of Loreto (1095 m), 14 vi 1995, leg. & det. KJT [KJT]; 2 ♂♂, same data but leg. & det. SWD [SWD]; 1 ♂, stream / waterfall, Loreto Road, 44 km E of Baeza-Tena Road, 15 vii 1996, leg. & det. KJT [KJT]; 1 ♂, stream / waterfall, Loreto Road, 40.2 km E of Baeza-Tena Road, 15 vii 1996, leg. & det. KJT [KJT]; 1 ♂, Baeza 10.6 km S on Hwy. 45, near Bermejo, seepage marsh, 24 vii 1996, leg. SWD, det. JJD [SWD]; PA, Mera: 4 ♂♂, small trib. of Río Anzu, 12.0 km N of Mera (- 1.4181, - 78.0528, 1190 m), 21 ix 2005, leg. & det. KJT [KJT]; 2 ♂♂, same data but 6 x 2009, leg. & det. JTJ [JTJ]; 3 ♂♂, Río Metolo, Los Copales cabanas between Mera and Shell, W of Puyo, Hwy. 50, 19 ix 2005, leg. & det. JJD [JJD]; 6 ♂♂, between Mera and Shell, Los Copales trails and forest N of Río Metolo (- 1.4886, - 78.0708, 1070 m), 20 ix 2005, leg. & det. WFM [FSCA]; 1 ♂, Mera, 12 km NE Río Anzu (- 1.4181, - 78.0528, 1190 m), 21 xi 2005, leg. & det. WFM [LSAM]; Pastaza: 1 ♂, stream 94.0 km N of Sucua, Hwy. 45, 9 xi 1997, leg. & det. JJD [JJD]; 2 ♂♂, stream 5.5 + km SE off Hwy. 45 (trib. of larger river), N of Puyo (- 1.3606, - 77.8653, 900 m), 9 x 2009, leg. & det. JTJ [JTJ]. Heteragrion angustipenne Selys, 1886. Publ. Recs. ECU: * Selys (1886), Kirby (1890), Campos (1922), Schmidt (1942), Kimmins (1970 — Lectotype from “ Ecuador ” [BMNH]), Davies & Tobin (1984), Tsuda (1986, 1991), Bridges (1993), Steinmann (1997 a), (L 05), (H 08), (P 11), (H 14). Also BRA, PER: (L 05), (H 08), (P 11). Addl. Data. MS: Huamboya [FSCA, KJT, RWG]; Limón Indanza: [JJD, RWG]; NA, Archidona: [FAL, KJT, RWG]; OR, Francisco de Orellana: [KJT, SWD]; Loreto: [FSCA, JTJ]; PA, Mera: [FSCA, JJD, KJT]; Pastaza: [FSCA, JTJ, KJT, USNM]; Santa Clara: [FSCA]; SU, Shushufindi: [JTJ]; ZC, El Pangui: [FSCA, KJT]; Yantzaza: [USNM]. Heteragrion bariai De Marmels, 1989. Publ. Recs. ECU: (L 05), * Daigle (2005), (H 08), (P 11), (H 14). Also BRA, PER, VEN: (L 05), (H 08), (P 11); plus COL: (P 16). Addl. Data. NA, Tena: 5 cop. pairs, 7 ♂♂, 2 ♀♀, Río Chinchipino, 27 km E of Puerto Napo (- 1.0728, - 77.5864, 385 m), leg. & det. KJT [FAL, KJT, RWG]; 1 ♂, 1 ♀, same data but 13 vii 1996, leg. & det. JJD [JJD]; 2 ♂♂, same data but 10 x 2009 [JTJ]; 1 ♂, Puerto Napo 11.2 km E of Hwy. 45 on J. S. B. S. Road, Río Tiyuyacu (- 1.0516, - 77.7017, 400 m), 23 vii 1996, leg. WFM, det. JJD [FSCA]; 5 ♂♂, 1 ♀, 23 km E of Puerto Napo, J. S. B. S. (400 m), 17 x 1988, leg. SWD, det. JJD [FSCA, SWD]; OR, Francisco de Orellana: 2 ♂♂, across Río Napo from La Selva, streams in hill forest, 13 x 1988, leg. SWD, det. JJD [SWD]; 1 ♂, P. N. Y., Río Savaleto, km marker 31, 17 vii 1996, leg. & det. KJT [KJT]; 1 ♂, P. N. Y., Yasuni Biological Research Station along trails and roads within 5 km of Res. Sta. (- 0.6719, - 76.3992, 244 m), 29 viii 2000, leg. T. Emmel, det. KJT [FSCA]; La Joya de los Sachas: 1 ♂, east of La Joya de los Sachas, small stream, (- 0.2838, - 6.7817, 265 m), 29 x 2013, leg. & det. KJT [FSCA]; Loreto: 9 ♂♂, Loreto, 8.5 km E on road to Coca Road rocky gravel stream (- 0.6183, - 77.2950, 400 m), 14 xi 1997, leg. WFM, det. JJD [FSCA, LSAM, QZAC, USNM]; 4 ♂♂, 1 ♀, same data but leg. & det. KJT [KJT]; 3 ♂♂, same data but 18 xi 1997, leg. & det. KJT [KJT]; 2 ♂♂, small streams and forest trails 12 km E of Loreto, 2 + km S of Hwy. 45 (- 0.6217, - 77.2600, 365 m), 12 x 2009, leg. & det. JTJ [JTJ]; SU, Cascales: 1 ♂, 37 km W of Lago Agrio, river, 20 viii 1980, leg. SWD, det. JJD [SWD]; Lago Agrio: 1 ♂, Santa Cecilia. Río Francisco de Orellana, 25 – 31 iii 1969, leg. P. & P. Spangler, det.? [USNM]; Shushufindi: 1 ♂, Limoncocha, forest stream (- 0.40, - 76.62, 300 m), 6 iii 1972, leg. D. Pearson, det. JJD [FSCA, DRP]; 14 ♂♂, Limoncocha, Río Napo (- 0.40, - 76.62, 300 m), 10 – 17 xi 1980, leg. MJW, det. JJD [FSCA]; 1 cop. pair, 7 ♂♂, 1 ♀, Limoncocha, 22 – 23 viii 1980, leg. SWD, det. JJD [SWD]; 1 ♂, 1 tan. pair, stream 1.2 km E of jct. Coca Road & Shushufindi Road, 16 vii 1996, leg. & det. KJT [KJT]; 1 ♀, small stream SW of Shushufindi, (- 0.2067, - 76.6919), 29 x 2013, leg. & det. KJT [FSCA]. Note: Previously often referred to as “ Ecuador sp. A. ” Heteragrion bickorum Daigle, 2005. Publ. Recs. ECU: (H 08), (P 11); NA: * Daigle (2005), OR: * Daigle (2005), von Ellenrieder & Garrison (2011), (H 14); SU: * Daigle (2005 — as “ Napo ” cop. pair Holotype & Allotype: Limoncocha (- 0.40, - 76.62, 300 m), 25 viii 1980, leg. SWD, [FSCA]). Also PER: (P 16); COL: Bota-Sierra et al. (2018). Addl. Data. MS: Huamboya: 3 ♂♂, 1 tan. pair, Huamboya N on Hwy. 45, small flowing stream in forest (- 1.80, - 77.95, 884 m), 16 ix 2005, leg. & det. WFM [FSCA, QZAC, USNM]; NA, Carlos Julio Arsemena Tola: 5 ♂♂, 19.3 km S of Puerto Napo at Zetzayco (- 1.0594, - 77.6428, 460 m), 21 vii 1996, leg. SWD, det. JJD [FSCA, RWG]; 1 ♂, same data but leg. & det. KJT [KJT]; Tena: 1 ♂, 1 ♀, Río Chinchipino, 27 km E of Puerto Napo (- 1.0728, - 77.5864, 385 m), 13 vii 1996, leg. & det. KJT [KJT]; 2 ♂♂, Puerto Napo, 2 nd stream E of Río Sinde on J. S. B. S. Road, 23 vii 1996, leg. SWD, det. JJD [FSCA]; 2 ♂♂, stream, Puyo Road, 7.1 km S of Río Napo, 21 vii 1996, leg. & det. KJT [KJT]; 3 ♂♂, stream, road to J. S. B. S., 16.8 km E of Puerto Napo (385 m), 22 vii 1996, leg. & det. KJT [KJT]; 2 ♂♂, same data but leg. SWD, det. JJD [FSCA]; 2 ♂♂, J. S. B. S., 23 km E of Puerto Napo; tropical wet forest, streams above flood plain (- 1.0500, - 77.7833, 450 m), 16 – 17 x 1988, leg. SWD, det. JJD [FSCA]; OR, Francisco de Orellana: 1 ♂, P. N. Y., small stream, 2.3 km E of Sci. Research Sta., 9 vi 1995, leg. & det. KJT; 2 ♂♂, 3 ♀♀, P. N. Y., stream, Maxus 29 & 30 road, 1.8 km W of jct. with Waorani Road, 10 vi 1995, leg. & det. KJT [KJT]; 6 ♂♂, same data but 18 vii 1996 [KJT]; 2 ♂♂, same data but leg. SWD, det. JJD [FSCA]; 3 ♂♂, P. N. Y., small stream, road to Waorani Reserve, 3.6 km S of turn to Sci. Res. Sta., 10 vi 1995, leg. & det. KJT [KJT]; 3 ♂♂, same data but 18 vii 1996 [KJT]; 1 ♂, P. N. Y., small stream, road to Waorani Reserve, 5.6 km S of turn to Sci. Res. Sta., 10 vi 1995, leg. & det. KJT [KJT]; 1 ♂, P. N. Y., small stream, km 38 on Yasuni Road, 12 vi 1995, leg. & det. KJT [KJT]; 1 ♂, P. N. Y., small stream, 5.1 km down Waorani Road (200 m), 16 xi 1997, leg. & det. KJT [KJT]; 1 ♂, T. B. S., USFQ, Parahuaco trail between markers 300 – 1300 (- 0.6397, - 76.1453, 229 m), 21 i 2009, leg. & det. RWG & NVE [USFQ]; 2 ♀♀, T. B. S., USFQ, along Guacamayo trail at marker 800 (- 0.6314, - 76.1608, 241 m), 16 i 2009, leg. & det. RWG & NVE [NVE, RWG]; 2 tan pairs, 15 ♂♂, T. B. S., USFQ, Parahuaco trail between markers 300 – 1300 (- 0.6397, - 76.1453, 229 m), 16 – 21 i 2009, leg. & det. RWG & NVE [DRP, RWG, USFQ]; 2 tan. pairs, 10 ♂♂, 3 ♀♀, T. B. S., USFQ, stream crossing Chorongo trail at marker 225 (- 0.6350, - 76.1503, 220 m), 16 – 18 i 2009, leg. & det. RWG & NVE [NVE, RWG, USFQ]; 1 ♂, T. B. S., USFQ, along Chichico trail (- 0.6300, - 76.1525, 250 m), 12 i 2009, leg. & det. RWG & NVE [USFQ]; 16 ♂♂, T. B. S., USFQ, stream crossing Chorongo trail (- 0.6378, - 76.1486, 224 m), 11 – 16 i 2009, leg. & det. RWG & NVE [NMNHL, NVE, RWG, USFQ]; 2 tan. pairs, 18 ♂♂, 2 ♀♀, T. B. S., USFQ, stream crossing Matapalo trail at marker 450 (- 0.6369, - 6.1458, 245 m), 14 – 18 i 2009, leg. & det. RWG & NVE NVE, RWG, USFQ]; 1 ♀, T. B. S., USFQ, stream crossing Numa trail at marker 100, muddy bottom, shady (- 0.6383, - 76.0103, 220 m), 10 i 2009, leg. & det. RWG & NVE [USFQ]; Loreto: 2 ♂♂, stream, Loreto Road, 8.5 km E of Loreto (- 0.6183, - 77.2950, 400 m), 20 vii 1996, leg. & det. KJT [KJT]; 1 ♂, same data but 14 xi 1997 [KJT]; 1 cop. pair, same data but leg. WFM, det. JJD [FSCA]; PA, Mera: 3 ♂♂, Río Metolo, Los Copales, between Mera and Shell, Hwy. 50 (- 1.4919, - 78.0725, 1070 m), 19 ix 2005, leg. & det. KJT [KJT]; 5 ♂♂, same data but leg. & det. WFM [FSCA, LSAM, QZAC, USNM]; Pastaza: 1 ♀, small stream, 2 km S of Comunidad Chuwitayo, road between Macas & Puyo, 9 xi 1997, leg. & det. KJT [KJT]; 1 ♂, Río Sandalias, 5 km S of Puyo, Hwy. 45 (- 1.5003, - 77.9522, 1000 m), 11 xi 1997, leg. & det. KJT [KJT]; SU, Shushufindi: 26 ♂♂, La Selva, 100 km E Coca, streams in hill forest S of Napo River (- 0.54, - 76.54, 262 m), 13 x 1988, leg. SWD, det. RWG [RWG, USNM]; 28 ♂♂, 1 ♀, same data but det. JJD [FSCA]; 1 ♀, Limoncocha, forest trail (- 0.4000, - 76.6167, 269 m), 10 i 1972, leg. D. Pearson, det. DRP [DRP]; 14 ♂♂, 3 ♀♀, same data but 28 viii 1980, leg. SWD, det. JJD [FSCA, SWD]; 3 ♂♂, 1 ♀, same data but Río Napo, 23 – 28 viii 1980, leg. SWD, det. RWG [FSCA, RWG]; 1 ♂, same data but 28 viii 1980, leg. & det. KWK [RWG]; 7 cop. pairs, 60 ♂♂, 1 ♀, same data but 1 – 25 xi 1980, leg. MJW, det. JJD [FSCA]; 2 ♂♂, trails, small streams 3 km W of Shushufindi, 2 + km N off Hwy. 26 at km marker 17 (- 0.1664, - 76.6917, 290 m), 14 x 2009, leg. & det. JTJ [JTJ]. Note: This species is taxonomically very close to H. bariai. Heckman (2008) suggested that more research is necessary to resolve the relationship between the two taxa. We are retaining it as a distinct species pending further study. It was often referred to as “ Ecuador sp. B. ” Heteragrion cooki Daigle & Tennessen, 2000. Publ. Recs. ECU: Paulson (2004 a), (H 08), (P 11); Lencioni (2014), (H 14); EO: * Daigle & Tennessen (2000), Rivas-Torres et al. (2017); GY: Tennessen & Johnson (2008); LJ: Tennessen & Johnson (2008); LR: * Daigle & Tennessen (2000 — as “ Pichincha ” Allotype: [Buena Fé], Río Palengue Biological Station, 47 km S of Santo Domingo de las Colorados (183 m), 8 x 1988, leg. SWD [FSCA]); SD: * Daigle & Tennessen (2000 — as “ Pichincha ”, Holotype: stream, W of Hotel Tinalandia, Hwy. 30, about 9 km W of Alluriquin, 31 i 1997, leg. JJD [FSCA]). Addl. Data. BO, Caluma: [JTJ, KJT]; CN, La Troncal: [RWG]; EO, Piñas: [RWG]; GY, Balao: [JTJ]; Naranjal: [FSCA]; LJ, Catamayo: [JTJ]; LR, Buena Fé: [DRP, FAL, FSCA, RWG, SWD]; Montalvo: [KJT]; Quevedo: [JTJ, KJT]; MN, El Carmen: [JTJ]; Flavio Alfaro: [KJT]; Pedernales: [JTJ, KJT]; SD: [DRP, FAL, FSCA, JJD, JTJ, KJT, RWG, SWD]. Note: This endemic, Pacific slope form was often referred to as “ Ecuador sp. E. ” Heteragrion erythrogastrum Selys, 1886. Publ. Recs. ECU: (H 08), (P 11), (H 14); ES: * Daigle et al. (1997). Also CEN, COL: (H 08), (P 11). Addl. Data. ES, San Lorenzo: 6 ♂♂, stream 36.5 km N [NW] of Lita, 3 ii 1997, leg. & det. KJT [KJT, SWD]; 2 ♂♂, same data but leg. E. Tapia, det. KJT [FAL, FSCA]; MN, Junin: 1 ♀, Palmar, Playones (- 0.9333, - 80.2333, 200 m), 6 v 1941, leg. D. Laddey, det. RWG [RWG]; 1 ♂, same data but 15 v 1941 [RWG]; 1 ♂, same data but 25 iv 1941 [RWG]; SD: 1 ♂, Santo Domingo de los Colorados, about 15 km east, “ Hacienda La Lorena ” (- 0.3097, - 79.0389, 700 m), 22 ii 1941, leg. D. Laddey, det. RWG [RWG]; 2 ♂♂, same data but 25 ii 1941 [RWG]; 1 ♂, same data but 26 ii 1941 [RWG]. Note: Southernmost records for this common Central American species. Heteragrion inca Calvert, 1909. Publ. Recs. ECU: (L 05), Daigle (2005), (H 08), (P 11), (H 14); OR: von Ellenrieder & Garrison (2011); Also BOL, BRA, PER: (L 05), (H 08), (P 11); plus COL: (P 16). Addl. Data. NA, Archidona: [FSCA]; Tena: [DRP, FSCA, JTJ, RWG, SWD, SWD]; Carlos Julio Arosemena Tola: [FSCA]; OR, Francisco de Orellana: [FSCA, KJT, RWG, SWD, TWD, USNM]; Loreto: [FSCA, TWD, USNM]; PA, Mera: [KJT]; Pastaza: [FSCA, JTJ]; Santa Clara: [RWG]; Shushufindi: [FAL, FSCA, RWG]. Note: Often referred to in collections as “ Ecuador sp. C ”; variation in H. inca male cerci throughout its range deserves further taxonomic research. Heteragrion peregrinum Williamson 1919, Publ. Recs. (H 14 as “ cf. ”) ES, Quininde: 1 ♂, 1 ♀, Río Canande Re- serve, 35 km NE of Quininde, tributary of Río Hoja Blanca, small gravel stream in forest, shady gap next to stream (0.5264, - 79.2122, 345 m), 21 x 2012, leg. & det. WAH, data provided by RWG, checked by RWG. Also: COL Palacino et al. (2017). Note: Bota-Sierra et al. (2016) reported this species as “ Critically Endangered, Possibly Extinct, ” but it has been found recently; therefore, “ possibly extinct ” no longer applies and we propose that it needs to be reassessed.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B540FF85FF23F32D2720F9EF.taxon	discussion	A genus of three species, one of which has been found in Ecuador (Fig. 44). See Garrison (2014) for identification. Adults are active in light rainfall. Habitat is small streams at various elevations. Oxystigma petiolatum (Selys, 1862). Publ. Recs. ECU: (H 14 — as Oxystigma sp. A: SU, Shushufindi: * Garrison (2014 – 1 ♂, Shushufindi pipeline, near oil pipeline, 3.5 km NW of Shushufindi, small stream at edge of forest – fly- ing in light rain (- 0.1661, - 76.6919, 295 m), 16 i 2011, leg. DLW, WAH, R. Vargus, det. RWG [WAH]). Also BRA, GUF, SUR, VEN: Garrison (2014).	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B540FF80FF23F0FC2630FE6F.taxon	discussion	A poorly known genus; with five species recorded for Ecuador; the group is in need of further study. See Garrison & von Ellenrieder (2005), Pérez-Gutiérrez & Montes-Fontalvo (2011 c) and Daigle (2014) for identification. Adults perch for most of the day in dark shade with wings together (Fig. 45), being difficult to spot. Nymphs live in sheet flow on nearly vertical rocks at the edges of small, forested, mountainous streams (Tennessen 2010). * Heteropodagrion croizati Pérez-Gutiérrez & Montes-Fontalvo, 2011. Publ. Recs. COL Pérez- Gutiérrez & Montes-Fontalvo (2011 c). Addl. Data. NA, Tena: Road to Pulini, 21 km SE of Tena, 1.8 km S of road junction, small forest stream, sand and mud with deep holes, over steep-walled creek in forest (- 1.0820, - 77.6500, 415 m), 22 i 2011, leg. R. Vargus, WAH & Fred Morrison, det. KJT [WAH]. Heteropodagrion nigripes Daigle, 2014. Publ. Recs. * Daigle (2014 — Holotype ♂: MS, Limón Indanza: stream, Hwy. 45, 25.3 km N of Indanza (- 2.8570, - 78.3700, 810 m), 7 xi 1997, leg. & det. JJD [FSCA]; Huamboya: * Daigle (2014); Morona: * Daigle (2014); PA, Mera: * Daigle (2014). Heteropodagrion sanguinipes Selys. 1885. Publ. Recs. ECU: (* Selys 1885 — Holotype ♀: lost, Syntypes “ Quito ” 2 ♂♂, 2 ♀♀, [Quito], Quito (- 0.2167, - 78.5, 2763 m), leg.? [IRSNB]), Bridges (1993 — Type-?), Karsch (1891 a — as Neuragrion mysticum, Type L = Ecuador), Bridges (1993), Paulson (1983), Davies & Tobin (1984), Tsuda (1986, 1991), Steinmann (1997 a — also as “ N. mysticum ”), (P 11), (H 14 — also as N. mysticum); Cotapaxi Province: Garrison & von Ellenrieder (2005); PC: Selys (1886), Kirby (1890), Campos (1922), Garrison & von Ellenrieder (2005) SD: Tennessen (2010 — nymph description). Addl. Data. CT, Pangua: 2 ♂♂, 75 km E of Quevedo, spring seep (- 0.93, - 79.22, about 900 m), 12 v 1975, leg. P. J. Spangler, det. RWG [USNM]; Sigchos: 11 ♂♂, 4 ♀♀, San Francisco de las Pampas Otonga Reserve, Orillas del Río Las Damas (- 0.42, - 79.0, 1600 m), 2 ii 1999, leg. L. Torres, det. JJD [QZAC]; 2 ♂♂, 2 ♀♀, same data but 5 ii 1999 [QZAC]; 1 ♂, same data but 12 iii 1999 [QZAC]; 2 ♂♂, same data but 19 iii 1999 [QZAC]; 1 ♀, same data but Orillas del Río Esmeraldas (- 0.43, - 79.0, 1400 m), 6 ii 1999, leg. E. Tapia, det. JJD [QZAC]; 2 ♂♂, same data but forest of Mr. Tapia property, 14 v 1999, leg. E. Tapia [QZAC]; 1 ♀, same data but in station near tube of water (- 0.42, - 79.0, 1800 m), 24 v 1999, leg. L. Torres, det.? [QZAC]; 1 ♂, same data but in station near the small house, 30 iv 1999, leg. T. Enriquez, det.? [QZAC]; 2 ♂♂, same data but on border of Las Damas River (- 0.42, - 79.0 W, 1450 m), 30 iv 1999, leg. L. Torres, det.? [QZAC]; 1 ♂, same data but on road at 6 m of Z River (1500 m), 1 v 1999 [QZAC]; PC, Mejía: 1 ♀, Cornejo Astorga, 3.5 km W, stream (- 0.0556, - 78.7714, 1520 m), 16 vii 1977, leg. DRP, det. RWG [DRP]; San Miguel Bancos: 10 ♂♂, 2.3 km S of Tandayapa (- 0.0144, - 78.6803, 1250 m), 6 ix 1990, leg. OSF, det. SWD [RWG, SWD, USNM]; 1 ♂, 1 ♀, Tandayapa, spring trickle in secondary forest (- 0.0167, - 78.7667, 1700 m), 6 v 2009, leg. & det. WAH [RWG]; 1 ♀, Mindo, Bosque Protector Mindo Nambillo (- 0.0556, - 78.7714, 1244 m), 28 i 2009, leg. & det. RWG & NVE [RWG]; SD: 2 ♂♂, about 84 km SW of Quito, Chiriboga Road, 9 vii 1990, leg. OSF, det. SWD [USNM]; 1 ♂, stream, Hwy. E 30 at km marker 61, E of La Unión del Toachi (- 0.5003, - 78.8700, 1210 m), 26 ii 2010, leg. & det. KJT [KJT]; 4 ♂♂, same data but 27 ii 2010 [KJT]; 2 ♂♂, same data but 25 ii 2010, leg. & det. JTJ [JTJ]; 1 ♂, same but, 26 ii 2010 [JTJ]; 1 ♂, 2 ♀♀, same data but 27 ii 2010; leg. & det. JJD [JJD]; 2 ♂♂, same data but leg. & det. JJD [FSCA]; 1 ♂, small waterfall stream, km marker 70.5, E of Santo Domingo de los Colorados, Hwy. E 30 (- 0.3139, - 78.9417, 910 m), 27 ii 2010, leg. & det. KJT [KJT]; 2 ♂♂, same data but leg. & det. JTJ [JTJ]; 2 ♂♂, 2 ♀♀, same data but leg. & det. JJD [JTJ]. Note: Endemic to Ecuador’s Pacific slope. * Heteropodagrion superbum Ris, 1918. Publ. Recs. COL (Ris 1918); plus PAN (H 08). Addl. Data. BO, Caluma: 2 ♂♂, 1 ♀, Caluma, 11 km E of stream, waterfall (- 1.5764, - 79.1919, 890 m), 5 iii 2010, leg. & det. KJT [KJT]; PC, Mejía: 1 ♂, Manuel Cornejo, 2.3 km S of Tandapi, waterfalls (- 0.43, - 78.78, 1738 m), 6 x 1990, leg. OSF, det. JJD [FSCA]. Note: Garrison et al. (2010) suggested that H. superbum may be a junior synonym of H. sanguinipes. We elect to retain it here as a valid species until further studies warrant otherwise. Heteropodagrion varipes Daigle, 2014. MS, Morona: Daigle (2014 — Holotype ♂, Macas 20.3 km W on Gua- mote Rd, 0.6 km W on Río Albanico, two rocky waterfall streams (- 2.2475, - 78.2067, 1536 m), 18 ix 05, leg. JJD [FSCA]). Note: All known specimens were collected along the road between Macas and Guamote at small waterfalls. Heteropodagrion unresolved: CT, Pujili: 2 ♂♂, 75 km E of Quevedo, spring seep (- 0.9333, - 79.2167, 900 m), 12 v 1975, leg. P. J. Spangler, det. RWG [USNM]; 1 ♂, 66 km E of Quevedo, spring seep (- 1.02, - 79.48, 1325 m), 15 i 1978, leg. P. J. Spangler et al., det. RWG [USNM]; PA, Mera: 1 ♂, Abitagua, Río Pastaza watershed (- 1.45, - 78.15, 1200 m), 2 x 1936, leg. WCM, det. RWG [UMMZ]; 1 ♂, Abitagua, on playa of Pastaza at mouth of Río Guiller- mina (- 1.45, - 78.15, 1200 m), 3 vii 1941, leg. WCM, det. RWG [USNM]; 2 ♂♂, Abitagua, Mayorga’s playa (- 1.45, - 78.15, 1200 m), 6 vi 1941, leg. WCM, det. RWG [UMNZ].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B545FF80FF23F5302629FB8B.taxon	discussion	Archilestes is mainly a North American genus; nine species have been described. There are few species and few records in South America (Garrison et al. 2010). The genus was not expected to occur in Ecuador, but recently a photograph of an Archilestes species was taken in Pichincha Province by Remco Hofland (Fig. 46); no specimen was collected and therefore its specific identity is unknown. It is possible that the species is A. chocoanus Pérez- Gutiérrez (2012), recently described from Colombia, or it is an undescribed species. Pertinent references on identification are Gloyd (1980), De Marmels (1982), Garrison (1982) and Pérez-Gutiérrez (2012). Archilestes typically inhabit slow-flowing streams with vegetated, pooled areas.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B545FF80FF23F37127BBF8EC.taxon	discussion	Publ. Recs. ECU: Calvert (1907), Soukop (1954), Paulson (1983), Tsuda (1986) — all as Lestes henshawi; * Donnelly (1996), (P 11), (H 14); MS: Tennessen (2006); NA: Tennessen (1996 b), Daigle (1996); PA: Kennedy (1942 b — ♀ described as L. henshawi. Also COL, PER, VEN: (H 08), (P 16). Addl. Data. MS, Morona: 1 ♂, rocky stream and seeps, 22.5 km W of Macas on Guamote Road, 18 ix 2005, leg. & det. JJD [JJD]; NA, Archidona: 1 ♂, stream 12.5 km N of Narupa, Baeza-Tena road (- 0.67, - 77.78, 1585 m), 14 vii 1996, leg. & det. JJD [JJD]; 1 ♂, same data but leg. & det. WFM [QZAC]; 1 ♂, same data but leg. & det. SWD [RWG]; Quijos; 1 ♂, [Bermejo], 10.6 km S of Baeza on Hwy. 45 near Bermejo, seepage marsh (- 0.5083, - 77.8789, 1800 m), 14 vi 1995, leg. & det. WFM [FSCA]; 1 ♂, same data but 16 vi 1995, leg. & det. KJT [KJT]; 1 cop. pair, same data but 24 vii 1996, leg. & det. KJT [KJT]; 7 ♂♂, Baeza 4.8 – 5.0 km S, small clear streams and seeps (- 0.4761, - 77.8711, 1830 m), 20 xi 1997, leg. & det. WFM [FSCA, QZAC, USNM]; 3 ♂♂, 2 ♀♀, same data but leg. & det. KJT [KJT]; 1 ♂, same data but 16 x 2009, leg. & det. JTJ [JTJ]; OR, Loreto: 1 ♀, Loreto 38.9 km W on road to Narupa, pond (- 0.7411, - 77.5569, 988 m), 14 vi 1995, leg. & det. WFM [FSCA]. Note: All records are from the east slope. Originally confused with L. henshawi, but the true L. henshawi is recorded only from Costa Rica (Donnelly 1996).	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B545FF80FF23F2752695FA8F.taxon	discussion	Long, slender damselflies, usually with blue and black markings, adults perch at an angle on stems and twigs (Fig. 47). Four species are known in Ecuador. No single reference addresses the South American species of Lestes; for identification of the Ecuador species, we recommend consulting Westfall & May (2006), Heckman (2008) and the papers in which the original species descriptions appeared as listed below. Lestes breed in vegetated pools, ponds and small lakes, occasionally in very slow areas of sluggish streams.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B545FF80FF23F43525D9FD73.taxon	discussion	Two genera of Lestidae occur in Ecuador. Lestes is a large genus with 45 New World species, although only four species are recorded for Ecuador. They are large to medium-sized damselflies that perch with the wings out at an angle.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B545FF81FF23F16D2732FDA7.taxon	discussion	Publ. Recs. ECU: (H 14); GY: * Campos (1922), Calvert (1928); EO: Campos (1927); LJ: Tennessen & Johnson (2008). Also ARG, BRA, CEN, GUF, PER, VEN: (H 08), (P 16); Plus TTO: (P 16); plus GUY: (H 08) and COL: P & P (P 11). Addl. Data. EO, Machala: 1 ♂, borrow pit pond, Hwy. 25, S of Machala (- 3.4750, - 79.9778), 10 iv 2008, leg. & det. KJT [FSCA]; Santa Rosa: 2 ♂♂, stream and associated pond, wetland about 12 km E of Arenillas, Hwy. 25 (- 3.5244, - 79.9831, 30 m), 29 iii 2008, leg. & det. JTJ [JTJ]; 1 ♂, borrow pit, Hwy. 25 S of Santa Rosa (- 3.4750, - 79.9778, 20 m), 10 iv 2008, leg. & det. JTJ [JTJ]; GY, Daule: 2 ♂♂, Daule, Nobol (5.5 km N), 30 m, 1 xii 1978, leg. P. Spangler & J. Anderson, det.? [USNM]; Duran: 1 ♂, Hwy. E 40, ditch complex E of Duran (- 2.1586, - 79.7828), 20 ii 2010, leg. & det. KJT [FSCA]; Guayaquil: 3 ♂♂, 1 ♀, Guayaquil (- 2.20, - 79.90, 8 m), iii vi, leg. F. Campos, det.? [FSCA]; 1 ♂, same data but iv 1904 [FSCA]; N aranjal: 1 ♂, Cerro Mas Vale E on Hwy. Río Churute (- 2.42, - 79.67, 9 m), 10 ii 2001, leg. & det. F. Sibley [FSCA]; LJ, Loja: 1 ♂, 1 cop. pair, pond, entrance to Podocarpus Nat. Pk. (- 4.0911, - 79.2025, 2475 m), 6 iv 2008, leg. & det. KJT [FSCA]; 1 cop. pair, same data but leg. & det. JTJ [JTJ]. Note: Confined to Pacific slope.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B544FF8EFF23F09B2710FE37.taxon	discussion	Publ. Recs. ECU: (H 08), (P 11), (H 14); OR: Loreto: * Tennessen (1998 — as “ NA ”, Holotype & Allotype: 1 cop. pair, Puerto Francisco de Orellana 12.3 km W on road to Loreto from Coca-Sachas Road, pond (- 0.4750, - 77.0747, 290 m), 13 vi 1995, leg. WFM FSCA]); PA: Tennessen (1998). Also PER, VEN: (P 11). Addl. Data. NA, Tena: 4 ♂♂, 1 ♀, Puerto Napo 5.3 km E on J. S. B. S. Road, Río Sinde, pond upstream (- 1.0517, - 77.7544, 450 m), 13 vii 1996, leg. & det. JJD [FSCA]; 1 ♂, 1 ♀, same data but leg. & det. SWD [RWG]; 1 ♂, same data but 16 x 1988, leg. SWD, det. WFM [SWD]; 2 ♂♂, 2 ♀♀, same data but leg. & det. JJD [DRP, FAL, JJD, RWG]; 2 ♂♂, 2 ♀♀, same data but 22 vii 1996 [JJD]; OR, Francisco de Orellana: 4 ♂♂, 1 ♀, trib. of Río Payamino, road to Loreto, 1.7 km W of jct. with Coca Road (- 0.4367, - 77.0017, 258 m), 13 vi 1995, leg. & det. KJT [KJT]; 1 ♂, same data but leg. & det. WFM [USNM]; 2 tan. pairs, 1 ♂, 1 ♀, same data but leg. SWD, det. WFM [SWD]; 3 cop. pairs, 1 paratype cop. pair, Puerto Francisco de Orellana, 12.3 km W on road to Loreto from Coca- Sachas Road, pond (- 0.4750, - 77.0747, 290 m), 13 vi 1995, leg. & det. WFM [FSCA, QZAC]; 3 ♂♂, same data but leg. & det. KJT [KJT]; 2 tan. pairs, 1 ♂, same data but leg. SWD, det. WFM [SWD]; PA, Pastaza: 2 ♂♂, Cuasha just N of Río Pastaza bridge, 90.5 km N of Sucua, small forested stream (- 1.88, - 77.80, 915 m), 9 xi 1997, leg. & det. WFM [FSCA, LSAM]; 1 ♂, pond about 3 km N of Comunidad Chuwitayo, S of Puyo, 9 xi 1997, leg. & det. KJT [KJT]. Note: Confined to East slope.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B54BFF8FFF23F069272DFF6B.taxon	discussion	Publ. Recs. ECU: Tsuda (1986, 1991), Paulson (1983, 2011), Steinmann (1997 a), (H 14); GY: * Ris (1918), Campos (1922), EO: Campos (1927). Also CEN, COL, SUR, VEN: (H 08), (P 11); Plus TTO (P 11). Addl. Data. GY, Daule: 1 ♀, Daule (- 1.85, - 79.98, 9 m), x 1987, leg. G. Onore, det. WFM [FSCA]; Guayaquil: 1 ♀, Puna Vieja (- 2.97, - 80.18, 30 m), leg. F. Campos, det. MJW [ANSP]; 1 ♂, Guayaquil vicinity, iii 1911, leg. F. Campos, det. P. Calvert [FSCA]; 1 ♂, Isla Puna, Puerto Grande, ii 1936, leg. F. Campos, det. WFM [FSCA]; Naranjal: 1 ♂, 1 ♀, San Rafael, iii iv, leg. F. Campos, det. P. Calvert [FSCA]; LR, Urandeta: 1 ♂, ditches / pools 2.8 km E of Hwy. E 25 from km marker 98 (- 1.5217, - 79.5000, 35 m), 21 ii 2010, leg. & det. JTJ [JTJ]; MN, El Carmen: 4 ♂♂, borrow pit lake, 18 km NW of El Carmen, 2 iii 2010, leg. & det. JJD [JJD]; Pedernales: 1 ♂, small stream about 2 km E of Pedernales (0.0836, - 80.0139, 90 m), 28 ii 2010, leg. & det. KJT [KJT]. Note: A Pacific slope species, apparently at relatively low elevations. Campos (1922) listed three specimens as Lestes sp. from GY, Milagro: Hacienda Venecia de Chimbo. These specimens could not be located but were probably either L. forficula or L. tenuatus.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B54AFF8FFF23F7CC2659F9B0.taxon	discussion	A monotypic genus recently discovered in Ecuador. See De Marmels (2001 a) for identification. Adults are mostly brown and tan with the tips of the abdomen grayish white (Fig. 48); they perch with their wings out. The nymphs and their habitat are apparently unknown. Megapodagrion megalopus Selys, (1869). Podagrion megalopus Selys, 1869. Synonym: Megapodagrion arachne Racenis, 1959. Publ. Recs. ECU (H 14). Also BRA, GUY, PER, VEN (H 08). Addl. Recs. SU, Cuyabeno: adult, Río San Francisco, Nicky Lodge on Río Cuyabeno (- 0.1977, - 75.9243, 210 m), date not provided, data provided by WAH & DLW [WAH].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B54AFF8FFF23F7392040FE37.taxon	discussion	Three genera are included in this Neotropical family (Dijkstra et al. 2014), two of which occur in Ecuador.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B54FFF8AFF23F55527E5FBAF.taxon	discussion	Perilestidae is a small family of extremely thin and elongate damselflies; the abdomen is approximately twice the length of the wings. Adults perch on small branches and leaf tips in shade, hanging vertically with wings slightly spread or sometimes closed together. Their habitat is small, sluggish, forested streams. Six species occur in Ecuador. For generic identification, see Garrison et al. (2010).	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B54FFF88FF23F274207FFF47.taxon	discussion	Only one of the eight species of this genus is reported for Ecuador (Fig. 50). Species identification is based on Kennedy (1941 a). Habitat appears to be slow, almost pooled stretches of small shaded streams, although very little is known of the biology. Perilestes kahli Williamson & Williamson, 1924. Publ. Recs. ECU: Tsuda (1986, 1991), Paulson (1983), Daigle (2005), (P 11), (H 14); NA: * Kennedy (1937 b); OR: von Ellenrieder & Garrison (2011). Also BOL, BRA, VEN: (H 08), (P 11); plus COL: P & P (2011) and SUR: (P 16). Addl. Data. NA, Tena: 1 ♂, 1 ♀, Río Chinchipino (- 1.0728, - 77.5864, 385 m), 13 xi 1997, leg. & det. TWD [SWD]; 2 ♂♂, 1 ♀, same data but leg. & det. JJD [JJD]; 1 ♂, same data but leg. & det. KJT [KJT]; 1 ♂, same data but 10 x 2009 [KJT]; 1 ♂, same data but leg. & det. JTJ [JTJ]; 1 ♂, 2 ♀♀, Jatun Yacu, Río Napo watershed (- 1.0167, - 77.8333, 700 m), xii 1934, leg. WCM, det. CHK [UMMZ]; 1 ♀, Río Jatunyacu, Ilupulin Creek (vicinity of Napo) (- 1.0167, - 77.8333, 700 m), i 1935, leg. W. Clarke-Macintyre, det. CHK [UMMZ]; OR, Francisco de Orellana: 1 ♀, S side of Río Napo, across from La Selva, streams in hill forest, 13 x 1988, leg. & det. SWD [SWD]; 1 ♂, 3 ♀♀, P. N. Y., 1.8 km W of Waorani Reserve Road on Markus Oil Co Road PC 29 - 30, small stream (- 0.6861, - 76.4726, 200 m), 16 xi 1997, leg. & det. WFM [FSCA, QZAC, USNM]; 1 ♂, P. N. Y., stream, (- 0.7003, - 76.4764, 200 m), 16 xi 1997, leg. & det. WFM [SWD]; 1 ♂, small streams and forest trails 12 km E of Loreto, 2 + km S of Hwy. 45 (- 0.6217, - 77.2600, 365 m), 12 x 2009, leg. & det JTJ [JTJ]; 1 ♀, T. B. S., USFQ, stream crossing Matapalo trail at marker 450 [in tandem with ♂, Perissolestes romulus] (- 0.6369, - 76.1466, 245 m), 14 i 2009, leg. & det. RWG & NVE [NVE]; 1 ♀, Reserva Etnica Waorani, 1 km S Onkone Gare Camp (- 0.6528, - 76.4333, 220 m), 4 x 1994, leg. T. Erwin, det.? [USNM]; 1 ♀, same data but 7 x 1994 [USNM]; 1 ♂, same data but 9 x 1994 [USNM]; 1 ♂, 1 ♀, same data but 10 x 1994 [USNM]; ZC, Nangaritza: 1 ♂, Mayaicu, Río Nangarizta (- 4.0290, - 78.6340, 900 m), 2 v 1941, leg. D. Laddey, det. L. Gloyd, from UMMZ [RWG]. Perissolestes Kennedy, 1941 Very similar to Perilestes in appearance (Fig. 51) and reclusive behavior. Male cerci are white instead of the dark col- or of Perilestes. Five of the 11 species are recorded in Ecuador. For identification, see Kennedy (1937 b, 1941 a).	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B54DFF88FF23F08E24D5F82D.taxon	materials_examined	Publ. Recs. ECU: Davies & Tobin (1984), Tsuda (1986, 1991), Paulson (1983), Steinmann (1997 a — as “ Perilestes castor ”), (H 08), (P 11), (H 14); PA: * Kennedy (1937 b — Holotype: [Santa Clara], “ Oriente, Río Napo watershed, El Partidero, Río Anzu ”, 14 xi 1935, leg. WCM, det. CHK, checked and data provided by RWG [UMMZ], Bridges (1993). Addl. Data. MS, Limón Indanza: 1 ♂, stream / waterfall, 25.3 km N of Indanza, 9 xi 1997, leg. & det. JJD [JJD]. Note: Ecuador endemic.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B54CFF89FF23F4342434FD17.taxon	materials_examined	Publ. Recs. ECU: (L 05), (H 08), (P 11), (H 14); SU: * Donnelly (1998). Also PER: (L 05), (H 08); plus BRA: (H 08), (P 11). Addl. Data. SU, Shushufindi: 1 ♂, La Selva Lodge; forest (- 0.4983, - 76.3733, 200 m), 23 xi 1997, leg. & det. TWD [SWD].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B54CFF89FF23F4C825C5FC1B.taxon	materials_examined	Publ. Recs. ECU: Paulson (1983), Davies & Tobin (1984), Tsuda (1986, 1991), Steinmann (1997 a), (H 08), (P 11), (H 14); SD: Kennedy (1941 a — Holotype ♂: as “ Pichincha ”, Santo Domingo de los Colorados, ca 15 km east, “ Hacienda La Lorena ” (- 0.3097, - 79.0389, 700 m), 28 ii 1941, leg. David B. Laddey, det. CHK 1941, data provided by RWG [UMMZ]), Bridges (1993), Garrison et al. (2003).	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B54CFF89FF23F4C825C5FC1B.taxon	description	Note: No additional records have been added for this endemic species since its description nearly 80 years ago.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B54CFF89FF23F5C42744FA7F.taxon	materials_examined	Publ. Recs. ECU: (H 08), (P 11); OR: Daigle (1996), von Ellenrieder & Garrison (2011), (H 14). Also PER: (H 08), (P 11). Addl. Data. OR, Francisco de Orellana: 1 ♂, stream, 3.6 km N of station on main road, Yasuni Preserve, 18 vii 1996, leg. & det. JJD [JJD]; 1 ♂, stream 1.8 km from main road on PC 29 - 30 road, Yasuni Preserve, 18 vii 1996, leg. & det. JJD [JJD]; 2 ♂♂, stream at Waorani road MP 2.0 cutoff, 16 xi 1997, leg. & det. JJD [JJD]; 1 ♂, stream, road to Waorani Reserve, 3.6 km S of Yasuni Sci. Res. Sta., 10 vi 1995, leg. & det. KJT [KJT]; 1 ♂, same data but 18 vii 1996 [KJT]; 2 ♂♂, stream, 5.1 km down road to Waorani Reserve, P. N. Y., 16 xi 1997, leg. & det. KJT [KJT]; 1 ♂, P. N. Y., km 46 on Waorani Reserve Road small forested stream (- 0.65, - 76.45, 244 m), 16 xi 97, leg. & det. WFM [FSCA]; 1 ♂, P. N. Y., 1.8 km W of Waorani Reserve Road on Markus Oil Co Road PC 29 - 30, small stream (- 0.6861, - 76.4726, 200 m), 16 xi 1997, leg. & det. WFM [FSCA]; 4 ♂♂, T. B. S., USFQ, stream crossing Matapalo trail at marker 450 [one ♂, in tandem with female Perilestes kahli] (- 0.6369, - 76.1458, 245 m), 14 i 2009, leg. & det. RWG & NVE [NVE, RWG, USFQ]; 3 ♂♂, same data but 15 i 2009 [NMNHL, RWG]; 1 ♂, same data but 18 i 2009 [NVE]; 4 ♂♂, La Selva, 100 km E Coca, streams in hill forest S of Napo River (- 0.54, - 76.54, 262 m), 13 x 1988, leg. & det. SWD [SWD, RWG].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B54CFF89FF23F005270CF927.taxon	discussion	Two genera are included in this Neotropical family (Dijkstra et al. 2014), both of which are cryptic and some of the most difficult to find damselflies.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B54CFFB7FF23F0FC2152FEFE.taxon	discussion	This genus was thought to be endemic to Ecuador, but recently a new species from Colombia was described by Bota-Sierra (2017 a). Adults are rather drab and difficult to detect as they perch in shady forested areas (Fig. 52). Three species are known from Ecuador, and two undescribed species have been collected (Tennessen, unpubl.; Haber, pers. comm.). The nymph is unknown. Archaepodagrion armatum Tennessen & Johnson, 2010. Publ. Recs. ECU: (H 14); ZC, Zamora: * Tennessen & Johnson (2010 — Holotype ♂: small forested stream S of Zamora (- 4.1217, - 78.9728, 1070 m), 2 iv 2008, leg. & det. KJT [FSCA]; 5 paratype ♂♂, paratype ♀, same data but leg. KJT & JTJ [DRP, FSCA, RWG]). Note: So far known only from the east side of Parque Nacional Podocarpus. Archaepodagrion bicorne Kennedy, 1939. Publ. Recs. ECU: Paulson (1983), Davies & Tobin (1984), Tsuda (1986, 1991), Steinmann (1997 a), (H 08), (P 11), (H 14); NA: * Kennedy (1939 a — Holotype: “ Oriente, on or near Río Napo, Napo ”, 12 xii 1935, leg. WCM [UMMZ]; Allotype: PA, Santa Clara: “ Partidero on the Río Anzu ”, [El Partidero, Indian village on east bank of Río Anzu, jct. of trails from Mera and Puyo northward to Napo, edge of tropical jungle], 700 – 1000 m, 8 ii 1936, leg. WCM, data provided by RWG [UMMZ]); Bridges (1993), Tennessen (2006). Addl. Data. PA, Mera: 1 ♂, tributaries of Río Anzu, 12 km N of Mera (- 1.4181, - 78.0528, 1190 m), 6 x 2009, leg. & det. JTJ [JTJ]; 1 ♂, 12 km NE small trib to Río Anzu just E of bridge near old foot bridge (- 1.4180, - 78.0527, 1190 m), 21 ix 2005, leg. & det. JJD [JJD]; 1 teneral ♂, same data but leg. & det. WFM [FSCA]; 1 ♂, 1 ♀, [Mera], W in Abitagua valley Guillermina Creek (- 1.45, - 78.15, 1310 m), 26 v 1941, leg. WCM, det. CHK [FSCA]; 1 ♂, [Mera], W at Abitagua, near Río Pastaza (- 1.45, - 78.15, 1100 m), 29 vi 1941, leg. WCM, det. CHK [FSCA]; Pastaza: 1 paratype ♂, Canelos – [a mission and jungle town on Río Bobonaza at headquarters of canoe navigation in Río Pastaza watershed) (- 1.5833, - 77.8000, 600 m), 12 xii 1938, leg. WCM, det. CHK, data provided by RWG [UMMZ]; Santa Clara: 2 paratype ♂♂, Guama Yacu, near El Partidero, Río Napo watershed (- 1.2833, - 77.9000, 800 m), 9 xi 1935; leg. WCM, det. CHK, data provided by RWG [UMMZ]. Note: Ecuador endemic found only within a 50 km radius of Mera. Archaepodagrion bilobatum Kennedy, 1946. Publ. Recs. ECU: Paulson (1983), Davies & Tobin (1984), Tsuda (1986, 1991), Steinmann (1997 a), (H 08), (P 11), (H 14); MS, Morona: * Kennedy (1946 a — Holotype: “ Santiago- Zamora, Macas, Río Upano ” (- 2.3167, - 78.1167, 1050 m), leg. L. Alonzo, data provided by RWG [UMNZ]); Bridges (1993), Garrison et al. (2003). Note: An Ecuador endemic that is currently known only from the type specimen.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B572FFB5FF23F78421E0FAEB.taxon	discussion	A large genus, with 36 described species, six of which are recorded in Ecuador with another species or two ques- tionably occurring. See Calvert (1924) and Bick & Bick (1988) for species identification, based mainly on males. The adults are drably colored (Fig. 53) and species identification relies mainly on male appendage morphology. Nymphs live in edges of very small streams and seepage runs, in forested areas. Five of the six species in Ecuador are endemic to the country. Philogenia cassandra (Hagen in Selys, 1862). Publ. Recs. ECU: Schmidt (1942), Racenis (1953, 1959), Paulson (1983), Tsuda (1986, 1991), (P 11), (H 14); PA: * Selys (1886 — Bobonaza (- 1.5833, - 77.8000, 600 m), leg.? det. Selys, 1886 data provide by RWG [RWG]), Bick & Bick (1988). Also COL, PER, VEN: (H 08), (P 11). Philogenia macuma Dunkle, 1986. Publ. Recs. ECU: Tsuda (1986, 1991), Steinmann (1997 a), Paulson (2009), (H 14); MS: * Dunkle (1986 — Holotype: [Taisha:] Macuma (867 m), 2 ii 1972, leg. D. Pearson, FSCA]), Bridges (1993), Bick & Bick (1988), NA: Bick & Bick (1988); PA: Bick & Bick (1988). Addl. Data. MS, Taisha: 2 ♂♂, Macuma, forest trail (- 2.17, - 77.65, 867 m), 2 ii 1972, leg. D. Pearson, det. SWD [FSCA, RWG]; NA, Tena: 1 ♂, Río Jatun-yacu, Ilupulin Creek (vicinity of Napo), headwater trib. Río Napo (- 1.0167, - 77.8333, 700 m), i 1935, leg. WCM, det. RWG, from UMMZ (RWG); 1 #, 3 ♀♀, Jatun Yacu, Río Napo watershed (- 1.0167, - 77.8333, 700 m), iii 1937, leg. WCM, det. RWG, from C. Kennedy / UMMZ [RWG]; 1 ♂, 1 ♀, J. S. B. S., camiño hacia o Napo (- 1.0638, - 77.6201, 451 m), 8 xi 2014, leg. & det. ACRP [ACRP]; PA, Pastaza: 3 ♂♂, stream 2 km S of Comunidad Chuwitayo road between Macas and Puyo (- 1.9000, - 77.8033, 800 m), 9 xi 1997, leg. & det. TWD [JJD, SWD]; Santa Clara: 1 ♂, Río Arajuno, Dayuma Camp (- 1.1000, - 77.5833, 434 m), 22 ix 1990, leg. B. Clark, det. RWG [RWG]. Note: Ecuador endemic. Philogenia mangosisa Bick & Bick, 1988. Publ. Recs. ECU: Tsuda (1991), Steinmann (1997 a), (H 08), (P 11), (H 14); MS: * Bick & Bick (1988 — Holotype: [Morano:] (- 2.5244, - 77.8947, 850 m), 30 xi 1945, leg. L. Alonzo, UMMZ]), Bridges (1993), Tennessen (1998 — as P. raphaella Selys, 1886), Garrison et al. (2003), Tennessen (2006); ZA: Bybee & Tennessen (2008). Addl. Data. MS, Limón Indanza: 4 ♂♂, stream, Hwy. 45, 25.3 km N of Indanza (- 2.9217, - 78.4031, 1110 m), 7 xi 1997, leg. & det. KJT [KJT]; 1 ♂, same data but 5 xi 1997, leg. & det. JJD [JJD]; 1 ♂, same data but 9 ix 1997, leg. JJD, det. WFM [SWD]; Huamboya: 5 ♂♂, 2 ♀♀, Río Pasanac waterfall, 57 km N of Macas, Hwy. 45, 16 viii 2005, leg. & det. JJD [JJD]; 1 ♂, small stream, near Río Pasanac, 80 km N of Sucua, 9 xi 2007, leg. & det. KJT [KJT]; Morona: 1 Paratype ♂, Salado, Río Upano (- 2.2500 - 78.2667, 2000 m), date?, leg. L. Alonzo, det. RWG, from UMMZ, [RWG]; Santiago: 1 ♂, Yuganza [just S of Yuganza], 25.3 km N of Plan de Milagro Jct [trib of Río Yuganza], small stream with mossy rocks (- 2.8683, - 78.3739, 580 m), 7 xi 1997, leg. & det. WFM [FSCA]; ZC, El Pangui: 1 ♂, 1 ♀, stream about 15 km S of El Pangui, 6 xi 1997, leg. & det. KJT [KJT]; 1 ♂, [Pincho] [about 3 km N on Hwy. 45 or], 15 km S of El Pangui, tanic rocky stream (- 3.7242, - 78.5964, 914 m), 6 xi 1997, leg. & det. WFM [FSCA]; Zamora: 1 ♂, Zamora (- 4.0692, - 78.9567, 1000 m), 2 xi 1941, leg. D. Laddey, det. RWG, from UMMZ [RWG]; 1 ♀, same data but 9 xi 1941 [RWG]. Note: Ecuador endemic, east slope. Philogenia minteri Dunkle, 1986. Publ. Recs. ECU: Tsuda (1986, 1991), Steinmann (1997 a), (H 08), (P 11), (H 14); NA: Bick & Bick (1988); OR: Daigle (1996); PA: Tennessen (2006); SU: * Dunkle (1986 — Holotype: “ Limoncocha, NA ” [Shushufindi:], (- 0.4000, - 76.6167, 269 m), 19 ii 1972, leg. D. Pearson [FSCA]; Allotype, same data but 24 i 1972 [FSCA]), Bick & Bick (1988), Bridges (1993), Donnelly (1998). Also COL: Bota-Sierra et al. (2018). Addl. Data. NA, Carlos Julio Arosemena Tola: 1 ♀, stream, Puyo Road, 19.3 km S of Río Napo, 21 vii 1996, leg. KJT, det. JJD [FSCA]; 2 ♂♂; Loreto: 1 ♂, 1 ♀, rocky river seeps, 34.2 km W of Loreto on Loreto Road, 19 xi 1997, leg. & det. JJD [JJD]; 1 ♂, forest seep, about 12 km NE of Loreto (- 0.6217, - 77.2600, 365 m), 12 x 2009, leg. & det. KJT [KJT]; 1 ♂, small streams and forest trails 12 km E of Loreto, 2 + km S of Hwy. 45 (- 0.6217, - 77.2600, 365 m), 15 x 2009, leg. & det. JTJ [JTJ]; 1 ♀, same data but leg. KJT, det. JJD [FSCA]; 1 ♂, river along Loreto-Ar- chidona road (- 0.7417, - 77.5517, 900 m), 19 xi 1997, leg. & det. TWD [SWD]; PA, Mera: 1 ♀, between Mera and Shell, Los Copales, trails and forest N of Río Metolo (- 1.4886, - 78.0708, 1020 m), 20 ix 2005, leg. KJT, det. JJD [FSCA]; SU, Shushufindi: 1 paratype ♂, 1 paratype ♀, Limoncocha (- 0.4000, - 76.6167, 269 m), 23 viii 1980, leg. & det. SWD [SWD]; 1 paratype ♂, same data but 28 viii 1980, leg. & det. SWD [RWG]; 1 ♀, same data but 16 viii 1972, leg. E. Stiles, det. DRP [DRP]; 1 ♂, same data but lake edge, 5 vii 1991, leg. D. Pearson, det. DRP [DRP]; 1 ♂, 1 ♀, same data but forest trail, 19 ii 1972, leg. D. Pearson, det. DRP [DRP, FSCA]; 1 ♂, same data but 1 iii 1972 [DRP]; 1 ♀, same data but 24 i 1972 [FSCA]; 1 ♂, same data but forest stream, 23 vii 1977, leg. & det. DRP [DRP]; 1 ♂, 1 ♀, same data but Río Napo, 18 xi 1980, leg. & det. MWJ [FSCA]; 1 ♂, same data but stream by nature trail, 24 viii 1980, leg. KWK, det. KJT [KJT]; 1 ♂, stream about 3 km SW of Shushufindi (- 0.2136, - 76.6786), 12 v 2009, leg. & det. KJT [KJT]; 1 ♂, trails, small streams 3 km W of Shushufindi, 2 + km N off Hwy. 26 at km marker 17 (- 0.1664, - 76.6917, 290 m), 14 x 2009, leg. & det. JTJ [JTJ]; 1 ♂, La Selva Lodge; forest (- 0.4983, - 76.3733, 200 m), 22 xi 1997, leg. & det. TWD [SWD]; 2 ♂♂, same data but 24 xi 1997 [SWD]. Philogenia monotis (Kennedy, 1941). Publ. Recs. ECU: Paulson (1983), Davies & Tobin (1984), Tsuda (1986, 1991 — both as Agnophilogenia monotis and P. tinalandia), Steinmann (1997 a — as A. monotis and P. tinalandia), (H 08 — as P. tinalandia), (P 11 — also as P. tinalandia), (H 14 — also as P. tinalandia); SD: * Kennedy (1941 b — Holo-type ♀: “ Pichincha ”, ♂, Santo Domingo de los Colorados, ca 15 km east, “ Hacienda La Lorena ” (- 0.3097, - 79.0389, 700 m), date?, leg. D. Laddey [UMMZ]), * Bick & Bick (1988 — as P. tinalandia Holotype & Allotype: Santo Domingo de Los Colorados, 12 km E on Hwy. 30 near Hotel Tinalandia (- 0.2839, - 79.05, 760 m), 13 v 1985, leg. J. Nation [FSCA]), Bridges (1993 — as both P. monotis and P. tinalandia), Garrison et al. (2003), von Ellenrieder (2003 b — sunk generic name Agnophilogenia and suggested P. tinalandia was a junior synonym); P. tinalandia was synonymized with P. monotis by Bota-Sierra (2018). Also COL: Bota-Sierra et al. (2018). Addl. Data. SD: 2 ♂♂, Santo Domingo de Los Colorados, 12 km E on Hwy. 30 near Hotel Tinalandia (- 0.2839, - 79.0500, 760 m), 13 v 1985, leg. J. Nation, det. Bick & Bick [FSCA, RWG]; 1 ♂, same data but leg. & det. SWD (as P. tinalandia) [SWD]; 1 ♀, small stream, Hwy. 30, km marker 61, E of La Union del Toachi (- 0.5003, - 78.8700, 1210 m), 25 ii 1010, leg. & det. JJD, [JJD]; 2 ♂♂, same data but 27 ii 2010, leg. & det. KJT [FSCA]; 1 ♀, same data but leg. & det. JTJ; 2 ♂♂, 1 ♀, small stream, Hwy. 30, near km marker 71, near La Unión del Toachi (- 0.3139, - 78.9436, 900 m), 27 ii 2010, leg. & det. JTJ [JTJ]. Philogenia redunca Cook, 1989. Publ. Recs. ECU: Tsuda (1991), Paulson (2009), (H 14); NA: * Cook (1989), Bridges (1993); OR: Daigle (1996), von Ellenrieder & Garrison (2011); PA * Cook (1989 — Holotype: “ Yanamanaca, Napo ” [Santa Clara], Yanamanaca [next Indian village up Río Anzu from Yanamanaca is El Partidero; coordinates for El Partidero] (- 1.2833, - 77.9000), 6 x 1935, leg. WCM, data provided by RWG [UMMZ]; Allotype: [Mera], Abitagua, (- 1.45, - 78.15, 200 m), iv 1937, leg. WCM, data provided by RWG [UMMZ]); Garrison et al. (2003), Tennessen (2006); TU: * Cook (1989). Note: This is “ Philogenia sp. A ” reported by Bick & Bick (1988) from MS and NA. Addl. Data. MS, Morona: [FAL, [RWG]; NA, Archidona: [KJT, USNM]; Carlos Julio Arsemena Tola: [RWG]; Quijos: [RWG]; Tena: [RWG, SWD]; OR: Francisco de Orellana: [KJT]; Loreto: [JTJ, KJT, RWG]; PA, Mera: [FSCA, KJT, RWG]; Pastaza: [FSCA, JJD, KJT, SWD]; Santa Clara: [RWG]; SU, Gonzalo Pizarro: [SWD]; Shushufindi: [KJT]; TU, Baños: [FSCA, RWG]. Note: This endemic species is the most abundant and widespread east slope representative of the genus. One SWD specimen from 1980 was labeled P. berenice. Philogenia unresolved. There are several specimens that have not been determined, some of which represent undescribed species: a. Philogenia cf. minteri. Undescribed according to Garrison (pers. comm., 19 viii 2011): OR, Francisco de Orellana: 1 ♂, stream, 3.6 km from Sci. Res. Sta., P. N. Y., 18 vii 1996, leg. & det. KJT [KJT]; T. B. S., USFQ, stream crossing Chorongo trail (- 0.6378, - 76.1486, 224 m), 11 i 2009 leg. & det. RWG & NVE [NVE, RWG]; 1 ♂, same data but along Chichico trail (- 0.6300, - 76.1525, 250 m), 12 i 2009, leg. & det. RWG & NVE [RWG]; 2 ♂♂, same data but along Chorongo trail (- 0.6350, - 76.1500, 232 m), 16 i 3009 [NVE, RWG]; 1 ♂, same data but Matapalo trail just S to Harpia trail (- 0.6358, - 76.1503, 237 m), 10 i 2009 [USFQ]; 1 ♂, same data but stream crossing Matapalo trail at marker 450 (- 0.6369, - 76.1458, 245 m), 14 i 2009 [NVE]; 1 ♀, Numa trail at station (- 0.6383, - 76.1492, 220 m), 18 i 2009 [RWG]; 1 ♀, stream crossing Harpia trail (- 0.6256, - 76.1494, 238 m), 11 i 2009 [RWG]; 1 ♂, black water stream crossing Guacamayo trail at marker 2050 (- 0.6383, - 76.1492, 220 m), 16 i 2009 [USFQ; Loreto: 1 ♂, trib. of stream, 34.2 km W of Loreto, 19 xi 1997, leg. & det. KJT [KJT]. b. Philogenia cf. peruviana Bick & Bick, 1988. Also PER: (H 08), (P 11). MS, Santiago; 1 ♂, pond on side road, between Sucua and Mendez (- 2.6683, - 78.2483, 600 m), 6 xi 1997, leg. & det. TWD [SWD].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B570FFB5FF23F3B92689F9EF.taxon	discussion	Platystictidae contains one species-rich genus in the New World. Adults remain perched with wings usually together in deep shade for most of the daylight hours, their flight activity apparently limited to very early morning. They require small forested streams in which to complete their life cycle.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B570FFB0FF23F0B42466F897.taxon	discussion	Forty-three species have been described in this strictly New World genus, with six known for Ecuador. However, the genus is in need of revision, and there are potentially undescribed species in Ecuador. The few references by which some species identifications can be made are Calvert (1931) and Kennedy (1938 a, 1942 a). Adults are mediumsized, mostly dark with small blue stripes and spots on the thorax; the abdomen varies from nearly all black to blue on the apical segments (Figs. 54 a & 54 b). Little work has been done on the biology of this genus, though all species have been found in the vicinity of small shaded streams and seepage areas. Palaemnema abbreviata Kennedy, 1938. Publ. Recs. ECU: Paulson (1983), Davies & Tobin (1984). Tsuda (1991), Steinmann (1997 a), (H 08), (P 11), (H 14); NA: * Kennedy (1938 a — Holotype & Allotype: Río Jatun Yacu, “ Prov. Oriente ” [UMMZ]); Garrison et al. (2003), PA: * Kennedy (1938 a), TU: * Kennedy, (1938 a). Addl. Data. MS, Huamboya: 9 ♂♂, 1 ♀, stream, 10 km S. of Comunidad Chuwitayo (- 1.9500, - 77. 8583, 800 m), 9 xi 1997, leg. & det. TWD [SWD]; 2 ♂♂, same data but leg. & det. JJD [JJD]; 1 ♂, same data but leg. TWD, det. KJT [KJT]; 7 ♂♂, 1 ♀, same data but 10 xi 1997, leg. & det. KJT [KJT, RWG]; 2 ♂♂, 1 ♀, same data but 10 xi 1997, leg. & det. TWD [SWD]; NA, Tena: 1 ♂, [Puerto Napo] [upstream near jct with Río Mulatos] Jatun Yacu Río Napo watershed (- 1.05, - 77.98, 700 m), iii 1937, leg. W. Clark-Macintyre, det. CHK [FSCA]; 1 ♂, Puerto Ahuano, Misicocha (389 m), 16 xi 2010, leg. & det. ACRP (pers. comm. ACRP); OR: Francisco de Orellana: 2 ♂♂, across Río Napo from La Selva, streams in hill forest, 13 x 1988, leg. & det. SWD [SWD]; Loreto: 2 ♂♂, small streams and forest trails, 12 km E of Loreto, 2 + km S of Hwy. 45 (- 0.6217, - 77.2600, 365 m), 12 x 2009, leg. & det. JTJ [JTJ]; 1 ♂, same data but 15 x 2009, leg. & det. JTJ. [JTJ]; PA, Pastaza: 1 ♂, Río Sandalias, 5 km S of Puyo (- 1.5003, - 77.9522, 1000 m), 11 xi 1997, leg. & det. TWD [SWD]; 1 ♂, 1 ♀, stream 2 km S. of Comunidad Chuwitayo road between Macas and Puyo (- 1.9000, - 77.8033, 800 m), 9 xi 1997, leg. & det. TWD [FSCA]; 1 ♂, Cuasha just N of Río Pastaza bridge (90.5 km N of Sucua), small forested stream (- 1.88, - 77.80, 915 m), 9 xi 1997, leg. & det. WFM [FSCA]; SU, Shushufindi: 1 ♂, small forest stream, about 4 or 5 km WNW of Shushufindi (- 0.1692, - 76.6889, 270 m), 14 v 2009, leg. & det. KJT [RWG]. Note: Ecuador endemic. Palaemnema brucelli Kennedy, 1938. Publ. Recs. ECU: Paulson (1983), Davies & Tobin (1984), Tsuda (1986, 1991), Steinmann (1997 a), (H 08), (P 11), (H 14); NA: * Kennedy (1938 a — Holotype: “ Oriente: Río Anzu ” [UMMZ]), Bridges (1993), Garrison et al. (2003), PA: * Kennedy (1938 a), TU: * Kennedy (1938 a, Allotype- UMMZ]. Also COL: Palacino et al. (2017). Addl: Data. NA, Archidona: 11 ♂♂, 4 ♀♀, Río Chontayacu, new Coca Road, 5 km E of junction with Baeza- Tena Road, 11 xi 1990, leg. OSF, det. SWD [SWD, USNM]; 2 ♂♂, Puerto Ahuano, Misicocha (389 m), 16 xi 2010, leg. & det. ACRP (pers. comm., ACRP); Carlos Julio Arsemena Tola: 1 ♀, Ila S at Hacienda Ila Río Anzu (- 1.12, - 77.50, 700 m), 18 xii 1936, leg. WCM, det. CHK [FSCA]; PA, Mera: 1 ♀, [Mera] W at Abitagua Río Pastaza (- 1.45, - 78.15, 1100 m), 29 x 1936, leg. W. Clark-Macintyre, det. CHK [FSCA]. Palaemnema clementia (Selys, 1886). Publ. Recs. ECU: Paulson (1983), Tsuda (1986, 1991), (H 08), (P 11), (H 14); GY: * Kennedy (1938 a), NA: * Kennedy (1938 a). Also COL, PER, VEN: (H 08), (P 11); plus BOL. Addl. Data. BO, Caluma: 3 ♂♂, rocky stream 5 km E of Caluma, Guaranda Road (- 1.5942, - 79.2308, 480 m), 5 iii 2010, leg. & det. JJD [JJD]; 1 ♂, 1 ♀, same data but leg. & det. KJT [FSCA]; 5 ♂♂, same data but leg. & det. JTJ [JTJ]; LR, Buena Fé: 1 ♂, [Consumulo] S Río Palenque (- 0.58, - 79.35, 175 m), 13 vi 1974, leg. J. Longino, det. GHB [FSCA]; MN, Pedernales: 4 ♂♂, 2 ♀♀, Río Nalpe, about 3 km E of El Achote (0.0547, - 79.9739, 130 m), 28 ii 2010, leg. & det. KJT [FSCA]; 1 ♂, 1 ♀, same data but leg. & det. JTJ [JTJ]; Portoviejo: 1 ♂, small rocky stream E of Portoviejo, Hwy. E 40 (- 1.0750, - 80.1764, 220 m), 4 iii 2010, leg. & det. KJT [FSCA]; 1 ♂, same data but leg. & det. JTJ [JTJ]; NA, Tena: 4 ♂♂, Puerto Ahuano, Ahuano (362 m), 16 xi 2010, leg. & det. ACRP (pers. comm., ACRP); SD: 1 ♂, 1 ♀, near Recinto Baba Otongo, about 10 km SE of Santo Domingo de los Colorados (- 0.3650, - 79.2072), 24 ii 2010, leg. & det. KJT [FSCA]. Palaemnema lorena Kennedy, 1942. Publ. Recs. ECU: Paulson (1983), Davies & Tobin (1984), Tsuda (1986, 1991), Steinmann (1997 a), (H 08), (P 11), (H 14); SD: Kennedy (1942 a — Holotype & Allotype, plus Paratype ♀: as “ Pichincha ”, Santo Domingo de los Colorados, ca 15 km east, “ Hacienda La Lorena ” (- 0.3097, - 79.0389, 700 m), 21 ii 1941, leg. David Laddey, det. CHK, checked and data provided by RWG [UMMZ]), Bridges (1993), Garrison et al. (2003). Addl. Data. ES, Quininde: 1 ♂, 1 ♀, Río Canande Reserve, 35 km NE of Quininde, tributary of Río Hoja Blanca, swampy trickle in forest, (0.5264, - 79.2122, 345 m), date not provided, data provided by WAH & DLW. Palaemnema melanocauda Kennedy, 1942. Publ. Recs. ECU: Paulson (1983), Davies & Tobin (1984), Tsuda (1986, 1991), Steinmann (1997 a), (H 08), (P 11), (H 14). SD: * Kennedy (1942 a — Holotype ♀: as “ PC ”, Santo Domingo de los Colorados (500 m), 4 i 1941, leg. D. Laddey [UMMZ]), Bridges (1993), Garrison et al. (2003). Note: No other data available; Ecuador endemic. Palaemnema picicaudata Kennedy, 1938. Publ. Recs. ECU: Paulson (1983), Davies & Tobin (1984), Tsuda (1986, 1991), Steinmann (1997 a), (H 08), (P 11), (H 14); NA: * Kennedy (1938 a — Holotype ♂: [Tena:], near Napo (- 1.0500, - 77.7833, 400 m), 24 ii 1934, leg. WCM, det. CHK, data provided by RWG [UMMZ]; Allotype ♀: Río Jatun Yacu near Hacienda Ila (- 1.2000, - 77.8333, 700 m), 24 xii 1936, leg. WCM, det. CHK, data provided by RWG [UMMZ]), Bridges (1993), Tennessen (1996 b), Garrison et al. (2003). Also COL: Bota-Sierra et al. (2018). Addl. Data. MS, Limón Indanza: 2 ♂♂, 1 ♀, stream / waterfall 25.3 km N of Indanza (- 2.9217, - 78.4031, 1110 m), 9 xi 1997, leg. & det. JJD [JJD]; NA, Archidona: 1 ♂, stream 47.4 km W of Loreto (- 0.7422, - 77.5489, 1100 m), 14 vi 1995, leg. & det. KJT [FSCA]; 2 ♀♀, Guamani, about 5 km W on Rt E 20, small stream, (- 0.7161, - 77.6645), 25 x 2013, leg. & det. KJT [FSCA]; Tena: 1 ♀, [Puerto Napo, upstream near jct with Río Mulatos], Jatun Yacu Río Napo watershed (- 1.05, - 77.98, 700 m), iii 1937, leg. WCM, det. CHK [FSCA]; Paratype series: Río Jatun Yacu, upper Ilapulin near small streams (- 1.0167, - 77.8333, 700 m), iii 1937, leg. WCM, det. CHK, data provided by RWG [UMMZ]; 2 ♂♂, J. S. B. S., rego comedor (- 1.0665, - 77.6165, 430 m), 6 xi 2014, leg. & det. ACRP [ACRP]; OR, Loreto: 1 ♂, Concepcion, Río Napo (- 1.6667, - 77.4167, 400 m), 28 xii 1939, leg. WCM, det. CHK [FSCA]; 2 ♂♂, stream 8.5 km N of Loreto (- 0.6183, - 77.2950, 400 m), 14 xi 1997, leg. & det. TWD [SWD]; PA, Santa Clara: Paratype series: El Partidero, Río Napo watershed (- 1.2833, - 77.9000, 800 m), 9 xi 1935, leg. WCM, det. CHK, data provided by RWG [UMMZ]; 1 ♂, small stream, 2 km S of Comunidad Chuwitayo (- 1.9000, - 77.8033, 800 m), 9 xi 1997, leg. & det. KJT [FSCA]; 4 ♂♂, 3 ♀♀, Río Sandalias, 5 km S of Puyo (- 1.5003, - 77.8653, 1000 m), 11 xi 1997, leg. & det. TWD [SWD]; SU, Shushufindi: 1 ♂, La Selva Lodge, forest (- 0.4983, - 76.3733, 200 m), 23 xi 1997, leg. & det. TWD [KJT]. Palaemnema unresolved: There are several specimens that have not been determined and may represent undescribed species, all labeled as sp.: MS, Huamboya: 4 ♂♂, small stream, near Río Pasanac, 80 km N of Sucua, 9 xi 1997, leg. & det. KJT [KJT]; OR, Loreto: 1 ♀, small streams and forest trails 12 km E of Loreto, 2 + km S of Hwy. 45 (- 0.6217, - 77.2600, 365 m), 15 x 2009, leg. and det. JTJ [JTJ]; PA, Mera: 3 ♂♂, Río Metolo / trails, Los Copales, between Mera and Shell (- 2.4917, - 78.0719, 1070 m), 7 x 2009, leg. & det. JTJ [JTJ]; SD: 1 ♂, pond / small stream, Hwy. 30, near km marker 72, near La Unión del Toachi (- 0.3133, - 78.9472, 900 m), 26 ii 2010, leg. & det. JTJ [JTJ]; 1 ♀, same data but Hwy. 30, near km marker 61, E of La Unión del Toachi (- 0.3336, - 78.8700, 1210 m), 25 ii 2010, leg. & det. JTJ [JTJ]; SU, Shushufindi: 1 ♂, 1 ♀, trails, small streams 3 km W of Shushufindi, 2 + km N off Hwy. 26 at km marker 17 (- 0.1664, - 76.6917, 290 m), 14 x 2009, leg. & det. JTJ (JTJ); 2 ♂♂, same but, 13 x 2009, leg. & det. JTJ [JTJ]. Also Haber (2014) lists Palaemnema sp. “ A ”, “ B ”, ‘ C’, and “ D ” of which three appear to be undescribed.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B575FFB1FF23F16D2424FEFF.taxon	discussion	Polythorids are mostly large damselflies with stocky thorax and wide wings that are held together when perched. Many species have beautiful wing patterns of dark and white or yellow to yellow-orange. This Neotropical family requires small, mostly shaded, moderately to fast-flowing streams with riparian woody vegetation. Five genera and 22 described species have been recorded in Ecuador; at least two new species await description. The nymphs of very few species have been discovered. Consult Garrison et al. (2010) for keys to genera and references to nymphal descriptions.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B574FFBFFF23F7852473FCAA.taxon	discussion	Large to mid-sized wary damselflies with clear wings (some species have brilliant blue reflective patches in the wings) and either a mostly bright blue and black striped thorax or an olive-drab and dark brown striped thorax (Fig. 55). Of the 22 known species, eight occur in Ecuador, most of which are very poorly known — further surveys for these rare damselflies is highly recommended. There is potential for at least one new species. A key for species identification was provided by Bick & Bick (1990 a). Habitat is mostly small, heavily to partly shaded streams. Cora confusa Kennedy, 1940. Publ. Recs. ECU: Paulson (1983), Davies & Tobin (1984), Tsuda (1986, 1991), Bick & Bick (1990 a), Steinmann (1997 a), (H 08), (P 11), (H 14); PA: * Kennedy (1940 — Holotype & Allotype: “ upper Río Pastaza, Oriente ” [Mera:] Abitagua (- 1.45, - 78.15, 1200 m), 1 x 1939, leg. WCM, det. CHK, data provided by RWG [UMMZ]), Montgomery (1967), Bridges (1993), Garrison et al. (2003). Also COL: (P 16). Addl. Data. PA, Mera: 1 ♂, [Mera] W at Abitagua [GPS & elevation from Brown (1941)], (- 1.45, - 78.15, 1300 m), xii 1948, leg. WCM, det. CHK [FSCA]. Note: Abitagua is the only known location in Ecuador; no specimens have been collected since 1948. Cora dorado Bick & Bick, 1991. Publ. Recs. ECU: (H 08), (P 11), (H 14); TU: * Bick & Bick (1991 — Holotype: Yungula, where Río Blanco flows into Río Pastaza (- 1.24, - 78.20, 1700 – 1900 m), 19 ix 1936, leg. WCM [UMMZ]; same data but Allotype: 15 ix 1937 [UMMZ]), Bridges (1993), Garrison et al. (2003). Note: Ecuador endemic; known only from the type locality. Cora dualis McLachlan, 1878. Publ. Recs. ECU: Kirby (1890), Davies & Tobin (1984), Tsuda (1986, 1991), Paulson (1983), Steinmann (1997 a), (H 08), (P 11), (H 14); IM: * McLachlan (1878 — Type series 3 ♂♂, 2 ♀♀: [Cotacachi], “ Intag ” Apuela, NE on Apuela-Cuicocha Road, along Río Intag and tributaries between 1800 – 2800 m (0.3588, - 78.4509 at jct of Río Intag and tributary), data provided by RWG [BMNH]), Selys (1879), Campos (1922), Montgomery (1967), Kimmins (1970 — designated the Lectotype [BMNH]), Bridges (1993), Paulson (2011). Note: Ecuador endemic; all references apparently refer to the original type series (Bick & Bick, 1990 a). We did not find this species during a recent search in the Intag area (Mauffray & Tennessen 2018). Cora inca Selys, 1854. Publ. Recs. ECU: Davies & Tobin (1984), Tsuda (1986, 1991), De Marmels (1999), Paulson (1983), (H 08), (P 11), (H 14); NA: Tennessen (1998); PC: * Selys (1873 a — types, “ Quito ” [BMNH, IRSN]), Hagen (1877 — as “ probably C. brasilinensis ”), McLachlan (1878), Kirby (1890), Campos (1922), Montgomery (1967), Bridges (1993); MS: Tennessen (2006); PA: Bick & Bick (1990 a), (L 05), Tennessen & Johnson (2008). Also BRA, COL, VEN: (L 05), (H 08), (P 11). Addl. Data. CT, Sigchos: 1 ♂, San Francisco de las Pampas Otongo Reserve, Río Las Damas Epoca Illuviosa, Bosque virgen (- 0.45, - 79.0, 1900 m), 10 ix 1997, leg. E. Tapia, det. WFM [QZAC]; IM, Cotacachi: 6 ♂♂, Santa Rosa, La Florida Reserve, Cock of the Rock Trail, Quebrada El Placer (0.3603, - 78.4791, 1862 m), 25 viii 2018, leg. & det. WFM [FSCA, USFQ]; same data but, 5 ♂♂, leg. & det. KJT [KJT]; same data but, 5 ♂♂, 2 ♀♀, 30 viii 2018 [FSCA, USFQ]; same data but 3 ♂♂, leg. & det. KJT [KJT]; same data but, 2 ♂♂, 2 ♀♀, small stream, trib. of El Placer (0.3606, - 78.4788, 1863 m), leg & det. KJT [KJT]; same data but, 3 ♂♂, 1 ♀, La Florida Reserve, lower public trail, below cascada of Quebrada El Placer (0.3653, - 78.4784, 1858 m), leg. & det. KJT [KJT]; MS, Huamboya: 1 ♂, small stream with waterfall, 0.2 km S of Río Pasanac, Hwy. 45 [about 57 km N of Macas] (- 1.9532, - 77.8519), 16 ix 2005, leg. & det. KJT [KJT]; 3 ♂♂, rocky stream about 2.4 km W of loc. # 1 [12 km W on Macas-Guamote Road], flowing over road (- 2.2342, - 78.2181), 18 ix 2005, leg. & det. KJT [KJT]; 1 ♂, Macas, 22.9 km W on Gua- mote Road, 3.3 km W on Río Albanico, rocky stream and flowing roadside ditches (- 2.2342, - 78.2181, 1690 m), 18 ix 2005, leg. & det. WFM [FSCA]; NA, Quijos: 15 ♂♂, 1 ♀, Baeza 4.8 – 5.0 km S, small clear streams and seeps (- 0.4761, - 77.8711, 1830 m), 20 xi 1997, leg. & det. WFM [FSCA]; 3 ♂♂, 2 ♀♀, same data but leg. & det. JJD [JJD, KJT]; 1 ♂, 2 ♀♀, same data but leg. & det. KJT [KJT]; 3 ♂♂, same data but leg. & det. TWD [SWD]; PA, Mera: 2 ♂♂, [Mera] W at Abitagua (- 1.45, - 78.15, 1200 m), 11 x 1937, leg. WCM, det. GHB [FSCA]; ZC, Zamora: 1 ♂, stream / waterfall, W of Zamora (- 3.9628, - 79.0508, 1700 m), 1 iv 2008, leg. & det. KJT [FSCA]: 1 ♂, same data but leg. & det. JTJ [JTJ]. Cora jocosa McLachlan, 1881. Publ. Recs. ECU: Kirby (1880), Campos (1922), Kimmons (1970), Paulson (1983), Davies & Torbin (1984), Tsuda (1986, 1991), Paulson (1983 — all as Josocora jocosa), Paulson (1983), Steinmann (1997 a), (H 08), (P 11), (H 14); NA: Tennessen (1998); PA: * McLachlan (1881 a — type locality: ” Río Bobonaza ” [BMNH]), Kennedy (1940 — as J. jocosa), Montgomery (1967), Bick & Bick (1990 a); SU: Bick & Bick (1990 a); TU: Bick & Bick (1990 a), Bridges (1993). Also PER: (H 08), (P 11). Addl. Data. CB, Alausí: 1 ♂, Huigra, Río Huigra 1 km upstream from trail (- 2.2783, - 78.9731, 1350 m), 1 iv 1941, leg. WCM, det. GHB, checked by WFM [FSCA]; OR, Loreto: 4 ♂♂, rocky stream, 8.5 km E of Loreto, Loreto Road (- 0.6183, - 77.2950, 400 m), 14 xi 1997, leg. & det. JJD [JJD]; 2 ♂♂, same data but leg. & det. KJT [KJT]; 2 ♂♂, same data but leg. & det. TWD [SWD]; PA, Pastaza: 1 ♀, Puyo (- 1.4667, - 77.9833, 1000 m), 2 iii 1950, leg. WCM, det. GHB, data provided by RWG [UMMZ]; 1 ♂, Puyo 5 km S on Hwy. 45, Río Sandalias and tribs. (- 1.5003, - 77.9522, 1000 m), 11 xi 1997, leg. & det. WFM [FSCA]; SU, Cascales: 3 ♂♂, 1 ♀, Lago Agrío 56 km W, rocky stream (- 0.0067, - 77.2986, 500 m), 20 viii 1980, leg. & det. SWD [DRP, RWG, SWD]. Cora klenei Karsch, 1891. Publ. Recs. ECU: * Karsch (1891 b), Campos (1922 — “ indicated by Calvert ”), Kennedy (1940 — as Josocora klenei), Montgomery (1967 — as J. klenei), Paulson (1983 — as J. klenei), Davies & Tobin (1984 — as J. klenei), Tsuda (1986, 1991 — both as J. klenei), Bick & Bick (1990 a), Steinmann (1997 a), (H 08), (P 11), (H 14). Also COL: (H 08), (P 11). Note: Karsch gave no specific locality for the Ecuador specimen; the type is lost according to Bridges (1993) and apparently there have been no other collections from Ecuador reported. Cora munda McLachlan, 1878. Publ. Recs. ECU: Kirby (1890), Davies & Tobin (1984), Tsuda (1986, 1991), Paulson (1983), Steinmann (1997 a), (H 08), (P 11), (H 14); IM: * McLachlan (1878 — type series, 6 ♂♂, 3 ♀♀ [Co- tacachi]; “ Intag ” [Apuela, NE on Apuela-Cuicocha Road, along Río Intag and tributaries between 1800 – 2800 m, 0.3588, - 78.4509 at jct of Río Intag and tributary], 1 x 1839, leg. C. Buckley, data provided by RWG [BMNH]), Selys (1879), Kennedy (1919) Campos (1922), Montgomery (1967 — designated the Lectotype ♀ [BMNH]), Kim- mins (1970), Bick & Bick (1990 a), Bridges (1993); EO: Bick & Bick (1990 a); LJ: Tennessen & Johnson (2008). Addl. Data: EO, Piñas: 2 ♂♂, Piñas (- 3.6667, - 79.65, 1000 – 1500 m), 26 vii 1941, leg. D. Laddey, det. Bick & Bick [FSCA]; LJ, Catamayo: 4 ♂♂, 2 ♀♀, stream, 25 km from Hwy. 50 on road to Zaruma (- 3.8819, - 79.5558, 1030 m), 7 iv 2008, leg. & det. KJT [FSCA, KJT]; 1 ♂, same data but leg. & det. JTJ [JTJ]. Note: Ecuador endemic. Cora terminalis McLachlan, 1878. Publ. Recs. ECU: * Campos (1922 — “ by Dr. Martin ”), Schmidt (1942), Bick & Bick (1990 a), Tsuda (1991), (H 08), (P 11), (H 14). Also COL, PER: (H 08), (P 11); plus BOL: (H 08), (P 16). Addl. Data. MS, Morona: 1 ♂, between Macas and Sucua Río Blanco (- 3.43, - 78.57, 1700 m), 1 x 1937, leg.?, det. KJT [FSCA]. Cora unresolved: Haber (pers. comm. 2017) reports a potential new species from far eastern Ecuador with no specific data provided.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B57AFFBCFF23F5782778FB8B.taxon	discussion	Relatively small damselflies compared to Cora and Polythore, with beautifully patterned wings from dark with white or yellow bands (Fig. 56), to hyaline with orange patches. Three of the six species are recorded for Ecuador with certainty; a fourth species possibly occurs. See Bick & Bick (1992) for identification. Habitat is mostly small, mountainous, forested streams; females require decaying wood for oviposition. Species of Euthore are rarely en- countered in the field. Euthore fasciata Hagen in Selys, 1853. Publ. Recs. ECU: * Selys (1873 a — as E. plagiata), McLachlan (1878), Selys (1879 — “ a race of E. fasciata ”), Schmidt (1942 — as E. fasciata inlactea), Tsuda (1986, 1991), Paulson (1983), Steinmann (1997 a — as “ E. fasciata inlactea ”), (P 11), (H 14); MN: Bick & Bick (1992); MS: Bick & Bick (1992 — “ E. fasciata form plagiata ”); NA: Bick & Bick (1992). Also COL, PER: (H 08), (P 11); plus BRA: (H 08); plus VEN: (De Marmels 2016). Addl. Data. MN, Chone: 1 ♂, Chone (- 0.68, - 80.07, 110 m), 4 viii 1939, leg.?, det. Bick & Bick [FSCA]; MS, Morona: adult?, Río Upano (- 2.2500, - 78.2667, 2550 m), date?, leg. L. Alonzo, det. Bick & Bick, data provided by RWG [UMMZ]; NA, Archidona: 1 ♂, Cotundo (- 0.8833, - 77.8000, 611 m), vi 1975, leg?, det. Bick & Bick [RWG]; 1 ♂, [Sarayacu], 12.3 km N of Narupa on Baeza Hwy, small rocky stream (- 0.67, - 77.78, 1570 m), 14 vii 1996, leg. & det. WFM [FSCA]; Quijos: 2 ♂♂, small stream, 5 km S of Baeza (- 0.4758, - 77.8714, 1820 m), 20 xi 1997, leg. & det. KJT [KJT]; 1 ♂, 1 ♀, same data but 16 x 2009, leg. & det. JTJ [JTJ]; 1 ♀, same data but 03 xi 2001, leg. Fred Sibley, det. WFM [FSCA]: Tena: 1 ♂, Río Napo (- 1.0, - 78.0, 900 m), 1937, leg. WCM, det. Bick & Bick, data provided by RWG [UMMZ]. Note: The type locality for E. fasciata was listed as Venezuela (Hagen in Selys 1853). De Marmels (1988) speculated that the type locality was most likely Ecuador, although he found 10 different South American rivers named “ Río Negro. ” It is possible that the type locality was the Ecuadorian Río Negro, a tributary of the Río Pastaza which was accessible during the mid- 19 th century and which fits into the probable range of this species. * Euthore fassli Ris, 1914. Publ. Recs: COL: Ris (1914). Addl. Data. IM, Cotacachi: Santa Rosa, La Florida Reserve, Cock of The Rock Trail, small stream, trib. of El Placer (0.3606, - 78.4788, 1863 m), 30 viii 2018, leg. & det. KJT [KJT]. Euthore mirabilis McLachlan, 1878. Publ. Recs. ECU: Kirby (1890), Kennedy (1919), Davies & Tobin (1984), Tsuda (1986, 1991), Paulson (1983), Steinmann (1997 a), (P 11), (H 14); IM: * McLachlan (1878 — type series 6 ♂♂, 6 ♀♀: [Cotacachi], “ Intag ” [Apuela, NE on Apuela-Cuicocha Road, along Río Intag and tributaries between 1800 – 2800 m, 0.3588, - 78.4509 at jct of Río Intag and tributary], 1 x 1839, leg. Clarence Buckley, det. R. McLachlan- 1881, data provided by RWG [BMNH], Selys (1879), Kennedy (1919), Campos (1922), Montgomery (1967), Kimmins (1970 — designated the lectotype [BMNH]); Kennedy (1970), Bick & Bick (1992 — as “ Cora mirabilis ”), Bridges (1993). Note: All references quote the type specimens from Intag. A recent search for this species in the Intag area came up negative (Tennessen & Mauffray 2018).	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B579FFBCFF23F29925F7F9CB.taxon	discussion	Medium-sized damselflies with clear wings, sometimes with an apical brown spot on the hindwing (Fig. 57). One of the two known species occurs in Ecuador. Habitat is sluggish small streams, oviposition taking place in damp logs. See Kennedy (1940) for identification. Miocora peraltica Calvert, 1917. Publ. Recs. ECU: (H 14), (P 16); * ES: Daigle et al. (1997). Also CEN, COL: (H 08), (P 11). Addl. Data. ES, San Lorenzo: 2 ♂♂, stream 5 km NW of Lita along RR tracks (0.8847, - 78.4667, 500 m), 5 ii 1997, leg. & det. JJD [JJD, KJT]; 1 ♂, Lita along RR track 1 – 5 km W adjacent to Río Baboso seeps, pools & streams (0.8792, - 78.4589, 535 m), 5 ii 1997, leg. & det. WFM [FSCA]; 1 ♂, Río Canande Reserve, 35 km NE of Quininde, tributary of Río Hoja Blanca, swampy trickle in forest (0.5264, - 79.2122, 345 m), 20 x 2012, leg. Fred Morrison & WAH, det. WAH [WAH]. Note: Questions remain concerning the synonymy of M. subapicalis with M. peraltica (Bota-Sierra, pers. comm.).	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B579FFB9FF23F0D82036FB1F.taxon	discussion	Mostly large damselflies with patterned wings (Figs. 58 a & 58 b), sometimes with only narrow black band at the tip. Twenty species have been described, of which 11 occur in Ecuador; two undescribed species in Ecuador are known. Species identification was provided by Bick & Bick (1985, 1986). Polythore inhabit small streams to seepage runs in forested areas, from the lowlands to high elevations. Females lay eggs in dead fallen branches above the flowing water. Polythore aurora (Selys, 1879). Publ. Recs. ECU: Campos (1922), Davies & Tobin (1984), Tsuda (1986, 1991), Steinmann (1997 a), (L 05), Paulson (1983), Heckman (2008), (P 11), (H 14); NA: * Selys (1879 — type series: (originally 4 adults) 2 ♂♂, 1 ♀, [now designated as lectotype and paralectotypes, Rojas-Riano (2011 a)] “ Río Napo ” [Tena], Río Napo (- 1, - 78), date?, leg.?, from McLachlan, data provided by RWG, [BMNH]), Kirby (1890), Schmidt (1942), Fraser (1946 a), Montgomery (1967), Bick & Bick (1986), Bridges (1993). Also PER: (P 11); plus BRA: (L 05), (H 08). Addl. Data. Rojas-Riano (2011 a) discusses the confusion of the designated types of this species. There are no other records of this species from Ecuador despite many field trips to the Río Napo area near Tena. There is a possibility of mislabeling, but we retain it on the list for now. Polythore concinna (McLachlan, 1881). Publ. Recs. ECU: Kirby (1890), Campos (1922), Fraser (1946 a), Davies & Tobin (1984), Tsuda (1986, 1991), Paulson (1983), Steinmann (1997 a), (H 08), (P 11), (H 14). MS: Bick & Bick (1986); Napo: Bick & Bick (1986) Tennessen (1996 b), Daigle (1996); PA: * McLachlan (1881 — as T. concinna, Type series, 15 ♂♂, 14 ♀♀: “ Río Bobonaza ” — [Pastaza], Río Bobonaza, (- 1.5833, - 77.8000, 600 m), [BMNH, IRSNB]), Kimmins (1970 — designated Lectotype [BMNH]), Montgomery (1967), Bick & Bick (1986), Bridges (1993); Tennessen (2006); Rojas-Riano (2011 a — designated five paralectotypes [BMNH]); PC: Bick & Bick (1986); TU: Bick & Bick (1986). Also COL: (H 08), (P 11); plus PER: Rojas-Riano (2011 a). Addl Data. MS, Huamboya: [FSCA, KJT, SWD]; Morona: [FSCA]; NA, Archidona: [DRP, FAL, FSCA, JJD, KJT, RWG, SWD, USNM]; Carlos Julio Arsemena Tola: [FSCA, RWG]; Tena: [DRP, FSCA, RWG]; OR: Francisco de Orellana: (Steve Rovell photo); Loreto: [FSCA, JTJ, KJT, RWG, SWD, TWD, USNM]; PA, Mera: [FSCA, JTJ, KJT, RWG]; Pastaza: [FSCA, UTIC, KJT, RWG, TWD, USNM]; Santa Clara: [KJT, RWG]; PC, Quito: [FSCA, RWG]; SU, Shushufindi: [FSCA, KJT]; TU, Baños: [FSCA, RWG]. Note: Polythore concinna was common at a number of the locations and usually outnumbered other members of the genus. Polythore derivata (McLachlan, 1881). Publ. Recs. ECU: Kirby (1890), Campos (1922), Davies & Tobin (1984), Tsuda (1986, 1991), Paulson (1983), Tennessen (1996 b), Steinmann (1997 a), (H 08), (P 11), (H 14); PA: * Mclachlan (1881 a — as T. derivata type series, 6 ♂♂, 2 ♀♀: “ Río bobonaza ” [Pastaza], Río Bobonaza (- 1.5833, - 77.8000, 600 m), [BMNH]), Ris (1918), Schmidt (1942), Fraser (1946 a — as P. picta derivata), Montgomery (1967), Kimmins (1970 — designated Lectotype [BMNH]), Bick & Bick (1985), Bridges (1993); Rojas-Riano (2011 a — designated 2 Paralectotype ♂♂ s [IRSNB]); MS: Bick & Bick (1985); Napo: Bick & Bick (1985); OR: Daigle (1996), von Ellenrieder & Garrison (2011); PC: Bick & Bick (1985); SU: Bick & Bick (1985); TU: Bick & Bick (1985). Also COL, PER: (H 08), (P 11); Plus BOL: (H 08). Addl. Data. MS, Huamboya: [FSCA, KJT, RWG, TWD, USNM]; Limón Indanza: [JJD, KJT]; Taisha: [RWG]; NA, Carlos Julio Arosemena Tola: [FSCA]; Tena: [DRP, FSCA]; OR, Francisco de Orellana: [FSCA, UTIC, KJT, RWG, SWD]; Loreto: [JTJ, KJT, RWG, SWD]; PA, Mera: [FSCA, JJD, KJT, RWG]; Pastaza: [FSCA, KJT, TWD, USNM]; Santa Clara: [RWG]; SU, Cascales: [KJT]; Gonzalo Pizarro: [FSCA, QZAC]; Lago Agrio: [RWG, SWD]; Shushufindi: [DRP, FAL, FSCA, RWG, SWD, USNM]; TU, Baños: [FSCA]; ZC: Zamora: (C. Willigalla). Polythore gigantea (Selys, 1853). Publ. Recs. ECU: Hagen (1877), McLachlan (1878), Ris (1918), Campos (1922 — por Dr. Martin, all as Thore gigantea), Montgomery (1967), Tsuda (1986, 1991), Paulson (1983), Steinmann (1997 a), (H 08), (P 11), Altamiranda-Saavedra et al. (2014), (H 14); BO: Bick & Bick (1985), Roja-Riano (2011 a); CB: * Selys (1873 a); CT: Bick & Bick (1985), Rojas-Riano (2011 a); IM: Bick & Bick (1985), Rojas-Riano (2011 a); LR: Bick & Bick (1985); PC: Bick & Bick (1985), Rojas-Riano (2011 a); SD: Bick & Bick (1985, 1986); TU: Rojas-Riano (2011 a). Also COL, PER: (H 08), (P 11), Rojas-Riano (2011 a). Note: Not reported from Ecuador in Rojas-Riano (2011 a). Addl. Data. BO, San Miguel: [FSCA, RWG]; CT, Latacunga: [RWG]; Pangua: [USNM]; Pujili: [RWG]; Sigchos: [FSCA]; IM, Ibarra: [FSCA, RWG]; LR, Quevedo: [USNM]; NA, Tena: [FSCA]; PC, Quito: [ACRP]; Mejía: DRP. RWG]; San Miguel Bancos: [SWD]; SD: [DRP, FSCA, KJT, JTJ, RWG, SWD, USNM, USFQ]; TU, Baños: [FSCA, QZAC]. Note: Primarily a high elevation species found also in the central valley, but most records are from the upper west slope. Polythore mutata (McLachlan, 1881). Publ. Recs. ECU: Kirby (1890), Campos (1922), Davies & Tobin (1984), Tsuda (1986, 1991), Paulson (1983), Steinmann (1997 a), (H 08), (P 11), (H 14); NA: Bick & Bick (1986); OR: von Ellenrieder & Garrison (2011); PA: * McLachlan (1881 — as Thore mutata, type locality: “ Río Bobonaza ” [BMNH]), Fraser (1946 a), Montgomery (1967), Kimmins (1970 — designated lectotype), Bick & Bick (1986), Bridges (1993), Rojas-Riano (2011 a — designated 7 paralectotypes IRSNB]); SU: Bick & Bick (1986), Donnelly (1998). Also COL: (H 08), (P 11); plus PER: (P 11). Addl. Data. NA, Carlos Julio Arsemena Tola: 1 ♂, Río Anzu (- 1.2000, - 77.8333, 700 m), viii 1934, leg. WCM, det. Bick & Bick, data provided by RWG [UMMZ]; Tena: 1 ♂, [Puerto Napo], Río Napo (- 1.04, - 77.78, 436 m), date?, leg.?, det. Bick & Bick [FSCA]; 2 ♂♂, 2 ♀♀, J. S. B. S., 23 km E Puerto Napo, tropical wet forest, streams above flood plain (450 m), leg. A. Jaramillo, det. DRP [DRP]; same data but 9 – 13 vi 1994, 3 ♂♂, 1 ♀, same data but 3 – 11 xi 1994 [DRP]; 4 ♂♂, 2 ♀♀, same data but 17 x 1988, leg. & det. SWD [FAL, FSCA, JJD, SWD]; OR, Francisco de Orellana: 1 ♂, 1 ♀, streams in hill forest S of Napo River from La Selva, 13 x 1988, leg. & det. SWD [SWD]; 2 ♂♂, P. N. Y., Yasuni Biological Research Sta. along trails and roads within 5 km of Res. Sta. (- 0.6719, - 76.3992, 244 m), 28 – 31 viii 2000, leg.?, det. JJD [FSCA]; same but 1 ♂, 4 x 1998, leg. L. N. Harris, det. WFM [FSCA]; 4 ♂♂, 1 ♀, T. B. S., USFQ, stream crossing Chorongo trail (- 0.6378, - 76.1486, 224 m), 11 i 2009, leg. & det. RWG & NVE [NVE, RWG, USFQ]; 3 ♂♂, 1 ♀, T. B. S., USFQ, stream crossing Chorongo trail at marker 225 (- 0.6350, - 76.1503, 220 m), 16 – 18 i 2009, leg. & det. RWG & NVE [RWG, USFQ]; 5 ♂♂, T. B. S., USFQ, stream crossing Matapalo trail at marker 450 (- 0.6369, - 76.1458, 245 m), 14 – 15 i 2009, leg. & det. RWG & NVE [NVE, RWG, USFQ]; 1 ♂, T. B. S., USFQ, blackwater stream crossing Guacamayo trail at marker 2050 (- 0.6383, - 76.1492, 220 m), 17 i 2009, leg. & det. RWG & NVE [NVE]; 3 ♀♀, T. B. S., USFQ, along Guacamayo trail at marker 1700 (- 0.6350, - 76.1533, 237 m), 17 i 2009, leg. & det. RWG & NVE [NVE, USFQ]; 2 ♂♂, T. B. S., USFQ, Parahuaco trail between markers 300 – 1300 (- 0.6397, - 76.1453, 229 m), 21 i 2009, leg. & det. RWG & NVE [USFQ]; 1 ♂, T. B. S., USFQ, along Chichico trail (- 0.6300, - 76.1525, 250 m), 12 i 2009, leg. & det. RWG & NVE, [USFQ]; Loreto: 1 ♀, Loreto, N at pond outlet (- 0.8203, - 77.3185), 2 xi 2013, leg. & det. KJT [FSCA]; SU, Lago Agrio: Santa Cecilia, Río Francisco de Orellana (0.0850, - 76.9925), 25 – 31 iii 1969, leg. P & P Spangler, det.? [USNM]; Cascales: 1 ♂, 37 km W of Lago Agrio, river, 20 viii 1980, leg. & det. SWD [SWD]; Shushufindi: 1 ♂, 1 ♀, Limoncocha (- 0.4000, - 76.6167, 269 m), 24 – 26 vii 1977, leg. & det. D. Paulson [DRP]; 1 ♂, same data but, river and stream, 23 viii 1978, leg. & det. KWK [RWG]; 2 ♂♂, 1 ♀, leg. & det. SWD [SWD]; 1 ♂, same data but Río Napo, 12 xi 1974, leg. B. Drummond III, det.? [FSCA]; 3 ♂♂, 1 ♀, same data but 20 v 1976, leg. E. Rogers, det.? [FSCA]; 46 ♂♂, 3 ♀♀, 3 – 28 xi 1980, leg. & det. MJW [FSCA]; 2 ♂♂, 1 ♀, Shushufindi, stream about 2 km W, 12 v 2009, leg. & det. KJT [KJT]; 1 ♂, trails, small streams 3 km W of Shushufindi, 2 + km N off Hwy. 26 at km marker 17 (- 0.1664, - 76.6917, 290 m), 14 v 2009, leg. & det. KJT [KJT]; 1 ♂, same data but 13 x 2009, leg. & det. JTJ [JTJ]; 1 ♂, same data but 14 x 2009 [JTJ]; 1 ♂, [Limoncocha] E, N of La Selva Lodge, Lago Mandicocha & outlet stream “ coca ” [about 100 km E on Río Napo] (- 0.4983, - 76.3733, 200 m), 12 x 1988, leg. & det. SWD [FSCA]; 4 ♂♂, 1 ♀, same data but, forest, 22 – 25 xi 1997, leg. & det. TWD [SWD]. Polythore neopicta Bick & Bick, 1990. Publ. Recs. ECU: (H 08), (P 11), (H 14); MS: * Tennessen (2006). Also PER: (H 08), (P 11). Addl. Data. GY, Guayaquil: 2 ♂♂, 1 ♀, Guayaquil (- 2.20, - 79.90, 8 m), ii 1921, leg. F. W. Goding, det. GHB, checked by KJT (doubtful locality data) [FSCA]; MS, Huamboya: 2 ♂♂, 2 ♀♀, Río Pasanac waterfall, 57 km N of Macas, Hwy. 45 (- 1.9532, - 77.8519), 16 ix 2005, leg. & det. JJD [JJD, KJT]; 5 ♂♂, 1 ♀, small stream about 1 km S of Río Pasanac, Hwy. 45 (- 1.9500, - 77.8583, 800 m), 16 ix 2005, leg. & det. KJT [KJT]; 1 ♂, Macas, 58 km N on Hwy. 45 [70 km N of Sucua], roadside ditch and tribs near Río Pasanac (- 1.9447, - 77.8536, 762 m), 9 xi 1997, leg. & det. WFM [LSAM]; Limón Indanza: 2 ♂♂, waterfall 25.3 km N of Indanza, Hwy. 45, 7 xi 1997, leg. & det. JJD [JJD, KJT]; Santiago: 1 ♂, Yuganza [just S of Yuganza], 25.3 km N of Plan de Milagro Jct [trib to Río Yuanza], small stream with mossy rocks (- 2.8683, - 78.3739, 680 m), 7 xi 1997, leg. & det. WFM [FSCA]; OR, Loreto: Loreto, 1 ♀, 8.5 km E on road to Coca Road, rocky gravel stream (- 0.6183, - 77.2950, 400 m), 18 xi 1997, leg. & det. WFM [FSCA]. Polythore ornata (Selys, 1879). Publ. Recs. BOL, PER: Heckman (2008), Paulson (2011). Addl. Data. MS, Morona: 1 ♀, Giulca, Río Upano, 1400 m, no date, leg. L. Alonzo, det. RWG from C. H. Ken- nedy – UMMZ [RWG]; 1 ♀, Macas, 15 km W on Macas-Guamote Road, small stream and roadside seepage pools (- 2.2731, - 78.1939, 1067 m), 17 ix 05, leg. WFM, det. JJD, checked by WFM [FSCA]. Polythore procera (Selys, 1869). Publ. Recs. ECU: * Selys (1873 a — as Thore picta var aequatorialis, Holotype of T. aequatorialis: “ Equador ”, from McLachlan [BMNH]), Hagen (1875 — as T. aequatorialis), McLachlan (1878 — as T. procera), Kirby (1890 — as T. aequatorialis), Campos (1922 — as T. aequatorialis), Fraser (1946 a — as P. picta), Ris (1918 — as T. aequatorialis), Kimmins (1970 — as T. picta var aequatorialis), Tsuda (1986, 1991), (L 05), Paulson (1983), Bridges (1993), (H 08), (P 11), (H 14); BO: Bick & Bick (1985); CB: Selys (1873 a — as T. gigantea race procera), Bick & Bick (1985); MS: Bick & Bick (1985), Tennessen (2006); Rojas-Riano (2011 a); NA: Bick & Bick (1985), Daigle (1996); PA: McLachlan (1881), Bick & Bick (1985), Bick & Bick (1985), Rojas-Riano (2011 a); PC: Selys (1873 a — as T. gigantean race procera), Bick & Bick (1985); SU: Bick & Bick (1985); TU: Bick & Bick (1985), Rojas-Riano (2011 a); ZC: Bick & Bick (1985), Tennessen & Johnson (2008). Also BRA, COL: (L 05), (H 08), (P 11). Addl. Data. CB, Camada: [FSCA, RWG]; EO, Piñas: [JTJ]; LJ, Loja: [UTIC]; MS, Morona: [FSCA, JJD, KJT, RWG]; NA, Archidona: [FSCA, JJD, KJT, RWG]; El Chaco: [SWD]; Tena: [RWG]; PA, Mera: [FSCA, JTJ, KJT, RWG]; Pastaza: [KJT]; SD: [FSCA, JTJ, KJT]; SU, Gonzalo Pizarro: [DRP, FAL, QZAC, RWG]; TU, Baños: [FSCA, RWG, SWD]; ZC, Zamora: [FSCA, KJT, USNM]. Polythore terminata Fraser, 1946. Publ. Recs. ECU: Tsuda (1991), De Marmels (1999), Daigle et al. (1996), (P 11), (H 14); MS: * Bick & Bick (1985), Tennessen (1998); NA: Bick & Bick (1985), Tennessen (1998); ZC: Bick & Bick (1985), Tennessen & Johnson (2008). Also PER, VEN: (H 08), (P 11). Addl. Data. MS, Limón Indanza: 2 ♂♂, stream, Hwy. 45, 25.3 km N of Indanza (- 2.9217, - 78.4031, 1110 m), 7 vi 1997, leg. & det. KJT [KJT]; Morona: 2 ♂♂, 1 ♀, Macas, Río Upano (- 2.3167, - 78.1167, 1050 m), i 1939, leg.?, det. Bick & Bick, data provided by RWG [UMMZ]; 6 ♂♂, 1 ♀, Mangosisa, Río Upano (- 2.5244, - 77.8947, 850 m), xi 1945, leg. L Alonzo, det. Bick & Bick, data provided by RWG [UMMZ]; 6 ♂♂, 3 ♀♀, same data but [FSCA]; 1 ♀, Macas, Río Upano (- 2.30, - 78.10, 1050 m), date?, leg. L. Alonzo, det. GHB [FSCA]; 3 ♂♂, 2 ♀♀, Macas 15.7 km W on Macas-Guamote Road, small stream (- 2.2689, - 78.1936, 1463 m), 17 ix 2005, leg. & det. WFM [FSCA, LSAM, QZAC]; 1 ♂, 1 ♀, same data but leg. & det. KJT [KJT]; Santiago: 2 ♂♂, Chupianza (- 2.6833, - 78.3000, 1364 m), ii 1939, leg.?, det. Bick & Bick, data provided by RWG [AMNH]; Sucua: 2 ♂♂, 2 ♀♀, Sucua (- 2.4667, - 78.1667, 800 m), ii 1939, leg.?, det. Bick & Bick, data provided by RWG [UMMZ]; NA, Archidona: 1 ♂, Archi- dona (- 0.91, - 77.81, 573 m), date?, leg. R. Haensch, det. GHB [FSCA]; 1 ♀, Jondachi (- 0.7167, - 77.8000, 1000 m), 15 xi 1976, leg. H. Real, det. GHB [RWG]; 2 ♂♂, 1 ♀, Archidona (- 0.9167, - 77.800, 604 m), 13 x 1976 – 18 xii 1976, leg. H. Real, det. GHB [RWG]; 1 ♂, [Narupa] E on road to Loreto 43.5 km W of Loreto seep near large rocky river [Río Hollin] (- 0.7022, - 77.7189, 1036 m), 15 vii 1996, leg. & det. KJT [KJT]; 1 ♂, same data but leg. & det. WFM [FSCA]; 1 ♂, same data but 14 xi 1997 [KJT]; 1 ♂, same data but leg. & det. TWD [SWD]; OR, Loreto: 2 ♂♂, rocky river seeps, 34.2 km W of Loreto on Loreto Road (- 0.7417, - 77.5517, 900 m), 19 xi 1997, leg. & det. TWD [SWD]; 1 ♂, same data but leg. & det. JJD [JJD]; 1 ♂, same data but leg. & det. KJT [KJT]; ZC, Zamora: 1 ♂, stream / waterfall, near entrance of Podocarpus Nat. Pk., S of Zamora (- 4.1053, - 78.9639), 2 iv 2008, leg. & det. KJT [FSCA]; 1 ♂, trails, Podocarpus Nat. Pk. (- 4.1144, - 78.9642, 1040 m), 2 iv 2008, leg. & det. JTJ [JTJ]; 1 ♀, stream / waterfalls on trails, Podocarpus Nat. Pk. (- 4.1217, - 78.9728, 1070 m), 2 iv 2008, leg. & det. JTJ [JTJ]; 1 ♂, stream 5 km N of Guaguayme Alto (- 3.8778, - 78.8928, 880 m), 4 iv 2008, leg. & det. KJT [FSCA]; 1 ♂, same data but leg. & det. JTJ [JTJ]; 4 ♂♂, 1 ♀, Zamora (- 4.0692, - 78.9567, 1000 m), 15 x 1941, leg. D. Laddey, det. Bick & Bick, data provided by RWG [UMMZ]; 1 ♀, same data but 25 x 1941 [FSCA]; 1 ♂, same data but xi 1941 [FSCA]; 1 ♂, stream crossing road just before the village of Guadalupe (- 3.9113, - 78.8575), 20 viii 2006, leg. & det. JCA [UTIC]; 2 ♂♂, 1 ♀, stream off road to village of Guadalupe, N of Zamora (- 3.8758, - 78.8923), 21 viii 2006, leg. & det. JCA [UTIC]; Nangaritza: 2 ♂♂, Cordillera de Condor, 20 km SE Paquisha, 21 ix 1990, leg. OSF, det. SWD [SWD, USNM]. Polythore victoria (McLachlan, 1869). Publ. Recs. PI: Rojas-Riano (2011 a); PA: Rojas-Riano (2011 a). Also BOL, COL, PER: Rojas-Riano (2011 a). Note: Not much is known about the distribution or biology of this species. Specimen data is from the early 19 th century. Polythore vittata (Selys, 1869). Publ. Recs. ECU: * Fraser (1946 a), Montgomery (1967 — as Thore vittata race aequatorialis), Paulson (1983), Tsuda (1986, 1991), Steinmann (1997 a), Lencioni (2005), (P 11), Rojas-Riano (2011 a), (P 14); MS: Bick & Bick (1986); PA: Bick & Bick (1986). Also BRA, PER: (L 05), (P 11), Rojas-Riano (2011 a). Addl. Data. EO, Piñas: 1 ♂, 1 ♀, Moromoro (- 3.72, - 79.73, 1500 m), 22 vii 1941, leg. D. Laddey, det. Bick & Bick 1985 [FSCA]; 1 ♂, same data but 14 vii 1941 [FSCA]; PA, Pastaza: Río Bobonaza (- 1.5833, - 77.8000, 600 m), date?, leg.?, det. Bick & Bick 1985 [RWG]. Note: The status of vittata, picta and their relationship to procera are still uncertain. A specimen labeled “ Macas via Riobamba ” was listed by Bick & Bick (1986) as “ Chimborazo or Morona-Santiago. ” We consider it to be MS. Polythore unresolved: There are several specimens in various collections that have not been determined and may represent undescribed species: Polythore n. sp. (to be described by KJT: AZ, Santa Isabela: 1 ♂, 1 ♀, stream, 21 km E of Hwy. 25, road from Balao Jct. (- 2.9400, - 79.5625), 11 iv 2008, leg. KJT [KJT]; 1 ♀, same data but leg. JTJ [JTJ]; Polythore sp.: PA, Santa Clara: adult, 25 km N id Puyo, 11 xi 1988, leg. R. K. Robbins [USNM].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B57CFFB9FF23F2E52128F8F3.taxon	discussion	A monotypic genus, medium-sized with clear wings; males are unusual in possessing a prominent, vertical projection on abdominal segment 10 (Fig. 59; also, see Garrison et al. 2010). Habitat is small rivers at medium elevations (approximately 1000 m). Stenocora percornuta Kennedy, 1940. Publ. Recs. ECU: Davies & Tobin (1984), Tsuda (1986, 1991), Paulson (1983 & 2004 a), Steinmann (1997 a), (P 11), (H 14); NA: Tena * Kennedy (1940 — Holotype: Río Jatun Yacu, Oriente Prov, Río Napo watershed (- 1.0167, - 77.8333, 700 m), 12 ix 1935, leg. WCM, det. CHK, data provided by RWG [ANSP]); PA: * Kennedy (1940 — Allotype: [Santa Clara], El Partidero, Indian village on east bank of Río Anzu, jct. of trails from Mera and Puyo northward to Napo, edge of tropical jungle (- 1.2833, - 77.8000, 800 m), 12 xi 1935, leg. WCM, det. CHK, data provided by RWG [UMMZ]), Bridges (1993), Garrison et al. (2003), Montgomery (1967), Garrison et al. (2003), Tennessen (2006). Also PER: (P 11); Plus BRA: (H 08). Addl. Data. OR, Loreto: 1 ♀, Loreto, 8.5 km E on road to Coca Road, rocky gravel stream (- 0.6183, - 77.2950, 400 m), 20 vii 1996, leg. & det. WFM [FSCA]; PA, Mera: 1 ♂, Río Metolo, Los Copales cabanas between Mera and Shell, W of Puyo, Hwy. 50, small streams / trails (- 1.4919, - 78.0725, 1070 m), 19 viii 2005, leg. & det. JJD [JJD]; 1 ♂, same data but leg. & det. KJT [JJD]; 2 ♀♀, same data but 22 viii 2005, leg. & det. KJT [KJT]. Note: IUCN Red List Status is NT, Near Threatened (von Ellenrieder 2009 c).	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B563FFA6FF23F25A210DFB49.taxon	discussion	Aeshnidae are commonly known as “ darners ” in the New World. They are medium to large in size and are strong fliers. Adults range in color pattern from mostly drab brown to intricately colored with blue, green and yellow marks on a black or brown background. Nine genera and 47 species occur in Ecuador.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B563FFA6FF23F3562081F93D.taxon	discussion	A genus of only two species that are very restricted in distribution (Garrison et al. 2006). See Ris (1904) and Martin (1909) for identification. These large dragonflies apparently inhabit small streamlets on steep mountain sides in cloud forest. One species (Fig. 60) has been found in Ecuador. Allopetalia pustulosa Selys, 1873. Publ. Recs. COL, VEN: (H 06), Garrison (2006), (P 11); plus PER: Garrison (2006). Addl. Data. PC, Mejía: 1 ♂, Chincha via Chiriboga 25 v 1981, leg. & det.? [QZAC]; TU, Baños: 1 ♀, Baños, “ in cane field, no nearby water ” (- 1.40, - 78.4, 1850 m), 1 xi 1937, leg.? det?. Data provided by RWG [UMMZ].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B563FFA7FF23F0C3262AF81E.taxon	discussion	Large, strong flying dragonflies (Fig. 61); most of the five New World species are widespread, including the two species recorded in Ecuador. Geijskes (1968) clarified the status of A. concolor; for a key to the species, see Needham et al. (2014). Habitat for the two species in Ecuador is mainly shallow, weedy ponds. Anax amazili (Burmeister, 1839). Publ. Recs. ECU: Paulson (1983), Tsuda (1986, 1991), (H 14); GA: * Calvert (1906), Currie (1901), Martin (1908), Martin (1911), Campos (1922), Schmidt (1938), Asahina (1961), Gloger (1964), Turner (1967), (Peck 1992 — nymph collections from brackish lagoons), (P 11); GY: Campos (1922), Calvert (1934), EO: Campos (1927). Also ARG, BRA, CEN, PRY, PER, TTO, URY, VEN: (H 06), (P 11); plus COL: (P & P); and GUF, SUR: (P 16). Addl. Data. GA, Santa Cruz: 1 ♀, Isla Santiago, Jaboncillo Camp, grasslands, 8 – 9 iv 1992, leg. J. Cook & S. Peck, det.? [USNM]; 1 ♀, same data but littoral mangrove, UV light, 28 iii 1992 [USNM]; 1 ♀, same data but Bahia Sullivan, arid zone, night UV light [USNM]; 1 ♀, San Cristobal Island (- 0.9243, - 89.0087, 191 m), 26 ii 2014, leg. & det. ACRP [ACRP]; GY, Guayaquil: 1 ♂, 1 ♀, Puerto Grande (- 2.93, - 80.08, 2 m), ii 1936, leg.?, det. P. Calvert [ANSP, FSCA]; 2 ♂♂, 5 ♀♀, Guayaquil, (- 2.20, - 79.90, 8 m), date?, leg. F. Campos, det. MJW [ANSP, FSCA]; 1 ♀, Milagro, ditch / pool S of Milagro, Hwy. E 25 (- 2.1650, - 79.6078, 35 m), 8 iii 2010, leg. & det. KJT [FSCA]; LR, Ventanas: 1 ♂, pond / stream 7.2 km E of Hwy. E 25, 9 km SW of Quinzaloma (- 1.2539, - 79.3703, 100 m), 21 ii 2010, leg. & det. JTJ [JTJ]; SU, Shushufindi: 1 ♂, Limoncocha, Río Napo (- 0.4000, 76.6167, 269 m), 25 xi 1980, leg. & det. MJW [FSCA]; 4 ♀♀, same data but 8 xi 1980 [FSCA]; 2 ♂♂, 2 ♀♀, same data but Thorn Palm Swamp on Río Napo [FSCA]; adult, Shushufindi-Southwest Station, 25 viii 1975, leg. A. Langley, det.? [USNM]. Anax concolor Brauer, 1865. Publ. Recs. ECU: (H 06), (P 11), (H 14). PA: * Tennessen (1998). Also ARG, BOL, BRA, CEN, COL, GUF, SUR, VEN: (H 06); (P 11). Addl. Data. MS, Santiago: 2 ♂♂, pond along road between Tayuza and Mendez (- 2.7283, - 78.1700, 600 m), 6 xi 1997, leg. & det. TWD [SWD]; 1 ♂, pond 5 km N of Bella Union, Hwy. 45, 8 xi 1997, leg. & det. JJD [JJD]; PA, Pastaza: 1 ♂, pond near Comunidad Chuwitayo, S of Puyo, 9 xi 1997, leg. & det. KJT [KJT]; NA, Tena, 2 ♂♂, as A. longipes concolor, J. S. B. S., Pond # 4, 24 xi 1991, leg. E. McCallie, Ellen, det.? [USNM]; Archidona: 3 ♂♂, as A. longipes, Cueva Jumandy, about 5 km N Archidona, 13 ix 1990, leg. OSF, det.? [USNM]; OR, Francisco de Orellana: 1 ♂, Indiyama 8 iii 1980, leg. & det.? [QZAC].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B561FFA2FF23F38C2162FDA7.taxon	discussion	Five of the eight species of Coryphaeschna occur in Ecuador. For identification, refer to Paulson (1994) and Tennessen (2001). These large dragonflies (Fig. 62) occupy shallow weedy ponds and slow reaches of streams, usually in forested areas. Coryphaeschna adnexa (Hagen, 1861). Publ. Recs. ECU: Martin (1909, 1911 — both as Aeschna adnexa), Hincks (1934), Geijskes (1943), Paulson (1983), Tsuda (1986, 1991), Steinmann (1997 b), (H 06 )) (P 11), (H 14); GY: * Calvert (1906 )), Campos (1922, 1931 — both as A. adnexa), Calvert (1956); EO: Campos (1927); NA: Calvert (1956), Daigle (1996). Also ARG, BRA, CEN, COL, GUF, PRY, PER, SUR, VEN: (H 06), (P 11); plus TTO (P 11). Addl. Data. LR, Buena Fé: 1 ♀, [Consumulo] S Río Palenque (- 0.58, - 79.35, 175 m), 11 vi 1974, leg. J. Longi- no, det. JJD [FSCA]; 1 ♂, Babahoyo, swamp at Babahoyo, 21 vi 1975, leg. J. Cohen, A. Langley & P. Monnig, leg.? [USNM]; NA, Tena: 1 ♂, Río Sinde, pond, 5 km E of Puerto Napo (- 1.0517, - 77.7544, 450 m), 13 vii 1996, leg. & det. KJT [KJT]; 1 ♂, same data but 22 vii 1996, leg. & det. JJD [JJD]; 1 ♂, 1 ♀, same data but leg. & det. WFM [FSCA, LSAM]; 1 ♂, J. S. B. S., 23 km E of Puerto Napo, Tropical Wet Forest, 18 x 1988, leg. & det. SWD [UTIC]; 3 ♂♂, 1 ♀, Río Napo slough, 3.2 km E of Puerto Napo, 23 vii 1996, leg. & det. JJD [JJD]; OR, Francisco de Orellana: 1 ♂, pond near trib. of Río Payamino, road to Loreto, 1.7 km W of Coca Road (- 0.4367, - 77.0017, 258 m), 13 vi 1995, leg. & det. KJT [FSCA]; 1 ♂, same data but 18 xi 1997 [KJT]; 1 ♀, P. N. Y., 2.1 km E of Yasuni Biological Research Sta., pond and swamp (- 0.6741, - 76.3685, 244 m), 15 xi 1997, leg. & det. WFM [FSCA]; SU, Shushufindi: Limoncocha (- 0.4000, - 76.6167, 269 m), 24 vii 1977, leg. & det. DRP [DRP]; 1 ♂, same data but 23 viii 1980, leg. & det. SWD [FSCA, KJT]; 1 ♀, same data but 25 viii 1980 [UTIC]; 1 ♂, same data but lawn, 23 viii 1980, leg. & det. SWD [JJD]; 1 ♂, same data but 24 viii 1980 [UTIC]; 1 ♀, same data but forest trail, 24 vii 1971, leg. D. Pearson, det. DRP [DRP]; 1 ♀, same data but Río Napo, 16 xi 1973, leg. B. Drummond III, det.? [FSCA]; 1 ♀, same data but 15 i 1964; 1 ♂, same data but 19 iv 1974 [FSCA]; same data but 6 xi 1980, leg. & det. MJW [FSCA]; 2 ♂♂, same data but 1 xi 1980 [FSCA]; 1 ♂, same data but 18 xi 1980; 1 ♂, same data but Thorn Palm Swamp “ on Río Napo, ” xi 1980, leg. & det. MJW [FSCA]; “ La Selva, 100 km E of Coca, Manicocha, ” 12 x 1988, leg. & det. SWD [FSCA]. Coryphaeschna amazonica De Marmels, 1989. Publ. Recs. ECU: * (H 14). Also BRA, CEN, GUF, PER, SUR, VEN: (H 06), (P 16). Addl. Data. ES, Quininde: 1 ♂, Río Hoja Blanca, 3 km N of Río Canande Reserve, 35 km NE of Quininde (0.5439, - 79.2035), date not provided, data provided by WAH & DLW; PA, Pastaza: 1 ♂, Comuna San Ramon (- 1.3786, - 77.8670), date not provided; data provided by WAH & DLW. Coryphaeschna diapyra Paulson, 1994. Publ. Recs. ECU: (H 06 )), (P 11), (H 14); SU: * Paulson (1994). Also CEN, VEN: (H 06), (P 11). Addl. Data. SU, Shushufindi: 3 ♂♂, Limoncocha, Thorn Palm Swamp “ on Río Napo ” (- 0.4000, - 76.6167, 269 m), 26 xi 1980, leg. MJW, det. DRP [FSCA]. Coryphaeschna huaorania Tennessen, 2001. Publ. Recs. ECU: (Paulson 2004 a) (P 11), (H 14); MS: * Tennessen (2001 — Holotype & Allotype: [Patuca], [about 1 ml N], 43.4 km S of Sucua, large pond [about 5 km N of Mendez road jct] (- 2.7283, - 78.2700, 600 m), 6 xi 1997, leg. TWD [FSCA]); NA: * Tennessen (2001); OR: * Tennessen (2001). Note: So far reported only from Ecuador. Addl. Data. NA, Tena: 1 ♀, pool near Río Napo, 3.2 km E of Puerto Napo (- 1.0456, - 77.7467, 420 m), 23 vii 1996, leg. & det. KJT [KJT]; OR, Francisco de Orellana: 1 ♂, P. N. Y., pool near Río Sabaleto, 17 vii 1996, leg. & det. KJT [KJT]; 1 ♂, pond, Loreto Road, 11 km W of Río Payamino ferry [now bridge] (- 0.4800, - 77.0800, 300 m), 18 xi 1997, leg. & det. KJT [KJT]; 1 ♂, 11.8 km W on Loreto Road, large pond (- 0.4742, - 77.0583, 285 m), 18 xi 1997, leg. WFM, det. KJT [FSCA]. Coryphaeschna viriditas Calvert, 1952. Publ. Recs. ECU: Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14); GY: Calvert (1956); LJ: * Campos (1925 — as C. virens). Also BOL, BRA, CEN, COL, GUF, GUY, PRY, PER, VEN, SUR: (H 06), Paulson (2009); plus TTO: (P 11). Addl. Data. PA, Pastaza: 1 ♀, old road to Tena, 22 km E of Puyo, 2 m wide polluted stream bordered by forest and cutover area (- 1.424, - 77.874), date not provided; data provided by WAH & DLW.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B567FFA1FF23F47C269CFB7B.taxon	discussion	Gynacantha are large, predominantly brown amd / or green dragonflies (Fig. 63) that are typically crepuscular. It is the most species-diverse genus of Aeshnidae in Ecuador, with 12 species recorded. The main reference for iden- tification is Williamson (1923 a). Only a few species are commonly encountered. The nymphs develop rapidly in temporary forest pools, although at least one species occupies phytotelmata. Gynacantha adela Martin, 1909. Publ. Recs. ECU: Schmidt (1942), (H 14); CR: * Campos (1922). Also ARG BOL, BRA, PER: (H 06), (P 11). Addl. Data. SU, Shushufindi: 1 ♀, Limoncocha, 2 xi 1980, leg. MJW, det. WFM [FSCA]; ZC, Nangaritza: 1 paratype ♂, “ Mayaicu ” — as “ Morona-Santiago Province: Mayalico on Río Naganitza (- 3.0592, - 77.9956, 490 m) ” [corrected to Mayaicu, Río Nangaritza, (- 4.0290, - 78.6340, 900 m), 27 xi 1941, leg. D. Laddey, det. CHK, rechecked by WFM [FSCA]. Gynacantha auricularis Martin, 1909. Publ. Recs. ECU: * Paulson (1983), (H 06), (P 11), (H 14). Also BOL, BRA CEN, GUF, GUY, PER, SUR, VEN: (H 06), (P 11). Addl. Data. OR, Francisco de Orellana: 1 ♀, Yasuni Scientific Station, Río Tiputini, 39 km SE of Pompeya, 61 km ESE of Coca (- 0.6741, - 76.3974), date not provided; data provided by WAH & DLW; SU, Shushufindi: 1 ♀, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 13 v 1975, leg. B. Drummond III, det. MJW [FSCA]; 1 ♂, same data but forest trail, 10 i 1972, leg. D. Pearson, det. DRP [DRP]; 1 ♂, same data but 24 viii 1980, leg. & det. SWD [SWD]; 1 ♀, same data but 28 viii 1980 [SWD]. Gynacantha bifida Rambur, 1842. Publ. Recs. ECU: (H 14); CR: * Campos (1922), EO: * Campos (1922, 1927), GY: * Campos (1922). Plus ARG, BRA, PRY, PER, VEN: (P 11); also COL: Palacino et al. (2017). Addl. Data. GY, Balzar: 1 ♂, Batan (- 1.17, - 79.92, 64 m), date?, leg. F. Campos, det. P. Calvert, checked by WFM [FSCA]; Guayaquil: 2 ♀♀, Guayaquil (- 2.20, - 79.90, 8 m), date?, leg. F. Campos, det. P. Calvert [FSCA]; LR, Quevedo: 1 ♀, Quevedo, xi 1985, leg. & det.? [QZAC]. Gynacantha caudata Karsch, 1891. Publ. Recs. ECU: * Karsch (1891 — Type “ Ecuador ”), Campos (1922 — “ pers. comm., Calvert ”), Paulson (1983), Davies & Tobin (1984), Tsuda (1986, 1991), Bridges (1993), Steinmann (1997 b), (H 06), (P 11), (H 14); CR: Williamson (1923 a). Also CEN: (H 06), (P 11). Addl. Data. GY, Playas: 1 ♂, Playas (- 2.63, - 80.38, 6 m), 20 iv 1938, leg.?, det.? [FSCA]; LR, Babahoyo: 1 ♂, 1 ♀, Playas de Montalvo (- 1.8167, - 79.2833, 15 m), 2 iv 1938, leg.?, det. CHK, from UMMZ [RWG]; 1 tan. pair, same data but [FSCA]; 2 ♂♂, Playas de Montalvo (- 1.6833, - 79.2833, 15 m), 15 iv 1938, leg. WCM, det. CHK; data provided by RWG [UMMZ]; 1 ♀, same data but 16 iv 1938 [UMMZ]; MN, Flavio Alfaro: 1 ♂, small forested stream 0.5 km N of Chindul, 1 iii 2010, leg. & det. JJD [JJD]; OR, Francisco de Orellana: 1 ♂, P. N. Y., Yasuni Biological Research Sta. along trails and roads within 5 km of Res. Sta. (- 0.6719, - 76.3992, 244 m), 1 ix 2000, leg. T. Emmel, det. KJT [FSCA]. Gynacantha gracilis (Burmeister, 1839). Publ. Recs. ECU: Hincks (1934), Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14); OR: von Ellenrieder & Garrison (2011). Also ARG, BOL, BRA, CEN, GUF, GUY, PER, SUR, VEN: (H 06), (P 11). Addl. Data. NA, Tena: 1 ♂, J. S. B. S., Pond # 2, 14 xi 1991, leg. E. McCallie, det.? [USNM]; OR, Francisco de Orellana: 1 ♀, T. B. S., USFQ, Guacamayo trail between markers 450 – 2100 (- 0.6350, - 76.1533, 237 m), 22 i 2009, leg. & det. RWG & NVE [NVE]; SU, Shushufindi: 1 ♂, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 5 xi 1980, leg. & det. MJW [FSCA]; 1 ♂, same data but forest trail, 27 vii 1971, leg. & det. D. Pearson [DRP]; 2 ♂♂, same data but 8 viii 1971 [DRP]; 1 ♂, 1 ♀, same data but 26 viii 1971 [DRP]; 1 ♀, same data but 28 i 1972 [DRP]; 1 ♀, same data but 24 viii 1980, leg. & det. SWD [SWD]. Gynacantha interioris Williamson, 1923. Publ. Recs. ECU: (H 06), (P 11), (H 14); OR: * Daigle (1996), von Ellenrieder & Garrison (2011). Also BOL, BRA, PER, GUF, VEN: (H 06), (P 11). Addl. Data. NA, Tena: 1 ♂, 1 ♀, Río Chinchipino, 27 km E of Puerto Napo (- 1.0728, - 77.5864, 385 m), 10 x 2009, leg. & det. KJT [FSCA]; 3 ♂♂, 1 ♀, same data but leg. & det. JTJ [JTJ]; OR, Francisco de Orellana: 1 ♀, P. N. Y., ditch near Sci. Research Sta. (- 0.6694, - 76.3975, 229 m), 9 vi 1995, leg. & det. KJT [KJT]; 1 ♀, same data but leg. & det. WFM [FSCA]; 1 ♂, same data but leg. & det. SWD [SWD]; 1 ♂, same data but 10 vi 1995, leg. & det. JJD [JJD]; 1 ♂, same data but leg. & det. KJT [KJT]; 1 ♂, same data but leg. & det. WFM [FSCA]; 1 ♂, same data but leg. & det. SWD [SWD]; 1 ♂, same data but 11 vi 1995 [SWD]; 1 ♂, 1 ♀, same data but 17 vii 1996, leg. & det. KJT [KJT]; 1 ♂, same data but leg. & det. WFM [QZAC]; 2 ♂♂, 2 ♀♀, same data but 18 vii 1996, leg. & det. JJD [JJD]; 1 ♂, 1 ♀, same data but 15 xi 1997, leg. & det. KJT [KJT]; 1 ♂, 1 ♀, same data but leg. & det. TWD [SWD]; 1 ♀, same data but 16 xi 1997 [SWD]; 1 ♂, same data but 17 xi 1997 [SWD]; SU, Shushufindi: 1 ♀, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 12 v 1975, leg. B. Drummond III, det. MJW [FSCA]; 1 ♀, same data but 24 viii 1980, leg. & det. SWD [FSCA]; 1 ♂, same data but 28 viii 1980 [SWD]; 1 ♂, same data but 2 xi 1980, leg. & det. MJW [FSCA]. Gynacantha litoralis Williamson, 1923. Publ. Recs. ECU: Paulson (1983), (H 14). Also: GUF, PER, SUR: (H 06), (P 11); plus BRA: (H 06); and COL: (P 16); and VEN: De Marmels (2016). Addl. Data. OR, Francisco de Orellana: 1 ♀, P. N. Y., Yasuni Biological Research Station, grassy ditches, (- 0.6694, - 76.3975, 229 m), 16 ii 1997, leg. & det. WFM [FSCA]; SU, Shushufindi: 1 ♂, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 25 xi 1980, leg. & det. MJW [FSCA]; 1 ♂, same data but forest trail, 8 vii 1971, leg. D. Pearson, det. DRP [DRP]; 1 ♂, same data but 8 viii 1971 [DRP]; 1 ♂, same data but 22 viii 1971 [DRP]; 1 ♀, same data but 18 xii 1971 [DRP]; 1 ♂, same data but 24 vii 1977, leg. & det. DRP [DRP]; 1 ♂, same data but 23 viii 1980, leg. & det. SWD [SWD]; 1 ♂, same data but 28 viii 1980 [SWD]; 1 ♂, forest near Río Napo, S of Limoncocha (- 0.4203, - 76.6108, 250 m), 11 v 2009, leg. & det. KJT [FSCA]. Gynacantha membranalis Karsch, 1891. Publ. Recs. ECU: Hincks (1934), Paulson (1983), Tsuda (1986, 1991), Steinmann (1997 b), (H 06), (P 11), (H 14); ES: Daigle et al. (1997); NA: Ris (1918); OR: von Ellenrieder & Garrison (2011); PA: * Calvert (1906). Also BOL, BRA, CEN, COL, GUF, GUY, PER, SUR, VEN: (H 06), (P 11); plus TTO: (P 11). Addl. Data. ES, San Lorenzo: 1 ♀, tiny pool near RR track, about 5 km N of Lita, 5 ii 1997, leg. & det. KJT [KJT]; MS, Morona: 1 ♂, Macas, “ Oriente, Acc. 788 ”, (- 2.30, - 78.12, 1006 m), date?, leg.? det.? [FSCA]; Taisha: 1 ♂, Macuma, forest trail (867 m), 2 ii 1972, leg. D. Pearson, det. DRP [DRP]; NA, Tena, 1 ♂, J. S. B. S., near Tena, inside station (450 m), 13 vi 1994, leg. A. Jaramillo, det. DRP [DRP]; 1 ♂, same data but 14 vi 1994 [DRP]; 1 ♀, same data but 22 vi 1994 [DRP]; 1 ♂, same data but 18 x 1988, leg. & det. SWD [SWD]; 1 ♀, Río Nushino, viii 1984, leg. Venodiocloff, det. JJD; OR, Francisco de Orellana: 1 ♂, T. B. S., USFQ, open grassy area behind dining room, surrounded by bushes and trees and some seepage areas (- 0.6383, - 76.1492, 220 m), 10 i 2009, leg. & det. RWG & NVE [USFQ]; 2 ♂♂, T. B. S., USFQ, along Harpia trail, marker 350 (- 0.6250, - 76.1633, 228 m), 18 i 2009, RWG & NVE [NVE, USFQ]; 1 ♀, Reserva Etnica Waorani, 500 m N of Onkonne Gare Camp, UV light (- 0.63 - 76.60), 10 i 1994, leg. T. Erwin, det.? [USNM]; 1 ♀, same data but 31 i 1994, leg. R. Medina, det.? [USNM]; SU: Shushufindi: 1 ♂, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 5 ix 1972, leg. T. Emmel, det. MJW [FSCA]; 1 ♂, same data but 3 – 16 vi 1977, leg. P. Spangler, det.? [USNM]; 1 ♂, same data but 1972, leg. P. Kazan, det. SWD [SWD]; 1 ♂, same data but forest trail, 19 vii 1971, leg. D. Pearson, det. DRP [DRP]; 1 ♂, same data but 24 vii 1971 [DRP]; 1 ♂, same data but 27 vii 1971 [DRP]; 1 ♀, same data but 10 viii 1971 [DRP]; 1 ♂, same data but 26 viii 1971 [DRP]; 1 ♀, same data but 17 i 1972 [DRP]; 1 ♂, same data but 24 i 1972 [DRP]; 1 ♀, same data but 13 iii 1972 [DRP]; 3 ♂♂, trails, small streams 3 km W of Shushufindi, 2 + km N off Hwy. 26 at km marker 17 (- 0.1664, - 76.6917, 290 m), 14 x 2009, leg. & det. KJT [FSCA, JTJ]; 1 ♂, La Selva Lodge; forest (- 0.4883, - 76.3733, 200 m), 22 xi 1997, leg. & det. TWD [SWD]. Gynacantha mexicana Selys, 1868. Publ. Recs. ECU: Martin (1909, 1911), Maria (1938), Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14); CR * Campos (1922), Williamson (1923 a); EO: * Campos (1922, 1927); GY: * Campos (1922), Campos (1931); LR: Williamson (1923 a). Also BRA, CEN, COL, GUF, GUY, PER, VEN: (H 06), (P 11); plus BOL, SUR: (H 06); and TTO: (P 11). Addl. Data. GY, Naranjal: 1 ♂, 1 ♀, Finca de la Gringa, 15 km E of Tres Cerritos (- 2.4456, - 79.5556, 25 m), 11 iv 2008, leg. & det. KJT [KJT]; 2 ♂♂, 1 ♀, same data but 12 iv 2008, leg. & det. JTJ [JTJ]; 1 ♀, Guayaquil (- 2.20, - 79.90, 8 m), date?, leg. F. Campos, det.? [FSCA]; LR, Montalvo: 1 ♀, small stream 1.8 km N of Hwy. 491, 25 km E of Babahoyo, 7 iii 2010, leg. & det. JJD [JJD]. Gynacantha nervosa Rambur, 1842. Publ. Recs. ECU: Hincks (1934), Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14); CR: * Campos (1922), Williamson (1923 a); LR: Williamson (1923 a). Also BOL, BRA, CEN, COL, GUF, GUY, SUR, VEN: (H 06), (P 11); plus PER: (H 06), (P 16); and TTO: (P 11). Addl. Data. EO, Piñas: 1 ♂, Buenaventura, Riachuelo 14 (- 3.6868, - 79.7438, 555 m), 23 vi 2016, leg. S. Vil- lar, det. ACRP [ACRP]; 1 ♂, 2 ♀♀, same data but “ casa ” (- 3.6542, - 79.7682, 555 m); ES, Esmeraldas: 2 ♂♂, 1 ♀, Hacienda Guinchele, 7.5 km E of Esmeraldas, 5 v 1963, leg. T. Hubbell & L. Pena, det. NVE, from CHK (NV, UMMZ]; Quininde: 1 ♀, Cupe (320 m), 23 i 1982, leg. & det.? [QZAC]; GY, Guayaquil: 1 ♂, Guayaquil (- 2.7333, - 79.75, 17 m), 20 iv 1963, leg. T. Hubbell & L. Pena, det. NVE [UMMZ]; 1 ♂, Guayaquil, leg. F. Campos, det. MJW [ANSP]; Naranjal: 8 ♂♂, 1 ♀, Cerro Mas Vale Fundaction Andrate (- 2.40, - 79.63, 15 m), 13 xi 2001, leg. & det. F. Sibley [FSCA]; 5 ♂♂, 2 ♀♀, same data but 14 xi 2001 [FSCA]; 2 ♂♂, 1 ♀, “ Finca de la Gringa, ” 15 km E of Tres Cerritos, near Cerro Cimalón (- 2.4456, - 79.5556, 25 m), 11 iv 2008, leg. JTJ [JTJ]; 1 ♂, 1 ♀, same data but 12 iv 2008, leg. & det. KJT [FSCA]; LR, Montalvo: 1 ♂, 1 ♀, small stream 1.8 km N of Hwy. 491, about 25 km E of Babahoyo (- 1.8111, - 79.3189, 60 m), 12 iv 2008, leg. & det. KJT [KJT]; 1 ♂, same data but 7 iii 2010, [FSCA]; NA, Tena: 1 ♀, J. S. B. S., near Tena, inside forest (450 m), 22 vi 1994, leg. A. Jaramillo, det. DRP [DRP]; OR, Francisco de Orellana: 1 ♀, Coca, iii 1988, leg. & det.? [QZAC]; SU, Lago Agrio: 1 ♀, Lago Agrio, at lights, 15 iv 1976, leg. A. Langley, det.? [USNM]; Shushufindi: 1 ♀, Limoncocha, forest trail, 12 i 1972, leg. D. Pearson, det. DRP [DRP]. Gynacantha tenuis Martin, 1909. Publ. Recs. ECU: * Paulson (1983), (H 06), (P 11), (H 14). Also BRA, GUF, GUY, PER, SUR, TTO, VEN: (H 06), (P 11); plus COL: (H 06), (P 11), (P & P); and BOL, CEN: (P 11). Addl. Data. NA, Tena: 1 ♂, Puerto Napo, 23 km E, J. S. B. S., 16 x 1988, leg. & det. SWD [SWD]; SU, Shushufindi: 1 ♂, Limoncocha, lake edge, 22 viii 1971, leg. D. Pearson, det. DRP, verified DRP [DRP]. Gynacantha tibiata Karsch, 1891. Publ. Recs. ECU: * Karsch (1891 c — type: “ Ecuador ” specimen lost), McLachlan (1896), Calvert (1906), Martin (1909, 1911), Campos (1922), Maria (1938), Paulson (1983), Tsuda (1991), Bridges (1993), Steinmann (1997 b), (H 06), (P 11), (H 14). Also CEN, PER, VEN: (H 06), (P 11). Addl. Data. LR, Quevedo: 1 ♀, Quevedo, Río Palenque (- 1.03, - 79.43, 100 m), 15 xi 1972, leg. D. Dodson, det. WFM [FSCA]; SD: 4 ♂♂, about 93 km SW Quito, Chiriboga Road, 9 vii 1990, leg. OSF, det.? [USNM]; 1 ♂, Tinalandia, 12 km E of Santo Domingo de los Colorados, streams S of river (- 0.2839, - 79.0500, 760 m), 5 x 1988, leg. & det. SWD [SWD]. Two additional specimens are probably this species: NA, Tena: adult, J. S. B. S., Pond # 2, 20 xi 1991, leg. E. McCallie, det. SWD “ tibiata group ” [USNM]; same data but J. S. B. S., Pond # 1, 24 xi 1991. Note: Williamson & Williamson (1930 a) discussed the type of tibiata which was lost at that time, and stated that “ the issue with the Ecuador specimen was not settled and additional specimens from Ecuador need to be collected. ” The additional records presented here verify its existence in Ecuador.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B564FFAFFF23F30821B7FE4B.taxon	discussion	A genus of 15 large, dark species (Figs. 64 a & 64 b), seven of which are recorded in Ecuador. For species identification, see Belle (1989). Habitat is small streams in swampy woodlands; adults are crepuscular and seldom seen, therefore being rarely collected. * Neuraeschna calverti Kimmins, 1951. Publ. Recs. ECU: (H 14 — as N. cf. calverti). Also BRA, PER: (H 06), (P 11); plus SUR, VEN: (P 11). Addl. Data. OR, Francisco de Orellana: 1 ♀, [Anangu] 100 km E Coca near La Selva, S across Río Napo, streams in hill forest (- 0.53, - 76.38, 290 m), 13 x 1988, leg. & det. SWD [SWD]; 1 ♀, cf. Río Shiripuno Lodge, 27 km SE Río Shiripuno bridge, (- 1.1047, - 76.7317, 235 m), date not provided; data provided by WAH & DLW; SU, Cuyabeno: 1 ♂, 1 ♀, Nicky Lodge on Río Cuyabeno, 210 m (- 0.1977, - 75.9243), date not provided; data provided by WAH & DLW; Shushufindi: 1 ♂, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 18 xi 1980, leg. D. Pat- terson, det. SWD, checked by WFM [FSCA]. Neuraeschna claviforcipata Martin, 1909. Publ. Recs. ECU: (H 06), (P 11), (H 14); SU: * Belle (1989 — as “ Napo ”), De Marmels & Neiss (2013). Also BRA, GUF, SUR, VEN: (H 06), (P 11). Addl. Data. SU, Shushufindi: 1 ♀, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 5 xi 1980, leg. & det. J. Belle [FSCA]. Neuraeschna costalis (Burmeister, 1839). Publ. Recs. ECU: (H 06), (P 11), (H 14); PA: * Belle (1989). Also BRA, GUF, GUY, SUR, VEN: (P 11). Addl. Data. ES, San Lorenzo: 1 ♀, Concepcion (1.05, - 78.83, 25 m), date?, leg. F. Campos, det. MJW [ANSP]; PA, Mera: 1 ♂, Shell, Río Pastaza, Puyo [presumably W, just downriver from Shell] (- 1.53, - 78.05, 1000 m), 21 xi 1936, leg. WCM, det. CHK [FSCA]. SU, Shushufindi: 1 ♀, cf. La Selva Lodge, (- 0.4978, - 76.3741), date not provided; data provided by WAH & DLW. Neuraeschna dentigera Martin, 1909. Publ. Recs. ECU: (H 14?). Also BRA, GUY, PER, SUR: (H 06). Addl. Data. OR, Francisco de Orellana: 1 ♀, Yasuni Scientific Station, Río Tiputini, 39 km SE of Pompeya, 61 km ESE of Coca (W- 550) (- 0.6741, - 76.3974), date not provided; data provided by WAH & DLW. * Neuraeschna harpya Martin, 1909. Also BRA, GUF, GUY, PER, SUR, VEN: (H 06), (P 11). Addl. Data. NA, Tena: 1 ♀, J. S. B. S., near Tena, inside station (450 m), 1 xii 1994, leg. A. Jaramillo, det. DRP [DRP]; OR, Francisco de Orellana: 1 ♀, T. B. S., Río Tiputini, 55 km SE of Pompeya, 94 km ESE of Coca (- 0.6369, - 76.1503), date not provided; data provided by WAH & DLW; SU, Cuyabeno: 1 ♂, Río San Francisco, 850 m N of Nicky Lodge on Río Cuyabeno (- 0.1977, - 75.9243, 210 m), date not provided; data provided by WAH & DLW. Neuraeschna maya Belle, 1989. Publ. Recs. ECU: * (H 06), (P 11). Also CEN: (H 06), (P 11). Addl. Data. MS, Santiago: 1 ♀, pond 5 km N of Mendez Junction, Hwy. 45, 8 xi 1997, leg. & det. JJD [JJD]. Neuraeschna titania Belle, 1989. Publ. Recs. ECU: Tsuda (1991), (H 06), (P 11), (H 14); NA: * Belle (1989); PC: * Belle (1989 — Holotype: [Quito], Jaruqui [probably Yarugui] (- 0.1500, - 78.3167, 2728 m), date?, leg. Staudinger [BMNH]); PA: * Belle (1989 — Allotype: [Pastaza] Puyo, Río Pastaza watershed (1000 m), 28 xi 1936, leg. WCM [UMMZ]; ZC: * Belle (1989), Bridges (1993). Also PER: (H 06), (P 11). Addl. Data. MS, Morona: 2 ♂♂, 1 ♀, Mangosisa, Río Upano (- 2.5244, - 77.8947, 850 m), date?, leg. L. Alonzo, det. NVE, from CHK; data provided by RWG [RWG, UMMZ]; NA, Archidona: 1 ♀, Archidona, at light (- 0.9167, - 77.8000, 604 m), 1 – 31 v 1971, leg. H. Real, det. J Belle [RWG]; OR, Francisco de Orellana: 1 ♂, Yasuni Scientific Station, Río Tiputini, 39 km SE of Pompeya, 61 km ESE of Coca (- 0.6741, - 76.3974), date not provided; data pro- vided by WAH & DLW; PA, Pastaza: 2 ♀♀, Puyo, Río Pastaza watershed [twilight] (- 1.4667, - 77.9833, 1000 m), 21 – 28 xi 1936, leg. WCM, det. J. Belle; Santa Clara: 1 ♀, headwaters of Río Arajuno, Napo watershed (- 1.0833, - 77.5333, 1000 m), 29 iv 1941, leg. WCM, det. RWG [UMMZ]; ZC, Zamora: 1 ♂, Zamora (- 4.000, - 78.9567, 700 m), 3 xii 1941, leg. D. Laddey, det. J. Belle, from CHK; data provided by RWG [UMMZ].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B56AFFADFF23F1B627CEFDDF.taxon	discussion	Only four species of Remartinia are known, two of which occur in Ecuador. These are large green aeshnids with small black markings (Fig. 65). See Calvert (1956) and Carvalho (1992) for species identification. Known mainly from small ponds with emergent vegetation along the edges. Remartinia luteipennis (Burmeister, 1839). Publ. Recs. LJ: * Tennessen & Johnson (2008), (H 14). Also ARG, BRA, CEN, COL, PRY, VEN: (H 06), (P 11). Addl. Data. ES, San Lorenzo: 1 ♂, roadside pool 36.5 km NW of Lita, near Urbina, 3 ii 1997, leg. WFM, det. KJT [KJT]; 1 ♂, [San Antonio], nearby, 36.5 km WNW of Lita on [old] San Lorenzo Hwy., pools / seeps / stream (0.95, - 78.68, 200 m), 3 ii 1997, leg. & det. WFM [FSCA]; 1 ♀, San Lorenzo, pools 5.6 km NW of Lita, Finca Con- suegros, 4 ii 1997, leg. & det. JJD [JJD]; 1 ♂, Lita along RR track 1 – 5 km W adjacent to Río Baboso seeps, pools & streams (0.8792, - 78.4589, 535 m), 5 ii 1997, leg. & det. WFM [LSAM]; GY, Naranjal: “ Finca de la Gringa, ” 15 km E of Tres Cerritos (- 2.4456, - 79.5556, 25 m), 12 iv 2008, leg. & det. KJT [KJT]; IM, Cotacachi: 1 ♂, Apu- ela, Ruta Plaza Gutierrez, seepage into Rio Tuabunchi (0.3575, - 78.4993, 1634 m], 27 viii 2018, leg. & det. WFM [FSCA]; same data, 1 ♂, 1 ♀, leg. & det. KJT [KJT]; Ibarra: 1 ♂, Salinas, 0.5 km E, pools / small irrigation stream (0.55, - 78.12, 1676 m), 6 ii 1997, leg. & det. WFM [FSCA]; LJ, Loja: 1 ♂, seep trib. of Río Chamba, Vilcabamba (- 4.2636, - 79.2172, 1555 m), 31 iii 2008, leg. & det. JTJ [JTJ]; MS, Santiago: 2 ♂♂, pond 5 km N of Bella Union, Hwy. 45, 7 xi 1997, leg. & det. JJD [JJD]; NA, Archidona: stream / pool 43.5 km W of Loreto, Loreto road, 15 vii 1996, leg. & det. JJD [JJD]; 1 ♀, small stream, Baeza-Tena Road, 12 km N Loreto Road, 14 vii 1996, leg. & det. KJT [KJT]; Tena: 1 ♀, Río Napo slough, 3.2 km E of Puerto Napo, 23 vii 1996, leg. & det. JJD [JJD]; PC, San Miguel de los Bancos: 1 ♂, Mindo, Mindo Gardens Ecolodge, vegetated cement pond (- 0.0720, - 78.7560, 1330 m], leg. & det. WFM [FSCA]; same data but 1 ♂, leg. & det. KJT [KJT]; SU, Shushufindi: 1 ♂, Limoncocha, Thorn Palm Swamp “ on Río Napo ” (- 0.4000, - 76.6167, 269 m), 26 xi 1980, leg. & det. MJW [FSCA]. Remartinia rufipennis (Kennedy, 1941). Publ. Recs. ECU: Paulson (1983 — as Coryphaeschna rufipennis), Tsuda (1986, 1991 — as C. rufipennis), Steinmann (1997 b — as C. rufipennis), (H 06), (P 11), (H 14); ES: Daigle et al. (1997) [Note: The ES specimens were later determined to be R. luteipennis]; MS: Tennessen (1998); PA: Calvert (1956) commented that he could not find “ Río Pupuyacu ”; we suspect it was Río Papayacu which is in Pastaza); ZC: Tennessen (2000). Also PER: (H 06), (P 11). Addl. Data. MS, Santiago: 1 ♀, pond along road between Tayuza and Mendez (- 2.7283, - 78.2700, 600 m), 6 iv 1997, leg. & det. TWD [SWD]; 1 ♀, same data but 7 xi 1997 [SWD]; 1 ♂, [Patuca], [about 1 ml N], 43.4 km S of Sucua, large pond [about 5 km N of Mendez road jct] (- 2.75, - 78.27, about 600 m), 7 xi 1997, leg. & det. KJT [KJT]; 1 ♂, same data but leg. & det. WFM [FSCA]; 1 ♂, 3 ♀♀, same data but 8 xi 1997, leg. & det. WFM [FSCA, LSAM, QZAC, USNM]; 2 ♀♀, same data but leg. KJT [KJT]; SU, Shushufindi: 1 ♂, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 31 viii 1974, leg. B. Drummond III, det. MJW [FSCA]; ZC, Zamora: 1 ♂, grassy marsh, 5.5 km E of Zamora, 5 xi 1997, leg. & det. KJT [KJT]; 1 ♀, ditch / pool near small stream, Bombuscaro Road S of Zamora, about 1000 m, 19 xiii 2006, leg. & det. KJT [KJT].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B568FFA8FF23F4A42442FB1F.taxon	discussion	A large genus (42 species), mainly found at higher elevations, over 1000 m. Adults are variously patterned with markings of blue, green, yellow, gray and black (Fig. 66). The recent revision by von Ellenrieder (2003 a) covers all 11 species occurring in Ecuador. A wide variety of habitats are utilized, including ponds, marshes, and small seeps to slow pools in small to mid-sized streams. Some species are relatively abundant. Rhionaeschna absoluta (Calvert, 1952). Publ. Recs. IM: * von Ellenrieder (2003 a). Also ARG, CHL, PER, URY: (H 06), (P 11). Addl. Data. IM, Ibarra: adult, Salinas (0.55, - 78.1333, 1544 m), leg. KJT, det. NVE [KJT]. Rhionaeschna biliosa (Kennedy, 1938). Publ. Recs. ECU: Paulson (1983), Tsuda (1991), Steinmann (1997 b — as Aeshna biliosa), (H 06), (P 11), (H 14); MN: von Ellenrieder (2003 a); TU: * Kennedy (1938 b — Holotype: La ventana near Baños ” [Baños], (2000 m), v 1936, leg. WCM UMMZ]; Kennedy (1939 d — Allotype: Baños, El Trablon (2000 m), 28 x 1938, leg. WCM [ANSP]), Calvert (1956), Bridges (1993), Garrison et al. (2003), von Ellenrieder (2003 a). Also PER: (H 06), (P 11). Addl. Data. MN, no other data, det. CHK, data provided by RWG [UMMZ]; TU, Baños: 1 ♂, Baños, Runtun Lake (- 1.40, - 78.42, 2200 m), 6 vi 1939, leg.?, det. WFM [FSCA]; 1 ♂, Santa Rosa [probably Baños, Lake Run- tun] (- 1.2833, - 78.6667, 2900 m), ix 1948, leg. WCM, det. NVE, from UMMZ [NVE]; 1 ♂, same data but xi 1948 [RWG]; 1 ♀, Baños, Río Ulba (- 1.4 - 78.3667, 3778 m), date?, leg.?, det. NVE from UMMZ [RWG]; adults, San- tiago Pillaro Chaupi, Mt. Tungurahua (- 1.1, - 78.65, 2600 – 3500 m), leg.?, det. P. Calvert 1956 [ANSP]. Rhionaeschna brevicercia (Muzón & von Ellenrieder, 2001). Publ. Recs. ECU: Paulson (2004 a), (H 06), (P 11), (H 14); ES: * Muzón & von Ellenrieder (2001), von Ellenrieder (2003 a); IM: * Muzón & von Ellenrieder (2001), von Ellenrieder (2003 a); NA: von Ellenrieder (2003 a); MN: * Muzón & von Ellenrieder (2001), von Ellenrieder (2003 a); PC: * Muzón & von Ellenrieder (2001 — Holotype ♂: as Aeshna brevicercia, [San Miguel Bancos], between Calacali and Nanegalito, about 30 km W of Quito, 2000 m, 4 xi 1990, leg. OSF [USNM]); SD: * Muzón & von Ellenrieder (2001 — Allotype ♀: “ as Pichincha ”, 24 i 1988, leg. Morales [NMNHL]; Muzón (2007); TU: Tennessen (2006), von Ellenrieder (2003 a). Also COL, VEN: (H 06), (P 11). Addl. Data. IM, Cotacachi: 1 ♀, Santa Rosa, El Refugio de Intag Cloud Forest Lodge, south along road at, roadside seeps (0.3675, - 78.4716, 1845 m), 28 viii 2018, leg. & det. KJT [KJT]; Otavalo: 1 ♂, Otavalo, ESE Lago San Pablo (0.20, - 78.22, 2800 m), 11 xii 1998, leg. H. Werning, det. NVE [QZAC]; NA, Archidona: 1 ♂, [Sa- rayacu], 13.5 km N of Narupa on Baeza Hwy., along road (- 0.65, - 77.78, 1676 m), 14 vii 1996, leg. & det. WFM [FSCA]; 1 ♂, [Sarayacu] 12.8 km N of Narupa on Baeza Hwy., small shallow sandy stream (- 0.67, - 77.78, 1585 m), 14 vii 1996, leg. & det. WFM [FSCA]; 1 ♀, Cosanga S on Hwy. 45, 26.7 km N of Narupa, road side ditch and seeps (- 0.6017, - 77.8338, 2240 m), 19 xi 1997, leg. & det. WFM [FSCA]; Quijos: 1 ♂, 1 ♀, Baeza, 4.8 – 5.0 km S, small clear streams and seeps (- 0.4761, - 77.8711, 1830 m), 20 xi 1997, leg. & det. WFM [FSCA]; 1 ♂, same data but leg. & det. TWD [SWD]; 1 ♂, small stream, 5 km S of Baeza (- 0.4758, - 77.8714, 1825 m), 10 v 2009, leg. S. Ingley & KJT, det. KJT [FSCA]; TU, Baños: 1 ♀, Ulba S along road to Runtun, small seepage / Eleocharis marsh (- 1.4074, - 78.4067, 1990 m), 13 ix 2005, leg. & det. WFM [FSCA]. Note: Specimens of this species were often labeled A. brevifrons or A. intricata prior to Muzón & von Ellenrie- der (2001). The majority of known collection localities can be found in Muzón & von Ellenrieder (2001). Rhionaeschna cornigera (Brauer, 1865). Publ. Recs. ECU: * Ris (1918), Campos (1922 — “ por Dr. Martin ”), Calvert (1947 a), Dunkle (1981), Paulson (1983), Tsuda (1986, 1991), Steinmann (1997 b — as Aeshna cornigera cornigera), (H 06), (P 11), (H 14); AZ: von Ellenrieder (2003 a); BO: von Ellenrieder (2003 a), CB: Calvert (1956), von Ellenrieder (2003 a); EO: Calvert (1956), von Ellenrieder (2003 a); LJ: von Ellenrieder (2003 a); MS: Tennessen (2006); LR: Calvert (1956); NA: Tennessen (1996 b), Daigle (1996), Tennessen (1998), von Ellenrieder (2003 a); PA: Calvert (1956), von Ellenrieder (2003 a); PC: von Ellenrieder (2003 a); SU: von Ellenrieder (2003 a); TU: Calvert (1956), von Ellenrieder (2003 a). Also BOL, CEN, COL, PER, VEN: (H 06), (P 11). Addl, Data. BO, San Miguel: [RWG]; CB, Alausí: [RWG]; Cumandá: [RWG]; Pallatanga: [RWG]; EO, Piñas: [RWG]; IM, Cotocachi: [FSCA, KJT, USFQ]; LJ, Loja: [RWG]; MS, Morona: [KJT, UMMZ]; NA, Archidona: [FSCA, JJD, KJT, QZAC, RWG, SWD]; El chaco: [SWD]; Quijos: [FSCA, JJD, KJT, LSAM, QZAC, SWD, SWD]; OR, Loreto: [FSCA JJD]; PA, Mera: [FSCA, RWG]; PC, Quito: [ACRP, RWG]; San Miguel: [NVE, RWG, QZAC]; SU, Gonzalo Pizarro: [QZAC]; TU, Baños: [ANSP, RWG]; ZA, Zamora: (C. Willigalla). Rhionaeschna elsia (Calvert, 1905). Publ. Recs. ECU: (H 06), (H 14); IM: Daigle et al. (1997). Also CHL, PER: (H 06), (P 11). Addl. Data. IM, Ibarra: 1 ♀, Salinas 1.5 km E small stream (0.4853, - 78.1333, 1615 m), 2 ii 1997, leg. & det. WFM [FSCA]; 1 ♀, same data but 6 ii 1997, leg. & det. KJT [KJT]; 1 ♂, pools / stream near sugarcane fields 0.5 km S of Salinas on road to Lita, 2 ii 1997, leg. & det. JJD [JJD]. Rhionaeschna galapagoensis (Currie, 1901). Publ. Recs. ECU: Tsuda (1986, 1991), (H 06), (P 11), GA, San Cristobal: * Currie (1901), Needham (1904), Martin (1908, 1911), Campos (1922), Schmidt (1938), Calvert (1956), Asahina (1961), Gloger (1964), Turner (1967), Belle (1990), Peck (1992), all as Aeshna galapagoensis; von Ellenrieder (2003 a), Cordero – Rivera et al. (2016); Isabela: Peck (1992), Santa Cruz: Peck (1992). Note: Endemic to the Galapagos; Peck (1992) recorded it from all cantons in “ arid to pampa zones with larval collections reported from brackish lagoons. ” Specific locality data can be found in Peck (1992) and other references listed above. Rhionaeschna intricata Martin, 1908. Publ. Recs. ECU: Martin (1908, 1911), Paulson (1983), Tsuda (1986, 1991), Steinmann (1997 b — as Aeshna intricata), (H 06), (P 11), (H 14); CB: von Ellenrieder (2003 a); NA: Tennessem (1998), PC: Martin (1908 — LECTOTYPE: [Quito], Nanegal (00.70, - 78.40, 2093 m), date?, data provided by NVE [IRSNB]), Calvert (1956); Muzón & von Ellenrieder (2001); TU: Calvert (1956) [could be R. brevicercia]. Also PER: (H 06), (P 11); plus COL: (H 11). Addl. Data. CB, Alausí: adults, Guasuntos (- 2.2167, - 78.8167, 2833 m), date?, checked by Muzón & NVE [NMNHL]; Pallatanga: adults, Pallatanga (- 1.9833, - 78.95, 2248 m), checked by Muzón & NVE [ANSP, NMNHL]; LJ, Loja: 1 ♂, stream and waterfall, 14 km S of Vilcabamba at Hwy. 39 (- 4.3439 - 79.1834), 17 viii 2006, leg. & det. JCA, checked by NVE [UTIC]; PC, Quito: adults, Quito (- 0.2167, - 78.5000, 2763 m), checked by Muzón & NVE [ANSP]. Note: According to von Ellenrieder (2003 a), most specimens in Martin’s type series were R. brevicercia. Previous records for ARG, BRA, CEN, CHL, and VEN are for other species. Rhionaeschna joannisi (Martin, 1897). Publ. Recs. ECU: Paulson (1983), Tsuda (1991), Steinmann (1997 b — as Aeshna joannisi), (H 06), (P 11), Bota-Sierra (2014 a), (H 14); LJ: Tennessen & Johnson (2008); NA: von Ellenrieder (2003 a); PA: * Calvert (1956), De Marmels (2001 b), von Ellenrieder (2003 a); PC: von Ellenrieder (2003 a). Also COL, BOL: (H 06), (P 11); plus VEN: (H 06). Addl. Data. CT, Sigchos: 1 ♂, Los Libres (2000 m), 5 xi 1994, leg. S. Espinoza, det. KJT [KJT]; IM, Cotacachi: 1 ♂, Santa Rosa, Ruta Siempre Verde, seeps and bog (0.3672, - 78.4378, 2226 m), 28 viii 2018, leg. & det. KJT [KJT]; LJ, Loja: 1 ♂, pond, entrance to Podocarpus Natl. Park (- 4.0911, - 79.2025, 2475 m), 6 iv 2008, leg. & det. KJT [KJT]; NA, Archidona: 1 ♀, roadside trib. 26.4 km N of Narupa jct. (2225 m), 19 xi 1997, leg. & det. KJT [KJT]; PA, Mera: adults, Abitagua (- 1.45, - 78.15, 1200 m), date?, leg.? det. P. Calvert [ANSP]; PC, Mejía: 1 ♂, 1 ♀, Cornejo Astorga, 9.6 km E, ditch (2012 m), 21 vii 1977, leg. & det. DRP [DRP]; San Miguel Bancos: 1 ♀, Tandayapa, above, on Tandayapa Valley Road, road through forest (1830 m), 1 vii 2008, leg. & det. DRP [DRP]. Rhionaeschna marchali (Rambur, 1842). Publ. Recs. ECU: * Martin (1909, 1911), Ris (1918), Schmidt (1942), Dunkle (1981), Paulson (1983), Tsuda (1986, 1991), Steinmann (1997 b — as Aeshna marchali), (H 06), (P 11), Bota- Sierra (2014 a), (H 14); CB: Campos (1922); CT: Campos (1922), Calvert (1956); EO: Campos (1922); IM: Calvert (1956); LJ: Calvert (1956), Tennessen & Johnson (2008); NA: Tennessen (1996 b); Daigle (1996); Tennessen (1998); PC: Campos (1922), Calvert (1956), Tennessen (1996 a); TU: Campos (1922), Navás (1934), Calvert (1956). Also COL, PER, BOL, VEN: (H 06), (P 11). Addl. Data. AZ, Cuenca: [ANSP, NMNHL]; Gualaceo: [KJT, NMNHL, TWD, QZAC, USNM]; Guano: [RWG]; Nabón: [FSCA, JJD, KJT, USNM]; Riobamba: [RWG]; BO, Sam Miguel: [RWG]; CR, Espejo: [ACRP]; Tulcan: [ANSP]; CB, Chambo: [RWG]; Colta: [FSCA, RWG]; CT, Latacunga: [ANSP, FSCA]; EO: [QZAC]; IM, Cotacachi: [ANSP, FSCA, KJT, USFQ]; Otavalo: [DRP, FSCA, JJD, KJT, QZAC, SWD, USFQ]; LJ, Loja: [ANSP, JJD, JTJ, RWG]; Saraguro: [NMNHL]; MN: [NMNHL]; NA, Archidona: [DRP, KJT, SWD]; El Chaco: [SWD]; Quijos: [FSCA, JJD, KJT, LSAM, QZAC, SWD, USNM]; PC, Cayambe: [DRP]; Mejía: [ANSP, DRP]; Quito: [FSCA, KJT, QZAC, SWD]; Rumiñahui: [KJT]; San Miguel Bancos: (KJT); SD: [FSCA, QZAC]; TU, Ambato: [RWG]; Baños: [ANSP, FSCA, UMMZ, RWG, QZAC, SWD, USNM]; Patate: [ANSP]; San Pedro: [QZAC]; ZC, Zamora: [RWG]. Rhinoaeschna planaltica (Calvert, 1952). Publ. Recs. ECU: Tsuda (1991), Steinmann (1997 b — as Aeshna cornigera planaltica), (H 06), (P 11), (H 14); AZ: Calvert (1952), von Ellenrieder (2003 a); BO: Calvert (1956), von Ellenrieder (2003 a); CB: * Calvert (1952), von Ellenrieder (2003 a); LJ: Tennessen & Johnson (2008); MS: * Calvert (1952), von Ellenrieder (2003 a); PA: Calvert (1952, 1956), von Ellenrieder (2003 a); PC: von Ellenrieder (2003 a). Also ARG, BOL, BRA, PRY, PER, URY, VEN: (P 11); plus COL: (H 06). Addl. Data. BO, San Miguel: Balzapamba (- 1.7667, - 79.1833, 700 m), no other data, det. P. Calvert, data from RWG [ANSP]; CB, Cumandá: Naranjapata (- 2.2667, - 79.0833, 650 m), no other data, det. P. Calvert, data from RWG [ANSP]; Alausí: Huigra (- 2.7833, - 79.4333, 3600 m), no other data, det. P. Calvert, data from RWG [ANSP]; LJ, Catamayo: 1 ♂, stream pool, 7 km from Hwy. 50 on road to Zaruma (- 3.9458, - 79.4953), 7 iv 2008, leg. & det. JTJ [JTJ]; 2 ♂♂, 1 ♀, same data but waterfall, [JTJ]; MS, Morona: Río Upano (- 2.20, - 78.45, 1050 m), no other data, det. NVE [UMMZ]; PA, Pastaza: 1 ♂, Puyo (- 1.4667, - 77.9833, 980 m), 4 iv 1976, leg. P. Spangler, det. NVE [USNM]; Mera: Abitagua (- 1.45, - 78.15, 1200 m), no other data, det. P. Calvert [ANSP]; PC, Mejía: 1 ♂, Cornejo Astorga 9.6 km E, ditch (2012 m), 16 vii 1977, leg. & det. DRP [DRP]; 1 ♀, same data but 21 vii 1977 [DRP]; 1 ♀, Tandapi (- 0.4169, - 78.7831, 1400 m), 15 – 20 vi 1965, leg.?, det. NVE [DRP]. Rhionaeschna psilus (Calvert, 1947). Publ. Recs. ECU: Paulson (1983), Tsuda (1991), (H 06), (P 11), (H 14); EO: von Ellenrieder (2003 a); MS: Tennessen (1998); NA: * Calvert (1947 a, 1956), von Ellenrieder (2003 a); MN: von Ellenrieder (2003 a); TU: * Calvert (1947 a & 1956), von Ellenrieder (2003 a). Also ARG, CEN, PER, VEN: (H 06), (P 011); plus COL: (P 16). Addl. Data. EO, Piñas: 2 ♂♂, Buenaventura, Río 20 (- 3.6792, - 79.7373, 880 m), 2 vii 2016, leg. & det. ACRP [ACRP]; 1 ♂, same data but, 4 vii 2016; 1 ♂, same data but Riachuelo 20 (- 3.6790, - 79.7378, 892 m), 29 vi 2016; MN, Jipijapa: 1 ♂, [Playa], San Jose, 3 mi inland (- 1.38, - 80.75, 60 m), 14 iii 1970, leg.?, det.? [FSCA]; Manta: 1 ♂, Santa Rosa (- 1.3167, - 80.6333, 352 m), 13 v 1946, leg. L. Alonzo, det. NVE, from CHK [UMMZ]; Portoviejo: 2 ♂♂, 1 ♀, concrete ditch, Hwy. E 40, E of Portoviejo (- 1.0672, - 80.1350, 420 m), 4 iii 2010, leg. & det. JTJ [JTJ]; MS, Santiago: pond, road between Tayuza and Mendez (- 2.7283, - 78.2700, 600 m), 6 xi 1997, leg. & det. TWD [SWD]; 2 ♀♀, [Patuca], [about 1 mi N], 43.4 km S of Sucua, large pond [about 5 km N of Mendez road jct] (- 2.75, - 78.27, 488 m), 7 xi 1997, leg. & det. JJD [JJD]; 1 ♂, same data but leg. & det. KJT; 1 ♂, same data but 8 xi 1997, leg. & det. WFM [FSCA]; NA, Tena: adults, Río Jatunyacu (- 1.0167, - 77.8333, 700 m), date?, leg.?, det. P. Calvert, data provided by RWG [ANSP]; OR, Loreto: 1 ♂, pond 38.9 km W of Loreto (990 m), 14 vi 1995, leg. & det. KJT [KJT]; 1 ♀, forest near Reserva Río Bigal, about 30 km N of Loreto (- 0.5642, - 77.4114, 900 m), 15 v 2009, leg. & det. KJT [FSCA]; TU, Baños: adults, La Palmera (- 1.4167, - 78.200, 1300 m), date?, leg.?, det. P. Calvert, data provided by RWG [ANSP].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B56DFFA8FF23F2E426EDF863.taxon	discussion	A genus containing five large species, two recorded in Ecuador. Male thorax is mostly dark green, that of female drab (Fig. 67). The genus was revised by Geijskes (1959), although clarification of some species is needed (Garrison et al. 2006). The habitat is small, slow streams in forested to partially open areas. Staurophlebia auca Kennedy, 1937. Publ. Recs. ECU: Geijskes (1959), Paulson (1983), Davies & Tobin (1984), Tsuda (1986, 1991), Steinmann (1997 b), (H 06), (P 11), (H 14); NA: * Kennedy (1937 a — Holotype: [Tena] “ Oriente: quebrada on the Sapallo Yacu ” [small trib. to Jatun Yacu] (800 m), iii 1937. leg. WCM [UMMZ]), Bridges (1993), Garrison et al. (2003 a). Also GUF PER, SUR, VEN: (H 06), (P 11). Staurophlebia reticulata (Burmeister, 1839). Publ. Recs. ECU: Hincks (1934), Geijskes (1959), Paulson (1983), Tsuda (1986, 1991), Steinmann (1997 b — as S. reticulata reticulata) (H 06), (P 11), (H 14; MN: * Ris (1916), Campos (1925); NA: Tennessen (1998). Also ARG, BRA, CEN, COL, GUF, GUY, PRY, PER, SUR, TTO, URY, VEN: (H 06), (P 11). Addl. Data. ES, San Lorenzo: 1 ♀, Concepcion (1.05, - 78.83, 25 m), date?, leg. R. Campos, det. GHB [FSCA]; LR, Buena Fé: 1 ♂, 1 ♀, [Consumulo], S Río Palenque (- 0.58, - 79.35, 175 m), 20 xi 1972, leg. D. Dodson, det.? [FSCA]; 1 ♂, Santo Domingo 47 km S, forest slough at Río Palenque Field Station (200 m), 18 vii 1977, leg. & det. DRP [DRP]; NA, Tena: 1 ♂, Río Chinchipino, oxbow pond, 27 km E Puerto Napo, 13 xi 1997, leg. & det. KJT [KJT]; 1 ♂, J. S. B. S., sendeiro 1 - rego (- 1.0690, - 77.6165, 394 m), 7 xi 2014, leg. & det. ACRP [ACRP]; 1 ♂, same data but, rego 11 (- 1.0686, - 77.6138, 423 m), 15 xii 2014; SU, Shushufindi: 1 ♂, small stream about 3 km NW of Shushufindi (- 0.1554, - 76.6917, 290 m), 14 v 2009, leg. & det. KJT [FSCA].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B56CFF57FF23F3D02495FD83.taxon	discussion	Of the nine species in this genus, five occur in Ecuador. Adults are mainly dark green and black and are relatively slender and small compared to most aeshnids (Fig. 68); they are crepuscular, sometimes flying at treetop level just before dark. Small, temporary, rainforest pools and tree holes are the preferred habitat. References are Williamson (1923 a) and von Ellenrieder & Garrison (2003). Triacanthagyna caribbea Williamson, 1923. Publ. Recs, — ECU: Paulson (2011), (H 14); LR: von Ellenrieder & Garrison (2003). Also BOL, BRA, CEN, COL, GUF, SUR, TTO, VEN: (H 06), (P 11). Addl. Data. LR, Babahoyo: 1 ♂, Playas de Montalvo (- 1.8167, - 79.2833, 15 m), 7 iv 1938, leg. WCM, det. NVE & Garrison, from CHK [UMMZ]; OR, Francisco de Orellana: 1 ♂, Taracoa, 8 i 1985, leg. H. Bohlen, det. NVE [RWG]. Triacanthagyna ditzleri Williamson, 1923. Publ. Recs. ECU: Paulson (2011), (H 14); MS: von Ellenrieder & Garrison (2003); NA: von Ellenrieder & Garrison (2003); PA: von Ellenrieder & Garrison (2003). Also BOL, BRA, CEN, COL, GUY, PER, SUR, TTO, VEN: (H 06), (P 11); Plus GUF, PRY: (H 06). Addl. Data. MS, Gualaquiza: 1 ♀, Bomboiza, 17 km S of Gualaquiza [should be N of Gualaquiza] (- 3.4333, - 78.5167, 880 m), 20 ix 1990, leg. OSF, det. NVE & RWG [USNM]; NA, Archidona: 1 ♀, Río Shicayacu [Parish of Contundo] (- 1.0833, - 78.0689, 900 m), 8 v 1941, leg. WCM, det. NVE & RWG [UMMZ]; Carlos Julio Arsemena Tola: 2 ♀♀, Río Pupuyacu (- 1.2000, - 77.8333, 500 m), 19 – 22 xi 1934, leg. D. Laddey, leg. NVE & RWG [UMMZ]; Tena: 2 ♀♀, Río Jatun Yacu, Ilupulin Creek (- 1.0167, - 77.8333, 700 m), 20 iii 1937, leg. WCM, det. NVE & RWG, from CHK [RWG, UMMZ]; PA, Pastaza: 1 ♀, Puyo (- 1.4667, - 77.9833, 1100 m), xii 1948, leg. WCM, det. NVE & RWG [UMMZ]; SU, Shushufindi: 1 ♂, forest near Río Napo, S of Limoncocha (- 0.4203, - 76.6108, 250 m), 11 v 2009, leg. S. Bybee, S. Ingley & KJT, det. KJT [FSCA]. Triacanthagyna obscuripennis (Blanchard, 1845). Publ. Recs. CR: * Campos (1922); GY: * Campos (1922). Also PER, BOL: (H 06), (P 11); plus CEN: (H 06). Note: von Ellenrieder & Garrison (2003) suggest that “ the apparently disjunct distribution may be due to the species being rare and more though collecting will show it to occur in intermediate localities. ” Triacanthagyna satyrus (Martin, 1909). Publ. Recs. ECU: (H 06), (P 11), (H 14); NA: von Ellenrieder & Garrison (2003); SU: von Ellenrieder & Garrison (2003). Also CEN, GUF, GUY, PER, TTO, VEN: (H 06), (P 11); plus BRA, PRY: (H 06); plus SUR: (P 16); and COL: Bota-Sierra et al. (2018). Addl. Data. NA, Tena: 2 ♂♂, J. S. B. S., Pond # 2 (- 1.0500, - 77.7833, 522 m), 14 xi 1991, leg. E. McCallie, det. NVE & RWG [UMMZ]; OR, Francisco de Orellana: 1 ♂, Reserva Etnica Waorani, Onkore Gare Camp, at night (- 0.6333, - 76.6000, 220 m), 10 i 1994, leg. T. Erwin, leg. NVE & RWG [UMMZ]; SU, Shushufindi: 1 ♀, Limon- cocha, forest trail (- 0.4000, - 76.6167, 269 m), 5 ix 1971, leg. D. Pearson, det. DRP [DRP]; 1 ♀, same data but 14 xii 1971 [DRP]; 1 ♂, 28 xii 1971 [DRP]; 1 ♂, 4 i 1972 [DRP]; 1 ♂, 10 i 1972 [DRP]; 1 ♀, same data but 1 iii 1972 [DRP]; 1 ♂, 1 ♀, same data but 24 – 25 i 1972, det. NVE & RWG [RWG]. Triacanthagyna septima (Selys in Sagra, 1857). Publ. Recs. ECU: Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14); AZ: von Ellenrieder & Garrison (2003); CR: * Campos (1922), Williamson (1923 a); EO: Campos (1927); ES: von Ellenrieder & Garrison (2003); GY: * Campos (1922), Williamson (1923 a), Campos (1931 — as G. septima), von Ellenrieder & Garrison (2003); LJ: von Ellenrieder & Garrison (2003); LR: Williamson (1923 a), von Ellenrieder & Garrison (2003). Also BOL, BRA, CEN, COL, GUF, GUY, PER, SUR, TTO, VEN: (H 06), (P 11). Addl. Data. ES, Esmeraldas: 2 ♂♂, 1 ♀, Esmeraldas, 4 v 1963, leg. T. Hubbell & L. Pena, det. NVE & RWG [RWG, UMMZ]; GY, Naranjal: 1 ♀, Hacienda Balao Chico (- 2.7333, - 79.75, 17 m), 23 iv 1963, leg. T. Hubbell & L. Pena, det. NVE & RWG [UMMZ]; GY, Guayaquil: 2 ♂♂, 3 ♀♀, Guayaquil (- 2.20, - 79.90, 8 m), date?, leg. F. Campos, det.? [USNM]; LJ, Loja: 1 ♀, Malacatos (- 4.2333, - 79.2833, 1500 m), vii 1941, leg. D. Laddey, leg. NVE & RWG [MNRJ]; Saraguro: 1 ♀, Pillangal (- 3.6000, - 79.2667, 2713 m), date?, leg.?, det. NVE [IRSNB]; LR, Babahoyo: 1 ♂, Playas de Montalvo (- 1.8167, - 79.2833, 15 m), 9 iv 1938, leg. WCM, det. NVE & RWG, from C. H. Kennedy [UMMZ].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B592FF57FF23F4802134FCE2.taxon	discussion	This large family is well-represented in the Neotropics (approximately 30 genera, 280 species), although most of the species are secretive and very seldom seen or collected. In general the adults hide in vegetation and are active near the water at unpredictable times. The majority of species are lotic. In addition to 11 genera and 29 named species in Ecuador, there are several unresolved species, a few of which may be undescribed.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B592FF54FF23F5A0272BFD3B.taxon	discussion	Small, slender gomphids occupying small streams in forest. Only one of the four known species has been recorded in Ecuador. There are three additional specimens which have not been determined to species. The only image we could find (Fig. 69) is of a species not reported from Ecuador but that might occur there. References include Belle (1966, 1972, 1975). Agriogomphus sylvicola Selys, 1869. Publ. Recs. ECU: (H 06), (P 11), (H 14); NA: * Tennessen (1998). Also BRA, COL, PER, SUR: (H 06), (P 11). Addl. Data. MS, Huamboya: 1 ♂, stream near Río Pasanac, Hwy. 45 (- 1.9532, - 77.8586), 16 ix 2005 (nymph, emerged 28 ix 2005), leg. & det. KJT [KJT]; NA, Tena: 1 ♂, Río Chinchipino, 27.2 km E of Puerto Napo (- 1.0728, - 77.5864, 385 m), 13 xi 1997, leg. & det. JJD [JJD]; 1 ♀, same data but leg. TWD [SWD]; 1 teneral ♀, 10 x 2009, leg. & det. KJT [KJT]; OR, Francisco de Orellana: 1 ♀, Río Shiripuno Lodge, 27 km SE Río Shiripuno bridge, (- 1.1047, - 76.7317, 235 m), date not provided, data provided by WAH & DLW; teneral ♀, Reserva Etnica Waorani, 1 km S Onkone Gare Camp, (- 0.63, - 76.60), 25 i 1994, leg. T. Irwin et. al. [USNM]; SU, Shushufindi: oil pipeline road, 2 km W of Shushufindi, small tributaries of Río Curiyacu and isolated rain pools. (- 0.1661, - 76.6916), date not provided, data provided by WAH & DLW. Agriogomphus unresolved: OR, Orellana: 1 ♂, T. B. S., Quebrada Harpia (- 0.6350, - 76.1460, 217 m), 6 xii 2012, leg. & det. ACRP [ACRP]; 1 ♀, same data but (- 0.6349, - 76.1463, 248 m), 9 xii 2012; 1 ♀, same data but sendero Harpia (- 0.66351, - 76.1466, 227 m).	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B591FF55FF23F4C8217AF8CF.taxon	discussion	A large genus of large dragonflies (Fig. 70) with at least five species occurring in Ecuador. For species identification see Belle (1970, 1992 b, 1994). Found mostly in mud-bottomed, slow-flowing or stagnant waters. Aphylla boliviana Belle, 1978. Publ. Recs. ECU: Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14); NA: Belle (1992 b); OR: Tennessen (1996 b); PA: Tennessen (1998), von Ellenrieder & Garrison (2011); SU: * Belle (1978 — description of ♂), Dunkle (1981); Belle (1992 b), ZC: Tennessen & Johnson (2008). Also BOL, PER: (H 06), (P 11); plus COL: (P 16). Addl. Data. MS, Santiago: 1 ♀, [Patuca], [about 1 ml N], 43.4 km S of Sucua, large pond [about 5 km N of Mendez road jct] (- 2.75, - 78.27, 488 m), 8 xi 1997, leg. & det. WFM [FSCA]; NA, Tena: 1 ♂, Río Sinde (- 1.0500, - 77.7481, 450 m), 12 xi 1997, leg. & det. TWD [SWD]; OR, Francisco de Orellana: 2 ♂♂, [Pompeya Sur], S on [Pompeya Sur-Iro road] at km marker 31, borrow pit near Río Sabaleto (- 0.60, - 76.47 W, 229 m), 12 vi 1995, leg. & det. WFM [FSCA]; 1 ♂, same data but leg. & det. SWD [SWD]; 2 ♂♂, ditch 20.5 km N Yasuni Sci. Sta., 11 vi 1995, leg. & det. KJT [KJT]; 2 ♂♂, swamp, Yasuni Road, km marker 8 (- 0.4825, - 76.5686, 250 m), 12 vi 1995, leg. & det. KJT [FSCA]; 1 ♂, 1 ♀, marsh 2.1 km E of Yasuni Road, Yasuni Preserve, 15 xi 1997, leg. & det. JJD [JJD]; P. N. Y., 6.2 km W of Yasuni Road on PC 29 - 30, pool / stream, 10 vi 1995, leg. & det. SWD [SWD]; 1 ♂, small lake 11.8 km W of Coca, Loreto Road, 18 xi 1997, leg. & det. JJD [JJD]; Loreto: 1 ♀, pond along Loreto-Archidona road (- 0.7417, - 77.5517, 900 m), 19 xi 1997, leg. & det. TWD [SWD]; PA, Pastaza: 1 ♀, open pond 91.5 km N of Sucua, 9 xi 1997, leg. & det. JJD [JJD]; 2 ♂♂, pond 3 km N of Comunidad Chuwitayo, S of Puyo, 9 xi 1997, leg. & det. KJT [KJT]; 1 ♂, pond on road between Macas and Puyo (- 1.8633, - 77.8150, 900 m), 9 xi 1997, leg. & det. TWD [SWD]; SU, Lago Agrio: 1 ♀, Lago Agrío (300 m), 1 vi 1976, leg. W. Schacht, det. J. Belle [SWD]; Shushufindi: 1 ♂, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 8 xi 1973, leg. B. Drummond III, det. MJW [FSCA]; 1 ♂, same data but 22 viii 1980, leg. & det. SWD [JJD]; 3 ♂♂, same data but 23 – 29 viii 1980, leg. & det. SWD [SWD]; 2 ♂♂, 1 ♀, same data but 2 xi 1980, leg. & det. MJW [FSCA]; 2 ♀♀, same data but 6 xi 1980 [FSCA]; 1 ♀, same data but 12 xi 1980 [FSCA]; 1 ♀, same data but 20 xi 1980 [FSCA]; 14 ♂♂, 3 ♀♀, same data but 21 xi 1980 [FSCA]; 3 ♂♂, 1 ♀, same data but 25 xi 1980 [FSCA]; 1 ♂, 1 ♀, same data but 27 xi 1980 [FSCA]; 1 ♀, same data but forest trail, 1 iii 1972 [DRP]; 1 ♂, same data but 23 vii 1977, leg. & det. DRP; 1 ♂, same data but lake edge, 16 viii 1971, leg. D. Pearson, det. DRP [DRP]; 2 ♂♂, same data but 6 i 1972 [DRP]; 1 ♀, same data but 14 i 1972 [DRP]; 1 ♀, same data but 25 viii 1971 [FSCA]; 2 ♂♂, same data but 26 vii 1977, leg. D. Paulson & S. Hills, det. DRP [DRP]; ZC, Zamora: 1 ♀, tilapia ponds and small streams, Hosteria El Arenal, about 1 km S of Cumbaratza (- 4.0017, - 78.8753, 950 m), 3 iv 2008, leg. & det. JTJ [JTJ]. Notes: There is a specimen collected by a student, labelled IM, Ibarra: 1 ♂, Ibarra 28 i 1989 [QZAC]. Since there are no records for A. boliviana at high elevations, we have not included it as a data record. Interestingly, males have either a red abdomen or a black abdomen. Aphylla dentata Selys, 1859. Publ. Recs. ECU: (H 14 — as A. cf. dentata), NA: * Belle (1992 b). Also ARG, BRA, GUF, GUY, PRY, SUR, VEN: (H 06), (P 11). Addl. Data. OR, Francisco de Orellana: 1 ♀, near Yasuni Scientific Station, Río Tiputini, 39 km SE of Pom- peya, 61 km ESE of Coca (- 0.6741, - 76.3974), date not provided, data provided by WAH & DLW; SU, Shushufindi: 1 ♀, Limoncocha, road through forest, 12 viii 1971, leg. D. Pearson, det. DRP, checked by J. Belle [DRP]; 1 ♀, same data but lake edge, 14 i 1972 [DRP]. Aphylla producta Selys, 1854. Publ. Recs. ECU: (H 14). Also ARG, BRA, GUY, PRY, PER, SUR, VEN, URY: (H 06), (P 11); plus TTO: (P 16). Addl. Data. SU, Lago Agrio: ♂, Santa Cecilia, Río Francisco de Orellana (0.0850 - 76.9925), 25 – 31 iii 1969, leg. P. Spangler, det. TWD [USNM]. Aphylla silvatica Belle, 1992. Publ. Recs. ECU: (H 06), (P 11), (H 14); SU: * Belle (1992 b — Holotype: “ Napo, ” Limoncocha, Río Playaco (- 0.4000, - 76.6167, 269 m), 24 viii 1980, leg. KWK [FSCA]), Dunkle (1993), Bridges (1993). Note: Endemic. Aphylla spinula Belle, 1992. Publ. Recs. ECU: (H 06), (P 11), (H 14); SU: Donnelly (1998). Also PER: (H 06), (P 11). Addl. Data. SU, Shushufindi: 1 ♂, La Selva Lodge; Lake Mandicocha and outlet stream (- 0.4983, - 76.3733, 200 m), 25 xi 1997, leg. & det. TWD [SWD]. Aphylla unresolved: There are several specimens yet to be specifically determined that may represent undescribed species: SU: * Belle (1992 b) listed a specimen as “ spec indet, near boliviana ”); “ Species E, ” Tiputini Biodiversity Station, oxbow laguna, 21 i 2012, Tim Kell, Vicki Liu, William Haber, David Wagner leg., det. WAH; Lago Agrio: 1 ♀, “ Napo Prov., ” Lago Agrio, 24 vii 1975, leg. A. Langley & J. Cohen, det.? [USNM].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B597FF52FF23F68825E3FB36.taxon	discussion	Small brownish gomphids (Fig. 71) rarely seen at water. Nymphs have been collected in Ecuador and one ♂ was reared but the species has not been identified. The unusual nymphs bury themselves in the shallow edges of small, sandy forest streams. Archaeogomphus unresolved: NA, Tena: 1 ♂, reared, Río Sinde, E of Puerto Napo, 12 xi 1997, leg. & det. KJT [KJT].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B597FF52FF23F2C125D8F98C.taxon	discussion	Only three species of Desmogomphus have been described, one of which was reported from a small sandy, gravel, clear-water stream (Fig. 72). Descriptions can be found in Williamson (1920) and Belle (1970, 1972). Desmogomphus paucinervis (Selys, 1873). Publ. Recs. ECU: Tsuda (1986), Steinmann (1997 b); PC: * Selys (1873 b — Type locality, “ Quito ”), Hagen (1875), Campos (1922 — “ from Kirby ”). Publ. Recs. ECU: (H 14). Also CEN, COL (P 16). Addl. Data. ES, Quininde: Nymphs collected at Río Canande Reserve, 35 km NE of Quininde, tributary of Río Hoja Blanca, swampy trickle in forest (0.5264, - 79.2122, 345 m), date not provided, data provided by WAH & DLW.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B597FF50FF23F0662017FBAF.taxon	discussion	The taxonomy of this genus is in need of synoptic update as reference to scattered papers is necessary for identification (Calvert 1920; Belle 1970, 1988 b, 1996; Donnelly 1989); at least four species are known from Ecuador (Fig. 73), but much more work is needed to clarify the status of others. Habitat includes small streams in forested areas. Epigomphus hylaeus Ris, 1918. Publ. Recs. ECU: * Tsuda (1986, 1991), (H 06), (H 11). Also BRA, GUF, PER, SUR: (H 06), (P 11). Addl. Data. ZC, Zamora: 1 ♂, 1 ♀, Zumbi, Río Zamora (- 3.3889, - 78.7752, 700 m), 1 x 1941, leg. D. Laddey, det. JJD [FSCA]; 1 ♂, same data but 3 x 1941 [FSCA]; 1 ♀, same data but 10 x 1941 [FSCA]; 1 ♂, same data but 12 x 1941 [FSCA]; 2 ♀♀, same data but 29 x 1941 [FSCA]; 2 ♀♀, same data but 31 x 1941 [FSCA]. Note: This species has not been collected in Ecuador in over 75 years. Epigomphus obtusus Selys, 1869. Publ. Recs. ECU: * Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14); MS: Tennessen (1998) Also BRA, COL, PER: (H 06), (P 11). Addl. Data. MS, Huamboya: 7 ♂♂, 3 ♀♀, stream 10 km S of Comunidad Chuwitayo (- 1.9500, - 77.8583, 800 m), 9 xi 1997, leg. & det. TWD [KJT, SWD]; 2 ♂♂, same but leg. & det. JJD [JJD]; 5 ♂♂, 1 ♀, same data but 10 xi 1997, leg. & det. KJT [KJT]; 1 ♂, Macas, 58 km N on Hwy. 45 (70 km N of Sucua), roadside ditch and tributaries near Río Pasanac (- 1.9447, - 77.8536, 762 m), 9 xi 1997, leg. & det. WFM [FSCA]; 1 ♂, same data but 10 xi 1997 [LSAM]; 1 ♀, same data but 16 ix 2005 [FSCA]; 1 ♂, small stream about 1 km S of Río Pasanac, Hwy. 45 (- 1.9500, - 77.8583, 820 m), 16 ix 2005, leg. & det. KJT [KJT]; NA, Tena: 1 ♂, J. S. B. S. (- 1.0655, - 77.6176, 421 m), leg. & det. ACRP [ACRP]; 1 ♂, same data but RA 4 - 5 (- 1.0722, - 77.6144, 434 m); SU, Shushufindi: 1 ♂, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 1 xi 1980, leg. & det. MJW [FSCA]; 1 ♂, same data but lake edge, 14 i 1972, leg. D. Pearson, det. DRP [DRP]: 2 ♂♂, same data but forest stream, 24 vii 1977, leg. & det. DRP [DRP]. * Epigomphus occipitalis Belle, 1970. Publ. Recs. ECU: (H 14). Also PER: (H 06), (P 11). Addl. Data. SU, Putumayo: 2 ♂♂, Río Cuyabeno, 60 km E Lago Agrío (- 0.0167, - 76.3167), 16 – 26 ii 1998, leg. G. Kareofelas, det. A. Rehn, checked by RWG [RWG]; Shushufindi: 2 ♂♂, trails, small streams 3 km W of Shushufindi, 2 + km N off Hwy. 26 at km marker 17 (- 0.1664, - 76.6917, 290 m), 13 x 2009, leg. & det. JTJ [JTJ]; 1 ♀, oil pipeline road, 2 km W of Shushufindi, small tributaries of Río Curiyacu and isolated rain pools (- 0.1661, - 76.6916), date not provided, data provided by WAH & DLW; 1 ♂, La Selva Lodge, Río Napo, 70 km E of Coca, in forest on Tower Trail. (- 0.4978, - 76.3741), date not provided, data provided by WAH & DLW. Epigomphus unresolved: There are several specimens in various collections that have not been determined and may represent undescribed species: ES, San Lorenzo: 1 ♀, along RR track, about 2 km N of Lita, near Río Mira (0.8847, - 78.4667, 500 m), 5 ii 1997, leg. and det. KJT [FSCA]; MN, Junin: 1 ♀, Palmar (- 0.17, - 79.47), 6 iv 1941, leg. D. B. Laddey [FSCA]; OR, Francisco de Orellana: 1 ♀, P. N. Y., 1.8 km W of Waorani Reserve Rd on Markus Oil Co Rd PC 29 - 30, small stream (- 0.6861, - 76.4726, 200 m), 10 vi 1995, leg. and det. WFM [FSCA].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B595FF51FF23F27427CEFF46.taxon	discussion	Small rare gomphids (Fig. 74), with one of the three known species recorded so far in Ecuador (Tennessen 2011). Habitat is small, shallow forest streams. Perigomphus angularis Tennessen, 2011. Publ. Recs. MS: * Tennessen 2011 — Holotype ♂: Huamboya: small stream 1 km S of Río Pasanac, Hwy. 45 (- 1.9500, - 77.8583, 820 m), 16 x 2005, leg. KJT [FSCA]). Note: Known only from this single collection.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B594FF51FF23F75C275DF8F1.taxon	discussion	Two of the three known species occur in Ecuador; a reared female specimen of an indeterminable species is in the FSCA (Belle 1992 a). For species identification, see Belle (1979) and Machado (2005). These medium-sized gom- phids (Fig. 75) occupy small lakes and ponds with muddy bottoms and are infrequently collected. Peruviogomphus moyobambus Klots, 1994. Publ. Recs. ECU: (H 14). NA: * Belle (1992 a) mentions a specimen he collected near the junction of the Río Napo and Río Arajuno in the discussion of the “ P. spec. indet. ” Listed only for PER: (H 06), (P 11). Addl. Data. NA, Tena: J. S. B. S., Pond # 16 A, 16 xi 1991, leg. B. Sharp, det. SWD [USNM]; 1 ♂, road to air- port and La Punta, E of Tena, shallow pond with lily pads and rocky stream outflow (- 1.0482, - 77.5861), date not provided, data provided by WAH & DLW; OR, Francisco de Orellana: 1 ♂, Río Shiripuno Lodge, 27 km SE Río Shiripuno bridge (- 1.1047, - 76.7317, 235 m), date not provided, data provided by WAH & DLW. Peruviogomphus pearsoni Belle, 1979. Publ. Recs. ECU: Paulson (1983), Davies & Tobin (1984), Tsuda (1986, 1991), Steinmann (1997 b), (H 06), (P 11), (H 14); SU: * Belle (1979 — description, as NA: Limoncocha, leg. DRP [FSCA]); Bridges (1993). Addl. Data. SU, Shushufindi: Limoncocha, lake edge (- 0.4000, - 76.6167, 269 m), 2 x 1971, leg. D. Pearson, det. DRP [DRP]; 1 ♀, same data but 22 viii 1971 [DRP]; 1 ♂, same data but 25 viii 1971 [DRP]; 1 ♂, [Limoncocha] [about 40 km E] or 100 km E of Coca [on Río Napo] Lago Garzacocha, La Selva Jungle Lodge (0.4819, - 76.3733, 235 m), 12 x 1988, leg. & det. SWD [SWD]. Note: Possibly an Ecuador endemic.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B594FF5FFF23F18E25EAFC53.taxon	discussion	A species-rich genus in the Neotropics, although only three species have been found in Ecuador (Fig. 76). The major reference for identification is Belle (1988 a). The nymphs live in slow portions of small to mid-sized streams where silt and sand have accumulated. Phyllocycla anduzei (Needham, 1943). Publ. Recs. ECU: * Dunkle (1981), (H 06), (P 11), (H 14); NA: Tennessen (1996 b); PA: Belle (1988 a). Also BOL, COL, PER, VEN, TTO: (H 06), (P 11). Addl. Data. NA, Tena: 1 ♂, Puerto Napo, 5.3 km E, Río Sinde on J. S. B. S. Road (- 1.0500, - 77.7481, 450 m), 15 vi 1995, leg. & det. SWD [DRP]; 2 ♂♂, same data but leg. & det. KJT [KJT]; 2 ♂♂, 1 ♀, Puerto Napo 11.2 km E of Hwy. 45 on J. S. B. S., Río Tiyuyacu (- 1.0500, - 77.7086, 420 m), 15 vi 1995, leg. & det. JJD [JJD]; 5 ♂♂, same data but leg. & det. KJT [KJT, RWG]; 4 ♂♂, same data but leg. & det. WFM [FSCA, QZAC, USNM]; 7 ♂♂, same data but leg. & det. SWD [RWG, SWD]; 1 ♀, Río Chinchipino, 27 km E of Puerto Napo (- 1.0728, - 77.5864, 385 m), 13 xi 1997, leg. & det. KJT [KJT]; SU, Lago Agrio: 1 ♂, Santa Cecilia, 24 km W of Lago Agrío on road to Baeza (0.12, - 77.07, 335 m), 20 viii 1980, leg. & det. SWD [SWD]; Shushufindi: 1 cop. pair, Limoncocha, forest edge, 26 vii 1977, leg. & det. DRP [DRP]; 1 ♂, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 24 xi 1980, leg. & det. MJW [FSCA]. Phyllocycla hespera (Calvert, 1909). Publ. Recs. ECU: Saint Quentin (1967), Paulson (1983 — as Aphylla hespera), Davies & Tobin (1984), Tsuda (1986, 1991), Steinmann (1997 b — as A. hespera), (H 06), (P 11), (H 14); GY: Tennessen & Johnson (2008); LR: * Calvert (1909 — as G. hesperus “ quevedo ”, Holotype [ANSP], Campos (1922 — as G. hesperus), Bridges (1993 — as G. hesperus), Belle (1988 a). Addl. Data. GY, Balao: 1 ♂, stream, banana plantation, 2 km E of Hwy. 25 & Balao jct. (- 2.9100, - 79.6919, 70 m), 28 iii 2008, leg. & det. KJT [KJT]; 2 ♂♂, same data but leg. & det. JTJ [JTJ]; LR, Babahoyo: 1 ♂, Playas de Juan Montalvo near Balzapamba, W of Andes (- 1.8167, - 79.2833, 15 m), 19 iv 1938, leg. WCM, det. J. Belle 1981, from UMMZ [RWG]; Buena Fé, 1 ♂, [Consumulo], S Río Palenque (- 0.58, - 79.35, 175 m), 11 vi 1974, leg. J. Longino, det.? [FSCA]; Quevedo: 1 ♂, 2 ♀♀, stream 16 km E of Hwy. E 25 from Formisa, 23 ii 2010, leg. & det. JJD [JJD]. Note: An Ecuador endemic. Phyllocycla malkini Belle, 1970. Publ. Recs. ECU: * Dunkle (1981), Tsuda (1991), (H 06), (P 11), (H 14); SU: Belle (1988 a). Also BRA, PER: (H 06), (P 11). Addl. Data. OR, La Joya de los Sachas: 1 ♂, Limoncocha [just W of village], Río Playaco (- 0.42, - 76.63, 300 m), 22 viii 1980, leg. & det. SWD [SWD]; SU, Lago Agrio: 1 ♂, [Santa Cecillia], 24 km W of Lago Agrío on road to Baeza, 20 viii 1980, leg. & det. SWD [SWD]; Shushufindi: 5 ♂♂, 1 ♀, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 10 xi 1980, leg. & det. MJW [FSCA]; 6 ♂♂, same data but 17 xi 1980 [FSCA]; 1 ♂, same data but forest stream, 6 iii 1972, leg. D. Pearson, det. DRP [DRP]; 1 ♂, 1 ♀, same data but 25 vii 1977, leg. & det. DRP (DRP).	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B59AFF5CFF23F250242CF854.taxon	discussion	Another species-rich Neotropical genus, with five species recorded for Ecuador (Fig. 77). See Belle (1984) for species identification. Habitat is small to large streams, open and forested. * Phyllogomphoides aculeus Belle, 1982. Publ. Recs. ECU: (H 14). Also PER: (P 16). Addl. Data. SU: Lago Agrio: 1 ♀, drainage of Río Aguarico, 33 km ESE of Lago Agrio (0.0006, - 76.6004), date not provided, data provided by WAH & DLW. Phyllogomphoides brunneus Belle, 1981. Publ. Recs. ECU: Paulson (1983), Davies & Tobin (1984), Tsuda (1986, 1991), Bridges (1993), Steinmann (1997 b), De Marmels (1999), (H 06), (P 11), (H 14); MS: * Belle (1981 — Holotype: [Morona], Macas, date?, leg.? [FSCA]; NA: Belle (1984); PA: Belle (1984), Tennessen (2006); SU * Belle (1981 — Allotype: [Shushufindi], Limoncocha, open path with puddles in forest, 16 ii 1972, leg. D. Pearson [DRP]; ZC: Belle (1984). Also VEN: (H 06), (P 11); plus PER: (H 06); plus COL: Bota-Sierra et al. (2018). Addl. Data. MS, Morona: Gualaquiza: 1 ♂, Bomboiza, 17 km S Gualaquiza, 20 ix 1990, leg. OSF, det.? [USNM]; 1 ♂, Macas Oriente, Acc. 788 (- 2.30, - 78.12, 1006 m), date?, leg.?, det.? [FSCA]; OR, Loreto: 1 ♂, small stream, near Reserva Río Bigal, about 30 km N of Loreto (- 0.5642, - 77.4117, 900 m), 15 v 2009, leg. & det. KJT [FSCA]; 1 ♂, stream about 12 km NE of Loreto (- 0.6217, - 77.2600, 365 m), 12 x 2009, leg. & det. KJT [FSCA]; 1 ♂, same data but 15 x 2009 [FSCA]; 1 ♂, NE of Loreto, streams, (- 0.6220 - 77.2596, 384 m), 26 x 2013, leg. & det. CDK [FSCA]; 1 ♂, S of Loreto, stream, (- 0.8141, - 77.3371, 305 m), 3 xi 2013, leg. & det. CDK [FSCA]; PA, Mera: 1 ♂, Río Metolo, Los Copales cabanas between Mera and Shell, W of Puyo, Hwy. 50 (- 1.4919, - 78.0725, 1070 m), 20 ix 2005, leg. & det. JJD [JJD]; SU, Lago Agrio: 1 ♂, Santa Cecillia 24 km W of Lago Agrío on road to Baeza, stream (- 0.12, - 77.07, 335 m), 20 viii 1980, leg. KWK, det.? [FSCA]; Shushufindi: 5 ♀♀, with exuviae, 1 ♂, with exuvia, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 29 viii 1978, leg. KWK, det. MJW [FSCA]; 1 ♀, same data but 5 xi 1980, leg. MJW & D. Robinson, det. MJW [FSCA]; 1 ♀, same data but 12 xi 1980 [FSCA]; 2 ♂♂, same data but 15 xi 1980 [FSCA]; 1 ♀, same data but 17 xi 1980 [FSCA]; 1 ♀, same data but forest path, 28 i 1972, leg. D. Pearson, det. D. Paulson [DRP]; 1 ♀, same data but puddles on path in forest, 16 ii 1972 [DRP]; 1 ♀, same data but forest stream, 25 vii 1977, leg. & det. DRP [DRP]; 1 ♀, same data but forest edge, 26 vii 1977 [DRP]; 2 ♀♀, same data but 22 viii 1980, leg. & det. SWD [SWD]; 1 ♂, stream about 3 km NW of Shushufindi (- 0.1664, - 76.6917, 290 m), 13 v 2009, leg. & det. KJT [KJT]. Phyllogomphoides camposi (Calvert, 1909). Publ. Recs. ECU: Saint Quentin (1967 — as Gomphoides camposi), Paulson (1983), Davies & Tobin (1984), Tsuda (1986, 1991), Steinmann (1997 b), (H 06), (P 11), (H 14); LR: * Calvert (1909 — HOLOYPE ♂: [Quevedo] [about 70 m], Quevedo, vi, leg. F. Campos [ANSP], Campion (1920), Campos (1922), Bridges (1993). Note: This species is named after F. Campos, who we consider the “ father of Ecuadorian Odonata studies. ” Campos (1922) listed the date as June (vi), but Calvert (1909) did not give the date. Phyllogomphoides lieftincki (Belle, 1970). Publ. Recs. ECU: * Tsuda (1986, 1991). (H 06). Also BOL: (H 06), (P 11); plus PER: (H 11). Data. NA: Tena: 1 ♂, 1 ♀, road to airport and La Punta, E of Tena, shallow pond with lily pads and rocky stream outflow (- 1.0482, - 77.5861), date not provided, data provided by WAH & DLW (— as P. cf. lieftincki); OR, Loreto: 1 ♂, about 2 km E of Loreto center; 2 – 3 m wide shallow rocky river; about 250 m to forest upstream (- 0.6756, - 77.3101), date not provided, data provided by WAH & DLW (— as P. cf. lieftincki); SU, Shushufindi: 1 ♀, [as “ Napo Prov], Limoncocha, 5 vi 1977, leg. W. E. Steiner, Jr., det. RWG [USNM]; 1 ♂, same except 3 – 16 vi 1977, leg. P. J. Spangler and D. R. Givens, det. RWG [USNM]. Phyllogomphoides semicircularis (Selys, 1854). Publ. Recs. none. Also COL, VEN: (H 06). Addl. Data. * SU, Shushufindi: 1 ♂, Shushufindi, SW, forest edge (- 0.2201, - 76.6988), 29 x 2013, leg. & det. KJT [FSCA].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B598FF5AFF23F6882776F95F.taxon	discussion	The largest genus of Gomphidae with 68 described species, at least seven of which are found in Ecuador. The genus is found only in the New World. Basic references are by Belle (1973, 1991 b); new species are likely to be found. Nymphs of these small to medium-sized gomphids bury themselves in sand and silt of mainly small streams to large rivers. The image (Fig. 78) from Eastern Ecuador is of uncertain specific identification. Progomphus abbreviatus Belle, 1973. Publ. Recs. ECU: * Belle (1983 a), (H 06), (H 14). Also COL, VEN: (H 06), (P 11). Addl. Data. IM, Ibarra: reared ♂, stream 45.8 km E of Lita, 6 ii 1997, leg. & det. KJT (KJT); NA, Quijos: 2 ♂♂, small stream, 5 km S of Baeza (- 0.4758, - 77.9714, 1825 m), 20 xi 1997, leg. & det. KJT [FSCA]; 1 ♂, Yanayacu Biological Station, 3.5 km SW of Cosanga, small forest stream (- 0.5804, - 77.8654), reared adult on 11 i 2011, data provided by WAH & DLW; PC, San Miguel Bancos: 1 ♂, Reserva Gralarias, 7 km NE of Mindo, small forest stream (- 0.0106, - 78.7307), date not provided, data provided by WAH & DLW. Progomphus anomalus Belle, 1973. Publ. Recs. ECU: Paulson (1983), Tsuda (1986, 1991), Steinmann (1997 b), (H 06), (P 11), (H 14); MS: * Belle (1973); NA * Belle (1973); PA * Belle (1973 — Holotype ♂: [Mera], Río Pastaza watershed, Abitagua (1000 m), 28 x 1036, leg. WCM [UMMZ]; Bridges (1993), Garrison et al. (2003). Also CEN, PER: (H 06), (P 11). Progomphus formalis Belle, 1973. Publ. Recs. ECU: Paulson (1983), Davies & Tobin (1984), Tsuda (1986, 1991), Steinmann (1997 b), (H 06), (P 11), (H 14); NA: * Belle (1973 — Holotype: Jatun Yacu (700 m), 12 iv 1935, leg. WCM [UMMZ]), Bridges (1993), Tennessen (1998). Also PER: (H 06), (P 011). Addl. Data. NA, Tena: 1 ♂, Río Chinchipino, 27 km E of Puerto Napo (- 1.0728, - 77.5864, 385 m), 13 xi 1997, leg. & det. KJT [KJT]; 1 ♂, same data but leg. & det. TWD [SWD]; OR, Francisco de Orellana: 1 ♂, Via Auca, 8 km SE of Coca, oil platform access road, tiny forest trickle (- 0.4991, - 76.9169), date not provided, data provided by WAH & DLW; SU, Shushufindi: 1 ♀, oil pipeline road, 2 km W of Shushufindi, small tributaries of Río Curiyacu and isolated rain pools (- 0.1661, - 76.6916), date not provided, data provided by WAH & DLW. Progomphus phyllochromus Ris, 1918. Publ. Recs. ECU: Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14); MN: * Belle (1973); MS: * Belle (1973); PA: * Belle (1973); ZC: * Belle (1973). Also ARG, BOL, COL, PER, VEN: (H 06), (P 11). Addl. Data. MN, Pedernales: 1 ♀, Cojimies (0.3667, - 80.0333), 9 iii 1949, leg. WCM, det. J. Belle, data from RWG [UMMZ]; MS, Morona: 1 ♀, Macas, Río Upano (- 2.3167, - 78.1167, 1050 m), date?, leg. L. Alonzo, det. J. Belle, data provided by RWG [UMMZ]; PA, Mera: 1 ♀, Abitagua (- 1.45, - 78.15, 1000 m), 7 xi 1936, leg. WCM, det. J. Belle, data provided by RWG [UMMZ]; Pastaza: 1 ♂, Puyo, 5 v 1977, leg. P. Spangler & D. Givens, det.? [USNM]; SD: 1 ♂, forest stream at Hotel Tinalandia, 7.3 km W of Alluriquin (- 0.2750, - 79.0833, 670 m), 20 vii 1997, leg. & det. DRP [DRP]; ZC, Zamora, 1 ♀, Zamora (- 4.0692, - 78.9567, 1000 m), 15 i 1941, leg. D. Laddey, det. J. Belle, data provided by RWG [UMMZ]. Progomphus pijpersi Belle, 1966. Publ. Recs. ECU: * Belle (1973), Tsuda (1986, 1991), (H 06), (P 11), (H 14). Also BRA, SUR, VEN: (H 06), (P 11). Addl. Data. NA, Tena: 1 ♀, Jatunyacu, Río Napo watershed, juvenile (- 1.0167, - 77.8333, 700 m), 21 iii 1937, leg. WCM, det. J. Belle, data provided by RWG [UMMZ]. Progomphus pygmaeus Selys, 1873. Publ. Recs. ECU: * Campos (1922 — “ por Dr. Martin), Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14); LR: Belle (1973); EO: Belle (1973); NA: Daigle (1996). Also BOL, BRA, CEN, COL, GUY, PER, VEN: (H 06), (P 11). Addl. Data. BO, Caluma: 1 ♀, rocky stream 5 km E of Caluma, Guaranda Road, 5 iii 2010, leg. & det. JJD [JJD]; EO, Piñas: 1 ♂, Buenaventura, Riachuelo 20 (- 3.6535, - 79.1466, 836 m), 29 vi 2016, leg. & det. ACRP [ACRP]; Portovelo: 1 ♂, Portovelo (- 3.7167, - 79.6167, 650 m), 1 viii 1941, leg. D. Laddey, det. J. Belle, data pro- vided by RWG [UMMZ]); LR, Babahoyo: 1 ♀, Playas de Juan Montalvo, near Balzapamba (- 1.6833, - 79.2833, 15 m), 6 iv 1938, leg.?, det. J. Belle, data provided by RWG [UMMZ]); Buena Fé: 1 ♀, [Consumulo], S Río Palen- que (- 0.58, - 79.35, 175 m), 11 vi 1974, leg. J. Longino, det. GHB [FSCA]; NA, Tena: 1 ♂, Río Sinde 5.3 km E of Puerto Napo (- 1.0500, - 77.7481, 450 m), 13 vii 1996, leg. & det. JJD [JJD]; 2 ♂♂, same data but leg. & det. WFM [FSCA]; 1 ♂, same data but 22 vii 1996 [FSCA]; 2 ♂♂, same data but leg. & det. JJD [JJD]; 1 ♂, Río Chinchipino, 27 km E of Puerto Napo (- 1.0728, - 77.5864, 385 m), 13 xi 1997, leg. & det. KJT [KJT]; SU, Shushufindi: 1 ♂, Li- moncocha, forest stream (- 0.4000, - 76.6167, 269 m), 18 xii 1971, leg. D. Pearson, det. DRP [DRP]; 1 ♂, same data but 6 iii 1972 [DRP]; 1 ♂, 1 ♀, stream about 3 km SW of Shushufindi (- 0.2136, - 76.6786, 270 m), 12 v 2009, leg. & det. KJT [KJT]. Progomphus suberbus Belle, 1973. Publ. Recs. ECU: Paulson (1983), Davies & Tobin (1984), Tsuda (1986, 1991), Steinmann (1997 b), (H 06), (P 11); LR: * Belle (1973 — Holotype ♂: (15 m), 4 iii 1938, leg. WCM [UMMZ]), Bridges (1993), Garrison et al. (2003). Also VEN: (H 06), (P 11). Note: All records so far refer to the type specimen. The allotype is from VEN. Progomphus unresolved: There are specimens in various collections that have not been determined and may represent undescribed species: GY, Balao: 1 ♀, small stream / ditches in banana plantation, about 2 km E of Hwy. 25 near Balao Jct., (- 2.9100, - 79.6919, 70 m), 28 iii 2008, leg. and det. JTJ [JTJ]; OR, Loreto: 1 ♂, 2 ♀♀, small streams and forest trails 12 km E of Loreto, 2 + km S of Hwy. 45 (- 0.6217, - 77.2600, 365 m), 12 x 2009, leg. & det. JTJ [JTJ]; 1 ♂, same data but 15 x 2009; PC, San Miguel: 1 ♀, Bancos, between Calacali and Nanegalito (about 30 km W Quito), 9 iv 1990, leg. OSF, det. SWD [USNM]; SU, Shushufindi: 1 ♂, 1 ♀, small streams / trails 3 km W of Shushufindi, 2 + km N off Hwy. 26 at km marker 17 (- 0.1664, - 76.6917, 290 m), 14 v 2009, leg. and det. KJT [KJT]; 1 ♂, same data but 14 x 2009, leg. and det. JTJ [JTJ].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B59FFF5BFF23F1242770FA7C.taxon	discussion	Only one of the ten known species in this genus (Fig. 79) has been recorded for Ecuador. See Belle (1963, 1983 b) for identification. These large gomphids are inhabitants mainly of forested streams. Zonophora calippus Selys, 1869. Publ. Recs. ECU: Tsuda (1991 — as ssp klugi), (H 14); NA: * Belle (1983 b — as ssp klugi). Also BOL, BRA, GUF, GUY, PRY, PER, VEN, SUR: (H 06), (P 11). Addl. Data. SU, Lago Agrio: 1 ♀, drainage of Río Aguarico, 33 km ESE of Lago Agrio (0.0006, - 76.6004), date not provided, data provided by WAH & DLW; Shushufindi: 1 ♂, stream about 3 km SW of Shushufindi (- 0.2136, - 76.6786, 250 m), 12 v 2009, leg. & det. KJT [FSCA]; 1 ♂, small streams / trails 3 km W of Shushufindi, 2 + km N off Hwy. 26 at km marker 17 (- 0.1664, - 76.6917, 290 m), 14 v 2009, leg. & det. KJT [FSCA]; 1 ♂, 1 ♀, oil pipeline road, 2 km W of Shushufindi, small tributaries of Río Curiyacu and isolated rain pools (- 0.1661, - 76.6916), date not provided, data provided by WAH & DLW.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B59EFF5BFF23F00124C8F924.taxon	discussion	The majority of New World Corduliidae occur in North America. Only two of the four South American genera occur in Ecuador (Paulson 2016).	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B59EFF58FF23F0F8245FFE37.taxon	discussion	A group of 12 species that occurs mostly in eastern South America, with one unidentified species (Fig. 80) collected in eastern Ecuador at moderate elevation. The basic reference is Machado & Costa (1995). The unknown species is probably a stream-dweller. Navicordulia sp. PA, Mera: 1 ♀, small streams / trails, Los Copales, between Mera and Shell (- 1.4919, - 78.0725, 1070 m), 6 x 2009, leg. & det. KJT, also examined by RWG [KJT]. Note: This specimen remains specifically undetermined and possibly represents an undescribed species; there are no previous records of Navicordulia from northwestern or western South America. The single female was collected in late afternoon in a small clearing; it flew incredibly fast and erratically. However, uncertainty still exists for a number of genera tentatively placed in the family Synthemistidae (Paulson & Schorr (2019), including the two following genera.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B59DFF58FF23F3A02189F8B3.taxon	discussion	This is a monotypic genus at present, the only described species to date being P. sericea (Selys, 1871) which ranges from Bolivia east to Venezuela and Brazil. For identification, see Geijskes (1970). The only reference to the genus in Ecuador appears to be based on an undescribed species (Fig. 81). This is a significant northwest extension of the range for this genus. Paracordulia sp. “ A ” Publ. Recs. ECU: (H 14). Addl. Data. SU, Cuyabeno: 1 ♂, 10 km NE of Tarapoa, road crossing at small forest stream (- 0.044, - 76.230, 235 m), 12 i 2014, leg. M. Veit, E. Wyman, WAH, DLW, & F. Morrison, det. WAH [WAH]: Haber (pers. comm. 2016) reports this new undescribed species with appendages quite distinct from P. sericea.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B59CFF59FF23F38426A9F9A6.taxon	discussion	Synthemistidae is mainly an Old World family that has recently been expanded to include three genera of Neotropical dragonflies previously placed in Corduliidae (Dijkstra et al. 2013).	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B59CFF47FF23F0572770FF6B.taxon	discussion	For species identification see von Ellenrieder & Garrison (2005). Only one species (Fig. 82) has been recorded for Ecuador with certainty, but a second, undescribed species has been discovered. Habitats appear to be restricted to mountain seepage streams and bogs; nymphs of some species may be semiaquatic, hiding under rocks or moss. Gomphomacromia fallax McLachlan, 1881. Publ. Recs. ECU: Selys (1882), Kirby (1890), Martin (1907, 1914), Campos (1922), Cowley (1934), Schmidt (1942), Paulson (1983), Davies & Tobin (1984), Tsuda (1986, 1991), von Ellenrieder & Garrison (2005), Steinmann (1997 b), (H 06), (P 11), (H 14); IM: * McLachlan (1881 — type series 3 ♂♂, 2 ♀♀: [Cotacachi] “ Intag ” [Apuela, NE on Apuela-Cuicocha Road, along Río Intag and tributaries between 1800 – 2800 m at jct of Río Intag and tributary (0.3588, - 78.4509), 1 x 1939, leg. Clarence Buckley, data provided by RWG [BMNH]); Kimmins (1969 a — established Lectotype), Bridges (1993); MS: Tennessen (2006); PA: von Ellenrieder & Garrison (2005). Also ARG, BOL, PER: (H 06), (P 11); plus COL: (P 16). Addl. Data. IM, Cotacachi: 1 ♂, Santa Rosa, along road south of El Refugio, roadside seeps & small open stream (0.3703, - 78.4745, 1866 m), 23 viii 2018, leg. & det. KJT [KJT]; 2 ♂♂, north of El Rosario, small trib and seepage to Rio Cristopamba (0.4880, - 78.5080, 2387 m), 24 viii 2018, leg. & det. WFM [FSCA, USFQ]; same data but 1 ♂, leg. & det. KJT [KJT]; MS, Morona: 1 ♂, near rocky waterfall stream 20.5 km W of Macas (- 2.2475, - 78.2067, 1535 m), 18 ix 2005, leg. & det. KJT [KJT]; PA, Mera: 1 ♂, Abitagua, Río Pastaza watershed (- 1.45, - 78.15, 1000 m), 11 v 1939, leg. WCM, det. M. May, data provided by RWG [RWG]; SD: 1 ♂, Tinalandia, 12 km E of Santo Domingo de los Colorados, SWD [SWD]; ZC, Zamora: Estacion scientifica San Francisco, Copalinga yellow trail (1100 – 1700 m), leg. & det. C. Willigalla.. Gomphomacromia sp. NA, Quijos: 1 ♂, hillside seep, about 5.5 km E of Papallacta, Rt. 28 (- 0.3739, - 78.0779, 2730 m), 5 xi 2013, leg. & det. KJT [KJT].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B582FF47FF23F70D2194FD1C.taxon	discussion	A genus of 12 species, one of which occurs in Ecuador (Fig. 83). See May (1992) for identification. The nymphs are found in small, clear mountainous streams. Neocordulia batesi (Selys, 1871). Publ. Recs. ECU: * Martin (1906, 1909), Campos (1922), Paulson (1983 — also as Neocordulia longipollex), Tsuda (1986, 1991 — also as N. longipollex), Steinmann (1997 b), Garrison (2006 — as N. longipollex as a ssp. of N. batesi). (H 06), (P 11), (H 14); PA: * McLachlan (1881 — as G. batesi); NA: May (1992); PA: May (1992); SD: * May (1992 — as PC). Also BRA, CEN: (H 06), (P 11); plus COL, PER: (H 06). Addl. Data. OR, Loreto: 1 ♂, forest stream about 12 km NE of Loreto (- 0.6217, - 77.2600, 365 m), 12 x 2009, leg. & det. KJT [FSCA]; same data but 1 ♂, 26 x 2013, leg. & det. KJT [FSCA]; PA, Mera: Río Metolo, Los Copales, between Mera and Shell (- 1.4919, - 78.0725, 1070 m), 5 x 2009, leg. & det. JTJ [JTJ].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B581FF45FF23F7A92467FEDB.taxon	discussion	Anatya are medium-sized dragonflies with blue eyes and pale blue thoracic sides (Fig. 84) that occupy open and shaded ponds. There are only two known species, one of which is found in Ecuador. For species identification see De Marmels (1992). Anatya guttata (Erichson, 1848). Publ. Recs. ECU: Heckman (2006), Paulson (2011), (H 14); GY * Campos (1922); NA: Tennessen (1998). Also BOL, BRA, GUF, GUY, PER, SUR, TTO, VEN: (H 06), (P 11). Plus CEN, COL: (P 11). Addl. Data. ES, Quininde: 1 ♀, Cuppa (300 m), 23 i 1982, leg. & det.? [QZAC]; GY, Naranjal: 1 ♀, Finca de la Gringa, 15 km E of Tres Cerritos (- 2.4456, - 79.5556, 25 m), 12 iv 2008. leg. & det. KJT [KJT]; LR, Buena Fé: 1 ♀, [Patricia Pilar] just S at Reserva Palenque [about 47 km S of Santo Domingo] (- 0.66, - 79.26, 107 m), 9 x 1988, leg. R. Miller & L. Stange, det. WFM [FSCA]; 2 ♂♂, same data but 8 x 1980, leg. & det. SWD [SWD]; Montalvo: 1 ♂, small stream 1.8 km N of Hwy. 491, about 25 km E of Babahoyo (- 1.8111, - 79.3189, 60 m), 7 iii 2010, leg. & det. JTJ [JTJ]; Quevedo: 1 ♂, 1 ♀, Río Palenque, 12 xi 1981, leg. & det.? [QZAC]; MN, El Carmen: 1 ♂, El Carmen, 22 i 1983, leg. & det.? [QZAC]; 1 ♂, borrow pits about 18 km NW of El Carmen (- 0.1350, - 79.5797, 185 m), 2 i 2010, leg. & det. JTJ [JTJ]; NA, Tena, 1 ♂, Río Sinde, pond, 5.3 km E of Puerto Napo (- 1.0517, - 77.7544, 450 m), 13 vii 1996, leg. & det. KJT [KJT]; 3 ♂♂, same data but 22 vii 1996, leg. & det. WFM [FSCA, LSAM, QZAC]; 1 ♂, same data but 12 xi 1997, leg. & det. KJT [KJT]; OR, Francisco de Orellana: 3 ♂♂, 4 ♀♀, Primavera, forest trail from Río Napo to Laguna Taracoa, 27 vii 1977, leg. DRP & S. Hills, det. DRP [DRP]; 2 ♀♀, same data but 26 viii 1980, leg. & det. SWD [SWD]; 1 ♂, Reserva Etnica Waorani, 1 km S Onkone Gare Camp (- 0.6528, - 76.4333, 220 m), 10 iv 1994, leg. T. Erwin, det.? [USNM]; 1 ♂, Yuturim 25 ii 1990, leg. & det.? [QZAC]; SD: 1 ♂, Tinalan- dia, 12 km E of Santo Domingo de los Colorados, streams S of river (- 0.2839, - 79.0500, 760 m), 5 x 1988, leg. & det. SWD [SWD].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B581FF44FF23F6892124FEDB.taxon	discussion	The Libellulidae comprise the most species-rich group of Anisoptera in the Neotropics. These are the dragonflies that most people see, as many are common pond inhabitants, although a few genera depend on flowing waters for their development. We have recorded 33 genera with 141 named species in Ecuador and there undoubtedly will be more species added to the list in future years. One species is endemic to Ecuador.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B580FF45FF23F78425D8FD4F.taxon	discussion	A monotypic genus (for identification see Garrison et al. 2006). Adults are mostly brown (Fig. 85). Habitat is small, shaded forest streams. * Argyrothemis argentea Ris, 1909. Publ. Recs. ECU: (H 14). Also. BRA, GUF, SUR, VEN: (P 11); plus COL: (P 16). Addl. Data. SU, Shushufindi: 1 ♂, trails, small streams 3 km W of Shushufindi, 2 + km N off Hwy. 26 at km marker 17 (- 0.1664, - 76.6917, 290 m), 14 x 2009, leg. & det. JTJ [JTJ]; 1 ♂, 2 km W of Shushufindi, small tribu- taries of Río Curiyacu and isolated rain pools (- 0.1661, - 76.6916), date not provided, data provided by WAH & DLW.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B580FF43FF23F530276AFEFF.taxon	discussion	Two of the three species in this genus are recorded for Ecuador. Species are variable in color pattern, being either brown and black or mostly red (Fig. 86). See Needham et al. (2014) for species identification. Habitat consists of various ponds and swamps, even brackish water, usually vegetated. Brachymesia furcata (Hagen, 1861). Publ. Recs. ECU: (H 06), (P 11), (H 14); SU: * Dunkle (1981). Also ARG, BOL, BRA, CEN, CHL, GUF, PRY, PER, SUR, TTO, VEN: (H 06, P 11); plus COL: (H 06, P & P). Addl. Data. EO, Arenillas: 1 ♂, farm pond, seeps, 11 km S of Arenillas, on road to Alamar (- 3.6436, - 80.0989, 95 m), 10 iv 2008, leg. & det. JTJ [JTJ]; Machala: borrow pit pond, Hwy. 25, S of Machala (- 3.4750, - 79.9778), 10 iv 2008, leg. & det. KJT [KJT]; SE, Salinas: 1 ♂, Salinas (- 2.95, - 80.13, 3 m), 30 vi 1989, leg. L. Stange & R. Miller, det. GHB [FSCA]; SU, Shushufindi: 10 ♂♂, 5 ♀♀, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 13 – 28 xi 1973, leg. B. Drummond III, det.? [FSCA, KJT]; 2 ♂♂, same data but 11 – 17 i 1974 [FSCA]; 45 ♂♂, 3 ♀♀, same data but 1 – 27 xi 1980, leg. & det. MJW [FSCA]; 2 ♂♂, same data but 23 viii 1980, leg. & det. SWD [UTIC]; 5 ♂♂, same data but 3 – 16 vi 1977, leg. P. Spangler & D. Givens, det. RWG [RWG, USNM]; 1 ♀, same data but lake edge, 5 vii 1971, leg. D. Pearson, det. DRP [DRP]; 3 ♂♂, same data but 16 – 25 viii 1971 [DRP]; 1 ♂, same data but 2 x 1971 [DRP]; 2 ♂♂, same data but 11 xii 1971; 1 ♂, same data but 13 i 1972; 1 ♂, same data but 20 ii 1972 [DRP]; 1 ♀, same data but 6 iii 1972; 13 ♂♂, 1 ♀, same data but 24 – 26 vii 1977, leg. & det. DRP [DRP]; 2 ♂♂, same data but forest edge, 25 – 26 vi 1977, leg. & det. DRP [DRP]. Brachymesia herbida (Gundlach, 1889). Publ. Recs. ECU: (P 11), (H 14); CB: Campos (1922 — as Cannacria batesi); GA: * Currie (1901 — as Holotype C. fumipennis: “ Galapagos Is ” [Isabela], Albemarle, 23 iii 1899, leg. R. Snodgrass, USNM], Calvert (1907), Ris (1912), Campos (1922), (P 11), (Peck 1992); NA: Tennessen (1998); SU: Dunkle (1981). Also ARG, BOL, BRA, CEN, COL, GUF, GUY, PRY, PER, TTO, VEN: (H 06), (P 11); plus SUR: (H 06). Addl. Data. GA. Isabela: 2 ♀♀, Isla Isabela, littoral zone, 2 iii 1989, leg. & det. S. Peck [USNM]; 1 ♂, same data but arid zone, 10 iii 1989 [USNM]; Santa Cruz: 2 ♀♀, Isla Santa Cruz, Tortuga Bay, 30 i 1989, leg. & det. S. Peck, et. al. [USNM]; GY, Guayaquil: 1 ♀, Puna Vieja (- 2.97, - 80.18, 30 m), leg. F. Campos, det. MJW [ANSP]; LR, Babahoyo: 1 ♀, swamp at Babahoyo, 12 vi 1975, leg. J. Cohen & A. Langley, det.? [USNM]; Buena Fé: 1 ♂, [Consumulo], S Río Palenque (- 0.58, - 79.35, 175 m), 16 vi 1974, leg. J. Longino, det.? [FSCA]; OR, Francisco de Orellana: 2 ♂♂, trib. of Río Payamino, road to Loreto, 1.7 km W of jct. w / Coca Road (- 0.4367, - 77.0017, 258 m), 18 xi 1997, leg. & det. KJT [KJT]; 1 ♂, same but leg. & det. WFM [FSCA]: SU, Shushufindi: 1 ♂, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 31 x 1980, leg. & det. MJW [FSCA]; 1 ♀, same data but 3 xi 1980 [FSCA]; 1 ♀, same data but lakeshore, 26 vii 1977, leg. DRP & S. Hills, det. DRP [DRP]; same data but 22 viii 1980, leg. & det. SWD [KJT]; 6 ♂♂, 1 ♀, same data but 23 viii 1980 [JJD, SWD]; 2 ♂♂, La Selva, 100 km E of Coca, Garzaco- cha, 12 x 1988, leg. & det. SWD [UTIC].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B586FF40FF23F7842074F8F3.taxon	discussion	A genus of 15 species that is in need of revision; species identification is difficult and new species probably remain to be discovered. The five species that occur in Ecuador can be identified using Calvert (1898) and Ris (1913 a). Brechmorhoga adults are mainly dark (Fig. 87) and inhabit streams in the small to medium-size range, usually with sand and rock substrates and medium to fast flow. The adults are fast fliers, often difficult to see over shaded streams. Brechmorhoga nubecula (Rambur, 1842). Publ. Recs. ECU: Schmidt (1942), Paulson (1983), Tsuda (1986, 1991), Steinmann (1997 b), (H 06), (P 11), (H 14); CR: Campos (1922); EO: Campos (1927); GY: Campos (1922); IM: * Ris (1913 a) Also ARG, BRA, CEN, COL, PRY, PER, TTO, VEN: (H 06), (P 11). Addl. Data. BO, Caluma: 1 ♂, rocky stream 5 km E of Caluma, Guaranda Road, 5 iii 2010, leg. & det. JJD [JJD]; GY, Naranjal: 1 ♂, San Rafael [as “ San Raphael ”] (- 2.18, - 79.23, 200 m), date?, leg. F. Campos, det. P. Calvert [FSCA]; NA, Tena: 1 ♀, Río Chinchipino, 27 km E of Puerto Napo (- 1.0728, - 77.5864, 385 m), 13 xi 1997, leg. & det. KJT [KJT]; 1 ♂, NE of Tena, Río Shicai-Yacu, 900 m, 6 iv 1941, leg. WCM, det. WFM [FSCA]; 1 ♀, [Puerto Napo, about 1 km up river on Río Anzu from the Río Napo], “ Las Palmas ” (- 1.07, - 77.80, 900 m), 11 xii 1946, leg. WCM, det. WFM [FSCA]; PA, Mera: 1 ♀, Río Metolo, Los Copales cabanas between Mera and Shell, W of Puyo, Hwy. 50 (- 1.4919, - 78.0725, 1070 m), 20 ix 2005, leg. & det. JJD [JJD]; SD: 1 ♂, Santo Domingo de Los Colorados, 10 km E on Hwy. 30, W of Hotel Tinalandia (- 0.28, - 79.03, 800 m), 19 v 1975, leg. T. Emmel, det.? [FSCA]; 1 cop. pair, 1 ♀, Santo Domingo de Los Colorados, 12 km E on Hwy. 30 near Hotel Tinalandia (- 0.30, - 79.67, 609 m), 25 vii 1986, leg. T. Emmel, det.? [FSCA]; 1 ♂, same data but 22 xii 1975, leg. J. Harper, det. KJT [KJT]; SU, Cascales: 1 ♀, 37 km W of Lago Agrío on Baeza Road, river, 20 viii 1980, leg. & det. SWD [SWD]; Shushufindi: 1 ♂, Limoncocha, forest path, 4 i 1972, leg. D. Pearson, det. DRP [DRP]; ZC, Zamora: 1 ♀, stream off road to village of Guadalupe N of Zamora (- 3.8939, - 78.8880), 21 viii 2006, leg. & det. JCA [UTIC]; 1 ♀, small stream 5 km N of Guaguayme Alto (- 3.8778, - 78.8928, 880 m), 4 iv 2008, leg. & det. KJT [JTJ]. * Brechmorhoga pertinax (Hagen, 1961). Publ. Recs. ECU: (H 14). Also BOL, CEN, COL, PER: (H 06), (P 11). Addl. Data. MS, Gualaquiza: 1 ♀, Gualaquiza, 800 m, leg. Campos, det. PPC [FSCA]; PC, [Quito]: Río Pacto, broad shallow river with rocks and boulders, 1 km SW Pacto village (- 0.1398, - 78.7741), date not provided, data provided by WAH & DLW. Brechmorhoga praecox (Hagen, 1861). Publ. Recs. ECU: (H 14); NA: * Tennessen (1996 b). Also BRA, CEN, COL, PER, VEN: (H 06), (P 11); plus TTO: (P 11). Addl. Data. EO, Piñas: 6 ♂♂, 1 ♀, Piñas (- 3.67, - 79.23, 1000 – 1500 m), 2 – 25 vii 1941, leg. D. Laddey, det.? [FSCA]; Portovelo: 2 ♀♀, Portovelo (- 3.72, - 79.62, 600 – 650 m), 3 – 4 viii 1941, leg. D. Laddey, det.? [FSCA]; IM, Ibarra: 1 ♂, pools / stream near sugarcane fields 0.5 km S of Salinas on road to Lita (0.4853, - 78.1333), 2 ii 1997, leg. & det. JJD, as B. postlobata [JJD]; 1 ♀, same data but leg. & det. KJT [KJT]; MS, Morona: 1 ♀, Mangosiza, Río Upano (- 2.60, - 77.77, 850 m), date?, leg. D. Laddey, det. WFM [FSCA]; NA, Tena: 1 ♂, Puerto Napo, 5.3 km E of Hwy. 45 on J. S. B. S. Road, Río Sinde (- 1.0500, - 77.7481, 450 m), 15 vi 1995, leg. & det. KJT [KJT]; 3 ♂♂, 1 ♀, same data but leg. & det. JJD [JJD]; 8 ♂♂, 3 ♀♀, same data but leg. & det. WFM [FSCA, LSAM, QZAC, USNM]; 1 ♂, same data but leg. & det. SWD [SWD]; 2 ♂♂, same data but 13 vii 1996 [SWD]; 2 ♂♂, same data but 22 vii 1996 [FSCA, QZAC]; 1 ♂, same data but 12 xi 1997, 1 ♂, same data but leg. & det. TWD [SWD]; 1 ♂, Puerto Napo 11.2 km E of Hwy. 45 on J. S. B. S. Road, Río Tiyuyacu (- 1.0516, - 77.7017, 425 m), 13 xi 1997, leg. & det, WFM [QZAC]; ZC, Yantzaza: 1 ♂, Yantzaza, 6.5 km [N on Hwy. 45], Río Chicana (- 3.7681, - 78.7350, 762 m), 6 xi 1997, leg. & det. WFM [QZAC]. Brechmorhoga rapax Calvert, 1898. Publ. Recs. ECU: * Ris (1913 a), Campos (1925 — “ por Ris 1916 ”), Ris (1918 — as B. rapax rapax), Maria (1938), Paulson (1983), Tsuda (1991), Steinmann (1997 b — as B. rapax rapax), (H 06), (P 11), (H 14); MS: Tennessen (2006 — as B. rapax crocosema). Also CEN, COL, VEN: (H 06), (P 11); plus PER: (P 16). Addl. Data. LJ, Loja: 1 ♂, small stream off road towards Zamora (- 4.0330, - 79.0132), 18 viii 2006, leg. & det. JCA [UTIC]; MS, Morona: 3 ♂♂, rocky stream 22.5 km W of Macas, Guamote Road, 18 viii 2005, leg. & det. JJD, as subsp. crocosema [JJD]; 1 ♂, same data but rocky stream about 2.4 km W of that location flowing over road (- 2.2342, - 78.2181), leg. & det. KJT as subsp. crocosema [KJT]; SD: 1 ♀, Santo Domingo de Los Colorados 12 km E on Hwy. 30 near Hotel Tinalandia (- 0.30, - 79.67, 610 m), 15 v 1986, leg. T. Emmel, det.? [FSCA]; 1 ♂, About 93 km SW Quito, Chiriboga Road, 9 vii 1990, leg. OSF, det.? [USNM]; 1 ♂, small stream, Hwy. 30, km marker 61, E of La Union de los Toachi, 27 ii 2010, leg. & det. JJD as subsp. crocosema [JJD]. * Brechmorhoga vivax Calvert, 1906. Publ. Recs. ECU: (H 14). Also CEN, PER, VEN, PRY: (H 06, P 11); plus COL: (P & P); and ARG (P 16). Addl. Data. CT, Pangua: 1 ♂, small stream 8 km E of Moraspungo, 22 ii 2010 m, leg. & det. JJD [JJD]; EO, Piñas: 1 ♀, Piedras (- 3.63, - 79.92, 300 m), 2 vii 1941, leg. D. Laddey, det.? [USNM]; ES, San Lorenzo: 2 ♂♂, stream near RR track, about 2 km N of Lita, 5 ii 1997, leg. E. Tapia & KJT, det. KJT [KJT]; PA, Mera: 2 ♂♂, Río Metolo, Los Copales, between Mera and Shell (- 1.4919, - 78.0725, 1070 m), 19 ix 2005, leg. & det. KJT [KJT, RWG]; 1 ♀, same data but 4 x 2009, leg. & det. JTJ [JTJ]; Pastaza: 1 ♂, Río Sandalias, 5 km S of Puyo, Hwy. 45 (- 1.5003, - 77.9522, 1000 m), 11 xi 1997, leg. & det. KJT [KJT]; 1 ♀, stream 5.5 + km SE off Hwy. 45 (trib. of larger river), N of Puyo (- 1.3606, - 77.8653, 900 m), 9 x 2009, leg. & det. JTJ [JTJ]; SD: 1 ♀, small streams, Hwy. 30, near km marker 70, near La Unión del Toachi (- 0.4806, - 78.9394, 920 m), 27 ii 2010, leg. & det. JTJ [JTJ]; TU, Baños: 1 ♀, stream, Hwy. 50, between Río Verde and Río Negro (- 1.4069, - 78.2553, 1515 m), 4 x 2009, leg. & det. JTJ [JTJ]; ZC, Zamora: 1 ♂, stream / waterfall W of Zamora (- 4.0456, - 79.0006, 1150 m), 1 iv 2008, leg. & det. JTJ [JTJ].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B584FF4EFF23F6882053FF6B.taxon	discussion	A genus of only three species, two of which occur in Ecuador. The most recent diagnosis was by Donnelly (1992). Cannaphila are medium-sized dragonflies (Fig. 88 a & b) that occupy muddy seepage areas of small slow streams, usually partly shaded, from low to medium elevations.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B584FF4EFF23F6882053FF6B.taxon	discussion	Addl. Data. ES, Quinindé: 1 ♀, road from Las Golondrinas to Canande Reserve, 30 km NE Quininde, drain- age pond for palm oil plantation, with small stream outflow (0.4669, - 79.2336), no date provided, data provided by WAH & DLW; 1 ♂, small gravel stream in forest, 38 km NE of Quininde, tributary of Río Hoja Blanca (- 0.5455, - 79.2007), date not provided, data provided by WAH & DLW. Cannaphila vibex (Hagen, 1861). Publ. Recs. ECU: * Calvert (1909 — “ teste Ris in Litt ”), Racenis (1954), Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14); BO: Ris (1910); CR: Campos (1922 — as Leptetrum merida); CB: Campos (1922 — as L. merida); ES: Ris (1910), Daigle et al. (1997); GY: Ris (1910), Campos (1922 — as L. merida); NA: Ris (1910), Tennessen (1996 b); OR: von Ellenrieder & Garrison (2011); ZC: Ris (1910). Also ARG, BOL, BRA, CEN, COL, PER, VEN: (H 06), (P 11). Addl. Data. CB, Alausí: [FSCA]; Camada: [FSCA]; BO, San Miguel: [ANSP]; CT, Sigchos: [QZAC]; EO, Piñas: [ACRP] ES, San Lorenzo: [FSCA, JJD, KJT, LSAM, QZAC, USNM]; GY, Naranjal: [ANSP, FSCA]; IM, Cotacachi: [FSCA, KJT, USFQ]; Ibarra: [KJT, QZAC]; San Miguel Urcuq: [SWD]; LJ, Loja: [FSCA, JTJ]; LR, Buena Fé: [DRP]; MS, Gualaquiza: [ANSP, FSCA]; Huamboya: [FSCA, TWD, USNM]; Limón Indanza: [FSCA, KJT]; Logrono: [FSCA]; Morona: [FSCA, QZAC, TWD, USNM]; NA, Archidona: [FSCA, UTIC, KJT, LSAM, QZAC, RWG, SWD, USNM]; El Chaco: [FSCA, UTIC, KJT, SWD]; Quijos: [FSCA, UTIC, JJD, KJT, LSAM, QZAC, TWD, USNM]; Tena: [LSAM, QZAC]; OR, Francisco de Orellana: [RWG, NVE, TWD, USFQ]; Loreto: [FSCA, KJT]; PA, Mera: [FSCA, JTJ]; Pastaza: [FSCA, SWD]; PC, San Miguel de los Bancos: [FSCA, USFQ]; Quito: [USNM]; SD: [DRP, FSCA, UTIC, USNM]; SU, Gonzalo Pizarro: [QZAC]; Shushufindi: [DRP, FSCA, KJT, SWD]; TU, Baños: [FSCA, QZAC]; ZC, Yantzaza: [FSCA, QZAC, USNM]; Zamora: [JTJ].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B58BFF4FFF23F7382698FD17.taxon	discussion	One of the four species of this genus occurs in Ecuador (Fig. 89). See Ris (1910) for identification. Adults are slight of stature; they perch near partly shaded ponds and swamps. Dasythemis esmeralda Ris, 1910. Publ. Recs. ECU: Campos (1925), Fraser (1946 b), Geijskes (1971), Paulson (1983), Davies & Tobin (1984), Tsuda (1986, 1991), Steinmann (1997 b), (H 06), (P 11), (H 14); ES: * Ris (1910 — type series, 2 ♂♂, Esmeralda, “ from Selys ”), Schmidt (1942), Bridges (1993); OR: von Ellenrieder & Garrison (2011); SU: Donnelly (1998). Also BOL, BRA, COL, GUF, PER, SUR, TTO, VEN: (P 11). Huamboya: 1 cop. pair, Macas, 58 km N on Hwy. 45 (70 km N of Sucua), roadside ditch and tribs. near Río Pasanac (- 1.9447, - 77.8536, 762 m), 10 xi 1997, leg. & det. WFM [FSCA]; NA, Carlos Julio Arosemena Tola: 1 ♂, [Ila], 49 km N of Puyo on road to Tena Río Anzu [mistakenly listed as “ Río Pastaza ”] (- 1.20, - 77.87, 512 m), 26 x 1980, leg. & det. MJW [FSCA]; Tena: 1 ♂, J. S. B. S., near Tena, clearing near stream (450 m), 19 vi 1994, leg. A. Jaramilo, det. DRP [DRP]; 1 ♂, same data but 16 x 1988, leg. & det. SWD [SWD]; 4 ♂♂, 5.3 km E Puerto Napo on J. S. B. S. Road, Río Sinde (- 1.0500, - 77.7481, 450 m), 16 x 1988, leg. & det. SWD [RWG, SWD]; 1 ♂, same data but 13 vii 1996, leg. & det. WFM [FSCA]; 1 ♂, Puerto Napo, 17.2 km E on J. S. B. S. road, pool near stream (340 m), 22 vii 1996, leg. & det. SWD [SWD]; OR, Francisco de Orellana: 4 ♂♂, seepage stream 2.1 km E of Yasuni station, Yasuni Preserve, 15 xi 1997, leg. & det. JJD [JJD]; 3 ♂♂, swamp, P. N. Y. (- 0.6542, - 76.3542, 200 m), 15 x 1997, leg. & det. KJT [KJT]; 1 ♂, P. N. Y., 8 km N of Waorani Road on Yasuni Road palm swamp just S of Río Bogi 2 Este, km marker 42.3 (- 0.6734, - 76.4397), 12 vi 1995, leg. & det. WFM [FSCA]; 1 ♂, P. N. Y., grassy semi-shaded pools / stream, 5.6 km W Yasuni Road on PC 29 - 30 (- 0.7133, - 76.3000, 200 m), 10 vi 1995, leg. & det. SWD [SWD]; 1 ♀, pond, Loreto Road, 11.8 km W of Río Payamino ferry, 18 xi 1997, leg. & det. KJT [KJT]; 4 ♂♂, 1 ♀, T. B. S., USFQ, Mauritia palm swamp off Guacamayo trail at marker 2100 (- 0.6331, - 76.1628, 220 m), 20 i 2009, leg. & det. RWG & NVE [NVE]; Loreto: 6 ♂♂, Pano, 12 ix 1990, leg. OSF, det.? [USNM]; 1 ♂, streams and forest trails 12 km E of Loreto, 2 + km S of Hwy. 45 (- 0.6217, - 77.2600, 365 m), 12 x 2009, leg. & det. KJT [KJT]; 2 ♂♂, 1 ♀, same data but 15 x 2009 [KJT, RWG]; 1 ♂, same data but 15 x 2009, leg. & det. JTJ [JTJ]; PA, Mera: 1 ♀, Mera, 12 km W of Puyo, 5 ii 1976, leg. P. Spangler, det.? [USNM]; SU, Cascales: 1 ♀, 37 km W of Lago Agrio, river, 20 viii 1980, leg. & det. SWD [SWD]; Lago Agrio: 1 ♀, 2 km N. Lago Agrio, 26 ix 1975, leg. A. Langley, det.? [USNM]; 2 ♀♀, Santa Cecilia, Río Francisco de Orellana, 25 – 31 iii 1969, leg. P & P Spangler, det.? [USNM]; 1 cop. pair, 26 km S of Lago Agrio, stream & marsh, 19 viii 1980, leg. & det. SWD [SWD]; Shushufindi: 1 ♂, 1 ♀, 1 cop. pair, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 9 xi 1972, leg. E. Stiles, det. DRP [DRP]; 1 ♂, same data but 10 xi 1980, leg. & det. MJW [FSCA]; 1 ♂, 2 ♀♀, La Selva Lodge, forest (- 0.4983, - 76.3733, 200 m), 25 xi 1997, leg. & det. TWD [SWD]; 2 ♀♀, stream about 3 km SW of Shushufindi (- 0.2136, - 76.6786, 250 m), 12 v 2009, leg. & det. KJT [KJT]; 2 ♂♂, stream about 3 km NW of Shushufindi (- 0.1664, - 76.6917, 290 m), 14 v 2009, leg. & det. KJT [KJT]; 1 ♂, 1 ♀, same data but 14 x 2009, leg. & det. JTJ [JTJ].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B58AFF4CFF23F05B2714FEA3.taxon	discussion	One of the eight species has been found in Ecuador. For identification, refer to Montgomery (1940). The adults are very distinctive, with black wings that usually have red or yellowish veins at the base (Fig. 90). Habitat includes ponds and marshes. Diastatops pullata (Burmeister, 1839). Publ. Recs. ECU: (H 06), (P 11), (H 14), Bota-Sierra et al. (2015); OR: * Dunkle (1981); SU: (Donnelly 1998). Also ARG, BOL BRA, GUF, GUY, PER, SUR, VEN: (H 06), (P 11); plus PRY: (P 11); and COL: (P 16). Addl. Data. OR, Francisco de Orellana: 3 ♂♂, 1 ♀, Anangu, SE across Río Napo from Providencia Lago Anangu “ on Río Napo ” (- 0.5213, - 76.4361, 300 m), 9 v 1975, leg. T. Emmel, det.? [FSCA]: 1 ♂, same data but leg. B. Drummond III, det.? [FSCA]; 1 ♂, [La Nueva Esperanza] [across Río Napo from] near Primavera “ Limonco- cha ”, Río Napo & Lake Taracoa (- 0.45, - 76.77, 300 m), 4 xi 1980, leg. B. Drummond III, det.? [FSCA]: 1 ♂, same data but leg. & det. MJW [FSCA]; 14 ♂♂, 1 ♀, same data but 27 vii 1977, leg. DRP & S. Hills, det. DRP [DRP, RWG]; 1 ♂, same but 26 viii 1978, leg. & det. KWK [SWD]; 2 ♂♂, same data but 26 viii 1980, leg. & det. SWD [SWD]; SU, Shushufindi: 2 ♂♂, La Selva Lodge, L. Mandicocha and outlet stream (- 0.4983, - 76.3733, 200 m), 25 xi 1997, leg. & det. TWD [SWD].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B589FF4DFF23F7602143FBAF.taxon	discussion	Two of the seven species are recorded for Ecuador; see Ris (1913 a, 1918) and Needham et al. (2014) for identification. Adults are slender and mostly dark (Fig. 91); they perch with wings typically set down and forward over the thorax. They breed in slow streams, mostly in pooled areas, that are either open or mostly shaded. Dythemis nigra Martin, 1897. Publ. Recs. ECU: * Kirby (1897); Maria (1938), Paulson (2011), (H 14) — all as D. multipunctata); ES: (Daigle et al. 1997), SD: (Daigle et al. 1997); ZC: Tennessen & Johnson (2008). Also ARG, BRA, CEN, GUF, GUY, PRY, PER, TTO, VEN: (P 11); plus COL: (P & P); and SUR (P 16). Addl Data: ES, San Lorenzo: [FSCA, KJT]; LR, Buena Fé: [DRP, SWD]; MN, Portoviejo: [JTJ]; MS, Huambioya: [FSCA. KJT]; NA, Tena: [FSCA, UTIC, JTJ, KJT, QZAC, SWD, TWD, USNM; OR, Francisco de Orellana: [FSCA, QZAC, USNM]; Loreto: [FSCA, QZAC, SWD]; PA, Pastaza: [USNM]; Santa Clara: [USNM]; PC, San Miguel de los Bancos: (FSCA, KJT, USFQ]; Quito: [ACRP]; SD: [FSCA, QZAC]; SU, Lago Agrio: [USNM]; Shushufindi: [KJT, SWD]; ZC, El Pangui: [KJT]. Note: Meurgey (2010) found the true D. multipunctata on St. Vincent in the Lesser Antilles and stated that Central and South American records for D. multipunctata really represent D. nigra. Dythemis sterilis (Hagen, 1861). Publ. Recs. ECU: von Ellenrieder & Muzón (2007), Paulson (2011), (H 14); CR: Campos (1922); EO: Campos (1927); GY: * Calvert (1906 — as D. velox form sterilis), Campos (1922); LR: * Calvert (1906 — as D. velox form sterilis), Campos (1922); ZC: Tennessen & Johnson (2008). Also CEN, COL, VEN: (P 11); Plus PER, TTO: (P 11); and ARG, BRA, GUF: (H 06). Addl. Data. CT, Pangua: 1 ♂, concrete pool, E side of Las Juntas (- 1.1831, - 79.2453, 330 m), 22 ii 2010, leg. & det. JTJ [JTJ]; EO, Piñas: 1 ♂, Piñas, 15 km S (- 3.80, - 79.73, 1100 m), 4 vii 1989, leg. R. Miller, det. GHB [FSCA]; Santa Rosa: 1 ♀, stream, 12 km E of Arenillas, Hwy. 25 (- 3.5244, - 79.9831, 30 m), 29 iii 2008, leg. & det. KJT [KJT]; ES, Esmeraldas: SW of Esmeraldas SW, muddy stream, 19 vii 1977, leg. & det. DRP [DRP]; ♀, 1 ♂, (300 m), 6 – 22 v 1983, leg. & det.? [QZAC]; GY, Balao: 1 ♂, small stream / ditches in banana plantation, about 2 km E of Hwy. 25 near Balao Jct. (- 2.9100, - 79.6919, 70 m), 28 iii 2008, leg. & det. JTJ [JTJ]; Naranjal: 2 ♂♂, Bucay (- 2.20, - 79.18, 233 m), 1911, leg. F. Campos, det. WFM [ANSP, FSCA]; 3 ♂♂, 3 ♀♀, Cerro Mas Vale E on Hwy. Río Churute (- 2.42, - 79.67, 9 m), 10 ii 2001, leg. & det. F. Sibley [FSCA]; 1 ♂, Las Mercedes SE at small bridge below thermal hot springs (- 2.6058, - 79.4886, 107 m), 13 iii 2001, leg. & det. F. Sibley [FSCA]; 1 ♂, Las Mercedes SE on road to Cuenca, base of mountain (- 2.6169, - 79.4542, 244 m), 13 ii 2001, leg. & det. F. Sibley [FSCA]; LR, Babahoyo: 1 ♂, Babahoyo, date?, leg. F. Campos, det.? [USNM]; Buena Fé: 2 ♂♂, Santo Domingo 47 km S, forest slough at Río Palenque Field Station (200 m), 18 vii 1977, leg. & det. DRP [DRP]; 2 ♂♂, same data but 8 x 1988, leg. & det. SWD [UTIC]; Quevedo: 1 ♀, Pichilingue, Río Palenque (- 1.10, - 79.48, 64 m), 2 ii 1955, leg. E. Schlinger, det. MJW [FSCA]; PC, San Miguel: between Puerto Vicente Maldonaldo and Puerto Quito (550 m), photo M. Reid, det. DRP; SD: 1 ♂, Santo Domingo de los Colorados, 11 v 1985, leg. & det.? [QZAC]; 1 ♂, pond near Hotel Colorados, Hwy. 30, 1 i 97, leg. E. Tapia, det. KJT [KJT]; ZC, Zamora: 1 ♂, Zamaro, 4.7 km SE [on Río Jambue Road], gravel quarry adjacent to Río Jambue (- 4.07, - 78.93, 914 m), 5 xi 1997, leg. & det. WFM [FSCA]; 1 ♂, same data but leg. & det. KJT [KJT]; 1 ♀, stream crossing road just before village of Guadalupe (- 3.9114 - 78.8575), 20 – 21 viii 2006, leg. & det. JCA [UTIC]. Note: Meurgey (2011) created the subspecies D. sterilis sterilis and D. sterilis multipunctata. Since the latter is restricted to the lesser Antilles, we simply use D. sterilis for the Ecuador fauna.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B58FFF4AFF23F688260AFDDF.taxon	discussion	A single species recorded in Ecuador; adults with reddish-orange abdomen (Fig. 92). For species identification, see Westfall (1988). The nymphs occupy flowing sections of small sand / rock bottomed streams. Elasmothemis cannacrioides (Calvert, 1906). Publ. Recs. ECU: * Westfall (1988), (H 06), (P 11), (H 14); ES: Daigle et al. (1997); NA: Tennessen (1996 b), Daigle (1996). Also ARG, BRA, CEN, COL, GUF, PER, TTO, VEN: (H 06), (P 11); plus SUR: (P 16). Addl. Data. CN, La Troncal: [JJD, SWD]; Cañar: [FSCA, TWD, USNM]; EO, Piñas: [JTJ]; ES, San Loren- zo: [JJD, KJT]: GY, Balao: [JTJ]; MN, El Carmen: [USNM]; MS, Gualaquiza: [USNM]; Huamboya: [SWD]; NA, Tena: [FSCA, JJD, KJT, LSAM, QZAC, SWD, TWD, USNM]; OR, Francisco de Orellana: [FSCA, KJT, QZAC, USNM]; Loreto: [USNM]; PA, Mera: [FSCA, QZAC, USNM]; Pastaza: [FSCA, KJT]; PC, Quito: [KJT]; SD: [FSCA]; SU, Cascales: [UTIC, KJT, SWD]; Lago Agrio: [UTIC, USNM]; Shushufindi: [DRP, FSCA, KJT, USNM]; ZC, Yantzaza: [FSCA, KJT, QZAC, USNM].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B58FFF4BFF23F049260EFEA3.taxon	discussion	One of the two species described in this genus has been reported from Ecuador. For the complicated history of species identification in this genus, see the papers by Machado (1954, 1992). Adults are small, slender dragonflies (Fig. 93) that frequent small forested streams. Elga leptostyla Ris, 1911. Publ. Recs. ECU: (H 06), (P 11), (H 14); NA: * Tennessen (1996 b, 1998). Also in BRA CEN, GUF, GUY, PER, SUR, VEN: (H 06, P 11); plus COL: (H 06), (P & P); plus TTO: (P 11). Addl. Data. NA, Tena: 1 ♀, Río Chinchipino, 27 km E of Puerto Napo (- 1.0728, - 77.5864, 385 m), 15 vi 1995, leg. & det. KJT [KJT]; 1 cop. pair, 3 ♂♂, same data but 13 xi 1997 [KJT]; 1 ♂, same data but 10 x 2009, leg. & det. JTJ [JTJ]; 1 ♀, same data but 22 vii 1996, leg. & det. SWD [FSCA]; OR, Francisco de Orellana: 1 ♀, P. N. Y., Yasuni Biological Research Sta. along trails and roads within 5 km of Res. Sta. (- 0.6719, - 76.3990, 245 m), 18 vii 1996, leg. & det. WFM [FSCA]; 1 ♂, T. B. S, TLE Plot T- 8, Sta. 10, Lot # 2879, fogging canopy (- 0.6319, - 76.1442, 220 - 250 m), 21 vii 2002, leg. T. Erwin, det. WFM [USNM]; 2 ♀♀, T. B. S., Quebrada Harpía (- 0.6350, - 76.1460, 217 m), 5 xii 2012, leg. & det. ACRP [ACRP]; SU, Shushufindi: 1 ♀, stream 3 km NW of Shushufindi (- 0.1664, - 76.6917, 290 m), 14 v 2009, leg. KJT, det. RWG [KJT].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B58EFF77FF23F7602081FEA3.taxon	discussion	Of the ten species in this genus, all eight that occur in South America have been recorded in Ecuador. See Kennedy (1923) and Williamson (1923 b) for species identification. Adults are medium to large with diverse body coloration, including green, red, orange, brown and black (Figs. 94 a & b). The habitats are mostly lentic, including ponds, lake edges and marshes, but they can be found also in slow streams. Erythemis attala (Selys, 1857). Publ Recs. ECU: Ris (1911 b), Maria (1938), Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14); CR: * Calvert (1906), Campos (1922); EO: Campos (1927); GY: * Calvert (1906), Campos (1922, 1931), Palacino-Rodríguez et al. (2015); LR: * Calvert (1906), Campos (1922); MN: Ris (1911 b). Also in ARG, BOL, BRA, CEN, COL, GUF, GUY, PRY, PER, SUR, VEN, URY: (H 06), (P 11). Addl. Data. EO, Santa Rosa: 1 ♂, 1 ♀, pond, 12 km E of Arenillas, Hwy. 25 (- 3.5244, - 79.9831, 30 m), 9 iv 2008, leg. & det. KJT [KJT]; GY, Daule: 1 ♂, Daule, date?, leg. F. Campos, det.? [USNM]; Duran: 1 ♀, Duran (- 2.17, - 79.83, 9 m), 12 ii 1911, leg. S. Rhoads, leg.? [ANSP]; LR, Buena Fé: 1 cop. pair, 3 ♂♂, as “ Pinchincha, ” Río Palenque Biological Station, 47 km S of Santo Domingo de los Colorados (- 0.5864, - 79.3647, 150 m), 8 x 1988, leg. & det. SWD [JJD, KJT, SWD, USNM]; 1 ♂, [Patricia Pilar], just S at Reserva Palenque [about 47 km S of Santo Domingo] (- 0.66, - 79.26, 110 m), 9 x 1988, leg. & det. SWD [FSCA, SWD]; OR, Francisco de Orellana: 1 ♂, Lake Teracoa, 26 viii 1980, leg. & det. SWD [UTIC]; 1 ♂, same data but 9 vii 1983, leg. T. & P. Gantenbein, det. RWG [CSCA]; SU, Shushufindi: 3 ♂♂, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 6 – 21 xi 1980, leg. & det. MJW [FSCA]; 2 ♂♂, same data but 23 viii 1980, leg. & det. SWD [SWD]; 1 ♂, same data but edge of clearing, 11 viii 1972, leg. E. Stiles, det. DRP [DRP]; 2 ♂♂, same data but lakeshore, 24 vii 1977, leg. & det. DRP [DRP]; 1 ♂, La Selva, 100 km E of Coca, Garzacocha, 12 x 1988, leg. & det. SWD [UTIC]. Erythemis carmelita Williamson, 1923. Publ. Recs. SU: * Dunkle (1981). Also BRA COL, PER, VEN: (H 06), (P 11). Addl. Data. SU, Shushufindi: 2 ♂♂, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 6 xi 1980, leg. & det. MJW [FSCA]; 1 ♂, same data but 8 xi 1980 [FSCA]; 1 ♂, same data but 25 xi 1980 [FSCA]; 1 ♂, same data but N end of boat dock, 22 viii 1980, leg. & det. SWD [SWD]; 1 ♀, same data but 25 viii 1980 [SWD]. * Erythemis credula (Hagen, 1861). Publ. Recs. ECU: (H 14). Also ARG, BRA, CEN, COL, GUF, GUY, SUR, VEN: (H 06), (P 11); plus BOL: (H 06); and TTO: (P 11). Addl. Data. PC, San Miguel Bancos: 1 ♀, Mindo (1500 m), 24 i 1994, leg. & det.? [QZAC]; OR, Francisco de Orellana: 1 ♂, pond, Loreto Road, 12.3 km W jct. with Coca Road (- 0.4750, - 77.0747, 290 m), 13 vi 1995, leg. & det. KJT [KJT]; 1 ♂, same data but leg. & det. SWD [SWD]; 2 ♂♂, pond, Loreto Road, 11.8 km W of Río Payamino ferry (0.4752, - 77.0583, 285 m), 18 xi 1997, leg. & det. KJT [KJT]; 1 ♂, same data but leg. & det. WFM [FSCA]; Anangu SE across Río Napo from Providencia Lago Anangu “ on Río Napo ” (- 0.5213, - 76.4361, 300 m), 9 v 1975, leg. T. Emmel, det.? [FSCA]; 1 ♂, same data but leg. B. Drummond III, det.? [FSCA]. Erythemis haematogastra (Burmeister, 1839). Publ. Recs. ECU: Maria (1938), Paulson (2011), (H 14); CR: * Calvert (1906), Campos (1922); EO: Campos (1927); GY: * Calvert (1906); Ris (1911 b), Campos (1922); LR: * Campos (1922); OR: Tennesen (1998), von Ellenrieder & Garrison (2011), Palacino-Rodríguez (2011 — as “ Napo ”). Also in BRA, CEN, COL, GUF, GUY, PRY, PER, SUR, VEN: (H 06), (P 11); plus ARG: (H 06); and TTO: (P 11). Addl. Data. LR, Babahoyo: Babahoyo (- 1.80, - 79.53, 10 m), date?, leg. F Campos, det. P. Calvert [FSCA]; OR, Francisco de Orellana: 1 ♂, [La Nueva Esperanza] [across Río Napo from], near Primavera “ Limoncocha ”, Río Napo & Lake Taracoa (- 0.45, - 76.77, 300 m), 27 vii 1977, leg. DRP & S. Hills, det. DRP (DRP); 3 ♂♂, 1 ♀, same data but 26 viii 1980, leg. & det. SWD [KJT, SWD]; 3 ♂♂, same data but 4 xi 1980, leg. & det. MJW [FSCA]; 2 ♂♂, P. N. Y., 2.1 km E of Yasuni Biological Research Sta., pond and swamp, (- 0.6741, - 76.3685, 245 m), 15 xi 1997, leg. & det. WFM [FSCA, QZAC]; 1 ♂, P. N. Y., small stream at km 38 on Yasuni Road (- 0.6333, - 76.5000), 12 vi 1995, leg. & det. SWD [SWD]; 1 ♂, pond on Coca-Loreto road (- 0.4800, - 77.0800, 300 m), 18 xi 1997, leg. & det. TWD [SWD]; La Joya de los Sachas: 1 ♂, La Joya de los Sachas, E, pond (- 0.2202, - 76.6987, 260 m), 29 x 2013, leg. & det. CDK [FSCA]; SU, Shushufindi: 1 ♀, Limoncocha, Río Napo (- 0.4000, - 76.6167, 290 m), 3 – 16 vi 1977, leg. P. Spangler & D. Givens, det.? [USNM]; 2 ♂♂, same data but 14 xi 1980, leg. & det. MJW [FSCA]; 3 ♂♂, 1 ♀, same data but forest edge & swamp, 25 – 26 vii 1977, leg. & det. DRP [DRP]; 2 ♂♂, 1 ♀, La Selva, Garzacocha, 100 km E Coca, 12 x 1988, leg. & det. SWD [FSCA, SWD]; 1 ♀, Shushufindi, SW, pond (- 0.2201, - 76.6988), 29 x 2013, leg. & det. KJT [FSCA]. Erythemis mithroides (Brauer, 1900). Publ. Recs. ECU: (H 14); SU: * Dunkle (1981). Also BOL, BRA, CEN, COL, GUF, GUY, PRY, PER, SUR, TTO, VEN: (H 06), (P 11). Addl. Data. EO, Santa Rosa: 1 ♂, pond, 12 km E of Arenillas, Hwy. 25 (- 3.5244, - 79.9831, 30 m), 9 iv 2008, leg. & det. KJT [KJT]; LR, Quevedo: 1 ♀, Pichilingue Río Palenque (- 1.10, - 79.48, 65 m), 2 ii 1955, leg. E Schlinger, det.? [FSCA]; SU, Shushufindi: 1 ♂, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 4 xi 1980, leg. & det. MJW [FSCA]; 1 ♂, same data but 27 viii 1980, leg. & det. SWD [SWD]; 2 ♂♂, La Selva, Garzacocha, 100 km E Coca, 12 x 1988, leg. & det. SWD [FSCA, SWD]. Erythemis peruviana (Rambur, 1842). Publ. Recs. ECU: Maria (1938), Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14); CR: * Calvert (1906), Campos (1922); EO: Campos (1927); GY: * Calvert (1906), Ris (1911 b), Campos (1922); NA: Ris (1911 b). Also in ARG BOL, BRA, CEN, COL, GUF, GUY, PRY, PER, SUR, TTO, VEN: (H 06), (P 11); plus TTO: (P 11). Addl. Data. EO, Santa Rosa: 1 ♂, pond, 12 km E of Arenillas, Hwy. 25 (- 3.5244, - 79.9831, 30 m), 9 iv 2008, leg. & det. KJT [KJT]; GY, Guayaquil: 5 ♂♂, Guayaquil (- 2.20, - 79.90, 8 m), ii, iv & viii, leg. F. Campos, det.? [FSCA]; Daule: 1 ♂, Nobol, 5.5 km N, 1 xii 1978, leg. P. Spangler & J. Anderson, det.? [USNM]; Yaguachi: 2 ♀♀, Casiguana, date?, leg. F. Campos, det.? [USNM]; LR, Babahoyo: 1 ♂, 1 ♀, swamp at Babahoyo, 21 vi 1975, leg. J. Cohen, A. & M. Langley, det.? [USNM]; 1 ♂, 20 km N Babahoyo, 22 vi 1975, leg. J. Cohen, A. & M. Langley, det.? [USNM]; OR, Francisco de Orellana: 1 ♂, 1 cop. pair, trib. of Río Payamino, road to Loreto, 1.7 km W of jct. with Coca Road (- 0.4367, - 77.0017, 258 m), 13 vi 1995, leg. & det. KJT [KJT]; 2 ♂♂, 1 ♀, same data but leg. & det. WFM [LSAM, QZAC, USNM]; 1 ♂, same data but leg. & det. SWD [SWD]; 2 ♂♂, same data but 18 xi 1997 [QZAC, USNM]; 2 ♂♂, same data but leg. & det. TWD [SWD]; 2 ♂♂, same data but leg. & det. JJD [JJD]; 1 ♀, same data but leg. & det. KJT [KJT]; PC, San Miguel Bancos: 1 ♂, Mindo, 7 viii 1994, leg. & det.? [QZAC]; SD: 1 ♂, Palmeras, 26 xi 1988, leg. & det.? [QZAC]; SU, Shushufindi: 1 ♀, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 1 ♂, same data but 22 viii 1971, leg. D. Pearson, det. DRP [DRP]; 1 ♀, same data but 31 xii 1973, leg. B. Drummond III, det. KWK [FSCA]; ♀, same data but 11 – 16 v 1975 [FSCA]; 2 ♂♂, same but leg. T. Emmel [FSCA]; 2 ♂♂, 1 ♀, same data but 3 – 16 vi 1977, leg. P. Spangler & D. Givens, det.? [USNM]; 1 ♂, 3 ♀♀, same data but 27 viii 1980, leg. & det. SWD [KJT, SWD]; 5 ♂♂, 4 ♀♀, same data but 31 x 1980, leg. & det. MJW [FSCA]; 11 ♂♂, 6 ♀♀, same data but 1 – 29 xi 1980 [FSCA]; 1 ♀, same data but lake edge, 5 vii 1971, leg. D. Pearson, det. DRP [DRP]. Erythemis plebeja (Burmeister, 1839). Publ. Recs. ECU: Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14); CR: * Calvert (1906), Campos (1922); CB: Campos (1922); EO: Campos (1927); GY: * Calvert (1906), Ris (1911 b), Campos (1922); LR: * Calvert (1906), Campos (1922). Also ARG, BOL, BRA, CEN, COL, GUF, GUY, PRY PER, SUR, VEN: (H 06), (P 11); Plus TTO: (P 11). Addl. Data. CN, La Troncal: [FSCA]; ES, Esmeraldas: [DRP]; GY, Duran: [FSCA, USNM]; Guayaquil: [ANSP, FSCA, USNM]; Naranjal: [FSCA]; Samborondon: [UTIC]; IM, Ibarra: [FSCA]; LR, Babahoyo: [USNM]; Buena Fé: [UTIC]; NA, Tena: [FSCA, UTIC, KJT, LSAM, QZAC]; SD: [FSCA]; SU, Shushufindi: [FSCA]. Erythemis vesiculosa (Fabricius, 1775). Publ. Recs. ECU: Paulson (1983), Tsuda (1986, 1991 — as Lepthemis vesiculosa), (H 06), (P 11), (H 14); CR: * Campos (1900), Calvert (1906 — as L. vesiculosa), Campos (1922 — as L. vesiculosa); CB: Campos (1922 — as L. vesiculosa); EO: Campos (1927); ES: Campos (1922 — as L. vesiculosa); GA: Muddeman (2007); GY: * Campos (1900 as L. vesiculosa), Calvert (1906 — as L. vesiculosa), Ris (1911 b — as L. vesiculosa), Campos (1922 — as L. vesiculosa); LR: Calvert (1906 — as L. vesiculosa), Campos (1922 — as L. vesiculosa). Also in ARG, BOL, BRA, CEN, COL, GUF, GUY, PRY, PER, SUR, VEN: (H 06), (P 11); plus TTO: (P 11). Addl. Data. EO, Piñas: [ACRP]; Santa Rosa: [KJT]; ES, San Lorenzo: [QZAC]; GA, Isabela: [USNM]; San Cristóbal: [ACRP]; GY, Duran: [ANSP]; Guayaquil: [ANSP, FSCA, USNM]; Naranjal [FSCA]; LR, Babahoyo: [USNM]; Montalvo: [JJD]; MN, El Carmen: [QZAC]; Sucre: [QZAC]; NA, Tena: [UTIC]; OR, Francisco de Orellana: [KJT]; SE, Santa Elena: [USNM]; SD: [LSAM]; SU, Shushufindi: [FSCA, UTIC].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B5B2FF71FF23F7602688F95F.taxon	discussion	This is the most species-rich genus (58 described species) of Anisoptera in the Neotropics. The basic reference is Borror (1942); see also Borror (1957) and Paulson (2003). At least 21 species are recorded in Ecuador and more may occur here. Adults are small to medium sized and variously colored in red, brown or black (Figs. 95 a, b, c, d). As expected, the habitats occupied are diverse, including ponds, lakes, sloughs, marshes, brackish water, phytotelmata, and pools in slow streams. A synoptic update of the genus is warranted. Erythrodiplax abjecta (Rambur, 1842). Publ. Recs. ECU: * Karsch (1891 b — as E. ponderosa, description, type locality “ Ecuador ”), Ris (1911 a), Campos (1922 “ por Dr. Calvert ”), Racenis (1954), Bridges (1993), (H 06), Paulson (2003), (P 11), Bota-Sierra (2014 a), (H 14); ES: (Ris, 1911 a); EO: Campos (1927 — as E. connata form?); GY: * Ris (1911 a); PC: Ris (1911 a), Paulson (2003). Also CEN, COL, VEN: (P 11). Addl. Data. CT, Sigchos: 2 ♂♂, Otonga, Esmeraldas and “ Z ” rivers area, in Naranjito, 1 v 1999, leg. E. Tapia, det. JJD [JJD]; 5 ♂♂, San Francisco de las Pampas, principal road near Esmeraldas River on Naranjito (- 0.42, - 79.0, 1450 m), 1 iv 1999, leg. E. Tapia, det.? [QZAC]; 1 ♂, San Francisco de las Pampas Road between Naranjito-San Francisco de Las Pampas (- 0.42, - 79.0, 1500 m), 14 v 1999; leg. E. Tapia, det.? [QZAC]; 3 ♀♀, San Francisco de las Pampas Otonga Reserve on B 2 River above the grass (- 0.42, - 79.0, 1650 m), 28 iv 1999, leg. L. Torres, det.? [QZAC]; 1 ♂, 4 ♀♀, San Francisco de las Pampas Otonga Reserve, Cerca del Río P- 1 (- 0.42, - 79.0, 2200 m), 11 iii 1999, leg. E. Tapia, det.? [QZAC]; GY, Guayaquil: 1 ♂, Guayaquil (- 2.20, - 79.90, 8 m), date?, leg. & det. MJW [FSCA]; Naranjal: 1 ♀, [Venecia], listed as “ Verecia ”, (- 2.17, - 79.52, 23 m), date?, leg. F. Campos, det. P. Calvert [FSCA]; 1 ♂, 4 ♀♀, [San Rafael], listed as “ San Raphael ” (- 2.18, - 79.23, 200 m), date?, leg. F. Campos, det. P. Calvert [FSCA]; Yaguachi: 1 ♀, Casiguana (- 2.13, - 79.78, 4 m), date?, leg. F. Campos, det. P. Calvert [FSCA]; IM: Cotacachi: 3 ♂♂, Santa Rosa, El Refugio de Intag Cloud Forest Lodge, south along road at large open seep (0.3773, - 78.4774, 1907 m), 23 viii 2018, leg. & det. WFM [FSCA, USFQ], same data but 2 ♂♂, south along road at roadside seeps (0.3693, - 78.4728, 1854 m), 23 viii 2018, leg & det. WFM [FSCA, USFQ]; 1 ♂, Apuela, Ruta Plaza Gutierrez, seepage into Rio Tuabunchi (0.3575, - 78.4993, 1634 m), 26 viii 2018, leg. & det. WFM [USFQ]; same data but 1 ♂, leg. & det. KJT [KJT]; same data but, 1 ♂, 27 viii 2018, leg. & det. KJT [KJT]; 2 ♂♂, Selve Alegre, Ruta Tolla Intag, roadside pool (0.2800, - 78.5160, 1751 m), 26 viii 2018, leg & det. WFM [FSCA]; same data but 1 ♂, Santa Rosa, Ruta Siempre Verde, seeps and bog (0.3672, - 78.4378, 2226 m), 28 viii 2018, leg. & det. WFM [FSCA]; same data but 2 ♂♂, 2 ♀♀, pool and seeps (0.3676, - 78.4306, 2322 m), 28 viii 2018, leg. & det. KJT [KJT]; PC, Mejía: 1 ♂, Cornejo Astorga, 9.6 km E, ditch (2012 m), 16 vii 1977, leg. & det. DRP [DRP]; 3 ♂♂, Santo Domingo de los Colorados, 32 km E, waterfall & trickle, 10 x 1988, leg. & det. SWD [DRP, SWD]; 16 ♂♂, 10 km E of Tandapi, 29 km E of Tinalandia (- 0.45, - 78.75, 1920 m), 24 x 1980, leg. & det. MJW [JJD]; Quito: 1 ♂, Maquipucuna Reserve, near Nanegalito, slow stream (1250 m), 28 v 1994, leg. A. Jaramillo, det. DRP [DRP]; San Miguel Bancos: 6 ♂♂, 2 ♀♀, between Calacali and Nanegalito, about 30 km W Quito, 9 vii 1990, leg. OSF, det.? [USNM]; 2 ♂♂, Mindo, Mindo Gardens Ecolodge, vegetated cement pond (- 0.0720, - 78.7560, 1330 m), leg. & det. WFM [FSCA, USFQ]; same data but 1 ♂, leg. & det. KJT [KJT]. * Erythrodiplax amazonica Sjöstedt, 1918. Publ. Recs. BRA, GUF, GUY, PER, TTO, VEN: (H 06), (P 11); plus SUR: (P 16). Addl. Data. OR, Orellana: 1 ♂, T. B. S., Quebrada Harpía (- 0.6351, - 76.1463, 221 m), 11 vi 2013, leg. & det. ACRP (pers. comm., ACRP); SU, Shushufindi: 1 ♂, 3 km SW Shushufindi, stream, (- 0.2136, - 76.6786, 260 m), 12 v 2009, leg. & det. KJT [FSCA]; 1 ♂, 1 ♀, stream, 3 km NW of Shushufindi (- 0.1664, - 76.6917, 290 m), 13 x 2009, leg. & det. JTJ [JTJ]. Erythrodiplax andagoya Borror, 1942. Publ. Recs. ECU: Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14); ES: Daigle et al. (1997); SD: * Borror (1942 — as “ Pichincha ”), Daigle et al. (1997 — as “ Pichincha ”). Also COL: (H 06, P 11, P & P); plus CEN: (P 11). Addl. Data. CT, Pangua: [JTJ]; ES, San Lorenzo: [FSCA, JJD, KJT, LSAM]; GY, Balao: [JTJ]; Guayaquil: [ANSP]; Naranjal: [FSCA]; LR, Buena Fé: [FSCA, SWD, USNM]; Quevedo: [ANSP]; MN, El Carmen: [FSCA, JJD, JTJ, KJT]; PA, Pastaza: [USNM]; PC, Quito: [FSCA, KJT, USFQ]; San Miguel Bancos: [RWG]; SD: [FSCA, JJD, KJT, LSAM, RWG, QZAC, SWD]; SU, Lago Agrio: [USNM]. Note: This often abundent species occurs mainly on the lower Pacific slope. The east slope records from the USNM (Pastaza) need to be checked as they are probably a different species. Erythrodiplax attenuata (Kirby, 1889). Publ. Recs. ECU: * Borror (1942), (H 06), (P 11), (H 14); NA: Tennessen (1996 b). Also BOL, BRA, COL, PER, VEN: (H 06), (P 11). Plus SUR: (H 06). Addl. Data. OR, Francisco de Orellana: 1 ♂, trib. of Río Payamino, road to Loreto, 1.7 km W of jct. with Coca Road (- 0.4367, - 77.0017, 258 m), 13 vi 1995, leg. & det. KJT [KJT]; 2 ♂♂, same data but leg. & det. WFM [FSCA]; La Joya de los Sachas: 1 ♂, Pompeya near Puerto Pompeya Ferry Landing (- 0.4400, - 76.6194, 240 m), 17 vii 1996, leg. & det. WFM [FSCA]. Erythrodiplax basalis (Kirby, 1897). Publ. Recs. ECU: Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14), Dalzochio et al. (2018); NA: * Borror (1942); OR: von Ellenrieder & Garrison (2011); PA: * Borror (1942). Also BRA, COL, GUF, GUY, PER, VEN: (H 06), (P 11); plus ARG BOL, PRY, URY, SUR, TTO: Paulson (2011). Addl. Data. ES, Esmeraldas: [QZAC]; Quininde: Q); LR, Buena Fé: [USNM]; MN, El Carmen: [QZAC]; MS, Huamboya: [FSCA, QZAC]; Santiago: [SWD]; Sucua: [FSCA]; NA, Archidona: [QZAC]; Carlos Julio Arsemena Tol: [FSCA, QZAC, USNM]; El Chaco: [FSCA, UTIC, KJT, LSAM, SWD]; Tena: [DRP, FSCA, UTIC, KJT, QZAC, RWG, TWD, SWD, USNM]; OR, Francisco de Orellana: [DRP, FSCA, KJT, LSAM, QZAC, RWG, TWD, SWD, USNM]; Loreto: [USNM]; PA, Pastaza: [JTJ, KJT, USNM]; Santa Clara: [FSCA]); PC, San Miguel Bancos: [DRP]; SD: [USNM]; SU, Cascales: [FSCA, JJD, KJT, QZAC, USNM]; Lago Agrio: [FSCA, UTIC, LSAM, QZAC, RWG, SWD, USNM]; Shushufindi: [CSCA, DRP, FSCA, JTJ, KJT, LSAM, QZAC, SWD, USNM]; ZC, El Pangui: [KJT]. Erythrodiplax castanea (Burmeister, 1839). Publ., Recs. ECU: Fraser (1946 b), Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14); ES: * Ris (1911 a), Campos (1922 — “ from Ris in Selys collection ”), Daigle et al. (1997); NA: Borror (1942); OR: von Ellenrieder & Garrison (2011); PA: Borror (1942); ZC: Tennessen & Johnson (2008). Also ARG, BOL, BRA, CEN, COL, GUF, GUY, PRY, PER, TTO, VEN: (H 06), (P 11); plus SUR: (P 16). Addl Data. EO, Piñas: [ACRP]; Santa Rosa: [JTJ]; ES, San Lorenzo: [FSCA, JJD, KJT, LSAM, QZAC, USNM]; MS, Huamboya: [FSCA, QZAC, USNM]; OR, Francisco de Orellana: [FSCA, JJD, KJT, LSAM, NVE, QZAC,); Loreto: [FSCA, KJT, SWD]; NA, Archidona: (MAR, USNM]; Tena: [DRP, KJT, LSAM, QZAC, SWD, USFQ, USNM]; SU, Cascales: [SWD]; Lago Agrio: [SWD]; Shushufindi: [FSCA, KJT, JTJ]; ZC, Zamora: [UTIC, JTJ, KJT]. * Erythrodiplax cleopatra Ris, 1911. Publ. Recs. CHL, PER: (H 06), (P 11). Addl. Data. EO, Santa Rosa: 1 ♂, stream and associated pond, wetland about 12 km E of Arenillas, Hwy. 25 (- 3.5244, - 79.9831, 30 m), 29 iii 2008, leg. & det. JTJ, checked by DRP [JTJ].). Note: This is a considerable range extension to the north from previous records. Erythrodiplax fervida (Erichson in Schomburgk, 1848). Publ. Recs. ECU: Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14); CR: * Campos (1922 — as E. ochracea); CB: * Campos (1922 — as E. ochracea); GY: * Campos (1922 — as E. ochracea). Also CEN, COL, GUF, GUY, SUR, VEN: (H 06), (P 11); plus TTO: (P 11). Addl. Data. EO, Piñas: 1 ♂, Buenaventura, Camino principal (- 3.6520, - 79.7922, 323 m), 28 vi 2016, leg. & det. ACRP [ACRP]; Santa Rosa: 1 ♂, 1 cop. pair, pond, 12 km E of Arenillas, Hwy. 25 (- 3.5244, - 79.9831, 30 m), 29 iii 2008, leg. & det. KJT [KJT]; 2 ♂♂, same data but 9 iv 2008, leg. & det. JTJ [JTJ]; MN, Chone: 1 ♂, Chone, W end of town, 9 i 1978, leg. P. Spangler & J. Anderson, det.? [USNM]. Note: According to Borror (1942), all previous references to E. ochracea (Burmeister, 1839) in northern South America should be referred to E. fervida; E. ochracea is a southern South American species. Erythrodiplax funerea (Hagen, 1861). Publ. Recs. ECU: * Ris (1911 a), Campos (1922 — “ por Dr. Ris ”), Paulson (1983), Tsuda (1986, 1991), Steinmann (1997 b), (H 06), (P 11), (H 14), Dalzochio et al. (2018). Also CEN, COL: (H 06), (P 11). Addl. Data. CB, Alausí: 2 ♂♂, Huigra, Río Huagra, 1 km upstream from trail (- 2.2783, - 78.9731, 1350 m), 13 ii 1911, leg. S. Rhoads, det.? [ANSP, FSCA]; ES, San Lorenzo: 3 ♂♂, 1 ♀, Carondelet, date?, leg. F. Campos, det. MJW [ANSP]; 4 ♂♂, 2 ♀♀, Concepcion (1.05, - 78.83, 25 m), leg. F. Campos, det. MJW [ANSP]; GY, Guayaquil: 1 ♂, Puna Vieja (- 2.97, - 80.18, 30 m), leg. F. Campos, det. MJW [ANSP]. Note: These are the southernmost records for this common Central American species. Erythrodiplax fusca (Rambur, 1842). Recs. ECU: * Calvert (1906 — as E. connata, “ form d ”), Campos (1922 — as E. connata fusca), Fraser (1946 b — as E. connata fusca), Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14); BO: Borror (1942); CB: Ris (1911 a), Navás (1934); ES: Daigle et al. (1997), Paulson (2003); LR: Borror (1942); NA: Borror (1942), Paulson (2003); OR: von Ellenrieder & Garrison (2011); PC: Paulson (2003); SD: Daigle et al. (1997 — as “ Pichincha ”), Dalzochio et al. (2018). Also ARG, BOL, BRA, CEN, COL, GUF, GUY, PRY, PER, SUR, URY, VEN: (H 06), (P 11); plus TTO: (P 11). Addl. Data. BO, San Miguel: [USNM]; CN, La Troncal: [KJT, SWD]; CB, Alausí: [FSCA]; Pallatanga: [ANSP]; CT, Pangua: [JTJ]; Sigchos: [QZAC]; EO, Piñas: [FSCA]; Santa Rosa: [JTJ]; ES, Esmeraldas: [DRP]; San Lorenzo: [ANSP, FSCA, KJT, USNM]; GY, Duran: [ANSP, FSCA]; Naranjal: [FSCA]; Yaguachi: [ANSP, FSCA, USNM]; Imbabura, Ibarra: [FSCA, KJT, QZAC, USNM]; LJ, Catamayo: [JTJ]; Loja: [FSCA, UTIC]; Macará: [ANSP, FSCA]; LR, Buena Fé: [DRP, SWD, USNM]; Quevedo: [ANSP, FSCA, USNM]; Ventanas: [JTJ]; MN, El Carmen: [QZAC]; Jipijapa: [FSCA]; Portoviejo: [JTJ]; MS, Gualaquiza: [ANSP, FSCA]; Santiago: [FSCA, KJT, SWD]; NA, Archidona: [FSCA]; El Chaco: [UTIC]; Tena: [ANSP, DRP, FSCA, KJT, LSAM, SWD, USNM]; OR, Francisco de Orellana: [FSCA, UTIC, KJT, QZAC, USFQ, USNM]; Loreto: (MAR); PA, Santa Clara: [KJT]; Pastaza: [FSCA]; PC, Mejía: [SWD]; San Miguel Bancos: [DRP, FSCA, KJT, NVE, RWG, USNM, USFQ]; Quito: [DRP]; SD: [CSCA, DRP, FSCA, UTIC, JJD, KJT, QZAC, SWD, USNM]; SU, Cascales: [KJT, SWD, USNM]; Lago Agrio: [FSCA, UTIC, QZAC, SWD, USNM]; Shushufindi: [DRP, FSCA, SWD, USNM]; ZC, Yantzaza: [KJT, USNM]; Zamora: [FSCA, UTIC, JTJ, KJT, LSAM, QZAC, USNM]. Note: All former records of E. connata (Burmeister, 1839) and E. connata connata from Ecuador are of E. fusca according to Paulson (2003). Also the various references to forms “ a – d ” of connata could be either E. fusca or even E. abjecta. Since these specimens were not available to study nor contribute additional provincial or cantonal records to either species, their data are not included in this study. The abdomen of west slope specimens is not pruinose and specimens are often referred to as “ red form ” whereas those from the eastern slopes are pruinose. Erythrodiplax ines Ris, 1911. Publ. Recs. ECU: Ris (1916 — as E. connata var ines), Schmidt (1942), Fraser (1946 b), Paulson (1983), Tsuda (1986, 1991), Steinmann (1997 b), (H 06), (P 11), (H 14); LJ: Tennessen & Johnson (2008); NA: Borror (1942), Tennessen (1996 b), Daigle (1996); PA: Borror (1942); SD: * Ris (1911 a — Holotype & Allotype: [SD:] “ Santa Inez ” [ZMUH]), Ris (1916, 1918), Campos (1925), Borror (1942), Bridges (1993), Tennessen (2006). Also BOL, COL, PER: (H 06), (P 11). Addl. Data. CB, Alausí: [ANSP, FSCA]; Cumandá: [FSCA]; Pallatanga: [DRP, FSCA]; LJ, Catamayo: [JTJ]; Loja: [FSCA, JTJ, KJT]; MS, Huamboya: [FSCA]; Limón Indanza: [FSCA, KJT, QZAC]; Morona: [FSCA, KJT, QZAC, USNM]; NA, Archidona: [FSCA, KJT, QZAC, LSAM, RWG, SWD, TWD, USNM]; El Chaco: [FSCA, KJT, SWD]; Quijos: [FSCA, JJD, KJT, QZAC, TWD, USNM]; OR, Loreto: [FSCA, JTJ, MAR); SU, Gonzalo Pizarro: [DRP, QZAC, SWD]; PA, Mera: [SWD, USNM]; TU, Baños: [FSCA, JTJ, RWG, QZAC, USNM]; ZC, Yantzaza: [USNM]; Zamora: [FSCA, UTIC, USNM]. Note: According to Heckman (2006), usually found at elevations about 1000 m; this statement seems to hold true; in addition, this species occurs on both sides of the Andes. Erythrodiplax kimminsi Borror, 1942. Publ. Recs. ECU: Fraser (1946 b), Paulson (1983), Tsuda (1986, 1991), (H 06), Bridges (1993), (P 11), (H 14); GY: * Campos (1922 — as E. erichsoni); CB: * Campos (1922 — as E. erichsoni); LR: Borror (1942 — description); OR: von Ellenrieder & Garrison (2011). Also BRA, CEN, COL, PER, VEN: (H 06), (P 11). Addl. Data. EO: Santa Rosa: [JTJ, KJT]; GY, Balao: [JTJ]; LR, Buena Fé: [DRP, FSCA, SWD, USNM]; Quevedo: [ANSP, FSCA]; NA, Tena: [DRP, FSCA, JJD, KJT, SWD, TWD, USNM]; OR, Francisco de Orellana: [FSCA, KJT, LSAM, QZAC, RWG, SWD, TWD, USNM]; Joya de los Sachas: [FSCA]; Loreto: [JTJ, KJT]; PA, Santa Clara: [RWG]; SD: [USNM]; SU, Lago Agrio: [FSCA, KJT, QZAC, SWD, USNM]; Shushufindi: [DRP, FSCA, SWD]. * Erythrodiplax lativittata Borror, 1942 Publ. Recs. ARG, BRA, COL, PER, VEN: (H 06), (P 11). Addl. Data. SU, Shushufindi: 1 ♀, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 2 viii 1974, leg. Boyce A Drummond III, det. WFM [FSCA]. * Erythrodiplax laurentia Borror, 1942. Publ. Recs. BRA, GUF, GUY, SUR, VEN: (H 06), (P 11). Addl. Data. SU, Cascales: labeled “ Napo Prov. ”, 31 km W Lago Agrío and 4 km S, 20 ix 1975, leg. A. Langley & J. Cohen, det. RWG with note “ in geographic range of E. kimminsi, but morphology of laurentia ” [USNM]. Erythrodiplax melanica Borror, 1942. Publ. Recs. ECU: (H 14). Also BRA, PER: (P 16). Addl. Data. OR, Francisco de Orellana: Shiripuno Lodge, 27 km SE Río Shiripuno bridge, (- 1.1047, - 76.7317, 235 m), date not provided, WAH; SU: Cuyabeno Reserve, Sendero Palma Roja, 1.4 km SE of Cuyabeno Lodge (- 0.0151, - 76.1698), date not provided, data provided by WAH. Erythrodiplax melanorubra Borror, 1942. Publ. Recs. ECU: Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14), Dalzochio et al. (2018); BO: * Borror (1942); MS: * Borror (1942 — as “ Azuay ”), Paulson (2003); NA: * Borror (1942), Tennessen (1996 b), Paulson (2003); PA: * Borror (1942); SD: * Borror (1942 — as “ Pichincha ”); TU: * Borror (1942). Also ARG, BOL, BRA, PRY, PER, VEN: (H 06), (P 11); plus COL: (H 06); and CHL, GUF: (P 11). Addl. Data. CT, Sigchos: [FSCA, QZAC]; MS, Huamboya: [FSCA]; Limón Indanza: [FSCA, KJT]; Morona: [FSCA, KJT]; Santiago: [KJT]; Taisha: [DRP]; NA, Archidona: [FSCA, KJT, TWD, USNM]; Carlos Julio Arosemena Tola: [FSCA]; Tena: [DRP, KJT, QZAC, SWD, TWD, USNM]; OR, Loreto: [JJD, KJT, LSAM, QZAC, SWD, TWD, USNM]; PA, Mera: [FSCA, KJT, USNM]; Pastaza: [USNM]; Santa Clara: [FSCA, USNM]; PC, Quito: [QZAC]; San Miguel Bancos: [QZAC]; SD: [QZAC, SWD]; SU, Cascales: [USNM]; Gonzalo Pizarro: [QZAC, USNM]; Lago Agrio: [USNM]; Shushufindi: [KJT]; TU, Baños: [FSCA, JTJ, SWD]; ZC, Yantzaza: [FSCA]; Zamora: [FSCA, KJT]. Erythrodiplax paraguayensis (Förster, 1904). Publ. Recs. ECU: (P 11), (H 14), Dalzochio et al. (2018); NA: * Daigle (1996). Also ARG, BOL, BRA, GUY, PRY, SUR, VEN: (H 06), (P 11); plus COL: (P & P). Addl. Data. OR, Francisco de Orellana: 1 ♂, Río Tiputini slough, km marker 31 at Yasuni Road, Yasuni Pre- serve, 17 xi 1997, leg. & det. JJD; verified ix 2011 - JJD [JJD]. Erythrodiplax parvimaculata Borror, 1942. Publ. Recs. ECU: (H 06), (P 11), (H 14); SU: * Donnelly (1998). Also BOL, VEN: (H 06), (P 11); plus BRA: (H 06); and COL Palacino et al. (2017). Addl. Data. OR, La Joya de los Sachas: 1 ♂, ditch, Pompaya, 0.5 km N of Río Napo ferry, 19 vii 1996, leg. & det. KJT [KJT]; Loreto: 2 ♂♂, Pano, 12 ix 1990, leg. OSF, det.? [USNM]; SU, Shushufindi: La Selva Lodge, forest (- 0.4983, - 76.3733, 200 m), 25 xi 1997, leg. & det. TWD [SWD]; 2 ♀♀, same data but 12 x 1988, leg. & det. SWD [SWD]; 1 ♂, same data but 13 x 1988 [SWD]; 1 ♂, Shushufindi, 3 km SW, stream (- 0.2136, - 76.6786), 12 v 2009, leg. & det. KJT [FSCA]. Erythrodiplax tenuis Borror, 1942. Publ. Recs. ECU: Paulson (1983), Tsuda (1986, 1991), (P 11), (H 14); NA: * Borror (1942 — description), Bridges (1993), Tennessen (1996 b), OR: Tennessen (1998), von Ellenrieder & Garrison (2011). Also PER, BRA (P 11). Addl. Data. NA, Carlos Julio Arosemena Tola: 9 ♂♂, 9 ♀♀, [Ila], 52 km N of Puyo on Road to Tena (- 1.17, - 77.85, 527 m), 26 x 1980, leg. & det. MJW [FSCA]; El Chaco: 1 ♀, stream, 53 km N of Chaco, 18 viii 1980, leg. & det. SWD [SWD]; Tena: 1 ♂, J. S. B. S., Pond # 18, 19 xi 1991, leg. E. McCallie, det.? [USNM]; 1 ♀, Río Chinchipino, 27 km E of Puerto Napo (- 1.0728, - 77.5864, 385 m), 15 vi 1995, leg. & det. KJT [KJT]; 3 ♂♂, same data but 13 xi 1997, leg. & det. JJD [JJD]; 2 ♂♂, sedge marsh, 1.9 km E of 2 nd stream E of Río Sinde, E of Puerto Napo on J. S. B. S. Road, 23 vii 1996, leg. & det. SWD [SWD]; OR, Francisco de Orellana: 1 ♀, trib. of Río Pay- amino, road to Loreto, 1.7 km W of jct. with Coca Road (- 0.4367, - 77.0017, 258 m), 13 vi 1995, leg. & det. KJT [KJT]; 1 ♂, Puerto Francisco de Orellana, 8.2 km W on road to Loreto from Coca-Sachas Road, pond (- 0.4578, - 77.0381, 288 m), 18 xi 1997, leg. & det. WFM [FSCA]; 2 ♂♂, 2 ♀♀, pond, P. N. Y., km marker 29.5, 17 xi 1997, leg. & det. KJT [KJT]; 1 ♂, 1 ♀, Anangu SE across Río Napo from Providencia Lago Anangu, “ on Río Napo ” (- 0.5213, - 76.4361, 300 m), 9 v 1975, leg. T. Emmel, det. MJW [FSCA]; PA, Pastaza: 4 ♂♂, 1 ♀, pond 3 km N of Comunidad Chuwitayo, S of Puyo, 9 xi 1997, leg. & det. KJT [KJT]; 7 ♂♂, 3 ♀♀, Cuasha, approx. 4 km N on Hwy. 45 (94 km N of Sucua), cut grass pond (- 1.83, - 77.82, 884 m), 9 x 1997, leg. & det. WFM [FSCA, LSAM, QZAC, USNM]; 2 ♂♂, pond on road between Macas and Puyo (- 1.8633, - 77.8150, 900 m), 9 xi 1997, leg. & det. TWD [SWD]; Santa Clara: adults, 39 km N of Puyo, 5 vi 1977, leg. P. Spangler & D. Givens, det.? [USNM]; 1 ♀, Puyo Road, 19.3 km S of Río Napo, near Zatzayaco (455 m), 21 vii 1996, leg. & det. KJT [KJT]; SU, Lago Agrio: adults, 2 km N of Lago Agrio, 26 ix 1975, leg. A. Langley, det.? [USNM]; adults, same data but 27 viii 1975 [USNM]; Shushufindi: 1 ♀, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 4 x 1974, leg. B Drummond III, det. MJW [FSCA]; ZC, El Pangui, 2 ♂♂, near stream about 15 km S of El Pangui, 6 xi 1997, leg. & det. KJT [KJT]. Erythrodiplax umbrata (Linnaeus, 1758). Publ. Recs. ECU: * Calvert (1906), Ris (1911 a), Paulson (1983), Tsuda (1991), (H 06), (P 11), (H 14), Dalzochio et al (2018); CR: Campos (1922); CB: Campos (1922); EO: Campos (1927); GY: Calvert (1906), Campos (1922, 1932), Borror (1942); LR: * Calvert (1906), Campos (1922); NA: Borror (1942). Also ARG, BOL, BRA, CEN, COL, GUF, GUY, PRY, PER, SUR, URY, VEN: (H 06), (P 11); plus TTO: (P 11). Addl. Data. ES, Esmeraldas: [DRP, QZAC]; San Lorenzo: [ANSP, FSCA]; GY, Daule: [USNM]; Duran: [ANSP]; Guayaquil: [ANSP, FSCA, USNM]; Naranjal: [FSCA]; Playas: [FSCA]; LJ, Macará: [ANSP]; LR, Babahoyo: [USNM]; Buena Fé: [FSCA]; Montalvo: [JJD]; MN, Chone: [USNM]; El Carmen: [QZAC]; Sucre: [USNM]; NA, Tena: [DRP, FSCA, UTIC, KJT, LSAM, USNM]; OR, Francisco de Orellana: [FSCA, KJT, QZAC, USFQ]; Loreto: [KJT]; SD: [USNM]; SU, Cascales: [USNM]; Lago Agrio: [UTIC, QZAC]; Shushufindi: [DRP, FSCA, UTIC, USNM]; ZC, Zamora: [USNM]. Erythrodiplax unimaculata (De Geer, 1773). Publ. Recs. ECU: Tsuda (1986), (H 06), (P 11), (H 14); PA: * Karsch (1890 — as Micrathyria hemimelaena Holotype ♂: [PA], “ Sarayacu ”), Ris (1911), Borror (1942). Also BOL, BRA, CEN, COL, GUF, GUY, PRY, PER, SUR, VEN: (H 06), (P 11); plus TTO: (P 11). Addl. Data. NA, El Chaco, 1 ♂, stream 39 km NE Chaco, 18 viii 1980, leg. & det. SWD [RWG]; Tena: 1 ♂, pool, Río Sinde, 5 km E of Puerto Napo (- 1.0517, - 77.7544, 450 m), 13 vi 1996, leg. & det. KJT [KJT]; 1 ♂, 1 ♀, same data but 12 xi 1997, leg. & det. KJT [KJT]; OR, Francisco de Orellana: 1 ♂, 4 ♀♀, Reserva Etnica Waorani, Onkone Gare Camp, along road (- 0.63 - 76.6), 30 i 1994, leg. P. Araujo, det.? [USNM]; 1 ♂, San Francisco de Orel- lana [downriver about 20 minutes on the south shore], [Arcoiris Jungle Lodge], San Carlos on Río Napo (- 0.4550, - 76.0042, 300 m), 8 v 1975, leg. T. Emmel det. MJW [FSCA]; 1 ♀, same data but leg. B. Drummond III, det. MJW [FSCA]; 4 ♀♀, Primavera, forest trail from Río Napo to Laguna Taracoa, 27 vii 1977, leg. & det. DRP [DRP]; 3 ♂♂, 2 ♀♀, [La Nueva Esperanza] [across Río Napo from] near Primavera “ Limoncocha ”, Río Napo & Lake Tara- coa (- 0.45, - 76.77, 300 m), 5 ♂♂, 4 ♀♀, same data but 26 viii 1980, leg. & det. SWD [JJD, KJT, SWD]; 3 ♂♂, 10 ♀♀, same data but 4 xi 1980, leg. & det. MJW [FSCA]; 1 cop. pair, same data but 26 xi 1980 [FSCA]; 3 ♀♀, same but 9 vii 1983, leg. T. & P. Gamtenbein, det. RWG [CSCA]; 2 ♂♂, 1 ♀, Yuturi, 25 ii 1990, leg. & det.? [QZAC]; 1 ♂, Payamino, km 13.7, 30 xii 1983, leg. & det.? [QZAC]; 1 ♂, P. N. Y., ditches around research station, 11 vi 1995, leg. & det. SWD [SWD]; 1 ♂, P. N. Y., 2.3 km E of Yasuni Biological Research Station, small stream (- 0.67, - 76.37, 183 m), 9 vi 1995, leg. & det. WFM [FSCA]; 3 ♂♂, [Pompeya Sur] S on [Pompeya Sur-Iro road] at km marker 31, borrow pit near Río Sabaleto (- 0.60, - 76.47 W, 229 m), 12 vi 1995, leg. & det. KJT [KJT]; 3 ♀♀, 12 vi 1995, leg. & det. WFM; 1 ♂, 1 ♀, same data but 17 vii 1996 [FSCA, QZAC, USNM]; 1 ♂, 1 ♀, same data but 17 xi 1997, leg. & det. TWD [SWD]; 1 ♂, 1.7 km W on Loreto Road from Coca-Sachas Road, stream, pools & flowing shallow marsh (- 0.4367, - 77.0017, 258 m), 13 vi 1995, leg. & det. WFM [FSCA]; La Joya de los Sachas: 1 ♂, 1 ♀, Pompeya near Puerto Pompeya Ferry Landing (- 0.4400, - 76.6194, 240 m), 17 vii 1996, leg. & det. WFM [QZAC]; 1 ♂, 1 ♀, 0.5 km W of Puerto Pompeya Ferry Landing roadside ditches (- 0.4444, - 76.6263, 238 m), 19 vii 1996, leg. & det. WFM [FSCA]; PC, Quito: 1 ♂, Guayllabamba (2000 m), 9 i 1993, leg. & det.? [QZAC]; San Miguel Bancos: 1 ♂, Puerto Quito Endega (750 m), xii 1983, leg. & det.? [QZAC]; SU, Cascales: 1 ♂, [El Dorado de Cascales], 58 km W of Lago Agrío on Hwy. 45 at “ El Paraiso ”, stream / marsh (- 0.08, - 77.20, 366 m), 8 vi 1995; leg. & det. WFM [FSCA]; Lago Agrio: 2 ♂♂, 1 ♀, 17 km S of Lago Agrio, 19 viii 1980, leg. & det. SWD [SWD]; 2 ♀♀, stream, E edge of Lago Agrio, 18 viii 1980, leg. & det. SWD [SWD]; 1 ♂, pond 3 km W of Lago Agrío on Hwy. 45 (- 0.97, - 76.83, 275 m), 8 vi 1995, leg. & det. KJT [KJT]; 1 ♂, same data but leg. & det. WFM [LSAM]; Shushufindi: 4 ♂♂, 1 ♀, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 4 – 6 ix 1972, leg. T. Emmel, det. MJW & SWD [FSCA]; 1 ♂, 1 ♀, same data but 21 iii 1974 [FSCA]; 6 ♂♂, 2 ♀♀, same data but 11 – 14 v 1975 [FSCA]; 5 ♀♀, same data but 5 – 24 xi 1973, leg. B. Drummond III, det. MJW & SWD [FSCA]; 1 ♂, same data but 11 i 1974 [FSCA]; 1 ♂, same data but 17 iii 1974 [FSCA]; 1 ♂, 2 ♀♀, same data but 1 – 31 viii 1974 [FSCA]; 7 ♂♂, 6 ♀♀, same data but 10 – 15 v 1975 [FSCA]; 1 ♀, same data but 21 xii 1979, leg. C. Burne, det. SWD [FSCA]; 3 ♂♂, 5 ♀♀, same data but 23 – 28 viii 1980, leg. & det. SWD [SWD]; 21 ♂♂, 24 ♀♀, same data but 31 x – 24 xi 1980, leg. & det. MJW [FSCA]; 1 ♂, 1 ♀, same data but lake edge, 5 vii 1971, leg. D. Pearson, det. DRP [DRP; 1 ♂, same data but 2 x 1971 [DRP]; 1 ♂, same data but 11 xii 1971 [DRP]; 1 ♀, same data but 13 i 1972 [DRP]; 1 ♂, same data but 20 ii 1972 [DRP]; 1 ♀, same data but 24 ii 1972 [DRP]; 6 ♀♀, same data but road through partially cleared forest, 12 viii 1971, leg. D. Pearson, det. DRP [DRP]; 1 ♀, same data but forest trail, 10 viii 1971, leg. D. Pearson, det. DRP [DRP]; 1 ♀, same data but 28 i 1972 [DRP]; 1 ♂, same data but 1 iii 1972; 3 ♀♀, same data but clearing, 11 / 12 viii 1972, leg. E. Stiles, det. DRP [DRP]; 11 ♀♀, same data but 23 – 26 vii 1977, leg. & det. DRP [DRP]; 3 ♂♂, same data but swamp, 26 vii 1977, leg. & det. DRP & S. Hills [DRP]; 2 ♂♂, 4 ♀♀, [Limoncocha] [about 40 km E] or 100 km E of Coca [on Río Napo], Lago Garzacocha La Selva Jungle Lodge (- 0.4819, - 76.3733, 235 m), 12 x 1988, leg. & det. SWD [FSCA, USNM]; 1 ♂, same data but 13 iv 1992, leg. & det. E. Smith [SWD]; 1 ♂, 1.2 km W on Hwy. 22, Río Shushufindi (- 0.1853, - 76.6617, 255 m), 16 vii 1996, leg. & det. WFM [LSAM].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B5B4FF7EFF23F12424D9F83A.taxon	discussion	A monotypic genus (Kirby 1889; De Marmels 1989). Very small dragonfly with white eyes, black thorax and red abdomen (Fig. 96), found in shaded pools with leaf litter on bottom. Fylgia amazonica Kirby, 1889. Publ. Recs. ECU: (H 06), (P 11), (H 14); NA: * Tennessen (1998); OR: * Tennessen (1998), von Ellenrieder & Garrison (2011). Also BRA, GUF, GUY, PER, SUR, VEN: (H 06), (P 11); plus COL: (P 16). Addl. Data. NA, Tena,: 3 ♂♂, oxbow pool, Río Chinchipino, 27.7 km E of Puerto Napo (- 1.0728, - 77.5864, 385 m), 13 xi 1997, leg. & det. KJT [KJT]; 1 ♂, J. S. B. S., zona encharcada camiño Napo (- 1.0652, - 77.6196, 404 m), 2 xi 2014, leg. & det. ACRP (pers. comm., ACRP); OR, Francisco de Orellana: 1 ♂, forest swamp, P. N. Y. (- 0.6508, - 76.3558, 200 m), 15 xi 1997, leg. & det. KJT [KJT]; 2 ♂♂, 1 ♀, same data but 16 xi 1997 [KJT]; 3 ♂♂, same data but leg. & det. TWD [SWD]; SU, Shushufindi: 1 ♂, [Limoncocha] [about 40 km E] or 100 km E of Coca [on Río Napo], Lago Garzacocha, La Selva Jungle Lodge (- 0.4819, - 76.3733, 235 m), 12 x 1988, leg. & det. SWD [SWD]. Gynothemis Calvert in Ris, 1909 Two species of this small genus of small dragonflies (Fig. 97) are found in Ecuador, one of which we have not identified yet. Some of the species now included in this genus had been placed in Macrothemis. The habitat is small forested pools containing leaf litter (Fleck 2004). * Gynothemis pumila (Karsch, 1890). Publ. Recs. ECU: (H 14). Also BRA, COL, GUF, GUY, PER, TTO, VEN: (H 06), (P 11); plus SUR: (P 16). Addl. Data. NA, Tena: 1 ♂, 2 ♀♀, Puerto Napo, 23 km E, J. S. B. S., tropical wet forest 18 x 1988, leg. & det. SWD [FSCA]; 1 ♀, same data but 16 x 1988; SU, Shushufindi: 1 ♂, 2 ♀♀, stream about 3 km SW of Shushufindi (- 0.2136, - 76.6786), 13 x 2009, leg. & det. KJT [KJT]; 2 ♀♀, trails, small streams 3 km W of Shushufindi, 2 + km N off Hwy. 26 at km marker 17 (- 0.1664, - 76.6917, 290 m), 13 x 2009, leg. & det. JTJ [JTJ]; same data but 1 ♂, 1 ♀, 28 x 2013, leg. & det. CDK [FSCA].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B5B4FF7EFF23F12424D9F83A.taxon	description	Gynothemis sp. unresolved: OR, Loreto: 1 ♀, forest about 12 km NE of Loreto (- 0,6217, - 77.2600, 365 m), 12 x 2009, leg. & det. KJT [KJT].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B5BAFF7CFF23F6882714FE37.taxon	discussion	Only one species of this small genus has been found in Ecuador. Another species, I. amazonica (Fig. 98), has yet to be recorded in Ecuador although it probably occurs here. A key to species was provided by Rácenis (1969). Habitats include ponds and marshes. Idiataphe longipes (Hagen, 1861). Publ. Recs. ECU: (H 14 — as I. cf. longipes) PA: * Tennessen (1998). Also BRA, COL, GUF, GUY, PRY, PER, VEN: (H 06), (P 11); Plus ARG: (P 11). Addl. Data. OR, Francisco de Orellana: 1 ♀, pond, Loreto Road, 12.3 km W of jct. with Coca Road (- 0.4750, - 77.0075, 290 m), 13 vi 1995, leg. & det. KJT [KJT]; 1 ♂, ponds near Comunidad El Rocio, (- 0.5054, - 77.1569, 288 m), 27 x 2013, leg. & det. CDK [FSCA]; PA, Pastaza: 1 ♂, 1 ♀, pond 3 km N of Comunidad Chuwitayo, S of Puyo, 9 xi 1997, leg. & det. KJT [KJT]; 1 ♀, pond on road between Macas and Puyo (- 1.8633, - 77.8150, 900 m), 9 xi 1997, leg. & det. TWD [SWD].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B5B9FF66FF23F7CC268BFEA3.taxon	discussion	Of the 28 described New World species, only one species has been found in Ecuador (Fig. 99), the nymphs of which develop in small shaded pools and ditches. Libellula herculea Karsch, 1889. Publ. Recs. ECU: Kennedy (1922), Racenis (1954), Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14); AZ: * Kirby (1889 — as Belonia longipennis), Calvert (1906), Campos (1922); BO: Ris (1910); ES: Calvert (1909), Campos (1922), Daigle et al. (1997); OR: von Ellenrieder & Garrison (2011); PC: * Calvert (1909), Campos (1922). Also ARG, BOL, BRA, CEN, COL, GUF, GUY, PRY, PER, VEN: (H 06), (P 11); plus SUR: (P 16). Addl. Data. AZ, Santa Isabel: 1 ♂, stream, 21 km E of Hwy. 25, road from Balao jct. (- 2.9400, - 79.5653, 730 m), 11 iv 2008, leg. & det. KJT [KJT]; 1 ♂, 1 ♀, same data but leg. & det. JTJ; BO, San Miguel: 1 ♂, Bal- zapamba (- 1.7667, - 79.1833, 700 m), ix 1935, leg. F. Campos, det. MJW [ANSP]; EO, Piñas: Buenaventura, Río Principal (- 3.6561, - 79.7735, 329 m), 19 vi 2016, leg. & det. ACRP [ACRP]; ES, San Lorenzo: Lita along RR track, 1 – 5 km W, adjacent to Río Baboso seeps, pools & streams (0.8792, - 78.4589, 535 m), 5 ii 1997, 5 ♂♂, 1 ♀, leg. & det. WFM [FSCA, LSAM, QZAC, USNM]; 1 ♂, 1 ♀, same data but leg. & det. JJD [JJD]; 1 ♂, same data but leg. & det. KJT [KJT]; MS, Huanboya: 1 ♂, small stream near Río Pasanac, Hwy. 45, S of river, E side of Hwy., 16 ix 2005, leg. & det. KJT [KJT]; 1 ♀, Macas, 58 km N on Hwy. 45 (70 km N of Sucua), roadside ditch and tribs near Río Pasanac (- 1.9447, - 77.8536, 762 m), 10 xi 1997, leg. & det. WFM [FSCA]; Taisha: 1 ♀, Macuma, small ponds (867 m), 1 ii 1972, leg. D. Pearson, det. DRP [DRP]; NA, Tena: 2 ♂♂, Tena (500 m), leg. F. Campos, det. MJW [ANSP]; 1 ♂, Tena (- 0.98, - 77.80, 500 m), date?, leg. F. Campos, det. MJW [FSCA]; adult, 1 km S of Tena, 28 v 1977, leg. P. Spangler & D. Givens, det.? [USNM]; 1 ♂, J. S. B. S., near Tena (450 m), 18 vi 1994, leg. A. Jaramillo, det. DRP [DRP]; OR, Francisco de Orellana: 1 ♂, T. B. S., open grassy area behind dining room, surrounded by bushes and trees and some seepage areas (- 0.6383, - 76.1492, 220 m), 10 i 2009, leg. & det. RWG & NVE [USFQ]; Loreto: 1 ♀, small streams and forest trails 12 km E of Loreto, 2 + km S of Hwy. 45 (- 0.6217, - 77.2600, 365 m), 12 x 2009, leg. & det. JTJ [JTJ]; 1 ♂, Cotapino, S along roadside stream (- 0.7076, - 77.4743, 530 m), 4 xi 2013, leg. & det. CDK [FSCA]; same data but 1 ♂, leg. & det. KJT [FSCA]; PC, San Miguel Bancos: 1 ♂, Puerto Quito (750 m), 28 v, leg.?, det. SWD [QZAC]; 1 ♂, 1 ♀, Mindo, swamps and streams near town (- 0.0556, - 78.7714, 1244 m), 28 i 2009, leg. & det. RWG & NVE [RWG]; 1 ♂, 1 ♀, Mindo, Via al Mariposario, Mindo Gardens Eco Lodge, vege- tated cement pond (- 0.0720, - 78.7560, 1390 m), 3 ix 2018, leg. & det. KJT [KJT]; Quito: 3 ♂♂, 1 ♀, Río Tulambi, Forestry Sta. Maquipucuna, about 5 km E of Nanegal (0.12, - 78.62), 9 v 1990, leg. OSF, det.? [USNM]; SD: 1 ♀, Santo Domingo, 12 km E at Tinalandia, 11 – 17 v 1986, leg. G. Edwards, det. SWD [SWD]; 2 ♂♂, about 93 km SW Quito, Chiriboga Road, 9 vi 1990, leg. OSF, det.? [USNM]; SU, Shushufindi: 1 ♀, Limoncocha, forest trail, 8 x 1971, leg. D. Pearson, det. DRP [DRP]. Note: There is a ♂ specimen in QZAC labeled “ Calvla ” 12 i 1977; we could not find this location in any reference. Macrothemis Hagen, 1868 A species-rich genus of 41 species with at least 14 species in Ecuador. The latest key to males was provided by May (1998) although species identification is difficult; females of very few species are identifiable. These are small to medium-sized mostly dark dragonflies (Fig. 100) that inhabit small streams to medium-sized rivers, both open and forested. Macrothemis delia Ris, 1913. Publ. Recs. E CU: * (H 14). Also CEN, SUR, VEN: (H 06), (P 16). Addl. Data. OR, Francisco de Orellana: 1 ♂, Río Shiripuno, bridge crossing 59 km SSE of Coca, (- 1.0079, - 76.9451), date not provided, data provided by WAH & DLW; 1 ♂, T. B. S., Río Tiputini, 55 km SE of Pompeya, 94 km ESE of Coca (- 0.6369, - 76.1503), date not provided, data provided by WAH & DLW. * Macrothemis extensa Ris, 1913. Publ. Recs. BRA, CEN, GUF, PER, VEN: (H 06), (P 11); plus SUR: (P 16). Addl. Data. LR, Buena Fé: 1 ♂, Río Palenque Biological Station, 47 km S of Santo Domingo de los Colorados (0.5864, - 79.3647, 150 m), 8 x 1988, leg. & det. SWD [SWD]; SD: 1 ♀, Alluriquín 7.3 km W of Hotel Tinalandia, 29 x 1994, leg. A. Jaramillo, det. DRP [DRP]. * Macrothemis fallax May, 1998. Publ. Recs. CEN: May (1998), (P 16); plus COL: (P & P), (P 16). Addl. Data. ES, San Lorenzo: 1 ♀, tiny stream near RR track, about 4 km N of Lita, 5 ii 1997, leg. & det. KJT [KJT]; MN, Portoviejo: 1 ♂, concrete ditch, Hwy. E 40, E of Portoviejo, 4 iii 2010, leg. & det. JJD [JJD]; 1 ♂, same data but leg. KJT, det. WFM [FSCA]; SD: 3 ♂♂, Tinalandia, 12 km E of Santo Domingo de los Colorados, streams S of river (- 0.2839, - 79.05, 760 m), 5 x 1988, leg. & det. SWD [RWG, SWD]. Note: The additional records are the southernmost for this species which is often confused with M. pseudimitans. Macrothemis hahneli Ris, 1913. Publ. Recs. ECU: (H 06), (P 11), (H 14). * Daigle et al. (1996). NA: Daigle (1996 — misidentified as M. musiva). Also in COL, PER, VEN: (H 06), (P 11); plus ARG: (Garrison pers. comm. 2019). Addl. Data. MS, Huamboya: 1 ♂, Macas, 58 km N on Hwy. 45 [70 km N of Sucua], roadside ditch and tribs near Río Pasanac (- 1.9447, - 77.8536, 762 m), 16 ix 2005, leg. & det. WFM [FSCA]; Morona: 3 ♂♂, 1 ♀, small stream and roadside seepage pool, 15 km W of Macas (- 2.2731, - 78.1939, 1070 m), 15 ix 2005, leg. & det. KJT [KJT]; 3 ♂♂, 1 ♀, leg. & det. WFM [FSCA, QZAC, USNM]; 3 ♂♂, rocky stream, 22.5 km W of Macas, Guamote Road, 18 ix 2005, leg. & det. JJD [JJD]; 2 ♂♂, 2 ♀♀, Macas, 22.9 km W on Guamote Road, 3.3 km W on Río Al- banico rocky stream and flowing roadside ditches (- 2.2342, - 78.2181, 1690 m), 18 xi 2005, leg. & det. WFM [FSCA, LSAM]; NA, Archidona: 5 ♂♂, 1 ♀, [Narupa], E on road to Loreto, 43.5 km W of Loreto seep near large rocky river [Río Hollin] (- 0.7022, - 77.7189, 1036 m), 15 vii 1996, leg, & det. JJD [JJD]; 2 ♂♂, same data but leg. & det. SWD [SWD]; 1 ♂, same data but 14 vi 1995, leg. & det. KJT [KJT]; 1 ♂, 1 ♀, same data but 15 vii 1996, leg. & det. WFM [FSCA, LSAM]; 5 ♂♂, same data but 14 xi 1997 [FSCA, LSAM, QZAC, USNM]; 2 ♂♂, [Sarayacu], 12.8 km N of Narupa on Baeza Hwy., small shallow sandy stream (- 0.67, - 77.78, 1585 m), 14 vii 1996, leg. & det. WFM [LSAM, QZAC]; 3 ♂♂, same data but leg. KJT [KJT]; 3 ♂♂, same data but leg. & det. SWD [SWD]; 2 ♂♂, seep by waterfall, Loreto Road, 44.1 km E of Narupa jct., 14 xi 1997, leg. & det. KJT [KJT]; 1 male same data but leg. & det. TWD [SWD]; 2 ♂♂, [Jondachi], 42 km N of Tena on road between Tena & Baeza (- 0.72, - 77.78, 1190 m), 27 x 1980, leg. & det. MJW [FSCA, RWG]; Quijos: 1 ♂, 1 ♀, Baeza 4.8 – 5.0 km S, small clear streams and seeps (- 0.4761, - 77.8711, 1830 m), 20 xi 1997, leg. & det. WFM [FSCA]; 2 ♂♂, same data but leg. & det. JJD [JJD]; 5 ♂♂, 2 ♀♀, same data but leg. & det. KJT [KJT]; 3 ♂♂, 2 ♀♀, same data but leg. & det. TWD [SWD]; 1 ♀, same data but 10 v 2009, [KJT]; 1 ♂, same data but 3 xi 2001, leg. & det. F. Sibley [FSCA]; 1 ♂, Baeza, 1 mile S on Hwy. 45 (- 0.4641, - 77.8842, 1830 m), 3 xi 2001, leg. & det. F. Sibley [FSCA]; 1 ♂, stream, Baeza-Tena Road, near San Isidore Road, 24 vii 1996, leg. & det. KJT [KJT]; PC, Mejía: 1 ♀, Tatatambo, about 5 km E on Hwy. 30, rocky stream (- 0.45, - 78.73, 2286 m), 30 i 1997, leg. & det. WFM [FSCA]; PA, Mera: 1 ♀, seep near Río Anzu, 12.0 km N of Mera (- 1.4181, - 78.0528, 1190 m), 21 ix 2005, leg. & det. KJT [KJT]; 1 ♂, [Mera] Baños-Puyo Road, [about 5 km W], small forested pond (- 1.4806, - 78.1444, 1097 m), 8 xi 2001, leg. & det. F. Sibley [FSCA]; ZC, Yantzaza: 1 ♀, Yantzaza, 10.8 km N [on Hwy. 45], rocky gravel stream (- 3.7669, - 78.6915, 762 m), 6 xi 1997, leg. & det. WFM [FSCA]; Zamora: 1 ♂, stream / waterfall W of Zamora (- 3.9583, - 79.0289, 1660 m), 1 iv 2008, JTJ [JTJ]; 1 ♀, 30 km E of Loja, foot of cascade, 23 ix 1990, leg. OSF, det.? [USNM]. Macrothemis hemichlora (Burmeister, 1839). Publ. Recs. ECU: * Calvert (1906), Ris (1913 a), Campos (1922), Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14). Also ARG CEN, COL, GUF, PER, SUR, TTO, VEN: (P 11); plus BRA: (H 06). Addl. Data. EO, Arenillas: [JTJ, KJT]; Piñas: [ACRP, FSCA]; Portovelo: [FSCA]; LR, Buena Fé: [SWD]; MN, Flavio Alfario: [FSCA]; MS, Morona: [FSCA, KJT]; Santiago: [SWD]; NA, Tena: [DRP, FSCA, UTIC, KJT, LSAM, QZAC, SWD, TWD, USNM]; OR, Francisco de Orellana: [KJT]; SU, Shushufindi: [FSCA, KJT]; ZC: Nangaritza: [FSCA]; Yantzaza: [FSCA]. * Macrothemis idalia? Ris, 1919. Publ. Recs. GUY, VEN: (H 06), (P 11). Addl. Data. OR, Loreto: 1 ♀, 12 km NE of Loreto, forest (- 0.6217, - 77.2600, 365 m), 12 x 2009, leg. KJT, det. RWG [FSCA]; SU, Lago Agrio: 2 ♀♀, [Santa Cecillia]. 24 km W of Lago Agrío on road to Baeza, stream, 20 viii 1980, leg. SWD, det. WFM [FSCA]. Note: Rosser Garrison (pers. comm.) compared the ♀ from near Loreto with ♀♀ in his collection and tenta- tively identified it as this species. A male from Sucumbíos Province, near Shushufindi, collected by KJT on 13 May 2009 was sent to R. W. Garrison who replied “ ... this could be the unknown ♂? of M. idalia Ris – size, body pattern, claws etc. consistent with known females; but until new material, association would be difficult. ” Further collecting, especially a mated pair, will be necessary to confirm this record. Heckman (2006) stated that “ without a description of the ♂, the status of this species is uncertain. ” * Macrothemis imitans Karsch, 1890. Publ. Recs. ECU: (H 14). Also ARG, BOL, BRA, CEN, GUY, PRY, TTO, URY, VEN: (H 06), (P 11); plus COL: (P & P). Addl. Data. NA, Tena: 3 ♂♂, 1 ♀, Puerto Napo 5.3 km E of Hwy. 45 on J. S. B. S. Road, Río Sinde (- 1.0500, - 77.7481, 450 m), 15 vi 1995, leg. & det. WFM [FSCA, QZAC, USNM]; 3 ♂♂, same data but leg. & det. KJT [KJT]; 2 ♂♂, same data but leg. & det. SWD [SWD]; 1 ♂, same data but 13 vii 1996 [KJT]; 2 ♂♂, 1 ♀, same data but leg. & det. WFM [FSCA, QZAC]; 3 ♂♂, Puerto Napo 11.2 km E of Hwy. 45 on J. S. B. S. Road, Río Tiyuyacu (- 1.0516, - 77.7017, 396 m), 13 xi 1997, leg. & det. WFM [FSCA, QZAC, USNM]; OR, Francisco de Orellana: 1 ♂, [Pom- peya Sur] S on [Pompeya Sur-Iro road] at km marker 31, borrow pit near Río Sabaleto (- 0.60, - 76.47, 229 m), 17 xi 1997, leg. & det. WFM [FSCA]; PA, Pastaza: 1 ♂, Río Sandalias, 5 km S of Puyo, Hwy. 45 (- 1.5003, - 77.9522, 1000 m), 11 xi 1997, leg. & det. KJT [KJT]; 1 ♂, same data but leg. & det. WFM [FSCA]; SU, Cascales: 1 ♂, river, 37 km W of Lago Agrio, 20 viii 1980, leg. & det. SWD [SWD]; Lago Agrio: 1 ♂, stream, 26 km S of Lago Agrio, 19 viii 1980, leg. & det. SWD [SWD]. Macrothemis inequiunguis Calvert, 1895. Publ. Recs. ECU: * Calvert (1898, 1906), Ris (1913 a), Campos (1922), Tsuda (1991), (H 06), (H 14). Also CEN, VEN: (H 06), (P 11); plus COL: (H 06). Addl. Data. SD: 2 ♂♂, (as M. cf. inequiunguis), Alluriquín 7.3 km W, forest stream at Hotel Tinalandia, 20 vii 1977, leg. & det. DRP [DRP]. Note: The distribution data indicate that M. inequiunguis is confined to the west slope, its range extending northward to Central America and extreme southern United States, whereas M. tessellata (see below), a very similar species considered by some to be synonymous with M. inequiunguis, is confined to the east slope. * Macrothemis ludia Belle, 1987. Publ. Recs. ECU: (H 14). Also VEN: (H 06), (P 11); plus SUR: (P 16). Addl. Data. NA, Tena, 1 ♀, J. S. B. S., near Tena, in garden (450 m), 12 vi 1994, leg. A. Jaramillo, det. DRP [DRP]; OR, Francisco de Orellana: 1 ♂, T. B. S., Río Tiputini, 55 km SE of Pompeya, 94 km ESE of Coca (- 0.6369, - 76.1503), date not provided, data provided by WAH & DLW; SU, Shushufindi: 1 ♂, Limoncocha, forest stream, 9 iii 1972, leg. D. Pearson, det. DRP [DRP]. Macrothemis mortoni Ris, 1913. Publ. Recs. ECU: (H 06), (P 11), (H 14); NA: * Tennessen (1996 b). Also BOL PER, VEN: (H 06), (P 11). Addl. Data. NA, Archidona: 1 ♂, stream 43.5 km W of Loreto (1100 m), 14 vi 1995, leg. & det. KJT [KJT]; Tena: 1 ♂, stream, road to J. S. B. S., 11.2 km E of Puerto Napo, 15 vi 1995, leg. & det. KJT [KJT]; 1 ♂, Puerto Napo, 3.2 km E on J. S. B. S. Road, sloughs along Napo River, 23 vii 1996, leg. & det. SWD [SWD]; MS, Huamboya: 1 ♂, stream 10 km S. of Comunidad Chuwitayo (- 1.9500, - 77.8583, 800 m), 10 xi 1997, leg. & det. TWD [SWD]. Macrothemis musiva Calvert, 1898. Publ. Recs. ECU: Schmidt (1942), (H 06), (P 11), (H 14); ES: * Ris (1913 a), Campos (1922). Also ARG, BOL, BRA, CEN, COL, GUF, PRY, PER, VEN: (H 06), (P 11). Addl. Data. MS, Gualaquiza: 3 ♂♂, 3 ♀♀, Gualaquiza (- 3.40, - 78.58, 800 m), iv – v 1935, leg. F. Campos, det. MJW [ANSP, FSCA]; NA, Tena: 4 ♂♂, 1 ♀, Puerto Napo 3.2 km E on J. S. B. S. Road ponds and flowing marsh (- 1.0456, - 77.7467, 420 m), 23 vii 1996, leg. & det. WFM [FSCA, LSAM, QZAC, USNM]; 2 ♂♂, same data but leg. & det. KJT [KJT]; 2 ♂♂, same data but leg. & det. SWD [SWD]; 1 ♀, Puerto Napo 5.3 km E of Hwy. 45 on J. S. B. S. Road, Río Sinde (- 1.0500, - 77.7481, 450 m), 22 vii 1996, leg. & det. WFM [FSCA]; 1 ♀, Puerto Napo 27.7 km E of Hwy. 45 on J. S. B. S. Road, Río Chinchipino (- 1.0728, - 77.5864, 385 m), 13 xi 1997 [FSCA]; 2 ♀♀, same data but leg. & det. KJT [KJT]; 1 ♂, Tena, 13 ix 1990, leg. OSF, det.? [USNM]; OR, Francisco de Orellana: 1 ♂, P. N. Y., 2.3 km E of Yasuni Biological Research Sta., small stream (- 0.67, - 76.37), 9 vi 1995, leg. & det. WFM [FSCA]; 1 ♂, P. N. Y., 3.6 km S of Yasuni Road on Waorani Reserve Road (- 0.6956, - 76.4594, 244 m), 10 vi 1995, leg. & det. WFM [QZAC]; 1 ♂, P. N. Y., Yasuni Biological Research Sta. along trails and roads within 5 km of Res. Sta. (- 0.6719, - 76.3992, 244 m), 18 vii 1996, leg. & det. WFM [FSCA]; 1 ♂, stream, Yasuni Road, 20.5 km N Sci. Res. Sta., 11 vi 1995, KJT [KJT]; Loreto: 1 ♂, pond & small stream 34 km W of Loreto (835 m), 14 vi 1995, leg. & det. SWD [JJD] 4 ♂♂, Pano, 12 ix 1990, leg. OSF, det.? [USNM]; SU, Cascales: 2 ♂♂, stream 58 km W of Lago Agrio, El Paraiso (455 m), 8 vi 1995, leg. & det. KJT [KJT]; Lago Agrio: 2 ♂♂, stream, E edge of Lago Agrio, 18 viii 1980, leg. & det. SWD [SWD]; PA, Pastaza: 1.5 km S of Puyo, 19 v 1977, leg. P. Spangler & D. Givens, det.? [USNM]. Macrothemis pseudimitans Calvert, 1898. Publ. Recs. ECU: Muttkowski (1910), Maria (1938), Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14); EO: Campos (1927); ES: Campos (1922); IM: Ris (1913 a), Campos (1922); GY: * Calvert (1906), Campos (1922). Also BRA, CEN, COL, TTO, VEN: (H 06), (P 11); plus PER: (P 16). Addl. Data. IM, Ibarra: 3 ♂♂, “ Guayupe ” [presumably “ Guallupe ”] (- 0.78, - 78.30, 1000 m), 13 ii 1946, leg. L. Alonzo, det. MJW [FSCA]; 1 ♂, Lita, 45.8 km E, rocky stream (- 0.78, - 78.32, 855 m), 6 ii 1997, leg. & det. WFM [FSCA]. Note: It is possible that some of the previous records for this species are actually for the very similar M. fallax, which was not described until 1998. Macrothemis tessellata (Burmeister, 1839). Publ. Recs. ECU: * Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14). Also ARG, BRA, CEN, COL, PRY: (P 11); plus SUR, VEN: (P 16). Addl. Data. NA, Tena: 1 ♂, Río Sinde, 5 km E of Puerto Napo (- 1.0500, - 77.7481, 450 m), 15 i 1995, leg. & det. KJT [KJT]; OR, Francisco de Orellana: 2 ♂♂, P. N. Y., 3.6 km S of Yasuni Road on Waorani Reserve Road (- 0.6956, - 76.4594, 244 m), 10 vi 1995, leg. & det. WFM [FSCA, QZAC]; 1 ♀, P. N. Y., Yasuni Biological Research Sta. along trails and roads within 5 km of Res. Sta. (- 0.6719, - 76.3992, 244 m), 18 vii 1996, leg. & det. WFM [FSCA]; PA, Mera: 1 ♂, Río Anzu 12.0 km N of Mera, 21 ix 2005, leg. & det. JJD [JJD]; SU, Shushufindi: 1 ♂, trails, small streams 3 km W of Shushufindi, 2 + km N off Hwy. 26 at km marker 17 (- 0.1664, - 76.6917, 290 m), 14 x 2009, leg. & det. JTJ [JTJ]; 1 ♂, Limoncocha, 23 viii 1980, leg. & det. SWD [SWD]; 1 ♂, Shushufindi, about 3 km W, small stream, (- 0.1662, - 76.6879), 28 x 2013, leg. & det. KJT [FSCA]. Note: We are treating these specimens from the east slope, previously determined to be M. inequiunguis, as M. tessellata; see also note under M. inequiunguis. Macrothemis spp. unresolved: IM, Cotacachi: 1 ♀, Santa Rosa, El Refugio de Intag Cloud Forest, south along road at roadside seeps (0.3693, - 78.4728, 1854 m), 23 viii 2018, leg. & det. WFM [FSCA]; same data but 1 ♂, roadside seeps and open small open stream (0.3703, - 78.4745, 1866 m), 23 viii 2018 [FSCA]; 2 ♂♂, 2 ♀♀, same data but leg. KJT [KJT]; PC, Quito: 1 ♂, 1 ♀, Mindo (1354 m), 15 v 2011, leg. & det. ACRP [ACRP]; SU, Shushufindi: 1 ♀, Shushufindi road, 2.5 km W of Shushufindi at km marker 18, 3.5 km S, (- 0.2136, - 76.6786, 280 m), 12 v 2009, leg. KJT (specimen labeled M. newtoni?); 1 ♂, 1 ♀, Shushufindi, road, about 3 km NW of Shushufindi, (- 0.1664, - 76.6167, 290 m), 13 v 2009, leg. KJT (both of these specimens were determined by RWG as M. cf. brevidens). Miathyria Kirby, 1889 A genus of just two species, easily distinguished (Ris 1913 b). Both species are found in lentic waters such as ponds and canals with floating aquatic plants, especially water hyacinth and water lettuce; the nymphs hide in the roots. See Fig. 101. Miathyria marcella (Selys in Sagra, 1857). Publ. Recs. ECU: Maria (1938), Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14); CR: Campos (1922); CB: Campos (1922); EO: Campos (1927); GY: * Therese (1900), Calvert (1906), Ris (1913 b), Campos (1922, 1930, 1932); LR: Calvert (1906), Campos (1922). Also ARG, BOL, BRA, CEN, COL, GUF, GUY, PRY, PER, TTO, URY, VEN: (H 06), (P 11); plus SUR: (P 16). Addl. Data. BO, San Miguel: [USNM]; CN, Cañar: [FSCA]; EO, Pasaje: [QZAC]; GY, Duran: [ANSP]; Guayaquil: [ANSP, FSCA, USNM]; Naranjal: [FSCA]; LR, Babahoyo: [USNM]; Buena Fé: [DRP]; Ventanas: [JJD]; MN: Chone: [USNM]; Sucre: [USNM]; SU, Shushufindi: [FSCA]. Miathyria simplex (Rambur, 1842). Publ. Recs. ECU: Ris (1913 b), Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14); EO: Campos (1927); GY: * Calvert (1906), Campos (1922, 1930). Also BRA, CEN, GUF, GUY, PER, VEN: (H 06), (P 11); plus TTO: (P 11); and COL, SUR (P 16). Addl. Data. GY, Guayaquil: 1 ♀, Guayaquil, 18 viii 1917, leg. F. Goding, det.? [USNM]; OR, Francisco de Orellana: 1 ♂, [La Nueva Esperanza], [across Río Napo from], near Primavera “ Limoncocha ”, Río Napo & Lake Taracoa (- 0.45, - 76.77, 300 m), 26 vii 1980, leg. & det. SWD [SWD]; 3 ♂♂, P. N. Y., Río Savaleto 2 & slough at km 31, 17 vii 1996, leg. & det. SWD [SWD]; 2 ♂♂, trib. of Río Payamino, road to Loreto, 1.7 km W of jct. with Coca Road (- 0.4367, - 77.0017, 258 m), 18 xi 1997, leg. & det. KJT [KJT]; 2 ♂♂, same data but leg. & det. WFM [FSCA, QZAC]; 1 ♀, Puerto Francisco de Orellana, 11.8 km W on Loreto Road, large pond (- 0.4742, - 77.0583, 285 m), 18 xi 1997, leg. & det. WFM [FSCA]; 1 ♀, same data but leg. & det. KJT [KJT]; SU, Lago Agrio: 1 ♀, 2 km N of Lago Agrio, 27 viii 1975, leg. A. Langley, det.? [USNM]; Shushufindi: 6 ♂♂, 2 ♀♀, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 6 xi 1980, leg. & det. MJW [FSCA]; 2 ♀♀, same data but 25 xi 1980 [FSCA]; 1 ♀, same data but 3 – 16 vi 1977, leg. P. Spangler & D. Givens, det.? [USNM]; 1 ♂, Limoncocha, clearing, 24 vii 1977, leg. & det. D. Paulson [DRP]; 2 ♂♂, Limoncocha, lakeshore, 26 vii 1977, leg. DRP & S. Hills, det. DRP [DRP]; 1 ♂, same data but 25 viii 1980, leg. & det. SWD [SWD].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B5A3FF63FF23F760201EF8CD.taxon	discussion	A large group of small to medium-sized dragonflies (Fig. 102) in need of revision. Heckman (2006) devised a key to species based on the scattered literature. Twenty-one species have been recorded for Ecuador, plus there are a number of specimens with questionable identity that could increase this number. The group occupies a wide variety of habitats including vegetated lakes, ponds, and sloughs, both open and shaded. Micrathyria aequalis (Hagen, 1861). Publ. Recs. ECU: Muttkowski (1910 — as M. septima and proposed synonymy with M. aequalis), Paulson (1983), Tsuda (1986, 1991), Silsby (2001), (H 06), (P 11), (H 14); GY: * Calvert (1906), Campos (1922). Also CEN, COL, GUF, GUY, SUR, VEN: (H 06), (P 11); plus TTO: (P 11). Addl. Data. EO, Arenillas: 1 ♂, farm pond, seeps, 11 km S of Arenillas, on road to Alamar (- 3.6436, - 80.0989, 95 m), 10 iv 2008, leg. & det. JTJ [JTJ]; 2 ♂♂, same data but leg. & det. KJT [KJT]; Machala: 1 ♀, borrow pit pond, Hwy. 25, S of Machala (- 3.4750, - 79.9778), 10 iv 2008, leg. & det. KJT [KJT]; Santa Rosa: 1 ♂, pond, 12 km E of Arenillas, Hwy. 25 (- 3.5244, - 79.9831, 30 m), 29 iii 2008, leg. & det. KJT [KJT]; 3 ♂♂, same data but leg. & det. JTJ [JTJ]; 2 ♂♂, same data but 9 iv 2008, leg. & det. KJT [KJT]; Piñas: 1 ♀, Piedras (- 3.63, - 79.92, 300 m), 6 vii 1941, leg. D. Laddey, det. KJT [FSCA]; GY, Daule: 1 ♂, Nobol, 5.5 km N, (30 m), 12 i 1978, leg. P. Spangler & J. Anderson, det.? [USNM]; Guayaquil: 1 ♂, Guayaquil, leg. F. Campos, det. MJW [ANSP]; LR, Babahoyo: adults, 20 km N of Babahoyo, 22 vi 1975, leg. J. Cohen, A. Langley & P. Monnig, det.? [USNM]; Ventanas: 1 ♂, pond / stream 7.2 km E of Hwy. E 25, 9 km SW of Quinzaloma (- 1.2539, - 79.3703, 100 m), 21 ii 2010, leg. & det. JTJ [JTJ]; MN, El Carmen: 1 ♂, El Carmen, i 1983, leg. & det.? [QZAC]; 1 ♂, borrow pits about 18 km NW of El Carmen (- 0.1350, - 79.5797, 185 m), 2 iii 2010, leg. & det. JTJ [JTJ]. * Micrathyria artemis Ris, 1911. Publ. Recs. ARG, BRA, GUF, GUY, PER, SUR, VEN: (H 06), (P 16). Addl. Data. NA, Tena: 1 ♂, Puerto Napo, 3.2 km E on Jatun Sacha Road, 23 vii 1996, leg. & det. SWD [FSCA]; SU, Shushufindi: 1 ♂, Limoncocha, lake edge (- 0.4000, - 76.6167, 269 m), 8 ii 1971, leg. D. Pearson, det. DRP [DRP]; 1 ♂, same data but 26 vii 1977, leg. DRP & S. Hills, det. DRP, checked by RWG [DRP]. Micrathyria atra (Martin, 1897). Publ. Recs. ECU: (H 06), (P 11), (H 14). Also ARG, BRA CEN, GUF, GUY, PER, SUR, TTO, VEN: (H 06), (P 11). Addl. Data. MS, Santiago: 1 ♂, [Patuca], [about 1 mi N], 43.4 km S of Sucua, large pond, [about 5 km N of Mendez road jct] (- 2.75, - 78.27, 488 m), 7 xi 1997, leg. & det. KJT [KJT]; 1 ♀, same data but leg. & det. WFM [FSCA]; 3 ♂♂, same data but 8 xi 1997 [FSCA, LSAM, QZAC]; 7 ♂♂, 2 ♀♀, same data but leg. & det. JJD [JJD]; 2 ♂♂, 1 ♀, same data but leg. & det. KJT [KJT]; NA, Tena: 1 ♂, forest pool 14.8 km E of Puerto Napo on J. S. B. S. Road, 15 vi 1995, leg. & det. SWD [JJD]; 1 ♂, same data but leg. & det. KJT [KJT]; 1 ♂, Río Sinde, 5.3 km E of Puerto Napo (- 1.0500, - 77.7481, 450 m), 15 vi 1995, leg. & det. KJT [KJT]; 1 cop. pair, Puerto Napo, 14.8 km E on J. S. B. S. Road, forest pool, 15 vi 1995, leg. & det. SWD [SWD]; 2 ♂♂, 1 ♀, J. S. B. S., Pond # 14, 18 – 23 xi 1991, leg. E. McCallie, det.? [USNM]; OR, Loreto: 2 ♂♂, S of Loreto, forest pond, (- 0.8141, - 77.3371, 350 m), 2 xi 2013, leg. & det. CDK [FSCA]; 1 ♂, N of Loreto, pond, (- 0.8205, - 77.3202), 2 xi 2013, leg. & det. KJT [FSCA]; SU, Cascales: 1 ♂, pool, 58 km W of Lago Agrio, 20 viii 1980, leg. & det. SWD [SWD]; Shushufindi: 26 ♂♂, 2 ♀♀, 1 cop. pair, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 2 – 26 xi 1980, leg. & det. MJW [FSCA]; 3 ♂♂, 2 ♀♀, same data but palm swamp, 25 vii 1977, leg. & det. DRP [DRP]. Micrathyria caerulistyla Donnelly, 1992. Publ. Recs. ECU: (H 06), (P 11), (H 14). Also CEN: (H 06), (P 11). Addl. Data. MS, Santiago: pond along road between Sucua and Mendez (- 2.6833, - 78.2283, 600 m), 6 xi 1997, leg. & det. TWD [SWD]; NA, Tena: 6 ♂♂, Puerto Napo, 3.2 km E on J. S. B. S. Road, ponds and flowing marsh (- 1.0456, - 77.7467, 420 m), 23 vii 1996, leg. & det. WFM [FSCA, JJD, LSAM, QZAC, USNM]; 4 ♂♂, same data but leg. & det. KJT [KJT]; 4 ♂♂, same data but leg. & det. SWD [RWG, SWD]; 3 ♂♂, 5.3 km E of Hwy. 45 on J. S. B. S. Road, Río Sinde (- 1.0500, - 77.7481, 450 m), 16 x 1988, leg. & det. SWD [FSCA, SWD]; 1 ♂, same data but 9 x 2009, leg. & det. JTJ [JTJ]; OR, Francisco de Orellana: 5 ♂♂, Puerto Francisco de Orellana, 1.7 km W on Loreto Road from Coca-Sachas Road, stream, pools & flowing shallow marsh (- 0.4367, - 77.0017, 258 m), 13 vi 1995, leg. & det. WFM [FSCA, QZAC, USNM]; 1 ♂, 1 cop. pair, same data but leg. & det. KJT [KJT]; 2 ♂♂, same data but leg. & det. SWD [SWD]; 1 ♂, same data but 18 xi 1997, leg. & det. WFM [FSCA]; 1 ♂, same data but leg. & det. TWD [SWD]; 2 ♂♂, [FSCA, JJD]; 1 ♂, 12.3 km W of Coca Road on Loreto Road (820), 13 vi 1995, leg. & det. SWD [SWD]; 1 ♂, P. N. Y., 6.2 km W of Yasuni Road on PC 29 - 30, pool / stream, 10 vi 1995, leg. & det. SWD [SWD]; 1 ♀, [Pompeya Sur] S on [Pompeya Sur-Iro road] at km marker 31, borrow pit near Río Sabaleto (- 0.60, - 76.47, 229 m), 17 vii 1996, leg. & det. SWD [SWD]; 1 ♂, P. N. Y., 6.3 km S of Yasuni Road on Waorani Reserve Road, marshy pond (- 0.7120, - 76.4745, 259 m), 10 vi 1995, leg. & det. WFM [FSCA]; 1 ♂, P. N. Y., 2.1 km E of Yasuni Biological Research Station pond and swamp (- 0.6741, - 76.3685, 244 m), 15 xi 1997, leg. & det. WFM [FSCA]; SU, Shushufindi, 1 ♂, Limoncocha, lakeshore & nearby areas (- 0.4000, - 76.6167, 269 m), 4 ix 1978, leg. KWK, det. MJW [FSCA]; La Selva Lodge, Lake Mandicocha & outlet stream (- 0.4983, - 76.3733, 200 m), 25 xi 1997, leg. & det. TWD [SWD]; ZC, Yantzaza: 1 ♂, slow stream 7 km N of El Pangui, 6 xi 1997, leg. & det. KJT [KJT]. Micrathyria catenata Calvert, 1909. Publ. Recs. ECU: (H 06), (P 11), (H 14); ES: Daigle et al. (1997); NA: Tennessen (1996 b); SU: * Westfall (1992). Also ARG, BRA, CEN, GUF, PER, SUR, TTO, VEN: (H 06), (P 11); plus COL: Palacino et al. (2017). Addl. Data. ES: San Lorenzo: [FSCA, JJD, KJT]; MS, Santiago: [FSCA, JJD, KJT, LSAM, QZAC, TWD, USNM]; Taisha: [DRP]; NA, Archidona: [FSCA]; Carlos Julio Arosemena Tola: [FSCA]; Chaco: (KWN), Tena: [DRP, FSCA, JJD, KJT, SWD, USNM]; OR, Francisco de Orellana: [FSCA, KWK, KJT, LSAM, QZAC, RWG, SWD, TWD, USNM]; SU, Cascales: [SWD]; Shushufindi: [FSCA, KJT]; PA, Pastaza: [FSCA, KJT, QZAC, USNM]. Micrathyria dictynna Ris, 1911. Publ. Recs. ECU: (H 06), (P 11), (H 14); MS: Tennessen (2006); OR: von Ellenrieder & Garrison (2011); SU * Donnelly (1998). Also CEN, VEN: (H 06), (P 11); plus COL: (P & P). Addl. Data. MS, Huamboya: 2 ♂♂, small stream w / pools, forested, near Río Pasanac [57 km N of Macas], Hwy. 45, N of river, E side of Hwy. (- 1.9550, - 77.8567), 16 ix 2005, leg. & det. JJD [JJD]; 1 ♂, 1 ♀, same data but leg. & det. KJT [KJT]; NA, Tena: 2 ♂♂, J. S. B. S., pond # 13, 23 xi 1991, leg. E. McCallie, det.? [USNM]; 1 ♂, Río Sinde, pond, 5.3 km E of Puerto Napo (- 1.0517, - 77.7544, 450 m), 13 vii 1996, leg. & det. KJT [KJT]; 1 ♂, Río Chinchipino, oxbow pool, 27 km E of Puerto Napo (- 1.0728, - 77.5864, 385 m), 13 xi 1997, leg. & det. KJT [KJT]; OR, Francisco de Orellana: 1 ♀, Yuturi, 2 iii 1990, leg. & det.? [QZAC]; 2 ♂♂, T. B. S., USFQ, Guaca- mayo trail between markers 450 – 2100 (- 0.6350, - 76.1533, 237 m), 2009, leg. & det. RWG & NVE [NVE, RWG]; Loreto: 1 ♂, forest about 12 km NE of Loreto (- 0.6217, - 77.2600, 365 m), 15 x 2009, leg. KJT [KJT]; same data but 1 ♂, 3 xi 2013, leg. & det. KJT [FSCA]; 1 ♂, S of Loreto, forest pond, (- 0.8141 - 77.3371, 350 m), 2 xi 2013, leg. & det. CDK [FSCA]; PA, Mera: 1 ♂, seepage stream, Los Copales Cabanas between Mera and Shell, W of Puyo, Hwy. 50, 22 ix 2005, leg. & det. JJD [JJD]; SU, Lago Agrio: 1 ♂, swamp, 17 km S of Lago Agrio, 19 viii 1980, leg. & det. SWD [SWD]; Shushufindi: 1 ♀, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 11 ix 1973, leg. B. Drummond III, det.? [FSCA]; same data but forest edge, 25 vii 1977, leg. & det. DRP [DRP]; 1 ♂, stream about 3 km NW of Shushufindi (- 0.1664, - 76.6917, 290 m), 13 v 2009, leg. & det. KJT [KJT]; 1 ♂, La Selva Lodge; forest (- 0.4983, - 76.3733, 200 m), 22 xi 1997, leg. & det. TWD [SWD]; 1 ♀, La Selva, Manicocha, 12 x 1988, leg. & det. SWD [SWD]; 1 ♂, 1 ♀, La Selva, Garzacocha, 12 x 1988, leg. & det. SWD [SWD]. Micrathyria didyma (Selys in Sagra, 1857). Publ. Recs. ECU: Paulson (1983), Tsuda (1986, 1991), Steinmann (1997 b as M. didyma didyma), Silsby (2001), (H 06), (P 11), (H 14); EO: Campos (1927); GY: * Calvert (1906), Ris (1911 a), Campos (1922); IM: Ris (1911 a — as M. didyma didyma), Campos (1922). Also CEN, COL, GUF, GUY, SUR, VEN: (H 06), (P 11); plus BRA: (H 06); plus TTO: (P 11). Addl. Data. GY, Guayaquil: 1 ♂, Guayaquil (- 2.20, - 79.90, 8 m), date?, leg. F. Campos, det. P. Calvert, checked WFM [FSCA]. Micrathyria hippolyte Ris, 1911. Publ. Recs. ECU: (H 06), (P 11), (H 14); NA: * Tennessen (1996 b). Also BRA, GUF, GUY, PER, SUR, VEN: (H 06), (P 11). Addl. Data. NA, Tena: 1 ♂, [Puerto Napo], [upstream near jct with Río Mulatos] Jatun Yacu Río Napo watershed (- 1.05, - 77.98, 700 m), 25 ii 1937, leg. WCM, det.? [FSCA]; 2 ♂♂, pond, road to J. S. B. S., 14.8 km E of Puerto Napo, 15 vi 1995, leg. & det. KJT [KJT]; 2 ♂♂, 1 ♀, same data but leg. & det. SWD [SWD]; SU, Shushufindi: 1 ♂, Limoncocha (- 0.4000, - 76.6176, 269 m), 28 viii 1980, leg. & det. SWD [SWD]; 1 ♀, same data but forest trail, 27 ix 1971, leg. D. Pearson, det. DRP [DRP]; 1 ♀, same data but 24 i 1972 [DRP]; 1 ♂, same data but 19 ii 1972 [FSCA]; 1 ♂, same data but forest stream, 9 iii 1972 [DRP]; 1 ♂, same data but forest edge, 24 vii 1977, leg. & det. DRP [DRP]. Micrathyria iheringi Santos, 1946. Publ. Recs. ECU: * Daigle et al. (1996), (H 06), (P 11), (H 14). Also BRA: (H 06), (P 11). Addl. Data. MS, Huamboya: 1 ♂, Macas, 58 km N on Hwy. 45 (70 km N of Sucua), roadside ditch and tribs. near Río Pasanac (- 1.9447, - 77.8536, 762 m), 10 xi 1997, leg. & det. WFM [FSCA]; 1 ♂, (labeled “ PA ”), small stream about 1 km S of Río Pasanac (- 1.9503, - 77.8631, 820 m), 16 ix 2005, leg. & det. KJT [FSCA]; 1 ♂, same data but 16 ix 2005 [KJT]; NA, Tena: 1 ♂, Tena (- 0.98, - 77.8, 500 m), date?, leg. F. Campos, det. KJT [FSCA]; 1 ♂, J. S. B. S., 18 x 1988, leg. & det. SWD [SWD]; 1 ♂, same data but Pond # 8, 25 xi 1991, leg. E. McCallie, det.? [USNM]; 2 ♂♂, Río Sinde, slough; 5.3 km E Puerto Napo (- 1.0500, - 77.7481, 450 m), 16 x 1988, leg. & det. SWD [RWG, SWD]; 1 ♂, same data but 13 vii 1996, leg. & det. SWD [SWD]; 4 ♂♂, pool near Río Napo, 3.2 km E of Puerto Napo (- 1.0456, - 77.7467, 420 m), 23 vii 1996, leg. & det. KJT [KJT]; 1 ♂, same data but leg. & det. SWD [SWD]; 1 ♂, Puerto Napo, 17.2 km E on J. S. B. S. Road, pond near stream (340 m), 22 vii 1996, leg. & det. SWD [SWD]; 1 ♂, Puerto Napo, 2 nd stream E of Río Sinde on J. S. B. S. Road, 23 vii 1996, leg. & det. SWD [SWD]; OR, Francisco de Orellana: 1 ♂, Comunidad El Rocio, E on Rt E 20, pond, (- 0.5056, - 77.1566), 27 x 2013, leg. & det. KJT [FSCA]; Loreto: 3 ♂♂, Pano, 12 ix 1990, leg. OSF, det.? [USNM]; SU, Shushufindi: 1 ♂, trails, small streams 3 km W of Shushufindi, 2 + km N off Hwy. 26 at km marker 17 (- 0.1664, - 76.6917, 290 m), 13 x 2009, leg. & det. JTJ [JTJ]; ZC, Zamora: 3 ♂♂, Zamora (- 4.07, - 78.95, 1000 m), 21 x 1941, leg. D. Laddey, det. JTJ [FSCA]. Micrathyria laevigata Calvert, 1909. Publ. Recs. ECU: Ris (1911 — as M. didyma laevigata), Paulson (1983), Tsuda (1986, 1991), Steinmann (1997 b — as M. didyma laevigata), (H 06), (P 11), (H 14); SD: Daigle et al. (1997). Also CEN, COL, PER, VEN, TTO, BRA: (H 06), (P 11). Addl. Data. OR, Francisco de Orellana: 1 ♂, [La Nueva Esperanza] [across Río Napo from] near Primavera “ Limoncocha ” Río Napo & Lake Taracoa (- 0.45, - 76.77, 300 m), 4 xi 1980, leg. & det. MJW [FSCA]; La Joya de los Sachas: 2 ♂♂, 2 ♀♀, Pompeya. 5 km W of Puerto Pompeya Ferry Landing, roadside ditches (- 0.4444, - 76.6263, 238 m), 19 vii 1996, leg. & det. WFM [FSCA, LSAM, QZAC]; 1 ♂, same data but leg. & det. JJD [JJD]; 3 ♂♂, same data but leg. & det. KJT [KJT]; SU, Shushufindi: 3 ♂♂, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 5 xi 1980, leg. David Robinson & MJW, det. MJW [FSCA]; 2 ♂♂, 4 ♀♀, same data but clearing, forest edge, swamp & lakeshore, 24 – 26 vii 1977, leg. & det. DRP [DRP]. Micrathyria mengeri Ris, 1919. Publ. Recs. ECU: * Dunkle (1995), (H 06), (P 11), (H 14). Also BRA, CEN, COL, GUF, GUY, PER, SUR, TTO, VEN: (P 11). Addl. Data. SU, Shushufindi: 2 ♂♂, Limoncocha, swamp, (- 0.4000, - 76.6167, 269 m), 26 vii 1977, leg. & det. DRP [DRP]; 3 ♂♂, same data but swamp channel, 25 viii 1980, leg. & det. SWD [RWG, SWD]. Micrathyria occipita Westfall, 1992. Publ. Recs. ECU: (H 06), (P 11), (H 14); NA: Tennessen (1996 b); OR: * Westfall (1992, Holotype & Allotype, 1 cop. pair, ” Lake Taracoa near Primavera ” = [Francisco de Orellana], [La Nueva Esperanza] [across Río Napo from] near Primavera “ Limoncocha ” Río Napo & Lake Taracoa (- 0.45, - 76.77, 300 m), 26 viii 1980, leg. SWD [FSCA], Bridges (1993), von Ellenrieder & Garrison (2011). Also PER: (H 06), (P 11); plus VEN: (P 11). Addl. Data. NA, Tena: [FSCA, UTIC, KJT, QZAC, TWD, USNM]; OR: Francisco de Orellana [DRP, FSCA, JJD, KJT, KWN, LSAM, QZAC, RWG, TWD, USNM]; SU, Lago Agrío: [DRP, FSCA, KJT, USNM]; Shushufindi: [FSCA, KJT, QZAC, RWG, USNM]; ZC, Zamora: [JJD]. Note: An east slope species; specific data for most records can be found in Westfall (1992). Micrathyria ocellata Martin, 1897. Publ. Recs. ECU: Muttkowski (1910), Paulson (1983), Tsuda (1986, 1991), Steinmann (1997 b — as M. ocellata ocellata), (H 06), (P 11), (H 14); CR: Calvert (1906), GY: * Martin (1897 — TYPE, Guayaquil, date?); Bridges (1993), Calvert (1906), (Ris, 1911 a — as M. ocellata ocellata), EO: Campos (1927); NA: Tennessen (1996 b). Also ARG, BOL, BRA, CEN, COL, GUF, GUY, PER, SUR, TTO, VEN: (H 06), (P 11). Addl. Data. EO: Machala: [KJT]; Quininde: [KWK, FSCA]; Santa Rosa [JTJ, KJT]; GY, Daule: [USNM]; Guayaquil: [FSCA, USNM]; LR: Quevedo: [FSCA]; MN, El Carmen: [JTJ]; Pedernales: [JTJ]; MS, Santiago: [SWD]; NA, Tena: [FSCA, JJD, KJT, LSAM, QZAC, SWD, USNM]; OR, Francisco de Orellana: [KJT, QZAC]; SU. Lago Agrío: [USNM]; Shushufindi: [DRP, FSCA, KJT, KWN, SWD]. Micrathyria pseudeximia Westfall, 1992. Publ. Recs. ECU: (H 06), (P 11), (H 14); SU: * Westfall (1992 — description). Also BOL, BRA, CEN, COL, GUF, PRY, PER, SUR, VEN, TTO: (H 06), (P 11); plus ARG: (P 11). Addl. Data. LR, Buena Fé: 7 ♂♂, 1 ♀, 1 cop. pair, Santo Domingo, 47 km S, forest slough at Río Palenque Field Station (200 m), 18 vii 1977, leg. & det. DRP [DRP]; 8 ♂♂, 1 ♀, paratypes, same data but 8 – 9 x 1988, leg. & det. SWD [SWD]; MN, El Carmen: 2 ♂♂, borrow pits about 18 km NW of El Carmen (- 0.1350, - 79.5797, 185 m), 2 iii 2010, leg. & det. JTJ [JTJ]; NA, Tena: 1 ♂, Puerto Napo 5.3 km E on Jatun Sacha Road Río Sinde swamp and pond approx. ½ km upstream (- 1.0517, - 77.7544, 450 m), 12 xi 1997, leg. & det. WFM [FSCA]; OR, Francisco de Orellana: 1 ♂, [Pompeya Sur] S on [Pompeya Sur-Iro Road] at km marker 31, borrow pit near Río Sabaleto (- 0.60, - 76.47, 238 m), 17 vii 1996, leg. & det. SWD [SWD]; 1 ♂, pond, road to Waorani Reserve, 6.2 km S of turn to Yasuni Sci. Res. Sta., 10 vi 1995, leg. & det. WFM [FSCA]; 1 ♂, swamp, P. N. Y. (- 0.6508, - 76.3558, 200 m), 16 xi 1997, leg. & det. KJT [KJT]; 1 ♂, 1 ♀, same data but leg. & det. TWD [SWD]; 1 ♂, stream 2.3 km E of Yasuni Sci. Res. Sta., 9 vi 1995, leg. & det. SWD [SWD]; 1 ♂, Puerto Francisco de Orellana 1.7 km W on Loreto Road from Coca-Sachas Road, stream, pools & flowing shallow marsh (- 0.4367, - 77.0017, 258 m), 13 vi 1995, leg. & det. SWD [SWD]; 1 ♂, pond 8.2 km W of Río Payamino, Coca, 18 xi 1997, leg. & det. JJD [JJD]; 1 paratype ♂, [La Nueva Esperanza] [across Río Napo from], near Primavera “ Limoncocha ”, Río Napo & Lake Taracoa (- 0.45, - 76.77, 300 m), 26 viii 1980, leg. & det. SWD [SWD]; Loreto: 1 ♂, S of Loreto, forest pond, (- 0.8141, - 77.3371, 350 m), 2 xi 2013, leg. & det. CDK [FSCA]; PC, Quito: 2 ♂♂, Pacto Parriquia, San Jose de Mashpi, SE on Mashpi road, small marshy pond & adjacent Tilapia ponds (0.1676, - 78.9024, 542 m), 3 ix 2018, leg. & det. KJT [KJT, USFQ]; SD: 1 paratype ♂, Santo Domingo de Los Colorados, 12 km E on Hwy. 30 near Hotel Tinalandia, pond with water lilies (- 0.30, - 79.67, 610 m), 6 x 1988, leg. & det. SWD [SWD]; 1 ♂, Al- luriquin, 7.3 km W, Hotel Tinalandia, 11 iii 1990, leg. C. Anderson, det. DRP [DRP]; SU, Lago Agrio: 1 paratype ♂, brushy ponds, 24 km W of Lago Agrio, 20 viii 1980, leg. & det. SWD [SWD]; 2 ♂♂, swamp, 17 km S of Lago Agrio, 19 viii 1980, leg. & det. SWD [SWD]; Shushufindi: 2 ♂♂, Limoncocha, swamp, 26 vii 1977, leg. DRP & S. Hills, det. DRP [DRP]; 1 paratype ♂, same data but 25 viii 1980, leg. & det. SWD [SWD]; 1 paratype ♂, [Limoncocha], [about 40 km E], or 100 km E of Coca [on Río Napo], Lago Garzacocha, La Selva Jungle Lodge (- 0.4819, - 76.3733, 200 m), 12 x 1988, leg. & det. SWD [SWD]. Micrathyria romani Sjöstedt, 1918. Publ. Recs. ECU: (H 06), (P 11), (H 14); SU: * Dunkle (1981), Westfall (1992). Also BRA, PER: (H 06), (P 11). Addl. Data. SU, Shushufindi: 3 ♂♂, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 3 – 16 vi 1977, leg. P. Spangler & D. Givens, det.? [USNM]; 23 ♂♂, 2 ♀♀, same data but 1 – 25 xi 1980, leg. & det. MJW [FSCA]; 1 ♂, same data but forest edge, 26 vii 1977, leg. & det. DRP [DRP]; 43 ♂♂, same data but lake edge, 26 xii 1971, D. Pearson, det. DRP [DRP]; 16 ♂♂, same data but 26 vii 1977, leg. DRP & S. Hills, det. DRP [DRP]; 1 ♂, same data but 11 vii 1977, leg. & det. KWK [SWD]; 1 ♀, same data but 29 viii 1978, det. JJD [FSCA]; 2 ♂♂, same data but 23 viii 1980, leg. & det. SWD [KJT]; 5 ♂♂, [Limoncocha], [about 40 km E], or 100 km E of Coca [on Río Napo], Lago Garzacocha, La Selva Jungle Lodge (- 0.4819, - 76.3733, 200 m), 11 x 1988, leg. & det. SWD [FSCA, SWD]; 1 ♂, same data but in forest, 6 v 1990, leg. & det. E. Smith [SWD]. Micrathyria spinifera Calvert, 1906. Publ Recs. ECU: * Paulson (1983), (H 06), (P 11), (H 14). Also BOL, BRA, GUF, GUY, PER, SUR, TTO, VEN: (H 06), (P 11). Addl. Data. NA, Archidona: 1 ♂, along Archidona-Loreto Road, stream, pasture to forested, (- 0.7164, - 77.6640, 1185 m), 25 x 2013, leg. & det. CDK [FSCA]. OR, Francisco de Orellana: 1 ♂, [La Nueva Esperanza] [across Río Napo from], near Primavera “ Limoncocha ”, Río Napo & Lake Taracoa (- 0.45, - 76.77, 300 m), 27 vii 1977, leg. DRP & S. Hills, det. DRP [DRP]; 1 ♂, same data but 4 xi 1980, leg. & det. MJW [FSCA]; La Joya de los Sachas: 1 ♂, road at Río Napo ferry, S of Limoncocha, 17 vii 1996, leg. & det. KJT [KJT]; SU, Shushufindi: 1 ♂, Limoncocha, Río Napo (- 0.4000, - 76.6167, 236 m), 6 ix 1972, leg. T. Emmel, det. JTJ [FSCA]; 1 ♂, same data but forest trail, 10 viii 1971, leg. D. Pearson, det. DRP [DRP]; 1 ♂, same data but 24 vii 1977, leg. & det. DRP [DRP]; 7 ♂♂, 1 ♀, same data but forest edge, 25 – 26 vii 1977 [DRP]; 1 ♂, same data but lake edge, 2 x 1971, leg. D. Pearson, det. DRP [DRP]; 1 ♂, same data but 29 viii 1978, leg. KWK, det. JJD [KWK]; 2 ♂♂, Limoncocha, small clearing near Playaco River, 24 viii 1980, leg. & det. SWD [SWD]. Micrathyria sympriona Tennessen, 2000. Publ. Recs. ECU: (H 06), (P 04), (P 11), (H 14); ZC: * Tennessen (2000 — Holotype & Allotype: cop. pair: [Zamora], cutgrass marsh 5.5 km SE of Zamora (- 4.17, - 78.94, 914 m), 5 xi 1997, leg. WFM [FSCA]). Also PER: (H 06), (P 11); plus COL: Palacino et al. (2017). Addl. Data. MS, Gualaquiza: 2 ♂♂, 1 ♀, Gualaquiza (- 3.40, - 78.58, 800 m), iv – v 1935, leg. F. Campos, det. WFM [ANSP, FSCA]; ZC, Zamora: 2 paratype ♂♂, Río Jamboe slough 4.2 km SE of Zamora, 5 xi 1997, leg. JJD, det. KJT [JJD]; 1 paratype ♂, 1 paratype ♀, cutgrass marsh 5.5 km SE of Zamora (914 m), 5 xi 1997, leg. JJD, det. KJT [JJD]; 6 paratype ♂♂, same data but leg. & det. KJT [KJT]; 7 ♂♂, same data but leg. WFM [FSCA, LSAM, QZAC, USNM]. 1 ♂, small stream 5 km N of Guaguayme Alto (- 3.8778, - 78.8928, 880 m), 4 iv 2008, leg. & det. JTJ [JTJ]. Note: The Gualaquiza specimens were labeled M. didyma. Micrathyria tibialis Kirby, 1897. Publ. Recs. ECU: * Paulson (1983), (H 06), (P 11), (H 14). Also ARG, BOL, BRA, CEN, GUY, PRY, PER, SUR, VEN: (H 06), (P 11); plus COL: (P & P); and GUF: (P 16). Addl. Data. OR, Francisco de Orellana: 14 ♂♂, 2 ♀♀, Anangu [SE across Río Napo from Providencia], Lago Anangu “ on Río Napo ” (- 0.5213, - 76.4361, 300 m), 9 v 1975, leg. B. Drummond III, det.? [FSCA]; SU, Shushufindi: 1 ♀, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 8 vi 1977, leg. S. Steiner & V. Warren Jr, det.? [USNM]; 13 ♂♂, same data but 3 – 16 vi 1977, leg. P. Spangler & D. Givens, det.? [USNM]; 1 ♂, same data but 25 viii 1980, leg. & det. SWD; 1 ♂, same data but 25 viii 1980 [SWD]; same data but 6 – 25 xi 1980, leg. & det. MJW [FSCA]; 1 ♂, same data but lake edge, 22 i 1972, leg. D. Pearson, det. DRP [DRP]; 2 ♂♂, same data but 16 ii 1972 [DRP]; 10 ♂♂, same data but 24 – 26 vii 1977, leg. & det. DRP [DRP]; 1 ♂, same data but 3 ix 1980, leg. & det. KWK [SWD]; 1 ♂, same data but 25 viii 1980, leg. KWK, det. JJD [FSCSA]. Micrathyria venezuelae De Marmels, 1989. Publ. Recs. ECU: * (H 06), (P 11). Also VEN: (H 06), (P 11); plus (ARG): (P 16); and COL: Palacino et al. (2017). Addl. Data. EO, Portovelo: 1 ♂, Portovelo (- 3.72, - 79.62, 600 – 650 m), 5 viii 1941, leg. D. Laddey, det. JJD [FSCA]; ES, Quininde: 1 ♂, Rosa Zarate, Río Cupa (0.42, - 79.42, 90 m), 10 i 1972, leg. D. Poveda, det. JJD [FSCA]; GY, Guayaquil: 1 ♂, Guayaquil (- 2.20, - 79.90, 8 m), date?, leg. N. Banks, det. JJD [FSCA]; LJ, Loja: 1 ♂, seep trib. of Río Chamba, Vilcabamba (- 4.2636, - 79.2172, 1555 m), 31 iii 2008, leg. & det. JTJ [JTJ]; LR, Buena Fé: 1 ♂, Santo Domingo, 47 km S, forest slough at Río Palenque Field Station (200 m), 18 vii 1977, leg. DRP, det. WFM [FSCA]; 6 ♂♂, 8 – 9 xi 1988, leg. & det. SWD [FSCA]; Urdaneta: 1 ♂, ditch / wooded area, 2.8 km E of Hwy. E 25 from km marker 98 (- 1.5217, - 79.5000, 35 m), 21 ii 2010, leg. & det. KJT [FSCA]; 2 ♀♀, 5 km E of Hwy. E 25 (from Formisa), small stream / secondary forest, (- 0.7253, - 79.4444), 23 ii 2010, leg. & det. KJT (FSCA); MN, Manta: 1 ♂, Santa Rosa, about 60 km SW of Manta near Río Seco (- 1.12, - 80.88, 40 m), 11 v 1946, leg. L. Alonzo, det. JJD [FSCA]; PA, Mera: 1 ♂, small streams / trails, Los Copales, between Mera and Shell (- 1.4919, - 78.0725, 1070 m), 20 ix 2005, leg. & det. KJT [KJT]; 1 ♂, same data but 22 ix 2005, leg. & det. JJD [JJD]; 1 ♂, Río Anzu, 12.0 km N of Mera (- 1.4181, - 78.0528, 1190 m), 21 ix 2005, leg. & det. KJT [KJT]; SD: 3 ♂♂, seeps at Hotel Colorados, Hwy. 30, about 12 km E of Santo Domingo, 30 i 1997, leg. & det. JJD [JJD]; 2 ♂♂, rocky pool, 8 km W of Alluriquin, Hwy. 30, 31 i 1997, leg. & det. JJD [JJD]. Micrathyria unresolved: a. Micrathyria cf. almeidai Santos, 1945. Publ. Recs. ECU: * (H 14). Also BRA: (H 06), (P 16). SU, Shushufindi: 1 ♂, Limoncocha, no addl. data provided, data provided by WAH & DLW). According to W. A. Haber (pers. comm. 2016), “ It likely is not almeidai, although it keys to almeidai in Heckman (2006), but the appendages are different. ” b. Micrathyria cf. eximia: Kirby, 1897. Publ. Recs. ECU: (H 14). Also COL, ARG, BRA: (H 06), (P 16). NA, Tena: 1 ♂, Puerto Napo, 5.3 km E, Río Sinde (- 1.0500, - 77.7481, 450 m), 16 x 1988, leg. & det. SWD [RWG]. According to Rosser Garrison (pers. comm. 2011), “ Probably a new species. ” c. Micrathyria cf. surinamensis: Geijskes, 1963. Publ Recs: GUF, SUR: (H 06, P 16) SU: Shushufindi: 1 ♂, La Selva, 100 km E Coca, Garzacocha (- 0.4983, - 76.3733, 200 m), 12 x 1988, leg. & det. SWD [RWG], data provided by RWG. d. Micrathyria sp.: ZC, Zamora: 1 ♂, shaded roadside ditch / pool, about 10 – 11 km N of Guaguayme Alto (- 3.8356, - 78.8939, 875 m), 4 iv 2008, leg. & det. JTJ [JTJ]. e. Micrathyria spp. (H 14): “ Three different species all in the laevigata group which needs to be revised; one of which is a new species according to Paulson and Garrison ” (Haber & Wagmer pers. comm. 2016).	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B5A5FF60FF23F6882125FA17.taxon	discussion	One of the four known species of this genus occurs in Ecuador (Fig. 103). Geijskes (1951) provided a key, which includes M. parana. Adults have been found flying over small, slow forested streams (pers. obs.), but the nymphs of the genus are unknown. Misagria parana Kirby, 1889. Publ. Recs. ECU: Geijskes (1951, 1971), Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14); PA: * Ris (1910). Also BRA, GUF, GUY, PER, SUR, VEN: (H 06), (P 11); plus COL (P 16). Addl. Data. OR, Francisco de Orellana: 1 ♂, 1 ♀, Reserva Etnica Waorani, 500 m N of Onkone Gare Camp (- 0.63, - 76.6), 10 – 13 i 1994, leg. M. Pogue, det.? [USNM]; Reserva Etnica Waorani, 1 km S of Onkone Gare Camp, fogging, Lot # 937 (- 0.6528, - 76.4333, 220 m), 7 x 1994, leg. T. Erwin, det. WFM [USNM]; 1 ♀, same data but 10 x 1994 [USNM]; 1 ♂, T. B. S. (- 0.6340, - 76.1499, 247 m), 13 xii 2012, leg. & det. ACRP [ACRP]; PA, Mera: 1 ♂, between Mera and Shell, Cabinas Los Copales along Río Metolo, trails and forest N of Río Metolo (- 1.4919, - 78.0725, 1070 m), 19 xi 2005, leg. & det. WFM [FSCA]; 1 ♂, same data but 22 ix 2005 [LSAM]; 1 ♂, same data but 7 x 2009, leg. & det. KJT [FSCA]; 2 ♂♂, same data but leg. & det. JTJ [JTJ].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B5A5FF6FFF23F3E22654FE13.taxon	discussion	Four of the seven known species of this genus occur in Ecuador. For species identification, see Santos (1950), Rácenis (1953) and von Ellenrieder (2014). These are small dragonflies, mostly black with yellow markings (Fig. 104); adults perch at open to sparsely shaded, vegetated ponds and marshes. Nephepeltia flavifrons (Karsch, 1889). Publ. Recs. ECU: (H 14); OR * — as “ Napo ” von Ellenrieder (2014). Also ARG, BRA, COL, GUF, PRY, PER, SUR, VEN: (P 11); also CEN, BOL: von Ellenrieder (2014). Addl. Data. OR, Francisco de Orellana: 6 ♂♂, Anangu SE across Río Napo from Providencia Lago Anangu “ on Río Napo ” (- 0.5256, - 76.4394, 300 m), 9 v 1975, leg. B. Drummond III, det. MJW [FSCA]; 1 ♂, same data but 12 v 1975, leg. T. Emmel, det. MJW [FSCA]. Nephepeltia flavipennis von Ellenrieder, 2014. Publ. Recs. ECU: (H 14); SU * von Ellenrieder (2014 — 1 paratype ♂, Baeza-Lago Agrío road, 53 km NE Chaco, ponds (0.0167, - 76.3167), 18 viii 1980, leg. SWD [FSCA]). Also BRA, PER: von Ellenrieder (2014). Note: This recently described species is known in Ecuador from the one paratype specimen which was collected near a vegetated pond (von Ellenrieder 2014). Nephepeltia leonardina Racenis, 1953. Publ. Recs. ECU: (H 06), (P 11), (H 14); NA: * Tennessen (1996 b), Daigle (1996), von Ellenrieder (2014), OR: von Ellenrieder (2014), SU: Donnelly (1998), von Ellenrieder (2014). Also PER, VEN: (H 06), (P 16); plus ARG, BOL, BRA, PRY: von Ellenrieder (2014). Addl. Data. MS, Santiago: 1 ♂, pond along road between Tayuza and Mendez (- 2.7283, - 78.2700, 600 m), 6 xi 1997, leg. & det. TWD [SWD]; NA, Tena: 1 ♂, [Puerto Napo], [upstream near jct with Río Mulatos] Jatun Yacu, Río Napo watershed (- 1.05, - 77.98, 700 m), iii 1937, leg. WCM, det. JJD [FSCA]; 2 ♂♂, same data but 24 ii 1937 [FSCA]; adult, 1 km S of Tena, 26 – 28 v 1977, leg. P. Spangler & D. Givens, det.? [USNM]; 2 ♂♂, same data but 8 km S of Tena [USNM]; 1 ♂, J. S. B. S., near Tena, large pond (- 1.0500, - 77.7833, 450 m), leg. A. Jaramillo, det. DRP [DRP]; 5 ♂♂, Puerto Napo, 23 km E at J. S. B. S., tropical wet forest (- 1.07, - 77.60, 450 m), 18 x 1988, leg. & det. SWD [FSCA, UTIC, SWD]; 1 ♀, Puerto Napo, 5.3 km E on Jatun Sacha Road, Río Sinde, swamp and pond approx. 1 – 2 km upstream (- 1.0517, - 77.7544, 450 m), 15 vi 1995, leg. & det. KJT [KJT]; 1 ♂, same data but 12 xi 1997 [KJT]; 3 ♂♂, same data but 22 vii 1996, leg. & det. WFM [FSCA, QZAC, USNM]; 2 ♂♂, same data but 12 xi 1997 [FSCA]; 1 ♂, same data but 12 xi 1997, leg. & det. TWD [SWD]; 1 ♂, Puerto Napo, 3.2 km E on J. S. B. S. Road, ponds and flowing marsh (- 1.0456, - 77.7467, 420 m), 23 vii 1996, leg. & det. WFM [FSCA]; 2 ♂♂, same data but leg. & det. SWD [SWD]; 1 ♂, Puerto Napo, 2 nd stream E of Río Sinde on J. S. B. S. Road, 23 vii 1996, leg. & det. SWD [SWD]; OR, Francisco de Orellana: 3 ♂♂, Puerto Francisco de Orellana, 1.7 km W on Loreto Road from Coca-Sachas Road, stream, pools &, flowing shallow marsh (- 0.4367, - 77.0017, 258 m), 13 vi 1995, leg. & det. WFM [FSCA, LSAM, QZAC]; 1 ♂, same data but leg. & det. KJT [KJT]; 4 ♂♂, Puerto Francisco de Orellana, 11.8 km w on Loreto Road, large pond (- 0.4742, - 77.0583, 285 m), 18 xi 1997, leg. & det. WFM [FSCA, QZAC, USNM]; 1 ♂, same data but leg. & det. KJT [KJT]; 2 ♂♂, 1 ♀, Puerto Francisco de Orellana, 12.3 km W on road to Loreto from Coca-Sachas Road, pond (- 0.4750, - 77.0747, 290 m), 13 vi 1995, leg. & det. WFM [FSCA]; 1 ♂, 1 ♀, same data but leg. & det. KJT [KJT]; 2 ♂♂, same data but leg. & det. SWD [SWD]; 3 ♂♂, [Pompeya Sur] S on [Pompeya Sur-Iro road] at km marker 31, borrow pit near Río Sabaleto (- 0.60, - 76.47, 229 m), 17 vii 1996, leg. & det. WFM [FSCA, QZAC, USNM]; 1 ♂, same data but leg. & det. KJT [KJT]; 2 ♂♂, same data but leg. & det. SWD [UTIC]; 1 ♂, same data but 17 xi 1997, leg. & det. JJD [JJD]; 2 ♂♂, 1 ♀, same data but leg. & det. TWD [SWD]; 1 ♂, pond, P. N. Y., km marker 29.5, 17 xi 1997, leg. & det. KJT [KJT]; 2 ♂♂, swamp pond at km 8, S of Río Napo on Yasuni road, 11 vi 1995, leg. & det. SWD [JJD]; 1 ♀, P. N. Y., swamp (- 0.6508, - 76.3558, 200 m), 16 xi 1997, leg. & det. TWD [SWD]; 1 ♂, P. N. Y., 2.1 km E of Yasuni Biological Research sta., pond and swamp (- 0.6741, - 76.3685, 244 m), 15 xi 1997, leg. & det. WFM [FSCA]; 2 ♂♂, [La Nueva Esperanza] [across Río Napo from], near Primavera “ Limoncocha ”, Río Napo & Lake Taracoa (- 0.45, - 76.77, 300 m), 4 xi 1980, leg. & det. MJW [FSCA]; 2 ♂♂, same data but 26 viii 1980, leg. & det. SWD [SWD]; SU, Shushufindi: 1 ♂, 15.7 km S of Shushufindi on Limoncocha road, swamp / marsh / pond, 19 vii 1996, leg. & det. SWD [SWD]; 2 ♂♂, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 3 – 16 vi 1977, leg. P. Spangler & D. Givens, det.? [USNM]; 1 ♂, same data but 6 – 25 xi 1980, leg. & det. MJW [FSCA]; 2 ♂♂, 1 ♀, same data but forest edge, 23 vii 1977, leg. & det. DRP [DRP]; 9 ♂♂, same data but lakeshore, 24 – 26 vii 1977, leg. & det. DRP [DRP]; 1 ♂, 1 ♀, same data but 1 ix 1978, leg. KWK, det. JJD [FSCA, SWD]; 1 ♂, La Selva Lodge; Lake Mandicocha and outlet stream (- 0.4983, - 76.3733, 200 m), 25 xi 1997, leg. & det. TWD [SWD]; 1 ♂, La Selva, inlet to Heron Lake, 14 iv 1992, leg. E. Smith, det. SWD [SWD]; 1 ♂, pond, about 3 km NW of Shushufindi (- 0.1822, - 76.6917, 265 m), 14 v 2009, leg. & det. KJT [FSCA]; 1 ♂, same data but 14 x 2009; 1 ♂, same data but leg. & det. JTJ [JTJ]. Nephepeltia phryne (Perty, 1834). Publ. Recs. ECU: (H 06), (P 11), (H 14); LR: von Ellenrieder (2014), NA: von Ellenrieder (2014); OR: * Tennessen (1996 b), von Ellenrieder & Garrison (2011), von Ellenrieder (2014); SD: von Ellenrieder (2014); SU: von Ellenrieder (2014). Also ARG, BRA, CEN, GUF GUY, PER, SUR, TTO, VEN: (H 06), (P 11); plus COL: (P & P), (P 16), von Ellenrieder (2014); and PRY: von Ellenrieder (2014). Addl. Data. ES, San Lorenzo: 3 ♂♂, 11 km SE San Lorenzo, La Chiquita, 20 viii 1976, leg. J. Cohen, det.? [USNM]; LR, Buena Fé: 4 ♂♂, 1 ♀, [Patricia Pilar], just S at Reserva Palenque [about 47 km S of Santo Domingo] (- 0.66, - 79.26, 107 m), 8 x 1988, leg. & det. SWD [FSCA, SWD]; MS, Santiago: 1 ♂, pond along road between Sucua and Mendez (- 2.6833, - 78.2283, 600 m), 6 xi 1997, leg. & det. TWD [SWD]; NA, Carlos Julio Arosemena Tola: 1 ♂, [Ila], Río Pucauyacu, 22 xi 1934, leg. WCM, det. D. Borror [FSCA]; 1 ♀, J. S. B. S., Pond # 19, 12 xi 1991, leg. E. McCallie, det.? [USNM]; 6 ♂♂, Puerto Napo, 5.3 km E of Hwy. 45 on J. S. B. S. Road, Río Sinde (- 1.0500, - 77.7481, 450 m), 13 vi 1996, leg. & det. WFM [FSCA, LSAM, QZAC, USNM]; 1 ♂, same data but leg. & det. KJT [KJT]; 1 ♂, 2 ♀♀, leg. & det. SWD [SWD]; 8 ♂♂, 1 cop. pair, same data but 22 vii 1996, leg. & det. WFM [FSCA, LSAM, QZAC, USNM]; 2 ♂♂, same data but 12 xi 1997 [FSCA, QZAC]; 1 ♂, same data but leg. & det. TWD [SWD]; 1 ♀, Río Chinchipino, 27 + km E of Puerto Napo (- 1.0728, - 77.5864, 385 m), 10 x 2009, leg. & det. JTJ [JTJ]; 1 ♂, same data but 22 vii 1996, leg. & det. SWD [SWD]; 1 ♂, Puerto Napo, 3.2 km E on J. S. B. S. Road, sloughs, 23 vii 1996, leg. & det. SWD [SWD]; 1 ♂, Puerto Napo, 2 nd stream E of Río Sinde on J. S. B. S. Road, 23 vii 1996, leg. & det. SWD [SWD]; OR, Francisco de Orellana: trib. of Río Payamino, road to Loreto, 1.7 km W of jct. with Coca Road (- 0.4367, - 77.0017, 258 m), 13 vi 1995, leg. & det. KJT [KJT]; 1 ♂, same data but leg. & det. WFM [FSCA]; 22 ♂♂, same data but leg. & det. SWD [SWD]; 1 ♂, [La Nueva Esperanza] [across Río Napo from], near Primavera “ Limoncocha ”, Río Napo & Lake Taracoa (- 0.45, - 76.77, 300 m), 27 vii 1977, leg. DRP & S. Hills, det. DRP [DRP]; 8 ♂♂, same data but 26 viii 1980, leg. & det. SWD [UTIC, SWD]; 7 ♂♂, same data but 4 – 8 xi 1980, leg. & det. MJW [FSCA]; 1 ♂, 1 ♀, [Pompeya Sur] S on [Pompeya Sur-Iro road] at km marker 31, borrow pit near Río Sabaleto (- 0.6569, - 76.4528, 216 m), 17 vii 1996, leg. & det. WFM [FSCA, QZAC]; 2 ♂♂, P. N. Y., swamp (- 0.6508, - 76.3558, 200 m), 16 xi 1997, leg. & det. KJT [KJT]; 4 ♂♂, P. N. Y., 2.1 km E of Yasuni Biological Research Station, pond and swamp (- 0.6741, - 76.3685, 244 m), 15 – 16 xi 1997, leg. & det. WFM [FSCA, QZAC, USNM]; 2 ♂♂, 1 ♀, stream 2.3 km E of Yasuni Sci. Res. Sta., 9 vi 1995, leg. & det. KJT [KJT]; 1 ♂, 1 ♀, same data but leg. & det. SWD [SWD]; SU, Lago Agrio: adults, 2 km. N of Lago Agrio, 27 vii 1975, A. Langley, det.? [USNM]; 2 ♂♂, 17 km S of Lago Agrio, swamp, 19 viii 1980, leg. & det. SWD [SWD]; Shushufindi: 2 ♂♂, 1 ♀, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 23 viii 1980, leg. & det. SWD [KJT, SWD]; 7 ♂♂, 2 ♀♀, same data but 14 xi 1980, leg. & det. MJW [FSCA]; 1 ♂, same data but lakeshore, 26 vii 1977, leg. DRP & S. Hills, det. DRP [DRP]; 1 ♂, La Selva, Garzacocha, 12 x 1988, leg. & det. SWD [SWD].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B5AAFF6DFF23F4102677FAB3.taxon	discussion	The main reference for this genus is Borror (1931); see also Geijskes (1948) and Rehn (2003). Oligoclada ranges over most of South America north of Paraguay, with nine species in Ecuador. They are small, dark dragonflies (Fig. 105) that inhabit small streams, backwaters of streams and adjacent swamps, canals, ponds and man-made reser- voirs. Only two of the 23 species have been reared. Oligoclada abbreviata (Rambur, 1842). Publ. Recs. ECU: Daigle (2005), (H 06), (P 11), (H 14); LR: * Campos (1922); NA: Tennessen (1998). Also BRA, GUF, GUY, PER, SUR, VEN: (P 11); plus COL Palacino et al. (2017 — as O. abbreviata limnophila). Addl. Data. LR, Quevedo: 1 ♀, Quevedo, Río Palenque (- 1.03, - 79.43, 100 m), date?, leg. F. Campos, det. WFM [FSCA]; NA, Tena: 1 ♂, Río Chinchipino, 27 km E of Puerto Napo (- 1.0728, - 77.5864, 385 m), 13 xi 1997, leg. & det. KJT [KJT]; 2 ♂♂, same data but leg. & det. TWD [SWD]; 2 ♂♂, same data but leg. & det. JJD [JJD]; OR, Francisco de Orellana: 1 ♂, stream (- 0.4625, - 77.1362, 282 m), 20 x 2013, leg. C. Kerst, det. WFM [FSCA]. Oligoclada crocogaster Borror, 1931. Publ. Recs. ECU: (H 14 — as O. cf. crocogaster); NA: * Tennessen (1996 b); OR: Rhen (2003). Also BRA: (H 06), (P 11). Addl. Data. OR, Francisco de Orellana: 1 ♂, swamp, P. N. Y. (- 0.6508, - 76.3558, 200 m), 16 xi 1997, leg. & det. KJT [KJT]; Loreto: 1 ♂, Loreto 34 km W on road to Narupa, pond (- 0.7428, - 77.5314, 835 m), 14 vi 1995, leg. & det. WFM [FSCA]; 2 ♂♂, same data but leg. & det. KJT [KJT]; SU, Shushufindi: 3 ♂♂, La Selva Lodge, for- est (- 0.4983, - 76.3733, 200 m), 23 xi 1997, leg. & det. TWD [SWD]; 1 ♂, Shushufindi, about 3 km W, small steam (- 0.1662, - 76.6879), 28 x 2013, leg. & det. KJT [FSCA]. Oligoclada monosticha Borror, 1931. Publ. Data. ECU: * Rhen (2003), (H 06), (P 11), (H 14). Also PER, VEN: (H 06), (P 11); plus COL: (P 16). Addl. Data. OR, Francisco de Orellana: 2 ♂♂, [La Nueva Esperanza] S of [across Río Napo from] Primavera Laguna Taracoa (- 0.45, - 76.77, 244 m), 27 vii 1977, leg. DRP & S. Hills, det. DRP [DRP, FSCA]; Loreto: 1 ♂, small streams and forest trails 12 km E of Loreto, 2 + km S of Hwy. 45 (- 0.6217, - 77.2600, 365 m), 12 x 2009, leg. & det. JTJ [JTJ]. Oligoclada pachystigma Karsch, 1890. Publ. Recs. ECU: (H 06), (P 11), (H 14); OR: * Rhen (2003). Also BRA, GUF, GUY, PRY, PER, SUR, VEN: (H 06), (P 11); plus COL: (P 16). Addl. Data. OR, Francisco de Orellana: 1 ♂, [Anangu] [SE across Río Napo from Providencia], Lago Anangu “ on Río Napo ” (- 0.5213, - 76.4361, 300 m), 9 v 1975, leg. T. Emmel, det.? [FSCA]; 10 ♂♂, [La Nueva Esperanza] [across Río Napo from] near Primavera “ Limoncocha ”, Río Napo & Lake Taracoa (- 0.45, - 76.77, 300 m), 27 vii 1977, leg. & det. DRP [DRP]; 16 ♂♂, same data but 4 xi 1980, leg. & det. MJW [FSCA]; 3 ♂♂, same data but 26 viii 1980, leg. & det. SWD [SWD]; 1 ♀, streams in hill forest S of Napo River across from La Selva, 13 x 1988, leg. & det. SWD [SWD]; 1 ♂, [Pompeya Sur] S on [Pompeya Sur-Iro road] at km marker 31, borrow pit near Río Sabaleto (- 0.6569, - 76.4528, 216 m), 12 vi 1995, leg. & det. WFM [FSCA]; 2 ♂♂, 17 vii 1996, leg. & det. SWD [SWD]; 1 ♂, pond, Loreto Road, 11.8 km W of Río Payamino ferry, 18 xi 1997, leg. & det. KJT [KJT]; Loreto: 1 ♂, pond along Loreto-Archidona Road (- 0.7417, - 77.5517, 900 m), 19 xi 1997, leg. & det. TWD [SWD]; 1 ♂, mid-sized stream 12 km E of Loreto, about 4 km S of Hwy. 45 (- 0.6353, - 77.2564, 350 m), 11 x 2009, leg. & det. JTJ [JTJ]; SU, Shushufindi: 1 ♀, Limoncocha, Río Napo (- 0.40, - 76.62, 300 m), 6 ix 1972, leg. T. Emmel, det.? [FSCA]; 1 cop. pair, 5 ♂♂, 11 ♀♀, same data but 23 viii 1980, leg. & det. SWD [UTIC, KJT, SWD]; 43 ♂♂, 4 ♀♀, same data but 2 – 25 xi 1980, leg. & det. MJW [FSCA]; 3 ♂♂, 2 ♀♀, same data but lake edge, 5 vii 1971, leg. D. Pearson, det. DRP [DRP]; 3 ♂♂, 2 ♀♀, same data but 16 – 25 viii 1971 [DRP]; 1 ♂, same data but 11 xii 1971 [DRP]; 1 ♀, same but 6 i 1972 [DRP]; 2 ♂♂, same data but 24 ii 1972 [DRP]; 7 ♂♂, same data but leg. DRP & S. Hills, det. DRP [DRP]; 1 ♀, same data but 3 ix 1978, leg. KWK, det. MJW [FSCA]; 1 ♂, same data but forest trail, 28 xii 1971, leg. D. Pearson, det. DRP [DRP]; 2 ♂♂, same data but 16 ii 1972 [DRP]; 1 ♂, 5 ♀♀, same data but for- est edge and clearing, 23 – 26 vii 1977, leg. & det. DRP [DRP]; 5 ♂♂, 1 ♀, La Selva, 100 km E Coca, Garzacocha, 11 – 12 x 1988, leg. & det. SWD [NVE, SWD]; 2 ♂♂, trails, small streams, 3 km W of Shushufindi, 2 + km N off Hwy. 26 at km marker 17 (- 0.1664, - 76.6917, 290 m), 13 x 2009, leg. & det. JTJ [JTJ]. * Oligoclada risi Geijskes, 1984. Publ. Recs. BRA, GUF, SUR, VEN: (H 06), (P 11). Addl. Data. NA, Tena: 2 ♂♂, Río Chinchipino, 27 km E of Puerto Napo (- 1.0728, - 77.5864, 385 m), 13 xi 1997, leg. & det. KJT [KJT]. Oligoclada stenoptera Borror, 1931. Publ. Recs. ECU: Rhen (2003), (H 06), (P 11), (H 14); NA: * Tennessen (1998). Also PER, BRA: (H 06), (P 11). Addl. Data. NA, Tena: 3 ♂♂, Río Sinde, pond, 5 km E of Puerto Napo (- 1.0517, - 77.7544, 450 m), 12 xi 1997, leg. & det. KJT [KJT]; OR, Francisco de Orellana: 1 ♂, [La Nueva Esperanza] [across Río Napo from] near Pri- mavera “ Limoncocha ”, Río Napo & Lake Taracoa (- 0.45, - 76.77, 300 m), 27 vii 1977, leg. & det. DRP [DRP]; 1 ♂, same data but 26 viii 1980, leg. & det. SWD [SWD]; 1 ♂, swamp pond, Yasuni Road, km marker 8, 12 vi 1995, leg. & det. KJT [KJT]; 1 ♂, same data but leg. & det. SWD [SWD]; 3 ♂♂, P. N. Y., swamp (- 0.6508, - 76.3558, 200 m), 15 – 16 xi 97, leg. & det. KJT [KJT]; 1 ♂, P. N. Y., swamp (- 0.6508, - 76.3558, 200 m), 16 xi 1997, leg. & det. TWD [SWD]; 1 ♂, [Pompeya Sur], S on [Pompeya Sur-Iro road] at km marker 31, borrow pit near Río Sabaleto (- 0.6569, - 76.4528, 216 m), 17 vii 1996, leg. & det. WFM [FSCA]; 1 ♂, same data but leg. & det. SWD [SWD]; 1 ♂, P. N. Y., 2.1 km E of Yasuni Biological Research Sta., pond and swamp (- 0.6741, - 76.3685, 244 m), 15 xi 1997, leg. & det. WFM [FSCA]; 1 ♂, same data but leg. & det. JJD [JJD]; 1 ♂, same data but 9 vi 1995, leg. & det. SWD [SWD]; SU, Lago Agrio: 1 ♂, stream, 44 km W of Lago Agrio, 20 viii 1980, leg. & det. SWD [SWD]. Oligoclada teretidentis Rehn, 2003. Publ. Recs. ECU: Paulson (2004 a), (H 06), (P 11), (H 14); OR: * Rhen (2003 — Holotype & Allotype: [Francisco de Orellana], forested shore of oxbow lagoon near Río Tiputini, approx. 1 km NW of the T. B. S., 11 x 2001, leg. A. Rehn [UMMZ]). Addl. Data. NA, [Tena:] 1 ♂, gravel road S from road from Tena to J. S. B. S. and Airport, tilapia pond (- 1.1280, - 77.7040), date not provided, data provided by WAH & DLW; OR, Francisco de Orellana: 1 ♂, T. B. S., Río Tipu- tini, 55 km SE of Pompeya, 94 km ESE of Coca (- 0.6369, - 76.1503), date not provided, data provided by WAH & DLW; 1 ♂, Shiripuno Lodge, 27 km SE Río Shiripuno bridge, (- 1.1047, - 76.7317, 235 m), date not provided, data provided by WAH & DLW. Note: Endemic to Ecuador. Oligoclada umbricola Borror, 1931. Publ. Recs. ECU: Paulson (1983), Tsuda (1991), Rhen (2003), (H 06), (P 11), (H 14); LR: * Kormondy (1958, plesiallotype); NA: Tennessen (1998); SD: Daigle et al. (1997). Also CEN, COL, VEN: (H 06), (P 11). Addl. Data. LR, Buena Fé: 4 ♂♂, Santo Domingo, 47 km S, forest slough at Río Palenque Field Station (200 m), 18 vii 1977, leg. & det. DRP [DRP]; 2 ♂♂, same data but 8 x 1988, leg. & det. SWD [SWD]; Montalvo: 1 ♂, stream 1.8 km N of Hwy. 491, about 25 km E of Babahoyo (- 1.8111, - 79.3189, 60 m), 7 iii 2010, leg. & det. KJT [KJT]; Quevedo: 3 ♀♀, Pichilingue, Río Palenque (- 1.10, - 79.48, 64 m), 2 ii 1955, leg. E. Schlinger & E. Ross, det. JJD [FSCA]; 2 ♂♂, stream 16 km E of Hwy. E 25 [from Formisa] (- 0.7256, - 79.3483, 200 m), 24 ii 2010, leg. & det. JTJ [JTJ]; MN, El Carmen: 1 ♂, 1 ♀, borrow pit / small slow stream, NW of El Carmen (- 0.1350, - 79.5797, 185 m), 28 ii 2010, leg. & det. JTJ [JTJ]; 2 ♂♂, same data but 2 iii 2010, leg. & det. JJD [JJD]; 1 ♂, same data but leg. & det. KTJ [FSCA]; SD: 1 ♂, Santo Domingo de Los Colorados, 12 km E on Hwy. 30 near Hotel Tinalandia (- 0.30, - 79.67, 610 m), 23 x 1980, leg. & det. MJW [FSCA]; 1 ♂, rocky pool 8 km W of Alluriquin, Hwy. 30, 31 i 1997, leg. & det. JJD [JJD]. Oligoclada walkeri Geijskes, 1931. Publ. Recs. ECU: * Rhen (2003), (H 06), (P 11), (H 14). Also BRA, GUF, GUY, PER, SUR, TTO, VEN: (H 06), (P 11); plus COL Palacino et al. (2017). Addl. Data. NA, Tena: 1 ♂, Río Chinchipino, 27 km E of Puerto Napo (- 1.0728, - 77.5864, 385 m), 23 vii 1996, leg. & det. KJT [KJT]; 2 ♂♂, same data but 22 vii 1996, leg. & det. SWD [SWD]; SU, Shushufindi: 1 ♂, Limon- cocha, forest stream, 1 iii 1972, leg. D. Pearson, det. DRP [DRP].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B5A8FF16FF23F37027B4FD73.taxon	discussion	A species-rich New World genus with 28 species of which 12 are recorded for Ecuador with mention of one more possibly occurring here. The most recent taxonomic treatments were by von Ellenrieder (2009 b, 2013 a). This is a diverse group color-wise (Figs. 106 a, b, c) that inhabits open and forested pools depending on species, often with muddy water and / or substrate, sometimes in slow streams. Orthemis aequilibris Calvert, 1909. Publ. Recs. OR: * von Ellenrieder (2012). Also BOL, BRA, COL, GUF, GUY, PRY, PER, SUR, VEN: (H 06); plus ARG, CEN: (P 16). Note: The von Ellenrieder (2012) record is a casual mention in a general discussion about the sympatry of Orthemis species noted at the T. B. S. * Orthemis aciculata von Ellenrieder, 2012. Publ. Recs. BRA, CEN, PER, SUR, TTO: von Ellenrieder (2012), (P 16). Addl. Data. SU: Cuyabeno Reserve, 16.7 km E of Cuyabeno village, 3 km SE of Cuyabeno Lodge, no date provided, data provided by WAH & DLW; Nicky Lodge, 8.6 km NW of Playas de Cuyabeno village, 120 km SE of Lago Agrio (215 m), no date provided, data provided by WAH & DLW. Orthemis ambirufa Calvert, 1909. Publ. Recs. ECU: (H 14), (P 16); * EO: Piedras (- 3.63, - 79.92), 6 vii 1941, leg. D. B. Laddey [USNM], von Ellenrieder (2012). Also ARG, BRA, GUF, SUR, VEN: (H 06), von Ellenrieder (2012); plus BOL, COL: von Ellenrieder (2012). Orthemis biolleyi Calvert, 1906. Publ. Recs. ECU: Campos (1925), Navás (1923), Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14); GY: * Ris (1910); NA: Daigle (1996); OR: Tennessen (1996 b), von Ellenrieder & Garrison (2011). Also BOL, BRA, CEN, COL, GUF, GUY, PER, SUR, TTO, VEN: (H 06), (P 11). Addl. Data. EO, Piñas: 10 km SE towards Portovelo (- 3.72, - 79.67, 610 m), 3 vii 1989, leg. R. Miller, det. GHB [FSCA]; GY, Guayaquil: 1 ♀, Guayaquil (- 2.7333, - 79.7500, 17 m), date?, leg.?, det. Ris, data provided by RWG; MN, El Carmen: 1 ♀, El Carmen, leg. & det.? [QZAC]; MS, Huamboya: 2 ♂♂, Macas, 58 km N on Hwy. 45 (70 km N of Sucua), roadside ditch and tribs. near Río Pasanac (- 1.9447, - 77.8536, 762 m), 10 xi 1997, leg. & det. WFM [FSCA, QZAC]; 7 ♂♂, same data but 16 ix 2005, leg. & det. WFM [FSCA, JJD, QZAC, USNM]; 2 ♂♂, Río Pasanac forest pools, 57 km N of Macas, Hwy. 45, 10 xi 1997, leg. & det. JJD [JJD]; NA, Archidona: photo, 1 ♂, Archidona, a few miles W at entrance to Playa Bosque Reserve, deep, semi-shaded, fishless hand-dug pond, 11 vii 2011, photo by Jerry Johnson, det. SWD; El Chaco: 1 ♂, stream, 52 km NE of Chaco, 18 viii 1980, leg. & det. SWD [SWD]; Tena: 2 ♂♂, J. S. B. S., near Tena (- 1.0500, - 77.7833, 450 m), 4 – 29 xi 1994, leg. A. Jaramillo, det. DRP [DRP]; 2 ♀♀, same data but 16 x 1988, leg. & det. SWD [SWD]; 1 ♀, pond, Puyo Road, 11 km S of Río Napo (500 m), 21 vii 1996, leg. & det. KJT [KJT]; 1 ♂, pond along Puyo-Tena Road (- 1.1150, - 77.8083, 600 m), 12 xi 1997, leg. & det. TWD [SWD]; OR, Francisco de Orellana: 3 ♂♂, pond, road to Waorani Reserve, 5.6 km S of turn to Yasuni Sci. Res. Sta., 10 vi 1995, leg. & det. KJT [JJD, KJT]; 2 ♀♀, P. N. Y., 8 km N of Waorani Road on Yasuni Road, palm swamp just S of Río Bogi 2 Este [km marker 42.3], (- 0.6734, - 76.4397), 12 vi 1995, leg. & det. WFM [FSCA, JJD]; 1 ♂, swamp, P. N. Y. (- 0.6508, - 76.3558, 200 m), 15 xi 1997, leg. & det. KJT [KJT]; 1 ♂, same data but 16 xi 1997, leg. & det. TWD [SWD]; 1 ♂, P. N. Y., 2.3 km E of Yasuni Biological Research Sta., small stream (- 0.67, - 76.37, 200 m), 9 vi 1995, leg. & det. WFM [FSCA]; 1 ♂, 1 ♀, same data but leg. & det. SWD [SWD]; 5 ♂♂, 1 ♀, P. N. Y., 2.1 km E of Yasuni Biological Research Sta., pond and swamp (- 0.6741, - 76.3685, 244 m), 15 – 16 xi 1997, leg. & det. WFM [FSCA, LSAM, QZAC, USNM]; 1 ♂, P. N. Y., Yasuni Biological Research Sta. along trails and roads within 5 km of Res. Sta. (- 0.6719, - 76.3992, 244 m), 18 vii 1996, leg. & det. WFM [QZAC]; 1 ♂, P. N. Y., 5.6 km W of Yasuni Road on PC 29 - 30, small stream / swamp (- 0.38, - 76 30), 10 vi 1995, leg. & det. SWD [SWD]; 1 ♂, T. B. S., USFQ, Mauritia palm swamp off Guacamayo trail at marker 2100 (- 0.6331, - 76.1628, 220 m), 20 i 2009, leg. & det. RWG & NVE [NVE]; 1 ♀, T. B. S., USFQ, open grassy area behind dining room, surrounded by bushes and trees and some seepage areas (- 0.6383, - 76.1492, 220 m), 11 i 1009, leg. & det. RWG & NVE [NVE]; La Joya de los Sachas: 1 ♂, Pompeya, 0.5 km W of Puerto Pompeya Ferry Landing roadside ditches (- 0.4444, - 76.6263, 238 m), 19 vii 1996, leg. & det. WFM [LSAM]; Loreto: 1 ♂, trail 12 km NE of Loreto (- 0.6217, - 77.2600, 365 m), 15 x 2009, leg. & det. KJT [KJT]; PC, San Miguel Bancos: 1 ♂, upper Mindo Valley, ix 2004, photo by M. Read; SD: 1 ♂, Alloriquin, i 1983, leg. & det.? [QZAC]; SU, Shushufindi: 1 ♂, Limoncocha, Río Napo (- 0.4000, - 76.6167, 296 m), 10 xi 1980, leg. & det. MJW [FSCA]; 1 ♀, same data but lake shore 12 viii 1972, leg. E. Stiles, det. DRP [DRP]; 1 ♀, Shushufindi, 2 km S of Yamaramsuku School on Limoncocha Road, stream and marsh (- 0.2147, - 76.6494, 255 m), 19 vii 1996, leg. & det. WFM [QZAC]; 1 ♂, near Río Blanco, S of Shushufindi, 19 vii 1996, leg. & det. KJT [KJT]; 1 ♂, trails, small streams 3 km W of Shushufindi, 2 + km N off Hwy. 26 at km marker 17 (- 0.1664, - 76.6917, 290 m), 13 x 2009, leg. & det. JTJ [JTJ]; 1 ♀, La Selva, Manicocha, 12 x 1988, leg. & det. SWD [SWD]; 1 ♂, NW of Shushufindi, shallow mud bottom pond, (- 0.1647 - 76.6880, 273 m), 28 x 2013, leg. & det. CDK [FSCA]. Orthemis cinnamomea von Ellenrieder, 2009. Publ. Recs. ECU: (P 11), (H 14). OR: von Ellenrieder (2012) SU: von Ellenrieder (2009 b, 2012). Also PER: von Ellenrieder (2009, 2012), (P 16). Addl. Data. OR: Llanchama village, Río Tiputini, 16 km W of Tiputini village (205 m), no date provided, data provided by WAH & DLW; SU, Shushufindi: 1 paratype ♂, Limoncocha, lake edge (- 0.4000, - 76.6167, 269 m), 13 i 1972, leg. D. Pearson, det. NVE [DRP]; 1 paratype ♀, same data but 16 i 1972 [DRP]; 1 paratype ♂, same data but 20 ii 1972 [RWG]; Nicky Lodge, 8.6 km NW of Playas de Cuyabeno village, 120 km SE of Lago Agrio (215 m), no date provided, data provided by WAH & DLW. Orthemis coracina von Ellenrieder, 2009. Publ. Recs. ECU: (P 11), (H 14); SU: * von Ellenrieder (2009 — Holotype ♂: Limoncocha, forest edge (- 0.24, - 76.36, 300 m), 23 vii 1977, leg. DRP, [USNM]. Also BRA, COL, PER, SUR: von Ellenrieder (2012), (P 16). Orthemis cultriformis (Hagen, 1861). Publ. Recs. ECU: Navás (1923), Campos (1925), Santos (1967 c), Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14); MS: von Ellenrieder (2012); NA: Daigle (1996), von Ellenrieder (2012); OR: Tennessen (1996 b), von Ellenrieder & Garrison (2011), von Ellenrieder (2012); PA: * Ris (1910). SU: Donnelly (1998). Also ARG, BOL, BRA, CEN, COL, GUF, GUY, PRY, PER, SUR, TTO, VEN: (H 06), (P 11). Addl. Data. ES, Esmeraldas: [IRSNB]; MS, Gualaquiza: [ANSP, FSCA]; Huamboya: [KJT]; NA, Tena: [DRP, FSCA, JJD, JTJ, LSAM, KJT, QZAC, RWG, SWD, SWD]; OR, Francisco de Orellana: [FSCA, KJT, LSAM, NVE, QZAC, SWD, TWD, USFQ, USNM]; Loreto: [JTJ]; PA, Mera: [KJT]; SU, Lago Agrio: [FSCA, SWD]; Shushufindi: [DRP, FSCA, KJT]; ZC, El Pangui: [FSCA, KJT, QZAC]; Yantzaza: [FSCA]. Orthemis discolor (Selys, 1857). Publ. Recs. ECU: Selys (1875 — as Libellula discolor), Racenis (1954), (P 11), (H 14); CR: Campos (1922); CB: Calvert (1906), Campos (1922); EO: Campos (1927); ES: Campos (1922); GY: * Hagen (1861), Ris (1910), Campos (1922); IM: Daigle et al. (1997); LR: Campos (1922); NA: Daigle (1996); OR: von Ellenrieder & Garrison (2011); PC: Ris (1910), Campos (1922). Also ARG, BOL, BRA, CEN, CHL, COL, GUF, GUY, PRY, PER, SUR, TTO, URY, VEN: (H 06), (P 11). Addl. Data. CN, La Troncal: [KJT]; CB, Cumandá: [FSCA]; Pallatanga: [ANSP]; EO, Piñas: [ACRP]; ES, Quininde: [QZAC]; San Lorenzo: [ANSP, KJT, LSAM, QZAC, USNM]; GY, Naranjal: [FSCA]; Guayaquil: [FSCA]; IM, Cotacachi: [KJT]; Ibarra: [FSCA, JJD, KJT, QZAC]; LR, Buena Fé: [FSCA]; Quevedo: [JJD]; MN, El Carmen: [JJD, QZAC]; Jipijapa: [FSCA]; Portoviejo: [JJD]; MS, Gualaquiza: [ANSP, FSCA]; Huamboya: [FSCA]; Morona: [FSCA]; Santiago: [FSCA, KJT, LSAM, QZAC, TWD, USNM]; NA, Archidona: [FSCA, QZAC]; El Chaco: [FSCA]; Tena: [ANSP, DRP, FSCA, QZAC]; OR, Francisco de Orellana: [FSCA, KJT, NVE, TWD, QZAC, USFQ, USNM]; Loreto: [QZAC]; PA, Santa Clara: [FSCA]; PC, Mejías: [QZAC]); San Miguel Bancos: [DRP, QZAC]; SD: [DRP, FSCA, JJD. QZAC]; SU, Shushufindi: [DRP, FSCA, KJT]; ZC, El Pangui: [FSCA]; Zamora: [FSCA, JJD, JTJ, KJT, LSAM]. Note: All records prior to 1996 are listed as “ Orthemis ferruginea ” except Hagen (1861); specimens from Ecuador labeled as O. ferruginea should be O. discolor. Orthemis paulsoni von Ellenrieder, 2012. Publ. Recs. ECU: (H 14); OR: * von Ellenrieder (2012); SU: von Ellenrieder (2012 — holoytpe ♂: forest trail about 3 km W of Shushufindi (- 0.1664, - 76.6917, 290 m), 14 x 2009, leg. KJT [FSCA]). Also PER: von Ellenrieder (2012); plus BOL (P 16). Addl. Data. SU, Shushufindi: 1 ♂, trail 3 km NW of Shushufindi (- 0.1664, - 76.6917, 290 m), 13 x 2009, leg. & det. KJT [KJT]; 3 ♂♂, 1 ♀, same data but 14 x 2009 [KJT]; 1 ♂, trails, small streams 3 km W of Shushufindi, 2 + km N off Hwy. 26 at km marker 17 (-. 0.1664, - 76.6917, 290 m), 13 x 2009, leg. & det. JTJ [JTJ]. Orthemis regalis Ris, 1910. Publ. Recs. ECU: (H 06), (P 11), * von Ellenrieder (2012), (H 14). Also BRA, PER, SUR, VEN: (H 06), (P 11), von Ellenrieder (2012); plus (GUF): (P 16). Addl. Data. SU, Shushufindi: 1 ♀, Limoncocha, forest trail (- 0.4000, - 76.6167, 269 m), 14 xii 1971, leg. D. Pearson, det. DRP, checked by RWG [DRP]. Orthemis schmidti Buchholz, 1950. Publ. Recs. ECU: (H 06), (P 11), (H 14); NA: Daigle (1996), von Ellenrieder (2009); (P 11), OR: * Tennessen (1996 b), von Ellenrieder & Garrison (2011), von Ellenrieder (2012). Also BOL, BRA, GUF, PER, SUR, VEN, TTO: (H 06), (P 11); Plus CEN: (P 11). Addl. Data. CT, Sigchos: 1 ♂, San Francisco de las Pampas Road between Naranjito-San Francisco de Las Pampas (- 0.42, - 79.0, 1500 m), 1 v 1999, leg. L. Torres, det.? [QZAC]; 2 ♀♀, same data but 14 v 1999, leg. E. Tapia, det.? [FSCA]; MS, Morona: 1 ♂, Macas, 15 km W on Guamote Road, small stream and roadside seepage pools. 15 ix 2005, leg. WFM, det. JJD [JJD]; NA, Tena: 3 ♂♂, Puerto Napo 5.3 km E of Hwy. 45 on J. S. B. S. Road, Río Sinde (- 1.0500, - 77.7481, 450 m), 13 vii 1996, leg. & det. WFM [FSCA, LSAM, QZAC]; 1 ♂, same data but leg. & det. JJD [DRP]; 1 ♂, same data but leg. & det. KJT [FSCA]; 3 ♂♂, Puerto Napo, 3.2 km E on J. S. B. S. Road, ponds and flowing marsh, 23 vii 1996, leg. & det. WFM [FSCA, JJD, USNM]; 1 ♂, same data but leg. & det. SWD [NVE]; OR, Francisco de Orellana: 1 ♂, trib. of Río Payamino, road to Loreto, 1.7 km W of jct. with Coca Road (- 0.4367, - 77.0017, 258 m), 13 vi 1995, leg. & det. KJT [KJT]; 1 ♂, same data but leg. & det. WFM [JJD]; 2 ♂♂, same data but 18 xi 1997, leg. & det. TWD [SWD]; 1 ♂, same data but leg. & det. JJD [JJD]; 2 ♂♂, swamp pond at km 8, S of Río Napo on Yasuni Road, 11 vi 1995, leg. & det. SWD [JJD]; 4 ♂♂, grassy swamp, Yasuni Road, km marker 31, near Río Savaleto 2, Yasuni Preserve, 12 vi 1995, leg. & det. KJT [JJD, KJT]; 1 ♀, stream, PC 29 - 30 Road, 1.8 km W of Yasuni Road, 10 vi 1995, leg. & det. KJT [KJT]; 2 ♂♂, 1 ♀, P. N. Y., Yasuni Biological Research Sta., along trails and roads within 5 km of Res. Sta., (- 0.6719, - 76.3992, 244 m), 18 vii 1996, leg. & det. WFM [FSCA, LSAM, USNM]; 1 ♀, T. B. S., USFQ, open grassy area behind dining room, surrounded by bushes and trees and some seep- age area (- 0.6383, - 76.1492, 220 m), 10 i 2009, leg. & det. RWG & NVE [NVE]; ZC: Zamora: Road from Zamora to Podocarpus National Park, (948 m), leg. & det. C. Willigalla. Orthemis sulphurata Hagen, 1868. Publ. Recs. ECU: Calvert (1909), Tsuda (1986), (H 06), (P 11), (H 14); CR: Campos (1922); CB: Campos (1922); GY: * Hagen (1868 — ♀ type, Guayaquil), Calvert (1899), Ris (1910), Cam- pos (1922), Donnelly (2001), Tennessen & Johnson (2008). Also PER: Hoffman (2009); TTO, VEN?, (H 06); plus (CEN). Addl. Data. BO, Caluma: 1 ♂, small forested stream 9 km E of Caluma, 5 iii 2010, leg. & det. JTJ [JJD]; EO, Arenillas: 1 ♂, farm pond, 11 km S of Arenillas, road to Alamar (95 m), 10 iv 2008, leg. KJT, det. JJD [JJD]; Piñas: 1 ♂, 10 km SE of Piñas, 3 vii 1989, leg. R. Miller & L. Stange, det. JJD [JJD]; Santa Rosa: 2 ♂♂, 1 ♀, stream and associated pond, wetland about 12 km E of Arenillas, Hwy. 25 (- 3.5244, - 79.9831, 30 m), 29 iii 2008, leg. & det. JTJ [JTJ]; 1 ♂, 1 ♀, same data but leg. & det. KJT [KJT]; 3 ♂♂, 1 ♀, same data but 9 iv 2008 [JJD, KJT]; ES, Esmeraldas: Esmeraldas, SW at muddy stream, 19 vii 1977, leg. & det. DRP [DRP]; GY, Balao, 2 ♂♂, small stream / ditches in banana plantation, about 2 km E of Hwy. 25 near Balao Jct. (- 2.9100, - 79.6919, 70 m), 28 iii 2008, leg. & det. JTJ [JJD, JTJ]; 1 ♂, same data but leg. KJT [JJD]; Guayaquil: 1 ♀, Guayaquil (- 2.20, - 79.90, 8 m), date?, leg. F. Cam- pos, det. P. Calvert [FSCA]; Naranjal: 3 ♂♂, 1 ♀, Bucay (- 2.2, - 79.18, 233 m), viii, leg. F. Campos, det. P. Calvert [FSCA, USNM]; 1 ♂, Horned Screamer Reserve, 5 miles E of Cerro Mas Vale (10 m), 10 ii 2001, leg. F. Sibley, det. RWG [RWG]; 1 ♀, flooded fields just W of Fundacion Andrate (- 2.3833, - 79.6333, 10 m), 14 ii 2001, leg. F. Sibley, det. TWD [RWG]; 1 ♀, Cerro Mas Vale, small pond at base of north flank (- 2.3833 - 79.6333, 10 m), 14 ii 2001, leg. F. Sibley, det. TWD [RWG]; 1 ♂, Cerritos just W of Fundacion Andrate, flooded field (- 2.3918, - 79.6283, 9 m), 14 ii 2001, leg. Fred C Sibley, det. TWD [FSCA]; 1 ♂, 1 ♀, Fundación Andrate, Mas Vale area thorn scrub (15 m), 13 xi 2001, leg. F. Sibley, det. TWD [DRP]; 3 ♂♂, 1 tan. pair, Cerro Mas Vale Fundacion Andrate (- 2.40, - 79.63, 15 m), 14 xi 2001, leg. F. Sibley, det. TWD [FSCA]; 1 ♂, Finca de la Gringa, 15 km E of Tres Cerritos (- 2.4456, - 79.5556, 25 m), 12 iv 2008, leg. & det. KJT [KJT]; LJ, Macará: 2 ♂♂, 4 ♀♀, Macará (- 4.38, - 79.95, 400 m), date?, F. Cam- pos R., det. MJW [ANSP, FSCA, JJD]; LR, Buena Fé: 1 ♀, [Consumulo], S Río Palenque (- 0.58, - 79.35, 175 m), 12 vi 1974, leg. J. Longino, det.? [FSCA]; 1 ♂, 2 ♀♀, same data but 12 – 18 viii 1975 [FSCA]; Quevedo: 2 ♂♂, 2 ♀♀, small stream 5 km E of E 25 / Formisa cutoff, 23 ii 2010, leg. & det. JJD [JJD]; Urdaneta: 8 ♂♂, ditches / pools 2.8 km E of Hwy. E 25 (- 1.5217, - 79.5000, 35 m), 21 ii 2010, leg. & det. JJD [JJD]; Ventanas: 1 ♂, 1 ♀, pond / stream 7.2 km E of Hwy. E 25 or 9 km SW of Quinzaloma, 21 ii 2010, leg. & det. JJD [JJD]; MN, Pedernales: 2 ♂♂, Río Nalpe, 3 km E of El Achote, road to Pedernales, 28 ii 2010, leg. & det. JJD [JJD]; 5 ♂♂, small stream 2 km E of Pedernales, 28 ii 2010, leg. & det. JJD [JJD]; PC, San Miguel Bancos: 2 ♂♂, Pedro Vicente Maldonado, road to Río Silanche Preserve, forested & open streams, (about 500 m), 1 vii 2008, leg. & det. DRP [DRP]; SD: 1 ♂, Santo Domingo de los Colorados, 6 iii 1973, leg. M. Deyrup, det. DRP [DRP]; 1 ♂, Tinalandia, 12 km E of Santo Domin- go de los Colorados, 11 v 1986, leg. G. Edwards, det. JJD [JJD]; 1 ♂, pineapple vendor puddle, Parroquia luz de America town, 24 ii 2010, leg. & det. JJD [JJD].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B5D3FF17FF23F5302123FD17.taxon	discussion	A genus of two very wide-ranging species, both migratory (Fig. 107). Key to species can be found in numerous references, including Needham et al. (2014). Habitats range from small ponds, including temporary pools, to ditches and slow parts of small streams, usually with little or no aquatic vegetation. Adults oviposit in pools formed after heavy rain events. Development from egg to adult is rapid, often less than 55 days (Hawking & Ingram 1994). Pantala flavescens (Fabricius, 1798). Publ. Recs. ECU: Paulson (1983), Tsuda (1991), (H 06), (P 11), (H 14); CR: Campos (1922); CB: Campos (1922); EO: Campos (1927); ES: Campos (1922); GA: * Currie (1901), Calvert (1907), Ris (1913 a), Campos (1922), Calvert (1947 b), Gloger (1964), Turner (1967), Peck (1992), (P 11); GY: Calvert (1906), Ris (1913 a), Campos (1922); LR: Calvert (1906), Campos (1922). Also ARG, BOL, BRA, CEN, CHL, COL, GUF, GUY, PRY, PER, SUR, URY, VEN: (H 06), (P 11); plus TTO: (P 11). Addl. Data. AZ, Ona: [FSCA, KJT]; CB, Alausí: [ANSP, FSCA]; EO, Piñas: [ACRP]; Portovelo: [USNM]; ES, Esmeraldas: [DRP]; Muisne: [QZAC]; San Lorenzo: [FSCA]; GA, Isabela: [USNM]; GY, Guayaquil: [ANSP, FSCA]; Naranjal: [FSCA]; IM, Ibarra: [QZAC]; LJ, Loja: [UTIC]; LR, Buena Fé: [DRP, FSCA]; NA, Tena: [FSCA, QZAC]; OR, Francisco de Orellana: [FSCA, KJT, QZAC, USNM]; PA, Santa Clara: [KJT]; Mera: [FSCA]; PC: Quito: [KJT] SD: [FSCA, SWD]; SU, Lago Agrío: [SWD]; Shushufindi: [DRP]; ZC, Zamora: [USNM]. Pantala hymenaea (Say, 1840). Publ. Recs. ECU: Tsuda (1986, 1991), (H 06), (P 11), (H 14); CR: Campos (1922); EO: Campos (1927); GA: * McLachlan (1877), Currie (1901), Ris (1913 a), Campos (1922), Gloger (1964), Turner (1967), Peck (1992), (P 11); GY: Ris (1913 a), Campos (1922). Also ARG, BRA, CEN, CHL, COL, PER, VEN: (H 06), (P 11); Plus TTO: (P 16). Addl. Data. EO, Santa Rosa: 1 ♀, ponds, S side of Hwy. 25 near Santa Rosa (- 3.4306, - 79.9592, 15 m), 29 iii 2008, leg. & det. JTJ [JTJ]; GA, San Cristóbal: 3 ♂♂, 8 ♀♀, Isla Espanola, Bahia Manzanillo, general collecting and UV light, 23 iv 1992 to 2 v 1992, leg. S. Peck & J. Cook, det.? [USNM]; GY, Guayaquil: 7 ♂♂, 5 ♀♀, Puerto Grande (- 2.93, - 80.08, 2 m), ii 1936, leg. Campos, det. P. Calvert [ANSP, FSCA].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B5D2FF12FF23F4EC24D9F844.taxon	discussion	For species identification see Ris (1930), Needham et al. (2014) and von Ellenrieder & Muzón (1999). Perithemis are small, amber-winged dragonflies (Fig. 108) that occupy open and shaded ponds, ditches and small, slow streams. The group is well-represented in Ecuador where nine of the 13 described species are recorded. Perithemis bella Kirby, 1889. Publ. Recs. ECU: Paulson (1983), (H 06), (P 11), (H 14). Also BRA, PER: (H 06), (P 11); Plus COL: (P 16). Addl. Data. SU, Shushufindi: 41 ♂♂, 9 ♀♀, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 1 – 27 xi 1980, leg. & det. MJW [FSCA]; 1 ♀, same data but lake edge, 5 vii 1971, leg. D. Pearson, det. DRP [DRP]; 6 ♂♂, 2 ♀♀, same data but 26 vii 1977, leg. & det. DRP [DRP]; 2 ♂♂, 2 ♀♀, same data but 23 – 24 viii 1980, leg. & det. SWD [SWD]; 3 ♂♂, same data but forest edge, 25 – 26 vii 1977, leg. & det. DRP [DRP]. Perithemis cornelia Ris, 1910. Publ. Recs. ECU: (H 06), (P 11), (H 14); OR: Tennessen (1998), von Ellenrieder & Garrison (2011); SU: Donnelly (1998). Also BOL, BRA, GUF, PER, VEN: (H 06), (P 11); Plus SUR: (P 16). Addl. Data. NA, Tena: 1 ♂, Puerto Napo, 23 km E at J. S. B. S., tropical wet forest (- 1.07, - 77.60, 450 m), 16 x 1988, leg. & det. SWD [SWD]; OR, Francisco de Orellana: 1 ♂, P. N. Y., 6.8 km E of Yasuni Biological Research Sta., small forested stream (- 0.6545, - 76.3417, 238 m), 15 xi 1997, leg. & det. WFM [FSCA]; 1 ♂, same data but leg. & det. KJT [KJT]; 1 ♂, P. N. Y., stream (- 0.7100, - 76.4750, 200 m), 16 xi 1997, leg. & det. TWD [SWD]; 2 ♂♂, P. N. Y., stream (- 0.6542, - 76.3542, 200 m), 15 xi 1997, leg. & det. TWD [SWD]; 1 ♂, P. N. Y., km 47 on [Pompeya Sur-Iro road] (- 0.65, - 76.45, 244 m), 1 ix 2000, leg. T. Emmel, det. JJD [FSCA]; 1 ♂, T. B. S., USFQ, stream crossing Matapalo trail at marker 450 (- 0.6369, - 76.1458, 245 m), 15 i 2009, leg. & det. RWG & NVE [RWG]; 5 ♂♂, T. B. S., USFQ, black water stream crossing Guacamayo trail at marker 2050, (- 0.6383, - 76.1492, 220 m), 17 i 2009, leg. & det. RWG & NVE [NVE, RWG, USFQ]; SU, Shushufindi: 1 ♂, NW of Shushufindi, stream near pond, (- 0.1647, - 76.6880, 273 m), 28 x 2013, leg. & det. CDK [FSCA]. Perithemis domitia (Drury, 1773). Publ. Recs. ECU: Muttkowski (1910 — as P. domitia tenera), (H 06), (P 11), (H 14); CR: Campos (1922 — as P. domitia var pocahontas); GY: Campos (1922); EO: Campos (1927); LR: * Calvert (1906 — as var. t enera), Campos (1922); SD: Daigle et al. (1997). Also BRA, CEN, COL, VEN: (P 11); plus PER: (H 06); and TTO: (P 11). Addl. Data. LR, Buena Fé: 1 ♀, Santo Domingo 47 km S, forest slough at Río Palenque Field Station (200 m), 18 v 1977, leg. & det. DRP [DRP]; 1 ♂, same data but 8 x 1988, leg. & det. SWD [JJD]; 4 ♂♂, same data but 9 x 1988 [FSCA]; Montalvo: 1 ♂, small stream 1.8 km N of Hwy. 491, about 25 km E of Babahoyo (- 1.8111, - 79.3189, 60 m), 7 iii 2010, leg. & det. JTJ [JTJ]; MN, Flavio Alfaro: 1 ♂, small stream 0.5 km N of Chindul, forested (- 0.2389, - 79.8742, 30 m), 1 iii 2010, leg. & det. JTJ [JTJ]; Junin: 1 ♂, stream just S of Junin (- 0.9317, - 80.2256, 70 m), 3 iii 2010, leg. & det. JTJ [JTJ]; Pedernales: 1 ♂, Río Nalpe, about 3 km E of El Achote (0.0547, - 79.9739, 130 m), 28 ii 2010, leg. & det. JTJ [JTJ]; 1 ♂, small stream about 2 km E of Pedernales (0.0836, - 80.0139, 90 m), 28 ii 2010, leg. & det. JTJ [JTJ]; SD: 3 ♂♂, rocky pool 8 km W of Alluriquin, Hwy. 30, 31 i 1997, leg. & det. JJD [JJD]. Note: Data available indicates a west slope species. Perithemis electra Ris, 1910. Publ. Recs. ECU: (H 06), (P 11), (H 14); OR: * Tennessen (1996 b), von Ellenrieder & Garrison (2011); SU: Donnelly (1998). Also BOL, BRA, CEN, COL, GUF, PRY, PER, TTO, VEN: (H 06), (P 11); Plus SUR: (P 16). Addl. Data. GY, Naranjal: 1 ♂, Bucay (- 2.2, - 79.18, 233 m), 1911, leg. F. Campos, det. WFM [FSCA]; LR, Quevedo: 1 ♂, stream 16 km E of Hwy. E 25 [from Formisa], (- 0.7256, - 79.3483, 200 m), 23 ii 2010, leg. & det. KJT [KJT]; MN, Pedernales: 1 ♂, small stream about 2 km E of Pedernales (0.0836, - 80.0139, 90 m), 28 ii 2010, leg. & det. KJT [KJT]; Flavio Alfaro: 1 ♂, small forest stream, 0.5 km N of Chindul, Hwy. E 15 (0.2389, - 79.8742, 30 m), 1 iii 2010, leg. & det. KJT [KJT]; MS, Huanboya: 1 ♂, 1 ♀, small stream with pools, forested, near Río Pasanac, Hwy. 45 [N of river, E side of Hwy.], (- 1.9550, - 77.8567), 16 ix 2005, leg. & det. KJT [KJT]; Santiago: 1 ♂, [Patuca], [about 1 ml N], 43.4 km S of Sucua, large pond, [about 5 km N of Mendez Road jct] (- 2.75, - 78.27, 488 m), 7 xi 1997, leg. & det. WFM [FSCA]; 1 ♂, 1 ♀, same data but 8 xi 1997, leg. & det. KJT [KJT]; NA, Tena: 1 ♂, Puerto Napo, 5.3 km E of Hwy. 45 on J. S. B. S. Road, Río Sinde (- 1.0500, 77.7481, 450 m), 13 vii 1996, leg. & det. WFM [FSCA]; 1 ♀, Río Chinchipino, 27 km E of Puerto Napo (- 1.0728, - 77.5864, 385 m), 13 xi 1997, leg. & det. KJT [KJT]; 1 ♂, same data but leg. & det. TWD [SWD]; OR, Francisco de Orellana: 1 ♂, P. N. Y., 5.6 km S of Yasuni Road on Waorani Reserve Road, small stream (- 0.7056, - 76.4727, 251 m), 10 vi 1995, leg. & det. WFM [FSCA]; 2 ♂♂, P. N. Y., 29 km N of Waorani Road on Yasuni Road, stream (- 0.4917, - 76.5584, 216 m), 11 vi 1995, leg. & det. WFM [FSCA, QZAC]; 2 ♂♂, ditch, Yasuni Road, 20.5 km N of Sci. Res. Sta., 11 vi 1995, leg. & det. KJT [KJT]; 2 ♂♂, P. N. Y., shallow swamp W of Yasuni road on PC 29 - 30, 10 vi 1995, leg. & det. SWD [SWD]; 1 ♂, Puerto Francisco de Orellana, 12.3 km W on road to Loreto from Coca-Sachas Road, pond (- 0.4750, - 77.0747, 290 m), 13 vi 1995, leg. & det. WFM [USNM]; 2 ♂♂, Puerto Francisco de Orellana, 1.7 km W on Loreto Road from Coca-Sachas Road, stream, pools and flowing shallow marsh (- 0.4367, - 77.0017, 258 m), 13 vi 1995, leg. & det. WFM [FSCA, LSAM]; 2 ♂♂, [Pompeya Sur], S on [Pompeya Sur-Iro road] at km marker 31, borrow pit near Río Sabaleto (- 0.6569, - 76.4528, 228 m), 17 vii 1996, leg. & det. WFM [FSCA, QZAC]; 1 ♂, P. N. Y., tiny stream (- 0.6508, - 76.3558, 200 m), 15 ix 1997, leg. & det. TWD [SWD]; 3 ♂♂, P. N. Y., 2.1 km E of Yasuni Biological Re- search Sta., pond and swamp (- 0.6741, - 76.3685, 244 m), 15 xi 1997, leg. & det. WFM [FSCA, QZAC, USNM]; 1 ♂, forest pool, near Río Savaleto, P. N. Y., 17 xi 1997, leg. & det. KJT [KJT]; Loreto: 1 ♂, small streams and forest trails 12 km E of Loreto, 2 + km S of Hwy. 45 (- 0.6217, - 77.2600, 365 m), 11 x 2009, leg. & det. JTJ [JTJ]; 1 ♂, same data but 15 x 2009, leg. & det. KJT [KJT]; SU, Lago Agrio: 1 ♂, Lago Agrío 7 km W on Hwy. 45, swamp & stream (- 0.08, - 76.93, 314 m), 8 vi 1995, leg. & det. WFM [FSCA]; Shushufindi: 1 ♂, Limoncocha Río Napo (- 0.4000, - 76.6167, 269 m), 5 ix 1972, leg. T. Emmel, det. MJW [FSCA]; 2 ♂♂, same data but 23 viii 1980, leg. & det. SWD [SWD]; 4 ♂♂, same data but swamp 26 vii 1977, leg. DRP & S. Hills, det. DRP [DRP]; 1 ♀, La Selva, Garzacocha, 12 x 1988, leg. & det. SWD [SWD]; ZC, El Pangui: 1 ♂, El Pangui 7 km N, slow muddy stream (- 3.5353, - 78.5333, 754 m), 6 xi 1997, leg. & det. WFM [FSCA]; Zamora: 1 ♂, gravel quarry near Río Jamboe, 4.2 km SE of Zamora, 5 xi 1997, leg. & det. KJT [KJT]; 2 ♂♂, same data but leg. & det. JJD [JJD]. Perithemis lais (Perty, 1933). Publ. Recs. ECU: (H 06), (P 11), (H 14); NA: von Ellenrieder & Muzón (1999); SU: Daigle (2003), * Donnelly (1998). Also ARG, BOL, BRA, COL, GUF, GUY, PER, SUR, VEN: (H 06), (P 11); plus TTO: (H 06). Addl. Data. OR, Francisco de Orellana: 1 ♂, Anangu SE across Río Napo from Providencia Lago Anangu “ on Río Napo ” (- 0.5213, - 76.4361, 300 m), 9 v 1975, leg. T. Emmel, det. MJW [FSCA]; 23 ♂♂, [La Nueva Esperanza], [across Río Napo from] near Primavera “ Limoncocha ”, Río Napo & Lake Taracoa (- 0.45, - 76.77, 300 m), 27 vii 1977, leg. DRP & S. Hills, det. DRP [DRP]; 1 ♂, same data but 13 vii 1977, leg. & det. KWK [SWD]; 16 ♂♂, same data but 26 viii 1980, leg. & det. SWD [KJT, SWD]; 47 ♂♂, 1 ♀, same data but 4 – 21 xi 1980, leg. & det. MJW [FSCA]; 2 ♂♂, same data but 9 vii 1983, leg. H. Gantenbein & P. Gantenbein, det. RWG [CSCA]; 2 ♂♂, [Pompeya Sur], S on [Pompeya Sur-Iro road] at km marker 31, borrow pit near Río Sabaleto (- 0.6569, - 76.4528, 229 m), 17 vii 1996, leg. & det. WFM [LSAM, QZAC]; 1 ♂, P. N. Y., Yasuni Biological Research Sta. along trails and roads within 5 km of Res. Sta. (- 0.6719, - 76.3992, 244 m), 18 vii 1996, leg. & det. WFM [QZAC]; 2 ♂♂, P. N. Y., swamp (- 0.6508, - 76.3558, 200 m), 15 xi 1997, leg. & det. KJT [FSCA]; 1 ♀, P. N. Y, tiny stream (- 0.6542, - 76.3542, 200 m), 15 xi 1997, leg. & det. TWD [SWD]; 1 ♂, Puerto Francisco de Orellana 11.8 km w on Loreto Road large pond (- 0.4742, - 77.0583, 285 m), 18 xi 1997, leg. WFM [FSCA]; 1 ♂, same data but leg. & det. TWD [SWD]; SU, Shushufindi: 1 ♂, Limoncocha, lake edge (- 0.4000, - 76.6167, 269 m), 2 x 1971, leg. D. Pearson, det. DRP [DRP]; 3 ♂♂, same data but 24 ii 1972 [DRP]; 2 ♂♂, same data but 26 vii 1977, leg. & det. DRP [DRP]; 2 ♂♂, same data but 25 vii 1980, leg. & det. SWD [SWD]; 1 ♂, La Selva Lodge; forest (- 0.4983, - 76.3733, 200 m), 23 xi 1997, leg. & det. TWD [NMNHL]; 1 ♂, same data but L. Mandicocha and outlet stream, 25 xi 1997, leg. TWD [SWD]; 6 ♂♂, same data but Garzacocha, 12 x 1988, leg. & det. SWD [SWD]. Perithemis mooma Kirby, 1889. Publ. Recs. ECU: (H 06), (P 11), (H 14); SU: * Donnelly (1998), von Ellenrieder & Muzón (1999); ZC: Tennessen & Johnson (2008). Also ARG, BRA, CEN, COL, GUF, PRY, PER, URY, VEN: (H 06), (P 011); plus TTO: (P 11) and SUR (P 16). Addl. Data. EO, Santa Rosa: [KJT]; Arenillas: [KJT, JTJ]; ES, Esmeraldas: [QZAC]; GY, Naranjal: [FSCA]; MN, El Carmen: [JTJ]; MS, Santiago: [FSCA, LSAM, QZAC, SWD, TWD, USNM]; NA, Tena: [DRP, FSCA, KJT, QZAC, SWD]; OR, Francisco de Orellana: [FSCA, KJT, QZAC, SWD, USNM]; SU, Lago Agrio: [SWD]; Shushufindi: [DRP, FSCA, SWD]; ZC, El Pangui: [FSCA, KJT]; Zamora: [KJT, JTJ]. Note: Garrison et al. (2006) suggest that further studies may reveal that P. mooma is the same as P tenera from North America. Perithemis parzefalli Hoffmann, 1990. Publ. Recs. ECU: (H 06), (P 11); NA: Daigle (1996); OR: Tennessen (1996 b), von Ellenrieder & Garrison (2011); SU: Donnelly (1998). Also PER: (H 06), (P 11). Addl. Data. MS, Huanboya: 3 ♂♂, small stream with pools, forested, near Río Pasanac, Hwy. 45 [N of river, E side of Hwy.] (- 1.9550, - 77.8567), 16 ix 2005, leg. & det. KJT [KJT]; 2 ♂♂, same data but leg. & det. JJD [JJD]; NA, Tena: 2 ♂♂, stream 14.6 km E of Puerto Napo, J. S. B. S. Road, 23 vii 1996, leg. & det. JJD [JJD]; 1 ♂, 5.3 km E of Hwy. 45 on J. S. B. S. Road, Río Sinde (- 1.0500, - 77.7481, 450 m), 13 vii 1996, leg. & det. WFM [FSCA]; 1 ♂, same data but 12 xi 1997, leg. & det. KJT [KJT]; 2 ♂♂, pond along Puyo-Tena Road (- 1.1150, - 77.8083, 600 m), 12 xi 1997, leg. & det. TWD [SWD]; 1 ♂, Río Chinchipino, oxbow pond, 27 km E of Puerto Napo (- 1.0728, - 77.5864, 385 m), 13 xi 1997, leg. & det. KJT [KJT]; 1 ♂, same data but leg. & det. TWD [SWD]; OR, Francisco de Orellana: 9 ♂♂, Primavera, Laguna Taracoa (- 0.4500, - 76.7833, 300 m), 27 vii 1977, leg. DRP & S. Hills, det. DRP [DRP]; 9 ♂♂, same data but 26 viii 1980, leg. & det. SWD [KJT, SWD]; 1 ♂, pond on Coca-Loreto road (- 0.4742, - 77.0583, 285 m), 18 xi 1997, leg. & det. TWD [SWD]; 1 ♂, [Pompeya Sur] S on [Pompeya Sur-Iro road] at km marker 8, swamp & stream (- 0.4694, - 76.5869, 253 m), 12 vi 1995, leg. & det. WFM [LSAM]; 1 ♂, P. N. Y., 2.3 km E of Yasuni Biological Research Station, small stream (- 0.67, - 76.37, 213 m), 9 vi 1995, leg. & det. WFM [FSCA]; 5 ♂♂, same data but leg. & det. SWD [JJD, SWD]; 2 ♂♂, 1 ♀, same data but leg. & det. KJT [KJT]; 1 ♂, P. N. Y., 5.6 km S of Yasuni Rd on Waorani Reserve Road, small stream (- 0.7056, - 76.4727, 251 m), 10 vi 1995, leg. & det. WFM [FSCA]; 1 ♂, same data but leg. & det. KJT [KJT]; 2 ♂♂, P. N. Y., 6.3 km S of Yasuni Road on Waorani Reserve Road, marshy pond (- 0.7120, - 76.4745, 259 m), 10 vi 1995, leg. & det. WFM [QZAC, USNM]; 1 ♂, P. N. Y., swamp (- 0.6508, - 76.3558, 200 m), 15 xi 1997, leg. & det. KJT [KJT]; 2 ♂♂, P. N. Y., small stream / pond, 3.5 km W of Yasuni Road on PC 29 - 30, 10 vi 1995, leg. & det. SWD [SWD]; 2 ♂♂, P. N. Y., grassy semi-shaded pools / stream, 6.2 km W of Yasuni Road on PC 29 - 30, 10 vi 1995, leg. & det. SWD [SWD]; 2 ♂♂, 1 ♀, swamp ponds, 2.3 km E on Yasuni Road, 9 vi 1995, leg. & det. SWD [SWD]; 1 ♂, [Pompeya Sur] S on [Pompeya Sur-Iro road] at km marker 31, borrow pit near Río Sabaleto (- 0.6569, - 76.4528, 229 m), 12 vi 1995, leg. & det. WFM [FSCA]; 4 ♂♂, same data but 17 vii 1996, [FSCA, QZAC, USNM]; 1 ♂, same data but leg. & det. SWD [SWD]; 1 ♂, same data but 17 xi 1997 [FSCA]; SU, Cascales: 2 ♂♂, stream, 44 km W of Lago Agrio, 20 viii 1980, leg. & det. SWD [SWD]; 1 ♂, swamp, 26 km S of Lago Agrio, 19 viii 1980, leg. & det. SWD [SWD]; Shushufindi: 1 ♂, Limoncocha (- 0.4000, - 76.6167, 269 m), 30 viii 1980, leg. & det. SWD [SWD]; 1 ♂, same data but swamp, 26 vii 1977, leg. DRP & S. Hills, det. DRP [DRP]; 1 ♂, [Limoncocha] [about 40 km E], or 100 km E of Coca [on Río Napo], Lago Garzacocha, La Selva Jungle Lodge (- 0.4819, - 76.3733, 235 m), 11 x 1988, leg. & det. SWD [FSCA]; 10 ♂♂, same data but 12 x 1988 [JJD, SWD — as “ P. midasi ”); ZC, Nangaritza: 1 ♂, Orquideas, 3 km S, Cabanas Yankuam, next to the Río Nangaritza (- 4.1625, - 78.6419, 914 m), late November, 2007, photo. by Martin Reid. Note: Originally to be described by L. K. Gloyd with the manuscript name “ P. midasi; ” there are specimens labeled as “ P. midasi ” in various collections. Perithemis rubita Dunkle, 1982. Publ. Recs. ECU: Paulson (1983), Davies & Tobin (1984), Tsuda (1986, 1991), Steinmann (1997 b), (H 06), (P 11), (H 14); OR: * Dunkle (1982 — Holotype & Allotype “ Lake Taracoa near Primavera, NA ” [FSCA]), Bridges (1993); SU: * Dunkle (1982), Daigle (2003). Addl. Data. OR, Francisco de Orellana: 6 ♂♂, 1 ♀, paratypes, [La Nueva Esperanza], Lake Taracoa, swamp on north side (- 0.45, - 76.77, 244 m), 26 viii 1980, leg. & det. SWD [FSCA, SWD]; SU, Shushufindi: 1 ♂, Limon- cocha, swamp, 26 vii 1977, leg. DRP & S. Hills, det. DRP [DRP]; 1 ♂, La Selva, Garzacocha, 13 x 1988, leg. & det. SWD [SWD]. Perithemis thais Kirby, 1889. Publ. Recs. ECU: (H 06), (P 11), (H 14); OR: * Tennessen (1996 b), von Ellenrieder & Garrison (2011); SU: Donnelly (1998). Also ARG, BRA, GUF, GUY, PER, TTO, VEN: (H 06), (P 11); Plus CEN: (H 06); and COL: Palacino et al. (2017). Addl. Data. MS, Huamboya: 2 ♂♂, Río Pasanac pools, 57 km N of Macas, Hwy. 45, 16 ix 2005, leg. & det. JJD [JJD]; NA, Tena: 3 ♂♂, Río Chinchipino marsh, 27.2 km E of Puerto Napo (- 1.0728, - 77.5864, 385 m), 13 xi 1997, leg. & det. JJD [JJD]; OR, Francisco de Orellana: 1 ♂, P. N. Y. (300 m), 13 x 2001, leg. D. Bogar, det. C. Shiffer [FSCA]; 1 ♂, 1 ♀, same data but leg. & det. TWD [SWD]; 1 ♂, P. N. Y., 3.6 km S of Yasuni Road on Wa- orani Reserve Road (- 0.6956, - 76.4594, 244 m), 10 vi 1995, leg. & det. WFM [FSCA]; 1 ♂, P. N. Y., 2.3 km E on Yasuni Road, 9 vi 1995, leg. & det. SWD [SWD]; 2 ♂♂, pond, road to Waorani Reserve, 5.6 km S of turn to Yasuni Sci. Res. Sta., 10 vi 1995, leg. & det. KJT [KJT]; 1 ♂, 1 ♀, stream, PC 29 - 30 Road, 3.2 km from jct. with Yasuni Road, 18 vii 1996, leg. & det. KJT [KJT]; 1 ♂, slow stream, 6.8 km from Yasuni Sci. Sta., 15 xi 1997, leg. & det. KJT [KJT]; 1 ♂, P. N. Y., swamp (- 0.6508, - 76.3558, 200 m), 15 xi 1997, leg. & det. KJT [KJT]; 1 ♂, same data but tiny stream, 15 xi 1997, leg. & det. TWD [SWD]; 1 ♂, T. B. S., USFQ, stream crossing Matapalo trail at marker 450 (- 0.6369, - 76.1458, 245 m), 14 i 2009, leg. & det. RWG & NVE [NVE]; 2 ♂♂, T. B. S., USFQ, stream crossing Cho- rongo trail at marker 225 (- 0.6350, - 76.1503, 220 m), 16 – 18 i 2009, leg. & det. RWG & NVE [USFQ]; 1 ♂, T. B. S., USFQ, blackwater stream crossing Guacamayo trail at marker 2050 (- 0.6383, - 76.1492, 220 m), 17 i 2009, leg. & det. RWG & NVE [NVE]; 1 ♂, T. B. S., USFQ, Lago trail crossing river (- 0.6381, - 76.1558, 219 m), 21 i 2009, leg. & det. RWG & NVE [RWG]; Loreto: 1 ♂, stream about 12 km NE of Loreto (- 0.6217, - 77.2600, 365 m), 15 x 2009, leg. & det. KJT [FSCA]; SU, Shushufindi: 1 ♂, Shushufindi; La Selva Lodge; forest (- 0.4983, - 76.3733, 200 m), 25 xi 1997, leg. & det. TWD [SWD]; 1 ♂, stream about 3 km NW of Shushufindi (- 0.1664, - 76.6917, 290 m), 13 v 2009, leg. & det. KJT [KJT].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B5D6FF13FF23F68824F3F9B1.taxon	discussion	For species identification see Ris (1912) and Santos (1949 b). A small group, mature males of most species with brilliant red on dorsum of abdomen (Fig. 109). Frequent ponds with emergent and floating vegetation. Very little is known about these wary dragonflies. Ecuador is a new country record for the genus. * Planiplax machadoi Santos, 1949. Publ. Recs. ECU: (H 14). Also BRA, PER: (H 06), (P 11). Addl. Data. OR, Francisco de Orellana, 3 ♂♂, pond on Coca-Loreto Road (- 0.4800, - 77.0800, 300 m), 18 xi 1997, leg. & det. TWD [SWD]; 1 ♂, Puerto Francisco de Orellana 11.8 km W on Loreto Road, large pond (0.4742, - 77.0583, 285 m), 18 xi 1997, leg. & det. WFM [FSCA]; 2 ♂♂, same data but leg. & det. KJT [KJT]; SU, Shushufindi: 2 ♂♂, pond about 3 km NW of Shushufindi (- 0.1822, - 76.6917, 265 m), 14 v 2009, leg. & det. KJT [FSCA]; Shushufindi, 2 km W on oil pipeline road, Río Shushufindi and isolated pond, date not provided, data provided by WAH & DLW. * Planiplax phoenicura Ris, 1912. Also BRA, CEN, GUF, GUY, SUR, TTO, VEN: (P 11); plus COL: (P & P). Addl. Data. MN, El Carmen: 1 ♂, borrow pits about 18 km NW of El Carmen (- 0.1350, - 79.5797, 185 m), 2 iii 2010, leg. & det. KJT [FSCA].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B5D6FF11FF23F04E2647FEDB.taxon	discussion	See Belle (1998) for species identification. The genus inhabits ponds, ditches and overflow areas along large rivers, usually with little or no aquatic vegetation. This is a small group of rather large, bright red or carmine dragonflies (Fig. 110), with three species in Ecuador. Rhodopygia cardinalis (Erichson in Schomburgk, 1848). Publ. Recs. ECU: (H 06), (P 11), (H 14); MS: Belle (1998); NA: * Tennessen (1996 b), Daigle (1996). Also BRA, GUF, GUY PER, SUR, VEN: (H 06), (P 11); plus COL: (P & P). Addl. Data. MS, Morona: 2 ♂♂, Macas, Río Upano (- 2.3167, - 78.1167, 1050 m), xii 1956, leg. L. Alonzo, det. J. Belle [NMNHL]; NA, Tena: adult, 3 km S. Tena, 13 v 1977, leg. P. Spangler, Paul J & D. Givens, det.? [USNM]; 2 ♂♂, 1 ♀, Puerto Napo, 5.3 km E of Hwy. 45 on J. S. B. S. Road, Río Sinde (- 1.0500, - 77.7481, 450 m), 16 x 1988, leg. & det. SWD [JJD]; 1 cop. pair, 4 ♂, same data but 18 x 1988 [FSCA]; 1 ♂, same data but 13 vii 1996, leg. & det. WFM [FSCA]; 1 ♂, same data but leg. & det. JJD [JJD]; 1 ♂, same data but leg. & det. SWD [UTIC]; 1 ♂, same data but 22 vii 1996, leg. & det. WFM [FSCA]; 1 ♂, same data but leg. & det. JJD [JJD]; 1 ♂, pool, road to J. S. B. S., 14.6 km E Puerto Napo, 22 vii 1996, leg. & det. KJT [KJT]; 11 ♂♂, 2 ♀♀, 1 cop. pair, Puerto Napo 3.2 km E on J. S. B. S. Road ponds and flowing marsh (- 1.0456, - 77.7467, 420 m), 23 vii 1996, leg. & det. WFM [FSCA, LSAM, QZAC, USNM]; 1 ♂, same data but leg. & det. JJD [JJD]; 3 ♂♂, 1 ♀, same data but leg. & det. KJT [KJT]; 1 ♂, same data but leg. & det. SWD [SWD]; 1 ♂, 1 ♀, [Puerto Napo] pool near stream, 17.2 km E on J. S. B. S. Road (341 m), 22 vii 1996, leg. & det. SWD [SWD]; OR, Francisco de Orellana: 1 ♂, trib. of Río Payamino, road to Loreto, 1.7 km W of jct. with Coca Road (- 0.4367, - 77.0017, 258 m), 13 vi 1995, leg. & det. SWD [SWD]; 1 ♂, [Pompeya Sur], S on [Pompeya Sur-Iro road] at km marker 31, borrow pit near Río Sabaleto (- 0.6569, - 76.4528, 229 m), 17 vii 1996, leg. & det. WFM [FSCA]; 1 ♂, same data but leg. & det. KJT [KJT]; 1 ♂, same data but leg. & det. SWD [SWD]; 1 ♂, marsh at Payamino (- 0.4367, - 77.0017, 258 m), 18 xi 1997, leg. & det. TWD; SU, Lago Agrio: 1 ♂, Santa Cecilia. Río Francisco de Orellana, 25 – 31 iii 1969, leg. P. & P Spangler, det.? [USNM]; 1 ♂, swamp, 17 km S of Lago Agrio, 19 viii 1980, leg. & det. SWD [SWD]; Shushufindi: 4 ♂♂, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 12 – 25 xi 1980, leg. & det. MJW [FSCA]; 1 ♂, same data but lake edge, 6 i 1972, leg. D. Pearson, det. DRP [DRP]; 1 ♂, same data but 24 ii 1972 [DRP]; 1 ♂, same data but 24 vii 1977, leg. & det. DRP [DRP]; 1 ♂, same data but 26 vii 1977, leg. DRP & S. Hills, det. DRP [DRP]; 1 ♀, same data but 28 viii 1980, leg. & det. SWD [SWD]; 2 ♂♂, swamp / stream 2 km S of Yamaramsuku, 15.7 km S Shushufindi on Limoncocha Road, 19 vii 1996, leg. & det. SWD [RWG, SWD]; 1 ♂, same data but leg. & det. KJT [KJT]. Rhodopygia hinei Calvert, 1907. Publ. Recs. ECU: * Paulson (19830, (P 11), (H 14). Also CEN: (H 06), (P 11). Addl. Data: LR, Buena Fé: 1 ♂, 1 ♀, Santo Domingo, 47 km S, forest slough at Río Palenque Field Station (- 0.5500 - 79.3667, 151 m), 18 vii 1977, leg. & det. DRP, checked by RWG [DRP]. Rhodopygia pruinosa Buchholz, 1953. Publ. Recs. ECU: (H 06), (P 11), (H 14); NA: * Daigle (1996). Also BOL, BRA, GUF, GUY, (H 06), (P 11); plus SUR (P 16). Addl. Data. NA, Tena: 1 ♂, pool, road to J. S. B. S., 14.6 km E of Puerto Napo, 22 vii 1996, leg. & det. KJT [FSCA]; 1 ♂, Puerto Napo, 5.3 km E of Hwy. 45 on J. S. B. S. Road, Río Sinde (- 1.0500, - 77.7481, 450 m), 18 x 1988, leg. & det. SWD [SWD]; 1 ♂, same data but 13 vii 1996, leg. & det. WFM [FSCA]; 3 ♂♂, Puerto Napo, 3.2 km E on J. S. B. S. Road, ponds and flowing marsh (- 1.0458, - 77.7467, 420 m), 23 vii 1996, leg. & det. WFM [LSAM, QZAC, USNM]; 3 ♂♂, same data but leg. & det. JJD [JJD]; 2 ♂♂, same data but leg. & det. KJT [KJT]; 1 ♂, 1 ♀, same data but leg. & det. SWD [SWD]; SU, Lago Agrio: 1 ♂, Lago Agrío (0.1000, - 76.8667, 396 m), 24 viii 1975, leg. A. Langley & J. Cohen, det. RWG, from USNM [RWG].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B5D4FF11FF23F7A92723F94E.taxon	discussion	A large genus of mostly red dragonflies widely distributed around the world but with only one species in Ecuador (Fig. 111), occurring only at high elevations. For identification see De Marmels (2001 b). Sympetrum gilvum (Selys, 1884). Publ. Recs. ECU: Maria (1938 — as S. illotum var. gilva), Paulson (1983 — as S. illotum), Tsuda (1991), (H 06), (P 11), (H 14); AZ: Martin (1896 — as Diplax illota), Calvert (1906), Campos (1922 — as D. illota var gilva); CB: Campos (1922 — as D. illota var gilva); LJ: Tennessen & Johnson (2008); NA: Tennessen (1996 b), Daigle (1996); PC: * Selys (1884 — as D. illota var gilva), Calvert (1906), Ris (1911 b), Campos (1922 — as D. illota var gilva); TU: Ris (1911 b), Tennessen (2006). Also ARG, CHL, COL, PER, VEN: (P 11). Addl. Data. AZ, Gualaceo: [SWD]; CB, Alausí: [FSCA]; Pallatanga: [ANSP, FSCA]; IM, Cotacachi: [FSCA, KJT, USFQ]; Ibarra: [FSCA]; Otavalo: [FSCA, KJT, QZAC, SWD]; LJ, Catamayo: [FSCA, JTJ, KJT]; NA, Archidona: [SWD]; El Chaco: [SWD]); Quijos: [FSCA, KJT, LSAM, QZAC, SWD, USNM]; PC, Mejía: [DRP, FSCA]; Quito: [FSCA, UTIC, QZAC, USNM]; San Miguel Bancos: [FSCA, KJT, USNM]; TU, Baños: [FSCA, KJT, QZAC, USNM]. Note: There are many references and specimens labeled S. illotum since S. gilvum was considered to be either a junior synonym or subspecies of it.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B5D4FF1FFF23F1542437FD83.taxon	discussion	Of the five species of Tauriphila, three were keyed by Needham et al. (2014), two of which are wide-ranging and are found in Ecuador (Fig. 112). Breeding sites are open ponds and small lakes with aquatic macrophytes. Tauriphila argo (Hagen, 1869). Publ. Recs. ECU: (H 06), (P 11), (H 14); GY: Tennessen & Johnson (2008); NA: Tennessen (1998); OR: * Dunkle (1981). Also ARG, BOL, BRA, CEN, COL, GUF, GUY, PRY, PER, VEN, TTO: (H 06), (P 11); plus SUR (P 16). Addl. Data. GY, Balao: 1 ♂, small stream / ditches in banana plantation, about 2 km E of Hwy. 25 near Balao Jct. (- 2.9100, - 79.6919, 70 m), 28 iii 2008, leg. & det. JTJ [JTJ]; 1 ♂, same data but leg. & det. KJT [KJT]; NA, Tena: 1 ♂, Puerto Napo, 5.3 km E of Hwy. 45 on J. S. B. S. Road, Río Sinde (- 1.0500, - 77.7481, 450 m), 22 vi 1996, leg. & det. WFM [LSAM]; 4 ♂♂, same data but 16 x 1988, leg. & det. SWD [SWD]; 1 ♂, Puerto Napo, 3.2 km E on J. S. B. S. Road, ponds and flowing marsh (- 1.0456, - 77.7467, 420 m), 23 vii 1996, leg. & det. WFM [QZAC]. OR, Francisco de Orellana: 2 ♂♂, pond on Coca-Loreto road (- 0.4750, - 77.0747, 290 m), 18 xi 1997, leg. & det. TWD [SWD]; 3 ♂♂, [La Nueva Esperanza] [across Río Napo from] near Primavera “ Limoncocha ” Río Napo & Lake Taracoa (- 0.45, - 76.77, 300 m), 27 vii 1977, leg. DRP & S. Hills, det. DRP [DRP]; 1 ♀, same data but 20 viii 1980, leg. & det. SWD [SWD]; 1 ♂, same data but 4 xi 1980, leg. & det. MJW [FSCA]; 1 ♂, trib. of Río Payamino, road to Loreto, 1.7 km W of jct. with Coca Road (- 0.4367, - 77.0017, 258 m), 18 xi 1997, leg. & det. KJT [KJT]; 1 ♂, 1 ♀, Puerto Francisco de Orellana, 8.2 km W on road to Loreto from Coca-Sachas Road, pond (- 0.4578, - 77.0381, 288 m), 18 xi 1997, leg. & det. WFM [FSCA]; 1 ♂, Puerto Francisco de Orellana, 12.3 km W on road to Loreto from Coca-Sachas Road, pond (- 0.4750, - 77.0747, 290 m), 13 vi 1995, leg. & det. WFM [FSCA]; 1 ♂, same data but leg. & det. SWD [SWD]; SU, Lago Agrio: 2 ♂♂, brushy ponds, 24 km W of Lago Agrio, 20 viii 1980, leg. & det. SWD [SWD]; Shushufindi: 1 ♂, pond about 3 km NW of Shushufindi (- 0.1822, - 76.6917, 290 m), 14 v 2009, leg. & det. KJT [FSCA]; 2 ♂♂, Shushufindi, SW, pond, (- 0.2201, - 76.6988), 29 x 2013, leg. & det. KJT [FSCA].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B5DAFF1CFF23F48021BAFEDB.taxon	discussion	A genus with only two species, one of which is widely distributed in the Neotropics and also occurs in subtropi- cal America (the other species occurs in the Old World tropics). Adults (Fig. 113) are normally crepuscular but sometimes fly during midday. Shallow mud-bottomed ponds and ditches are the preferred habitat. The nymph was described by Fleck et al. (2004). Tholymis citrina Hagen, 1867. Publ. Recs. ECU: Ris (1913 a), Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14); CR: Campos (1922); EO: Campos (1927); LR: * Calvert (1906), Campos (1922); OR: Dunkle (1981). Also ARG, BRA, CEN, CHL, COL, GUF, GUY, PRY PER, VEN: (H 06), (P 11); plus TTO: (P 11) and SUR: (P 16). Addl. Data. SD: 1 ♀, Santo Domingo de los Colorados, 8 i, leg. & det.? [QZAC]; SU, Shushufindi: 1 ♂, Li- moncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 1 xi 80, leg. & det. MJW [FSCA]; 1 ♂, same data but 25 xi 1980 [FSCA]; 1 ♂, ditch S of Limoncocha (- 0.4203, - 76.6108), 11 v 2009, leg. & det. KJT [FSCA].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B5D9FF1AFF23F7A92646FB8B.taxon	discussion	See Ris (1913) and De Marmels & Rácenis (1982) for species identification. Adults (Fig. 114) perch in open areas near lentic habitats. Females oviposit in tandem. Four species of Tramea occur in Ecuador. Tramea abdominalis (Rambur, 1842). Publ. Recs. ECU: (H 06), (P 11), (H 14): GY: * Campos (1922); LR; * Campos (1922). Also ARG, BRA, CEN, COL, PRY, PER, TTO, VEN: (H 06), (P 11); plus SUR: (P 11). Addl. Data. ES, San Lorenzo: 1 ♂, pool, 36.5 km NW of Lita, near Urbina, 3 ii 1997, leg. & det. KJT [KJT]; MS, Santiago: 2 ♂♂, 1 ♀, [Patuca] [about 1 km N], 43.4 km S of Sucua, large pond [about 5 km N of Mendez road jct] (- 2.7283, - 78.2700, 600 m), 6 xi 1997, leg. & det. TWD; 2 ♂♂, same data but 7 xi 1997, leg. & det. WFM [FSCA, QZAC]; 1 ♂, same data but leg. & det. KJT [KJT]; 1 ♂, same data but 8 xi 1997; 2 ♂♂, 1 ♀, 1 tan. pair, same data but leg. & det. WFM [FSCA, QZAC]; NA, Tena: 1 ♂, Puerto Napo, 23 km E, J. S. B. S., tropical wet forest (- 1.07, - 77.60, 450 m), 18 x 1988, leg. & det. SWD [SWD]; 1 ♂, J. S. B. S., Pond # 14, 23 xi 1991, leg. E. McCallie, det.? [USNM]; 2 ♂♂, J. S. B. S., Pond # 16 A, 16 – 23 xi 1991, leg. B. Sharp, det.? [USNM]; 1 ♂, J. S. B. S., near Tena on building (450 m), 20 vi 1994, leg. A. Jaramillo, det. DRP [DRP]; 1 ♂, Puerto Napo, 5.3 km E on Jatun Sacha Road, Río Sinde, swamp and pond approx. 1 – 2 km upstream (- 1.0517, - 77.7544, 450 m), 13 vii 1996, leg. & det. KJT [KJT]; 2 ♂♂, same data but 12 xi 1997, leg. WFM [FSCA, QZAC]; 1 ♀, same data but 15 vi 1995, leg. & det. SWD [SWD]; 1 cop. pair, pool near Río Napo, 3.2 km E of Puerto Napo (- 1.0456, - 77.7467, 420 m), 23 vii 1996, leg. & det. KJT [KJT]; 1 tan. pair, same data but leg. & det. WFM [QZAC]; 1 ♂, same data but leg. & det. SWD [SWD]; OR, Francisco de Orellana: 1 ♀, P. N. Y., 6.2 km W of Yasuni Road on PC 29 - 30, pool / stream, 10 vi 1995, leg. & det. SWD [SWD]; PC, San Miguel de los Bancos: 1 ♂, Mindo, Mindo Gardens Ecolodge, vegetated cement pond (- 0.0270, - 78.7560, 1330 m), 3 ix 2018, leg. & det. WFM [FSCA]; SU, Shushufindi: 1 ♂, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 11 v 1975, leg. B. Drummond III, det. WFM [FSCA]; 11 ♂♂, 1 ♀, same data but 8 – 26 xi 1980, leg. & det. MJW [FSCA]; ZC, Zamora: 1 ♂, tilapia ponds and small streams, Hosteria El Arenal, about 1 km S of Cumbaratza (- 4.0017, - 78.8753, 950 m), 3 iv 2008, leg. & det. JTJ [JTJ]. Tramea binotata (Rambur, 1842). Publ. Recs. ECU: * Navás (1923), Campos (1925), Maria (1938), Racenis (1954 — as Trapezostigma binotata), Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14); GY: Ris (1913 b). Also ARG, BRA, CEN, COL, GUF, PRY, PER, SUR, TTO, VEN: (H 06), (P 11). Addl. Data. MS, Santiago: 1 ♀, [Patuca], [about 1 km N], 43.4 km S of Sucua, large pond [about 5 km N of Mendez road jct] (- 2.75, - 78.27, 488 m), 7 xi 1997, leg. & det. WFM [QZAC]; 2 ♂♂, same data but 8 xi 1997 [FSCA, QZAC]; 2 ♂♂, 1 ♀, same data but 6 xi 1997, leg. & det. TWD [SWD]; NA, Tena: 2 ♂♂, Tena (500 m), leg. F. Campos, det. MJW [ANSP]; 1 ♂, J. S. B. S., Pond # 4, 14 xi 1991, leg. E. McCallie, det.? [USNM]; 1 ♂, J. S. B. S., near Tena, pond (450 m), 18 vi 1994, leg. A. Jaramillo, det. DRP [DRP]; 1 ♂, Puerto Napo, 5.3 km E of Hwy. 45 on J. S. B. S. Road, Río Sinde (- 1.0500, - 77.7481, 450 m), 22 vii 1996, leg. & det. WFM [LSAM]; 1 ♀, same data but 16 x 1988, leg. & det. SWD [SWD]; 1 ♂, same data but 9 x 2009, leg. & det. JTJ [JTJ]; 1 cop. pair, Puerto Napo, 31.7 km E of Hwy. 45 on J. S. B. S. Road, marshy area (- 1.0572, - 77.5923, 396 m), 13 vii 1996, leg. & det. WFM [QZAC]; OR, Francisco de Orellana: 1 ♂, ditch, Yasuni Road, 20.5 km N of Sci. Res. Sta., 11 vi 1995, leg. & det. KJT [KJT]; 1 ♂, same data but leg. & det. SWD [SWD]; 1 ♂, P. N. Y., 29 km N of Waorani Road on Yasuni Road, road- side ditch and pond (- 0.4917, - 76.5584, 216 m), 11 vi 1995, leg. & det. WFM [FSCA]; 2 ♂♂, Puerto Francisco de Orellana, 12.3 km W on road to Loreto from Coca-Sachas Road, pond (- 0.4750, - 77.0747, 290 m), 13 vi 1995, leg. & det. WFM, [FSCA, QZAC]; 1 ♂, same data but leg. & det. KJT [KJT]; 3 ♂♂, Puerto Francisco de Orellana 11.8 km W on Loreto Road, large pond (- 0.4742, - 77.0583, 285 m), 18 xi 1997, leg. & det. WFM [FSCA, LSAM. QZAC]; 1 ♂, same data but leg. & det. KJT [KJT]; PA, Santa Clara: 1 ♂, 39 km N of Puyo, 5 vi 1977, leg. P. Spangler & D. Givens, det.? [USNM]; Pastaza: 1 ♂, pond 3 km N of Comunidad Chuwitayo, S of Puyo, 9 xi 1997, leg. & det. KJT [KJT]; 1 ♀, Cuasha, approximately 1.5 km N on Hwy. 45, [91.5 km N of Sucua], hard bottomed pond (- 1.85, - 77.82, 884 m), 9 xi 1997, leg. & det. WFM [FSCA]; ZC, Zamora: 1 ♂, tilapia ponds and small streams, Hosteria El Arenal, about 1 km S of Cumbaratza (- 4.0017, - 78.8753, 950 m), 3 iv 2008, leg. & det. JTJ [JTJ]. Tramea calverti Muttowski, 1911. Publ. Recs. ECU: Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14); EO: Campos (1927 — as T. longicauda var.?); GA: Dunkle (1989), Peck (1992), (H 06), (P 11); GY: * Campos (1922 — as T. longicauda? var.), De Marmels & Racenis (1982); MN: De Marmels & Racenis (1982); NA: De Marmels & Racenis (1982); ZC: De Marmels & Racenis (1982). Also ARG, BOL, BRA, CEN, COL, GUF, GUY, PER, SUR, TTO, VEN: (H 06), (P 11); plus URG: (P 16). Addl. Data. ES, Atacames: 1 ♂, Castelnuovo, iv 1987, leg. & det.? [QZAC]; GY, Duran: Duran, date?, leg. F. Campos, det.? [USNM]; Guayaquil: 1 ♂, 30 xi 1990, C. Anderson, det. DRP [DRP]; 3 ♂♂, 2 ♀♀, Puna Veija (- 2.97, - 80.18, 30 m), leg. F. Campos, det. MJW [ANSP, FSCA]; Naranjal, 1 ♂, 3 ♀♀, Bucay (- 2.2, - 79.18, 233 m), date?, leg. F. Campos, det. MJW [FSCA]; 1 ♀, [San Rafael] as “ San Raphael ”, (- 2.18, - 79.23, 200 m), date?, F. Campos, det. JJD [FSCA]; 1 ♂, 1 ♀, “ Finca de la Gringa, ” 15 km E of Tres Cerritos (- 2.4456, - 79.5556, 25 m), 12 iv 2008, leg. & det. KJT [KJT]; 4 ♂♂, 1 ♀, Cerro Mas Vale Fundaction Andrate (- 2.40, - 79.63, 15 m), 9 ii 2001, leg. & det. F. Sibley [FSCA]; 1 ♂, Cerro Mas Vale E on Hwy. Río Churute (- 2.42, - 79.67, 9 m), 10 ii 2001, leg. & det. F. Sibley [FSCA]; LR, Babahoyo: 1 ♀, Babahoyo, date?, leg. F. Campos, det.? [USNM]; 1 ♀, 6 km N of Babahoyo, 21 vi 1975, leg.? det.? [USNM]; Buena Fé: 1 ♂, 2 ♀♀, Santo Domingo, 47 km S, clearing at Río Palenque Field Station (200 m), 18 vii 1977, leg. & det. DRP [DRP]; 1 ♀, [Consumulo] S, Río Palenque (- 0.58, - 79.35, 175 m), 20 xi 1972, leg. D. Dodson, det. GHB [FSCA]; Ventanas: 1 ♂, ditches 2.8 km E of Hwy. E 25, about 13 km SW of Quinzaloma, 21 ii 2010, leg. & det. JJD [JJD]; MN, Chone: 1 ♀, 4.4 km W, near stream, 23 x 1978, leg. E. Long, det.? [USNM]; NA, Tena: 1 ♂, Tena (- 0.98, - 77.80, 500 m), leg. F. Campos, det. MJW [FSCA]; 1 ♀, J. S. B. S., 400 m, v 1991, leg. & det.? [QZAC]); 2 ♂♂, same data but 18 x 1988, leg. & det. SWD [SWD]; 2 ♂♂, 1 ♀, Puerto Napo, 5.3 km E of Hwy. 45 on J. S. B. S. Road, Río Sinde (- 1.0500, - 77.7481, 450 m), 16 – 18 x 1988, leg. & det. SWD [SWD]; OR, Francisco de Orellana: 1 ♂, trib. of Río Payamino, road to Loreto, 1.7 km W of jct. with Coca Road (- 0.4367, - 77.0017, 250 m), 13 vi 1995, leg. & det. KJT [KJT]; SU, Shushufindi: 1 ♂, Limoncocha, Río Napo (- 0.4000, - 76.6167, 269 m), 14 xi 1980, leg. & det. MJW [FSCA]. Note: Due to the confusion between this and the next species, T. cophysa, some of the historical records listed may refer to either species. Tramea cophysa (Hagen, 1861). Publ. Recs. ECU: Paulson (1983), Tsuda (1986, 1991), (H 11), (P 11), (H 14); CR: Campos (1922), EO: Campos (1927); GA: * Kirby (1889 — description of ♀,), Kirby (1890), Currie (1901), Calvert (1906), Ris (1913 b — as “ form b ”), Campos (1922), Davies & Tobin (1984 — all as T. darwini), Calvert (1947 b), Asahina (1961 — as T. cophysa darwini), Gloger (1964), Turner (1967), De Marmels & Racenis (1982), Peck (1992), (H 06), (P 11). The Tramea sp. reported by McLachlan (1877) from Galapagos is probably this species according to Currie (1901). GY: Calvert (1906), Ris (1913 b), Campos (1922), Campos (1930), Campos (1931 — as T. basalis), Turner (1967); LR: Calvert (1906), Campos (1922); MN: De Marmels & Racenis (1982); All the Galapagos records are for T. cophysa, De Marmels & Racenis (1982). Also ARG, BOL, BRA, GUY, PRY, PER, TTO, URY, VEN: (H 06), (P 11); plus COL: (P & P); and SUR: (P 16). Addl. Data. AZ, Cuenca: 1 ♀, La Iberia, N of Machala, banana plantation, 3 vii 1990, leg. P. Donahue, det. DRP [DRP]; GA, Isabela: 1 ♂, 1 ♀, Isabela Is., Tagus Cove, 14 xii 1973, leg. J. Harper, det. KJT [KJT]; 3 ♂♂, Isla Isabela, littoral zone, 2 – 4 iii 1989, leg. S. Peck, det.? [USNM]; San Cristóbal: 3 ♂♂, 1 ♀, Hacienda el Cafetal, charca (- 0.9242, - 89.5388, 191 m), 21 – 23 ii 2014, leg. & det. ACRP [ACRP]; Santa Cruz: ♀, Santa Cruz Island, 1948, leg. K. Vinton, det.? [USNM]; 2 ♀♀, Isla Floreana, Sea Lion Pt., littoral, Black Beach, 21 iii 1989, leg. S. Peck, det.? [USNM]; 1 ♂, Isla Santiago, Playa Espumilla, arid zone, UV and night collection, 4 iv 1992, leg. J. Cook & S. Peck, det.? [USNM]; 4 ♂♂, 4 ♀♀, Isla Santiago, Jaboncillo Camp, grasslands, 8 - 9 iv 1992, leg. J. Cook & S. Peck, det.? [USNM]; Isla Santiago, Central Camp, grasslands, general and UV light collection, 9 – 11 iv 1992, leg. J. Cook & S. Peck, det.? [USNM]; 3 ♀♀, Isla Santiago, Playa Espumilla, arid zone, 12 – 24 iv 1992, leg. J. Cook & S. Peck, det.? [USNM]; 1 ♀, Isla Sta. Cruz, lagoon 2 km E. CDRS, littoral zone, UV light & gen. collection, 9 v 1992, leg. J. Cook & S. Peck, det.? [USNM]; GY, Guayaquil: 1 ♀, Guayaquil (- 2.20, - 79.90, 8 m), i 1936, leg.?, det. JJD [FSCA]; LR, Montalvo: 1 ♂, pond in rice field 0.75 km N of Hwy. 491, about 25 km E of Babahoyo (- 1.8083, - 79.3111, 60 m), 7 iii 2010, leg. & det. JTJ [JTJ]; SD: 1 ♂, 1 ♀, Río Blanco, 23 km NNW Santo Domingo de los Colorados, 21 xii 1975, leg. J. Harper, det. KJT [KJT].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B5DFFF1BFF23F26E2660F97A.taxon	discussion	See Ris (1911 a) and Costa & Santos (1997) for species identification. Adults (Fig. 115) perch in shaded forest for the most part but can be found also in partly shaded spots near the forest edge. Females oviposit in damp mud in forested swales that will flood and hold water for a short period of time. No nymphs have been described. Five species of Uracis occur in Ecuador. Uracis fastigiata (Burmeister, 1839). Publ. Recs. ECU: Campos (1922 “ por comm. Ris), Maria (1938), Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14); ES: Daigle et al. (1997); IM: * Ris (1911 a); OR: von Ellenrieder & Garrison (2011); PA: (Ris 1911 a); SU: Donnelly (1998). Also BOL, BRA, CEN, COL, GUF, GUY, PER, TTO, VEN: (H 06), (P 11); plus SUR: (P 16). Addl. Data. ES, San Lorenzo [FSCA, QZAC, USNM]; NA, Carlos Julio Arsemena Tola: [FSCA]; Tena: [DRP, FSCA, SWD, USNM]; OR, Francisco de Orellana: [DRP, FSCA, KJT, LSAM, NVE, QZAC, SWD, USFQ, USNM]; PA, Mera: [FSCA, JTJ, KJT, QZAC, USNM]; Santa Clara: [USNM]; SU, Cascales: [FSCA]; Lago Agrío: [SWD, USNM]; Putumayo: [USNM]; Shushufindi: [DRP, FSCA, UTIC, SWD, SWD]; ZC, Zamora: [UTIC]. Uracis imbuta (Burmeister, 1839). Publ. Recs. ECU: * Ris (1911), Campos (1922), Racenis (1954), (H 06), (P 11), (H 14); OR: von Ellenrieder & Garrison (2011); ZC: Tennessen & Johnson (2008). Also ARG, BOL, BRA, CEN, COL, GUF, GUY, PER, SUR, TTO, VEN: (H 06), (P 11). Addl. Data. ES, San Lorenzo: [ANSP, FSCA, JJD, KJT, QZAC]; MS, Santiago: [SWD]; NA, Carlos Julio Arsemena Tola: [FSCA, USNM]; El Chaco: [SWD]; Tena: [DRP, FSCA, UTIC, KJT, SWD, USNM]; OR, Francisco de Orellana: [FSCA, KJT, LSAM, NVE, QZAC, SWD, TWD, USFQ, USNM]; Loreto: [FSCA, KJT]; SU, Cascales: [UTIC, KJT, SWD, USNM]; Lago Agrío: [FSCA, KJT, QZAC, SWD, USNM]; Putumayo: [USNM]; Shushufindi: [DRP, FSCA, KJT, SWD, USNM]; ZC, Zamora: [KJT]. * Uracis infumata (Rambur, 1842). Publ. Recs. ECU: (H 14): BOL, BRA, GUF, GUY, PER, VEN: (H 06), (P 11); plus COL: (P & P); and SUR (P 16). Addl. Data. OR, Francisco de Orellana: 1 ♀, forest pool, Río Savaleto, P. N. Y., 17 xi 1997, leg. & det. KJT [KJT]; adult, Shiripuno Lodge, Río Shiripuno, 26 km SE of bridge on the Auca Road from Coca. [- 1.1047, - 76.4827, 273 m), no date provided, data provided by WAH & DLW; SU: Cuyabeno Reserve, 16.7 km E of Cuyabeno village, 3 km SE of Cuyabeno Lodge, no date provided, data provided by WAH & DLW. Uracis ovipositrix Calvert, 1909. Publ. Recs. ECU: Paulson (1983), Tsuda (1986, 1991), (H 06), (P 11), (H 14). Also BRA, GUF, GUY, PER, SUR, VEN: (H 06), (P 11). Addl. Data. OR, Francisco de Orellana: 1 ♂, 1 ♀, Yasuni Scientific Station, Río Tiputini, 39 km SE of Pom- peya, 61 km ESE of Coca, muddy trail in forest (- 0.674, - 76.397, 205 m), 18 i 2012. leg. DLW, WAH, Fred Morrison, Tim Kell, Vicki Liu, det. WAH [image vetted by DRP] [WAH]. Uracis siemensi Kirby, 1897. Publ. Recs. ECU: * Paulson (1983), (H 06), (P 11), (H 14). Also BRA, GUF, SUR, PER, VEN: (H 06), (P 11); plus COL: (P & P). Addl. Data. NA, El Chaco: 1 ♂, El Chaco, 52 km NE on road to Lago Agrio, marsh, 18 viii 1980, leg. & det. SWD [FSCA]; OR, Francisco de Orellana: 1 ♂, [La Nueva Esperanza], [across Río Napo from] near Primavera “ Limoncocha ” Río Napo & lake Taracoa, forest trail to lake (- 0.45, - 76.77, 300 m), 27 vii 1977, leg. & det. DRP [DRP]; 1 ♂, same data but 4 xi 1980, leg. & det. MJW [FSCA].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B5DEFF19FF23F107272BFB8B.taxon	discussion	Identification of Zenithoptera species depends on wing coloration and structure of the vesica spermalis (Jurzitza 1982), (Pujol-Luz & Rodrigues da Fonseca 1997). Adults inhabit small to medium-sized vegetated ponds and pools of small, slow streams; the surrounding habitat is usually open to somewhat shaded by trees. The species in Ecuador have drab, mottled bodies, but the wings are spectacularly colored, brilliant iridescent blue on top and black underneath with white bands and spots (Figs. 116 a & b). The genus is peculiar in that adults perch with their wings either held slightly downward or together upright over the abdomen, somewhat similar to Zygoptera that fold their wings. However, at times one pair of wings will be held upright, while the other pair is held downward. This behavior of holding the wings upright is unique in the New World Anisoptera fauna (Paulson 2004 b). Two species are known from Ecuador. Zenithoptera fasciata (Linnaeus, 1758). Publ. Recs. ECU: (H 06), (P 11), (H 14); NA: * Tennessen (1996 b — as Z. americana); OR: Tennessen (1998), von Ellenrieder & Garrison (2011). Also BRA, CEN, COL, GUF, GUY, PER, SUR, VEN, TTO: (H 06), (P 11). Addl. Data. NA, Tena: 1 ♂, Puerto Napo, 5.3 km E of Hwy. 45 on J. S. B. S. Road, Río Sinde (- 1.0500, - 77.7481, 450 m), 22 vii 1996, leg. & det. WFM [LSAM]; 5 ♂♂, 1 ♀, J. S. B. S., Poza 3 (- 1.0655, - 77.6127, 437 m), 10 xi 2014, leg. & det. ACRP [ACRP]; OR, Francisco de Orellana: 1 ♀, Reserva Etnica Waorani, 1 km S of Onkone Gare Camp (- 0.63, - 76.60), 20 i 1994, leg. T. Erwin, det.? [USNM]; 2 ♂♂, swamp, P. N. Y. (- 0.6508, - 76.3558, 200 m), 15 xi 1997, leg. & det. KJT [KJT]; 1 ♂, same data but 16 xi 1997, leg. & det. TWD [SWD]; 2 ♂♂, pond, P. N. Y., km marker 29.5, 17 xi 1997, leg. & det. KJT [KJT]; 1 ♂, [Pompeya Sur], S on [Pompeya Sur-Iro road] at km marker 31, borrow pit near Río Sabaleto (- 0.6569, - 76.4528, 200 m), 17 xi 1997, leg. & det. KJT [KJT]; 1 ♂, Puerto Francisco de Orellana, 8.2 km W on road to Loreto from Coca-Sachas Road, pond (- 0.4578, - 77.0381, 288 m), 18 xi 1997, leg. & det. WFM [FSCA]; 3 ♂♂, Puerto Francisco de Orellana, 11.8 km W on Loreto Road, large pond (- 0.4742, - 77.0583, 285 m), 18 xi 1997, leg. & det. WFM [FSCA, LSAM, QZAC]; 2 ♂♂, same data but leg. & det. TWD [SWD]; 1 ♂, Puerto Francisco de Orellana, 12.3 km W on road to Loreto from Coca-Sachas Road, pond (- 0.4750, - 77.0747, 290 m), 13 vi 1995, leg. & det. WFM [FSCA]; 1 ♂, same data but leg. & det. SWD [KJT]. * Zenithoptera lanei Santos, 1941. Publ. Recs. ARG, BRA, PRY, PER, VEN: (H 06), (P 11); plus COL: (P 16). Addl. Data. NA, Tena: 1 ♂, J. S. B. S., near Tena, pond (450 m), 18 vi 1994, leg. A. Jaramillo, det. DRP [DRP]; OR, Francisco de Orellana: 1 ♀, T. B. S., fogging, TLE Plot T- 1, Sta. 8. FCS Work no. 095 (- 0.6319, - 76,1442, 220 – 250 m), 2 x 2000, leg. T Erwin, det. WFM [USNM]; 3 ♂♂, ponds near Comunidad El Rocio, (- 0.5054, - 77.1569, 290 m), 27 x 2013, leg. & det. CDK [FSCA]; same data but 2 ♂♂, leg. & det. KJT [FSCA]; La Joya de Los Sachas: 1 ♂, La Joya de Los Sachas, E, ponds, (- 0.2202, - 76.6987, 262 m), 29 x 2013, leg. & det. CDK [FSCA]; SU, Shushufindi, 1 ♂, stream about 3 km NW of Shushufindi, 2 + km N off Hwy. 26 at km marker 17 (- 0.1664, - 76.6917, 290 m), 13 v 2009, leg. & det. KJT [FSCA]; 1 ♂, same data but 14 v 2009 [FSCA]; 1 ♀, same data but 14 x 2009 [KJT]; 1 ♂, same data but 13 x 2009, leg. & det. JTJ [JTJ]; 1 ♂, Shushufindi, SW, pond (- 0.2201, - 76.6989), 29 x 2013, leg. & det. KJT [FSCA].	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B5DEFF19FF23F107272BFB8B.taxon	description		en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B5F4FF31FF23F5DB266DFB4B.taxon	description	Lestes henshawi Calvert, 1907. Publ. Recs. ECU: Soukop (1954), Paulson (1983), Tsuda (1986); PA: Kennedy (1942 b — ♀, described). These are all Lestes apollinaris Navás, 1934 according to Donnelly (1996) since Lestes henshawi is known only from Costa Rica. Lestes minutus Selys, 1862. Reported by Campos (1922) from “ Posorja ”. Unlikely since it is an Atlantic coast species listed from BRA, COL, GUF VEN, TTO: (H 08), (P 11). Lestes striatus Selys, 1862. Synonym of Lestes forficula Rambur, 1842.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B5FBFF3FFF23F42C212CFDBE.taxon	description	Cora cyane Selys, 1853. Synonym Cora incana Hagen in Selys, 1869. Publ. Recs. ECU: (H 08). Also VEN: (H 08), (P 11). We are not sure of the source of information that (H 08) used for the Ecuador record. Since there are no modern records in Ecuador, it is not included in our list until a voucher specimen or verifiable record is obtained. Cora mirabilis (McLachlan, 1878). See Euthore mirabilis McLachlan, 1878. Cora modesta Selys, 1869. Listed from Ecuador in Campos (1922), “ pers. comm. Dr. Martin ”, Bick & Bick (1990 a) and “? ” Tsuda (1991). Selys (1873 a) listed it in the generic summary from “ Quito ” but in the discussion he listed it only from Bogotá, Colombia. (P 16) listed it only from Colombia. We have not been able to resolve this record; therefore it is not included in our list until a verifiable record from Ecuador is obtained. Euthore fasciata inlactea Calvert, 1909. Synonym of Euthore fasciata Hagen in Selys, 1853. Euthore fasciata form plagiata Selys, 1873. Synonym of Euthore fasciata Hagen in Selys, 1853. Euthore fasciata form sulfurata De Marmels, 1982. Synonym of Euthore fasciata Hagen in Selys, 1853. Euthore hyalina (Selys, 1853), as “ thore hyalina ” from “ Bahia ” which was presumed to be Brazil, but Bridges (1993) suggested it “ might be Bahia de Carraquez, Ecuador. ” Lectotype by Montgomery (1967) [IRSN]. Also BRA, COL, PER: (P 11). We think that “ Bahia ” was actually in Brazil, since the Ecuadorian location proposed by Montgomery (1967) is on the Pacific coast totally disjunct from the other recorded localities of this species. It is not included in our list until a verifiable record is obtained. Euthore plagiata Selys, 1873. Synonym of Euthore fasciata Hagen in Selys, 1853. Josocora jocosa (McLachlan, 1881). See Cora jocosa McLachlan, 1881. Josocora klenei (Karsch, 1891). See Cora klenei Karsch, 1891. Miocora subapicalis Kennedy, 1940. Possible synonym of Miocora peraltica Calvert, 1917. Polythore albovittata (Selys, 1873). Synonym of Polythore vittata (Selys, 1869). Polythore beata McLachlan, 1869 as Thore beata. Campos (1922) listed it from Ecuador: “ pers. comm. Dr. Martin. ” Also Schmidt (1942). Bick & Bick (1986) stated that all west slope records are doubtful. Lencioni (2005), (H 08), (P 11), and Rojas-Riano (2011 a) did not list it but did report it from Brazil, Columbia and Peru. Type locality is “ Pebas, upper amazon ” which is in Peru (McLachlan 1869). It is not included in our list. If it occurs in Ecuador, it would be in the eastern lowlands. Polythore derivata derivata Fraser, 1946. Synonym of Polythore derivata (McLachlan, 1881). Polythore derivata race adjuncta Fraser, 1946. Synonym of Polythore derivata (McLachlan, 1881). Polythore derivata race ambigua Fraser, 1946. Synonym of Polythore derivata (McLachlan, 1881). Polythore derivata prisinata Fraser, 1946. Synonym of Polythore derivata (McLachlan, 1881). Polythore derivata terminata Fraser, 1946. Synonym of Polythore terminata Fraser, 1946. Polythore picta (Rambur, 1842). Publ. Recs. ECU: Hagen (1875), Montgomery (1967), Geijskes (1971), Paulson (1983), Tsuda, (1986, 1991), Steinmann (1997 a). IM: McLachlan (1878). Neither Bick & Bick (1985), (L 05), (H 08) nor (P 11) list it from Ecuador. (L 05) listed it from Brazil, Peru, Guyana & and French Guyana. Due to the confusion with procera and vittata, we do not think that P. picta occurs in Ecuador. Polythore picta race originata Fraser, 1946. Synonym of Polythore derivata (McLachlan, 1881). Sapho pulchella Kirby, 1889. Synonym of Polythore concinna (McLachlan, 1881). Thore acostai Navás, 1924. Synonym of Polythore vittata (Selys, 1869). Thore aequatorialis Selys, 1873. Synonym of Polythore procera (Selys, 1869). Thore albovittata (Selys, 1873). Synonym of Polythore vittata (Selys, 1869). Thore aurora Selys, 1879. See Polythore aurora (Selys, 1879). Thore beata McLachlan, 1869. See Polythore beata (McLachlan, 1869). Thore concinna McLachlan, 1881. See Polythore concinna (McLachlan, 1881). Thore derivata McLachlan, 1881. See Polythore derivata (McLachlan, 1881). Thore fasciata (Hagen in Selys, 1853). See Euthore fasciata Hagen in Selys 1853. Thore gigantea Selys, 1853. See Polythore gigantea Selys (1853). Thore gigantea race procera Selys, 1869. Synonym of Polythore procera (Selys, 1869). Thore montana Förster, 1914. Synonym of Polythore ornata (Selys, 1879). Thore mutata McLachlan, 1881. See Polythore mutata (McLachlan, 1881) Thore ornata Selys, 1879. See Polythore ornata (Selys, 1879). Thore pozuzina Förster, 1914. Synonym of Polythore ornata (Selys, 1879). Thore picta var. aequatorialis Selys, 1873. Synonym of Polythore procera (Selys, 1869). Thore picta race vitatta Selys, 1869. Synonym of Polythore vittata (Selys, 1869). Thore saundersii Selys, 1873. Synonym of Polythore picta (Rambur, 1842). Reported under this name from “ Baños ” Ecuador by Selys (1873), Hagen (1875) and by Campos (1922), but these were probably P. procera or P. neopicta. Thore tincta (Navás, 1924). Synonym of Polythore vittata (Selys, 1869). Thore victoria McLachlan, 1869. See Polythore victoria (McLachlan, 1869). Thore vittata race ” aequatorialis Selys, 1873. Synonym of Polythore vittata (Selys, 1869).	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B5F9FF3CFF23F031278EF851.taxon	description	Cordulia setifera (Hagen in Selys, 1871). See Neocordulia setifera (Hagen in Selys, 1871) in this appendix. Gomphomacromia batesi (Selys, 1871). See Neocordulia batesi (Selys, 1871). Gomphomacromia setifera (Hagen in Selys, 1871). See Neocordulia setifera (Hagen in Selys, 1871) in this appendix. Lauromacromia dubitalis (Fraser, 1939). Gomphomacromia dubitalis Fraser, 1939. Publ. Recs. ECU: (Tsuda 1986, 1991), (H 06). Also GUF, VEN: (H 06), (P 11). Garrison et al. (2006) indicates an eastern South America range for the genus; since there are no collection data or specific published records for Ecuador, it is not included on the list. Neocordulia longipollex Calvert, 1909. Synonym of Neocordulia bates i (Selys, 1871). Listed by Paulson (1983) and Tsuda (1986). Garrison et al. (2006) considered it a subspecies of N. batesi (Selys, 1871). Neocordulia setifera (Hagen in Selys, 1871). Publ. Recs. ECU: Martin (1906, 1914), Paulson (1983), Tsuda (1986, 1991), Steinmann (1997 b), (P 11). Also BRA: (H 06), (P 11); plus ARG: (P 11). May (1991) states that setifera is confined to southern Brazil and Paraguay and that there are no records or specimens from northern South America. Martin (1906, 1914) listed it from “ Selys collection, Amazonas, Ecuador. ” This was probably a mistake. Other published records simply echoed the Martin records. It is not included on this list.	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
D076D419B5F8FF27FF23F68827C2FBDF.taxon	description	Anatya anomala Kirby, 1889. Synonym of Anatya guttata (Erichson, 1848). Anatya normalis Calvert, 1899. Synonym of Anatya guttata (Erichson, 1848). Anatya theresiae Selys in Therese, 1900. Synonym of Anatya guttata (Erichson, 1848). Belonia funerea (Hagen, 1861). See Erythrodiplax funerea (Hagen, 1861). Belonia longipennis Kirby, 1889. Synonym of Libellula herculea Karsch, 1889. Brachymesia australis Kirby, 1889. Synonym of Brachymesia furcata (Hagen, 1861). Brechmorhoga inequiunguis (Calvert, 1895). See Macrothemis inequiunguis Calvert, 1895. Brechmorhoga pertinax eurysema Ris, 1913. Synonym of Brechmorhoga pertinax (Hagen, 1861). Brechmorhoga pertinax pertinax (Hagen, 1861). Synonym of Brechmorhoga pertinax (Hagen, 1861). Brechmorhoga pertinax peruviana Ris, 1913. Synonym of Brechmorhoga pertinax (Hagen, 1861). Brechmorhoga rapax crocosema Ris, 1913. Synonym of Brechmorhoga rapax Calvert, 1898. Brechmorhoga rapax rapax (Calvert, 1898). Synonym of Brechmorhoga rapax Calvert, 1898. Cannacria batesii Kirby, 1889. Synonym of Brachymesia herbida (Gundlach, 1889). Cannacria fumipennis Currie, 1901. Synonym of Brachymesia herbida (Gundlach, 1889). Cannacria smithii Kirby, 1894. Synonym of Brachymesia furcata (Hagen, 1861). Cannaphila insularis Kirby, 1889. Publ. Recs. ECU: (Tsuda, 1991). Unlikely to occur in Ecuador, probably based on a misidentification of C. vibex (Hagen, 1861). Dasythemis liriopa Karsch, 1889. Synonym of Dasythemis venosa (Burmeister, 1839). Publ. Recs. Campos (1922 — “ pers comm. Dr. Ris ”). This specimen is not discussed in Ris (1910) and it is reported only from southern South America by Paulson (2011). Diastatops fuliginea Rambur, 1842. Synonym of Diastatops pullata (Burmeister, 1839). Diplacodes abbreviata (Rambur, 1842). See Oligoclada abbreviata (Rambur, 1842). Diplacodes paraguayensis Förster, 1904. See Erythrodiplax paraguayensis (Förster, 1904). Diplacodes sp. Publ. Recs. GY: (Campos 1922). We are not sure what this species was since we could not locate the specimen; it was probably a species of Erythrodiplax. Diplax abjecta Hagen, 1861. See Erythrodiplax abjecta (Rambur, 1842). Diplax chloropleura (Brauer, 1865). See Erythrodiplax connata (Burmeister, 1839) in this appendix. Diplax credula Hagen, 1861. See Erythemis credula (Hagen, 1861). Diplax fraterna Hagen, 1873. See Erythrodiplax connata (Burmeister, 1839) in this appendix. Diplax fusca (Rambur, 1842). See Erythrodiplax fusca (Rambur, 1842). Diplax fuscofasciata Blanchard, 1845. Synonym of Erythrodiplax umbrata (Linnaeus, 1758). Diplax gilva Selys, 1884. See Sympetrum gilvum (Selys, 1884). Diplax illotum var. gilvum Selys, 1884. Synonym of Sympetrum gilvum (Selys, 1884). Diplax imbuta Burmeister, 1839. See Uracis imbuta (Burmeister, 1839). Diplax orchracea Hagen, 1861. Synonym of Erythrodiplax fervida (Erichson, 1848). Diplax pulla (Burmeister, 1839). Synonym of Erythrodiplax unimaculata (De Geer, 1773). Diplax unimaculata (De Geer, 1773). See Erythrodiplax unimaculata (De Geer, 1773). Dythemis aequalis Hagen, 1861. See Micrathyria aequalis (Hagen, 1861). Dythemis apicalis Hagen, 1861 (nomen nudum). Synonym of Nephepeltia phryne (Perty, 1834). Dythemis atra Martin, 1897. See Micrathyria atra (Martin, 1897). Dythemis broadwayi Kirby, 1894. Synonym of Dythemis sterilis (Hagen, 1861). Dythemis cannacrioides Calvert, 1906. See Elasmothemis cannacrioides (Calvert, 1906). Dythemis dicrota Hagen, 1861. Synonym of Micrathyria didyma (Selys in Sagra, 1857). Dythemis didyma (Selys in Sagra, 1857). See Micrathyria didyma (Selys in Sagra, 1857). Dythemis guttata (Erichson, 1848). See Anatya guttata (Erichson, 1848). Dythemis hemichlora (Burmeister, 1839). See Macrothemis hemichlora (Burmeister, 1839). Dythemis multipunctata Kirby, 1894. Meurgey (2011) found the true D. multipunctata is limited to the Lesser Antilles and stated that Central and South American records for D. multipunctata are really Dythemis nigra. Dythemis musiva (Hagen, 1861) See Macrothemis musiva Calvert, 1898. Dythemis multipunctata reducta De Marmels, 1989. Synonym of Dythemis nigra Martin, 1897. Dythemis pertinax, Hagen, 1961. See Brechmorhoga pertinax (Hagen, 1961). Dythemis sallaei Selys, 1868. Synonym of Brechmorhoga pertinax (Hagen, 1861). Dythemis tabida Hagen, 1861 (nomen nudum). Synonym of Dythemis nigra Martin, 1897. Dythemis velox Hagen, 1861. Reported by Tsuda (1986, 1991). The subspecies of D. velox according to Ris (1913): D. multipunctata and D. sterilis were raised to specific rank and then Meurgey (2011) showed that D. multipunctata references for mainland South America should be D. nigra (in the FSCA, Campos’ specimens of D. sterilis are labeled D. velox). Dythemis velox var sterilis (Hagen, 1861). Synonym of Dythemis sterilis (Hagen, 1861). Elga santosi Machado, 1954. Synonym of Elga leptostyla Ris, 1911. Ephidatia longipes (Hagen, 1861). See Idiataphe longipes (Hagen, 1861). Erythemis furcata Hagen, 1861. See Brachymesia furcata (Hagen, 1861). Erythemis longipes Hagen, 1861. See Idiataphe longipes (Hagen, 1861). Erythemis verbenata (Hagen, 1861). Synonym of Erythemis plebeja (Burmeister, 1839). Erythrodiplax amazonica melanica Borror, 1942. See Erythrodiplax melanica Borror, 1942. Erythrodiplax atroterminata Ris, 1911. A record at QZAC, NA, Archidona: 1 ♂, Archidona (705 m), 29 iii 1991, leg. & det.? [QZAC], could not be verified for this southern South American species; therefore it is not included on the list. Erythrodiplax attenuata forma hyalina Sjöstedt, 1918. Synonym of Erythrodiplax attenuata (Kirby, 1889). Erythrodiplax basalis basalis (Kirby, 1897). Synonym of Erythrodiplax basalis (Kirby, 1897). Erythrodiplax basifusca (Calvert, 1895). Publ. Recs. GY: Campos (1922). Paulson (2003) provided evidence that the true E. basifusca is found only in Mexico and the southwestern United States. The Ecuador records were for some other species in the connata - fusca group. Erythrodiplax chloropleaura (Brauer, 1865). See Erythrodiplax connata (Burmeister, 1839) in this appendix. Erythrodiplax communis (Rambur, 1842). See Erythrodiplax connata (Burmeister, 1839) in this appendix. Erythrodiplax connata (Burmeister, 1839). Publ. Recs. ECU (Racenis 1954; Paulson 1983; Tsuda 1986, 1991). CR: (Campos 1922), EO: (Campos 1927), GY: (Calvert 1906 “ form betweem d / e ” and form e’; (Campos 1922), (Borror, 1942 “ connata d / e by Calvert ”), LR: (Calvert 1906 “ form between d / e ”; (Campos 1922). The true E. connata is restricted to the west slope of the Andes in Chile (Paulson 2003); all of these historical records are presumed to be either E. abjecta or E. fusca. Erythrodiplax connata abjecta (Rambur, 1842). See Erythrodiplax abjecta (Rambur, 1842). Erythrodiplax connata connata (Burmeister, 1839) See Erythrodiplax connata (Burmeister, 1839) in this appendix. Erythrodiplax connata fraterna (Hagen, 1873). See Erythrodiplax connata (Burmeister, 1839) in this appendix. Erythrodiplax connata var “ 100 c ”. Publ Recs. IM: Campos (1922 “ forma incerta por Dr. Ris ”). We suspect that this was either E. abjecta or E. fusca. Erythrodiplax connata var “ 100 d ”. Publ. Recs. ES: Campos (1922 “ forma dudosa por Dr. Ris ”). We suspect that this was either E. abjecta or E. fusca. Erythrodiplax connata form d. Calvert, 1906. Probable reference to Erythrodiplax fusca (Rambur, 1842). Erythrodiplax connata fusca (Rambur, 1842). See Erythrodiplax fusca (Rambur, 1842). Erythrodiplax erichsoni (Kirby, 1894). Synonym of Erythrodiplax kimminsi Borror, 1942. Erythrodiplax lenti Ris, 1919 (part). Synonym of Erythrodiplax amazonica Sjöstedt, 1918. Erythrodiplax leontina (Brauer, 1865). See Erythrodiplax connata (Burmeister, 1839) in this appendix. Erythrodiplax media Borror, 1942. Publ. Recs. ECU: Tsuda (1986, 1991). We have no idea what the source of these records are. Paulson (2003) stated that the true E. media is restricted to southern Brazil, Paraguay and eastern Bolivia. Erythrodiplax ochracea (Burmeister, 1839). Publ. Recs. - CR: Campos (1922); CB: Campos (1922); GY: Campos (1922). Synonym of Erythrodiplax fervida (Borror, 1942). Erythrodiplax ponderosa Karsch, 1891. Synonym of Erythrodiplax abjecta (Rambur, 1842). Fylgia amazonica lychnitina De Marmels, 1989. Synonym of Fylgia amazonica Kirby, 1889. Leptetrum merida (Selys, 1868). Synonym of Cannaphila vibex (Hagen, 1861). Leptetrum vibex (Hagen, 1861). See Cannaphila vibex (Hagen, 1861). Lepthemis cultriformis Hagen, 1861. See Orthemis cultriformis (Hagen, 1861). Lepthemis haematogastra (Burmeister, 1839). See Erythemis haematogastra (Burmeister, 1839). Lepthemis verbenata Hagen, 1861. Synonym of Erythemis plebeja (Burmeister, 1839). Libellula abbreviata Rambur, 1842. See Oligoclada abbreviata (Rambur, 1842). Libellula abdominalis Rambur, 1842. See Tramea abdominalis (Rambur, 1842). Libellula abjecta Rambur, 1842. See Erythrodiplax abjecta (Rambur, 1842). Libellula americana Linnaeus, 1758. Synonym of Zenithoptera fasciata (Linnaeus, 1758). Libellula attala Selys in Sagra, 1857. See Erythemis attala (Selys, 1857). Libellula annulata Rambur, 1842 (nec Palisot de Beauvois, 1805). Synonym of Erythemis attala (Selys, 1857). Libellula analis Burmeister, 1839. Synonym of Pantala flavescens (Fabricius, 1798). Libellula annulosa Selys in Sagra, 1857. Synonym of Erythemis attala (Selys, 1857). Libellula basalis Hagen, 1875. Synonym of Tramea cophysa (Hagen, 1861). Libellula basalis Burmeister, 1839 (nec Stephens, 1835). Synonym of Tramea abdominalis (Rambur, 1842). Libellula bicolor Erichson in Schomburgk, 1848. Synonym of Erythemis peruviana (Rambur, 1842). Libellula binotata Rambur, 1842. See Tramea binotata (Rambur, 1842). Libellula cardinalis Erichson in Schomburgk, 1848. See Rhodopygia cardinalis (Erichson in Schomburgk, 1848). Libellula castanea Burmeister, 1839. See Erythrodiplax castanea (Burmeister, 1839). Libellula chloropleura Brauer, 1865. See Erythrodiplax connata (Burmeister, 1839) in this appendix. Libellula communis Rambur, 1842. See Erythrodiplax connata (Burmeister, 1839) in this appendix. Libellua connata Burmeister, 1839. See Erythrodiplax connata (Burmeister, 1839) in this appendix. Libellula cophysa Hagen, 1861. See Tramea cophysa (Hagen, 1861). Libellula discolor Selys, 1857. See Orthemis discolor (Selys, 1857). Libellula distinguenda Rambur, 1942 (pars). Synonym of Erythrodiplax fervida (Erichson, 1848). Libellula didyma Selys in Sagra, 1857. See Micrathyria didyma (Selys in Sagra, 1857). Libellula domitia Drury, 1773. See Perithemis domitia (Drury, 1773). Libellula erratica Erichson in Schomburgk, 1848. Synonym of Erythrodiplax basalis (Kirby, 1897). Libellula fallax Burmeister, 1839. Synonym of Erythrodiplax umbrata (Linnaeus, 1758). Libellula fasciata Linnaeus, 1758. See Zenithoptera fasciata (Linnaeus, 1758). Libellula fastigiata Burmeister, 1839. See Uracis fastigiata (Burmeister, 1839). Libellula fervida Erichson, 1848. See Erythrodiplax fervida (Erichson in Schomburgk, 1848). Libellula flavescens Fabricius, 1798. See Pantala flavescens (Fabricius, 1798). Libellula flavicans Rambur, 1842. Synonym of Erythrodiplax umbrata (Linnaeus, 1758). Libellula funerea Hagen, 1861. See Erythrodiplax funerea (Hagen, 1861). Libellula fusca Rambur, 1842. See Erythrodiplax fusca (Rambur, 1842). Libellula fuscofasciata (Blanchard, 1845). Synonym of Erythrodiplax umbrata (Linnaeus, 1758). Libellula guttata Erichson, 1848. See Anatya guttata (Erichson, 1848). Libellula haematogastra Burmeister, 1839. See Erythemis haematogastra (Burmeister, 1839). Libellula hemichlora Burmeister, 1839. See Macrothemis hemichlora (Burmeister, 1839). Libellula herbida Gundlach, 1889. See Brachymesia herbida (Gundlach, 1889). Libellula hymenaea Say, 1840. See Pantala hymenaea (Say, 1840). Libellula imbuta Burmeister, 1839. See Uracis imbuta (Burmeister, 1839). Libellula incompta Rambur, 1842. Synonym of Erythrodiplax fusca (Rambur, 1842). Libellula infumata Rambur, 1842. See Uracis infumata (Rambur, 1842). Libellula justinea Selys in Sagra, 1957. Synonym of Erythrodiplax fervida (Erichson, 1848). Libellula lais Perty, 1834. See Perithemis lais (Perty, 1934). Libellula leontina Brauer, 1865. See Erythrodiplax connata (Burmeister, 1839) in this appendix. Libellula marcella Selys in Sagra, 1857. See Miathyria marcella (Selys in Sagra, 1857). Libellula merida Selys, 1868. Synonym of Cannaphila vibex (Hagen, 1861). Libellula metella Selys in Sagra, 1857. Synonym of Perithemis domitia (Drury, 1773). Libellula mithra Selys in Sagra, 1857. Synonym of Erythemis attala (Selys, 1857). Libellula nubecula Rambur, 1842. See Brechmorhoga nubecula (Rambur, 1842). Libellula ochracea (Burmeister, 1839). Synonym of Erythrodiplax fervida (Borror, 1942). Libellula plebeja Burmeister, 1839. See Erythemis plebeja (Burmeister, 1839). Libellula pulla Burmeister, 1839. Synonym of Erythrodiplax unimaculata (De Geer, 1773). Libellula pullata Burmeister, 1839. See Diastatops pullata (Burmeister, 1839). Libellula phryne Perty, 1834. See Nephepeltia phryne (Perty, 1834). Libellula rubriventris Blanchard (1845). Synonym of Erythemis peruviana (Rambur, 1842). Libellula ruralis Burmeister, 1839. Synonym of Erythrodiplax umbrata (Linnaeus, 1758). Libellula sallaei (Selys, 1868). Synonym of Brechmorhoga pertinax (Hagen, 1961). Libellula simplex Rambur, 1842. See Miathyria simplex (Rambur, 1842). Libellula subfasciata Burmeister, 1839. Synonym of Erythrodiplax umbrata (Linnaeus, 1758). Libellula terminalis Burmeister, 1839. Synonym of Pantala flavescens (Fabricius, 1798). Libellula tessellata Burmeister, 1839, See Macrothemis tessellata (Burmeister, 1839). Libellula tripartita Burmeister, 1839. Synonym of Erythrodiplax umbrata (Linneaus, 1758). Libellula umbrata (Linneaus, 1758). See Erythrodiplax umbrata (Linneaus, 1758). Libellula unimaculata De Geer, 1773. See Erythrodiplax unimaculata (De Geer, 1773). Libellula unifasciata De Geer, 1773. Synonym of Erythrodiplax umbrata (Linnaeus, 1758). Libellula vibex Hagen, 1861. See Cannaphila vibex (Hagen, 1861). Libellula violacea De Geer, 1773. Synonym of Zenithoptera fasciata (Linnaeus, 1758). Libellula viridula Beauvois, 1807. Synonym of Pantala flavescens (Fabricius, 1798). Macrothemis catharina Karsch, 1890. Synonym of Brechmorhoga nubecula (Rambur, 1842). Macrothemis cydippe Calvert, 1898. Synonym of Macrothemis hemichlora (Burmeister, 1839). Macrothemis leucozona Ris, 1913. Synonym of Macrothemis imitans Karsch, 1890. Macrothemis newtoni Costa, 1990. Publ. Recs. BRA: (H 06), (P 11). We have not seen any specimens of this species and do not know the source of the records. It is possible that these records were obtained from an article in the Dragonfly Society of the Americas newsletter ARGIA mentioning a specimen tentatively identified as this species then later determined to be possibly M. idalia. Macrothemis pumila Karsch, 1890. See Gynothemis pumila (Karsch, 1890). Macrothemis pumila var. axillata Navás, 1924. Synonym of Gynothemis pumila (Karsch, 1890). Macrothemis tessellata inequiunguis Calvert, 1895. Synonym of Macrothemis inequiunguis Calvert, 1895. Macrothemis tessellata tessellata (Burmeister, 1939). Synonym of Macrothemis tessellata (Burmeister, 1839). Macrothemis tessellata (Burmeister, 1939) was recognized as a separate species distinct from M. inequiunguis by both Paulson (2011) and Garrison et al. (2006), but recently Rosser Garrison (pers. comm., 2019) thinks they might represent one species. Macrothemis uniseries Calvert, 1909. Synonym of Macrothemis musiva Calvert, 1898. Macrothemis vulgipes Calvert, 1898. Synonym of Macrothemis inequiunguis Calvert, 1895. Mesothemis abbreviata (Rambur, 1842). See Oligoclada abbreviata (Rambur, 1842). Mesothemis annulata (Martin, 1896). Synonym of Erythemis attala (Selys, 1857). Mesothemis attala (Hagen, 1861). See Erythemis attala (Selys, 1857). Mesothemis chloropleaura (Brauer, 1865). See Erythrodiplax connata (Burmeister, 1839) in this appendix. Mesothemis connata (Burmeister, 1839). See Erythrodiplax connata (Burmeister, 1839) in this appendix. Mesothemis leontina (Brauer, 1865). See Erythrodiplax connata (Burmeister, 1839) in this appendix. Mesothemis mithra (Hagen, 1861). Synonym of Erythemis attala (Selys, 1857). Mesothemis mithroides Brauer, 1900. See Erythemis mithroides (Brauer, 1900). Mesothemis poeyi Scudder, 1866. Synonym of Micrathyria didyma (Selys in Sagra, 1857). Mesothemis verbenata (Hagen, 1861). Synonym of Erythemis plebeja (Burmeister, 1839). Miathyria pusilla Kirby, 1889 Synonym of Miathyria simplex (Rambur, 1842). Micrathyria attenuata (Kirby, 1889). See Erythrodiplax attenuata (Kirby, 1889). Micrathyria basalis Kirby, 1897. See Erythrodiplax basalis (Kirby, 1897). Micrathyria carlota Needham, 1942. Synonym of Micrathyria ocellata Martin, 1897. Micrathyria didyma didyma (Selys, 1861). Synonym of Micrathyria didyma Selys, 1861. Micrathyria didyma laevigata Calvert, 1909. Synonym of Micrathyria laevigata (Calvert, 1909). Micrathyria dythemoides Calvert, 1909. Publ. Recs. ECU: (H 06), (P 11), (H 14). Also PER, SUR, TTO: (H 06), (P 11). Paulson (pers. comm., 2011) reports that this was a mis-identification and therefore it is not included on the list. WAH (pers. comm., 2017) indicated that he copied it from Dennis Paulson’s list. Micrathyria hemimelaena Karsch, 1890. Synonym of Erythrodiplax unimaculata (De Geer, 1773). Micrathyria mengeri mengeri Ris, 1919. Synonym of Micrathyria mengeri Ris, 1919. Micrathyria ocellata ocellata Martin, 1897. Synonym of Micrathyria ocellata Martin, 1897. Micrathyria ocellata dentiens Calvert, 1909. Synonym of Micrathyria ocellata Martin, 1897. Micrathyria ocellata quicha Calvert, 1909. Synonym of Micrathyria ocellata Martin, 1897. Micrathyria pruinosa Kirby, 1894. Synonym of Micrathyria didyma (Selys in Sagra, 1857). Micrathyria septima Selys, 1900. Synonym of Micrathyria aequalis (Hagen, 1861). Neothemis flavifrons Karsch, 1889. See Nephepeltia flavifrons (Karsch, 1889). Nannophya semiaurea Hagen, 1861 (nomen nudum). Synonym of Nephepeltia phryne (Perty, 1834). Nannothemis inermis Hagen, 1867 (nomen nudum). Synonym of Nephepeltia flavifrons (Karsch, 1889). Nannothemis prodita Hagen, 1861 (nomen nudum). Synonym of Nephepeltia flavifrons (Karsch, 1889). Nannothemis semiaurea Karsch, 1889. Synonym of Fylgia amazonica Kirby, 1889. Neurothemis affinis Kirby, 1889. Synonym of Erythrodiplax funerea (Hagen, 1861). Nephepeltia phryne phryne (Perty, 1834). Synonym of Nephepeltia phryne (Perty, 1834). Nephepeltia phryne tupiensis Santos, 1950. Synonym of Nephepeltia phryne (Perty, 1834). Nothemis apollinaris Navás, 1915. Synonym of Brechmorhoga rapax Calvert, 1898. Nothifixis laxa Navás, 1916. Synonym of Miathyria simplex (Rambur, 1842). Oligoclada abbreviata abbreviata (Rambur, 1842). Synonym of Oligoclada abbreviata (Rambur, 1842). Oligoclada abbreviata limnophila Machado & Machado, 1993. Synonym of Oligoclada abbreviata (Rambur, 1842). Oligoclada raineyi Ris, 1911. Synonym of Oligoclada abbreviata (Rambur, 1842). Orthemis anthracina De Marmels, 1989. Publ. Recs. ECU: (H 14 — as O. cf. anthracina). Also PER, VEN: (H 06), (P 16); plus CO, SU, TTO: (P 16). von Ellenrieder (2009) did not list it from Ecuador, but its range might include eastern Ecuador. WAH (pers. comm 2016) “ When I first keyed this one out, I had only one specimen. Now that we have more, I am convinced these are all O. aciculata, and that we have not yet collected O. anthracina in Ecuador. ” Orthemis concolor Ris, 1919. Publ. Recs. ECU: (H 06). Also BRA, GUF, GUY, PER, SUR, TTO, VEN: (H 06), (P 16); plus PER (H 06). The source of this single published record is unknown. It is similar in some respects to O. cinnamomea and the distribution is eastern Atlantic coastal areas of South America (von Ellenrieder 2009). It is not included in our list. Orthemis ferruginea (Fabricius, 1775). Publ. Recs. ECU: Paulson (1983), Tsuda (1986, 1991). Prior to the 1990 ’ s all references to O. ferruginea in South America (except var sulphurata) should be O. discolor (Burmeister, 1839). Donnelly (1995), De Marmels (1988) and Paulson (1998) all showed that Costa Rica is the southern-most limit for O. ferruginea. Hoffmann (2010) however reported a collection of O. ferruginea in Peru in 2001 and then a breeding population discovered in 2006 on the Pacific slope at 1200 m. There is no mention who vetted these Peruvian records. Orthemis ferruginea var sulphurata (Hagen, 1868). See Orthemis sulphurata (Hagen, 1868). Orthemis flavopicta Kirby, 1889. Publ Recs. ES: Campos (1922 “ por Dr. Ris ” in the Selys collection). Also BRA, CEN, GUF, PER: (H 06), (P 11). This is a dubious record according to von Ellenrieder (2009); she listed it from eastern Brazil and from Central America. Orthemis sibylla Ris, 1919. Synonym of Orthemis ambirufa Calvert, 1909. Orthemis tambopatae von Ellenrieder, 2009. Publ. Recs. ECU: (H 14). W. A. Haber (pers. comm., 2016) wrote “ 1 ♂, Shiripuno Lodge, det by Dennis Paulson (from photo only). I do not have the specimen, so cannot verify it with Natalia’s [von Ellenrieder] key. Maybe best to leave it out until we get another record or I can key out this specimen. ” von Ellenrieder (2009, 2012) reports it only from southern Peru. Perithemis austeni Kirby, 1897. Synonym of Perithemis bella Kirby, 1889. Perithemis cloe Hagen, 1861 (nomen nudum). Synonym of Perithemis mooma Kirby, 1889. Perithemis domitia var. iris Hagen, 1861. Synonym of Perithemis domitia (Drury, 1773). Perithemis domitia tenera (Say, 1840). Synonym of P. domitia (Drury, 1773). Subspecies name used by Muttkowski (1910). No other reference could be found. Perithemis intensa Kirby, 1889. Listed from the Galapagos * Calvert (1906). Needham et al. (2014) list it from the southwestern United States and northern Mexico south to Jalisco. Unlikely to be found in Ecuador. Perithemis midasi ms. Gloyd manuscript name which was never published but there are specimens in various museums which may have labels with this name. Later described by Hoffman (1990) with the name Perithemis parzefalli. Perithemis mooma var. octoxantha Ris, 1910. Synonym of Perithemis mooma Kirby, 1889. Perithemis naias Ris, 1910. Synonym of Perithemis lais (Perty, 1934). Perithemis piperi Hoffmann, 1987. Synonym of Perithemis mooma Kirby, 1889. Perithemis pocahontas Kirby, 1889. Synonym of Perithemis domitia (Drury, 1773). Perithemis tenera tenera (Say, 1840). Listed by Steinmann (1997 b). This is an incorrect reference to P. domitia (Drury, 1883); P. tenera is a North American species. Pronomaja mimetica Förster, 1909. Synonym of Uracis ovipositrix Calvert, 1909. Rhodopygia cardinalis var. colorata Navás, 1923. Synonym of Rhodopygia cardinalis (Erichson, 1848). Sympetrum castaneum (Burmeister, 1839). See Erythrodiplax castanea (Burmeister, 1839). Sympetrum illotum (Hagen, 1861). See discussion under Sympetrum gilvum (Selys, 1884). Sympetrum illotum gilvum (Selys, 1884). See Sympetrum gilvum (Selys, 1884). Sympetrum tandicola Singh, 1955. Synonym of Pantala flavescens (Fabricius, 1798). Tauriphila nycteris Karsch, 1890. Synonym of Tauriphila argo (Hagen, 1869). Tramea argo Hagen, 1869. See Tauriphila argo (Hagen, 1869). Tramea australis Hagen, 1867. See Tauriphila australis (Hagen, 1867). Tramea basalis (Hagen, 1875). Synonym of Tramea cophysa (Hagen, 1861). Tramea brasiliana Brauer, 1867. Synonym of Tramea binotata (Rambur, 1842). Tramea cophysa darwini Kirby, 1889. See Tramea cophysa (Hagen, 1861). Tramea cophysa form “ a ” Ris, 1913. This is Tramea calverti Muttkowski, 1911. Tramea cophysa? (in part) Calvert, 1909. This is Tramea calverti Muttkowski, 1911. Tramea darwini Kirby, 1889. Synonym of Tramea cophysa (Hagen, 1861). Tramea huanacina Förster, 1909. Synonym of Pantala hymenea (Say, 1840). Tramea iphigenia (Kirby, 1890). Synonym of Tauriphila australis (Hagen, 1867). Tramea longicauda Brauer, 1867. Synonym of Tramea binotata (Rambur, 1842). Tramea marcella Hagen, 1867. See Miathyria marcella (Selys in Sagra, 1857). Tramea paulina Förster, 1910. Synonym of Tramea binotata (Rambur, 1842). Tramea simplex (Rambur, 1842). See Miathyria simplex (Rambur, 1842). Tramea subbinotata Brauer, 1867. Synonym of Tramea binotata (Rambur, 1842). Tramea walkeri Whitehouse, 1943. Synonym of Tramea binotata (Rambur, 1842). Trapezostigma binotata (Rambur, 1844). See Tramea binotata (Rambur, 1842). Trapezostigmas cophysa (Hagen, 1861). See Tramea cophysa (Hagen, 1861). Trithemis abbreviata (Rambur, 1842). See Oligoclada abbreviata (Rambur, 1842). Trithemis abjecta Kirby, 1890). See Erythrodiplax abjecta (Rambur, 1842). Trithemis attenuata Kirby, 1889. See Erythrodiplax attenuata (Kirby, 1889). Trithemis basifusca Calvert, 1895. See Erythrodiplax basifusca (Calvert, 1895) in this appendix. Trithemis communis (Rambur, 1842). See Erythrodiplax connata (Burmeister, 1839) in this appendix. Trithemis connata (Burmeister, 1839). See Erythrodiplax connata (Burmeister, 1839) in this appendix. Trithemis erichsoni Kirby, 1894 (pars). Synonym of Erythrodiplax kimminsi Borror, 1942. Trithemis fraterna (Hagen, 1873). See Erythrodiplax connata (Burmeister, 1839) in this appendix. Trithemis funerea (Hagen, 1861). See Erythrodiplax funerea (Hagen, 1861). Trithemis fusca (Rambur, 1842). See Erythrodiplax fusca (Rambur, 1842). Trithemis montezuma Calvert, 1899. Synonym of Erythrodiplax umbrata (Linnaeus, 1758). Trithemis ochracea Kirby, 1890. Synonym of Erythrodiplax fervida (Borror, 1942). Trithemis pulla (Burmeister, 1839). Synonym of Erythrodiplax unimaculata (De Geer, 1773). Trithemis tyleri Kirby, 1899. Synonym of Erythrodiplax funerea (Hagen, 1861). Trithemis umbrata (Linneaus, 1758). See Erythrodiplax umbrata (Linneaus, 1758). Trithemis unimaculata (De Geer, 1773). See Erythrodiplax unimaculata (De Geer, 1773). Uracis fastigiata aberration pura Förster, 1909. Synonym of Uracis fastigiata (Burmeister, 1839). Uracis fastigiata forma machadina Förster, 1910. Synonym of Uracis fastigiata (Burmeister, 1839). Uracis quadra Rambur, 1842. Synonym of Uracis imbuta (Burmeister, 1839). Uracis guttata (Erichson, 1848). See Anatya guttata (Erichson, 1848). Uracis ovata Calvert, 1909. Synonym of Uracis infumata (Rambur, 1842). Urothemis guttata (Erichson, 1848). See Anatya guttata (Erichson, 1848). Zenithoptera americana (Linnaeus, 1758). Synonym of Zenithoptera fasciata (Linnaeus, 1758). APPENDIX B. Localities often listed in literature: A partial list identifying province and / or Canton for each. (Source- Google maps and / or Maplandia) Abitagua: Pastaza, near Mera, 43 km E of Baños, 5 km W of Mera Agua Clara: Guayas, near Perlengue Aguamo: Pastaza, Pastaza Alluriquin: Santo Domingo de los Tsáchilas (formerly Pichincha) Angamarca: Cotopaxi, Pujili Canton Babahoyo: Los Ríos Balzapampa: Bolivar Province, San Miguel Canton Balzar: Guayas Barraganetal: Guayas Batan: Guayas Bobonaza: Pastaza Bombaini-yacu: — as “ Napo ” by Garrison (2006). “ trail between main road and Río Bombaini-Yacu, 25 Mar. 1941 ” (Wm. Clarke MacIntyre), {probably pastaza?} Bucay: Guayas Province, Naranjal Casiguana: just east of Guayaquil, Guayas Cerro Azul: just W of Guayaquil, Guayas Chambo: Chimborazo Chimbo: Guayas, Milagro Canton, Hacienda Venecia de Chimbo Chonta-Yacu: Napo Cotocollao: Pinchincha, Quito Canton Cotopino: Cotapino, Napo W of Loreto Cotundo: Napo, Archidona on Baesa – Tena Hwy Daule: Guayas Duran: Guayas, just east of Guayaquil Gauilanes: near Río Yana Yacu where it flows into the Río Anzu Gualaquiza: Morano-Santiago (formerly part of Azuay) Guayupe: Imbabura Hacienda Ila: Napo, on the Río Anzu Intag, Intaj, Intaq: Imbabura, Cotacachi Canton: “ Intag ” - [Apuela, NE on Apuela-Cuicocha road, along Río Intag and tributaries between 1800 - 2800 m, 0.358817, - 78.450904 at jct of Río Intag and tributary] El Morro: Guayas, W of Guayaquil El Salada: Carchi Jatun Yacu: Napo La Palmera: Tungurahua, - 1.25, - 78.12 on Río Pastaza, a hacienda Las Palmas: Napo, SW of Tena on Río Anzu, - 1.03, - 77.48 from Clark-Mcintyre notes Latacunga: Cotopaxi Limoncocha: Sucumbíos (formerly Napo Province) Macuma: Morano-Santiago, E of Macas Mangosisa: Morano-Santiago Manual Cornego: Pichincha, near Tandapi Machachi: Pichincha, Mejía Canton Nanegal: Pichincha Naranjapata: Chimborazo Province, Cumandá Naranjito: Guayas Quevedo: Los Ríos Quebrada: Napo, on the Sapalloyacu off of the Jatun Yuca Pacayacu: Pastaza, 20 km downstream from Canelos Palanque: Los Ríos Palmar: Manabí Paramba: Imbabura Province, Ibarra Canton Pifo: Pichincha, on road to Baeza Posorja: Guayas, SW of Guayaquil Primavera: — as Napo by Dunkle. Could be Sucumbíos or Orelano Reventador: Sucumbíos [N of Baeza just past provincial border on road to Lago Agrío = Sucumbíos Pacai-yacu: Pastaza on Río Bobonaza Partidero: Pastaza on Río Anzu Puna Vieja: Guayas Proovine, Guayaquil Canton Riobamba: Chimborazo Río Anzu: Río Napo watershed, headwaters in Pastaza N of Mera, larger “ white-water ” section in Napo Province NW of Tena. It is presumed that Clark-Macintyre collected in the Pastaza portion Río Bobonaza: Pastaza Río Chucapi: Napo, near Hacienda Ila Río Churi-Yacu: Pastaza Río Huagra-Yacu: Pastaza Río Jatun Yacu: Napo, approx. 20 km W of Tena Río Leon: Azuay, — as “ Río Negro ” by mistake during 1997 trip Río Papayacu: Pastaza, E of Puyo Río Shicai-yacu: Río Chaca-yacu, Pastaza watershed, NE of Tena, 900 m Río Salado: Napo, El Chaco, between Baeza and San Rafael Río Topo: Tungurahua, 31 km E of Baños Río Tuyano: Napo, 30 km SE of Tena Río Upano: Morano-Santiago, near Mangosisa Río Yanamanaca: Pastaza, trib to Río Anzu near El Partidero Tandapi: Pichincha, Mejía Canton Tinalandia: Santo Domingo de los Tsáchilas (formerly Pichincha) Sabanilla: Guayas, Pedro-Carbo Canton, near Pedro-Carbo Sapallo Yacu: Napo, Tena, near Jatun Satcha San Antonio de Curaray: Pastaza, 3100 m San Rafael: Carchi Santa Elena: Guayas, western part near Pacific Santa Inez: Tungurahua, Santa ines on the Río Pastaza, 27 km E of Baños Sarayacu: Pastaza, 35 km downstream from Canelos Venecia: Guayas, (also mistakenly listed as “ Verecia ” on some labels) Yaguachi: Guayas, just E of Guayaquil Yanamanaca: Pastaza, near Partidero on the Río Anzu Zumba: Zamora-Chinchipe APPENDIX C. Useful resources for further studies	en	Mauffray, William F., Tennessen, Kenneth J. (2019): A Catalogue and Historical Study of the Odonata of Ecuador. Zootaxa 4628 (1): 1-265, DOI: 10.11646/zootaxa.4628.1.1
