taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
F5E72BFF597459FCB21C1A27E2A64C0A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1428291	https://doi.org/10.3897/herpetozoa.38.e165173.figure1	Figure 1. Distribution of the lineages of the Ingerophrynus parvus species complex as identified in the phylogenetic analyses. Stars denote type localities, with their corresponding taxon names in colored clouds; colors correspond to those used in Figs 2–5; symbol shapes denote mtDNA lineages shown in Fig. 2. Base map created using simplemappr. net. For locality information, see Suppl. material 1: table S 2. Photographs by P. Pawangkhanant and N. A. Poyarkov.	Figure 1. Distribution of the lineages of the Ingerophrynus parvus species complex as identified in the phylogenetic analyses. Stars denote type localities, with their corresponding taxon names in colored clouds; colors correspond to those used in Figs 2–5; symbol shapes denote mtDNA lineages shown in Fig. 2. Base map created using simplemappr. net. For locality information, see Suppl. material 1: table S 2. Photographs by P. Pawangkhanant and N. A. Poyarkov.	2025-10-01	Arkhipov, Dmitriy V.;Pawangkhanant, Parinya;Sarker, Goutam;Gorin, Vladislav A.;Bragin, Andrey M.;Nguyen, Tan Van;Hamidy, Amir;Muin, Mohd Abdul;Kurniawan, Nia;Brown, Rafe M.;Suwannapoom, Chatmongkon;Smith, Eric;Poyarkov, Nikolay A.		Zenodo	biologists	Arkhipov, Dmitriy V.;Pawangkhanant, Parinya;Sarker, Goutam;Gorin, Vladislav A.;Bragin, Andrey M.;Nguyen, Tan Van;Hamidy, Amir;Muin, Mohd Abdul;Kurniawan, Nia;Brown, Rafe M.;Suwannapoom, Chatmongkon;Smith, Eric;Poyarkov, Nikolay A.			
F5E72BFF597459FCB21C1A27E2A64C0A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1428292	https://doi.org/10.3897/herpetozoa.38.e165173.figure2	Figure 2. Phylogenetic relationships among the Ingerophrynus parvus species complex and other related Ingerophrynus species based on the mtDNA fragment including the 12 S rRNA, 16 S rRNA, and ND 1 gene sequences (up to 4,479 bp). For voucher specimen information and GenBank accession numbers, see Suppl. material 1: table S 2. Numbers at tree nodes correspond to ML UFBS / BI PP support values, respectively. Photographs by P. Pawangkhanant and N. A. Poyarkov.	Figure 2. Phylogenetic relationships among the Ingerophrynus parvus species complex and other related Ingerophrynus species based on the mtDNA fragment including the 12 S rRNA, 16 S rRNA, and ND 1 gene sequences (up to 4,479 bp). For voucher specimen information and GenBank accession numbers, see Suppl. material 1: table S 2. Numbers at tree nodes correspond to ML UFBS / BI PP support values, respectively. Photographs by P. Pawangkhanant and N. A. Poyarkov.	2025-10-01	Arkhipov, Dmitriy V.;Pawangkhanant, Parinya;Sarker, Goutam;Gorin, Vladislav A.;Bragin, Andrey M.;Nguyen, Tan Van;Hamidy, Amir;Muin, Mohd Abdul;Kurniawan, Nia;Brown, Rafe M.;Suwannapoom, Chatmongkon;Smith, Eric;Poyarkov, Nikolay A.		Zenodo	biologists	Arkhipov, Dmitriy V.;Pawangkhanant, Parinya;Sarker, Goutam;Gorin, Vladislav A.;Bragin, Andrey M.;Nguyen, Tan Van;Hamidy, Amir;Muin, Mohd Abdul;Kurniawan, Nia;Brown, Rafe M.;Suwannapoom, Chatmongkon;Smith, Eric;Poyarkov, Nikolay A.			
F5E72BFF597459FCB21C1A27E2A64C0A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1428298	https://doi.org/10.3897/herpetozoa.38.e165173.figure8	Figure 8. Holotype of Ingerophrynus chrysolophus sp. nov. from Wat Tham Sanook, Tha Sae, Chumphon, Thailand, in life — specimen ZMMU A- 8030 (adult female). Dorsal aspect (A); ventral aspect (B); lateral view of head (C); dorsal view of head (D); volar view of the left hand (E); plantar view of the right foot (F). Photographs by N. A. Poyarkov. Scale bars: 5 mm.	Figure 8. Holotype of Ingerophrynus chrysolophus sp. nov. from Wat Tham Sanook, Tha Sae, Chumphon, Thailand, in life — specimen ZMMU A- 8030 (adult female). Dorsal aspect (A); ventral aspect (B); lateral view of head (C); dorsal view of head (D); volar view of the left hand (E); plantar view of the right foot (F). Photographs by N. A. Poyarkov. Scale bars: 5 mm.	2025-10-01	Arkhipov, Dmitriy V.;Pawangkhanant, Parinya;Sarker, Goutam;Gorin, Vladislav A.;Bragin, Andrey M.;Nguyen, Tan Van;Hamidy, Amir;Muin, Mohd Abdul;Kurniawan, Nia;Brown, Rafe M.;Suwannapoom, Chatmongkon;Smith, Eric;Poyarkov, Nikolay A.		Zenodo	biologists	Arkhipov, Dmitriy V.;Pawangkhanant, Parinya;Sarker, Goutam;Gorin, Vladislav A.;Bragin, Andrey M.;Nguyen, Tan Van;Hamidy, Amir;Muin, Mohd Abdul;Kurniawan, Nia;Brown, Rafe M.;Suwannapoom, Chatmongkon;Smith, Eric;Poyarkov, Nikolay A.			
F5E72BFF597459FCB21C1A27E2A64C0A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1428299	https://doi.org/10.3897/herpetozoa.38.e165173.figure9	Figure 9. Ingerophrynus chrysolophus sp. nov. in life in situ. A. Holotype ZMMU A- 8030 (adult female) from Wat Tham Sanook, Tha Sae, Chumphon, Thailand; B. Paratype ZMMU A- 8034 (adult male) from Wat Tham Sanook, Tha Sae, Chumphon, Thailand; C. ZMMU A- 8020 (adult male) from Pa Klok, Phuket, Thailand; D. ZMMU A- 8059 (adult male) from Mt. Khao Kra Jom, Suan Phueng, Ratchaburi, Thailand. Photographs by N. A. Poyarkov.	Figure 9. Ingerophrynus chrysolophus sp. nov. in life in situ. A. Holotype ZMMU A- 8030 (adult female) from Wat Tham Sanook, Tha Sae, Chumphon, Thailand; B. Paratype ZMMU A- 8034 (adult male) from Wat Tham Sanook, Tha Sae, Chumphon, Thailand; C. ZMMU A- 8020 (adult male) from Pa Klok, Phuket, Thailand; D. ZMMU A- 8059 (adult male) from Mt. Khao Kra Jom, Suan Phueng, Ratchaburi, Thailand. Photographs by N. A. Poyarkov.	2025-10-01	Arkhipov, Dmitriy V.;Pawangkhanant, Parinya;Sarker, Goutam;Gorin, Vladislav A.;Bragin, Andrey M.;Nguyen, Tan Van;Hamidy, Amir;Muin, Mohd Abdul;Kurniawan, Nia;Brown, Rafe M.;Suwannapoom, Chatmongkon;Smith, Eric;Poyarkov, Nikolay A.		Zenodo	biologists	Arkhipov, Dmitriy V.;Pawangkhanant, Parinya;Sarker, Goutam;Gorin, Vladislav A.;Bragin, Andrey M.;Nguyen, Tan Van;Hamidy, Amir;Muin, Mohd Abdul;Kurniawan, Nia;Brown, Rafe M.;Suwannapoom, Chatmongkon;Smith, Eric;Poyarkov, Nikolay A.			
F5E72BFF597459FCB21C1A27E2A64C0A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1428300	https://doi.org/10.3897/herpetozoa.38.e165173.figure10	Figure 10. Variation in dorsal coloration of the paratypes and referred material of Ingerophrynus chrysolophus sp. nov. from Wat Tham Sanook, Tha Sae, Chumphon, Thailand, in life. Paratypes: adult male, ZMMU A- 8034, ZMMU A- 8039; adult female, ZMMU A- 8031–8033, ZMMU A- 8035–8037. Referred material: subadult topotypic males, ZMMU A- 8038, RIM NAP- 11589; subadult topotypic females, ZMMU A- 8040–8044; subadults (sex not determined), ZMMU A- 8045, ZMMU A- 8046, RIM NAP- 11591, RIM NAP- 11593. Photographs by N. A. Poyarkov.	Figure 10. Variation in dorsal coloration of the paratypes and referred material of Ingerophrynus chrysolophus sp. nov. from Wat Tham Sanook, Tha Sae, Chumphon, Thailand, in life. Paratypes: adult male, ZMMU A- 8034, ZMMU A- 8039; adult female, ZMMU A- 8031–8033, ZMMU A- 8035–8037. Referred material: subadult topotypic males, ZMMU A- 8038, RIM NAP- 11589; subadult topotypic females, ZMMU A- 8040–8044; subadults (sex not determined), ZMMU A- 8045, ZMMU A- 8046, RIM NAP- 11591, RIM NAP- 11593. Photographs by N. A. Poyarkov.	2025-10-01	Arkhipov, Dmitriy V.;Pawangkhanant, Parinya;Sarker, Goutam;Gorin, Vladislav A.;Bragin, Andrey M.;Nguyen, Tan Van;Hamidy, Amir;Muin, Mohd Abdul;Kurniawan, Nia;Brown, Rafe M.;Suwannapoom, Chatmongkon;Smith, Eric;Poyarkov, Nikolay A.		Zenodo	biologists	Arkhipov, Dmitriy V.;Pawangkhanant, Parinya;Sarker, Goutam;Gorin, Vladislav A.;Bragin, Andrey M.;Nguyen, Tan Van;Hamidy, Amir;Muin, Mohd Abdul;Kurniawan, Nia;Brown, Rafe M.;Suwannapoom, Chatmongkon;Smith, Eric;Poyarkov, Nikolay A.			
F5E72BFF597459FCB21C1A27E2A64C0A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1428301	https://doi.org/10.3897/herpetozoa.38.e165173.figure11	Figure 11. Cranial morphology of Ingerophrynus chrysolophus sp. nov.: adult female, paratype ZMMU A- 8032 (A – C); adult male, paratype ZMMU A- 8059 (D – F), visualized via micro-CT scanning. Skulls are shown in dorsal (A, D), ventral (B, E), and lateral (C, F) aspects. For the lateral views only, the left half of the cranium is shown. Abbreviations: nas – nasal, sphen – sphenethmoid, prsph – parasphenoid, prm – premaxilla, smx – septomaxilla, pal – palatine, vom – vomer, max – maxilla, pt – pterygoid, fp – frontoparietal, qj – quadratojugal, soc – supraorbital crest, sq – squamosal, stc – supratympanic crest, pro – prootic, exoc – exoccipital, st – stapes, mnm – mentomeckelian, dnt – dentary, ansp – angulosplenial. Visualization by D. V. Arkhipov and V. A. Gorin. Scale bar: 5 mm; all pictures are to scale.	Figure 11. Cranial morphology of Ingerophrynus chrysolophus sp. nov.: adult female, paratype ZMMU A- 8032 (A – C); adult male, paratype ZMMU A- 8059 (D – F), visualized via micro-CT scanning. Skulls are shown in dorsal (A, D), ventral (B, E), and lateral (C, F) aspects. For the lateral views only, the left half of the cranium is shown. Abbreviations: nas – nasal, sphen – sphenethmoid, prsph – parasphenoid, prm – premaxilla, smx – septomaxilla, pal – palatine, vom – vomer, max – maxilla, pt – pterygoid, fp – frontoparietal, qj – quadratojugal, soc – supraorbital crest, sq – squamosal, stc – supratympanic crest, pro – prootic, exoc – exoccipital, st – stapes, mnm – mentomeckelian, dnt – dentary, ansp – angulosplenial. Visualization by D. V. Arkhipov and V. A. Gorin. Scale bar: 5 mm; all pictures are to scale.	2025-10-01	Arkhipov, Dmitriy V.;Pawangkhanant, Parinya;Sarker, Goutam;Gorin, Vladislav A.;Bragin, Andrey M.;Nguyen, Tan Van;Hamidy, Amir;Muin, Mohd Abdul;Kurniawan, Nia;Brown, Rafe M.;Suwannapoom, Chatmongkon;Smith, Eric;Poyarkov, Nikolay A.		Zenodo	biologists	Arkhipov, Dmitriy V.;Pawangkhanant, Parinya;Sarker, Goutam;Gorin, Vladislav A.;Bragin, Andrey M.;Nguyen, Tan Van;Hamidy, Amir;Muin, Mohd Abdul;Kurniawan, Nia;Brown, Rafe M.;Suwannapoom, Chatmongkon;Smith, Eric;Poyarkov, Nikolay A.			
F5E72BFF597459FCB21C1A27E2A64C0A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1428296	https://doi.org/10.3897/herpetozoa.38.e165173.figure6	Figure 6. The call of Ingerophrynus. parvus s. str. from Selangor, Malaysia (A) and Ingerophrynus сhrysolophus sp. nov. (B) from Chumphon, Thailand. A. 3 s waveform of relative amplitude (Rel. amp.) and corresponding spectrogram over time (above); 1 s waveform of relative amplitude (Rel. amp.) and corresponding spectrogram over time, obtained from the last two calls in A (below); B. 6.5 s waveform of relative amplitude (Rel. amp.) and corresponding spectrogram over time (above); 0.6 s waveform of relative amplitude (Rel. amp.) and corresponding spectrogram over time, obtained from the last two calls in B (below). Data on acoustic characters are summarized in Suppl. material 1: table S 6.	Figure 6. The call of Ingerophrynus. parvus s. str. from Selangor, Malaysia (A) and Ingerophrynus сhrysolophus sp. nov. (B) from Chumphon, Thailand. A. 3 s waveform of relative amplitude (Rel. amp.) and corresponding spectrogram over time (above); 1 s waveform of relative amplitude (Rel. amp.) and corresponding spectrogram over time, obtained from the last two calls in A (below); B. 6.5 s waveform of relative amplitude (Rel. amp.) and corresponding spectrogram over time (above); 0.6 s waveform of relative amplitude (Rel. amp.) and corresponding spectrogram over time, obtained from the last two calls in B (below). Data on acoustic characters are summarized in Suppl. material 1: table S 6.	2025-10-01	Arkhipov, Dmitriy V.;Pawangkhanant, Parinya;Sarker, Goutam;Gorin, Vladislav A.;Bragin, Andrey M.;Nguyen, Tan Van;Hamidy, Amir;Muin, Mohd Abdul;Kurniawan, Nia;Brown, Rafe M.;Suwannapoom, Chatmongkon;Smith, Eric;Poyarkov, Nikolay A.		Zenodo	biologists	Arkhipov, Dmitriy V.;Pawangkhanant, Parinya;Sarker, Goutam;Gorin, Vladislav A.;Bragin, Andrey M.;Nguyen, Tan Van;Hamidy, Amir;Muin, Mohd Abdul;Kurniawan, Nia;Brown, Rafe M.;Suwannapoom, Chatmongkon;Smith, Eric;Poyarkov, Nikolay A.			
F5E72BFF597459FCB21C1A27E2A64C0A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1428297	https://doi.org/10.3897/herpetozoa.38.e165173.figure7	Figure 7. Climatic models of ranges of Ingerophrynus parvus s. str. (A) and Ingerophrynus chrysolophus sp. nov. (B) built in MaxEnt. Red to green indicates a high probability of suitable conditions for the species; shades of blue indicate less suitable conditions. White dots correspond to known localities for both species (see Suppl. material 1: table S 5 for details on locality information and data sources).	Figure 7. Climatic models of ranges of Ingerophrynus parvus s. str. (A) and Ingerophrynus chrysolophus sp. nov. (B) built in MaxEnt. Red to green indicates a high probability of suitable conditions for the species; shades of blue indicate less suitable conditions. White dots correspond to known localities for both species (see Suppl. material 1: table S 5 for details on locality information and data sources).	2025-10-01	Arkhipov, Dmitriy V.;Pawangkhanant, Parinya;Sarker, Goutam;Gorin, Vladislav A.;Bragin, Andrey M.;Nguyen, Tan Van;Hamidy, Amir;Muin, Mohd Abdul;Kurniawan, Nia;Brown, Rafe M.;Suwannapoom, Chatmongkon;Smith, Eric;Poyarkov, Nikolay A.		Zenodo	biologists	Arkhipov, Dmitriy V.;Pawangkhanant, Parinya;Sarker, Goutam;Gorin, Vladislav A.;Bragin, Andrey M.;Nguyen, Tan Van;Hamidy, Amir;Muin, Mohd Abdul;Kurniawan, Nia;Brown, Rafe M.;Suwannapoom, Chatmongkon;Smith, Eric;Poyarkov, Nikolay A.			
