taxonID	type	description	language	source
F5E72BFF597459FCB21C1A27E2A64C0A.taxon	description	Figs 1, 2, 8, 9, 10, 11, Table 1	en	Arkhipov, Dmitriy V., Pawangkhanant, Parinya, Sarker, Goutam, Gorin, Vladislav A., Bragin, Andrey M., Nguyen, Tan Van, Hamidy, Amir, Muin, Mohd Abdul, Kurniawan, Nia, Brown, Rafe M., Suwannapoom, Chatmongkon, Smith, Eric, Poyarkov, Nikolay A. (2025): A taxonomic revision of the Ingerophrynus parvus species complex (Amphibia, Anura, Bufonidae) with the description of a new species from Thailand. Herpetozoa 38: 271-297, DOI: 10.3897/herpetozoa.38.e165173
F5E72BFF597459FCB21C1A27E2A64C0A.taxon	diagnosis	Diagnosis. A member of the genus Ingerophrynus with the following combination of morphological characters: a medium-sized species (SVL 30.3 – 35.7 mm in males, 34.0 – 42.4 mm in females); head large and wide (HL / HW 0.81 – 0.98 in males, 0.80 – 0.96 in females); parotoid elongate, narrow, and sharply raised; parotoid not continuous with an oblique row of conspicuously enlarged warts; warts on flanks less elevated than those of dorsum; cranial crests not thickened behind eyes; lores vertical; tympanum distinct, its diameter slightly exceeding two-thirds of eye length (TD / ED 0.53 – 0.64 in males, 0.51 – 0.77 in females); tibia relatively short (TIL / SVL 0.40 – 0.44 in males, 0.39 – 0.43 in females); males with a subgular vocal sac; no tarsal ridge or tibial gland; first finger longer than second; tip of third toe not reaching median subarticular tubercle of fourth toe; subarticular tubercles not enlarged; tarsal spine bases small; nuptial pads present; venter with low warts; ground color of flanks and dorsum light brown; dark brown stripes along the midline of the back; cranial ridges well-developed, bright orange.	en	Arkhipov, Dmitriy V., Pawangkhanant, Parinya, Sarker, Goutam, Gorin, Vladislav A., Bragin, Andrey M., Nguyen, Tan Van, Hamidy, Amir, Muin, Mohd Abdul, Kurniawan, Nia, Brown, Rafe M., Suwannapoom, Chatmongkon, Smith, Eric, Poyarkov, Nikolay A. (2025): A taxonomic revision of the Ingerophrynus parvus species complex (Amphibia, Anura, Bufonidae) with the description of a new species from Thailand. Herpetozoa 38: 271-297, DOI: 10.3897/herpetozoa.38.e165173
F5E72BFF597459FCB21C1A27E2A64C0A.taxon	description	Description of holotype (Fig. 8). Adult female in a good state of preservation, body stout (SVL 42.4 mm); head relatively large, wider than long (HL / HW 0.88); head with a pair of straight, cranial crests not thick, diverging and raised posteriorly (Fig. 8 D); snout projecting beyond lower jaw, gently rounded in lateral view (Fig. 8 C), truncated in dorsal view (Fig. 8 D); upper eyelid lacking supraciliary tubercles; protuberant nostrils at tip of snout with lateral orientation; canthus rostralis sharp, lores vertical (Fig. 8 C); tympanum distinct, approximately half diameter of eye (TD / ED 0.51); paratoid gland elongate, rounded, low in profile, separated from eyelid by supratympanic crest (Fig. 8 D); obliquely oriented row of enlarged, round to elongate dorsolateral warts extend posteriorly from paratoid gland approximately two-thirds way down flanks (Fig. 8 A). Forelimbs relatively long and slender (28.7 mm), fingers moderately long, tips blunt, not swollen; relative finger lengths: II <I <IV <III; fingers free of webbing (Fig. 8 E); finger subarticular tubercles distinct, conical; finger subarticular tubercle formula 1: 1: 2: 1; inner metacarpal tubercle slightly elongate; outer metacarpal tubercle triangular-shaped, dilated, bigger than inner metacarpal tubercle (IMC / OMC 0.63); two metacarpal tubercles not in touch with each other (Fig. 8 E). Hindlimbs robust and short; tibia relatively short (TIL / SVL 0.42); relative toe lengths: IV <II <I <III <IV; tips of toes like those of fingers; fourth toe nearly three times the length of third and fifth; third and fifth toes with nearly two phalanges free of web; fourth toe with four free phalanges; second toe nearly completely webbed; first toe completely webbed; webbing formula: i 1 - 1 ½ ii 1 ½ - 2 ½ iii 2 ½ - 3 iv 3 - 2 v; subarticular tubercles conspicuous, somewhat conical, much smaller than metatarsal tubercles, toe subarticular tubercle formula 1: 1: 2: 3: 1; inner metatarsal tubercle oval, shorter than length of first toe (IMT / T 1 0.37); outer metatarsal tubercle smaller, rounded (Fig. 8 F). Skin on flanks and dorsum covered with numerous conical warts, those of flanks lower in profile than those of dorsum; warts of dorsum capped by numerous spinules; enlarged series of nearly symmetrical paravertebral warts on dorsum beginning posterior to orbit and extending posteriorly beyond sacrum; venter covered with coarsely spinose granules.	en	Arkhipov, Dmitriy V., Pawangkhanant, Parinya, Sarker, Goutam, Gorin, Vladislav A., Bragin, Andrey M., Nguyen, Tan Van, Hamidy, Amir, Muin, Mohd Abdul, Kurniawan, Nia, Brown, Rafe M., Suwannapoom, Chatmongkon, Smith, Eric, Poyarkov, Nikolay A. (2025): A taxonomic revision of the Ingerophrynus parvus species complex (Amphibia, Anura, Bufonidae) with the description of a new species from Thailand. Herpetozoa 38: 271-297, DOI: 10.3897/herpetozoa.38.e165173
F5E72BFF597459FCB21C1A27E2A64C0A.taxon	description	Osteological description. The following description of adult skull morphology is based on the tomographic data obtained for the adult male (ZMMU A- 8059, paratype) and the adult female (ZMMU A- 8032, paratype) (Fig. 11).	en	Arkhipov, Dmitriy V., Pawangkhanant, Parinya, Sarker, Goutam, Gorin, Vladislav A., Bragin, Andrey M., Nguyen, Tan Van, Hamidy, Amir, Muin, Mohd Abdul, Kurniawan, Nia, Brown, Rafe M., Suwannapoom, Chatmongkon, Smith, Eric, Poyarkov, Nikolay A. (2025): A taxonomic revision of the Ingerophrynus parvus species complex (Amphibia, Anura, Bufonidae) with the description of a new species from Thailand. Herpetozoa 38: 271-297, DOI: 10.3897/herpetozoa.38.e165173
F5E72BFF597459FCB21C1A27E2A64C0A.taxon	etymology	Etymology. The species name “ сhrysolophus ” is a Latinized adjective in the nominative singular, masculine gender, derived from the Ancient Greek words “ χρυσός ” or “ chrysos, ” meaning “ gold, ” and “ λόφος ” or “ lophos, ” meaning “ crest ” or “ ridge. ” The species name is given in reference to the characteristic golden-orange coloration of supratympanic crests in the new species. We suggest the following common names for the new species: Golden-crested Dwarf Toad (in English), Khang kok khrae hua tong (คางคกแคระห ั วทอง, in Thai), and Zlatogrebnistaya shlemonosnaya zhaba (Златогребнистая шлемоносная жаба, in Russian).	en	Arkhipov, Dmitriy V., Pawangkhanant, Parinya, Sarker, Goutam, Gorin, Vladislav A., Bragin, Andrey M., Nguyen, Tan Van, Hamidy, Amir, Muin, Mohd Abdul, Kurniawan, Nia, Brown, Rafe M., Suwannapoom, Chatmongkon, Smith, Eric, Poyarkov, Nikolay A. (2025): A taxonomic revision of the Ingerophrynus parvus species complex (Amphibia, Anura, Bufonidae) with the description of a new species from Thailand. Herpetozoa 38: 271-297, DOI: 10.3897/herpetozoa.38.e165173
F5E72BFF597459FCB21C1A27E2A64C0A.taxon	distribution	Distribution. Ingerophrynus chrysolophus sp. nov. is reliably known from central, eastern, western, and southern Thailand (Mae Hong Son, Tak, Kamphaeng Phet, Uthai Thani, Kanchanaburi, Ratchaburi, Phetchaburi, Prachuap Khiri Khan, Chumpon, Ranong, Surat Thani, Phang Nga, Phuket, Krabi, Nakhon Si Thamarat, Trang, Songkhla, Satun, Trat, and Chanthaburi provinces); the adjacent parts of southern Myanmar (Tanintharyi Region and Yangon State); and southwest Cambodia (Cardamom Mountains) (Fig. 1). According to the results of the species distribution modeling (Fig. 7 B), the occurrence of the new species is expected in the northernmost Peninsular Malaysia (Perlis State); further studies are required to clarify the extent of its distribution in the Thai-Malay Peninsula.	en	Arkhipov, Dmitriy V., Pawangkhanant, Parinya, Sarker, Goutam, Gorin, Vladislav A., Bragin, Andrey M., Nguyen, Tan Van, Hamidy, Amir, Muin, Mohd Abdul, Kurniawan, Nia, Brown, Rafe M., Suwannapoom, Chatmongkon, Smith, Eric, Poyarkov, Nikolay A. (2025): A taxonomic revision of the Ingerophrynus parvus species complex (Amphibia, Anura, Bufonidae) with the description of a new species from Thailand. Herpetozoa 38: 271-297, DOI: 10.3897/herpetozoa.38.e165173
