taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
02B523B8A84257E99251CBE50DFBBA48.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1062327	https://doi.org/10.3897/zookeys.1204.117261.figure1	Figure 1. Map showing sampling sites of four new Marphysa species in Terengganu mangrove forest, east coast of Peninsular Malaysia A location of Terengganu on the east of Peninsular Malaysia B symbols indicate each sampling site; Setiu Wetlands (red star), Kuala Ibai (red rhombus), Merchang (red triangle), and Kerteh (red oval).	Figure 1. Map showing sampling sites of four new Marphysa species in Terengganu mangrove forest, east coast of Peninsular Malaysia A location of Terengganu on the east of Peninsular Malaysia B symbols indicate each sampling site; Setiu Wetlands (red star), Kuala Ibai (red rhombus), Merchang (red triangle), and Kerteh (red oval).	2024-06-04	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat		Zenodo	biologists	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat			
02B523B8A84257E99251CBE50DFBBA48.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1062328	https://doi.org/10.3897/zookeys.1204.117261.figure2	Figure 2. The schematic drawing of type of pectinate chaetae present in Marphysa from Terengganu A thin, narrow isodont with short and slender inner teeth (type 1) B thin, wide isodont with short and slender inner teeth (type 2) C thin, narrow heterodont with short and slender inner teeth (type 3) D thick, wide isodont with short and slender inner teeth (type 4) E thick, wide isodont with short and slender inner teeth (type 5) F thick, narrow anodont with long and thick inner teeth (type 6) G thick, wide anodont with long and slender inner teeth (type 7) H thick, wide anodont with long and thick inner teeth (type 8). Scale bars: 18 µm (A); 35 µm (B, G); 20 µm (C); 38 µm (D); 32 µm (E); 13 µm (F); 30 µm (H).	Figure 2. The schematic drawing of type of pectinate chaetae present in Marphysa from Terengganu A thin, narrow isodont with short and slender inner teeth (type 1) B thin, wide isodont with short and slender inner teeth (type 2) C thin, narrow heterodont with short and slender inner teeth (type 3) D thick, wide isodont with short and slender inner teeth (type 4) E thick, wide isodont with short and slender inner teeth (type 5) F thick, narrow anodont with long and thick inner teeth (type 6) G thick, wide anodont with long and slender inner teeth (type 7) H thick, wide anodont with long and thick inner teeth (type 8). Scale bars: 18 µm (A); 35 µm (B, G); 20 µm (C); 38 µm (D); 32 µm (E); 13 µm (F); 30 µm (H).	2024-06-04	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat		Zenodo	biologists	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat			
02B523B8A84257E99251CBE50DFBBA48.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1062340	https://doi.org/10.3897/zookeys.1204.117261.figure14	Figure 14. Marphysa ibaiensis sp. nov. Holotype UMTAnn 2179 (A – I). Light microscopy images and digital drawing A anterior end, lateral view. Arrow shows shallow groove B anterior end, dorsal view C maxillae, dorsal view D mandibles, dorsal view E bidentate hook, chaetiger 97 F parapodium, chaetiger 10 G parapodium, chaetiger 97 H parapodium chaetiger 145 I posterior segments and pygidium, dorsal view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 2 mm (A, D); 1 mm (B, C, I); 20 µm (E), 0.1 mm (F – H).	Figure 14. Marphysa ibaiensis sp. nov. Holotype UMTAnn 2179 (A – I). Light microscopy images and digital drawing A anterior end, lateral view. Arrow shows shallow groove B anterior end, dorsal view C maxillae, dorsal view D mandibles, dorsal view E bidentate hook, chaetiger 97 F parapodium, chaetiger 10 G parapodium, chaetiger 97 H parapodium chaetiger 145 I posterior segments and pygidium, dorsal view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 2 mm (A, D); 1 mm (B, C, I); 20 µm (E), 0.1 mm (F – H).	2024-06-04	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat		Zenodo	biologists	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat			
02B523B8A84257E99251CBE50DFBBA48.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1062341	https://doi.org/10.3897/zookeys.1204.117261.figure15	Figure 15. SEM images of Marphysa ibaiensis sp. nov. Holotype UMTAnn 2179 (B, C, E, F), paratype AM W. 54052 (A, D, G) A pectinate chaetae, chaetiger 97 B pectinate chaetae, chaetiger 10 C pectinate chaetae, chaetiger 88 D pectinate chaetae, chaetiger 88 E – F pectinate chaetae, chaetiger 125 G serrations and projections on limbate chaetae, chaetiger 31. Numbers denoted by arrows indicate the type of pectinate chaetae; 1. Thin, narrow isodont; 3. Thin, narrow heterodont; 4, 5. Thick, wide isodont; 7. Thick, wide anodont. Scale bars: 5 µm (A – B, D); 10 µm (C, F – H); 20 µm (E).	Figure 15. SEM images of Marphysa ibaiensis sp. nov. Holotype UMTAnn 2179 (B, C, E, F), paratype AM W. 54052 (A, D, G) A pectinate chaetae, chaetiger 97 B pectinate chaetae, chaetiger 10 C pectinate chaetae, chaetiger 88 D pectinate chaetae, chaetiger 88 E – F pectinate chaetae, chaetiger 125 G serrations and projections on limbate chaetae, chaetiger 31. Numbers denoted by arrows indicate the type of pectinate chaetae; 1. Thin, narrow isodont; 3. Thin, narrow heterodont; 4, 5. Thick, wide isodont; 7. Thick, wide anodont. Scale bars: 5 µm (A – B, D); 10 µm (C, F – H); 20 µm (E).	2024-06-04	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat		Zenodo	biologists	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat			
02B523B8A84257E99251CBE50DFBBA48.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1062342	https://doi.org/10.3897/zookeys.1204.117261.figure16	Figure 16. Sampling site in Kuala Ibai (estuary and lagoon area) A habitat of Marphysa ibaiensis B found in sediment deposited inside driftwood C live M. ibaiensis sp. nov.	Figure 16. Sampling site in Kuala Ibai (estuary and lagoon area) A habitat of Marphysa ibaiensis B found in sediment deposited inside driftwood C live M. ibaiensis sp. nov.	2024-06-04	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat		Zenodo	biologists	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat			
BF9FE410DC1E56E49E0865806C6310C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1062327	https://doi.org/10.3897/zookeys.1204.117261.figure1	Figure 1. Map showing sampling sites of four new Marphysa species in Terengganu mangrove forest, east coast of Peninsular Malaysia A location of Terengganu on the east of Peninsular Malaysia B symbols indicate each sampling site; Setiu Wetlands (red star), Kuala Ibai (red rhombus), Merchang (red triangle), and Kerteh (red oval).	Figure 1. Map showing sampling sites of four new Marphysa species in Terengganu mangrove forest, east coast of Peninsular Malaysia A location of Terengganu on the east of Peninsular Malaysia B symbols indicate each sampling site; Setiu Wetlands (red star), Kuala Ibai (red rhombus), Merchang (red triangle), and Kerteh (red oval).	2024-06-04	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat		Zenodo	biologists	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat			
BF9FE410DC1E56E49E0865806C6310C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1062328	https://doi.org/10.3897/zookeys.1204.117261.figure2	Figure 2. The schematic drawing of type of pectinate chaetae present in Marphysa from Terengganu A thin, narrow isodont with short and slender inner teeth (type 1) B thin, wide isodont with short and slender inner teeth (type 2) C thin, narrow heterodont with short and slender inner teeth (type 3) D thick, wide isodont with short and slender inner teeth (type 4) E thick, wide isodont with short and slender inner teeth (type 5) F thick, narrow anodont with long and thick inner teeth (type 6) G thick, wide anodont with long and slender inner teeth (type 7) H thick, wide anodont with long and thick inner teeth (type 8). Scale bars: 18 µm (A); 35 µm (B, G); 20 µm (C); 38 µm (D); 32 µm (E); 13 µm (F); 30 µm (H).	Figure 2. The schematic drawing of type of pectinate chaetae present in Marphysa from Terengganu A thin, narrow isodont with short and slender inner teeth (type 1) B thin, wide isodont with short and slender inner teeth (type 2) C thin, narrow heterodont with short and slender inner teeth (type 3) D thick, wide isodont with short and slender inner teeth (type 4) E thick, wide isodont with short and slender inner teeth (type 5) F thick, narrow anodont with long and thick inner teeth (type 6) G thick, wide anodont with long and slender inner teeth (type 7) H thick, wide anodont with long and thick inner teeth (type 8). Scale bars: 18 µm (A); 35 µm (B, G); 20 µm (C); 38 µm (D); 32 µm (E); 13 µm (F); 30 µm (H).	2024-06-04	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat		Zenodo	biologists	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat			
BF9FE410DC1E56E49E0865806C6310C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1062331	https://doi.org/10.3897/zookeys.1204.117261.figure5	Figure 5. Marphysa kertehensis sp. nov. Holotype UMTAnn 2181 (A – I). Light microscopy images and digital drawing A anterior end, lateral view B anterior end, dorsal view C maxillae, dorsal view D mandibles, dorsal view E parapodium, chaetiger 10 F parapodium, chaetiger 295 G parapodium, chaetiger 462 H median region, dorsal view. Arrows indicate black dot at the base of dorsal cirri I posterior segments and pygidium, ventral view. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Bf: branchial filament. Scale bars: 2 mm (A, H, I); 1 mm (B – D); 0.1 mm (E – G).	Figure 5. Marphysa kertehensis sp. nov. Holotype UMTAnn 2181 (A – I). Light microscopy images and digital drawing A anterior end, lateral view B anterior end, dorsal view C maxillae, dorsal view D mandibles, dorsal view E parapodium, chaetiger 10 F parapodium, chaetiger 295 G parapodium, chaetiger 462 H median region, dorsal view. Arrows indicate black dot at the base of dorsal cirri I posterior segments and pygidium, ventral view. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Bf: branchial filament. Scale bars: 2 mm (A, H, I); 1 mm (B – D); 0.1 mm (E – G).	2024-06-04	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat		Zenodo	biologists	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat			
BF9FE410DC1E56E49E0865806C6310C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1062332	https://doi.org/10.3897/zookeys.1204.117261.figure6	Figure 6. Scanning Electron Microscopy (SEM) images of Marphysa kertehensis sp. nov. Holotype UMTAnn 2181 (B – E), paratype AM W. 54059 (A, F) A limbate chaetae, chaetiger 10 B parapodia, chaetiger 300 C – D pectinate chaetae, chaetiger 300 E pectinate chaetae, chaetiger 462 F pectinate chaetae, chaetiger 525. Numbers denoted by arrows indicate the type of pectinate chaetae: 1. Thin, narrow isodont; 2. Thin, wide isodont; 4. Thick, wide isodont. Scale bars: 50 µm (A); 100 µm (B); 20 µm (C – D); 10 µm (E – F).	Figure 6. Scanning Electron Microscopy (SEM) images of Marphysa kertehensis sp. nov. Holotype UMTAnn 2181 (B – E), paratype AM W. 54059 (A, F) A limbate chaetae, chaetiger 10 B parapodia, chaetiger 300 C – D pectinate chaetae, chaetiger 300 E pectinate chaetae, chaetiger 462 F pectinate chaetae, chaetiger 525. Numbers denoted by arrows indicate the type of pectinate chaetae: 1. Thin, narrow isodont; 2. Thin, wide isodont; 4. Thick, wide isodont. Scale bars: 50 µm (A); 100 µm (B); 20 µm (C – D); 10 µm (E – F).	2024-06-04	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat		Zenodo	biologists	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat			
BF9FE410DC1E56E49E0865806C6310C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1062333	https://doi.org/10.3897/zookeys.1204.117261.figure7	Figure 7. Sampling site in Kerteh mangrove forest (river area) A habitat of Marphysa kertehensis sp. nov. B Marphysa kertehensis sp. nov. found inside driftwood and C in the sediment D live and complete M. kertehensis sp. nov.	Figure 7. Sampling site in Kerteh mangrove forest (river area) A habitat of Marphysa kertehensis sp. nov. B Marphysa kertehensis sp. nov. found inside driftwood and C in the sediment D live and complete M. kertehensis sp. nov.	2024-06-04	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat		Zenodo	biologists	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat			
37BB3E0577F650AD9CA20FA6E8FDE9F4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1062327	https://doi.org/10.3897/zookeys.1204.117261.figure1	Figure 1. Map showing sampling sites of four new Marphysa species in Terengganu mangrove forest, east coast of Peninsular Malaysia A location of Terengganu on the east of Peninsular Malaysia B symbols indicate each sampling site; Setiu Wetlands (red star), Kuala Ibai (red rhombus), Merchang (red triangle), and Kerteh (red oval).	Figure 1. Map showing sampling sites of four new Marphysa species in Terengganu mangrove forest, east coast of Peninsular Malaysia A location of Terengganu on the east of Peninsular Malaysia B symbols indicate each sampling site; Setiu Wetlands (red star), Kuala Ibai (red rhombus), Merchang (red triangle), and Kerteh (red oval).	2024-06-04	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat		Zenodo	biologists	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat			
37BB3E0577F650AD9CA20FA6E8FDE9F4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1062328	https://doi.org/10.3897/zookeys.1204.117261.figure2	Figure 2. The schematic drawing of type of pectinate chaetae present in Marphysa from Terengganu A thin, narrow isodont with short and slender inner teeth (type 1) B thin, wide isodont with short and slender inner teeth (type 2) C thin, narrow heterodont with short and slender inner teeth (type 3) D thick, wide isodont with short and slender inner teeth (type 4) E thick, wide isodont with short and slender inner teeth (type 5) F thick, narrow anodont with long and thick inner teeth (type 6) G thick, wide anodont with long and slender inner teeth (type 7) H thick, wide anodont with long and thick inner teeth (type 8). Scale bars: 18 µm (A); 35 µm (B, G); 20 µm (C); 38 µm (D); 32 µm (E); 13 µm (F); 30 µm (H).	Figure 2. The schematic drawing of type of pectinate chaetae present in Marphysa from Terengganu A thin, narrow isodont with short and slender inner teeth (type 1) B thin, wide isodont with short and slender inner teeth (type 2) C thin, narrow heterodont with short and slender inner teeth (type 3) D thick, wide isodont with short and slender inner teeth (type 4) E thick, wide isodont with short and slender inner teeth (type 5) F thick, narrow anodont with long and thick inner teeth (type 6) G thick, wide anodont with long and slender inner teeth (type 7) H thick, wide anodont with long and thick inner teeth (type 8). Scale bars: 18 µm (A); 35 µm (B, G); 20 µm (C); 38 µm (D); 32 µm (E); 13 µm (F); 30 µm (H).	2024-06-04	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat		Zenodo	biologists	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat			
37BB3E0577F650AD9CA20FA6E8FDE9F4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1062334	https://doi.org/10.3897/zookeys.1204.117261.figure8	Figure 8. Marphysa merchangensis sp. nov. Holotype UMTAnn 2149 (B – H), paratype UMTAnn 2148 (A). Light microscopy images and digital drawing A anterior end, lateral view B anterior end, dorsal view. Arrows indicate eyes C maxillae, dorsal view D mandibles, dorsal view E parapodium, chaetiger 10 F parapodium, chaetiger 134 G parapodium, chaetiger 250 H posterior segments and pygidium, ventral view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 1 mm (A – D, H); 0.1 mm (E – G).	Figure 8. Marphysa merchangensis sp. nov. Holotype UMTAnn 2149 (B – H), paratype UMTAnn 2148 (A). Light microscopy images and digital drawing A anterior end, lateral view B anterior end, dorsal view. Arrows indicate eyes C maxillae, dorsal view D mandibles, dorsal view E parapodium, chaetiger 10 F parapodium, chaetiger 134 G parapodium, chaetiger 250 H posterior segments and pygidium, ventral view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 1 mm (A – D, H); 0.1 mm (E – G).	2024-06-04	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat		Zenodo	biologists	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat			
37BB3E0577F650AD9CA20FA6E8FDE9F4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1062335	https://doi.org/10.3897/zookeys.1204.117261.figure9	Figure 9. SEM images of Marphysa merchangensis sp. nov. Holotype UMTAnn 2149 (A, B, E, H), paratype AM W. 54044 (C, D, F, G) A, B pectinate chaetae, chaetiger 10 C pectinate chaetae, chaetiger 164 D pectinate chaetae, chaetiger 245 E pectinate chaetae, chaetiger 250 F pectinate chaetae, chaetiger 265 G spiniger chaetae, chaetiger 10 H subacicular hook, chaetiger 128. Numbers denoted by arrows indicate the type of pectinate chaetae; 1. Thin, narrow isodont; 4, 5. Thick, wide isodont; 6. Thick, narrow anodont; 8. Thick, wide anodont. Scale bars: 5 µm (A – C, E); 10 µm (D, F, H); 50 µm (G).	Figure 9. SEM images of Marphysa merchangensis sp. nov. Holotype UMTAnn 2149 (A, B, E, H), paratype AM W. 54044 (C, D, F, G) A, B pectinate chaetae, chaetiger 10 C pectinate chaetae, chaetiger 164 D pectinate chaetae, chaetiger 245 E pectinate chaetae, chaetiger 250 F pectinate chaetae, chaetiger 265 G spiniger chaetae, chaetiger 10 H subacicular hook, chaetiger 128. Numbers denoted by arrows indicate the type of pectinate chaetae; 1. Thin, narrow isodont; 4, 5. Thick, wide isodont; 6. Thick, narrow anodont; 8. Thick, wide anodont. Scale bars: 5 µm (A – C, E); 10 µm (D, F, H); 50 µm (G).	2024-06-04	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat		Zenodo	biologists	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat			
37BB3E0577F650AD9CA20FA6E8FDE9F4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1062336	https://doi.org/10.3897/zookeys.1204.117261.figure10	Figure 10. Sampling site in Merchang mangrove estuary A habitat of M. merchangensis sp. nov. B – C worm found in decayed root of Exoecaria agallocha (Malay: Bebuta) D live worms.	Figure 10. Sampling site in Merchang mangrove estuary A habitat of M. merchangensis sp. nov. B – C worm found in decayed root of Exoecaria agallocha (Malay: Bebuta) D live worms.	2024-06-04	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat		Zenodo	biologists	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat			
37BB3E0577F650AD9CA20FA6E8FDE9F4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1062339	https://doi.org/10.3897/zookeys.1204.117261.figure13	Figure 13. Sampling in Setiu Wetlands (river area) A, C habitat of M. setiuense sp. nov. and M. merchangensis sp. nov. within Talipariti tiliaceum B Marphysa setiuense sp. nov. in situ D live M. setiuense sp. nov.	Figure 13. Sampling in Setiu Wetlands (river area) A, C habitat of M. setiuense sp. nov. and M. merchangensis sp. nov. within Talipariti tiliaceum B Marphysa setiuense sp. nov. in situ D live M. setiuense sp. nov.	2024-06-04	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat		Zenodo	biologists	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat			
01A9E8EEC19B5AB194C50570A564565C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1062327	https://doi.org/10.3897/zookeys.1204.117261.figure1	Figure 1. Map showing sampling sites of four new Marphysa species in Terengganu mangrove forest, east coast of Peninsular Malaysia A location of Terengganu on the east of Peninsular Malaysia B symbols indicate each sampling site; Setiu Wetlands (red star), Kuala Ibai (red rhombus), Merchang (red triangle), and Kerteh (red oval).	Figure 1. Map showing sampling sites of four new Marphysa species in Terengganu mangrove forest, east coast of Peninsular Malaysia A location of Terengganu on the east of Peninsular Malaysia B symbols indicate each sampling site; Setiu Wetlands (red star), Kuala Ibai (red rhombus), Merchang (red triangle), and Kerteh (red oval).	2024-06-04	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat		Zenodo	biologists	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat			
01A9E8EEC19B5AB194C50570A564565C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1062328	https://doi.org/10.3897/zookeys.1204.117261.figure2	Figure 2. The schematic drawing of type of pectinate chaetae present in Marphysa from Terengganu A thin, narrow isodont with short and slender inner teeth (type 1) B thin, wide isodont with short and slender inner teeth (type 2) C thin, narrow heterodont with short and slender inner teeth (type 3) D thick, wide isodont with short and slender inner teeth (type 4) E thick, wide isodont with short and slender inner teeth (type 5) F thick, narrow anodont with long and thick inner teeth (type 6) G thick, wide anodont with long and slender inner teeth (type 7) H thick, wide anodont with long and thick inner teeth (type 8). Scale bars: 18 µm (A); 35 µm (B, G); 20 µm (C); 38 µm (D); 32 µm (E); 13 µm (F); 30 µm (H).	Figure 2. The schematic drawing of type of pectinate chaetae present in Marphysa from Terengganu A thin, narrow isodont with short and slender inner teeth (type 1) B thin, wide isodont with short and slender inner teeth (type 2) C thin, narrow heterodont with short and slender inner teeth (type 3) D thick, wide isodont with short and slender inner teeth (type 4) E thick, wide isodont with short and slender inner teeth (type 5) F thick, narrow anodont with long and thick inner teeth (type 6) G thick, wide anodont with long and slender inner teeth (type 7) H thick, wide anodont with long and thick inner teeth (type 8). Scale bars: 18 µm (A); 35 µm (B, G); 20 µm (C); 38 µm (D); 32 µm (E); 13 µm (F); 30 µm (H).	2024-06-04	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat		Zenodo	biologists	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat			
01A9E8EEC19B5AB194C50570A564565C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1062337	https://doi.org/10.3897/zookeys.1204.117261.figure11	Figure 11. Marphysa setiuense sp. nov. Holotype UMTAnn 2177 (A – J). Light microscopy images and digital drawing A anterior end, lateral view B anterior end, dorsal view. Arrows indicate eyes C maxillae, dorsal view D mandibles, dorsal view E unidentate hook, chaetiger 136 F bidentate hooded hook, chaetiger 140 G parapodium, chaetiger 10 H parapodium, chaetiger 77 I parapodium, chaetiger 136 J posterior segments and pygidium, dorsal view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 1 mm (A, C, D, J); 1 mm (B); 20 µm (E – F); 0.1 mm (G – I).	Figure 11. Marphysa setiuense sp. nov. Holotype UMTAnn 2177 (A – J). Light microscopy images and digital drawing A anterior end, lateral view B anterior end, dorsal view. Arrows indicate eyes C maxillae, dorsal view D mandibles, dorsal view E unidentate hook, chaetiger 136 F bidentate hooded hook, chaetiger 140 G parapodium, chaetiger 10 H parapodium, chaetiger 77 I parapodium, chaetiger 136 J posterior segments and pygidium, dorsal view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 1 mm (A, C, D, J); 1 mm (B); 20 µm (E – F); 0.1 mm (G – I).	2024-06-04	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat		Zenodo	biologists	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat			
01A9E8EEC19B5AB194C50570A564565C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1062338	https://doi.org/10.3897/zookeys.1204.117261.figure12	Figure 12. SEM images of Marphysa setiuense sp. nov. Holotype UMTAnn 2177 (B, D – G), paratype AM W. 54050 (A, C, H) A pectinate chaetae, chaetiger 30 B pectinate chaetae, chaetiger 77 C pectinate chaetae, chaetiger 203 D pectinate chaetae, chaetiger 77 E, F pectinate chaetae, chaetiger 136 G spiniger chaetae, chaetiger 25 H subacicular hook, chaetiger 183. Numbers denoted by arrows indicate the type of pectinate chaetae; 1. Thin, narrow isodont; 2. Thin, wide isodont; 7, 8. Thick, wide anodont. Scale bars: 5 µm (A – C); 10 µm (D – F); 20 µm (G, H).	Figure 12. SEM images of Marphysa setiuense sp. nov. Holotype UMTAnn 2177 (B, D – G), paratype AM W. 54050 (A, C, H) A pectinate chaetae, chaetiger 30 B pectinate chaetae, chaetiger 77 C pectinate chaetae, chaetiger 203 D pectinate chaetae, chaetiger 77 E, F pectinate chaetae, chaetiger 136 G spiniger chaetae, chaetiger 25 H subacicular hook, chaetiger 183. Numbers denoted by arrows indicate the type of pectinate chaetae; 1. Thin, narrow isodont; 2. Thin, wide isodont; 7, 8. Thick, wide anodont. Scale bars: 5 µm (A – C); 10 µm (D – F); 20 µm (G, H).	2024-06-04	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat		Zenodo	biologists	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat			
01A9E8EEC19B5AB194C50570A564565C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1062339	https://doi.org/10.3897/zookeys.1204.117261.figure13	Figure 13. Sampling in Setiu Wetlands (river area) A, C habitat of M. setiuense sp. nov. and M. merchangensis sp. nov. within Talipariti tiliaceum B Marphysa setiuense sp. nov. in situ D live M. setiuense sp. nov.	Figure 13. Sampling in Setiu Wetlands (river area) A, C habitat of M. setiuense sp. nov. and M. merchangensis sp. nov. within Talipariti tiliaceum B Marphysa setiuense sp. nov. in situ D live M. setiuense sp. nov.	2024-06-04	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat		Zenodo	biologists	Che Engku Abdullah, Che Engku Siti Mariam;Idris, Izwandy;Fahmi, Afiq Durrani Mohd;Flaxman, Beth;Hutchings, Pat			
