taxonID	type	description	language	source
DB728788FF86FFD7FF6EFB6BFE31FB56.taxon	diagnosis	Diagnosis. Wing distinguished from those of other Cimbrophlebia species by the following: 1), 2 A morphology: four pectinate branches separates it from C. bittaciformis, [seven branches: three basal branches pectinate, four distal branches dichotomous], C. leahyi [six branches: two basal branches pectinate, minimum of four dichotomous distal branches], not known to be distinct from C. brooksi, C. westae, where 2 A is little known by preservation; 2), Shape: length about four times width separates it from C. brooksi [distinctly narrower: length about five times width], C. bittaciformis [similarly slender (see remarks)], (C. leahyi, C. westae length also about four times width); 3), Colouration distinct from that of C. brooksi, C. westae, C. bittaciformis (see below); closest to that of C. leahyi (as below) differing from C. leahyi by more overall dark colouration, including 2 A region [C. leahyi: light], further differentiated from C. leahyi by distinctly round / oval light spots [C. leahyi: differing position, sizes of more irregularly shaped light spots], (see descriptions, remarks, Figs. 1 A; 2 A, B; 4 A, C; 6 A, C; 7 A).	en	Archibald, Bruce (2009): New Cimbrophlebiidae (Insecta: Mecoptera) from the Early Eocene at McAbee, British Columbia, Canada and Republic, Washington, USA. Zootaxa 2249: 51-62, DOI: 10.5281/zenodo.190747
DB728788FF86FFD7FF6EFB6BFE31FB56.taxon	description	Description: holotype wing. As in diagnosis, Fig. 1 A, B, and the following. Length 28 mm, width 7 mm. Colouration as in Fig. 1 A: dark, with eight light spots (possibly more in basal-most region?) four along anterior portion, basal two smaller, distal two larger; four smaller spots along posterior portion. Sc: Sc 1 terminates on C ~ 2 / 3 wing length; Sc 2 separates from R 1 at pterostigmal region, joins with C at steep angle. R 1 branched, curved around pterostigma to join C. Rs with four dichotomous branches. M with four dichotomous branches; not preserved in basal quarter of wing. Branching of Cu 1, Cu 2 not preserved; Cu 1 long, straight, until 2 / 3 wing length, bent posteriad at m-cu to hind margin; Cu 2 straight for preserved portion, close to Cu 1, not preserved distad bend of Cu 1. 1 A: long, straight, close to Cu 2 in basal 2 / 3 of wing, apicad gently bent posteriad, meeting hind margin. 2 A with four pectinate branches (second branch with crossvein to posterior margin). 3 A: not detected. Few crossveins detected as preserved (Fig. 1 B).	en	Archibald, Bruce (2009): New Cimbrophlebiidae (Insecta: Mecoptera) from the Early Eocene at McAbee, British Columbia, Canada and Republic, Washington, USA. Zootaxa 2249: 51-62, DOI: 10.5281/zenodo.190747
DB728788FF86FFD7FF6EFB6BFE31FB56.taxon	materials_examined	Type material. Holotype: UCCIPR L- 18 F- 763 (part), UCCIPR L- 18 F- 764 (counterpart). A well preserved fore- or hind wing; housed at TRU; labelled: Holotype Cimbrophlebia flabelliformis, Archibald, 2009. Collected by unknown person at McAbee in 2002, donated to TRU by David Langevin, 2002. Locality and age. McAbee, British Columbia, Canada; Early Eocene.	en	Archibald, Bruce (2009): New Cimbrophlebiidae (Insecta: Mecoptera) from the Early Eocene at McAbee, British Columbia, Canada and Republic, Washington, USA. Zootaxa 2249: 51-62, DOI: 10.5281/zenodo.190747
DB728788FF86FFD7FF6EFB6BFE31FB56.taxon	etymology	Etymology. The specific epithet flabelliformis is formed from the Latin flabellum, “ fan ”, and forma, “ shape ”, referring to the overall impression of the somewhat evenly separated wing veins, radiating apicad in a fan-like manner.	en	Archibald, Bruce (2009): New Cimbrophlebiidae (Insecta: Mecoptera) from the Early Eocene at McAbee, British Columbia, Canada and Republic, Washington, USA. Zootaxa 2249: 51-62, DOI: 10.5281/zenodo.190747
DB728788FF86FFD7FF6EFB6BFE31FB56.taxon	discussion	Remarks. The configuration of the basal portion of 2 A seen in C. flabelliformis and C. sp. A corresponds with the vein fragment of C. bittaciformis previously identified as 3 A (Willmann, 1977: Fig. 2) and subsequently with uncertainty (Willmann, 1989: Fig. 135), indicating that this similarly-placed vein on C. bittaciformis may also be a portion of 2 A. The wing of C. flabelliformis is about four times as long as wide, as is that of C. leahyi (see bottom wing in Fig. 2 C), C. westae, and apparently C. sp. A judging by the shape of its preserved portion. This is distinct from the more slender wings of C. brooksi, length / width about 4.9, and C. bittaciformis (width not reported, but these appear distinctly long and slender: see Willmann, 1977: Fig. 1, complete wings of the holotype, and Fig. 2, incomplete wing of the paratype). These are confidently not simply forewing / hind wing differences, as shape appears reasonably consistent between them (comparable in C. leahyi and C. bittaciformis). The lengths of complete (or mostly so) cimbrophlebiid wings vary from about 25 to 38 mm: C. westae ~ 25 mm; C. flabelliformis, 28 mm; C. leahyi: ~ 31 mm; C. bittaciformis, hind wing 31.5 mm and forewing 34.5 mm; C. brooksi, ~ 38 mm. These differences, however, fall within variation found in the scorpionfly Panorpa communis (Ohm, 1961); further, where known, hind wings are a bit smaller than forewings. Only the great size differences between that modelled for the incomplete wing of C. sp. A (estimated as> 40 mm long) and the much smaller C. flabelliformis and C. westae are here considered meaningfully distinctive, given current knowledge. The holotype of C. flabelliformis bears four branches of Rs, unlike those of C. leahyi, C. brooksi, and C. westae, which have five branches. Willmann (1977) found this character to be plastic within C. bittaciformis (and see P. communis: Ohm, 1961), and so this is not considered diagnostic here. Wing colouration of the Okanagan Highlands species may be separated into three groups: 1), that of the McAbee species C. flabelliformis and C. leahyi; 2), of C. brooksi (Republic); and 3), of C. westae (Republic). C. flabelliformis and C. leahyi have variations of a general pattern consisting of mostly dark wing, with a series of large, irregular to oval light spots extending along the length of the anterior portion of the wing, touching the anterior margin: the smallest one or two in the basal quarter, a large one at about mid-wing, and a large one between this and the apex. There are various smaller spots along the posterior half of the wing. The preserved portion of C. sp. A indicates that it may belong to this group. The colouration of the C. westae wing is somewhat similar to that of the McAbee species, but with more, exaggerated light regions, which extend the length of much of the posterior wing as known, and a further light band apicad the large spot in the corresponding position of the apical-most one of C. flabelliformis and C. leahyi. The resultant remaining dark colouration is arranged as a lengthwise zigzag touching the anterior, but not posterior margin (Fig. 6 C). Much of this colour pattern is evident on regions of the wing not preserved in the single specimen of C. sp. A, which cannot be confidently excluded from this colouration group. The colour of C. brooksi is distinctly different from the above, with four dark bands extending from the anterior margin across about 2 / 3 of the wing width to about Cu 2 or 1 A. These are separated by light bands that connect with a light region that extends the length of the posterior portion of the wing. The apical region is preserved as generally light, and may have had complex light colouration in life. The wing of the Danish C. bittaciformis has a complex series of light and dark bands and spots (Willmann 1977: Figs. 1 – 2); the undescribed German species figured by Ansorge (2003: Fig 4 G) has a dark wing densely covered with small light spots.	en	Archibald, Bruce (2009): New Cimbrophlebiidae (Insecta: Mecoptera) from the Early Eocene at McAbee, British Columbia, Canada and Republic, Washington, USA. Zootaxa 2249: 51-62, DOI: 10.5281/zenodo.190747
DB728788FF84FFD1FF6EFAA5FF23F986.taxon	diagnosis	Diagnosis. Wing distinguished from those of other Cimbrophlebia species by the following: 1), 2 A morphology: six branches: two basal branches pectinate, minimum of four dichotomous distal branches separates it from C. bittaciformis [seven: three basal branches pectinate, four distal branches dichotomous], C. flabelliformis [four pectinate branches], not known to be distinct from C. brooksi, C. westae (2 A is little known by preservation); 2), Shape: wide wing, length about four times width is similar to that of C. westae, C. flabelliformis, separates it from C. brooksi [distinctly narrower: length about five times width], C. bittaciformis [also slender (see C. flabelliformis remarks)]; 4), Colouration: distinct from that of C. flabelliformis, C. brooksi, C. westae, C. bittaciformis (see descriptions, C. flabelliformis remarks, Figs. 1 A; 2 A, B; 4 A, C; 6 A, C; 7 A).	en	Archibald, Bruce (2009): New Cimbrophlebiidae (Insecta: Mecoptera) from the Early Eocene at McAbee, British Columbia, Canada and Republic, Washington, USA. Zootaxa 2249: 51-62, DOI: 10.5281/zenodo.190747
DB728788FF84FFD1FF6EFAA5FF23F986.taxon	description	Description: holotype (sex undetermined). As in diagnosis, Figs. 2 A – C; 3 A – C, and the following. Head: preserved characters as known in C. bittaciformis: long rostrum; large compound eyes; long antennae, ~ 14 mm preserved (incomplete). Thorax: poorly preserved. Legs: long, thin; tibial spurs, single, large tarsal claw preserved; dense, small hairs, some interspersed coarse, larger hairs detected on femur, tibia. Wing: length ~ 31 mm, width ~ 8 mm (likely, by preservation). Colouration (as preserved, see Fig. 2 A, B): wing mostly dark, but 2 A region light, light spots as above, Fig 2 A – B. Membrane rugose, except possibly in apical portion. Sc, R 1 as in C. flabelliformis. Rs with five branches. M with four branches; Cu 1, Cu 2, 1 A: little known by preservation; 2 A, long, branched as in diagnosis; 3 A not detected. Crossveins detected: one r 1 - sc, three csc 2, two rs-rs, one rs-m. Abdomen: partially, indistinctly preserved.	en	Archibald, Bruce (2009): New Cimbrophlebiidae (Insecta: Mecoptera) from the Early Eocene at McAbee, British Columbia, Canada and Republic, Washington, USA. Zootaxa 2249: 51-62, DOI: 10.5281/zenodo.190747
DB728788FF84FFD1FF6EFAA5FF23F986.taxon	materials_examined	Type material. Holotype: TRUIPR L- 018 F- 1160 (part) and TRUIPR L- 018 F- 1161 (counterpart). A rather complete specimen, in the collection of TRU. Labelled: Holotype Cimbrophlebia leahyi Archibald, 2009. Collected and donated to TRU by John Leahy, 2008. Locality and age. McAbee, British Columbia, Canada; unnamed formation of the Kamloops Group, Okanagan Highlands; Early Eocene.	en	Archibald, Bruce (2009): New Cimbrophlebiidae (Insecta: Mecoptera) from the Early Eocene at McAbee, British Columbia, Canada and Republic, Washington, USA. Zootaxa 2249: 51-62, DOI: 10.5281/zenodo.190747
DB728788FF84FFD1FF6EFAA5FF23F986.taxon	etymology	Etymology. The specific epithet is a patronym formed from the surname of the collector, John Leahy, recognizing his generosity by the loan of the holotype (and numerous other specimens) to the author and subsequent donation to the TRU collection.	en	Archibald, Bruce (2009): New Cimbrophlebiidae (Insecta: Mecoptera) from the Early Eocene at McAbee, British Columbia, Canada and Republic, Washington, USA. Zootaxa 2249: 51-62, DOI: 10.5281/zenodo.190747
DB728788FF84FFD1FF6EFAA5FF23F986.taxon	discussion	Remarks. The holotype is the most complete specimen of the family reported here, with body parts and all four wings attached. Preserved body parts conform to those reported for C. bittaciformis (Willmann, 1977) and Malmocimbrophlebia buergeri (Bechly & Schweigert, 2000), although preservation does not allow determination of whether the small hairs on the leg are arranged in rows as in C. bittaciformis (Willmann’s Fig. 6).	en	Archibald, Bruce (2009): New Cimbrophlebiidae (Insecta: Mecoptera) from the Early Eocene at McAbee, British Columbia, Canada and Republic, Washington, USA. Zootaxa 2249: 51-62, DOI: 10.5281/zenodo.190747
DB728788FF82FFD3FF6EF9F5FCB8FDC4.taxon	diagnosis	Diagnosis. Wing distinguished from those of other Cimbrophlebia species by the following: 1), Shape: narrow, length about five times width separates it from wider wings of C. leahyi, C. westae, C. flabelliformis [length about four times width], (C. bittaciformis similarly slender); 2), Colouration: distinct from that of C. flabelliformis, C. westae, C. bittaciformis, C. leahyi (see descriptions, C. flabelliformis remarks, Figs. 1 A; 2 A, B; 4 A, C; 6 A, C; 7 A).	en	Archibald, Bruce (2009): New Cimbrophlebiidae (Insecta: Mecoptera) from the Early Eocene at McAbee, British Columbia, Canada and Republic, Washington, USA. Zootaxa 2249: 51-62, DOI: 10.5281/zenodo.190747
DB728788FF82FFD3FF6EF9F5FCB8FDC4.taxon	description	Description: holotype wing. As in diagnosis, Figs. 4 A – B, and the following. Length ~ 31 mm long as preserved (incomplete, basal portion missing), 7 mm wide. Colouration as in Fig. 4 A, above. Membrane rugose, except possibly in apical portion. Sc as for genus. Rs with five branches, Rs 1, Rs 2 both branched apicad pterostigmal region, Rs 3 + 4 apparently not branched. M apparently with four branches, but region poorly preserved. Cu, Cu 1, Cu 2, 1 A poorly, fragmentarily preserved. 2 A: with four branches evident, most likely more branches. 3 A: not preserved. Crossveins: none detected. Paratype wing. As in diagnosis, Fig. 4 C – D, 5, and the following. Length ~ 38 mm as preserved, width not reliably measurable. Colouration as in Fig. 4 C, as in holotype, except more completely preserved (cf. Fig. 4 A, C). Membrane rugose, except possibly in apical portion. Sc: Sc 1 as for genus, Sc 2 not preserved. R 1 poorly preserved. Rs with four branches detected, region poorly preserved. Veins posteriad poorly, fragmentarily, or not preserved. Crossveins: none detected.	en	Archibald, Bruce (2009): New Cimbrophlebiidae (Insecta: Mecoptera) from the Early Eocene at McAbee, British Columbia, Canada and Republic, Washington, USA. Zootaxa 2249: 51-62, DOI: 10.5281/zenodo.190747
DB728788FF82FFD3FF6EF9F5FCB8FDC4.taxon	materials_examined	Type material. Holotype: SR 062005 A, B, part and counterpart. A well-preserved fore- or hind wing, missing basal portions; housed at SR; labelled: Holotype Cimbrophlebia brooksi, Archibald, 2009. Collected by Karl Volkman at Republic locality B 4131, June 2006. Paratype: SR 990405, part only. A mostly wellpreserved fore- or hind wing, but posterior region indistinct; housed at SR; labelled: Paratype Cimbrophlebia brooksi, Archibald, 2009. Collected by Caleb Brooks at Republic locality B 4131, October 1999. Locality and age. Republic, Washington, USA, University of Washington / Burke Museum locality B 4131; Early Eocene.	en	Archibald, Bruce (2009): New Cimbrophlebiidae (Insecta: Mecoptera) from the Early Eocene at McAbee, British Columbia, Canada and Republic, Washington, USA. Zootaxa 2249: 51-62, DOI: 10.5281/zenodo.190747
DB728788FF82FFD3FF6EF9F5FCB8FDC4.taxon	etymology	Etymology. The specific epithet is a patronym formed from the surname of Caleb Brooks, the collector of the paratype specimen, recognizing his generosity in this donation to SR.	en	Archibald, Bruce (2009): New Cimbrophlebiidae (Insecta: Mecoptera) from the Early Eocene at McAbee, British Columbia, Canada and Republic, Washington, USA. Zootaxa 2249: 51-62, DOI: 10.5281/zenodo.190747
DB728788FF82FFD3FF6EF9F5FCB8FDC4.taxon	discussion	Remarks. The rugose membrane of much of the wings of C. brooksi (Fig. 5) is not mentioned as present in C. bittaciformis by Willmann (1977) nor seen in his figures, nor on the wings of C. sp. A or C. flabelliformis. It is found in C. leahyi, and C. westae, however.	en	Archibald, Bruce (2009): New Cimbrophlebiidae (Insecta: Mecoptera) from the Early Eocene at McAbee, British Columbia, Canada and Republic, Washington, USA. Zootaxa 2249: 51-62, DOI: 10.5281/zenodo.190747
DB728788FF80FFDCFF6EFA03FD34F92C.taxon	diagnosis	Diagnosis. Wing distinguished from those of other Cimbrophlebia species by the following: 1), Shape: length about four times width separates it from C. brooksi [distinctly narrower: length about five times width], C. bittaciformis [similarly slender (see C. flabelliformis remarks)], (C. leahyi, C. westae length also about four times width); 2), Colouration: distinct from that of C. flabelliformis, C. brooksi, C. leahyi, C. bittaciformis (see C. flabelliformis remarks); closest to that of C. leahyi, further differentiated from C. leahyi by distinctly round / oval light spots [C. leahyi: differing position, sizes of more irregularly shaped light spots (see descriptions, C. flabelliformis remarks, Figs. 1 A; 2 A, B; 4 A, C; 6 A, C; 7 A).	en	Archibald, Bruce (2009): New Cimbrophlebiidae (Insecta: Mecoptera) from the Early Eocene at McAbee, British Columbia, Canada and Republic, Washington, USA. Zootaxa 2249: 51-62, DOI: 10.5281/zenodo.190747
DB728788FF80FFDCFF6EFA03FD34F92C.taxon	description	Description: holotype wing. Wing as in diagnosis, Fig. 6, and the following. Length ~ 25 mm, width ~ 6 mm. Colouration (basal third colouration poorly preserved) as in diagnosis. Membrane rugose, except in apical quarter, anal region. Sc, R 1 generalized for genus. Rs with five branches. M with three branches evident, region partly not preserved; Cu 1, Cu 2, 1 A generalized for genus; 2 A with five branches known (only apical portion clearly preserved); 3 A region not preserved. Crossveins: few detected as preserved; one r 1 - rs, one m-m, one m-cu 1.	en	Archibald, Bruce (2009): New Cimbrophlebiidae (Insecta: Mecoptera) from the Early Eocene at McAbee, British Columbia, Canada and Republic, Washington, USA. Zootaxa 2249: 51-62, DOI: 10.5281/zenodo.190747
DB728788FF80FFDCFF6EFA03FD34F92C.taxon	materials_examined	Type material. Specimen SRUI 0 99600, part only. A mostly well-preserved wing missing part of the apical posterior portion, with basal portion of 2 A region obscured, and basal third colouration obscured by mineral staining. Collected by Joanne West, April 19, 2009 at Republic. Housed at SR. Locality and age. Republic, Washington, USA, University of Washington / Burke Museum locality B 4131; Early Eocene.	en	Archibald, Bruce (2009): New Cimbrophlebiidae (Insecta: Mecoptera) from the Early Eocene at McAbee, British Columbia, Canada and Republic, Washington, USA. Zootaxa 2249: 51-62, DOI: 10.5281/zenodo.190747
DB728788FF80FFDCFF6EFA03FD34F92C.taxon	etymology	Etymology. The specific epithet is a patronym formed from the surname of Joanne West, the collector of the holotype specimen, recognizing her generosity in this donation to SR and valued participation in a collecting project on behalf of the author’s research.	en	Archibald, Bruce (2009): New Cimbrophlebiidae (Insecta: Mecoptera) from the Early Eocene at McAbee, British Columbia, Canada and Republic, Washington, USA. Zootaxa 2249: 51-62, DOI: 10.5281/zenodo.190747
DB728788FF8FFFDEFF6EF88AFC78FF60.taxon	description	Description. Wing as in Fig. 7 A – C, and the following. Length of portion preserved ~ 28 mm (estimated complete> 40 mm, see remarks, below), width ~ 7.5 mm (preserved portion, not maximum). Colouration as in Fig. 7 A, similar (preserved portion) to C. flabelliformis, C. leahyi; possibly, but less likely C. westae. Sc, R, Rs, M, small basal portions preserved. Cu 1, Cu 2, 1 A: preserved portions generalized as for genus. 2 A: six pectinate branches. Crossveins: none detected as preserved.	en	Archibald, Bruce (2009): New Cimbrophlebiidae (Insecta: Mecoptera) from the Early Eocene at McAbee, British Columbia, Canada and Republic, Washington, USA. Zootaxa 2249: 51-62, DOI: 10.5281/zenodo.190747
DB728788FF8FFFDEFF6EF88AFC78FF60.taxon	materials_examined	Material. UCCIPR L- 18 F- 998 (part), F- 1137 (counterpart). Fore- or hind wing, missing apical third and about two thirds of the anterior portion, and a small portion of basal hind margin; in the collection of TRU. Labelled: hypotype, Cimbrophlebia sp. A, Archibald, 2009. Collected by unknown person at McAbee in 2002, donated to TRU by David Langevin, 2002. Locality and age. McAbee, British Columbia, Canada; Early Eocene.	en	Archibald, Bruce (2009): New Cimbrophlebiidae (Insecta: Mecoptera) from the Early Eocene at McAbee, British Columbia, Canada and Republic, Washington, USA. Zootaxa 2249: 51-62, DOI: 10.5281/zenodo.190747
DB728788FF8FFFDEFF6EF88AFC78FF60.taxon	discussion	Remarks. The preserved portions of the wing of C. sp. A indicate that its shape is consistent with that of C. flabelliformis (Fig. 7 C). If so, its complete length would be likely over 40 mm. The width of the preserved portion (not maximum) is ~ 7.5 mm. By its large size, this is confidently separated from C. flabelliformis [length ~ 28 mm; width 7 mm] and C. westae [length ~ 25 mm, width ~ 6 mm]. The 2 A with six pectinate branches preserved separates it from C. bittaciformis [seven branches: three basal branches pectinate, four distal branches dichotomous], from C. leahyi [six branches: two basal branches pectinate, minimum of four dichotomous distal branches], and from C. flabelliformis [four pectinate branches], although not from C. brooksi and C. westae, where 2 A is little known by preservation. Cimbrophlebia sp. A apparently has a similar shape as known to C. flabelliformis, C. leahyi and C. westae, which are relatively wide, differing from the narrower wings of C. brooksi and C. bittaciformis (see above). Colouration, when known from a complete wing will surely be informative (see C. flabelliformis remarks above). Better-preserved specimens are needed to clarify supposition that this represents a distinct, separate species.	en	Archibald, Bruce (2009): New Cimbrophlebiidae (Insecta: Mecoptera) from the Early Eocene at McAbee, British Columbia, Canada and Republic, Washington, USA. Zootaxa 2249: 51-62, DOI: 10.5281/zenodo.190747
