taxonID	type	description	language	source
DA75665BFFA4FF87FF43FE9EFE09FA4E.taxon	description	1. Helochares anchoralis Sharp, 1890 Distribution: China: Fujian, Guangdong, Hainan, Hubei, Jiangxi, Sichuan, Taiwan, Yunnan; Cambodia, India, Philippines, Sri Lanka, Sumatra, Thailand, Vietnam. 2. Helochares crenatus Régimbart, 1903 Distribution: China: Yunnan. 3. Helochares densus Sharp, 1890 Distribution: China: Fujian, Guangdong, Guangxi, Hainan, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang. 4. Helochares hainanensis sp. n. Distribution: China: Hainan. 5. Helochares lentus Sharp, 1890 Distribution: China: Fujian, Guangdong, Guangxi, Guizhou, Hunan, Jiangxi, Sichuan, Taiwan, Yunnan; Bangladesh; Cambodia; India; Indonesia; Malaysia; Nepal; Sri Lanka; Thailand; Vietnam. 6. Helochares minor d’Orchymont, 1925 Distribution: China: Hainan (new record for China); India; Vietnam. 7. Helochares neglectus (Hope, 1845) Distribution: China: Fujian, Guangdong, Guangxi, Hainan, Hubei, Jiangsu, Jiangxi, Shanghai, Sichuan, Yunnan, Zhejiang; Cambodia; Indonesia; Malaysia; Philippines; Thailand; Vietnam. 8. Helochares nipponicus Hebauer, 1995 Distribution: China: Jilin (new record for China); Japan; South Korea. 9. Helochares sauteri d’Orchymont, 1943 Distribution: China: Guangdong, Guizhou, Hubei, Jiangxi, Sichuan, Taiwan, Zhejiang. 10. Helochares wuzhifengensis sp. n. Distribution: China: Jiangxi. 11. Helochares tengchongensis sp. n. Distribution: China: Yunnan.	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFA7FF84FF43FE76FA74F816.taxon	description	(Figs. 1 – 4)	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFA7FF84FF43FE76FA74F816.taxon	materials_examined	Material examined. Holotype (IAECAS): 1 male: “ CHINA: Hainan Province, / Qionghai City, Wanquan Town, | 2007.11.14, 19 ° 11 ’ N, 110 ° 23 ’ E, / 38 m, leg. Bian & Tong ”.	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFA7FF84FF43FE76FA74F816.taxon	diagnosis	Diagnosis. This new species is similar to Helochares (Hydrobaticus) minor in habitus, but can be distinguished by the following characteristics: (1) dorsal punctures on head, pronotum and elytra are slightly smaller and shallower than in H. minor (Figs. 1, 4); (2) elytra with ten rows of impressed serial punctures, not forming striae (Fig. 1); (3) serial punctures are slightly larger than interstrial punctures (Fig. 1); (4) interstriae on elytra wide and flat (Fig. 1); (5) aedeagus with median lobe very slender and spine-like in appearance, as long as parameres (Fig. 3); (6) parameres wider in basal 0.85, then suddenly narrowed, apices narrowly rounded (Fig. 3).	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFA7FF84FF43FE76FA74F816.taxon	description	Description. Habitus as in Figs. 1 – 2. BL: 4.5 mm, BW: 2.3 mm. Body oval, moderately convex. Head brown, pronotum and elytra yellowish-brown, apices of tarsal claws black, ventral side dark brown to black. Head (Fig. 4). Labrum brown, anterior margin straight. Clypeus brown, broad, anterior margin strongly emarginate, moderately expanded laterally in front of eyes, with somewhat rounded lateral margin; densely punctate, lateral area with slightly sparse punctures. Eyes slightly bulging, somewhat prominent; several setae anteriad to eyes. Frontoclypeal suture distinct. Systematic punctures small and distinct. Antennae with nine antennomeres, yellowish-brown; club loosely articulated, densely pubescent. Maxillary palpi moderately long and slender, about 1.2 times long as the width of head, yellow, the second palpomere curved inwards. Mentum and submentum with coarse punctures and wrinkles. Thorax (Fig. 1 – 2, 4). Pronotum about as twice wider than long; distinctly and densely punctate; anterior and posterior angles broadly rounded. Anterior margin slightly projected anteriad in mid-length, with a row of setae. Lateral margin slightly lighter in color; with a median dark mark closed to anterior margin and some larger punctures. Posterior margin slightly corrugate, with a transverse groove; near the posterior margin with a pair of small pits which overlap with black marks (Figs. 1, 4). Prosternum (Fig. 2) slightly bulging at the middle, not carinate; protruding posteriorly; pubescent and rugulose. Mesoventrite strongly convex at postmedian area. Metaventrite pubescent, with fine punctures, somewhat convex at middle portion. Elytra (Fig. 1) together slightly wider than pronotum, widest behind midpoint; with scutellary stria and ten impressed series of punctures, but not striate; interstrial punctures small and dense, as on head and pronotum; serial punctures slightly larger and deeper than interstrial punctures; interstriae wide and flat; posterior region with slightly sparse punctures. Scutellum triangular, with sparsely fine punctures. Legs. Femora dark brown, tibiae and tarsi brown, apices of tarsal claws black. Femora densely pubescent with apical fifth glabrous. Tarsi with some dense white scattered setae. Abdomen. All visible ventrites with dense pubescences and fine punctures; apex of the fifth ventrite strongly emarginate which is wider than deep (Fig. 2). Aedeagus (Fig. 3). Slender, 1.1 mm long. Basal piece symmetrical, very short, about 0.15 - times of the total length, the manubrium indistinct. Median lobe as long as parameres, very slender appearing as a long spine along its distal third; apophyses of median lobe extending beyond basal margin of basal piece. Parameres robust, wider in basal 0.85, then abruptly narrowed, apices narrowly rounded.	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFA7FF84FF43FE76FA74F816.taxon	distribution	Distribution. Known only from the type locality in Hainan Province, China.	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFA7FF84FF43FE76FA74F816.taxon	etymology	Etymology. Named after the type locality, Hainan Province, China. Habitat (Fig. 25). Slow flowing river, with pebbles, silt, deadwood and sides with short shrubs and fireweed.	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFA1FF83FF43FF57FA6FFADD.taxon	description	(Figs. 5 – 8)	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFA1FF83FF43FF57FA6FFADD.taxon	materials_examined	Material examined. Holotype (IAECAS): 1 male: “ CHINA: Jiangxi Province, Ganzhou City, / Shangyou County, Wuzhifeng Town, | 25 ° 57 ’ N, 114 ° 05 ’ E, 554 m, / 2009.10.05, leg. Bian & Tong ”.	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFA1FF83FF43FF57FA6FFADD.taxon	diagnosis	Diagnosis. This new species is similar to Helochares nipponicus in habitus, but can be distinguished by the following characteristics: (1) elytra gradually narrowed from midpoint (Fig. 5); broadly rounded along posterior half in H. nipponicus (Fig. 15); (2) elytra and ventral side lighter than H. nipponicus in color; (3) aedeagus (Fig. 7) with median lobe elongate, gradually widening towards apex, distal 0.25 narrowing to the apex. Median lobe slightly longer than parameres. Membranous inner sac with several small dentiform bulges on distal 0.2 to distal 0.4 (Fig. 7 a).	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFA1FF83FF43FF57FA6FFADD.taxon	description	Description. Habitus as in Figs. 5 – 6. BL: 4.5 mm, BW: 2.2 mm. Body oval, moderately convex. Head, pronotum, elytra and ventral side dark brown, apex of tarsal claws black. Head (Fig. 5). Labrum black, sparsely punctate, anterior margin straight. Clypeus dark brown, broad, densely punctate, anterior margin broadly emarginate; area in front of eyes with a few sparse punctures, moderately expanded laterally. Eyes rounded in dorsal view, slightly bulging, somewhat prominent. Frontoclypeal suture distinct. Systematic punctures small and distinct. Antennae with nine antennomeres, yellowish-brown; club loosely articulated, densely pubescent. Maxillary palpi moderately long and slender, about 1.2 times long as the width of head, yellow, second palpomere curved inwards. Mentum and submentum dark brown, with coarse punctures and wrinkles. Thorax (Fig. 5 – 6, 8). Maximum width of pronotum about twice as its length; distinctly and densely punctate; anterior and posterior angles broadly rounded. Anterior margin slightly projected anteriad in mid-length; a median black mark near anterior margin. Lateral margin yellowish-brown, with large punctures on ventral surface. Posterior margin mildly corrugate, with a transverse groove; a pair of small pits which overlap with black marks in posterior area (Fig. 5). Prosternum (Fig. 6) slightly bulging at the middle, not carinate, protruding posteriorly; pubescent and rugulose, with fine and dense punctures. Mesoventrite strongly convex at postmedian area. Metaventrite pubescent, with fine punctures, somewhat convex at middle portion, convex part with long and dense setae. Elytra (Fig. 8) together slightly wider than pronotum, subparallel in basal 0.6, then gradually narrowed posteriorly; with scutellary stria and ten impressed series of punctures; interstriae with small and dense punctures, serial punctures distinctly larger and deeper than the interstrial punctures; interstriae wide and flat; large punctures on underside of elytra usually visible from dorsal view on distal quarter. Scutellum shield triangular, without punctures. Legs. Femora dark brown, tibiae and tarsi brown, apex of tarsal claws black. Femora densely pubescent with apical fifth glabrous. Tarsi with dense white setae. Abdomen (Fig. 6). All visible ventrites with dense pubescences and fine punctures; apex of ventrite five strongly emarginate. Aedeagus (Fig. 7). Elongated, 1.2 mm long. Basal piece very short, about 0.17 - times of aedeagus length. Median lobe gradually widened from base to apex, distal 0.25 narrowed, slightly longer than parameres, apex narrowly rounded; Membranous inner sac with several small dentiform bulges on distal 0.2 to distal 0.4 (Fig. 7 a). Median lobe with two strong spines medially (Fig. 7 b).	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFA1FF83FF43FF57FA6FFADD.taxon	distribution	Distribution. Known only from the type locality in Jiangxi Province, China.	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFA1FF83FF43FF57FA6FFADD.taxon	etymology	Etymology. Named after the type locality, Wuzhifeng Town, Jiangxi Province, China. Habitat (Fig. 26). Slow flowing stream, with pebbles, silt, mossy stones, deadwood and sides with fireweed.	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFA0FF81FF43FA6AFAAAFAD6.taxon	description	(Figs. 9 – 12)	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFA0FF81FF43FA6AFAAAFAD6.taxon	materials_examined	Material examined. Holotype (IAECAS): 1 male, “ CHINA: Yunnan Province, Tengchong City, / Lianghe County, Longhe Village, 1074 m, | 24 ° 48 ′ 21.158 ″ N, 98 ° 17 ′ 51.522 ″ E, / 2016.10.22, leg. Wang, Peng & Sun ”. Paratypes (47): (IAECAS): 17 males, 8 females: “ CHINA: Yunnan Province, the road from / Nanleng village, Mangshi to Ruili City, | 839 m, 24 ° 9 ′ 37.812 ″ N, 98 ° 6 ′ 24.198 ″ E, / 2016.10.16, leg. Wang, Peng & Sun ”; 6 males, 6 females: “ CHINA: Yunnan Province, the road from / Nanleng village, Mangshi to Ruili City, | 827 m, 24 ° 13 ’ 50.83 ” N, 98 ° 15 ’ 51.534 ” E, / 2016.10.16, leg. Wang, Peng & Sun ”; 3 males: “ CHINA: Yunnan Province, Ruili City, / Kema Village, 839 m, | 24 ° 3 ′ 54.540 ″ N, 97 ° 57 ′ 47.813 ″ E, / 2016.10.17, leg. Wang, Peng & Sun ”; 1 male, 3 females: “ CHINA: Yunnan Province, Mangshi, / Nongkan Village, 780 m, | 24 ° 9 ′ 46.692 ″ N, 98 ° 6 ′ 5.142 ″ E, / 2016.10.18, leg. Wang, Peng & Sun ”; 1 female: “ CHINA: Yunnan Province, Tengchong City, / Mingguang Village bridge downstream, 1826 m, | 25 ° 28 ′ 39.318 ″ N, 98 ° 31 ′ 53.016 ″ E, / 2016.10.23, leg. Wang, Peng & Sun ”; 1 female: “ CHINA: Yunnan Province, Baoshan City, / Mangkuan Town, Xinguang Village, 1026 m, | 98 ° 50 ’ 53 ” E, 25 ° 28 ’ 18 ” N, / 2018.10.23, leg. Peng, Zhu & Dong ”; 1 male: “ CHINA: Yunnan Province, Baoshan City, / Mangkuan Town, Mangkuan Village, 847 m, | 98 ° 51 ’ 44 ” E, 25 ° 26 ’ 53 ” N, / 2018.10.23, leg. Peng, Zhu & Dong ”.	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFA0FF81FF43FA6AFAAAFAD6.taxon	diagnosis	Diagnosis. This new species is similar to Helochares lentus in habitus, but can be distinguished by the following characteristics: (1) H. tengchongensis (3.6 – 4.2 mm) is usually smaller than H. lentus (4.0 – 5.5 mm); (2) serial punctures on elytra are smaller, interstrial punctures are denser than in H. lentus (Fig. 9); (3) aedeagus (Fig. 11) broad. Apex of median lobe with a pair of cylindric protuberances. Membranous inner sac with a torch-shaped structure medially (Fig. 11 b), and strongly sclerotized with some dentate structures (Fig. 11 c).	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFA0FF81FF43FA6AFAAAFAD6.taxon	description	Description. Habitus as in Figs. 9 – 10. BL: 3.6 – 4.2 mm, BW: 1.8 – 2.2 mm. Body oval, moderately convex. Head and elytra dark brown, pronotum brown, apex of tarsal claws black, ventral side dark brown. Head (Fig. 9). Labrum black, anterior margin straight, with sparse and fine punctures. Clypeus dark brown, broad, densely punctate; anterior margin broadly emarginate; the area in front of eyes moderately expanded laterally, with sparse punctures. Eyes rounded in dorsal view, slightly bulging; somewhat prominent; several setae anteriad to eyes. Frontoclypeal suture distinct, a pair of splayed black marks on frons. Systematic punctures small and distinct. Antennae with nine antennomeres, yellowish-brown; club loosely articulated, with densely pubescences. Maxillary palpi moderately long and slender, about 1.0 times long as the width of head, yellow, second palpomere curved inwards. Mentum and submentum dark brown, with coarse punctures and wrinkles. Thorax (Figs. 9 – 10, 12). Maximum width of pronotum about as twice as length; punctures distinct and dense; anterior and posterior angles broadly rounded. Anterior margin slightly projected anteriad in mid-length, with a row of setae, a median black mark near anterior margin. Lateral margin slightly lighter in color. Posterior margin slightly corrugate; posterior area with a pair of small pits which overlap with black marks (Fig. 9). Prosternum (Fig. 12) slightly bulging at the middle, not carinate, protruding posteriorly, rugulose, with fine and dense punctures. Mesoventrite strongly convex posteromedially. Metaventrite pubescent, with fine punctures and somewhat convex at middle portion, with convex part densely pubescent (Fig. 10). Elytra (Fig. 9) together slightly wider than pronotum, widest at middle, then distinctly attenuate posteriorly; elytra with scutellary stria and ten impressed series of punctures, but not striate; interstrial punctures small, serial punctures slightly larger than interstrial punctures; large punctures on underside of elytra usually visible from dorsal view on distal quarter. Elytral suture distinct; scutellum shield triangular, with several fine punctures. Legs. Femora dark brown, tibiae and tarsi brown, apex of tarsal claws black. Femora densely pubescent with apical fifth glabrous. Tarsi with dense white setae. Abdomen. All visible ventrites with dense pubescences and fine punctures; apex of ventrite five strongly emarginate. Aedeagus (Fig. 11). Broad, 1.0 mm long. Basal piece short, about 0.2 - times of aedeagus length. Parameres nearly parallel-sided on outer margins, apical third gradually narrowed and apex narrowly rounded. Median lobe longer than parameres, about 4.5 times as long as basal piece, apex with two cylindric protuberances (Fig. 11 a); apophyses of median lobe extending beyond base of basal piece. Membranous inner sac with a torch-sharped structure medially (Fig. 11 b), and strongly sclerotized with some dentate structures (Fig. 11 c).	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFA0FF81FF43FA6AFAAAFAD6.taxon	distribution	Distribution. Known only from Yunnan Province, China.	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFA0FF81FF43FA6AFAAAFAD6.taxon	etymology	Etymology. Named after the type locality, Tengchong City, Yunnan Province, China. Habitat (Fig. 27). Slow flowing pool, with mud, deadwood and sides with mossy stones, and fireweed.	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFA2FF8EFF43FA2EFBF8FF7A.taxon	description	(Figs. 13 – 14, 21)	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFA2FF8EFF43FA2EFBF8FF7A.taxon	materials_examined	Material examined: 4 exs. (IAECAS): Hainan Province, Qionghai, Baishiling, 2007.11.14, 19 ° 11 ′ N, 110 ° 24 ′ E, 38 m, leg. Bian & Tong.	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFA2FF8EFF43FA2EFBF8FF7A.taxon	diagnosis	Diagnosis. Habitus as in Figs. 13 – 14. BL: 3.3 – 3.5 mm, BW: 1.6 – 1.7 mm. Body oval, moderately convex. Head, pronotum and elytra brown, apex of tarsal claws and ventral side black. Punctures on head dense. Punctures on pronotum denser and larger than those on head; pronotum with dark mark and several large punctures near anterior margin medially. Elytra with scutellary stria and ten striae; interstriae flat, with small punctures; strial punctures much larger and deeper than interstrial punctures. Aedeagus (Fig. 21) 0.7 mm long, distal 1 / 3 gradually narrowed. Basal piece short, about 0.2 - times of aedeagus length. Parameres short. Median lobe broadly membranous, with a sea anemone-like medial structure as apical region (Fig. 21 a). Apophyses of median lobe extending to base of basal piece. Membranous inner sac with dentiform bulges on each side (Fig. 21 b).	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFA2FF8EFF43FA2EFBF8FF7A.taxon	distribution	Distribution. China: Hainan (new record species to China); India; Vietnam.	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFACFF8FFF43FF57FBD4FD64.taxon	description	(Figs. 15 – 16, 22)	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFACFF8FFF43FF57FBD4FD64.taxon	materials_examined	Material examined: 7 exs. (IAECAS): Jilin Province, Baishan City, Baoquanshan Town, Bapandao Village, 2017.07.02, 41 ° 29 ’ 35 ” N, 127 ° 34 ’ 7 ” E, 962 m, leg. Peng.	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFACFF8FFF43FF57FBD4FD64.taxon	diagnosis	Diagnosis. Habitus as in Figs. 15 – 16. BL: 4.8 – 5.0 mm, BW: 2.5 – 2.8 mm. Body oval, moderately convex. Head and pronotum dark brown, elytra, ventral side and apex of tarsal claws black. Clypeus dark brown, broad; anterior margin broadly emarginate; frontoclypeal suture distinct; a pair of splayed black marks on frons. Pronotum with a dark mark near anterior margin medially. Elytra widest at midpoint, then slightly attenuated posteriorly. Strial punctures distinctly recognized, and interstrial punctures dense which smaller than the strial punctures. Aedeagus (Fig. 22) 1.0 mm long, strongly narrowed in distal 1 / 3. Basal piece short, about 0.3 - times of aedeagus length. Parameres with long spines crossing or nearly crossing at apex (Fig. 22 a). Median lobe broadly membranous, accompanied with a medial sea anemone-like structure (Fig. 22 b).	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFACFF8FFF43FF57FBD4FD64.taxon	distribution	Distribution. China: Jilin (new record species to China); Japan; South Korea.	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFACFF8DFF43FC40FBB9FC02.taxon	description	(Figs. 17 – 18, 23)	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFACFF8DFF43FC40FBB9FC02.taxon	materials_examined	Material examined: 6 exs. (IAECAS): Guangdong Province, Yunfu City, Yunan County, upper reaches of Dahe Reservoir, 90 m, 111 ° 24 ′ 53 ″ E, 23 ° 11 ′ 27 ″ N, 2017. 11.16, leg. Peng & Sun; 2 exs. (IAECAS): Guangdong Province, Yunfu City, Yunan County, water source upper reaches of Dahe Reservoir, 133 m, 111 ° 25 ′ 7 ″ E, 23 ° 12 ′ 2 ″ N, 2017.11.16, leg. Peng & Sun; 5 exs. (IAECAS): Guangdong Province, Yunfu City, Yunan County, Tongle Dashan Reserve, 269 m, 111 ° 23 ′ 6 ″ E, 23 ° 11 ′ 49 ″ N, 2017.11.16, leg. Peng & Sun; 2 exs. (IAECAS): Guangdong Province, Shaoguan City, Ruyuan County, lower reaches of Nanling Mountains reserve, 476 m, 113 ° 5 ′ 4 ″ E, 24 ° 55 ′ 46 ″ N, 2017.11.26, leg. Peng & Sun; 12 exs. (IAECAS): Guangxi Province, Jingxi City, Wuling Forest Park, 781 m, 106 ° 26 ’ 31 ” E, 22 ° 4 ’ 22 ” N, 2018.3.30, leg. Peng & Zhu; 10 exs. (IAECAS): Guangxi Province, Yuewei Town, 494 m, 106 ° 36 ’ 1 ” E, 22 ° 56 ’ 35 ” N, 2018.3.28, leg. Peng & Zhu; 1 ex. (IAECAS): Guangxi Province, Jingxi City, Longbang Town, Damao Village, 782 m, 106 ° 16 ’ 35 ” E, 22 ° 52 ’ 31 ” N, 2018.3.29, leg. Peng & Zhu; 1 ex. (IAECAS): Guangxi Province, Jingxi City, Longbang Town, Jiebang Village, 671 m, 106 ° 18 ’ 4 ” E, 22 ° 52 ’ 56 ” N, 2018.3.29, leg. Peng & Zhu; 1 ex. (IAECAS): Hainan Province, Qiongzhong County, lower reaches of Baihualing Waterfall, 347 m, 109 ° 49 ’ 15 ” E, 19 ° 0 ’ 30 ” N, 2018.3.15, leg. Tong, Peng & Zhu; 12 exs. (IAECAS): Hainan Province, Qiongzhong County, Shenyun Village, 253 m, 109 ° 36 ’ 21 ” E, 18 ° 59 ’ 40 ” N, 2018.3.17, leg. Tong, Peng & Zhu; 4 exs. (IAECAS): Hainan Province, Lingshui County, Diaoluo Mountain Forest Park, 720 m, 109 ° 36 ’ 13 ” E, 18 ° 40 ’ 26 ” N, 2018.3.22, leg. Tong, Peng & Zhu; 7 exs. (IAECAS): Yunnan Province, Tengchong City, Lianghe County, Longhe Village, 1074 m, 24 ° 48 ′ 21.158 ″ N, 98 ° 17 ′ 51.522 ″ E, 2016.10.22, leg. Wang, Peng & Sun; 5 exs. (IAECAS): Yunnan Province, Mangshi, Nongkan Village, 780 m, 24 ° 9 ′ 46.692 ″ N, 98 ° 6 ′ 5.142 ″ E, 2016.10.18, leg. Wang, Peng & Sun; 5 exs. (IAECAS): Yunnan Province, the road from Nanleng village, Mangshi to Ruili City, 839 m, 24 ° 9 ′ 37.812 ″ N, 98 ° 6 ′ 24.198 ″ E, 2016.10.16, leg Wang, Peng & Sun; 1 ex. (IAECAS): Yunnan Province, Tengchong City, Mingguang Village bridge downstream, 1826 m, 25 ° 28 ′ 39.318 ″ N, 98 ° 31 ′ 53.016 ″ E, 2016.10.23, leg. Wang, Peng & Sun; 2 exs. (IAECAS): Yunnan Province, Tongchong City, Zhongtang Village, 1803 m, 25 ° 26 ′ 3.102 ″ N, 98 ° 31 ′ 16.566 ″ E, 2016.10.23, leg. Wang, Peng & Sun; 26 exs. (IAECAS): Yunnan Province, the road from Nanleng village, Mangshi to Ruili City, 827 m, 24 ° 13 ′ 50.83 ″ N, 98 ° 15 ′ 51.534 ″ E, 2016.10.16, leg. Wang, Peng & Sun; 2 exs. (IAECAS): Yunnan Province, Ruili City, Kema Village, 839 m, 24 ° 3 ′ 54.540 ″ N, 97 ° 57 ′ 47.813 ″ E, 2016.10.17, leg. Wang, Peng & Sun; 1 ex. (IAECAS): Yunnan Province, Tongchong City, Mingguang Village bridge upstream, 1940 m, 25 ° 28 ′ 39.318 ″ N, 98 ° 31 ′ 53.016 ″ E, 2016.10.24, leg. Wang, Peng & Sun; 2 exs. (IAECAS): Yunnan Province, Tongchong City, Chenjia Village, 1615 m, 24 ° 52 ′ 59.268 ″ N, 98 ° 34 ′ 32.322 ″ E, 2016.10.26, leg. Wang, Peng & Sun; 3 exs. (IAECAS): Yunnan Province, Dali City, Yongping County, the road of Haichong, 1570 m, 99 ° 33 ’ 03 ” E, 25 ° 23 ’ 13 ” N, 2018.10.18, leg. Peng, Zhu & Dong; 1 ex. (IAECAS): Yunnan Province, Dali City, Yongping County, Changjie Village, 1533 m, 99 ° 34 ’ 22 ” E, 25 ° 19 ’ 23 ” N, 2018.10.18, leg. Peng, Zhu & Dong; 10 exs. (IAECAS): Yunnan Province, Baoshan City, Mangkuan Town, Mangkuan Village, 847 m, 98 ° 51 ’ 44 ” E, 25 ° 26 ’ 53 ” N, 2018.10.23, leg. Peng, Zhu & Dong; 1 ex. (IAECAS): Yunnan Province, Baoshan City, Xinguang Village, 1026 m, 98 ° 50 ’ 53 ” E, 25 ° 28 ’ 18 ” N, 2018.10.23, leg. Peng, Zhu & Dong.	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFACFF8DFF43FC40FBB9FC02.taxon	diagnosis	Diagnosis. Habitus as in Figs. 17 – 18. BL: 4.0 – 5.5 mm, BW: 2.0 – 2.8 mm. Body oval, moderately convex. Head, pronotum and elytra yellowish-brown, ventral side dark brown, apex of tarsal claws black. Clypeus brown, broad; anterior margin broadly emarginate; frons dark brown, with a pair of splayed black marks. Elytra slightly wider than pronotum; serial punctures distinct and sparse, much larger than the interstrial punctures, posterior punctures larger; interstriae wide and flat. Aedeagus (Fig. 23) 0.9 mm long. Basal piece very short, about 0.2 - times of aedeagus length. Median lobe distinctly longer than parameres, robust in basal 0.8, then distinctly narrowed, lateral margin strongly sclerotized in distal 0.2 (Fig. 23 a); membranous inner sac strongly sclerotized, with two long spines on each side (Fig. 23 b); apophyses of median lobe extending to base of basal piece. Parameres broad in basal 0.6, then gradually narrowed towards the apices (Fig. 23 c).	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFACFF8DFF43FC40FBB9FC02.taxon	distribution	Distribution. China: Fujian, Guangdong, Guangxi, Guizhou, Hunan, Jiangxi, Sichuan, Taiwan, Yunnan; Bangladesh; Cambodia; India; Indonesia; Malaysia; Nepal; Sri Lanka; Thailand; Vietnam.	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFAEFF8DFF43FC07FBF3F942.taxon	description	(Figs. 19 – 20, 24)	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFAEFF8DFF43FC07FBF3F942.taxon	materials_examined	Material examined: 2 exs. (IAECAS): Hainan Province, Qionghai City, Wanquan Town, 2007.11.14, 19 ° 11 ’ N, 110 ° 23 ’ E, 38 m, leg. Bian & Tong; 3 exs. (IAECAS): Hainan Province, Wanning City, 50 m, 1996.1.25, leg. Ji & Wang; 1 ex. (IAECAS): Yunan Province, Xishuangbanna, 1500 m, 1999.11, leg. Wang & Wei; 1 ex. (IAECAS): Guangxi Province, Nanning City, Huangjiang County, 208 m, 2011.11.14, leg. Bian, Guo & Tong.	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFAEFF8DFF43FC07FBF3F942.taxon	diagnosis	Diagnosis. Habitus as in Figs. 19 – 20. BL: 6.0 – 7.5 mm, BW: 3.00 – 3.65 mm. Body oval, large, moderately convex. Head, pronotum, elytra and ventral side dark brown, apex of tarsal claws black. Clypeus dark brown, broad; anterior margin broadly emarginate; maxillary palpi moderately long and slender, the second palpomere curved inwards. Anterior area of pronotum with a black mark shaped as an umbrella; posterior margin weakly corrugate, with a transverse groove. Elytra widest at mid-length, then slightly attenuated posteriorly; strial punctures deeply impressed, distinctly larger than the interstrial punctures; interstriae wide and flat. Aedeagus (Fig. 24) 1.5 mm long, elongate and parallel-sided. Basal piece short, about 0.2 - times of aedeagus length. Median lobe distinctly longer than parameres, apex of median lobe with a structure just liking “ horn ”, membranous inner sac strongly sclerotized (Fig. 24 a); apophyses of median lobe extending to base of basal piece. Parameres vestigial on basal 0.6.	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
DA75665BFFAEFF8DFF43FC07FBF3F942.taxon	distribution	Distribution. China: Fujian, Guangdong, Guangxi, Hainan, Hubei, Jiangsu, Jiangxi, Shanghai, Sichuan, Yunnan, Zhejiang; Cambodia; Indonesia; Malaysia; Philippines; Thailand; Vietnam.	en	Dong, Xue, Bian, Dongju (2021): Three new species and two new records of Helochares (Hydrobaticus) MacLeay 1871 from China (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4950 (1): 166-180, DOI: 10.11646/zootaxa.4950.1.9
