taxonID	type	description	language	source
D909CC267C02041A6D2DFEA0FD47202F.taxon	description	(Figs. 1 – 5, 17)	en	Miller, Kelly B., Gibson, James R., Alarie, Yves (2009): North American Stygobiontic Diving Beetles (Coleoptera: Dytiscidae: Hydroporinae) with Description of Ereboporus naturaconservatus Miller, Gibson and Alarie, New Genus and Species, from Texas, U. S. A. The Coleopterists Bulletin 63 (2): 191-202, DOI: 10.1649/1124.1, URL: http://dx.doi.org/10.1649/1124.1
D909CC267C04041D6C89FBADFCF8240F.taxon	description	(Figs. 1 – 5, 17)	en	Miller, Kelly B., Gibson, James R., Alarie, Yves (2009): North American Stygobiontic Diving Beetles (Coleoptera: Dytiscidae: Hydroporinae) with Description of Ereboporus naturaconservatus Miller, Gibson and Alarie, New Genus and Species, from Texas, U. S. A. The Coleopterists Bulletin 63 (2): 191-202, DOI: 10.1649/1124.1, URL: http://dx.doi.org/10.1649/1124.1
D909CC267C04041D6C89FBADFCF8240F.taxon	materials_examined	Type Locality. United States of America, Texas, Terrell County, Independence Creek Nature Conservancy Preserve, Caroline Springs.	en	Miller, Kelly B., Gibson, James R., Alarie, Yves (2009): North American Stygobiontic Diving Beetles (Coleoptera: Dytiscidae: Hydroporinae) with Description of Ereboporus naturaconservatus Miller, Gibson and Alarie, New Genus and Species, from Texas, U. S. A. The Coleopterists Bulletin 63 (2): 191-202, DOI: 10.1649/1124.1, URL: http://dx.doi.org/10.1649/1124.1
D909CC267C04041D6C89FBADFCF8240F.taxon	diagnosis	Diagnosis. This is the only member of the genus known and is characterized by the diagnostic combination unique to the genus (see above).	en	Miller, Kelly B., Gibson, James R., Alarie, Yves (2009): North American Stygobiontic Diving Beetles (Coleoptera: Dytiscidae: Hydroporinae) with Description of Ereboporus naturaconservatus Miller, Gibson and Alarie, New Genus and Species, from Texas, U. S. A. The Coleopterists Bulletin 63 (2): 191-202, DOI: 10.1649/1124.1, URL: http://dx.doi.org/10.1649/1124.1
D909CC267C04041D6C89FBADFCF8240F.taxon	description	Description. Measurements (in mm). TL 5 2.3 – 2.4; TL / GW 5 2.2 – 2.4; HL 5 0.7; HW 5 1.0; HL / HW 5 0.7; PL 5 0.4; PW 1 5 1.0 – 1.1; PW 2 5 0.8 – 0.9; PL / PW 1 5 0.4; PW 1 / PW 2 5 0.8 – 1.2; EW 5 0.9; EL 5 1.4; FL 5 0.6; RL 5 0.2; RL / FL 5 0.3. Habitus. Body elongate, greatest body width at head, dorso-ventrally depressed; body lightly sclerotized; coloration yellow over entire body. Sculpture and structure. Head (Figs. 1, 2) very large in proportion to body (L / W); eyes absent; anterior margin of clypeus concave, with continuous marginal bead; length of scape and pedicel each about 2 X width; length of flagellomeres I – VIII about 1.2 X width, apical flagellomere length about 2.5 X width. Pronotum (Figs. 1, 2) short and broad, greatest width anteriorly; anterolateral corners sharply pointed, extending anteriorly along lateral margins of head; submedially prominently constricted; posterolateral corners acute. Scutellum not externally visible. Elytron (Figs. 1, 2) elongate, elytra medially weakly fused along elytral suture, though not strongly so; lateral margin evenly and broadly curved anteriorly, elytra together slightly tapered posteriorly; elytral epipleuron very broad throughout length, extending medially, covering significant portions of lateral areas of metacoxae and abdominal sternites. Metathoracic wings absent. Head, pronotum, and elytron covered with microsculpture composed of fine, isodiametric cells making surface matte in appearance. Prosternum (Fig. 2) narrow; prosternal process very small, apically rounded, length about equal to width, not extending between mesocoxae to metasternum. Metacoxa (Fig. 2) large; metacoxal lines absent; apical lobes of posterior metacoxal process short and rounded; metacoxal bridge not differentiated (posterior margin of metacoxal process without delimiting margin), medially evenly sloped to abdomen; metatrochanter (Fig. 2) large, distinctly offset from metafemur, apex bluntly pointed, remote from metafemur; metafemur (Fig. 2) slender and elongate; metatibia (Fig. 2) slender, slightly and gradually expanded apically. Visible abdominal sternites II and III fused with suture absent medially (Fig. 2). Conspicuous sensory setae absent over entire body. Series of spinous setae present only on legs, but natatory setae entirely absent. Male genitalia. Median lobe in lateral aspect (Fig. 3) short, robust, broad, apex abruptly curved dorsad, apically broadly pointed; in ventral aspect (Fig. 4) broad, lateral margins evenly convergent to apex which is distinctly but shallowly emarginate. Lateral lobe (Fig. 4) relatively narrow, medially expanded, apex rounded. Female genitalia. Gonocoxosternite (Fig. 5) with posterior portion rhomboid, anterior portion elongate, moderately broad, slightly curved medially near anterior end; gonocoxa (Fig. 5) elongate, slender, medially bent; bursa copulatrix (Fig. 5) elongate, anteriorly broadly expanded, with elongate ring-like sclerotized structure on right side (receptacle); spermathecal duct (Fig. 5) elongate, slender; spermatheca (Fig. 5) robust, expanded on end near fertilization duct; fertilization duct (Fig. 5) elongate, slender.	en	Miller, Kelly B., Gibson, James R., Alarie, Yves (2009): North American Stygobiontic Diving Beetles (Coleoptera: Dytiscidae: Hydroporinae) with Description of Ereboporus naturaconservatus Miller, Gibson and Alarie, New Genus and Species, from Texas, U. S. A. The Coleopterists Bulletin 63 (2): 191-202, DOI: 10.1649/1124.1, URL: http://dx.doi.org/10.1649/1124.1
D909CC267C04041D6C89FBADFCF8240F.taxon	etymology	Etymology. The species name, naturaconservatus, comes from the Latin words natura, meaning ‘‘ nature’ ’ and conservatus, meaning ‘‘ keep’ ’ or ‘‘ conserve, ’’ in honor of The Nature Conservancy for their extensive work in the promotion of biodiversity and specifically in honor of the Independence Creek Nature Conservancy Preserve, the type locality for the species.	en	Miller, Kelly B., Gibson, James R., Alarie, Yves (2009): North American Stygobiontic Diving Beetles (Coleoptera: Dytiscidae: Hydroporinae) with Description of Ereboporus naturaconservatus Miller, Gibson and Alarie, New Genus and Species, from Texas, U. S. A. The Coleopterists Bulletin 63 (2): 191-202, DOI: 10.1649/1124.1, URL: http://dx.doi.org/10.1649/1124.1
D909CC267C04041D6C89FBADFCF8240F.taxon	distribution	Distribution and Habitat. Same as for the genus (see above).	en	Miller, Kelly B., Gibson, James R., Alarie, Yves (2009): North American Stygobiontic Diving Beetles (Coleoptera: Dytiscidae: Hydroporinae) with Description of Ereboporus naturaconservatus Miller, Gibson and Alarie, New Genus and Species, from Texas, U. S. A. The Coleopterists Bulletin 63 (2): 191-202, DOI: 10.1649/1124.1, URL: http://dx.doi.org/10.1649/1124.1
D909CC267C04041D6C89FBADFCF8240F.taxon	materials_examined	Material Examined. Holotype: Oi labeled, ‘‘ U. S. A.: Texas, Terrell Co., Independence Creek Preserve, Caroline Springs, 24.3 km S Sheffield, 30 ° 27 9 10 0 N 101 ° 44 9 10 0 W, 19 May 2007, J. R. Gibson and E. Chappell, colrs. / HOLOTYPE: Miller, Gibson and Alarie, 2008 [red label with double black line border]. ’’ Paratypes: 2 labeled same as holotype except ‘‘ / PARATYPE: Miller, Gibson and Alarie, 2008 [blue label with black line border]. ’’ Other subterranean Dytiscidae in North America	en	Miller, Kelly B., Gibson, James R., Alarie, Yves (2009): North American Stygobiontic Diving Beetles (Coleoptera: Dytiscidae: Hydroporinae) with Description of Ereboporus naturaconservatus Miller, Gibson and Alarie, New Genus and Species, from Texas, U. S. A. The Coleopterists Bulletin 63 (2): 191-202, DOI: 10.1649/1124.1, URL: http://dx.doi.org/10.1649/1124.1
D909CC267C05041F6D33FA39FEC0251F.taxon	description	(Figs. 6 – 8)	en	Miller, Kelly B., Gibson, James R., Alarie, Yves (2009): North American Stygobiontic Diving Beetles (Coleoptera: Dytiscidae: Hydroporinae) with Description of Ereboporus naturaconservatus Miller, Gibson and Alarie, New Genus and Species, from Texas, U. S. A. The Coleopterists Bulletin 63 (2): 191-202, DOI: 10.1649/1124.1, URL: http://dx.doi.org/10.1649/1124.1
D909CC267C05041F6D33FA39FEC0251F.taxon	type_taxon	Type Species. Comaldessus stygius Spangler and Barr, 1995 by original designation.	en	Miller, Kelly B., Gibson, James R., Alarie, Yves (2009): North American Stygobiontic Diving Beetles (Coleoptera: Dytiscidae: Hydroporinae) with Description of Ereboporus naturaconservatus Miller, Gibson and Alarie, New Genus and Species, from Texas, U. S. A. The Coleopterists Bulletin 63 (2): 191-202, DOI: 10.1649/1124.1, URL: http://dx.doi.org/10.1649/1124.1
D909CC267C05041F6D33FA39FEC0251F.taxon	materials_examined	Type Locality. U. S. A., Texas, Comal County, New Braunfels, Landa Park, Comal Spring # 3.	en	Miller, Kelly B., Gibson, James R., Alarie, Yves (2009): North American Stygobiontic Diving Beetles (Coleoptera: Dytiscidae: Hydroporinae) with Description of Ereboporus naturaconservatus Miller, Gibson and Alarie, New Genus and Species, from Texas, U. S. A. The Coleopterists Bulletin 63 (2): 191-202, DOI: 10.1649/1124.1, URL: http://dx.doi.org/10.1649/1124.1
D909CC267C05041F6D33FA39FEC0251F.taxon	diagnosis	Diagnosis. This species can be distinguished from other taxa within Bidessini using the following character combination: 1) eyes rudimentary, indicated by only a few ommatidia; 2) metacoxal lines absent; 3) occipital line absent (Fig. 6); 4) basal pronotal and basal elytral striae (‘‘ plicae’ ’) present (Fig. 6).	en	Miller, Kelly B., Gibson, James R., Alarie, Yves (2009): North American Stygobiontic Diving Beetles (Coleoptera: Dytiscidae: Hydroporinae) with Description of Ereboporus naturaconservatus Miller, Gibson and Alarie, New Genus and Species, from Texas, U. S. A. The Coleopterists Bulletin 63 (2): 191-202, DOI: 10.1649/1124.1, URL: http://dx.doi.org/10.1649/1124.1
D909CC267C05041F6D33FA39FEC0251F.taxon	discussion	Discussion. Comaldessus stygius was described from twelve specimens collected in drift nets from Comal Springs (Spangler and Barr 1995), a system of artesian springs. Relationships. This species is clearly a member of the tribe Bidessini based on the 2 - segmented parameres (Biström 1988) and the presence of basal pronotal and elytral striae (Fig. 6) which are characteristic of many Bidessini (Biström 1988). Spangler and Barr (1995) speculated that the species is related to Trogloguignotus Sanfilippo and Uvarus Guignot and that both Trogloguignotus and Comaldessus are derived from within the species-rich, plesiomorphic Uvarus, though without a cladistic analysis of this problematic tribe, it is difficult to make definitive conclusions.	en	Miller, Kelly B., Gibson, James R., Alarie, Yves (2009): North American Stygobiontic Diving Beetles (Coleoptera: Dytiscidae: Hydroporinae) with Description of Ereboporus naturaconservatus Miller, Gibson and Alarie, New Genus and Species, from Texas, U. S. A. The Coleopterists Bulletin 63 (2): 191-202, DOI: 10.1649/1124.1, URL: http://dx.doi.org/10.1649/1124.1
D909CC267C0704106D4BFB2AFE032301.taxon	description	(Figs. 9 – 12)	en	Miller, Kelly B., Gibson, James R., Alarie, Yves (2009): North American Stygobiontic Diving Beetles (Coleoptera: Dytiscidae: Hydroporinae) with Description of Ereboporus naturaconservatus Miller, Gibson and Alarie, New Genus and Species, from Texas, U. S. A. The Coleopterists Bulletin 63 (2): 191-202, DOI: 10.1649/1124.1, URL: http://dx.doi.org/10.1649/1124.1
D909CC267C0704106D4BFB2AFE032301.taxon	type_taxon	Type Species. Stygoporus oregonenesis Larson and Labonte, 1994 by original designation.	en	Miller, Kelly B., Gibson, James R., Alarie, Yves (2009): North American Stygobiontic Diving Beetles (Coleoptera: Dytiscidae: Hydroporinae) with Description of Ereboporus naturaconservatus Miller, Gibson and Alarie, New Genus and Species, from Texas, U. S. A. The Coleopterists Bulletin 63 (2): 191-202, DOI: 10.1649/1124.1, URL: http://dx.doi.org/10.1649/1124.1
D909CC267C0704106D4BFB2AFE032301.taxon	materials_examined	Type Locality. U. S. A., Oregon, Polk County, 7 km NNE Dallas, Smithfield Road, 44.973 ° N, 123.275 ° W, 90 m, well.	en	Miller, Kelly B., Gibson, James R., Alarie, Yves (2009): North American Stygobiontic Diving Beetles (Coleoptera: Dytiscidae: Hydroporinae) with Description of Ereboporus naturaconservatus Miller, Gibson and Alarie, New Genus and Species, from Texas, U. S. A. The Coleopterists Bulletin 63 (2): 191-202, DOI: 10.1649/1124.1, URL: http://dx.doi.org/10.1649/1124.1
D909CC267C0704106D4BFB2AFE032301.taxon	diagnosis	Diagnosis. This species is unique within Hydroporini in having the following combination of characters: 1) compound eyes absent; 2) prosternal process inserted dorsad of the anterior margin of the metasternum; 3) with longitudinal impressions laterally on the pronotum (Fig. 9); 4) metacoxal processes together truncate posteriorly; 5) with natatory setae on the metatarsus and metatibia.	en	Miller, Kelly B., Gibson, James R., Alarie, Yves (2009): North American Stygobiontic Diving Beetles (Coleoptera: Dytiscidae: Hydroporinae) with Description of Ereboporus naturaconservatus Miller, Gibson and Alarie, New Genus and Species, from Texas, U. S. A. The Coleopterists Bulletin 63 (2): 191-202, DOI: 10.1649/1124.1, URL: http://dx.doi.org/10.1649/1124.1
D909CC267C0704106D4BFB2AFE032301.taxon	discussion	Discussion. This species was described from nine specimens collected from a shallow well. The owners of the well, thinking the beetle may be a pest, chlorinated the well and the species has not been seen since (Larson and Labonte 1994). Larson and Labonte (1994) suggested that the presence of specimens in this particular well (an old well with no previous indication of a diving beetle fauna) may have been the result of an unusually large amount of precipitation that year in the region, possibly resulting in a change in the nature of the aquifer and temporarily allowing beetles into the well. Relationships. Larson and Labonte (1994) speculated that Stygoporus is the sister group to Sanfilippodytes Franciscolo based on the enlarged metatrochanters and produced posterior margin of the medial portions of the metacoxa. Sanfilippodytes occur in seeps and springs, and seem to be a likely candidate for occurrence in underground waters. One species, S. sbordonii Franciscolo, was described from a cave in Mexico (Franciscolo 1979). Although not anophthalmic like Stygoporus, S. sbordonii does have smaller eyes than typical Sanfilippodytes (Larson and Labonte 1994).	en	Miller, Kelly B., Gibson, James R., Alarie, Yves (2009): North American Stygobiontic Diving Beetles (Coleoptera: Dytiscidae: Hydroporinae) with Description of Ereboporus naturaconservatus Miller, Gibson and Alarie, New Genus and Species, from Texas, U. S. A. The Coleopterists Bulletin 63 (2): 191-202, DOI: 10.1649/1124.1, URL: http://dx.doi.org/10.1649/1124.1
D909CC267C0804106D27FD3CFE272427.taxon	description	(Figs. 13 – 16)	en	Miller, Kelly B., Gibson, James R., Alarie, Yves (2009): North American Stygobiontic Diving Beetles (Coleoptera: Dytiscidae: Hydroporinae) with Description of Ereboporus naturaconservatus Miller, Gibson and Alarie, New Genus and Species, from Texas, U. S. A. The Coleopterists Bulletin 63 (2): 191-202, DOI: 10.1649/1124.1, URL: http://dx.doi.org/10.1649/1124.1
D909CC267C0804106D27FD3CFE272427.taxon	type_taxon	Type Species. Haideoporus texanus Young and Longley, 1976 by original designation.	en	Miller, Kelly B., Gibson, James R., Alarie, Yves (2009): North American Stygobiontic Diving Beetles (Coleoptera: Dytiscidae: Hydroporinae) with Description of Ereboporus naturaconservatus Miller, Gibson and Alarie, New Genus and Species, from Texas, U. S. A. The Coleopterists Bulletin 63 (2): 191-202, DOI: 10.1649/1124.1, URL: http://dx.doi.org/10.1649/1124.1
D909CC267C0804106D27FD3CFE272427.taxon	materials_examined	Type Locality. U. S. A., Texas, Hays County, San Marcos, Southwest Texas State University, Aquatic Station, artesian well.	en	Miller, Kelly B., Gibson, James R., Alarie, Yves (2009): North American Stygobiontic Diving Beetles (Coleoptera: Dytiscidae: Hydroporinae) with Description of Ereboporus naturaconservatus Miller, Gibson and Alarie, New Genus and Species, from Texas, U. S. A. The Coleopterists Bulletin 63 (2): 191-202, DOI: 10.1649/1124.1, URL: http://dx.doi.org/10.1649/1124.1
D909CC267C0804106D27FD3CFE272427.taxon	diagnosis	Diagnosis. Haideoporus texanus can be distinguished from other Hydroporini by the following: 1) eyes small, located anterolaterally on head; 2) setae of body and legs extremely long; 3) pro- and mesocoxae large, globular; 4) prosternal process not extending posteriorly between mesothoracic legs to mesosternum.	en	Miller, Kelly B., Gibson, James R., Alarie, Yves (2009): North American Stygobiontic Diving Beetles (Coleoptera: Dytiscidae: Hydroporinae) with Description of Ereboporus naturaconservatus Miller, Gibson and Alarie, New Genus and Species, from Texas, U. S. A. The Coleopterists Bulletin 63 (2): 191-202, DOI: 10.1649/1124.1, URL: http://dx.doi.org/10.1649/1124.1
D909CC267C0804106D27FD3CFE272427.taxon	discussion	Discussion. This species was discovered in a well on the campus of Texas State University in San Marcos, Hays County, Texas (Young and Longley 1976) and subsequently also found at Comal Spring, Comal County, Texas (Bowles and Stanford 1997), the type locality for C. stygius (see above). The larva of Haideoporus has been described (Longley and Spangler 1977), making it the only subterranean dytiscid larva from North America described to date. Relationships. The species is currently placed in the Hydroporini (Nilsson 2001), but Young and Longley (1976) were cautious to make definitive statements about the relationships of this species to others in its subfamily. They pointed out close similarities of Haideoporus with other subterranean diving beetles, such as Morimotoa and Siettitia, but suspected that these similarities were due to convergence in similar habitats.	en	Miller, Kelly B., Gibson, James R., Alarie, Yves (2009): North American Stygobiontic Diving Beetles (Coleoptera: Dytiscidae: Hydroporinae) with Description of Ereboporus naturaconservatus Miller, Gibson and Alarie, New Genus and Species, from Texas, U. S. A. The Coleopterists Bulletin 63 (2): 191-202, DOI: 10.1649/1124.1, URL: http://dx.doi.org/10.1649/1124.1
