taxonID	type	description	language	source
99E47CE891444F8770E69F4BBA8EE4ED.taxon	description	Description. Size range: Length 1.7 - 2.0 mm; width 1.2 - 1.5 mm; Body: rufescent to rufobrunneus, somewhat narrowly elongate, widest at humeri, abruptly narrowed anteriorly at pronotal middle, with exaggerated sculpturing throughout. Head: head deflexed relative to anterior pronotal margin; frons flat, sides weakly rounded, longitudinally strigose, with fine setigerous punctures between strigae, setae minute; frontal stria complete along margin of eye and across front, prominent, descending onto epistoma anteriorly, subangulate at middle; supraorbital stria absent; epistoma narrowed to front, apex rather narrowly emarginate, with lateral striae meeting frontal stria, convergent, nearly meeting anterad; labrum small, about twice as wide as long, apex weakly emarginate; mandibles with incisor edges evenly curved to apex, basal teeth inconspicuous; submentum broadly triangular, weakly produced into base of oral cavity, sparsely setose; mentum subtrapezoidal, apex weakly sinuate; labial palpifers prominent; labial palps 3 - segmented, with basal palpomere very short, apical palpomere widest near base, subacute; maxillary cardines short, semicircular, glabrous, stipes with two setae along lateral margin; maxillary palpi 4 - segmented with basal palpomere very short, palpomeres 2 and 3 similar in length and breadth, ultimate palpomere about twice as long as penultimate, widest near base, narrowed apically; antennal scape stout, anterior surface becoming longitudinally carinate in apical half, with few apical setae; funicle widening slightly to short, disclike 8 th antennomere; antennal club about 2.5 x as long as wide, densely setose, with indistinct, denser subapical setose sensory patches on dorsal and ventral surfaces. Pronotum: widest near base, sides sinuate, strongly narrowed anterad midpoint, basal margin uneven; lateral marginal pronotal stria complete around lateral and anterior margins, though strongly sinuate at sides, submarginal stria present along sides, not parallel to marginal, joining it near anterior corner; pronotal disk with prominent dorsal process arising from entire anterior margin, narrowing and arcing more or less evenly to middle of posterior margin, sides of process longitudinally creased to deeply invaginated, dorsal surface of process coarsely reticulostrigose, sides more or less smooth; pronotal gland openings, if present, obscured by sculpturing, possibly incorporated into lateral invaginations of pronotal process. Elytra: elytron with striation generally carinate and exaggerated; epipleuron with complete marginal stria and additional stria along upper edge, paralleling outer subhumeral stria, continuing around elytral apex, variably meeting apices of dorsal striae; outer subhumeral, inner subhumeral, and dorsal striae 1 - 4 all complete, increasingly more strongly impressed toward suture, apices meeting apical marginal elytral suture; elytral intervals smooth to strongly microsculptured. Prosternum: prosternal keel rather narrow, base weakly produced, carinal striae complete, subparallel, united basally, meeting presternal suture anteriorly, which varies from indistinct to deeply impressed; lateral striae diverging to sides, delimiting anterior leg depression; prosternal lobe extremely reduced, no longer at midline than at sides, marginal stria obsolete. Mesoventrite: mesoventrite short, shallowly emarginate at middle, with complete marginal stria; mesometaventral stria paralleling or diverging anterad from mesometaventral suture at middle. Metaventrite: postmesocoxal and lateral metaventral striae parallel, arching toward metacoxa then anterad to metepisternum; metaventral disk weakly depressed at middle. Abdomen: 1 st abdominal ventrite with anterior marginal stria continued to posterior margin by lateral striae, disk rather simply and finely punctate; propygidium slightly to distinctly wider than long, rather strongly convex, sparsely to densely reticulostrigose; propygidium apparently with single pair of gland openings very close to anterior corners (obscured by sculpturing in most species); pygidium longer than basal width, sculptured as propygidium, generally smoother apicomedially. Legs: each trochanter with single seta; profemur subparallel-sided to expanded at middle of anterior margin, with anterior marginal stria delimiting microsculptured marginal area; protibiae widened from base, sides subparallel to slightly narrowing, bearing 3 - 5 marginal setae in apical half; protarsal groove very weakly developed; meso- and metafemora rather large, produced beyond epipleurae in repose, broad, variously widened along posterior margins; meso- and metatibiae long, widened apically, bearing 3 - 4 longitudinal striae on anterior surfaces; mesotibia only bearing 2 - 4 characteristic long, thin subapical setae, at least one of which is inserted on the posterior surface; all tarsi laterally compressed, bearing simple ventral setae, with relatively large weakly curved claws. Male genitalia (Fig. 4): Paired accessory sclerites present; T 8 with broad basal and narrower apical emarginations, line of basal membrane attachment complete, just distad basal emargination, ventral apodemes well developed, nearly meeting along midline; S 8 with halves separated, apical guides moderately and evenly developed from base to apex, each apex with single prominent seta; 9 th tergite with very weak ventrolateral apodemes, apices narrow, subacute; T 10 completely divided; S 9 broad at base, narrowest near apex, head broad, subquadrate apically, with small apical emargination, more strongly sclerotized along midline; tegmen broad basally, strongly narrowed to apex, narrowly divided apically, median foramen basad apical narrowing, moderately to strongly curved ventrad in apical half; basal piece about one-third tegmen length; median lobe short, simple, from one-fourth to one-third tegmen length. Female genitalia: T 8 forming a single plate, apically emarginate; S 8 tripartite, basal baculi convergent proximally; S 9 elongate, articulated with strap-shaped extension from apex of S 8; T 10 entire; overall ovipositor rather short; valvifers paddle-shaped, paddles nearly one-half total length; coxites strong, slightly longer than broad, two-thirds length of valvifers, strongly bidentate, with strengthening ridge on inner face; gonostyle present; bursa copulatrix membraneous, weakly expanded; spermatheca gradually expanded, apically bulbous, with slightly expanded spermathecal gland attached near its base.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
99E47CE891444F8770E69F4BBA8EE4ED.taxon	diagnosis	Diagnosis. This highly distinctive genus scarcely needs a diagnosis. Its prominent pronotal process (Figs 1, 2) is unique and unmistakable. Its assignment perhaps to tribe could be problematic, as it exhibits no hint of an emargination of the prosternal keel (Fig. 3 A-B). However, both male and female genitalia, as well as DNA sequences place it unambiguously as deeply nested within the neotropical Exosternini.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
99E47CE891444F8770E69F4BBA8EE4ED.taxon	etymology	Etymology. The genus name means ' conical helmet' referring to the anterior process of the pronotum. The gender of the name is feminine.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
7E8540C7D11E693167DA193AD72B3311.taxon	etymology	Etymology. This species name refers to the fact that it is the smallest known member of the genus.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
DB073FE231DAE6B041DB8159C8B87F07.taxon	etymology	Etymology. This species is named for the camel-like hump on the pronotum, from the specific name of the one-humped dromedary.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
C665FBD8D647CAAD6E4301434A5677AB.taxon	etymology	Etymology. This species is named for the three distinct lateral pronotal creases.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
CE40E1BAA79332A0867E1916063E9535.taxon	etymology	Etymology. This species is named for the fact that it has the deepest pronotal invaginations in the genus.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
7293B4716D59666B2DD410C6954F9C0E.taxon	description	Description. This genus differs from other Exosternini in the following combination of characters: body widest behind middle, subdepressed, somewhat flattened dorsally, metaventrite rather abruptly convex ventrally, glabrous, lacking secondary punctation, and almost entirely lacking typical striae; frons more or less coplanar with vertex; labrum broad, apically emarginate, lateral margins with conspicuous setal fringe; mandibles with short incisor edges; antennal scape elongate, slender, curving posteroventrad beneath eye; funicle about as long as scape, weakly widened distally, the 8 th antennomere cupuliform, no shorter than preceding antennomeres; antennal club elongate oval, completely tomentose, with two nearly complete setose annuli more distinctly interrupted on dorsal surface, only slightly curved basad at middle; eye substantially reduced; pronotum lacking prescutellar impression, though fine prescutellar fovea may be present, with three median gland openings on each side, one in anterior angles, one behind eye about one funicle width behind anterior margin, one displaced posterad on disk; three distinct gland openings also present along lateral pronotal margin; elytra with few striae extremely fine, inconspicuous; prosternal keel depressed, rather broad, produced at base; prosternal lobe deflexed; mesoventrite anteriorly emarginate; metaventrite strongly convex posteriorly, posterior margin arcuate; propygidium transverse, with single gland openings in anterolateral corners; pygidium large, apically rounded, without gland openings or marginal striae; trochanter with 2 - 3 short apical setae; femora moderately broad, slightly flattened; protibia subtriangular, with outer edge weakly outwardly rounded, not distinctly toothed, with numerous distinct marginal spines; two small protibial spurs present; all tarsi rather long and curving, with numerous ventral spines, those of protarsus (of male only?) strongly spatulate; mesotibia moderately expanded toward apex, metatibia less so, both with nearly complete series of marginal spines; male genitalia (Fig. 6) with paired accessory sclerites present; apices of S 8 bearing only very fine, inconspicuous setae; T 10 completely divided; S 9 with head broad, subquadrate, with complete apical flange; tegmen narrow and elongate, sides parallel in basal half, narrowed to thin, ventrally curved apex, median foramen basad apical narrowing, ventral surface with basal tooth formed by thin median keel; Female not known.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
7293B4716D59666B2DD410C6954F9C0E.taxon	etymology	Etymology. From the Greek, literally ' pregnant body', referring to the strongly convex metaventrite.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
BF8FAC38D9E13629EB5443211413A5C2.taxon	description	Description. Size range: Length 3.0 mm; width 2.7 mm; Body: body elongate ovoid, widest behind middle, subdepressed, dorsally somewhat flattened, more convex ventrally, dark rufescent, glabrous, with conspicuous ground punctation throughout, lacking secondary punctation, and almost entirely lacking typical striae. Head: frons elongate, largely coplanar with vertex, weakly depressed at middle; frontal and supraorbital striae absent; epistoma wide, somewhat deflexed, weakly emarginate apically; labrum large, about one-third as long as wide, apical margin deeply emarginate, lateral margins with conspicuous setal fringe; mandibles with short incisor edges, left bearing prominent basal tooth, right mandible with weaker basal tooth; submentum wide, slightly depressed relative to surrounding genae, anterior margin weakly outwardly arcuate; mentum trapezoidal, narrowed anteriorly, apical margin more or less entire; labial palps three segmented, basal segment short, penultimate and ultimate palopmeres elongate; submentum and mentum bearing numerous elongate setae; maxillary cardo shining, glabrous, stipes microsculptured and with numerous long setae; maxillary palp four segmented, rather stout; antennal scape elongate, not markedly widened apically, curving posteroventrad beneath somewhat reduced compound eye; funicle about as long as scape, weakly widened distally, the 8 th antennomere cupuliform, no shorter than preceding antennomeres; antennal club elongate oval, completely tomentose, with two nearly complete setose annuli more distinctly interrupted on dorsal surface, only slightly curved basad at middle. Pronotum: pronotum weakly convex, sides faintly arcuate from base to apices; prescutellar impression absent, though fine prescutellar fovea may be present; pronotal disk with three median gland openings on each side, one in anterior angles, close to corner, one behind eye about one funicle width behind anterior margin, one directly posterad this one, just behind pronotal midpoint; three distinct gland openings also present along lateral pronotal margin; marginal stria complete along lateral and anterior margins; fragments of lateral submarginal stria visible near anterior corners. Elytra: elytra with few striae extremely fine, inconspicuous; epipleuron with single complete marginal stria and fragments of additional epipleural striae detectable, oblique humeral stria visible near base, 1 st dorsal stria finely impressed in apical two-thirds, 2 nd stria visible in basal half, slightly abbreviated from base, 3 rd stria present in basal third, 4 th, 5 th and sutural striae entirely absent; elytral disk with very fine granular microsculpture faintly impressed toward apices. Prosternum: prosternal keel depressed, rather broad, produced at base, lacking striae; prosternal lobe about three-fourths length of keel, deflexed, lacking marginal stria; both prosternal keel and lobe rather densely microsculptured. Mesoventrite: mesoventrite broad, subquadrate, anterior margin emarginate, marginal stria faintly impressed. Metaventrite: metaventrite strongly convex posteriorly, posterior margin arcuate; mesometaventral, postcoxal and inner lateral metaventral striae absent, metaventral disk smooth at middle, more distinctly microsculptured at sides. Abdomen: abdominal ventrites 1 - 4 lacking striae, more or less distinctly microsculptured throughout; propygidium short, transverse, with single gland openings in anterolateral corners; pygidium large, apically rounded; propygidium and pygidium with very fine ground punctation and transverse waves of microsculpture. Legs: each trochanter with 2 - 3 short apical setae; femora moderately broad, slightly flattened; protibia subtriangular, with outer edge weakly outwardly rounded, not distinctly toothed, but set with about 9 distinct marginal spines; two small protibial spurs present; all tarsi rather long and curving, with numerous ventral spines, those of protarsus (of male only?) strongly spatulate; mesotibia moderately expanded toward apex, metatibia much less so, both with nearly complete series of marginal spines. Male genitalia (Fig. 6): Paired accessory sclerites present; T 8 with broad basal and narrower apical emarginations, line of basal membrane attachment complete distad basal emargination, ventral apodemes well developed, narrowing ventrally, slightly separated along midline; S 8 with halves approximate at base, apical guides widening from base to apex, apices narrowly rounded, bearing only very fine, inconspicuous setae; T 9 with base of dorsal flaps rather protuberant above proximal apodemes, ventrolateral apodemes well developed, T 9 apices narrow, convergent subacute; T 10 elongate, completely divided; S 9 with sides subparallel in basal half, narrowest distad midpoint, head broad, subquadrate apically, with complete apical flange, apex not emarginate; tegmen narrow and elongate, sides parallel in basal half, narrowed to thin, ventrally curved apex, median foramen basad apical narrowing, ventral surface with basal tooth formed by thin median keel; basal piece nearly one-half tegmen length; median lobe about one-third tegmen length, proximal arms thinned basally. Female: not known.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
BF8FAC38D9E13629EB5443211413A5C2.taxon	etymology	Etymology. We name this species for Mr. Marvin Rockwell, one of the original Quaker founders of the Costa Rican community of Monteverde, who has helped many visitors (the senior author included) better appreciate the biodiversity of Costa Rica.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
5BF33E80EAF77A2E3D06D9F2B52CBEC6.taxon	description	Description. This genus differs from other Exosternini in the following combination of characters: body round, strongly convex, with strong elytral ridges, rufescent, glabrous; frons and epistoma depressed along midline, frontal stria present; labrum wide, shallowly emarginate apically; mandibles with weakly arcuate incisor edges lacking basal teeth; antennal scape slightly wider near base, narrowed apically, with longitudinal carina along inner anterior edge; funicle shorter than scape, weakly widening apically, antennomere 8 short, cupuliform, not disc-like; antennal club slightly elongate oval, largely tomentose, with only indistinct subapical sensory patches; pronotum rather strongly convex, with two gland openings on each side, anterior opening simple, along anterior margin behind eye, posterior opening with secondary annulus, displaced posterad to near middle of disk; prescutellar impression absent; elytra with strong longitudinal ridges, apparently corresponding to alternate interstriae; prosternal keel emarginate at base; prosternal lobe short, apically truncate; mesoventrite narrowly produced at middle; propygidium slightly shorter than pygidium along midline, with gland openings near anterolateral corners; pygidium equilaterally subtriangular, apex rounded, simple; each trochanter with single long seta; femora narrow; protibia rounded apically, lacking marginal teeth, with marginal spines inserted only along apical half of edge; protibial spurs present, reduced; meso- and metatibiae thin, simple, with single longitudinal stria along inner edge, completely lacking teeth or spines along outer margin; all tarsi slightly compressed, with slightly spatulate ventral setae; male not known; female T 8 divided; S 8 forming a single plate, basal baculi articulated with basolateral corners, convergent, separate at base; S 9 not evident, though elongate articulating strap from median apex of S 8 is present; T 10 not observed; valvifers paddle-shaped, basal paddles just over one-third entire valvifer length; coxites over two-thirds valvifer length, apically tridentate, with lateral teeth rather weak; gonostyle slightly shorter than median tooth, setose; bursa copulatrix membraneous, lacking sclerites, not obviously expanded; spermatheca forming a gradually enlarged, elongate sac, an elongate, slightly kinked spermathecal gland inserted near its midpoint.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
5BF33E80EAF77A2E3D06D9F2B52CBEC6.taxon	etymology	Etymology. The name of this genus refers to the series of ridges on the elytra; feminine.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
9EC4BF79ADDC1627360EB7F7E3FB4082.taxon	description	Description. Size range: Length 1.7 mm; width 1.5 mm; Body: body round, strongly convex, with strong elytral ridges, rufescent, glabrous. Head: frons slightly longer than wide, depressed along midline, smooth, with complete frontal stria more or less rounded across front; epistoma depressed along midline, weakly emarginate apically; labrum about 4 x as wide as median length, shallowly emarginate apically; mandibles with weakly arcuate incisor edges lacking basal teeth; antennal scape slightly wider near base, narrowed apically, with longitudinal carina along inner anterior edge; funicle shorter than scape, weakly widening apically, antennomere 8 short, cupuliform, not disc-like; antennal club slightly elongate oval, largely tomentose, with only indistinct subapical sensory patches. Pronotum: pronotum rather strongly convex, sides narrowed evenly from base to apex, only faintly sinuate at base and middle; marginal pronotal stria complete along lateral and apical margins; sublateral stria present along entire lateral margin, just curving mediad anteriorly, pronotal disk shallowly depressed along its inner edge; pronotal disk with two gland openings on each side, anterior opening simple, along anterior margin behind eye, posterior opening with secondary annulus, situated directly posterad anterior opening, just in front of midline; prescutellar impression absent; posterior margin of disk simple. Elytra: elytra dominated by several strong longitudinal ridges, epipleuron with single submarginal stria, continued along apical margin to apex of 2 nd dorsal stria; outer subhumeral stria complete; other dorsal striae very fine, running near upper edge of elevated ridges, clearly visible only near apices; apices of 3 rd- 4 th, and 5 th-sutural striae joined along posterior margin. Prosternum: prosternal keel shallowly but subacutely emarginate at base, carinal striae obsolete basally, joined by anterior arch short of presternal suture; prosternal lobe less than half as long as keel, apically truncate, lacking marginal stria. Mesoventrite: mesoventrite narrowly produced at middle, with complete marginal stria, disk shallowly depressed behind. Metaventrite: mesometaventral stria well impressed, coincident with mesometaventral suture; postcoxal stria directed laterad toward middle of metepisternum, ending freely; lateral metaventral stria running obliquely toward outer third of metacoxa, slightly abbreviated apically. Abdomen: 1 st abdominal ventrite with complete anterior marginal stria continued by postmetacoxal stria which curves laterad behind coxa, ending freely; ventrites 2 - 4 impunctate; propygidium only slightly shorter than pygidium along midline, smooth, with inconspicuous gland openings near anterolateral corners; pygidium similar in basal width and midline length, apex rounded, simple. Legs: each trochanter with single long seta; femora rather narrow, metafemur particularly elongate; protibia gradually widened to rounded apical half, lacking marginal teeth, with marginal spines inserted only along apical half of edge; protibial spurs present, slightly reduced; meso- and metatibiae thin, simple, with single longitudinal stria along inner edge, completely lacking teeth or spines along outer margin; all tarsi slightly compressed, with slightly spatulate ventral setae. Male: not known.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
9EC4BF79ADDC1627360EB7F7E3FB4082.taxon	etymology	Etymology. The name of this species refers to its superficial resemblance to the histerid genus Onthophilus Leach, owing to the parallel ridges on the elytra.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
67D712F1F11C0E0A23F497F3A3384183.taxon	description	Description. Size range: Length 2.4 - 2.8 mm; width 1.6 - 1.9 mm; Body: body elongate, subcylindrical, sides parallel, rufobrunneus, variably punctate, glabrous. Head: frons weakly to strongly depressed at middle, subangulate at sides in front of eyes, frontal stria complete to strongly reduced, supraorbital stria present, detached; epistoma and labrum varied; mandibles strongly toothed; mouthparts rather strongly recessed in oral cavity; submentum flat, produced in front; mentum subquadrate, nearly as long as broad, bearing sparse long setae; ultimate labial palpomeres elongate, slightly compressed; cardo glabrous, stipes with few long setae on lateral margin; ultimate maxillary palpomere slightly compressed; antennal scape elongate, curved, widest near midpoint; funicle shorter than scape, widening from antennomere 4 - 8, 8 th antennomere cupuliform, more or less disclike; antennal club short, tomentose, with single, slightly elongate, axial sensory patch on ventral surface and longer patch on dorsal surface. Pronotum: pronotal sides more or less straight, slightly convergent to apex; prescutellar impression faintly impressed to obsolete; pronotal disk with two median gland openings on each side, one very close to margin behind eye, one posterad just behind pronotal midpoint; marginal pronotal stria present on lateral and anterior margins, may be complete or interrupted at sides; submarginal pronotal stria may be present at sides. Elytra: elytra slightly depressed along suture; epipleuron with single, complete marginal stria, outer subhumeral stria interrupted at middle, inner subhumeral stria absent, striae 1 - 5 present, 1 and 5 may be abbreviated, sutural stria complete. Prosternum: prosternal keel emarginate at base, carinal striae present, more or less complete, convergent anterad, joined in anterior arch; prosternal lobe short, slightly deflexed, marginal stria present. Mesoventrite: mesoventrite wide, short, with marginal stria fine, close to edge, may be interrupted; mesometaventral stria strongly angulate forward nearly to margin. Metaventrite: postmesocoxal stria poorly developed, short; lateral metaventral stria extending from inner corner of mesocoxa toward middle of metacoxa, abbreviated or not. Abdomen: 1 st abdominal ventrite with single, oblique lateral stria; propygidium slightly convex, about two-thirds length of pygidium, gland openings may be visible near anterolateral corners; pygidium moderately to strongly convex, apical margin rounded. Legs: trochanters with single long seta; femora moderately elongate; protibia with outer margin rounded, moderately strongly dentate, with 5 - 6 spinose teeth; two protibial spurs present, meso- and metatibiae narrow to moderately widened to apex, mesotibia with entire margin spinose, metatibia spinose toward apex; protarsi (of male only?) with spatulate ventral setae. Male genitalia: accessory sclerites absent; T 8 with narrow subacute apical emargination, ventral apodemes well developed but separated beneath, basal membrane attachment line intersecting slightly sinuate basal emargination; S 8 with halves approximate in basal half, apical guides increasingly wide toward apex, rather abruptly narrowed to narrowly rounded apices, with only very fine inconspicuous setae; T 9 with basal apodemes short, apex narrowly subacute, ventrolateral apodeme weakly hooked; T 10 undivided; S 9 desclerotized along midline, with deep apical emargination, apical flange interrupted, apical corners produced; tegmen with sides subparallel in basal two-thirds, slightly widened to subquadrate, subtruncate apex, lacking medioventral process; median lobe simple, about one-third tegmen length; basal piece short, with strong apicoventral articulating process. Female genitalia: T 8 entire, with narrow apical emargination; S 8 undivided, only emarginate apicolaterally, basal baculi strongly, arcuately convergent at base; S 9 present, short, connected to S 8 by sclerotized strap; T 10 entire; valvifers gradually expanded to base, paddles about half total length; coxites large, about two-thirds valvifer length, strongly bidentate; gonostyle slightly shorter than median tooth, setose; bursa copulatrix entirely membraneous, weakly expanded; spermatheca globose; spermathecal gland not evident in preparation.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
67D712F1F11C0E0A23F497F3A3384183.taxon	etymology	Etymology. Pyxis = canister or casket, referring loosely to the cylindrical body form, masculine.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
2FAB7E4F6FEEDBDBF0F7F5EBBFFBBB45.taxon	etymology	Etymology. The name of this species means ' devourer', alluding to its strong mandibles.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
62CCB624A4354C9EFB10CD7CCDDE119B.taxon	etymology	Etymology. This species' name refers to its very distinctive, strongly convex labrum.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
0B5F0A12A22621CF37127D614CD801A6.taxon	description	Description. This genus differs from other Exosternini in the following combination of characters: body elongate, cylindrical, parallel-sided, glabrous; frons subangulate in front of eyes, moderately produced above antennal insertions, weakly depressed at middle; epistoma convex, apex truncate; labrum about twice as wide as long; left mandible with outwardly arcuate incisor edge, right mandible with small acute basal tooth; submentum slightly depressed, outwardly arcuate along anterior margin; mentum subtrapezoidal, very narrowly emarginate apically, sparsely setose; cardo glabrous, smooth; stipes with few long setae; all palpi relatively stout, ultimate palpomeres, particularly maxillary palps, thickened and with numerous conspicuous punctures; antennal scape elongate, sides sinuate, narrowed at middle in anterior aspect; funicle gradually but slightly widened to apex, 8 th antennomere as long as preceding antennomeres; antennal club small, about as long as preceding 4 antennomeres, lacking complete annuli, sensoria poorly defined, apparently with two widely interrupted annuli close to dorsal apex; pronotal disk with only single distinct pair of gland openings, present between anterior margin and anterior submarginal stria; prescutellar impression absent; prosternal keel very shallowly emarginate at base; prosternal lobe slightly deflexed; anterior margin of mesoventrite broadly emarginate, but very weakly produced at center; propygidium rather long, more or less flat, with small gland openings near anterolateral corners; pygidium slightly longer than propygidium along midline, apical margin simple, rounded; protrochanter glabrous, meso- and metatrochanters each with two very short apical setae; femora simple; protibia with outer margin rounded, weakly dentate, strongly spinose; protibial spurs very short; protarsus with ventral setae simple; meso- and metatibiae elongate, outer margins with spinose, mesotibia faintly dentate; meso- and metatarsi short, each with single pair apicoventral spines; male genitalia with accessory sclerites present; T 8 rather short; S 8 halves approximate only at base, inner margins divergent to apex, apically with very fine, inconspicuous setae; T 9 with prominent, strongly hooked ventrolateral apodemes, dorsal lobes more strongly sclerotized along sides; T 10 divided; S 9 with apical emargination distinct, apical flange interrupted; tegmen with very large medioventral process; median lobe simple; female not known.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
0B5F0A12A22621CF37127D614CD801A6.taxon	etymology	Etymology. The name of this genus is from the Greek, meaning ' pill-shaped'; feminine.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
9F7C2D82177CFB31E2612E0AB80BB9E3.taxon	description	Description. Size range: Length 2.2 mm; width 1.3 mm; Body: body elongate, cylindrical, parallel-sided, rufobrunneus, ground punctation rather inconspicuous, glabrous. Head: frons rather broad, subangulate at sides in front of eyes, moderately produced in front above antennal insertions, beneath complete, subangulate frontal stria, weakly depressed at middle, with sparse ground punctures and very faint microsculpture within frontal depression; supraorbital stria weak but complete, narrowly detached from frontal stria at sides; epistoma convex, apex truncate; labrum about twice as wide as long, apex arcuate, but dorsal surface increasingly depressed to apex, making it appear somewhat emarginate; mandibles coarsely punctate on sides. Pronotum: pronotal sides subparallel in basal half, slightly arcuately narrowing to apex; marginal pronotal stria present on lateral and anterior margins, but narrowly interrupted behind eye, anterior portion continuous with lateral submarginal stria, which is complete and deeply impressed along sides; detached anterior submarginal stria transversely impressed behind head; pronotal disk with only single distinct pair of gland openings, located between anterior margin and free ends of transverse anterior submarginal stria; pronotal disk rather strongly convex, with few, small sparse secondary punctures near anterolateral corners; prescutellar impression absent. Elytra: elytral epipleuron with single epipleural stria present in basal half only, obsolete apically, outer subhumeral stria complete, inner subhumeral stria absent, dorsal striae 1 - 3 complete, rather crowded toward side, 4 th and 5 th striae absent, sutural stria thin, fragmented, obsolete in basal half; elytral disk with sparse secondary punctures in basal eighth. Prosternum: prosternal keel very shallowly emarginate at base, carinal striae complete, close together at middle, divergent anteriorly and posteriorly, free at base, united in narrow arch in front; short secondary striae present between carinal striae and procoxae; lateral prosternal striae weakly divergent; prosternal lobe very slightly deflexed, about two-thirds as long as keel, marginal stria present medially, divergent from edge, abbreviated at sides. Mesoventrite: anterior margin of mesoventrite broadly emarginate, but very weakly produced at center, marginal stria complete; mesometaventral stria coarsely crenulate, arched slightly forward onto basal third of mesoventrite. Metaventrite: postmesocoxal stria curved loosely behind coxa, extending nearly to mesepimeral-metepisternal corner; lateral metaventral stria curved laterad posteriorly toward posterior sixth of metepisternum; metaventral disk impunctate at middle, with few coarse secondary punctures at sides. Abdomen: 1 st abdominal ventrite with lateral striae obliquely impressed along inner edge of metacoxa; ventrites 2 - 4 with few coarse punctures only at extreme sides; propygidium rather long, about two-thirds as long as broad, more or less flat, with small gland openings near anterolateral corners, disk with few small, very irregularly sparse secondary punctures; pygidium elongate, about one third longer than basal width, slightly longer than propygidium along midline, apical margin rounded, disk weakly depressed along sides, punctation similar to but finer and sparser than that of propygidium. Legs: protrochanter glabrous, meso- and metatrochanters each with two very short apical setae; femora simple; protibia with outer margin rounded, weakly 5 - dentate, each tooth with moderately strong spine; posterior surface rather coarsely dimpled; protibial spurs very short; protarsus with ventral setae simple; meso- and metatibiae elongate, widened apically to about twice basal width, outer margins with 4 - 5 rather strong spines, mesotibia in particular faintly dentate; meso- and metatarsi short, each with single pair apicoventral spines. Male genitalia (Fig. 12): accessory sclerites present; T 8 rather short, with sides subparallel, apex very deeply, narrowly emarginate, basal membrane attachment line complete distad basal emargination, ventrolateral apodemes moderately developed, well separated beneath; S 8 halves approximate only at base, inner margins divergent to apex, apical guides widest near apex, with very fine inconspicuous setae; T 9 with broad basal apodemes, prominent, strongly hooked ventrolateral apodemes, dorsal lobes more strongly sclerotized along sides; T 10 weakly sclerotized, completely divided; S 9 with sides sinuate, subparallel, apical emargination distinct, rounded, apical flange interrupted, apical corners moderately prominent; tegmen widest near middle, slightly narrowed to base, more strongly narrowed to apex, with very large medioventral process; median lobe simple, about half tegmen length; basal piece long, about two-thirds tegmen length.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
9F7C2D82177CFB31E2612E0AB80BB9E3.taxon	etymology	Etymology. This species' name refers to its extreme similarity to the sympatric Pyxister devorator.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
2A278A900C2A1E2BE5EA08B92F17870A.taxon	description	Description. This genus differs from other Exosternini in the following combination of characters: body elongate, sides subparallel, distinctly convex; frons widened to rounded anterior corners produced over antennal bases; epistoma bituberculate along anterior margin; labrum with apical margin truncate, weakly produced and carinate; left mandible with straight, edentate incisor edge, right mandible curved to apex, with small, acute basal tooth; submentum rather short, slightly depressed, weakly produced anteriorly into oral cavity; mentum broadly subquadrate, weakly emarginate apically, labial palpi 3 - segmented, basal palpomere very short, penultimate and ultimate palpomeres rather thin and elongate; maxillary cardines very smooth, stipes faintly microsculptured, with few scattered setae, maxillary palpi 4 - segmented, basal palpomere short, 2 nd and 3 rd palpomeres subequal, ultimate palpomere about twice as long as penultimate, simply fusiform; antennal scape elongate, very slightly widened toward apex, with few setae on anterior surface; funicle slightly shorter than scape, widened slightly to cupuliform 8 th antennomere; antennal club elongate oval, about as long as funicle, tomentose, with median and subapical annuli crowded into apical half, both interrupted on dorsal surface, free ends enlarged into sensory patches, the apical-most more so; pronotal gland openings not evident; pronotal sides subparallel, abruptly bent to apical corners; transverse elevated carina present behind and parallel to anterior pronotal margin, curving posterad briefly at sides, abruptly bent to lateral margin behind anterior corner; prosternal keel overlapped at base by projecting mesoventrite, weakly convex, short, anteriorly displaced by prominent prosternal lobe, which is strongly produced, deflexed, with raised median ridge; mesoventrite subtrapezoidal, projecting anteriorly; mesometaventral stria absent; propygidium transverse, with small gland openings in anterolateral corners; pygidium about one third wider than long, apical margin simply rounded; protrochanter lacking setae, meso- and metatrochanters with pair of very short setae near apex of posterior edge; profemur rather narrow, anterior edge sinuate; protibia narrow at base, outer margin more or less rounded apically, weakly dentate, strongly spinose; protibial spurs present, short; protarsus somewhat laterally compressed, bearing spatulate ventral setae in both sexes; meso- and metafemora narrow; meso- and metatibiae slightly widened to apices, bearing few thin spines toward apex of outer margins; male genitalia with paired accessory sclerites present; T 8 with ventral apodemes nearly meeting along midline; S 8 with halves divergent apically, apical guides well developed, with subapical setae; T 9 with broad, blunt ventrolateral apodemes; T 10 completely divided; S 9 head broad, with prominent lateral flanges, apical flange low, continuous, not interrupted medially; tegmen lacking medioventral tooth or process; median lobe long, basal processes strongly differentiated; female T 8 forming a single plate; S 8 tripartite, basal baculi narrowly articulated with lateral plates, thin and convergent proximally; S 9 well developed, about twice as long as broad, articulated with strap-shaped extension from apex of S 8; T 10 entire; valvifers about as long as coxites, weakly paddle-shaped, weakly expanded in basal third; coxites nearly three times as long as wide, bidentate, with median apical tooth rather thin and elongate, outer tooth weakly developed, the two well separated; gonostyle flattened, shorter than median tooth, apically setose; bursa copulatrix membranous, barely expanded; spermatheca rather short, sausage-shaped, with basal stem short, with slightly thin, very elongate spermathecal gland attached near its base.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
2A278A900C2A1E2BE5EA08B92F17870A.taxon	etymology	Etymology. The name of this genus translates to ' shoulder ridge' referring to its most diagnostic feature.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
857EFCD03C2606F61F4994A667E91352.taxon	description	Description. Size range: Length 2.2 - 2.5 mm; width 1.9 - 2.1 mm; Body: body rufobrunneus, elongate, sides subparallel, distinctly convex. Head: frons nearly as long as broad, more or less flat, weakly depressed in middle, sides widened very slightly to rounded anterior corners produced over antennal bases; frontal stria fine, present close to sides and along anterolateral edges, absent from middle; supraorbital stria absent; frontal disk with fine but conspicuous ground punctation throughout, with few coarser punctures along dorsal margin, faintly microsculptured; epistoma bituberculate along anterior margin, tubercles subtended by weak carinae to ends of frontal stria, epistomal disk depressed along midline; labrum about one-third as long as broad, apical margin truncate, weakly produced and carinate above flattened supraoral surface. Pronotum: pronotal sides subparallel in basal two-thirds, abruptly bent to apical corners; pronotal gland openings not evident; marginal stria complete along lateral and anterior margins; sinuate, transverse elevated carina present one-fourth behind and parallel to anterior pronotal margin, curving posterad briefly at sides, abruptly bent to lateral margin one-third from anterior corner; ground punctation of disk fine posterad carina, markedly denser anterad carina, with few coarser, shallow punctures at sides; prescutellar impression absent. Elytra: convexity of elytra slightly greater than that of pronotum, i. e., lateral profile not a smooth curve; epipleuron smooth, with single, complete marginal stria; outer subhumeral stria carinate, forming distinct lateral margin to dorsal surface, inner subhumeral stria present only in basal half, striae 1 - 4 complete, 5 th stria obsolete in basal one-third, rarely complete, sutural stria obsolete in basal one-third; all dorsal striae rather shallowly impressed, but 4 th, 5 th and sutural striae broad, at least apically, delimited on inner and outer edges. Prosternum: prosternal keel overlapped at base by projecting mesoventrite, lacking carinal striae, weakly convex, short, anteriorly displaced by prominent prosternal lobe, which is strongly produced, deflexed, with raised median ridge, marginal stria absent. Mesoventrite: mesoventrite subtrapezoidal, narrowed, projecting anteriorly, lacking marginal stria; mesometaventral stria absent. Metaventrite: postcoxal stria present, curved around mesocoxa to middle of mesepimeron; lateral metaventral stria extending from inner corner of mesocoxa toward middle of metacoxa, bent laterad apically toward metepisternum; metaventral punctation fine and sparse at middle, with few coarser punctures at sides. Abdomen: 1 st abdominal ventrite with ground punctation rather dense, with parallel lateral striae along inner edge of metacoxa; ventrites 2 - 4 simply punctate, lacking transverse striae; propygidium wide, rather short, with small gland openings in anterolateral corners; pygidium about one third wider than long, apical margin simply rounded. Legs: protrochanter lacking setae, meso- and metatrochanters with pair of very short setae near apex of posterior edge; profemur rather narrow, anterior edge sinuate, partial stria along posterior margin; protibia narrow at base, outer margin widened to more or less rounded apical half, weakly dentate, each tooth bearing thin but rather long spine; protibial spurs present, short; protarsus somewhat laterally compressed, bearing spatulate ventral setae in both sexes; meso- and metafemora narrow, slightly elongate; meso- and metatibiae slightly widened to apices, bearing few thin spines toward apex of outer margins. Male genitalia (Fig. 15): Paired accessory sclerites present; T 8 with broad basal and narrower apical emarginations, line of basal membrane attachment complete, distad basal emargination, ventral apodemes well developed, nearly meeting along midline; S 8 with halves nearly meeting only at base, divergent apically, apical guides well developed from base to near apex, rather abruptly narrowed to subacute apex, each side with single inconspicuous subapical seta; T 9 with broad, blunt ventrolateral apodemes, apices narrow, obliquely subtruncate; T 10 completely divided; S 9 broad, truncate at base, narrowed toward apex, head broad, with prominent lateral flanges, apical flange low, continuous, not interrupted medially; tegmen rather narrow, widest at middle, evenly narrowed to base and apex, weakly curved in lateral aspect, lacking medioventral tooth or process; basal piece about one-third tegmen length; median lobe almost half tegmen length, gonopore rather wide, basal processes strongly differentiated, with thin proximal arms over half overall length.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
857EFCD03C2606F61F4994A667E91352.taxon	etymology	Etymology. The name of this species refers to pyramidal swellings on the apex of the epistoma.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
51689999DFE5C9309EF24A96B31E7F9A.taxon	description	Description. Size range: Length 1.8 - 2.2 mm; width 1.5 - 1.8 mm; Body: body depressed, sublimuloid, with sides rounded and pygidia variably prolonged, generally lightly colored, rufescent to rufo-brunneus, smooth, with fine ground punctation but very little secondary punctation. Head: frons and epistoma convex, prominent, frons rather broad with frontal stria complete, weakly recurved at middle; supraorbital stria fine, usually detached at sides; epistoma weakly emarginate apically; labrum short inwardly arcuate, about 4 x wider than long; mandibles rather short, basal denticles on incisor very small to obsolete; submentum transversely depressed, posterolateral margins raised, apical margin produced slightly into base of oral cavity; mentum about twice as wide as midline length, sides narrowed, apex weakly emarginate; maxillary cardo glabrous, stipes with two setae along lateral margin, basal palpomere short, 2 nd and 3 rd palopmeres short, subequal, ultimate palpomere narrowed apically, about 1.5 x as long as penultimate; antennal scape weakly expanded to apex, weakly carinate along inner margin, funicle narrow at base, weakly widened to 7 th and disc-like 8 th antennomere; antennal club tomentose, basal annulus obsolete, middle annulus with slightly basally expanded sensory patch on upper surface, simple on lower surface, apical annulus poorly defined, transverse. Pronotum: pronotal sides arcuate, strongly convergent to anterior corners; prescutellar impression absent; median pronotal gland openings very fine, one pair along anterior margin laterad eye, one pair behind eye about two-thirds pronotal length from anterior margin; marginal pronotal stria complete and continuous along lateral and anterior margins; submarginal stria complete laterally, not extending mediad behind head, very close to marginal stria, intervening disk narrowly convex. Elytra: elytral striation strongly reduced, single epipleural stria present, complete, diverging from margin in anterior half, outer subhumeral stria complete, nearly meeting 1 st dorsal stria apically, inner subhumeral stria absent, oblique humeral stria faint, 1 st dorsal stria more or less complete, may be slightly abbreviated basally, may be extended mediad along posterior margin of elytron, 2 nd and third dorsal striae weakly impressed, present in basal half or less, 4 th and 5 th striae completely absent, sutural stria usually represented only by extremely short striole at posteromedian corner of elytron, may be extended laterad by apical marginal stria. Prosternum: prosternal keel narrow, acutely emarginate at base, carinal striae weak to absent; prosternal lobe about half as long as keel, marginal stria present or absent. Mesoventrite: mesoventrite acutely produced in front, marginal stria complete, with varied fine strioles in anterolateral corners; mesometaventral stria absent. Metaventrite: mesoventrite with postmesocoxal stria present, varied in length, lateral metaventral stria absent; median portion of metaventral disk with fine ground punctation only, grading to coarser punctures laterad coxae, punctures along metaventral-metepisternal suture may coalesce into stria; metepisternum often with longitudinal stria. Abdomen: 1 st abdominal ventrite with single faint to abbreviated stria along inner margin of metacoxa, generally curved laterad behind coxa, disk faintly strigose at sides; ventrites 2 - 4 with posterior marginal stria along lateral thirds or more; ventrite 5 variously prolonged, apical margin strongly arcuate; propygidium flat or faintly depressed at sides, with basal marginal stria, complete or not; propygidial disk without obvious gland openings; pygidium with apex subacute to very prolonged and acuminate, with lateral marginal striae or not. Legs: all femora flattened and slightly expanded, arcuate on anterior and posterior edges; each trochanter with single seta; protibia with inner and especially outer edges arcuate, narrowing apically, the outer edge bearing 6 - 7 strong spines, denser near apex, lacking emarginations between, two apical protibial spurs present, strongly reduced, anterior surface of protibia with tarsal groove almost obsolete; protarsus of both sexes bearing spatulate ventral setae; meso- and metatibiae very narrow, parallel-sided, bearing a few thin spines toward apex of inner and outer edges; meso- and metatarsi as long or longer than corresponding tibia, with long, ventral setae that may be vaguely spatulate. Male genitalia: accessory sclerites present, basal; T 8 with weakly developed ventrolateral apodemes, apical margin may be slightly desclerotized, basal membrane attachment line intersecting basal emargination; S 8 divided, inner edges divergent in apical half, lateral guides weakly to moderately developed, apices narrowed, bearing a few conspicuous setae near apical corners; T 9 with ventrolateral apodemes only very weakly developed, not hooked, apices narrowed, acute at inner corners; T 10 weakly sclerotized, completely divided; S 9 broad, sclerotized along edges, with small apical emargination and weak apical flanges; tegmen flattened, moderately broad basally, slightly narrowed apically, lacking ventromedial process; median lobe more than half as long as tegmen, with proximal apodemes prominent, abruptly narrowed at extreme proximal end; basal piece long, about half as long as tegmen, with prominent apicoventral point. Female genitalia: T 8 forming a single plate, apically desclerotized, with shallow, arcuate basal emargination; S 8 tripartite, with median sclerite weakly divided from lateral sclerties, basal baculi narrowly attached to lateral sclerites, evenly convergent proximally; S 9 weakly sclerotized, elongate, articulated with strap-shaped extension from apex of S 8; T 10 broad, apically arcuate; valvifers paddle-shaped, paddles nearly one-half total length; coxites elongate, two-thirds length of valvifers, tridentate, with very prominent median tooth dwarfing teeth on either side; gonostyle long, bisetose, inserted between two lateral-most apical teeth; bursa copulatrix membranous, weakly expanded; spermatheca weakly sclerotized, approximately spherical, borne on long thin stalk inserted at base of common oviduct, with elongate, weakly spiraled spermathecal gland attached near its base.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
51689999DFE5C9309EF24A96B31E7F9A.taxon	diagnosis	Diagnosis. This genus is easy to recognize based on its sublimuloid shape (Fig. 16), with the body depressed, the sides rounded, and the pygidium variably prolonged and subacute. Its convex frons is also unusual, as are the rounded, spinose protibiae, the very narrow meso- and metatibiae, the reduced elytral striation, and the almost complete lack of secondary punctation. Lacrimorpha is resolved as the sister group of the genus Mecistostethus in our recent analysis of Exosternini relationships (Caterino and Tishechkin in review).	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
51689999DFE5C9309EF24A96B31E7F9A.taxon	etymology	Etymology. The name of this genus means ' tear-drop shaped', resulting from its tapered posterior end. The genus is feminine.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
642A8BD5B46EE9E9C18A6954878D2B67.taxon	etymology	Etymology. This species' name refers to its very smooth, glabrous body surface.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
B4EDF6F1125011AC4CD16DA705B86D54.taxon	etymology	Etymology. The name of this species refers to its moderately subdepressed body form.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
70D18172CB550379AAD281D35F069F25.taxon	etymology	Etymology. This species is named for its type locality, close to the dam of this name northeast of Manaus.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
ADF65F42D25060CA90E9B18C5A862157.taxon	etymology	Etymology. This species' name refers to the acuminate pygidium.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
8138632621224056E283BAF411FC0DB4.taxon	description	Description. Size range: Length 1.7 - 3.2 mm; width 1.5 - 2.8 mm; Body: body rather broadly ovoid, variably subdepressed to rather strongly depressed, rufescent to rufopiceous. Head: frons broad, frontal corners rounded, rather prominent over antennal bases; frontal disk depressed in common with epistoma, frontal stria recurved dorsad within depression, usually complete; sides of epistoma variably ridged, carinate and / or striate; labrum broad, apical margin generally carinate, truncate to weakly emarginate; mandibles with small or no basal teeth on incisor edge; submentum flat to slightly depressed, sparsely setose; mentum about half as long as wide, arcuately narrowed anteriorly, apical margin acutely emarginate; cardo smooth and glabrous, stipes with few setae along lateral margin; ultimate palpomeres fusiform; antennal scape elongate, curved, weakly carinate along anterior margin; funicle weakly widened beyond 5 th antennomere, 8 th antennomere slightly shorter than preceding; antennal club elongate, widest just beyond midpoint, tomentose, with interrupted basal annulus near midpoint and complete annulus between midpoint and apex, slightly enlarged basad at middle, particularly on dorsal surface. Pronotum: pronotal sides weakly arcuate, convergent to anterior corners, slightly to distinctly explanate at sides, marginal stria usually complete along lateral and apical margins, submarginal stria present very close to sides, absent across front, weakly crenulate; pronotum with pair of gland openings very close to anterior margin behind eye and glands variably displaced posterad onto pronotal disk, usually multiplied along a visible track bearing up to 5 distinct openings along its length; pronotal disk with secondary punctures, when present, strongly concentrated across basal half. Elytra: elytral disk weakly to moderately convex, all striae coarsely impressed, each stria comprising two alternating series of interconnected punctures, appearing chain-like at their most dense; epipleuron usually with one complete marginal stria, with fragments of a second in posterior half or not, inner subhumeral stria usually complete, dorsal striae 1 - 4 complete, 5 th and sutural striae rarely obsolete basally, bases of 4 th or 5 th and sutural striae rarely connected by weak basal arch; ground punctation of elytral disk fine to coarse, with at least a few coarse secondary punctures usually present in most interstriae. Prosternum: prosternal keel generally distinctly, subacutely emarginate at base, with complete carinal striae usually united in narrow anterior arch, short secondary lateral strioles frequently present between carinal striae and procoxae; lateral prosternal striae present, divergent in front of coxae; prosternal lobe one-half to two-thirds length of keel, apically rounded to subtruncate, with marginal stria present at middle, variably obsolete at sides. Mesoventrite: mesoventrite subacutely produced at middle, with complete marginal stria smooth to crenulate; mesometaventral stria crenulate, usually strongly arched to angulate anterad onto middle of mesoventral disk, disk frequently with sparse secondary punctures. Metaventrite: metaventral disk with coarse secondary punctures usually over most of surface, postmesocoxal stria present, recurved anterad to mesepimeron or ending freely posterolaterad coxa, lateral metaventral stria sinuate, extending from inner corner of mesocoxa toward middle of metacoxa, frequently abbreviated apically; coarse punctures of metepisternum may coalesce into vague to distinct striae. Abdomen: ventrites mostly coarsely punctate, punctures tending to coalesce into striae along apical margins of ventrites 2 - 4; 1 st ventrite with poorly developed lateral stria along inner edge of metacoxa, abbreviated or curving laterad behind coxa; propygidium transverse, 2 - 3 x as wide as long, with single gland opening on each side, often borne on weak convexity behind anterolateral corner, sometimes associated with weak oblique striole, disk generally sparsely covered with shallow secondary punctures, punctures often sparser in apical half; pygidium more or less equilateral, apex rounded to weakly subacute, gland openings generally present near lateral margin one-fifth to one-fourth from base, these often associated with lateral marginal pygidial striae; marginal stria when present rarely complete around apex; pygidial disk variably punctate. Legs: all trochanters bearing single long seta; profemur rather dilated at middle, narrowed, slightly emarginate at inner apex, protibia generally rounded, strongly spinose, tibial margin only rarely emarginate between to form marginal teeth; protibial spurs present but generally weak; protarsus of both sexes with strongly spatulate setae; meso- and metafemora weakly dilated basad middle, with complete posterior marginal stria; meso- and metatibiae similar, weakly widened to apex, with series of long, rather thin marginal spines; meso- and metatarsi with 4 ventral spines along apical margins only. Male genitalia: accessory sclerites vestigial or absent; T 8 with sides evenly tapered to subtruncate apex, basal membrane attachment line generally tangential to deep, rounded basal emargination, ventrolateral apodemes produced most strongly at base, narrowed apically, separated by about one-third T 8 width beneath; S 8 short along midline, with longer divergent lateral guides bearing several strong setae along apicoventral margin, with weak membraneous velum across entire apex; T 9 with ventrolateral apodemes not strongly tapered, inner apices subquadrate beneath, T 9 apices subacute to obliquely subtruncate; T 10 entire, may be emarginate basally and / or apically; S 9 rather short, with broad truncate to emarginate base, head variably subquadrate, apically emarginate, with apical flanges separate; aedeagus rather broad, flattened, sides rounded, apical division conspicuous, apices often distinctly separate, medioventral process strong, produced beneath near midpoint; basal piece one-fourth to one-third tegmen length, with distinct medioventral tooth; median lobe short, proximal arms strongly narrowed basally. Female genitalia: T 8 forming single broad plate; S 8 forming separate median and lateral plates, basal baculi thin, narrowly attached to lateral sclerites, convergent proximally; S 9 slightly elongate, articulated with strap-shaped extension from apex of S 8; T 10 weakly sclerotized; valvifers paddle-shaped, paddles rather short, about one-third valvifer length; coxites 2.5 - 3 x as long as wide, tridentate, with weak inner tooth, prominent median tooth, and intermediate lateral tooth; gonostyle nearly as long as median tooth, setose; bursa copulatrix membranous, lacking sclerites, weakly expanded; spermatheca weakly sclerotized, globose, borne on long thin stalk inserted at base of common oviduct, with elongate, spiraled spermathecal gland attached near its midpoint.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
8138632621224056E283BAF411FC0DB4.taxon	diagnosis	Diagnosis. In most respects this genus is highly distinctive. However, many of the distinguishing characters vary considerably within the group, in addition to appearing in unrelated species, and it is only through a combination of external characters and male genitalia can an unambiguous diagnosis be provided. For most species, the pattern of pronotal punctation, with coarse punctures restricted to the basal half of the pronotal disk (Figs 18 A, 19 A, C, 22 A, 23 A), is distinctive, and this in combination with coarsely impressed elytral striae (Fig. 18 C) and sparse secondary punctures in the elytral interstriae will separate most species. The longitudinal tracks of multiplied pronotal gland openings (Fig. 19 G) are also very unusual, and will put most species here easily. If the above are true, and the protarsi have spatulate setae, the diagnosis is unambiguous. Finally, the male genitalia of Crenulister also exhibit several distinctive characteristics, in particular the rather broad, flat aedeagus, with strong, acute ventromedial process (see Fig. 20 E), frequently with the tegmen apices separated, the few strong setae at the apex of the 8 th sternite (Fig. 20 B), and the broad, medially subquadrate ventrolateral apodeme of the 9 th tergite (Fig. 20 C). A single (undescribed) species in a distantly related group shares all of these external characters, and represents an amazingly similar overall body form. The only good characters to distinguish it are its lack of spatulate protarsal setae, and its entirely different male genitalia, with its aedeagus strongly narrowed and hooked apically, completely unlike the short, broad and flattened aedeagus of Crenulister. Phylogenetically, Crenulister emerges from within a diverse group of mostly undescribed taxa that we loosely term the ' scutellar impression group', particularly a small subgroup related to the named species Phelister blairi Hinton.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
8138632621224056E283BAF411FC0DB4.taxon	etymology	Etymology. The genus name refers to the ubiquitously crenulate elytral striae found in the species of this genus, in conjunction with the common histerid ending - ister. The gender of the genus is masculine.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
742FB4FC834050F9814C459F83137B28.taxon	etymology	Etymology. This species name refers to its relatively large size, and secondarily its occurrence in Mato Grosso, Brazil.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
5D4EF0BDE40475BCCBD32C0D86ED2326.taxon	etymology	Etymology. The name of this species refers to its conspicuously spinose protibiae.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
9892F14ACD72C8B5759BFFAFD7D1865A.taxon	etymology	Etymology. This species' name refers to its particularly conspicuous series of pronotal gland openings, against a largely impunctate background.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
D6EB85CFA0A4D2D8D8C7D236EEF74486.taxon	etymology	Etymology. The name of this species refers to the relative paucity of punctures in the elytral interstriae.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
EAE2B736FCCC3926AB662BF859D5233B.taxon	etymology	Etymology. This species' name refers to its relatively dark coloration.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
7142000EE7C013EBA524E01B068B833E.taxon	etymology	Etymology. This species' name refers to the ' unequal' distribution of pronotal punctures, distinctly increasing in density posterad.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
FBDEB3124781AFF85769806A04CCEEDB.taxon	etymology	Etymology. This species' name refers to the slightly explanate lateral pronotal margins (Fig. 23 B).	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
4087D4B5A15C65261F4D6589AD11D87A.taxon	etymology	Etymology. This species is named for its distinctively dentate protibiae.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
85C305993A732B49EC16CB36BEFC5926.taxon	etymology	Etymology. This species' name refers to its relatively impunctate, simple external sculpturing.	en	Caterino, Michael S., Tishechkin, Alexey K. (2014): New genera and species of Neotropical Exosternini (Coleoptera, Histeridae). ZooKeys 381: 11-78, DOI: http://dx.doi.org/10.3897/zookeys.381.6772, URL: http://dx.doi.org/10.3897/zookeys.381.6772
